Yan Wang's research while affiliated with Liaoning Technical University and other places

Publications (40)

Article
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Jeholornis is a representative of the earliest-diverging bird lineages, providing important evidence of anatomical transitions involved in bird origins. Although ~100 specimens have been reported, its cranial morphology remains poorly documented owing to poor two-dimensional preservation, limiting our understanding of the morphology and ecology of...
Article
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The Early Cretaceous diversification of birds was a major event in the history of terrestrial ecosystems, occurring during the earliest phase of the Cretaceous Terrestrial Revolution, long before the origin of the bird crown-group. Frugivorous birds play an important role in seed dispersal today. However, evidence of fruit consumption in early bird...
Article
Significance Owls, with their largely nocturnal habits, contrast strikingly with the vast majority of diurnal birds. A new spectacular late Miocene owl skeleton from China unexpectedly preserves the oldest evidence for daytime behavior in owls. The extinct owl is a member of the clade Surniini, which contains most living diurnal owl species. Analys...
Preprint
Full-text available
The Early Cretaceous diversification of birds was a major event in the history of terrestrial ecosystems, occurring during the earliest phase of the Cretaceous Terrestrial Revolution. Frugivorous birds play an important role in seed dispersal today, and may have done so since their origins. However, evidence of this has been lacking. Jeholornis is...
Article
Dinosaur tracks have been known and reported sporadically from the coal-bearing beds of the Fuxin Formation (upper Aptian–Albian, upper Lower Cretaceous) of Liaoning Province, northeastern China for some time but have not been described in detail. Here available material is illustrated suggesting a diversity of theropod track morphotypes, including...
Preprint
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In order to analyze the genetic evolution of 29 Lilii Bulbus (Baihe), high-density single nucleotide polymorphism (SNPs) marks, and a specific length amplified fragment sequences (SLAFs) were constructed. As a result of sequencing, 76.92 Mb reads were detected. The data of reads for each sample ranged from 1,356,347 to 5,736,545. The average of Q30...
Article
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Yishui Lily 140 (Lilium lancifolium) is a hybrid lily species which was bred from wild lily varieties due to its edible and medicinal value. In this study, we have sequenced the complete chloroplast (cp) of L. lancifolium. The complete cp sequence is 152,643 bp long, with a large single copy (LSC) region of 82,084 bp, a small single copy (SSC) regi...
Article
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The keratinous beak is inferred to have evolved multiple times in the Archosauria and in Aves. Unfortunately, this feature rarely preserves in the fossil record. Here we examine a collection of 603 specimens belonging to the Confuciusornithiformes, a clade of edentulous basal avians, only two of which preserve visible traces of the rhamphotheca. Pr...
Article
Many specimens of the basal bird Jeholornis from the Early Cretaceous Jehol Biota of northeast China include one or two distinctive paddle-shaped skeletal elements preserved in the thoracic region. These ossifications have generally been identified as lateral trabeculae, paired processes of the sternum that are common within the derived avian clade...
Article
Protopteryx fengningensis is from the 130.7 Ma Huajiying Formation making it one of the oldest known enantiornithines. Contributing to its significance, this taxon is also commonly resolved as the basal-most enantiornithine in phylogenetic analyses. Protopteryx preserves several unusual morphologies that are otherwise absent in the Enantiornithes b...
Article
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Background: Tooth morphology within theropod dinosaurs has been extensively investigated and shows high disparity throughout the Cretaceous. Changes or diversification in feeding ecology, i.e., adoption of an herbivorous diet (e.g., granivorous), is proposed as a major driver of tooth evolution in Paraves (e.g., Microraptor, troodontids and aviala...
Article
Direct indicators of diet and predator-prey relationships are exceedingly rare in the fossil record [1, 2]. However, it is through such traces that we can best understand trophic interactions in ancient ecosystems [3], confirm dietary inferences derived from skeletal morphologies [4], and clarify behavioral and ecological interpretations [5]. Here,...
Article
Significance Archaeorhynchus spathula is a basal member of the Ornithuromorpha, the lineage that includes neornithines. Although this is the fifth reported specimen, unlike the others it preserves significant soft tissue, revealing a tail morphology previously unknown in Mesozoic birds and an exceptional occurrence of fossilized lung tissue. This i...
