FIGURE 10-- - uploaded by Gilbert Klapper
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Distribution and number of conodont species in the Twin Falls Formation (HR26-LEF3), uppermost Fort Simpson Shale (JM1, JM2), Jean-Marie Member (JM3, PR1, BIR1, TR1) and upper member (TR2, RRI-4) of the Redknife Formation, and Kakisa Formation (TR3, TR4-7, BR2, BR3, RR1-6 and above). Lithostratigraphy shown in Figure 8. Approximate correlations with the standard zonation are as follows. The sample with Palmatolepis semichatovae and Polygnathu unicornis (HR26) is interpreted as Lower gigas Zone. The long interval beginning with the lowest occurrence of Polygnathus ettremae (JM 1) may be correlated with the Upper gigas Zone (see discussion in text). Generic abbreviations as in Figure 9.
Source publication
The Polygnathus biofacies characterizes the Frasnian sequence in the Hay River, Trout River, and adjacent areas ofs the southwestern Northwest Territories. The sequence spans almost the entire Frasnian. Only at certain levels are there minor incursions of species of Palmatolepis, Ancyrodella, and Ancyrognathus that elsewhere characterize the Palmat...
Contexts in source publication
Context 1
... next stratigraphic interval, which ex- tends from the Hay River Formation up into the Twin Falls Formation, from the first oc- currence of Polygnathus n. sp. F (Figure 9) up to that of P. evidens (Figure 10), includes Palmatolepis subrecta in HR22-1. This species, according to Ziegler (1971, chart 5) ranges from the Middle asymmetricus Zone to the Middle Palmatolepis triangularis Zone, and thus is of no help in refined correlation. ...
Context 2
... succeeding interval, from the first oc- currence of Polygnathus evidens up to that of P. planarius (Figure 10), and thus within the Twin Falls Formation, has Palmatolepis semichatovae (HR26-1) and Polygnathus unicornis (HR26-1, HR20-1). P. semicha- tovae occurs in the upper Perdrix and lower Mount Hawk formations in the Nikanassin Range, western Alberta, in a part of the se- quence interpreted as Lower gigas Zone (Lane and Klapper, in prep.). ...
Context 3
... next stratigraphic interval, from the first occurrence of Polygnathus planarius up to the first occurrence of P. imparilis and/or P. brevicarina, is in the upper Fort Simpson Shale, the Jean-Marie Member and lower part of the upper member of the Redknife For- mation ( Figure 10). Marking this interval, in addition to P. planarius, is the first occur- rence of P. samueli and P. ettremae. ...
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... highest Frasnian interval of the pres- ent study is marked at the base by the first occurrence of either Polygnathus imparilis or P. brevicarina ( Figure 10) and occurs in the uppermost beds of the upper member of the Redknife Formation and throughout the pro- ductive part of the Kakisa Formation. Nei- ther new species of Polygnathus has been il- lustrated previously. ...
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... ther new species of Polygnathus has been il- lustrated previously. Palmatolepis unicornis, P. gigas, and P. subrecta occur in several samples within the TR3 section ( Figure 10); in combination with Polygnathus ettremae the association is suggestive of the Upper gi- gas Zone. ...
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... number of indeterminate coniform elements occur at one locality of this study (JM3, Figure 10). Although some of the specimens (e.g., Figure 11.4) superfi- cially resemble such Ordovician genera as Multioistodus Cullison, all of the specimens under study differ from chirognathacean con- odonts in having well-developed white mat- ter throughout the cusp. ...
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... 1949. PELEKYSGNATHUS sp. Material.-Pelekysgnathus sp. occurs in the Famennian Trout River Formation (Table 3) Remarks. --Inasmuch as the apparatus of Mehlina gradata, as reconstructed by Uyeno (in Norris, Uyeno and McCabe, 1982, p. 7677), is closely similar to that of Pandorinel- lina insita (Stauffer) (Klapper in Clark et al., 1981, p. W166, fig. 110, 1), Mehlina should be included within the ...
Context 13
... platform of Polygna- thus samueli is very broad and represents three-fourths of the unit length; the anterior and that in Figure 10 from Remarks.-There is a similarity between Polygnathus samueli and P. brevis Miller and Youngquist (1947) in the character of the blade and the short, wide platform. In P. brevis (holotype reillustrated in Klapper, 1973, p. 341, Polygnathus Pl. 1, fig. 2 and topotypes herein on Figure 17.1, 17.6, from the type Sweetland Creek Shale), the adcarinal troughs are wide and extend about half to two-thirds the platform length; thus there is no tendency towards convergence of the troughs as in the new species. ...
Context 14
... differ in details of platform ornamen- tation and the tendency to develop a con- spicuous sinus in the outer posterior margin. Two specimens similar to those of Szulczew- ski are present in the Trout River collections (TR1-4; Figure 10) and are designated as P. cf. P. samueli (Figure 17.11). ...
