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Bol. Mus. Nac. Hist. Nat. Parag. Vol. 29
1680-4031/2025 Museo Nacional de Historia Natural del Paraguay. Este es un artículo de acceso abierto bajo la
licencia CC BY 4.0 (https://creativecommons.org/licenses/by/4.0/).
Editor responsable: / Recibido: / Aceptado: / Publicado online:
, Art. e2025003: 10 pp.
24.i.2023 2.iv.2025 9.iv.2025Bolívar R. Garcete-Barrett*,
*Museo Nacional de Historia Natural del Paraguay, San Lorenzo, Paraguay.
New and interesting records for longhorn beetles (Coleoptera: Cerambycidae) from
southern Paraguay
Nuevos e interesantes registros para escarabajos longicornios (Coleoptera: Cerambycidae)
del sur de Paraguay
John M. Leavengood, Jr.1,*,, Larry G. Bezark2, & Roy F. Morris, II3
1
John.M.Leavengood@usda.gov.
2521 46th Street Sacramento, California 95819, USA. E-mail: bezbycids@gmail.com.
3beetlesandbirds@gmail.com.
*Author for correspondance.
Abstract.- Twelve new country records are presented to increase our knowledge of the longhorn beetles
(Coleoptera: Cerambycidae) of Paraguay. Additional interesting records and updated distributions for
each respective species, a new plant association for Lophopoeum carinatulum
record of the genus Eupromera Westwood, 1846 from Paraguay are presented.
Key words: Neotropical Cerambycidae, Eupromera, new distribution records, biodiversity, host plant.
Resumen.- Se presentan doce nuevos registros de país para aumentar el conocimiento de los escarabajos
longicornios (Coleoptera: Cerambycidae) de Paraguay. Se presentan registros interesantes adicionales y
distribuciones actualizadas para cada especie respectiva, una nueva asociación de plantas para Lopho-
poeum carinatulum Bates, 1863, y el primer registro del género Eupromera de Paraguay.
Palabras clave: Cerambycidae Neotropical, Eupromera, nuevos registros de distribución, biodiversidad,
planta huésped.
collections of longhorn beetles were made
species of the country. To date, the longhorn
beetles of Paraguay have received consider-
able attention. Early work by Bosq (1945;
1947) and Viana (1972) laid the foundation
for the study of Cerambycidae in Paraguay.
(2004) added 96 species to the known fauna.
More recently, Bezark & Tyson (2020) in-
creased Paraguay’s known cerambycid fauna
from 774 to 863 species, and Cubilla et al.
(2021; 2023; 2024) subsequently added an-
other twelve species.
Additional work on general Neotropical
longhorn beetles has contributed to the known
longhorn fauna as well. Cerambycidae is just
one such family recently receiving compre-
by Garcete-Barrett (2016) catalogued the
Coreidae, Liu et al. (2022) produced a pre-
liminary checklist to Paraguay’s Bostrichidae,
Smith et al. (2022) catalogued Paraguay’s
Sphingidae, and Eger (pers. corr.) has been
developing a manuscript on the country’s
Pentatomidae. An annotated summary of
Paraguay’s faunal inventory project can be
found in Kochalka (2014).
As in Bezark & Tyson (2020), the speci-
mens collected represent the southern half of
the country comprising the departments of
Amambay, Canindeyú, Concepción, Guairá,
Itapúa, Misiones, Paraguarí and San Pedro.
Expeditions to the northern departments of
Alto Paraguay, Boquerón and Presidente
Hayes—together forming more than half of
the country’s landmass—should produce great
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29
2, Art. e2025003
John M. Leavengood, Jr., Larry G. Bezark & Roy F. Morris, II
contributions to Paraguay’s known beetle
fauna.
The present work adds twelve species to
the known fauna of Paraguay and presents up-
dated distributions for each respective species.
Methods
In addition to the aforementioned literature,
Bezark (2021; 2022), Monné (2024) and Ta-
vakilian and Chevillotte (2024) were used to
establish the validity of new records.
One or two specimens of each species in
suitable condition was photographed. Speci-
Nikon SMZ-18 stereomicroscope. Photograph
layers were stacked using Helicon Focus 6
(www.heliconsoft.com/heliconsoft-products/
helicon-focus) and edited using Adobe Photo-
shop Elements 12 Editor (https://www.adobe.
com/products/photoshop-elements.html),
plates.
