K.Yu. Iltsevich’s research while affiliated with St. Petersburg Research Center of the Russian Academy of Sciences and other places

The species composition of the mid-Early Pleistocene fauna in northwestern Azerbaijan has been elucidated through the analysis of fossil material recovered from the Palan-Tyukan site. This research also included a review of the relevant literature. The paleontological material has been fossilized to a similar extent, indicating similar burial conditions. Animal bones at the site were accumulated during a single sedimentation cycle in sediments of what was once a shallow, heavily silted body of water with slow-flowing or standing water. We have identified 16 species of large mammals, whose presence has allowed us to determine the time period in which this faunal complex existed, as well as the characteristics of the natural environment at the time of its formation. The order Carnivora is represented by 10 individuals, the order Proboscidea by 2 individuals, the order Perissodactyla by 3 individuals, and the order Artiodactyla by 18 individuals. The Palan-Tyukan paleo-landscape can be described as a mosaic of different environments. Flat riverine areas with dense vegetation gradually transitioned into the hilly steppe with mixed vegetation of varying densities. Our reconstruction of the paleogeographic environment indicates that large mammals of Palan-Tyukan lived there under conditions of a relatively humid subtropical climate. The Palan-Tyukan faunal complex belongs to the Mediterranean palaeobiogeographical subregion, biozone MNQ18, at the beginning of the Late Villafranchian period, and falls within the time interval from 1.93 to 1.77 million years ago.

The Early Pleistocene site of Palan-Tyukan (MNQ18, ca 1.85 Mya) is located in Transcaucasia, northwestern Azerbaijan. In total, more than 300 mammalian bones were collected there by one of the authors (M.V. Sablin) in 1986 and 1990. The remains were laid close to each other in a 25 m2 lens-like accumulation, in a stratum of normally magnetized (the upper part of the Olduvai subchron) yellowish-grey Lower Apsheron loams. No any excavations have been carried here out since. The present study is based on the analysis of the Bovidae remains. The material is represented by fragments of skulls and horn cores, isolated teeth and bones of the postcranial skeleton. As a result of the revision of the collection, Leptobos (Smertiobos) cf. etruscus (Falconer, 1859), Gazellospira torticornis (Aymard, 1854) and Gazella cf. bouvrainae (Kostopoulos, 1996) were identified. The degree of the unworn M3 tooth hypsodonty of Leptobos, as well as the size and morphology of limb bones indicate a greater similarity with those of L. etruscus from Olivola and Tasso. The characteristics of the horn cores, as well as the upper teeth and limb bones of the medium-sized spiral-horned antelope from Palan-Tyukan indicate a greater similarity with those of the nominative subspecies G. torticornis torticornis. The dimensions and degree of flattening of the horn cores, as well as the parameters of the p4 tooth, are similar to those of the medium-sized gazelle G. bouvrainae, described from several Early Pleistocene localities of Greece. The taxonomic composition of the Bovidae association from Palan-Tyukan is consistent with the previously determined stratigraphic position of the locality, the fauna from which belongs to the Psekups faunal complex of Eastern Europe, correlated with the Late Villafranchian of Western Europe. The composition of these herbivores community testifies to the spread of savanna-like forest-steppe landscapes in the west of Transcaucasia during this period of the Early Pleistocene.

We present here the results of a study of woolly mammoth remains from Yudinovo (Bryansk oblast, Russia) and argue for a revised interpretation of Late Upper Palaeolithic mammoth bone structures in Eastern Europe. Five distinct mammoth bone accumulations have been identified in the main (lower) archaeological layer of Yudinovo, dated to ca. 14.9–14.5 ¹⁴C kya BP (ca. 18.2–17.6 kya cal BP). Taphonomic and zooarchaeological analyses have shown that the mammoth remains found at Yudinovo derive from the butchery of animal carcasses, supporting the hypothesis of mammoth hunting during the Upper Palaeolithic. Our analyses indicate that hunting of both adult and young mammoths took place. Yudinovo was occupied at intervals during both the cold and warm periods of the year, and evidence for various activities is found at the site. The most recently excavated mammoth bone structure was built from a large number of bones, and had a circular or oval form, 4–9.5 m in diameter. The mammoth bone structures have traditionally been interpreted as the remains of dwellings, but we interpret them instead as ritualised middens. Such middens were probably an important part of the socio-symbolic systems of the Palaeolithic hunter-gatherers who built them.

