Alain Waller’s research while affiliated with French National Centre for Scientific Research and other places

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Publications (30)


True Flies (Insecta: Diptera) from the Late Eocene Insect Limestone (Bembridge Marls) of the Isle of Wight, England, UK
  • Article

December 2019

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143 Reads

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27 Citations

Earth and Environmental Science Transactions of the Royal Society of Edinburgh

Wiesław Krzemin´ski

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Krzemin´ Krzemin´ski

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[...]

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David B Nicholson

The Diptera fauna from the late Eocene of the Isle of Wight (Bembridge Marls) is studied including redescriptions of formerly described material. The fauna includes the following


Evolution and palaeosynecology of the Mesozoic earwigs (Insecta: Dermaptera)

August 2012

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59 Reads

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18 Citations

Cretaceous Research

Sinoprotodiplatys zhangi gen. et sp. nov. is described from the Lower Cretaceous Yixian Formation in Liaoning Province, near Beipiao City, China. It is one of the most recent representatives of the suborder Archidermaptera. The classification and synonymies among the archidermapteran families are reassessed. The palaeosynecological evolution of the Mesozoic earwigs is analysed using the new tool of the ‘Wagner Parsimony using Morphological Characters’ method (‘WAPUM’), suggesting that the evolution of the morphology of these insects was very progressive, perhaps owing to their way of life in the soil.


Fig. 1. Photographs of braconid wasp Palaeocharmon basalis gen. et sp. nov., holotype of female, MNHN A32944, from the French Amber (Farm Le Quesnoy, Oise, France), lowermost Eocene. A. Habitus in dorsal view. B. Habitus in lateral view. C. Part of the body and wing in dorsal view.  
Fig. 3. Photographs of braconid wasp Phanerotoma menieri sp. nov., holotype of female, A32945, from the French Amber (Farm Le Quesnoy, Oise, France), lowermost Eocene. A. Habitus in lateral view. B. Metasoma in lateral view. C. Head and mesosoma in lateral view. D. Head in front view. E. Habitus in dorsal view. F. Metasoma in dorsal view.  
Fig. 4. Drawings of braconid wasp Phanerotoma menieri sp. nov., holotype of female, A32945, from the French Amber (Farm Le Quesnoy, Oise, France), lowermost Eocene. A. Habitus in lateral view. B. Part of the left fore wing. C. Part of the right fore wing.  
New Fossil Non-Cyclostome Braconid Wasps from the Lowermost Eocene Amber of Paris Basin
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  • Full-text available

September 2010

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279 Reads

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11 Citations

Acta Palaeontologica Polonica

G. 2010. New fossil non−cyclostome braconid wasps from the lowermost Eocene amber of Paris Basin. Acta Palaeontologica Polonica 55 (3): 519–527. Two braconid wasp taxa with enlarged eyes and ocelli indicative of probable nocturnal activity are discussed and de− scribed from the lowermost Eocene amber of the Paris Basin.The new tribe Palaeocharmontini nov., for new genus Palaeocharmon with type species Palaeocharmon basalis sp. nov. is described and illustrated in the subfamily Char− montinae. The similarity of this taxon with members of subfamilies Helconinae, Homolobinae, and Brachistinae is shown and the peculiar character [presence of hind wing recurrent vein (m−cu)] unknown previously in non−cyclostome braconids is discussed. A third fossil species of the genus Phanerotoma (Ph. menieri sp. nov.) is described and compared with known species from the Baltic amber.], Museum and Institute of Zoology Polish Academy of Sci− ences, ul. Wilcza 64, 00−679 Warszawa, Poland; André Nel [anel@mnhn.fr], Alain Waller, and Gael De Plöeg, Laboratoire d'Entomologie, Muséum National d'Histoire Naturelle, 45 rue Buffon, F−75005 Paris, France.