Locality data are presented below under
“Specimens Examined.” These label data are
standardized to present country, department,
standardized date (in the form of day-month-
year as 1-II-1903), collector, depository and
number of specimens, in that order. Otherwise,
these data are presented verbatim.
Specimens examined reside in the deposi-
tories listed below, recognized by codens in
the “specimens examined” sections as follows:
IBNP—Inventorio Biologico Nacional [Museo
Nacional de Historia Natural del Paraguay],
San Lorenzo, Paraguay; JMLC—John M.
Leavengood, Jr., private collection, Brandon,
Florida; LGBC—Larry G. Bezark, private col-
lection, Sacramento, California; RFMC—Roy
(previously Lakeland, Florida).
Species are presented alphabetically by their
Results and Discussion
Subfamily Cerambycidae
Tribe Dichophyiini
Solangebira laetus (Blanchard, 1851)
(Figs. 1, 9)
Remarks: Perhaps the most surprising
record from Paraguay, the closest previ-
ously known records are from Argentina
in the provinces Neuquén, Rio Negro and
Chubut (Napp & Martins, 2002). More-
over, this variably colored species typi-
cally has some orange-reddish markings
on the pronotum, whereas the Paraguay
voucher is entirely monochromatic. Prior
to the generic reassignment in Santos-
Silva et al. (2023), this species was known
by the combinations Eryphus laetus and
Callideriphus laetus.
Distribution: Chile and Argentina (Napp
& Martins, 2002). Paraguay is a new country
record.
Specimens examined: PARAGUAY:
: San Ignacio, vic. Hotel Rural, S
26°52.508’ W 056°59.355’, elev. 451 ft, 5-8-
XII-2019, Eger, Tyson & Leavengood (JMLC,
1).
Tribe Eburiini
Eburodacrys inaequalis Galileo & Martins,
2009
(Fig. 2)
Distribution: Bolivia (Galileo & Martins,
2009). Paraguay is a new country record.
Specimens examined: PARAGUAY: -
: P.N. Paso Brayo, Carocal Station,
Tribe Elaphidiini
Pantonyssus nigriceps Bates, 1870
(Fig. 3)
Distribution: Colombia, Bolivia and Brazil
(Barros et al., 2019; Bezark, 2021). There are
also questionable records from Panama and
Guyana (Santos-Silva et al., 2019). Paraguay
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29 3
, Art. e2025003
New and interesting records for longhorn beetles from southern Paraguay
is a new country record.
Specimens examined: PARAGUAY:
: vic. Pro Cosara Nature Reserve, S
26°38.271’ W 55°39.850’, 933 ft., 9-10-XII-
2019, JE Eger & JM Leavengood (JMLC, 3;
LGBC, 1).
Figures 1–9. Species of Cerambycidae treated in this paper and probable host tree species of Lophopoeum carinatulum
Bates. 1) Solangebira laetus (Blanchard, 1851). 2) Eburodacrys inaequalis Galileo & Martins, 2009. 3) Pantonyssus
nigriceps Bates, 1870. 4) Sphalloeme costipennis Melzer, 1928. 5) Lophopoeum carinatulum Bates, 1863. 6–7) Ficus
lyrata Warburg at Hotel Independencia (Guairá). 8) Elytral humeral tubercles (or centrobasal crests) of L. carinatulum.
9) Elytral apices of E. laetus.
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29
4, Art. e2025003
John M. Leavengood, Jr., Larry G. Bezark & Roy F. Morris, II
Tribe Oemini
Sphalloeme costipennis Melzer, 1928
(Fig. 4)
Distribution: Argentina and Brazil (Galileo et
al., 2015). Monné (2024) included a dubious
(with a “?”) record from Honduras. Paraguay
is a new country record.
Specimens examined: PARAGUAY:
: P.N. Caazapá Est Cristal 9-XII-2014,
Subfamily Lamiinae
Tribe Acanthocinini
Lophopoeum carinatulum Bates, 1863
(Figs. 5-8)
Remarks:
twelve were collected from the same tree (Figs.
6-7), Ficus lyrata Warburg, that was visited
daily over the course of ten days at Hotel In-
Western Hemisphere, this tree is likely an op-
portunistic host for L. carinatulum and could
be added to its list of known hosts (Bezark et
al., 2022).
Distribution: Monné (2024) presented an
updated distribution for L. carinulatum includ-
ing Paraguay.