The Early Pleistocene site of Palan-Tyukan (MNQ18, ca 1.85 Mya) is located in Transcaucasia, northwestern Azerbaijan. More than 300 mammalian bones were laid close to each other in a 25 m² lens-like accumulation, in a stratum of normally magnetised (the upper part of the Olduvai subchron) yellowish-grey Lower Apsheron loams. The Palan-Tyukan fauna includes two species of sabertoothed cats. The medium-sized sabertoothed cat remains are ascribed here to Megantereon cf. cultridens. The large size of the massive fossil ulna from Palan-Tyukan is comparable to that of a broadly lion-sized felid. We attribute the bone to the species Homotherium cf. crenatidens. The morphological characters and the size of the Panthera remains from Palan-Tyukan suggest its similarities with the P. gombaszogensis remains from Olivola, Upper Valdarno and Dmanisi localities (the initial phase of Early Pleistocene). The occurrence of P. cf. gombaszogensis in Palan-Tyukan provides information on stratigraphic range of these taxa: it represents one of the earliest records of the genus Panthera in all of Eurasia. The hyaenid remains are ascribed here to Pliocrocuta perrieri. A Feliformia community of the Palan-Tyukan type presents evidence of the wider variety of environments ranging from wooded areas and savanna landscape.

The Early Pleistocene site of Palan-Tyukan (MNQ18, ca 1.85 Mya) is located in Transcaucasia, northwestern Azerbaijan. In total, more than 300 mammalian bones were collected there by M.V. Sablin in 1986 and 1990. The remains were laid close to each other in a 25 m2 lens-like accumulation, in a stratum of normally magnetized (the upper part of the Olduvai subchron) yellowish-grey Lower Apsheron loams. No any excavations have been carried here out since. The present study is based on the analysis of the Caniformia remains. The occurrence of raccoon dog in Palan-Tyukan provides knowledge about the Eurasian stratigraphic range of this taxon: it represents one of the latiest records of the species Nyctereutes megamastoides (Pomel, 1842) in all of Europe. The bear tooth (was found earlear by N.A. Lebedeva nearby) is ascribed here to the species Ursus etruscus Cuvier, 1823. The taxonomic scenario of the Plio-Pleistocene tribe Galictini of Eurasia is intricate and has been debated since the beginning of the last century. The dispute arose mainly due to the scantiness and the sparse nature of their fossil record. The main conclusion about the Pannonictis nestii (Martelli, 1906) remains from Palan-Tyukan was reached after a detailed morphological and metrical analysis. Palearctic badger remains were assigned to the species Meles thorali Viret, 1950 – the ancestor of the European and Asian badgers. The occurrence of M. thorali in Palan-Tyukan represents one of the latiest records of the species. The otter material from Palan-Tyukan is here referred to Lutraeximia cf. umbra Cherin et al., 2016. A Caniformia community of the Palan-Tyukan presents evidence of the wider variety of environments ranging from wooded areas with bodies of water to scrub and even savanna landscape in a relatively humid subtropical climate. The fauna of Palan-Tyukan is an important biostratigrafic reference fauna for mammal zonation in Europe. New discoveries can also help to clarify the taxonomic status of many of these extinct forms.