Article
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Birds have a highly specialized and efficient digestive system, but when this system originated remains uncertain. Here we report six gastric pellets attributable to the recently discovered 160-million-year-old troodontid dinosaur Anchiornis, which is among the key taxa for understanding the transition to birds. The gastric pellets contain lightly...
Article
Representatives of the froghopper family Sinoalidae were exclusively known from Jurassic deposits in northeastern China. A new taxon, Fangyuania xiai Chen, Szwedo and Wang, gen. et sp. nov., is erected from mid-Cretaceous Burmese amber and assigned to this family. The remarkable new sinoalid distinctly differs from its con-familial Jurassic relativ...
Article
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STM35-3 from the Yixian Formation is the only Early Cretaceous ornithuromorph preserving direct evidence of granivory. The crop contains numerous seeds and the preservation of gastroliths presumably within the ventriculus indicates this diet was paired with the presence of a gastric mill as in living granivorous birds. STM35-3 was originally referr...
Article
The Qingquan dinosaur tracksite, from the Lower Cretaceous Dasheng Group, Shandong Province, China adds to the growing record of saurischian-dominated ichnofaunas of the region. The site reveals the presence of avian theropods (Koreanaornis) and non-avian theropods tentatively referred to Jialingpus. Sauropod tracks are referred to Brontopodus. One...
Article
The lattice parameters, electronic structure, magnetism and multiferroicity of compound BiMn3Fe4O12 were investigated by using the first-principle calculations. The total energy calculations indicated that BiMn3Fe4O12 was a G-type antiferromagnetic insulator. The ferroelectricity was induced by magnetism and originated from the hybridization betwee...
Article
The KBiFe2O5 (KBFO) compound was synthesized by using a conventional solid-state reaction method and its structural, optical, electrical, and magnetic properties were thoroughly gone through. It was found that KBFO possessed a monoclinic structure with P2/c space group and could be prepared within the calcination temperature range from 1023 K to 10...
Article
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We report on an exceptional specimen of Eoconfuciusornis preserving rare soft-tissue traces of the ovary and wing. Ovarian follicles preserve a greater hierarchy than observed in Jeholornis and enantiornithines, suggesting confuciusornithiforms evolved higher rates of yolk deposition in parallel with the neornithine lineage. The preserved soft tiss...
Article
Here we describe the well-preserved skull of a juvenile specimen of Sapeornis, STM 16-18. Only the tail and hindlimb plumage of this specimen have been previously described. It preserves what we consider to be the complete dentition of Sapeornis with four premaxillary teeth, three maxillary teeth and two tiny dentary teeth on each side, the latter...
Article
There are a growing number of Early Cretaceous avian tracks and trackways from around the world, with Asia (China and Korea) having the largest reported number and diversity of Mesozoic avian traces to date, and these new discoveries are increasing the Early Cretaceous avian ichnodivesrity of Laurasia. Here we report on a new Lower Cretaceous avian...
Article
Bi0.9A0.1FeO3 (A = Ca, Sr and Ba) ceramics are synthesized by using the conventional solid state reaction method. X-ray diffraction analysis shows that all the ceramics have a hexagonal distorted perovskite structure with the space group R3c. It is found that A-site substitution with the smallest ionic radius ions Ca2+ among (Ca2+, Sr2+and Ba2+) ef...
Data
Supplementary Figures 1-15, Supplementary Table 1, Supplementary Discussion and Supplementary References
Article
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Our knowledge of Cretaceous plumage is limited by the fossil record itself: compression fossils surrounding skeletons lack the finest morphological details and seldom preserve visible traces of colour, while discoveries in amber have been disassociated from their source animals. Here we report the osteology, plumage and pterylosis of two exceptiona...
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The Chinese Lower Cretaceous Jehol Group is the second oldest fossil bird-bearing deposit, only surpassed by Archaeopteryx from the German Upper Jurassic Solnhofen Limestones. Here we report a new bird, Chongmingia zhengi gen. et sp. nov., from the Jehol Biota. Phylogenetic analyses indicate that Chongmingia zhengi is basal to the dominant Mesozoic...