Specimens examined: PARAGUAY: -
: Hotel Independencia, vic. Independencia,
10-20-XII-2019, JE Eger & JM Leavengood,
617 ft, S 25°43.069’ W 56°16.443’ (JMLC,
1), Hotel Independencia, vic. Independencia,
elev. 617 ft beating Ficus lyrata Warb., S
25°43.069’ W 56°16.443’, 10-20-XII-2019 JM
Leavengood (IBNP, 2; JMLC, 8; LGBC, 2);
: vic. Hotel Gabriela, nr. Paraguarí,
S 25°35.630’ W 057°07.495’, Elevation 504
ft., 1-4-XII-2019, JE Eger & JM Leavengood
(JMLC, 2).
Urgleptes recki (Melzer, 1934)
(Fig. 10)
Distribution: Brazil (Bezark, 2021). Paraguay
is a new country record.
Specimens examined: PARAGUAY: -
: San Ignacio, vic. Hotel Rural, S 26°52.508’
W 056°59.355’, elev. 451 ft, 5-8-XII-2019, Eger,
Tyson & Leavengood (JMLC, 1).
Tribe Aerenicini
Recchia acutipennis (Gahan, 1889)
(Fig. 11)
Distribution: Argentina, Bolivia, Brazil and
Paraguay (Monné, 2024).
Specimens examined: PARAGUAY: -
: P.N. Cerra Cora, 255m, 19-20-XI-2014,
Tribe Apomecynini
Bisaltes roseiceps Breuning, 1939
(Fig. 12)
Remarks: Breuning (1971) separated B .
roseiceps from B. strandi Breuning, 1940
based on a questionable feature (the general
color of the pubescence): frons, genae, and
apical area of the elytra with pinkish pu-
bescence, leading to B. roseiceps; and these
areas without pinkish pubescence, leading to
B. strandi. Based on the photographs of the
holotypes, original descriptions (Breuning,
1939; 1940a), and redescriptions by Breuning
(1971), we believe that B. strandi is a junior
synonym of B. roseiceps. However, we prefer
not to establish synonymy until we have ex-
amined a larger number of specimens. These
Antonio Santos-Silva.
Distribution: Brazil, Bolivia and Paraguay
(Breuning, 1960; Martinez, 2000; Bezark,
2021).
Specimens examined: PARAGUAY:
: Reserva Natural del Bosque
Mbaracayu, 2-5-XII-2017, J. E. Eger (RFMC,
1).
Tribe Colobotheini
Colobothea lateralis Bates, 1865
(Fig. 13)
Distribution: Brazil (Bezark, 2021). Paraguay
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29 5
, Art. e2025003
New and interesting records for longhorn beetles from southern Paraguay
is a new country record.
Specimens examined: PARAGUAY:
: Reserva Natural del Bosque
Mbaracayu, 2-5-XII-2017, J. E. Eger (RFMC,
1).
Tribe Eupromerini
Eupromera similis Breuning, 1940
(Figs. 14, 15)
Remarks: This is the first record of the
genus Eupromera from Paraguay.
Distribution: -
lis in the Brazilian state of Rio de Janeiro
(Breuning, 1940b). It is known from Bra-
zil and Argentina (Galileo & Martins,
1995). Paraguay is a new country record.
Specimens examined: PARAGUAY:
: San Ignacio, vic. Hotel Rural,
S 26°52.508’ W 056°59.355’, elev. 451
ft, BL + MV lights, Coll: JE Eger, W Ty-
son & JM Leavengood, Jr., 5-8-XII-2019
(JMLC, 1).
Figures 10–17. Species of Cerambycidae treated in this paper. 10) Urgleptes recki (Melzer, 1934). 11) Recchia acuti-
pennis Gahan, 1889. 12) Bisaltes roseiceps Breuning, 1939. 13) Colobothea lateralis Bates, 1865. 14–15) Eupromera
similis Breuning, 1940. 16) Adesmus nevisi (Gounelle, 1909). 17) Taeniotes amazonum Thomson, 1857.
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29
6, Art. e2025003
John M. Leavengood, Jr., Larry G. Bezark & Roy F. Morris, II
Tribe Hemilophini
Adesmus nevisi (Gounelle, 1909)
(Fig. 16)
Distribution: Bolivia and Brazil (Martins
& Galileo, 2004). Paraguay is a new country
record.
