Keith Bensusan's research while affiliated with University of Gibraltar and other places

Publications (55)

Article
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Invasive non‐native species (INNS) are recognized as a major threat to island biodiversity, ecosystems, and economies globally. Preventing high‐risk INNS from being introduced is the most cost‐effective way to avoid their adverse impacts. We applied a horizon scanning approach to identify potentially INNS in the United Kingdom Overseas Territories...
Article
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Three new exotic species of ants are recorded for the Macaronesian island of Madeira: Cardiocondyla obscurior Wheeler, 1929, Lepisiota capensis (Mayr, 1862) and Pheidole navigans Forel, 1901. The well-known invasive Pheidole megacephala is now rare on Madeira, possibly because of competition with P. navigans. We did not come across the Argentine an...
Conference Paper
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We tracked pallid swifts (Apus pallidus brehmorum) from a single breeding colony in Gibraltar over two years. Our results show movement of birds between specific regions within the non-breeding geographical area at specific times of the year. The tracking of a single individual showed remarkable fidelity to the areas visited between years. Furtherm...
Article
Full-text available
We tracked pallid swifts (Apus pallidus brehmorum) from a single breeding colony in Gibraltar over two years. Our results show movement of birds between specific regions within the non-breeding geographical area at specific times of the year. The tracking of a single individual showed remarkable fidelity to the areas visited between years. Furtherm...
Article
Phenological mismatch is often cited as a putative driver of population declines in long-distance migratory birds. The mechanisms and cues utilised to advance breeding ground arrival will impact the adaptability of species to further warming. Furthermore, timing of post-breeding migration potentially faces diverging selective pressures, with earlie...
Conference Paper
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Gibraltar has a rich fossil record. The Gorham’s Cave Complex is a UNESCO World Heritage Site that is best known for its occupation by Neanderthals, but many other species are documented to have been present alongside them. Some species present in the fossil record may continue to exist at an archaeological site, allowing study that can fine-tune o...
Article
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Species present in the fossil record may continue to exist at an archaeological site, allowing study that fine-tunes our picture of the ecological past. A large wintering population of Eurasian Crag Martins Ptyonoprogne rupestris (ECM) roosts at the 'Gorham's Cave Complex' UNESCO World Heritage site in Gibraltar, which is best known for its occupat...
Preprint
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Understanding the behaviour of hosts of SARS-CoV-2 is crucial to our understanding of the virus. A comparison of environmental features related to the incidence of SARS-CoV-2 with those of its potential hosts is critical. We examine the distribution of coronaviruses among bats. We analyse the distribution of SARS-CoV-2 in a nine-week period followi...
Article
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This review summarizes current knowledge of the status and threats to the plants and fungi of Great Britain and the UK Overseas Territories (UKOTs). Although the body of knowledge is considerable, the distribution of information varies substantially, and we highlight knowledge gaps. The UK vascular flora is the most well studied and we have a relat...
Book
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Kew’s State of the World’s Plants and Fungi project provides assessments of our current knowledge of the diversity of plants and fungi on Earth, the global threats that they face, and the policies to safeguard them. Produced in conjunction with an international scientific symposium, Kew’s State of the World’s Plants and Fungi sets an important inte...
Article
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Technomyrmex vexatus Santschi, 1919 is recorded for the first time in mainland Spain, from Tarifa in the province of Cádiz. The species was found during a Bioblitz within the Parque Natural del Estrecho. T. vexatus was previously known only from Gibraltar, Northern Morocco and Ceuta (Spain).
Article
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A number of interesing ant species are shared between southern Iberia and northern Morocco. Here, we report two new species to Europe from southern Iberia, both of which were described from North Africa. Temnothorax convexus is an arboreal species that we report from Gibraltar and Cadiz Province in southern Andalusia. Stenamma punciventre is an unu...