The Early Pleistocene site of Palan-Tyukan is located in Transcaucasia, northwestern Azerbaijan. More than 300 mammalian bones were laid close to each other in a 25 m2 lens-like accumulation, in a stratum of normally magnetised (the upper part of the Olduvai subchron) yellowish-grey Lower Apsheron loams. The present study is based on the analysis of the Equidae and Suidae remains. The taxonomic scenario of the Early Pleistocene Equidae is intricate and has been a matter of long debate with a multitude of hypotheses. The small-sized horse remains from the locality are ascribed here to species Equus (Allohippus) senezensis. The species likely used here are mostly open landscapes for their main habitat. The main conclusion about the Suidae remains from Palan-Tyukan was reached after a detailed morphological and metrical analysis. The fossil material is here referred to Sus strozzii. Seemingly, during the initial phase of the Early Pleistocene the species lived in humid subtropical Transcaucasian riverine forests and swamps. An Equidae/Suidae community of the Palan-Tyukan type presents evidence of the wider variety of environments ranging from wooded areas including bodies of water to scrub and even savannas as a landscape in a relatively humid subtropical climate.

The article presents the results of the study of osteological material from the early Iron Age hillfort Anashkino (Upper Dvina region, Pskov oblast, Russia). We present here a series of new facts, elaborating our ideas about the economic life, customs and traditions of the local population in this epoch. In total, 20647 determinate bones had been examined from the Anashkino hillfort from the VIII—I century BC cultural layers in 1999—2017. The osteological material is relatively well preserved. Freshwater fish played a significant role in the diet of people in the early Iron Age. In total, 407 individuals of eighteen species mammals were identified at Anashkino. Our analysis showed that the ratio of the live weight of wild and domestic mammals in the diet was approximately equal, and the butchering of animals took place directly at the settlement.

The Epigravettian site of Yudinovo (Bryansk oblast, Russia) was discovered in 1930 by K.M. Polikarpovich. It is located in the Sudost’ river valley and has a unique stratigraphy. This article presents the results of the study of the remains of large mammals from Yudinovo and also discusses their significance in revising the former interpretation of the existence of the site during the very end of the final part of the Late Pleistocene. In total, 38 268 mammalian bones were identified from cultural layers excavated between 1947–2019. The faunal assemblage is relatively small with a dominance of woolly mammoth and arctic fox, typical of a cold and dry tundra-steppe environment. We undertook stable isotopic tracking from samples of bones. Our analyses confirm the hunting of both adult and juvenile larger mammals by ancient humans. Based on the eruption sequence and wear of the milk teeth from young animals, we were able to clarify the season of their death. It seems that these individuals were hunted during the late spring or early autumn. Traces of gnawing by dogs were recorded on a few bones. We also present in this article the results of the study of so-called “dwellings”, constructed by stacking up body parts and bones that were extracted from carcasses of freshly killed mammoths. We interpret these structures as middens representing the remains of ritually deposited hunted game. It can be assumed that these “dwellings” were probably an important part of the socio-symbolic system of the peoples, who created them.

The Early Pleistocene locality Muhkai 2 was discovered in Central Dagestan, northeastern Caucasus, Russia, in 2006. Archaeological investigations of the site have been directed by Corresponding member of RAS H.A. Amirkhanov (Institute of Archaeology RAS, Moscow). The article presents the results of the study of osteological material from the excavations of 2006–2017. The cultural layers with stone tools and mammal bones are found within stratum of brownish-grey petrified loams with reversed magnetization. We identified 14 large species of mammals, which are indicators for the dating of the faunal assemblage and crucial in identifying the specific characteristics of the natural environment at the time of site formation. Stenon horse and Etruscan wolf dominate the spectrum of species at Muhkai 2. All large mammal species from the site are inhabitants of open and semi-open landscapes and animals indicative of closed biotopes are absent. The biotope appears to have been a dry savannah-steppe with small areas of forest vegetation. The climate at this time was warm and quite arid. Most likely, the animals died here due to natural causes and their carcasses were buried at the bottom of an ancient, slightly saline and shallow, temporary body of water. Data from the theriofauna place the age of the site Muhkai 2 within the chronological range of 2.1–1.77 Mya.