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Citations
... In sample A8, the frequency of Palmatolepis and (1) Polygnathus aequalis Klapper and Lane, 1985 Thomas, 1949; Lateral view of IUMC 124, sample A10, X 140; (9) Ancyrognathus sinelaminus (Branson and Mehl, 1934); Upper view of IUMC 100, sample A10, X 127; (10) Icriodus alternathus alternathus Branson and. Mehl, 1934 Polygnathus genera is equal and have Palmatolepidpolygnathid biofacies (related to the outer and middle ramp), and in the sample A9, the frequency of Polygnathus is about 85%, while Palmatolepis frequency is about 15%, representing Polygnathid biofacies. ...
The Late Devonian‐Early Carboniferous deposits of the Anarak section in northeastern Isfahan, Central Iran, evaluated based on conodont biostratigraphy, sedimentary environment and sequence stratigraphy. According to the field observations, five lithological units were identified. Investigating the conodont fauna of the Late Devonian‐Carboniferous (Mississippian‐Pennsylvanian) deposits of Bahram, Shishtu, and Qaleh (Sardar 1) formations in Anarak section led to the identification of 67 species of 18 conodont genera, and accordingly 22 conodont biozones were differentiated. The correlation of sea‐level change curves, regarding to the conodont biofacies with the global sea‐level curve, demonstrates the relative correlation in the mentioned times due to the shallow condition of the central Iran basin compared to the European and American basins. The microfacies analysis led to the identification of 12 microfacies related to the open sea, bioclastic barrier, lagoon and tidal flat sub‐sedimentary environments in a homoclinal carbonate ramp environment. Based on sequence stratigraphy studies, three 3 rd order sequences were identified. The first sequence, which is of the Late Devonian (upper part of the Bahram Formation, 32.5 m), the second sequence (12.5 m) is the Late Devonian (uppermost part of the Bahram Formation), and the third sequence (68 m) is the Early Carboniferous (the Shishtu I Formation).
... Many studies have revealed that the distribution of conodont elements is controlled by various environmental factors (e.g. depth, temperature, salinity), leading to diverse biofacies in different paleotectonic settings (Seddon and Sweet 1971;Sandberg 1976;Sandberg and Ziegler 1979;Dreesen and Thorez 1980;Sandberg and Dreesen 1984;Klapper and Lane 1985;Matyja 1987;Sandberg et al. 1988;Zhen and Percival 2003). For the Upper Devonian, numerous investigations of pelagic conodont biostratigraphy were carried out and several pelagic standard conodont zonation schemes were proposed, which are established based on the taxa of globally distributed deep-water biofacies (e.g. ...
... Among the conodonts of this assemblage, almost all of them are known to appear in the Frasnian, indicating that it is a typical Frasnian conodont fauna. Po. brevis generally occurs in the Lower or Upper rhenana Zones (Klapper and Lane 1985;Savage 1992;Ji and Ziegler 1993;Ovnatanova and Kononova 2008;Ovnatanova et al. 2017). Its upper range can be traced to the linguiformis Zone in America, Poland and South China (Ziegler and Sandberg 1990;Woroncowa-Marcinowska 2006;Huang and Gong 2016). ...
... In Italy, it occurs in the Upper rhenana Zone (Mossoni et al. 2012). Po. robustus, which has been reported only in Poland and Canada so far (Szulczewski 1971;Klapper and Lane 1985), ranges from the punctata Zone to a level no higher than jamieae Zone. As for Pn. ...
Pelagic standard conodont biozonation is less applicable to nearshore shallow-water depositional settings close to the peritidal, where it is needed to establish its own conodont zonation. The Upper Devonian Xiejingsi Formation in South China is an important unit for nearshore conodont biostratigraphy investigation as its deposits in the nearshore argillaceous-carbonate mixed facies which yield numerous marine fossils. 25 conodont species belonging to 6 genera of platform elements were described in four sections of this formation in western Hubei, including 9 new species. Based on the conodont distribution in these sections, six nearshore conodont biozones are introduced in ascending order: Polygnathus brevis – Polynodosus changyangensis sp. nov. Assemblage Zone, Icriodus cornutus – Pelekysgnathus arcuatus sp. nov. – Polygnathus brevilaminus Assemblage Zone, Palmatolepis angulata Zone, Palmatolepis angulata – Palmatolepis quadrantinodosalobata Assemblage Zone, Palmatolepis quadrantinodosalobata Zone and Polygnathus planirostratus Zone. The correlation of these biozones with those of pelagic standard ones indicates an age of the Xiejingsi Formation in western Hubei from at least the Frasnian linguiformis Zone to the Famennian Lower rhomboidea Zone. Through constructing nearshore shallow-water conodont biozones based on the establishment of comprehensive conodont succession, this paper provides a case study for high-resolution conodont biostratigraphy investigation in the shallow-water settings.
