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We document the occurrence of Early Triassic (Olenekian) crinoid ossicles in exotic blocks contained within the black limestone unit of the Thaynes Formation, which overlies the Dinwoody Formation at the classic Crittenden Springs Smithian ammonoid locality. Crinoid ossicles include two species, i.e., Holocrinus sp. and Articulata ord., fam., gen. et sp. indet. Furthermore, two co-occurring, age diagnostic conodonts, i.e., Neospathodus pakistanensis and Ns. posterolongatus, constrain the age of the crinoids from the early Smithian to the earliest middle Smithian. This discovery represents the third report of Smithian Holocrinidae in the Panthalassan area and it provides important data for the study of crinoid recovery during the Early Triassic.
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Abstract We document an extensive biostratigraphic investigation of a section of the Phukhaothong Dolomite Member of the marine Triassic Chaiburi Formation (part of the Sibumasu [or Shan-Thai] Block), which is exposed on the north side of an isolated mountain at Khao Thong in the Phatthalung area of southern Thailand. The 104 m+ thick section, consisting of bedded to massive, light grey dolomite, contains ammonoids in the middle to upper parts, and conodonts occur in abundant throughout the section. Seven distinct early Spathian (late Olenekian) ammonoid assemblages, a late Smithian (early Olenekian) conodont zone and three early Spathian conodont zones are recognized in ascending order as follows: ammonoids-Columbites sp. indet. beds, Arctomeekoceras? sp. indet. beds, Tirolites sp. indet. B beds, Tirolites sp. indet. C beds and Tirolites sp. indet. D bed in the Tirolites-Columbites Zone, and the Idahocolumbites cheneyi beds and Idahocolumbites phatthalungensis beds in the Idahocolumbites Zone; conodonts-late Smithian, Hadrodontina
aequabilis-Staeschegnathus perrii Zone, early Spathian Icriospathodus crassatus Zone, Triassospathodus
symmetricus-Novispathodus anhuiensis Zone and Novispathodus sp. I-Novispathodus sp. J Zone. The age of the primitive ichthyopterygian Thaisaurus chonglakmanii collected from
the Idahocolumbites cheneyi beds is constrained to the early Spathian, thus suggesting it is the oldest known ichthyopterygian, because the range of Marcouxia and Idahocolumbites is limited to the Columbites parisianus Subzone of the lower Spathian in the western USA. The Spathian ammonoid faunas exhibit a very strong relationship with other Tethyan as well as eastern Panthalassa
faunas in the low paleolatitudes, but bear very little or no relationship with middle and higher latitudinal faunas, suggesting the existence of a strong latitudinal diversity gradient during the Spathian. Late Smithian and early Spathian conodont faunas also exhibit a strong relationship with low paleolatitudinal faunas. Fifty-eight taxa (ammonoids: 26, conodonts: 32) are documented and one new ammonoid species, i.e., Idahocolumbites phatthalungensis, is described.
The ammonoid-bearing limestone blocks at the classic Crittenden Springs ammonoid site belong to the lower part of the Lower Triassic Thaynes Group. These 0.5–1.2 m thick blocks, consisting mainly of bioclastic floatstone and rudstone, contain abundant macro- and micro-fossils such as ammonoids, gastropods, bivalves and scaphopod shells, as well as conodont elements, fish and echinoid remains. Ammonoid and conodont assemblages obtained from three blocks are utilized for biostratigraphical analysis. Twelve informal Smithian ammonoid biostratigraphic intervals from a previous study are condensed into four ammonoid beds, based on the range of age-diagnostic taxa, in ascending order: Meekoceras millardense-M. aff. olivieri beds, Owenites koeneni beds, Anasibirites multiformis bed, and Condensoceras youngi bed. Conodonts recovered from the three blocks consist of 30 species distributed among 15 genera, including one newly described taxon, i.e., Guangxidella minuta Maekawa and Jenks sp. nov. The blocks are divided into two conodont interval zones, i.e., the Novispathodus ex gr. waageni Interval Zone and Nv. pingdingshanensis Interval Zone, based on the first occurrences of their eponymous taxa. Additionally, four conodont range zones, i.e., the Paullella meeki Range Zone, Guangxidella bransoni Range Zone, Scythogondolella milleri Range Zone, and Borinella buurensis Range Zone are recognized, based on the ranges of these four index species. Conodonts within these interval and range zones vary in age from middle Smithian to latest Smithian. The presence of key ammonoid and conodont taxa regarding the Smithian-Spathian boundary (SSB) such as the AW (Anasibirites and Wasatchites) and GXP (Glyptophiceras, Xenoceltites and Pseudosageceras) ammonoid assemblages and the conodonts B. buurensis, Nv. pingdingshanensis and S. milleri demonstrate that the study area is an important reference site for the SSB in the eastern Panthalassa area.
Abstract—We document a relatively small but very important late Griesbachian ammonoid and nautiloid assemblage from the Dinwoody Formation at Crittenden Springs, Elko County, Nevada. This discovery represents the first significant report of late Griesbachian ammonoids in the low-paleolatitudes of eastern Panthalassa, and it also signifies the first report of Wordieoceras wordiei and two co-occurring taxa outside of the Boreal Realm. This similarity in ammonoid faunas, irrespective of paleolatitude, provides support for the concept of weak latitudinal diversity gradients following the end-Permian extinction. The finding is even more noteworthy given the Dinwoody Formation’s reputation for poor fossil preservation and a near complete absence of documented and identifiable ammonoid and nautiloid occurrences. Consisting of four taxa of which two are newly described, the ammonoid fauna includes Wordieoceras wordiei (Spath), Kyoktites cf. K. hebeiseni Ware and Bucher, Wordieoceras mullenae n. sp. and a new taxon belonging to the Mullericeratidae family, Ophimullericeras paullae n. gen., n. sp. The nautiloids are attributed to a newly described species, i.e., Xiaohenautilus mulleni n. sp., a genus heretofore unknown in eastern Panthalassa but commonly reported from the late Griesbachian of South Primorye and the late Griesbachian/early Dienerian of South China.
... North America and West Australia became important regions for the study of Lower Triassic sections of conodont biostratigraphy in the second half of the last century, but there have been only a few continuing studies in this century (Sweet et al., 1971;McTavish, 1973;Paull, 1982Paull, , 1988Paull et al., 1997;Orchard and Zonneveld, 2009;Metcalfe et al., 2013). More recently, a couple of related studies of Smithian-Spathian conodont biostratigraphy have been completed (Maekawa and Jenks, 2021;Golding, 2021;Orchard, 2021). ...
... The Thaynes Formation of northeastern Nevada, which consists of various sedimentary rocks such as limestone, mudstone, shale and sandstone beds, contains abundant marine fossils, i.e., ammonoids, gastropods, bivalves, conodonts and echinoderms (Goodspeed and Lucas, 2007;Lucas et al., 2007a, b). These sediments were traditionally divided into the Dinwoody and Thaynes formations and their geological ages were generally defined by age-diagnostic ammonoids and conodonts (Clark, 1957(Clark, , 1959Kummel and Steel, 1962;Sweet et al., 1971;Solien, 1979;Paull, 1980;Carr, 1981;Clark and Carr, 1984;Mullen, 1985;Jenks, 2007;Jenks et al., 2010Jenks et al., , 2013Jenks et al., , 2021Jenks and Brayard, 2018;Maekawa and Jenks, 2021). Considerable biostratigraphical research has been conducted on the ammonoids and conodonts contained within the discontinuous ammonoid-bearing blocks in recent years, but taxonomic work on other groups, e.g. ...