Article
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We provide the first records of the subfamily Leptotyphlinae (Coleoptera: Staphylinidae) from Gibraltar. A new species is described, Paratyphlus tristancanoi sp. n., and Gynotyphlus perpusillus (Dodero, 1900) is recorded for the first time. The latter is a parthenogenetic species that was possibly imported accidentally, at the roots of plants.
Article
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Specimens belonging to forty-seven species of Anthicidae from Cadiz province have been studied. Thirteen of them are new records for the fauna of Cadiz and one species (Anthicus crinitus La Ferté-Sénectère, 1849) is recorded for the first time from the Iberian Peninsula. All previous records from literature on the fauna of Cadiz have been compiled....
Conference Paper
A number of specialized cave- and cliff-dwelling vertebrates use the cliffs around Gibraltar, for breeding or roosting. These are comprised of birds and bats. Several are faced with conservation and management issues. Recent advances in tracking may allow us to investigate reasons for declines and facilitate management where this is desirable or ne...
Article
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The Mediterranean Sea is known as an ecological barrier for numerous migratory birds flying from European breeding grounds to African wintering sites. Birds generally avoid migration over open sea and fly over land. In the Mediterranean Basin, few land bridges or bottlenecks for migratory birds exist. The narrowest are at the western and eastern ex...
Article
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A group of friends from the Gibraltar Botanic Gardens and the Science Faculty of Tetouan - Abdelmalek Essaâdi University went in search of Morocco's enigmatic Dragon Trees. They report on their thrilling expedition, the habitats and other plants encountered.
Article
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Authors summarise knowledge of the primarily tropical ant genus Dorylus (Hymenoptera, Formicidae) in Morocco. Until now, only one species has been recorded from Morocco : Dorylus fulvus. We add a second species, Dorylus aethiopicus recorded from Mirleft (southwestern Morocco). This species has already been recorded in the Maghreb, from Tunisia. We...
Article
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Authors summarise knowledge of the primarily tropical ant genus Dorylus (Hymenoptera, Formicidae) in Morocco. Until now, only one species has been recorded from Morocco : Dorylus fulvus. We add a second species, Dorylus aethiopicus recorded from Mirleft (southwestern Morocco). This species has already been recorded in the Maghreb, from Tunisia. We...
Article
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In order to understand patterns of abundances of migratory Lepidoptera in southernmost Europe and contrast this with those of sedentary species, we studied butterflies surveyed along transects during three spring migration seasons at the Rock of Gibraltar. Overall, 2508 butterflies belonging to 19 species were recorded. Of these, the four most nume...
Article
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Under the framework of the DAISIE consortium, whose main mission is to make an inventory of the alien invasive species of Europe and its islands, we review the current state of knowledge and provide an up-to-date catalogue and distributional status for alien seed beetles (Coleoptera: Chrysomelidae: Bruchinae) in Europe. This work is based on studie...
Data
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BIRDS IN GIBRALTAR 2007: impact of a cold spell on populations of Sardinian Warblers Sylvia melanocephala and Zitting Cisticolas Cisticola juncidis at Windmill Hill flats, Gibraltar:Mobbing of migrant Short-toed Eagles Circaetus gallicus by Yellow-legged Gulls Larus michahellis during their northbound passage over Gibraltar: Migrants killed by thun...
Data
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The Eagle Owl Bubo bubo in Gibraltar: Winter Wetland Bird Count in Northern Morocco, January 28th–30th 2005: Probable nesting of the Blue-crowned Conure Aratinga acuticaudata in Gibraltar: A winter roost of White Wagtails Motacilla alba alba in Gibraltar: The ‘Carrefour’ Yellow-browed Warbler: Gibraltar Ringing Report 2005
Data
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the strong and very active European and Central Asian Regional Office in the capital of the European Union. Today, 14 staff members coordinate the activities of the Partnership and work especially hard to improve biodiversity and bird conservation in the European Union, on behalf of and in close cooperation with all of its partners. GONHS was estab...