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Figure 1 Photograph of Eatonisca tertiaria Meunier 1905, Plesiotype num. 2105.
Figure 5 Drawing of the head of Eatonisca tertiaria Meunier 1905, frontal view; scale bar represents 0.3 mm.
Figures 2-4 Eatonisca tertiaria Meunier 1905. 2, photograph of head showing the eye bridge; 3, photograph of antennal fl agellomeres, arrows show the ascoids; 4, frontal view of the head, the arrow shows the depression on the scape.
Redescription of Eatonisca tertiaria Meunier 1905 from Baltic amber (Diptera: Psychodidae: Trichomyiinae)

January 2010

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85 Reads

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10 Citations

Annales de la Société entomologique de France (N S )

Eatonisca tertiaria Meunier 1905, is redescribed from the Eocene Baltic amber, with details of the eyes, antennae and male genitalia structure provided. This fossil is placed within Trichomyiinae, and shares several plesiomorphies with Psychodinae, Phlebotominae, and Sycoracinae.



A phantom midge from the Lower Cretaceous Lebanese amber (Diptera, Chaoboridae)

January 2009

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322 Reads

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6 Citations


Mantophasmatodea now in the Jurassic

July 2008

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974 Reads

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70 Citations

The Science of Nature

The Mantophasmatodea is the most recently discovered insect order. The fossil records of all other 'polyneopteran' orders extend far in the past, but the current absence of pre-Cenozoic fossils of the Mantophasmatodea contradicts a long evolutionary history, which has to be assumed from the morphological distinctness of the group. In this paper, we report the first Mesozoic evidence of a mantophasmatodean from the Middle Jurassic of Daohugou, Inner Mongolia, China. Furthermore, the new fossil shares apomorphic characters with Cenozoic and recent Mantophasmatodea, suggesting a longer evolutionary history of this order.


March flies and European Cenozoic palaeodimates (Diptera : Bibionidae)

April 2008

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72 Reads

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28 Citations

Annales de la Société entomologique de France (N S )

The proportions of the different bibionid genera are used to estimate the temperature changes between the Late Eocene and the Late Miocene in Western Europe. The genus Plecia is a good indicator of the warm Cenozoic palaeoclimates. We propose a new approach for the estimation of the value of the palaeoclimatic information that could be obtained by actualistic inferences using extinct taxa. Other taxa (i.e. Mastotermitidae) that could appear a priori better palaeoclimatic indicators result to be less sensitive than bibionids to temperature changes. Intraspecific and intrageneric morphological diversity is explored for the genera Bibio, Penthetria and Plecia, showing the difficulties to characterize the fossil bibionid species.


The first fossil Compsocidae from Cretaceous Burmese amber (Insecta, Psocoptera, Troctomorpha)

December 2007

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88 Reads

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34 Citations

Cretaceous Research

The first fossil record of the Compsocidae, Burmacompsocus perreaui gen. et sp. nov., is described from Late Albian Burmese amber. Its strong similarity to the two extant compsocid genera suggests a remarkable morphological stability within this group of 100Ma. This family, now known only in Central America, was certainly more widespread in the past.



Citations (26)


... Mouthpart morphology can be useful to assess the trophic position of Dermaptera species (Coll & Guershon, 2002). In carnivorous species the incisive area occupies two thirds of the total mandibular length, while it is restricted to half the mandibular length in phytophagous and omnivorous species (Waller, Caussanel & Jamet, 1996). The few existing stable isotope data revealed high variability within and among species (Okuzaki et al., 2009), confirming an omnivorous diet with a wide range of consumed resources and diet switching. ...

Reference:

Feeding habits and multifunctional classification of soil-associated consumers from protists to vertebrates
Variation morphologique des pièces buccales chez quelques Dermaptères
  • Citing Article
  • January 1996

Bulletin de la Société entomologique de France

... Identification keys (Braet et al. 2012;Koffi et al. 2020c) and characteristics of insects already identified using molecular barcodes from Africa (Agboyi et al. 2020;Durocher-Granger et al. 2021;Otim et al. 2021) were used to identify emerged parasitoids and collected predators (Brindle 1967;Waller et al. 1999;Kwadjo et al. 2012;Nicolas et al. 2015;Girod and Lassalle 2017). ...

