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1. Map of the Pachitea study area. Land cover classification was carried out in the area shaded gray. Polygons delimit indigenous communities. 500 m buffers around rivers are dark gray. The communities of Laguna-Raya and Santa Maria are labeled. Analysis of nutrient and organic matter chemistry along runoff pathways in a headwater stream at 2400 m asl in the Pachitea basin demonstrated strong terrestrial controls on N, P, and dissolved organic carbon. During the wet season, the narrow riparian zone buffered significant increases in nitrate (NO 3) and dissolved organic nitrogen (DON) originating in ground waters flowing from adjacent forested uplands (2)(Saunders et al. 2006). The riparian zone held a 'reserve buffering capacity' that significantly reduced increases in NO 3and DON concentrations in the river. Retention of soluble reactive phosphorus, ammonium (NH 4 + ), and dissolved organic phosphorus and carbon was not significant. Although absolute values of different parameters varied according to season (wet or dry), patterns of groundwater nitrogen retention were consistent throughout the year. Another important finding was that storm runoff pathways such as overland flow and throughfall carried higher concentrations of inorganic nitrogen to streams, thereby negating the retention capabilities of riparian soils (Ramos et al. 2003, Saunders et al. 2006). Nutrient monitoring data from larger rivers in 

1. Map of the Pachitea study area. Land cover classification was carried out in the area shaded gray. Polygons delimit indigenous communities. 500 m buffers around rivers are dark gray. The communities of Laguna-Raya and Santa Maria are labeled. Analysis of nutrient and organic matter chemistry along runoff pathways in a headwater stream at 2400 m asl in the Pachitea basin demonstrated strong terrestrial controls on N, P, and dissolved organic carbon. During the wet season, the narrow riparian zone buffered significant increases in nitrate (NO 3) and dissolved organic nitrogen (DON) originating in ground waters flowing from adjacent forested uplands (2)(Saunders et al. 2006). The riparian zone held a 'reserve buffering capacity' that significantly reduced increases in NO 3and DON concentrations in the river. Retention of soluble reactive phosphorus, ammonium (NH 4 + ), and dissolved organic phosphorus and carbon was not significant. Although absolute values of different parameters varied according to season (wet or dry), patterns of groundwater nitrogen retention were consistent throughout the year. Another important finding was that storm runoff pathways such as overland flow and throughfall carried higher concentrations of inorganic nitrogen to streams, thereby negating the retention capabilities of riparian soils (Ramos et al. 2003, Saunders et al. 2006). Nutrient monitoring data from larger rivers in 

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Ecosystem services are fundamental to the development of sustainable landscapes but are largely ignored or taken for granted in land management strategies. Ecosystems, and the ecological processes that define them, form the natural infrastructure supporting human activities to enhance the economic and social well-being of communities. This chapt...

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