Gary G. Anweiler’s research while affiliated with University of Alberta and other places

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Publications (23)


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The Moths of North America. Fascicle 25.4. Noctuoidea, Noctuidae (part): Pantheinae, Raphiinae, Balsinae and Acronictinae.
  • Book

February 2020

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361 Reads

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2 Citations

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Gary G. Anweiler

The New World Noctuidae of the subfamilies Pantheinae (10 genera, 34 species), Raphiinae (1 genus, 1 species), Balsinae (1 genus, 3 species), and Acronictinae (5 genera, 92 species) are revised. With the exception of the Neotropical Pantheinae genera Lichnoptera Herrich-Schäffer, Gaujonia Dognin, and Bathyra Walker, 130 species are treated, 102 of which occur in the United States and/or Canada; 26 are restricted to Mexico and/or Central America; one Eurasian species that could occur in Alaska/Yukon, and one European species that has been introduced as a biological control agent. Four new genera are described, Arctioptera (Type species: Charadra cavillator Morrison), Neopanthea (Type species: Panthea apanthea Anweiler), Characasia (type species: Charadra nigracreta Edwards), and Semixanthia (type species: Charadra franclemonti Anweiler and Schmidt). Two new subgenera of the genus Acronicta Ochsenheimer are proposed: Magnicosta (type species: Anytus atristrigatus Smith) and Dossena (type species: Microcelia retardata Walker). Sixteen new species are described: Panthea grande Anweiler and Schmidt, Semixanthia baja Anweiler and Schmidt, Charadra sonorensis Anweiler and Schmidt, Charadra insperata Anweiler and Schmidt, Charadra landolti Anweiler and Schmidt, Charadra ochrea Anweiler and Schmidt, Chloronycta vulcanica Schmidt and Anweiler, Acronicta nigrosuffusa Schmidt and Anweiler, Acronicta vermiforma Schmidt and Anweiler, Acronicta wiklei Schmidt and Anweiler, Acronicta immodica Schmidt and Anweiler, Acronicta cryptica Schmidt and Anweiler, Acronicta magnirena Schmidt and Anweiler, Acronicta texana Schmidt and Anweiler, Acronicta mexicana Schmidt and Anweiler, and Acronicta maya Schmidt and Anweiler. Acronicta vinnula floridensis Schmidt and Anweiler is described as a new subspecies. Six neotypes and 18 lectotypes are designated, and eleven species are subsumed into synonymy. Two species are raised from synonymy. Genitalia, dot range maps, and adults are illustrated, the latter in color. Color larval photographs for 77 species are also provided.


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  • Data
  • File available

July 2014

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195 Reads

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Brigette Zacharczenko

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Gary Anweiler
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Geographic distribution and phenotypic variation of Raphia frater subspecies. Circles indicate specimens examined during this study: green – subsp. piazzi; pink – subsp. abrupta; black – subsp. frater; blue – subsp. coloradensis; yellow – subsp. cinderella. Multi-coloured circles indicate transitional populations and/or phenotypically intermediate specimens between respective subspecies. a
Raphia frater piazzi (Zavallo Co., TX) b
Raphia frater abrupta (Oktibeha Co., MS) c
Raphia frater abrupta (Cottle Co., TX) d
Raphia frater abrupta (Cottle Co., TX) e
Raphia frater abrupta (Montgomery Co., MD) f, g
Raphia frater frater (Edmunston, NB) h
Raphia frater abrupta – frater intermediate (Anne Arundel Co., MD) i
Raphia frater abrupta – frater – coloradensis intermediate from highly variable population in Cherry Co., NE j
Raphia frater coloradensis (Alamosa Co., CO) k
Raphia frater coloradensis (Milk River valley, AB) l
Raphia frater coloradensis (Sanpete Co., UT) m
Raphia frater coloradensis (Elko Co., NV) n
Raphia frater cinderella (Ventura Co., CA) o, p
Raphia frater coloradensis – frater intermediates (Chelan Co., WA) q
Raphia frater elbea (Cochise Co., AZ) r
Raphia frater elbea (San Juan Co., UT) s
Raphia frater elbea (Santa Cruz Co., AZ). All specimens are males.
Distribution of Raphia frater subspecies (circles) relative to range of Populus larval host plants (coloured shading) in eastern North America. Black circles - subsp. frater; white circles – subsp. abrupta; grey circles – subsp. piazzi; half-circles represent transitional populations and/or phenotypically intermediate specimens. Blue shading – combined ranges of Populus tremuloides and Populus grandidentata; yellow shading – Populus deltoides; range overlap depicted in green. Populus ranges adapted from USGS (2013).
Distribution of Raphia frater subspecies (circles) relative to range of Populus species (coloured shading) in western North America. Half-circles represent transitional populations and/or phenotypically intermediate specimens. Ranges for Populus trichocarpa + Populus balsamifera and Populus angustifolia + Populus deltoides are combined, with both Populus balsamifera and Populus deltoides occuring in Alberta – Montana (upper right). Populus ranges adapted from USGS (2013).
Neighbour-joining tree and associated sampling sites of mtDNA barcode haplotypes in Raphia frater. Haplogroup colour corresponds to that of sampling sites. Subspecies assignment based on morphology and sample size is indicated for each haplogroup. Width of triangles is proportional to number of haplotypes, height represents maximum divergence within haplogroup.
Genitalia of Raphia frater. a female (Konza Prairie Biol. Stn., Riley Co., KS; leg. Metlevski), with inset showing apical spatulate setae and subapical dense setal crown of papillae anales b male vesica c male genital capsule (Pitchfork Ranch, Grant Co., NM; leg. C. Ferris); note magnification difference between sexes.
Taxonomy and biogeography of the Nearctic Raphia Hübner (Lepidoptera, Noctuidae, Raphiinae)

