Patricia K. Holmgren’s research while affiliated with New York Botanical Garden and other places

A name (Eremogone loisiae) is provided for those plants from northern Utah and southeastern Idaho that have gone under the misapplied name Eremogone kingii subsp. uintahensis. Eremogone loisiae, named in honor of Lois A. Arnow, is distinguished from E. kingii by its longer and narrower, flexuous leaves and its longer sepals, petals, styles, capsules, and seeds.

During preparation of manuscript for volume 2A of Intermountain Flora, we recognized the need to publish 11 new combinations and three second-step lectotypifications. The new combinations are: Eremogone fendleri var. plateauensis, E. fendleri var. porteri, E. fendleri var. brevicaulis, and E. kingii var. compacta (Caryophyllaceae), Atriplex gardneri var. robusta (Chenopodiaceae), Corydalis caseana var. brachycarpa (Fumariaceae), Claytonia parviflora var. utahensis (Montiaceae), Argemone parva and Eschscholzia californica var. mexicana (Papaveraceae), and Aquilegia micrantha var. loriae and Ranunculus adoneus var. caespitosus (Ranunculaceae). Second-step lectotypifications are provided for Talinum confertiflorum and Talinum pygmaeum (Montiaceae) and Platystemon termini (Papaveraceae). Key WordsCaryophyllaceae-Chenopodiaceae-Fumariaceae-Montiaceae-Papaveraceae-Ranunculaceae- Intermountain Flora

Lectotypes are selected for 10 names inHypericum (Clusiaceae),Elatine (Elatinaceae),Arctostaphylos andVaccinium (Ericaceae),Androsace (Primulaceae), andViola (Violaceae).

A lectotype for the nameMentzelia multiflora var.integra is selected and a new combination inMentzelia (Loasaceae) is made. A key to the varieties ofMentzelia marginata is included.

Phylogenetic hypotheses for the largely South African genus Pelargonium L'Hér. (Geraniaceae) were derived based on DNA sequence data from nuclear, chloroplast and mitochondrial encoded regions. The datasets were unequally represented and comprised cpDNA trnL­F sequences for 152 taxa, nrDNA ITS sequences for 55 taxa, and mtDNA nad 1 b/c exons for 51 taxa. Phylogenetic hypotheses derived from the separate three datasets were overall congruent. A single hypothesis synthesising the information in the three datasets was constructed following a total evidence approach and implementing dataset specific stepmatrices in order to correct for substitution biases. Pelargonium was found to consist of five main clades, some with contrasting evolutionary patterns with respect to biogeographic distributions, dispersal capacity, pollination biology and karyological diversification. The five main clades are structured in two (subgeneric) clades that correlate with chromosome size. One of these clades includes a "winter rainfall clade" containing more than 70% of all currently described Pelargonium species, and all restricted to the South African Cape winter rainfall region. Apart from (woody) shrubs and small herbaceous rosette subshrubs, this clade comprises a large "xerophytic" clade including geophytes, stem and leaf succulents, harbouring in total almost half of the genus. This clade is considered to be the result of in situ proliferation, possibly in response to late­Miocene and Pliocene aridification events. Nested within it is a radiation comprising c. 80 species from the geophytic Pelargonium section Hoarea , all characterised by the possession of (a series of) tunicate tubers.

Three new species and two new varieties of Mentzelia belonging to the multistemmed subshrub group of section Bartonia are described. Typical of the group, they are narrow endemics occurring in relatively barren clay soils in the Intermountain Region of interior western U.S. Mentzelia tiehmii and Mentzelia argillicola are from Nevada, Mentzelia memorabalis from northern Arizona, Mentzelia multicaulis variety uintahensis from northeastern Utah and northwestern Colorado, and Mentzelia multicaulis variety flumensevera from central Utah.

Nineteen additional herbaria have been called to our attention since the last installment of this series was published in February 2002. Information on these herbaria is presented below in alphabetical order by country. This brings the total number of herbaria added since the publication of Edition 8 to 393. Information for 3175 herbaria and 9623 people in 165 countries is available for searching by country, institution, city, state, acronym, staff member, correspondent, and research specialty at http://www.nybg.org/bsci/ih/ih.html. Telephone and fax numbers as well as e­mail and URL addresses are included. Note that the Index is fully searchable on research specialty, so it also serves as a PLANT SPECIALISTS INDEX.