Data
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The needs of the environment, and of our health and well being must run as a central theme through all Government policies, and within all Government departments, and which should be regularly measured against environ-mental parameters. Energy New generating capacity needs to be pro-vided with minimum delay, but this needs to be coupled with seriou...
Article
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Des prospections entomologiques au Maroc ont permis la découverte pour la première fois d'une fourmi appartenant à la sous famille Cerapachyinae Forel, 1893, élevant ainsi à 10, le nombre de sous familles de fourmis connues au Maroc. Le spécimen a été localisé près de Goulmima, province d'Errachidia. Il appartient au genre Cerapachys Smith, 1857, d...
Article
Explanations for the coexistence of migratory and sedentary (resident) birds are much debated: some authors have suggested that migrants make use of surplus resources not exploited by residents, whereas others have recorded ecological shifts in residents during periods when migrants are present. These shifts have been ascribed to the presence of mi...
Article
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The Eurasian Penduline Tit Remiz pendulinus was until recently considered to be an accidental winter visitor to Morocco. Regular wintering was suspected in the north-west of the country but had been poorly documented. The ringing data collected in the Smir marshes (north-west Morocco) during the period 2004-2008 indicate that Eurasian Penduline Tit...
Article
Raptor migration has been monitored regularly at Gibraltar since the mid-1960s. Long-term trends in abundance of spring migration were investigated for the ten most frequent species, five of which changed significantly in abundance during the study period. Black Kites Milvus migrans, Eurasian Sparrowhawks Accipiter nisus and Booted Eagles Hieraaetu...
Article
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The queen of Anochetus ghilianii is redescribed with specimens from southern Spain, as descriptions and illustrations of this caste provided so far are incomplete. Some observations on the ecology and distribution of the species are also provided.
Article
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The recently described Tetramorium parvioculum was known only from Gibraltar, its type locality. Here, we record it from two new sites around the Strait of Gibraltar, in Ceuta (North Africa) and mainland Spain. In both cases, the species was found in natural habitats, as is the case in Gibraltar.
Article
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A complete moult north of the Sahara has never been reported conclusively in the Reed Warbler Acrocephalus scirpaceus complex. During a ringing program carried out in the Smir marshes (northern Morocco) 140 adults and 292 juvenile Reed Warblers were examined during four autumns in the period 2004–2008. Of these birds, 8.9% of juveniles were either...
Article
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An account is given of the Syrphidae of Gibraltar with a brief description of their habitats: 42 species are recorded, 19 in the subfamily Syrphinae and 23 in the subfamily Milesiinae. Eumerus hungaricus Szilády, 1940 and Merodon eques Fabricius, 1805 are new to Iberia.
Article
Many insects undertake long-distance migrations to exploit seasonally variable conditions at high latitudes, but the mechanisms used by migrants to select and maintain beneficial flight headings are poorly understood. Using computerized flight simulators, we performed controlled experiments to test the ability of an obligate migrant butterfly (Vane...
Article
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Together with the Gibraltar Ornithological & Natural History Society (GONHS), the Gibraltar Botanic Gardens have formed close ties with scientific institutions in the neighbouring country of Morocco, especially with regard to the taxonomy and distribution of plant species that are shared between Gibraltar and the north of Morocco. In addition, the...
Article
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Brown-throated Martin or Plain Martin (Riparia paludicola) breeds across Africa south of the Sahara and southern Asia, with an isolated population in Morocco (the sole population in the Western Palaearctic). During the winter of 2007/2008, we observed the breeding of the species at lower Loukkos, northern Morocco. These observations concern the fir...
Article
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Wetlands are characterized by their high biodiversity. However, these habitats are very vulnerable and are often altered by human activity, which often includes drainage. Many wetlands have disappeared and this trend will continue until ecological awareness increases. The Smir marshes are presently targeted by the tourism industry. Knowledge of the...