... simplaZiegler and Sandberg, 1990, GMM B9A.13-60, Sample R-Q base. g Po. aequalisKlapper and Lane, 1985, GMM B9A.13-61, Sample R-Q base. h Po. aequalis Klapper and Lane, 1985, GMM B9A.13-62, Sample R-Q base. ...
The famous Martenberg section of the eastern Rhenish Massif, Germany, type-section of classical Frasnian goniatite and conodont zonations, has been restudied in order to document the microfacies development and to refine the conodont stratigraphy around the global semichatovae Event/Transgression, the proposed level to define a future upper Frasnian substage. More than 8.000 platform elements were identified and include new taxa. Palmatolepis jamieae is subdivided into the subspecies Pa. jamieae jamieae, Pa. jamieae savagei n. ssp., Pa. jamieae rosa n. ssp., and Pa. jamieae ssp. δ. Another new species, Pa. adorfensis n. sp., was previously partly identified as Pa. jamieae, while Pa. descendens n. sp. has previously been described in open nomenclature from Inner Mongolia. Morphotypes are defined in Icriodus symmetricus, Pa. ljaschenkoae, and Pa. proversa. A global literature survey shows that the eustatic semichatovae Event can be recognised in more than 20 regions of all continents with (sub)tropical Upper Devonian outcrops. At Martenberg, the transgression is preceded by a thin but distinctive interval with unconformities, microbial mats, sheet cracks, and currents that brought in the regionally youngest volcaniclastics. The new conodont data confirm that no typical Pa. jamieae (sensu the holotype) occur in the two beds originally supposed to represent the jamieae Zone in its reference section. We fully support the conclusion of Ovnatanova and Kononova (2020) that the jamieae Zone should be abandoned. Early Pa. jamieae subspecies and the related new taxa enter at Martenberg and in a few other regions in the globally easily recognisable Frasnian Zone 10 (= plana Zone). Frasnian Zone 11 (feisti Zone) is subdivided into subzones FZ 11a (= feisti Subzone) and FZ 11b (= nasuta Subzone). The base of the latter coincides with the semichatovae Transgression, the semichatovae Subzone of more shallow shelf settings, and is proposed to define in future the upper Frasnian substage base. On a global scale, the Martenberg section is currently the best bed-by-bed documented section for facies changes, conodont and goniatite biostratigraphy at the middle/upper Frasnian transition. Therefore, it is a prime candidate for a future GSSP selection. A global literature survey identified more than 20 other pelagic conodont successions that have the potential for precise correlation and a better understanding of the environmental changes associated with the semichatovae Event.
... -The form 2 morphotype of Ancyrodella gigas has the outer posterior platform margin formed by a convex curve without development of a sinus and an incurved posterior carina. In contrast, the holotype of A. gigas (re illu s trated by Klapper & Lane 1985, fig. 14.15) has well deve l oped sinuses at slightly posterior of platform midlength and the posterior carina is sinuous rather than incurved. ...
... 12-Polygnathus dubius Hinde 1879; upper view of EUIC S573, Sample S12. Klapper & Lane 1985; upper view of EUIC S595, Sample S13. 14-Icriodus symmetricus Branson & Mehl 1934; upper view of EUIC S580, Sample S12. ...
... Polygnathus and Palmatolepis are two of the most important conodont genera used for biostratigraphy and facies analysis, making up most of the conodont faunas in many Upper Frasnian pelagic facies sections (e.g., Klapper and Lane, 1985;Klapper and Foster, 1993;Ji and Ziegler, 1993;Zhang et al., 2019a) (Fig. 3). ...
... 32-LL-2.5-05. (4-5) Polygnathus evidens Klapper and Lane (1985) specimen number 32- Polygnathus lodinensis Pölsler (1969) specimen number 32-P1-02, 35-P1-1, 35-LL-38.5-01. (9) Palmatolepis bohemica Klapper and Foster (1993) specimen number 1-LL-6-04. ...
... They show the characteristic development of the rows of nodes parallel to the posterior carina. Such node rows are not present in the holotype of A. gigas (reillustrated in Klapper and Lane, 1985, fig. 14.15). ...
... The interval of the subterminus Zone ranges from sample S38-S103. The lower boundary of the subterminus Zone corresponds to the first occurrence of Icriodus subterminus Youngquist, 1947(Narkiewicz and Bultynck, 2007, 2010Bultynck and Gouwy 2008) and the upper boundary coincides with the first appearance of Ancyrodella pristina (Klapper and Lane, 1985). Other accompanying conodonts in the subterminus Zone are the following: Icriodus excavatus, Icriodus expansus, Icriodus cedarensis, Icriodus lilliputensis, Polygnathus xylus, Polygnathus pollocki, Polygnathus alatus, Polygnathus lodinensis, Icriodus difficilis, Icriodus norfordi and Icriodus latecarinatus. ...
