Bryn J. Mader's research while affiliated with City University of New York,- Queensborough Community College and other places

Publications (12)

Article
The peculiar sphenoidal “pits” of the common warthog (Phacochoerus africanus) long have been recognized as a feature that distinguishes that species from the closely related desert warthog (P. aethiopicus). Authors seem to regard these structures as blind pockets that do not extend beyond the basicranium. However, these structures actually are open...
Article
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The brontothere genera Metarhinus and Sphenocoelus (= Dolichorhinus) are both characterized by an unusual redirection of the air passageway in which the internal nares are effectively closed off and new openings that serve as the functional internal nares have appeared posteriorly. Rather than passing lateral to the vomer, the airstream appears to...
Article
In 1968 Schultz and Falkenbach described a new oreodont subspecies, Brachycrus laticeps mooki, from a locality 11 km (7 miles) southeast of Fort Logan, Meagher County, Montana. As a holotype for this new taxon, Schultz and Falkenbach (1968:369) designated both American Museum of Natural History (AMNH) specimen number 21321 (a skull) and AMNH 21322...
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Although pterosaurs from Africa are still rare, in recent years several specimens have been described from the Kem Kem beds (Upper Cretaceous, Cenomanian) of Morocco. Here we describe four additional specimens from this informal lithostratigraphic unit: a jaw fragment, two mid-cervical vertebrae, and a humerus. All these specimens show three-dimens...
Article
A systematic revision based on a morphological and statistical analysis recognizes two species of Eotitanops (E. borealis, and E. gregoryi) and three species of Palaeosyops (P. paludosus, P. fontinalis, and P. robustus) as valid. The name P. robustus is selected as having priority over the name P. laticeps under the Principle of First Reviser. Eoti...
Article
An extraordinarily well-preserved skull of the Uintan brontothere genus Metarhinus, supplemented by a second, less complete specimen, permits a detailed description of the cranial anatomy, including aspects that have never before been described. Of particular interest is the structure of the nasal cavity and internal nares, which may be virtually u...
Article
A remarkably well-preserved primitive horned brontothere skull from the Wiggins Formation of western Wyoming is described. Noteworthy characters of this specimen include an extensive system of large paranasal sinuses, a distinct vomerine plate, and a confluence of the anterior lacerate foramen and foramen rotundum. A nearly complete endocranial cas...
Article
Diplacodon progressum Peterson is not referable to the genus Diplacodon Marsh and is assigned the new generic name Pseudodiplacodon. Diplacodon is regarded as a nomen dubium, but may be a senior synonym of Eotitanotherium, a genus that is clearly distinct from Pseudodiplacodon. Only the type species of Pseudodiplacodon is accepted as valid, althoug...
Article
A systematic revision based on a morphological and statistical analysis recognizes two species of Eotitanops (E. borealis, and E. gregoryi) and three species of Palaeosyops (P. paludosus, P. fontinalis, and P. robustus) as valid. The name P. robustus is selected as having priority over the name P. laticeps under the Principle of First Reviser. Eoti...
Article
A systematic revision based on a morphological and statistical analysis recognizes the following Bridgerian and Uintan brontothere species as valid: Mesatirhinus megarhinus, Metarhinus fluviatilis, Metarhinus diploconus, Sphenocoelus uintensis, Sphenocoelus intermedius, Sphenocoelus hyognathus, Telmatherium validus, Protitanotherium emarginatum, Ps...
Article
Full-text available
5 ABSTRACT: We report the occurrence of an isolated avian dorsal vertebra in Cretaceous strata of Morocco. The specimen was found near Erfoud oasis in the Tafilalt region of southeastern Morocco preserved in a reddish sandstone. It has a large neural canal (ratio height neural canal/cranial articulation surface = 0.67), a feature regarded as a syna...