Article
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ABSTRACTWe report a new enantiornithine bird, Linyiornis amoena gen. et sp. nov., from the Lower Cretaceous Jiufotang Formation in northeastern China. Traces of ovarian follicles indicate that the specimen represents a female individual. The nearly three-dimensional preservation of the new specimen reveals morphological details rarely visible in ot...
Article
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The historically-famous Lotus Fortress site, a deep 1.5-3.0-meter-high, 200-meter-long horizonal notch high up in near-vertical sandstone cliffs comprising the Cretaceous Jiaguan Formation, has been known since the 13th Century as an impregnable defensive position. The site is also extraordinary for having multiple tetrapod track-bearing levels, of...
Article
The midline pattern of sternal ossification characteristic of the Cretaceous enantiornithine birds is unique among the Ornithodira, the group containing birds, non-avian dinosaurs and pterosaurs. This has been suggested to indicate that Enantiornithes is not the sister group of Ornithuromorpha, the clade that includes living birds and their close r...
Article
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Ornithuromorpha is the most inclusive clade containing extant birds but not the Mesozoic Enantiornithes. The early evolutionary history of this avian clade has been advanced with recent discoveries from Cretaceous deposits, indicating that Ornithuromorpha and Enantiornithes are the two major avian groups in Mesozoic. Here we report on a new ornithu...
Article
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The wings of birds and their closest theropod relatives share a uniform fundamental architecture, with pinnate flight feathers as the key component. Here we report a new scansoriopterygid theropod, Yi qi gen. et sp. nov., based on a new specimen from the Middle-Upper Jurassic period Tiaojishan Formation of Hebei Province, China. Yi is nested phylog...
Article
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A crop adapted for an herbivorous diet of seeds has previously been documented in the Early Cretaceous birds Sapeornis and Hongshanornis. Here we report on several specimens of Yanornis that preserve a crop containing fish. One specimen preserves two whole fish in the oesophagus, indicating that Early Cretaceous birds shared trophic specializations...
Article
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The two groups of archosaurs, crocodilians and birds, form an extant phylogenetic bracket for understanding the reproductive behaviour of dinosaurs. This behaviour is inferred from preserved nests and eggs, and even gravid individuals. Data indicate that many 'avian' traits were already present in Paraves-the clade that includes birds and their clo...
Article
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Four-Winged Birds? Recently, nonavialan dinosaurs with feathers on their fore- and hindlimbs have been described. Zheng et al. (p. 1309 ) describe eleven basal avialan fossils with clear evidence of feathered hindlimbs. Together these fossils show that early avialans possessed four wings, rather than two. A gradual reduction in hindlimb feathering...

Citations

... Jeholornis STM 3-8 was collected from the deposits of the Lower Cretaceous Jiufotang Formation at the Dapingfang locality near Chaoyang, Liaoning Province, China (He et al., 2004), and was referred to J. prima by Hu et al. (2022). This study also includes data from IVPP V13274 (J. ...
... Many new species of Middle Triassic marine reptiles and fishes often represented by complete skeletons, including the earliest turtle, have also been discovered in Guizhou and Yunnan provinces [33]. The Miocene Hezheng Fauna has also produced abundant and diverse mammals and birds, sometimes with exceptional preservation of soft tissues and gut contents [34]. Lastly, the Miocene Zhangpu Biota from Fujian represents another amber treasure trove, recording a diverse fauna and flora of the tropical forest ecosystem [35]. ...
... www.nature.com/scientificreports/ instead, pits and grooves occur dispersed on the labial surface (Fig. 3a) of the dentary and face caudally, as those in Confuciusornis and other edentulous theropods, suggesting the presence of a rhamphotheca [36][37][38] . A small plate-like triangular fragment was interpreted as the left splenial (Fig. 2h). ...
... The first pair of sternal ribs is more robust than the others and shows an elongate fanshaped morphology expanded laterodistally, differing from the slender shaft and the spoon-shaped distal end shared by other dromaeosaurids 5,25 . In some avialans 26 , the caudalmost sternal rib is the most robust and shows an expanded distal end similar to the condition in Daurlong. As in other paravians, the gastralia contact the pubis shaft in a position more proximal than in other theropods, where instead such contact is usually placed on the pubic foot 27 . ...