Etude comparée des pièces thoraciques et de leurs appendices chez quelques Dermaptères
  • Citing Article
  • January 1999

Bulletin de la Société entomologique de France

... It is worth mentioning that two very similar species described from females preserved in Burmese amber are placed in two different genera and subfamilies, i.e., Burmaculex Borkent & Grimaldi, 2004 in the fossil subfamily Burmaculicinae (Borkent & Grimaldi, 2016) and Priscoculex Poinar et al., 2019 in the extant subfamily Anophelinae (Poinar et al., 2019). Fossil mosquitoes from the Tertiary were reviewed by Edwards (1923), Statz (1944), Evenhuis (1994), Szadziewski (1998), Krzemiński et al. (2019) and Giłka et al. (2021). Recently, Azar et al. (2023) described males of a chaoborid-like Libanoculex intermedius from Lower Cretaceous Lebanese amber (Barremian,, which they placed, as the presumed oldest fossil mosquito, in its own subfamily Libanoculicinae of the Culicidae. ...

True Flies (Insecta: Diptera) from the Late Eocene Insect Limestone (Bembridge Marls) of the Isle of Wight, England, UK
  • Citing Article
  • December 2019

Earth and Environmental Science Transactions of the Royal Society of Edinburgh

... The discovered biodiversity of Psocodea in the Mesozoic and Cenozoic is significantly higher, with fossils collected from various localities and horizons. Taxa from the Mesozoic are reported from the Late Jurassic of Kazakhstan [48] and the Cretaceous ambers of Canada, France, Lebanon (Table 1), Myanmar, Russia, Spain, and the USA [37,[49][50][51][52]. Taxa from the Cenozoic are reported from the Eocene limestone of England (Isle of Wight) [53], the Eocene ambers of the Baltic region, China (Fushun), France (Oise), Germany (Bitterfeld amber) and Ukraine (Rovno amber) [54][55][56][57][58][59][60], and the Miocene ambers of Mexico, Costa Rica and the Dominican Republic [29,[61][62][63][64][65]. In the last two decades, the earliest definitive undisputed record of parasitic lice was Megamenopon rasnitsyni Wappler, Smith and Dalgleish, 2004, from the Eocene of Germany (Eckfeld maar) [66,67], although, the recent discovery of Archimenopon myanmarensis Zhang, Rasnitsyn, Zhang, Song, Shih, Ren, Wang, Li and Gao, 2024, assigned to the new stem family Archimenoponidae, from Burmese amber set back their oldest record to the mid-Cretaceous [68]. ...

The first fossil Myopsocidae (Psocoptera) in Dominican amber
  • Citing Article
  • November 2006

Zootaxa

... Nel et al. (2006) described Myopsocus arthuri from Dominican amber based on a single male specimen. We herein describe the female of this species, give a species diagnosis (missing in the original description) and recognize additional characters. ...

The First Fossil Myopsocidae (Psocoptera) In Dominican Amber

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A. Waller

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Jr

... La présente étude exclut la faune entomologique de Madagascar qui, du fait de ses affi nités multiples, indo-australiennes et afrotropicales, a une histoire propre laquelle a été longuement traité au cours de colloques (Lourenço 1996 ;Vachon 1953), à travers les très nombreuses faunes publiées dans la collection « Faunes de Madagascar » et dans de très nombreuses publications (Noyes & Prinsloo 1998 ;Jamet et al. 1999;Betsch et al. 2000 ;Lecoq 2000 ;Gibon 2006 ;Gibon & Randriamasimanana 2007 ;Le Goff et al. 2007). ...

