Gilbert Klapper

Northwestern University | NU · Department of Earth and Planetary Sciences

This paper establishes the ranges of late Givetian and Frasnian phacopid trilobites with reference to the Frasnian conodont zonation. The ranges integrate both hitherto formally known and numerous new taxa that were recovered from outer shelf limestones primarily in the Montagne Noire (southern France) and incidentally in related areas; i.e. Thurin...

The conodont species Polygnathus tuberculatus Hinde is best known in condensed conodont-bone bed lag deposits in western New York and southern Indiana. These occurrences, containing conodonts ranging from zones of the Middle GivetianMiddle varcus (= ansatus) Zone to Lower Frasnian Zone 2, have influenced a postulated range extension above the varcu...

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Klapper, G. and Vodražkova, S. 2013. Ontogenetic and intraspecific variation in the late Emsian - Eifelian (Devonian) conodonts Polygnathus serotinus and P. bultyncki in the Prague Basin (Czech Republic) and Nevada (western U.S.). Acta Geologica Polonica, 63 (2), 153-174, Warszawa. Samples from populations of Polygnathus serotinus Telford 1975 and...

The Devonian phacopid trilobite Plagiolaria kitabi Crônier & Tsmeyrek was described from a part of the Akbasai Formation in Uzbekistan that had been considered tentatively to be of late Famennian age based on the misidentification of Polygnathus inornatus Branson, a late Famennian? – Tournaisian conodont species. The revised identification herein o...

Conodont faunas from the uppermost part of the Coils Creek Member of the McColley Canyon Formation and primarily the lower part of the Denay Limestone (Middle Devonian, Eifelian) from the northern Roberts Mountains and adjacent central Nevada areas are the subjects of the present study. Four new species from the Denay Limestone (Eifelian, costatus...

The outline of the platform in Pa elements of conodont genera such as Palmatolepis is commonly one of the most heavily weighted taxonomic characters. But a taxonomy based only on visual discrimination of Pa elements of Frasnian species of Palmatolepis has resulted in widespread misidentification. Although their Pa elements can be confused, Palmatol...

The Ogilvie Formation, formerly dated as Eifelian and Givetian, is locally confined to the Early Devonian, Emsian. Regionally the top of the Ogilvie is markedly diachronous ranging in age from Emsian to Givetian. The Ogilvie Formation conformably overlies the Late Silurian to Early Devonian Gossage and/or early Emsian Michelle and Cranswick Formati...

The lower member of the Henryhouse Formation (Ludlow-Přídolí) is a thin (<14 m) succession of extremely argillaceous carbonate mudstones and shales that are restricted to the central Arbuckle Mountain region in southern Oklahoma. Conodont faunas of the Kockelella crassa, Ancoradella ploeckensis and Polygnathoides siluricus zones indicate that the l...

The boundary-stratotype for the base of the Famennian Stage at Coumiac in the Montagne Noire, southern France, was defined so as to coincide with the lower boundary of the Lower triangularis conodont zone. The definition of that biostratigraphic zone has to be modified in light of the present taxonomic distinction between Palmatolepis triangularis...

George Sanderson (with George Verville, background) collecting fusulines at the type Derryan (Pennsylvanian) section in Sierra County, New Mexico. Photo by G. P. Wahlman, 1983. George Albert Sanderson, Jr. was born August 22, 1926 in New Haven, Connecticut. He was raised by his mother and grandparents, who emigrated from Germany at the beginning...

A principal point of disagreement in conodont ecology between the pelagic depth stratification model and the benthic model of lateral segregation is the mode of life of the conodont animal. A study of the literature on certain Recent benthic and planktic organisms that may serve as ecologic analogues (chaetognaths, isopods, foraminifers) indicates...

P harciceras Hyatt, 1884 is the diagnostic ammonoid of the late middle Givetian Stage of the Middle Devonian Series. It occurs in the Rhenish Massif in Germany, the Montagne Noire in southern France, and in equivalent strata in the Anti-Atlas in southern Morocco. Verified North American occurrences of Pharciceras are confined to the New York succes...

Recognition of four types of skeletal apparatuses of Devonian compound and platform conodonts, which conform in their basic plan to those already established in the Ordovician, Silurian, and Carboniferous, emphasizes the persistence of but a few structural types in the Paleozoic. The vicarious nature primarily of compound but also of platform eleme...