... The abundant fossil birds from the Jehol Biota not only have increased the taxonomic diversity of the Cretaceous birds, but also have provided critical information regarding aspects of the ontogeny, sexual dimorphism, reproduction, and ecology (Zhang and Zhou, 2000;Zhou and Zhang, 2006;O'Connor et al., 2009;Zheng et al., 2011Zheng et al., , 2014Wang and Zhou, 2019;Bailleul et al., 2020). With increasing information provided by continuous discoveries of enantiornithines, the analysis and interpretation of their evolutionary history, interrelationships, and ecology have been proved challenging tasks Chiappe et al., 2019;O'Connor et al., 2020). The majority of the Jehol Biota enantiornithines have been referred to four main groups: the small-bodied and relatively unspecialized cathayornithids (sensu Wang and Liu, 2016; see also O'Connor and Dyke, 2010), the long-snouted longipterygids (O'Connor et al., 2009), and two groups of large-bodied taxa-the bohaiornithids (M. and the pengornithids (X.-L. . ...
... Diet appears rather conservative in Longipterygidae, with all taxa studied most adapted for invertivory or generalist feeding (except for Shengjingornis, whose diet is indeterminate). This conservation is consistent with previous propositions that adaptations for taking easy-toacquire foods are ancestral in Avialae [104]. Avialans are generally reconstructed as generalist mid-order consumers in Jehol ecosystems [10], and our results do not contradict this. ...
... Instead, the main grip of digit II was probably assisted by Microraptor resting its body weight on the tarsometatarsus 34 . This hypothesis is supported by the small size of its preserved prey 54,55,61,62 , which would be effectively pinned under Microraptor's weight. ...
... The Jehol Biota, one of the most important Mesozoic fossil Lagerstätten, comprises extraordinary taxonomic diversity of micro-and megaplants, invertebrates, and vertebrates; moreover, it provides the most informative source for understanding Mesozoic ecology (Chang et al., 2003;Wang and Zhou, 2003;Jiang et al., 2011;Pan et al., 2012;Carvalho et al., 2015a;Carvalho et al., 2015b). An exceptionally set of well-preserved soft tissues are reported for myriad taxa, especially internal organs and dermal appendices (including feathers and furs) of early birds, feathered dinosaurs, and primitive mammals, which contain most important and irreplaceable information for understanding the early evolution of biological and ecological characteristics (Zhang and Zhou, 2004;Zhang et al., 2010;Pan et al., 2016;Wang et al., 2017a;Wang et al., 2018;McNamara et al., 2018;Pan et al., 2019;Serrano et al., 2020;Ullmann et al., 2020;Cincotta et al., 2022). For example, the reported specimens of basal birds with feathers, including the Jeholornis, confuciusornithidormes, and Enantiornithes present the morphology, arrangement, and even three-dimensional structure of early feathers (Chuong et al., 2003;Zhang and Zhou, 2004;Zhang et al., 2006;Foth, 2012;O'Connor et al., 2012;Wang et al., 2014;Carvalho et al., 2015a). ...
... The remains of insects and arachnids sufficient to diagnose them to family level have been recovered from pellets of some small extant owls (Mrykalo et al., 2009), though, suggesting the possibility they could be recovered in theropods. As yet the only pellets found in association with a theropod are for Anchiornis (Zheng et al., 2018), most recently recovered as a basally branching avialan (Pei et al., 2020). Recently, Freimuth et al. (2021) described pellets bearing the remains of mammals and tentatively referred them to Troodon. ...
... As the first sinoalid from the mid-Cretaceous Kachin amber of northern Myanmar, Fangyuania xiai Chen, Szwedo et Wang, 2018 is distinctly different from its co-familial Jurassic relatives by a series of tegminal characteristics and body structures. To date, ten sinoalid genera with thirteen species have been reported from Kachin amber (Chen et al., , 2019a(Chen et al., ,c,d,e, 2020aHuang, 2019a, 2020;. ...