Article
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We describe a new species of Tetramorium, which has so far only been found in Gibraltar. Tetramorium parvioculum sp. n. belongs to the simillimum-group, which is of Afrotropical origin. It is likely that this species is a non native and was imported with plant material originating from tropical Africa. The ecology of this new species is discussed.
Article
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Three species belonging to the genus Pyramica are recorded from the area of the Strait of Gibraltar in southern Cádiz province (Andalusia), Spain: Pyramica argiola (Emery, 1869), Pyramica baudueri (Emery, 1875) and Pyramica membranifera (Emery, 1869). For two of these, P. argiola and P. baudueri, this is the first record for the province. The metho...
Article
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The ant species Technomyrmex vexatus (SANTSCHI, 1919) (Formicidae: Dolichoderinae) is recorded from Gibraltar on the Iberian Peninsula. This comprises the first record of the species in Europe and of the genus Technomyrmex MAYR, 1872 in Iberia. The species was found on the Rock of Gibraltar, where it is apparently common in thick maquis vegetation.
Article
Raptor migration has been monitored regularly at Gibraltar since the mid-1960s. Long-term trends in abundance of spring migration were investigated for the ten most frequent species, five of which changed significantly in abundance during the study period. Black Kites Milvus migrans, Eurasian Sparrowhawks Accipiter nisus and Booted Eagles Hieraaetu...
Conference Paper
There are a number of plant species which are peculiar to the Rock of Gibraltar, or which are found there and nowhere else on the European Continent. There are other species claimed to be found on Gibraltar and northern Morocco but which do not occur in the Spanish mainland around Gibraltar. A number of these species were selected for study, namely...
Data
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BIRDS IN GIBRALTAR 2006: Ruppell's Vulture, Moroccan White Wagtail and Icterine Warbler in Gibraltar in 2006: Nesting sites of the Yellow-legged Gull Larus michahellis in north-eastern Morocco: The Migrant Falls of Spring 1968:
Article
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A population of Buprestis (Yamina) sanguinea F. was found on the Rock of Gibraltar during the summer of 2000, feeding on Ephedra fragilis Desf. This confirms a record by Obenberger in 1930, and provides a marked expansion of this species' known distributional range. Adults are active from approximately the 25th July to the 14th August. Photographs...
Article
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Sandy beaches provide a transitional zone between the sea and land. They are regarded as extreme environments due to the harshness of some of their habitat characteristics. These have produced considerable adaptations in insects, many of which are restricted to these habitats. Despite the vast extent of coastline in Morocco (3500 km), insects of sa...
Article
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RESUMEN Se cita por vez primera la presencia de Cymindis suturalis Dejean 1825 en el sureste y sur de la Península Ibérica. Esta especie es afín a C. lineola Dufour 1820 aunque se distingue de la misma por varios caracteres morfológicos, de los cuales la coloración de los élitros es el más conspicuo. Se incluyen algunas consideraciones biogeográfic...

Citations

... 9-15% (Total = 39; Endemic = 3, possibly up to 6; Native = 24; Introduced = 9; Wetterer and Espadaler 2021), Madeira ca. 3% (Total = 31; Endemic = 1; Macaronesian endemic = 1; Native = 8; Introduced = 21; Wetterer et al. 2007;Guillem and Bensusan 2022) although it could also be 0% (see Guillem and Bensusan 2022) and one shared macaronesian endemic in the Azores (Total = 14; Endemic = 0; Macaronesian endemic = 1; Native = 5; Introduced = 8; Wetterer et al. 2004). The low number or even the possible lack of endemic ant species in the Madeira, the Azores and Cabo Verde archipelagos, when compared to the Canary Islands, could be interpreted due to past failure of natural migration, colonisation and evolution of ants, or the rivalry and elimination of endemic and native ants by exotic ants brought by human settlers (Wetterer et al. 2004(Wetterer et al. , 2006(Wetterer et al. , 2007Espadaler 2010, 2021;Guillem and Bensusan 2022). ...