June 2014

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227 Reads

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1 Citation

Citation: Schmidt BC, Anweiler GG (2014) Taxonomy and biogeography of the Nearctic Raphia Hübner (Lepidoptera, Noctuidae, Raphiinae). In: Schmidt BC, Lafontaine JD (Eds) Contributions to the systematics of New World macro-moths V. ZooKeys 421: 91–113. doi: 10.3897/zookeys.421.7517 Abstract The taxonomic status and biogeography of the North American Raphia species is reviewed using adult morphology, larval host plants, geographic phenotypic variation, and variation of mtDNA COI barcode sequences. Lack of diagnostic morphological differences, combined with relatively low mtDNA barcode divergences and clinal phenotypic variation in key geographic regions indicate that the six previously recognized species of North American Raphia are best interpreted as parapatric subspecies. Raphia frater abrupta Grote, stat. n., R. f. coloradensis Putnam-Cramer, stat. r., R. f. piazzi Hill, stat. n., and R. f. elbea Smith, stat. n., are accordingly revised to subspecies of R. frater Grote. Type locality restrictions are pro-vided for Raphia abrupta and Raphia frater and a neotype is designated for Raphia frater var. coloradensis.






Citations (6)


... The group has not been revised for many decades but all three Canadian species were treated by Forbes (1923) and are reasonably well known. However, it was only fairly recently that one of our species was recognized as Holarctic rather than non-native, based on a remote collection locality in northern Alberta (Pohl et al. 2005). Although there are more BINs available than there are documented species, it is thought that this represents intraspecific variation, and no undocumented species are expected (Table 1). ...

Reference:

Lepidoptera of Canada
New records of microlepidoptera in Alberta, Canada

Journal of the Lepidopterists’ Society

... This might be the product of the park's close proximity to shipping ports, the high diversity of ornamental plants and other non-native hosts, and the disturbed condition of the park following the windstorms that allowed the populations to increase to a level that could be detected. Another contributing factor could simply be the addition of DNA barcoding to the arsenal of detection ( Armstrong and Ball 2005;Chown et al. 2008;deWaard et al. 2008b). Typically an introduced species persists at low population densities before becoming established (Tilman 2004) so it is expected that few if any individuals will be collected. ...

First Canadian records of Lampropteryx sujfumata ([Denis & Schiffermiiller], 1775) (Geometridae: Larentiinae)

... The European skipper (Thymelicus lineola) was introduced at London, Ontario, in about 1910 and has spread widely in eastern North America but more locally in central and western North America. The spread of this species in central Alberta is relatively well-documented (Schmidt et al. 2003) and continues today. It feeds on a variety of grasses, but timothy grass (Phleum pratense L.), a common pasture grass, is preferred. ...

Additional butterfly records from Alberta, 1999-2002

Blue Jay

... Noctuoidea, the largest of the lepidopteran superfamilies, is of specific interest, having been studied by multiple authors following the discovery of the NGP (Sokolov 1936, Henke andKruse 1941). Our focus on the Acronictinae (Lepidoptera: Macroheterocera: Noctuidae) is occasioned by several recent systematic studies (Wagner 2007, Wagner et al. 2008, Schmidt et al. 2014, Rota et al. 2016) and by a number of biological attributes amenable to the study of wing pattern. The Acronictinae are an attractive study system for at least two reasons. ...

Polyphyly of Lichen-cryptic Dagger Moths: Synonymy of Agriopodes Hampson and description of a new basal acronictine genus, Chloronycta, gen. n. (Lepidoptera, Noctuidae)

... Further research on historical tree presence in the Great Plains may help clarify how typical these results are. In any event, cottonwoods are species of cultural value, providing vibrant fall colors, and an important resource for wildlife species such as pollinators [51]. Recently, cottonwood trees have been dying without replacement, due to altered hydrology in combination with other factors [52], which may represent a reset within the range of historical variation but reinitiation of establishment may require a specific combination of disturbance regimes. ...

Moths and Butterflies of the Prairies Ecozone in Canada

... Panthea is a small genus comprising 14 species worldwide, distributed mainly in the Nearctic and Palearctic regions (Anweiler 2009;Behounek et al. 2013;Schmidt and Anweiler 2020). Only three species are known in the Neotropical region (Panthea furcilla (Packard, 1864), P. reducta Anweiler, 2009, andP. ...

Revision of the New World Panthea Hübner (Lepidoptera, Noctuidae) with descriptions of 5 new species and 2 new subspecies