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... The study area overlaps with two ecoregions: the Espinal in the midwest and the Pampas in the east of La Pampa province (Cabrera 1976;Brown et al. 2006), comprising ∼ 94.635 km 2 in central Argentina (Fig. 1). The Espinal is a xerophilic forest ecosystem characterized by the dominant tree known as caldén (Neltuma caldenia) with grassland and bush covers. ...