... Pallid swifts tracked the gradual progression of the ITCZ along a decreasing enhanced vegation index (EVI) trend. Even though our sample included only five birds, we found similar patterns in foraging behaviour from GPS logged data presented by Finlayson et al. (2021) corroborating our findings (Supporting information). Crossing the ITCZ and its increasing likelihood of rain results in the first indications of increasing vegetation greenness (EVI), and potential food sources, which seemed to serve as a cue to stop migration. ...
... Seasonal migrations occur annually and are commonly linked to transitions between seasons. For instance, numerous bird species seasonally migrate between breeding ecosystems and wintering ecosystems, which coincides with transitions between seasons (Lawrence et al., 2021). Diel migrations include the return migration of organisms between ecosystems within a 24-h period. ...
... www.nature.com/scientificdata/ seed banking, with 72% of the UK's native and archaeophyte angiosperm species (see Online-only Table 1 for a glossary of terms used) currently conserved in seed banks 7 . ...
... Climate change and the resulting alterations in water regimes also seriously impact the state of water bodies and wetlands [1,8]. The effects of these factors are altered habitats, reduced biodiversity, and threat of extinction in certain species [9][10][11][12]. ...
... The habitat was an open, sun-exposed shrubland with scattered Q. faginea and Q. suber L. trees. The investigated sample was identified as T. convexus according to the original description, the morphologic description provided by Guillem and Bensusan (2019) and by comparison with the imaged syntype of the species in Antweb.org (CASENT0909022). ...
... La sottofamiglia Leptotyphlinae, considerata monofiletica nell'ambito degli Staphylinidae (GREBENNIKOV & NEWTON, 2008), è costituita da elementi di minute dimensioni (0,8-2,10 mm: cfr. PACE, 1996), dal corpo esile e allungato, atteri, depigmentati e anoftalmi, strettamente endogei, prevalentemente a vagilità assai bassa e spesso endemici di aree fortemente circoscritte, ad eccezione di Gynotyphlus perpusillus (Dodero, 1957), specie partenogenetica largamente diffusa nel Mediterraneo e segnalata anche di sporadiche stazioni della Francia atlantica e dell'Europa centrale (COIFFAIT, 1972;HERNANDO et al., 2018;ASSING, 2018aASSING, , 2018b. Di supposta origine gondwaniana (SAÍZ, 1973), la sottofamiglia è distribuita a livello globale in tutte le regioni zoogeografiche (PACE, 2013), tranne quella Orientale (GREBENNIKOV & NEWTON, 2008;OROUSSET, 2016), benché il maggior numero dei suoi rappresentanti (circa il 98%, cfr: OROUSSET, 2016) sia insediato nella sola Regione Paleartica occidentale e in particolare nell'area mediterranea (COIFFAIT, 1972;PACE, 1996; Figg. ...
... Migrating populations remained relatively stable, despite declines in key European breeding populations, in some cases exceeding 30% in 17.1 years (three generations) (BirdLife International, 2018). Bensusan et al. (2007) suggested to compare trends of migrating raptors at bottlenecks with trends on breeding grounds for monitoring the population sizes. However, this requires knowledge of source populations, which is not known for Akrotiri Peninsula. ...
... P. testacea se conoce en las provincias españolas de Barcelona (Menozzi, 1922;Csösz y Seifert, 2003;García et al., 2009;Herraiz, 2010;Bernal, 2016), Burgos (García y Cuesta-Segura, 2017), Cádiz (Guillem et al., 2011; Tabla II. Media de medidas biométricas de cada muestra de P. testacea y P. coarctata. ...
... The faunistic list of the Moroccan ant species was constructed from the results of Cagniant [57] and also included the additions provided in different publications [53,[58][59][60][61][62][63][64][65][66]. ...