Specimens examined: PARAGUAY: -
: Parque Nacional Cerro Cora, 7-9-XII-2017,
J. E. Eger (RFMC, 1).
Tribe Monochamini
Taeniotes amazonum Thomson, 1857
(Fig. 17)
Remarks: Santos-Silva & Botero (2018) did
not include Paraguay in the distribution of T.
amazonum. In most cases, it was impossible to
T. ama-
zonum. Thus, in several cases, Santos-Silva &
Botero (2018) omitted certain distributional
references. For example, raising uncertainty, it
T.
amazonum by Breuning (1943) included more
than one species. The distribution reported by
Breuning (1943 [1961 repeated this report])
for T. amazonum
Catarina et au Paraguay”) does not allow posi-
-
ing (1943) did not say if the narrow lateral
bands on pronotum are from base to apex on a
specimen by specimen basis, nor did he give
the collection locality of examined specimens
such that one could confirm vouchers. We
therefore consider the record presented herein
a new country record for Paraguay because
likely included more than one species. These
Antonio Santos-Silva.
Distribution: Argentina, Bolivia and Brazil
(Santos-Silva & Botero, 2018). Paraguay is a
new country record.
Specimens examined: PARAGUAY:
: Hotel Tirol, 13-15-XII-2017, J. E. Eger
(RFMC, 1).
Tribe Onciderini
Lochmaeocles sladeni (Gahan, 1903)
(Fig. 18)
Distribution: Brazil, Bolivia, Paraguay, Argen-
tina, and Uruguay (Monné, 2024).
Specimens examined: PARAGUAY:
Hotel Tirol, 13-15-XII-2017, J. E. Eger
(RFMC, 2).
Oncideres vicina Thomson, 1868
(Fig. 19)
Distribution: Brazil (Monné, 2024). Paraguay
is a new country record.
Specimens examined: PARAGUAY: -
: P.N. Paso Brayo, Carocal Station,
Tribe Polyrhaphidini
Polyrhaphis confusa Lane, 1978
(Fig. 20)
Distribution: Brazil and Paraguay (Bezark &
Tyson, 2020).
Specimens examined: PARAGUAY:
Hotel Tirol, 13-15-XII-2017, J. E. Eger
(RFMC, 1).
Tribe Xenofreini
Xenofrea albofasciata Galileo and Martins,
2001
(Fig. 21)
Distribution: Brazil (Galileo and Martins,
2005). Paraguay is a new country record.
Specimens examined: PARAGUAY:
Reserva Natural del Bosque
Mbaracayu, 2-5-XII-2017, J. E. Eger (RFMC,
1).
Subfamily Prioninae
Tribe Callipogonini
Ctenoscelis coeus (Perty, 1832)
(Fig. 22)
Distribution: French Guiana, Brazil and Bo-
livia (Bezark, 2021). Paraguay is a new country
record.
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29 7
, Art. e2025003
New and interesting records for longhorn beetles from southern Paraguay
Specimens examined: PARAGUAY:
: Laguna Blanca, (E Santa Rosa) 200m,
Acknowledgments
Our thanks are extended to Antonio Santos-Silva
(Museu de Zoologia da Universidade de São
Paulo, Brazil) and Eugenio Nearns (National
Museum of Natural History, Smithsonian In-
with literature. Antonio Santos-Silva also
provided helpful insights and commentary on
several species. For collecting and export per-
mits, we are most grateful to Luis Morán (for-
R. Garcete-Barrett (Curator of Entomology)
Figures 18–22. Species of Cerambycidae treated in this paper. 18) Lochmaeocles sladeni (Gahan, 1903). 19) Oncideres
vicina Thomson, 1868. 20) Polyrhaphis confusa Lane, 1978. 21) Xenofrea albofasciata Galileo and Martins, 2001. 22)
Ctenoscelis coeus (Perty, 1832).
Boletín del Museo nacional de Historia natural del Paraguay Vol. 29
8, Art. e2025003
John M. Leavengood, Jr., Larry G. Bezark & Roy F. Morris, II
(IBNP, Museo Nacional de Historia Natural del
Paraguay, San Lorenzo, Paraguay). For location
thanks extend to Carlos Aguilar Julio (IBNP
volunteer), Marcelo Cortés (IBNP volunteer)
-
tions, Gainesville, Florida, United States). And
we appreciate the specimens, support, guidance
-
Tampa, Florida) and Kevin Johns (United States
United States) for their continued support of his
Field Operations’ port reference collections.
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