The Horse Spring conodont succession in the Canning Basin replicates zones 6 through 13 of the thirteen-fold Frasnian zonation. Horse Spring, together with other Canning Basin sections, demonstrates the widespread application of the zonation. Furthermore. the recently formalized three-fold subdivision of Zone 13, which has been recognized in the Mo...

The terminal zone of the Frasnian standard conodont zonation, the linguiformis Zone has been subdivided into alower part defined by the total range of Palmatolepis linguiformis and an upper part defined by the absence of the nominal species together with the rare occurrences of Pa. ultima (= Pa. praetriangularis) and Ancyrognathus (= Ancyroides) ub...

The upper part of cores of the Onakawana B Drillhole in the Moose River Basin in northern Ontario includes the upper part of the upper member of the Williams Island Formation (22.5 m, 16 samples), and the entire overlying Long Rapids Formation (75.1 m, 49 samples). The sequence of conodonts from the drillhole was analyzed by graphic correlation as...

CONODONT BASED TRILOBITE BIOZONATION IN THE LATE DEVONIAN OFF-SHORE REALM FEIST, Raimund1, KLAPPER, Gilbert2, and LEROSEY-AUBRIL, Rudy1, (1) Paleontology, paleobiology and phylogeny, Institut des Sciences de l'Evolution de Montpellier (UMR5554), University Montpellier II, Place E. Bataillon, Cc062, Montpellier, 34095, France, rfeist@isem.univ-montp...

The development of the Frasnian (Upper Devonian) reef complexes of the southern Timan and Pechora region of northern European Russia is described. Barrier reef complexes progressively prograded eastwards during the Frasnian but the carbonate complexes were interrupted many times by regressive events. Using new conodont and ammonoid biostratigraphic...

The level of the Frasnian/Famennian boundary was accepted by the International Subcommission on Devonian Stratigraphy in 1991 to coincide with the lower boundary of the Lower Palmatolepis triangularis conodont Zone. The recommended GSSP for the stage boundary was accepted by IUGS in 1993. The boundary is drawn at the base of Bed 32a in an Upper Dev...

Ancyrodella rotundiloba is the defining species used to position the GSSP for the mutual bases of the Frasnian Stage and the Upper Devonian Series at Col du Puech de la Suque section E in the Montagne Noire. The taxonomic interpretation of this species advocated herein is centered around the primary types and coincides with that used in papers by i...

Previously it has not been possible to correlate the thirteen-fold Frasnian conodont zonation first developed in the Montagne Noire, southern France, with the nine-fold standard zonation of the Frasnian. The impasse has been due to the fact that the taxonomies underlying the two zonations are so disparate that comparisons of faunal lists and range...

A total of 13 conodont zones and 11 rugose coral faunal assemblages currently provides a subdivision of the Frasnian of the Western Canada Sedimentary Basin. The conodont zonation adopted is that first developed in the Montagne Noire area of southern France and subsequently found to have widespread applicability, especially when supplemented throug...

Through graphic correlation (Shaw's method) of Frasnian sections in the Montagne Noire, southwestern France, with those in the Alberta Rockies and Hay-Trout Rivers area of western Canada, the zonal and/or informal conodont faunal sequences developed for the three areas can be aligned with some precision for the first time. All of the zones are scal...

The physical boundaries of global Devonian transgressive-regressive (T-R) cycles are located for the first time at reference sections in central Nevada. Early Devonian (Pragian and Emsian), deepening events defining T-R cycle bases resulted in deeper subtidal facies overlying peritidal or shallow subtidal facies. Middle and earliest Late Devonian d...

A Frasnian composite standard provides a refined scaling for the thirteen-fold conodont zonation first developed in the Montagne Noire, France, but since replicated in North America, Australia, and now the Timan-Pechora region of Russia. Zones 4–13 are identifiable in seven cores from the Ukhta area of southern Timan and a core from the Bagan Field...

Although conodont endemism was not as high in the Frasnian as in earlier Devonian intervals, analysis of a limited number of intensively collected sections in the Montagne Noire, the United States and Canada, Western Australia, and the Russian Platform demonstrates levels of endemism ranging from about 30 to 35% in three composites of two zones eac...