Dermaptera of Madagascar. Ecological and historical factors of their distribution
  • Citing Article
  • April 2000

Annales de la Société entomologique de France (N S )

... The new damselfly belongs to the family Calopterygidae, an ancient group (diversifying during the Cretaceous after Dumont et al., 2005), but with a rather reduced and strictly Cenozoic fossil record, viz. Sinocalopteryx shangyongensis Lin et al., 2010 from the lowermost Eocene of China; two calopterygid nymphs from the middle Eocene of Baltic amber (Bechly & Wichard, 2008;Fleck et al., 2009); Sapho armissani Nel, 1987 from the Oligocene of Armissan (France); Sapho sannoisiensis (Meunier, 1914) and Sapho legrandi Nel & Petrulevičius, 2010 from the Oligocene of Aix-en-Provence (France); Calopteryx andancensis Nel & Brisac, 1994 from the upper Miocene of Ardèche (France), and a 'Calopterygidae (?) (Species G)' described and figured by Gentilini & Bagli (2004) from the upper Miocene of Monte Castellaro (Italy). ...

The oldest Calopterygidae in the Eocene Baltic amber (Odonata: Zygoptera)
  • Citing Article
  • January 2009

Zootaxa

... Little is known about the biology of extant evaniids, but their larvae are considered predators of cockroach eggs in oothecae (Huben, 1995). The Evaniidae have a good fossil record with the oldest species known from the late Hauterivian, numerous species documented up to the Miocene, and an important diversity in Burmese amber (e.g., Rasnitsyn et al., 1998;Nel et al., 2002;Deans et al., 2004;Jennings et al., 2012;Shih et al., 2020). Nevertheless, their past diversity is still underestimated. ...

An ensign-fly in the lowermost Eocene amber from the Paris basin (Hymenoptera: Evaniidae)
  • Citing Article
  • July 2002

Annales de la Société entomologique de France (N S )

... In other ambers, only Dryinus vetustus Olmi, 1995 (uncertain group), from Dominican amber, and Dryinus balticus (Olmi, 1984) 1.8 mm)). In the unique fossil species of Pseudodryinus known by females, P. parisiensis Peinado, Nel & Waller, 2006, from Oise amber, the size is not small (4.0 mm). Overall, in Burmese amber, species of Dryinus and Hybristodryinus represented exclusively by small females are minority (2 of 10 species of Dryinus known by females (20%); 3 of 11 species known by females in Hybristodryinus (27%)). ...

A dryinid wasp in Early Eocene amber from the Paris basin (Hymenoptera: Dryinidae)
  • Citing Article
  • April 2006

Zootaxa

... Among the remaining three extant genera, Sisyrina may have closer relationship with Sisyborina than Sisyra by the similar feature of gradate crossveins (i.e., seven subapical gradates present, and most of them interrupted centrally by a hyaline spot). Although the presence of gradate crossveins may be plesiomorphic in Sisyridae as many fossil genera possess this character state (Nel et al. 2003;Wichard et al. 2016;Yang et al. 2018), the peculiar interruption on many gradate crossveins probably stands to be apomorphic, suggesting the sister-group relationship between Sisyrina and Sisyborina. Besides, among the fossil genera that definitely belong to Sisyridae (Wichard et al. 2016;Lu & Liu 2021), i.e., Paleosisyra Nel et al., 2003, Prosisyrina Perkovsky & Makarkin, 2015, and Stictosisyra Yang et al., 2018, Sisyrina resembles Paleosisyra by having similar mouthpart characters (terminal segment of maxillary and labial palpus triangular in lateral view), wing venation (subapical gradates present) and male genitalia (sternum 9 short and simple) (Nel et al. 2003;Perkovsky & Makarkin 2015;Wichard et al. 2016;Yang et al. 2018). ...

New fossil spongilla-flies from the lowermost Eocene amber of France (Insecta, Neuroptera, Sisyridae)
  • Citing Article
  • April 2003

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