Citations

... The dataset includes all taxa belonging to Brontotheriidae that produce unique combinations of character data. All brontotheres were included as species except Palaeosyops due to uncertainty about the species-level taxonomy of this genus (Gunnell and Yarborough, 2000;Mihlbachler, 2008;Mader, 2009). Data for 15 characters (5, 30, 71, 72, 74-82, 88, 94), previously missing for Protitanops curryi, were entered based upon the newly described material. ...
... Peterson also more fully described the anatomy (p. 409) noting peculiar pouch-like structures (choanal pouches of Mader, 2008, andMihlbachler, 2008) that close off the internal nares and re-direct the air stream posteriorly to exit at a new opening between the pterygoid processes. Osborn (1929) identified the choanal pouches of Sphenocoelus as maxilloturbinates, as did Mihlbachler (2008). ...
... The Epapatelo otyikokolo (MGUAN-PA650) humerus displays an entepicondyle that is wider than the ectepicondyle in dorsoventral width, with the ectepicondyle of the humerus projecting anteriorly (a pteranodontid feature) [17,33]. The entepicondyle projects distally, a feature shared with nyctosaur A. elainus [17] and azhdarchoid CMN 50814 [60] (CMN: Canadian Museum of Nature, Ottawa, Canada). This gives the humerus a strong distal expansion that is also similar to Pteranodon sp., T. regalis, and A. elainus, a derived feature of non-Nyctosaurus pteranodontoids, but different from azhdarchoids that only have an expansion of the distal one-third of the humerus [17]. ...
... Herbivores are rare, with only occasional remains of sauropod dinosaurs reported (Lamanna and Hazegawa, 2014;Ibrahim et al., 2016), of which Rebbachisaurus is the only named taxon (Lavocat, 1954a). A diverse assemblage of crocodylomorphs is present (Sereno and Larsson, 2009), in addition to turtles (Gaffney et al., 2002;Gaffney et al., 2006), snakes (Rage andDutheil, 2008;Klein et al., 2017) amphibians (Rage andDutheil, 2008) and possibly birds (Riff et al., 2004). ...
... They are not the same as the sphenoidal pits in P. aethiopicus, but rather open in the center of the sphenoidal pits. We apply the term "sphenoidal apertures" to these openings, a term that was first used by Mader (2009) to describe similar openings in the nasopharyngeal region of an extinct perissodactyl (see Discussion, below). It is the purpose of the present work to provide a detailed description of the sphenoidal apertures in P. africanus as well as the various cranial sinuses. ...
... Lucas (1983) and Lucas and Schoch (1989) considered B. transylvanicum a synonym of the North American Diplacodon. Although Diplacodon is currently recognized as a nomen dubium (Mader, 2000), it was considered at the time to be a senior synonym of another North American genus, Protitanotherium (Lucas, 1983). Protitanotherium is currently recognized as a valid genus (Mader, 1989); however , Brachydiastematherium differs from Protitanotherium in two significant ways; there is no postcanine diastema, and the p3 possesses a large molariform metaconid. ...
... A bony structure projecting from the internal nares between the pterygoids in the same general location as the choanal pouches of Sphenocoelus was interpreted by Mihlbachler (2008) as posteriorly displaced turbinates. However, Mader (2009) identified this same structure as a greatly enlarged vomer. Mader noted that the internal nares were essentially closed off and that an opening posterior to the enlarged vomer was the functional exit for the airstream. ...
... It follows that as detailed in Paul et al. (2022 Suppl.) in modern peer reviewed literature as few as one skeletal character, often minor, has been used to separate species extant and fossil known from well documented remains often including complete skulls and skeletons, with many cases involving just two, three or a few features, among continental and marine tetrapods, with proposed species sometimes being contemporary, or at least partly separated stratigraphically (tyrannosaurids --Currie 2003a; ornithopod H. multidens - Barrett et al. 2005;Panthera spelaea, P. atrox --Sotnikova and Nikolskiy 2006;hadrosaurs Corythosaurus, Lambeosaurus --Evans and Reisz 2007;ichthyosaur Stenopterygius --Maisch 2008;Maxwell 2010;brontotheres Eotitanops, Palaeosyops, Metarhinus, Rhinotitan, --Mihlbacher 2008;Mader 2010;Psittacosaurus -Sereno 2010;plesiosaur Pliosaurus -Knutsen 2012;diplodocids Apatosaurus, Brontosaurus, Diplodocus --Tschopp et al. (2015); chasmosaurines - Campbell et al. 2016;Fowler and Freedman 2020;ornithomimids Ornithomimus, Dromiceiomimus --MacDonald & Currie 2018;proboscideans Mammut, Palaeoloxodon --Dooley et al. 2019;Larramendi et al. 2020). In a reversal of the proposition that species best documented by fossil remains should be distinguished by lots of characters, the Psittacosaurus known by the largest number of specimens, P. mongoliensis, is distinguished from sister species by the least number of attributes, two, in Sereno (2010). ...
... The Kem Kem assemblage has yet to yield diagnostic remains of a bird or mammal. Riff et al. (2002) described a small isolated vertebra from the Kem Kem Group as avian solely on the basis of its capacious neural canal, drawing comparisons to other Gondwanan taxa now regarded as non-avian (alvarezsaurids, unenlagines). Cavin et al. (2010), probably on that basis, added indeterminate avians to their faunal list. ...