The line of correlation (LOC) in the graphic correlation of the 27 sections in a Frasnian composite standard conforms empirically to the pattern of biostratigraphic events, the bases and tops of ranges of conodont species. The guiding principle is to position the LOC so as to avoid range overlaps of well understood species that have not been observ...

ABSTRACT-Multielemetnatx onomy and shape analysish ave led to substantialr evisions in the taxonomic concepts of 13 Frasnian species of Palmatolepis, which previously have been based largely on visual discrimination of the Pa elements. Septimembrate apparatuses have been recognized in three species and diagnostic Pb elements are associated with the...

Detailed qualitative analysis of the Pa element and multielement analysis emphasizing the associated Pb element have led to the present revision of the Frasnian species of Ancyrognathus. Diagnostic characters of the Pa element, such as the platform outline, carinal pattern, and shape of the pit, as well as salient features of the Pb element, are th...

The F‐F extinction is known on four continents by the sudden disappearance of shelly benthos, although questions remain regarding precisely when and therefore how suddenly it took place. Studies show that this extinction occurred at the gigas/triangularis zone boundary in Utah and Nevada. U.S.A., Hony and Sinsin sections. Belgium. Montagne Noire. F...

Details are given of the succession of conodonts, goniatites and trilobites across the Frasnian-Famennian boundary, documenting extinction and appearance levels of the Kellwasser Events at Coumiac, north of Cessenon (Herault). The sequence may be considered as a potentially global stratotype boundary section for the definition of this boundary. -Au...

The Middle-Upper Devonian Series Global Stratotype Section and Point (GSSP), which defines the basal boundary of the Upper Devonian Series and the Frasnian Stage, has now been agreed upon. It is exposed at Col du Puech de la Suque section in the southeastern Montagne Noire, France. Full details are given in a following contribution. -from Authors

A mainly Late Devonian (late Givetian to Early Carboniferous) major transgressive-regressive cycle, or depophase, is represented in the sedimentary succession of W Canada, W US, New York, Belgium and Germany. This major cycle comprises 6 smaller cycles characterized by initial abrupt deepening events followed by upward shallowing. The major and sma...

The Devonian System of Euramerica contains at least 14 transgressive-regressive (T-R) cycles of eustatic origin. These are separated into three groups (or depophases) and from Carboniferous cycles by three prominent regressions. Twelve post-Lochkovian T-R cycles are recognized, and they commonly appear to result from abrupt deepening events followe...

The Polygnathus biofacies characterizes the Frasnian sequence in the Hay River, Trout River, and adjacent areas ofs the southwestern Northwest Territories. The sequence spans almost the entire Frasnian. Only at certain levels are there minor incursions of species of Palmatolepis, Ancyrodella, and Ancyrognathus that elsewhere characterize the Palmat...

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Conodonts are zoologically enigmatic, toothlike phosphatic microfossils occurring in marine sedimentary rocks ranging in age from Cambrian to Triassic. Dimpled spheres of less than 1 millimeter in diameter are sporadic associates of conodonts and have identical chemical composition and microstructure. Mineralogy, morphology, and occurrence of these...

Note an error on p. 301, first complete paragraph, last line: The reference should not be to Huddle (1934) but should read 'Huddle (this paper).' This was an error introduced by an editor after we saw the proofs.

A regressive episode near the end of the Early Devonian caused a nearshore quartzose carbonate facies to shift to the edge of the carbonate shelf in central Nevada. Transgression followed in the latest Early Devonian and continued in the early Middle Devonian. Seaward of the shelf-wide unconformity produced by regression, a dolomitized, dark-gray l...

The writers note a convergence of opinion on a world-wide scale favoring placement of the Siluro-Devonian boundary at the base of the Gedinnian. This boundary is coincident with the base of the Monograptus uniformis and Icriodus woschmidti zones. It is recognizable in the brachiopod-coraltrilobite succession by the disappearance of pentamerids, Atr...

Conodont fauna, ages, and distribution of a new formal stratigraphic unit that previously was called the dark shale unit

44 p., 6 pl., 2 fig. http://paleo.ku.edu/contributions.html