... This 63-m-thick interval covers the samples S102-S167 and conodont diversity increases. The lower boundary of the biozone corresponds to the first occurrence of Ancyrodella pristina (Klapper and Lane, 1985). The upper boundary of this biozone coincides to the first appearance of Polygnathus aequalis Klapper and Lane, 1985 in the transitans Zone (Ziegler and Sandberg 1996); the first presence of this species represents the lower boundary of the subsequent biozone. ...
... The lower boundary of the biozone corresponds to the first occurrence of Ancyrodella pristina (Klapper and Lane, 1985). The upper boundary of this biozone coincides to the first appearance of Polygnathus aequalis Klapper and Lane, 1985 in the transitans Zone (Ziegler and Sandberg 1996); the first presence of this species represents the lower boundary of the subsequent biozone. ...
The Middle to Upper Devonian Kuh-e-Bande-Abdol-Hossein section in eastern Central Iran is an overall shallow marine, nearshore to open marine facies setting that contains a highly variable conodont record generally characterised by an Icriodid-Polygnathid biofacies. The lithology and the palaeoenvironmental setting is similar to other localities in Central Iran and exhibits numerous hiatuses. A continuous biostratigraphic record could not be established, but the section preserves the Givetian/Frasnian boundary. Based on the conodont record, major gaps also occur in the Famennian which confirms earlier results reported from other sections of similar palaeoenvironments in Central Iran.
... C o m p a r i s o n. This species is similar to Po. planarius Klapper et Lane, 1985 in the elongated lanceolate platform and thin transverse ridges covering the whole platform surface, but differs in the wider platform, the longer median ridge terminating near the posterior end of the platform, and the posterior end of the platform ornamented with discontinuous ridges split into nodes. ...
The conodont succession in the Upper Devonian facially different sections of northeastern European Russia (Chernyshev Ridge, Subpolar and Polar Urals, Pai-Khoi) are studied. The sections are subdivided and correlated taking into account the global standards for stages. The Frasnian Montagne Noire zonation (Klapper, 1989) is used for the first time for northeastern European Russia; the authors correlated this zonation with the scale of Ziegler and Sandberg (1990). Famennian deposits are subdivided using scale of Ziegler and Sandberg (1984) and miospore zonation. The age of local geological units (formations) is specified; the formations of different regions of northeastern European Russia are correlated to each other and to the regional subdivisions of EEP and Southern Urals. A total of 93 species of Ancyrodella, Mesotaxis, Palmatolepis, Polygnathus, and Zieglerina are described. Two new species Ctenopolygnathus parallelus sp. nov. and Polygnathus masonae sp. nov. are distinguished and five species are described in the open nomenclature. Special chapter is devoted to the phylomorphogenesis of Ancyrodella.
... The platform of Neo. huijunae n. sp. is more reminiscent of the Frasnian Po. aequalis Klapper and Lane, 1985, which is also variable. A few intermediate specimens between Neo. huijunae n. sp. and Neo. ...
Conodont faunas from the Wulankeshun section in the northwestern Junggar Basin of Xinjiang, NW China, are characterized by shallow-water assemblages with many endemic taxa. This study presents separate local icriodid and polygnathid conodont zonations for the Lower Member of the Hongguleleng Formation, which can only roughly be correlated with the pelagic “standard zonation”. The basal beds of the Hongguleleng Formation are assigned to a low diverse Icriodus praealternatus ferus n. ssp. Zone, which possibly includes latest Frasnian strata but it continues into the basal Famennian. At present, the Frasnian/Famennian boundary cannot be defined unequivocally by conodonts and there are no obvious Upper Kellwasser equivalents. However, the carbon isotope data are comparable with the Bulongguor Reservoir stratotype, where the top-Frasnian lies within the basal Hongguleleng Formation. In the early Famennian, partly endemic icriodid lineages provide successive I. cornutus, I. stenoancylus junggarensis n. ssp., and I. plurinodosus n. sp. (sub)zones. The basal Famennian is possibly marked by the oldest, endemic species of Neopolygnathus, Neo. huijunae n. sp. It is followed by local Neo. communis communis, “Polygnathus” pomeranicus, Po. cf. szulczewskii, and “Po.” pseudocommunis n. sp. zones. The new local conodont zonations provide a high biostratigraphic resolution for further future studies. The Wulankeshun shallow-water conodont faunas show restricted similarities with contemporaneous faunas from eastern Europe, Kazakhstan, and Iran. There are a few taxa that were originally described from other distant regions, such as NW Canada and eastern Australia. The local high number of endemic taxa (17 (sub)species, 46%) points to a significant plate tectonic isolation of the Junggar Basin in the early Famennian.
