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Acta Biol. Univ. Daugavp. 24(1) 2024
https://doi.org/10.59893/abud.24(1).004 ISSN 1407 - 8953
37
CALLIMETOPUS DAGTUMANUS S P. N O V., A NEW SPECIES
OF PTEROPLIINI THOMSON, 1860 (COLEOPTERA:
CERAMBYCIDAE: LAMIINAE) FROM MT. CANDALAGA
MOUNTAIN RANGE IN MARAGUSAN DAVAO DE ORO
PHILIPPINES
Milton Norman D. Medina*, Daven Jayson D. Agbas, Graden G. Obrial,
Jhonnel P. Villegas, Analyn A. Cabras
Medina M.N., Agbas D.J.D., Obrial G., Villegas J., Cabras A. 2024. Callimetopus
dagtumanus sp. nov., a new species of Pteropliini Thomson, 1860 (Coleoptera:
Cerambycidae: Lamiinae) from Mt. Candalaga mountain range in Maragusan Davao de
Oro Philippines. Acta Biol. Univ. Daugavp., 2024(1): 37-45.
Abstract
A new species of Callimetopus Blanchard, 1853, C. dagtumanus sp. nov. is described and
illustrated. This is the first species of Callimetopus in Mt. Candalaga range in Maragusan,
Davao de Oro Province, and the first for Eastern Mindanao Biodiversity Corridor,
Philippines. A short note on the species ecology and images of known Callimetopus
species in Mindanao is also presented.
Keywords: Cerambycidae, New Species, Description, Habitat, Mindanao, Philippines.
*Corresponding author: Milton Norman D. Medina. Faculty of Agriculture and Life
Sciences and URESCOM, Tropical Genomics Laboratory, Davao Oriental State
University, City of Mati, 8200 Philippines. National Museum of Natural History
Philippines, Ermita Notes on habitat specifcity and occurrence of some Philippine tger
beetles: Input to tropical biodiversity conservaton, Manila, Philippines. Email:
miltonnormanmedina@gmail.com, ORCID: https://orcid.org/0000-0001-6858-8048
Daven Jayson D. Agbas. Faculty of Computing, Data Science, and Engineering,
Information Technology Program, Davao Oriental State University, Mati City,
Philippines.
Graden G. Obrial. Faculty of Agriculture and Life Sciences, Biology Program, Davao
Oriental State University, City of Mati, 8200 Philippines.
Jhonnel P. Villegas. Faculty of Teacher Education, Davao Oriental State University, City
of Mati, 8200 Philippines. Center for Futures Thinking and Regenerative Development,
Davao Oriental State University, City of Mati, 8200 Philippines.
Analyn A. Cabras. Faculty of Agriculture and Life Sciences and URESCOM, Tropical
Genomics Laboratory, Davao Oriental State University, City of Mati, 8200 Philippines.
National Museum of Natural History Philippines, Ermita Notes on habitat specifcity and
Milton Norman D. Medina, Daven Jayson D. Agbas, Graden G. Obrial, Jhonnel P. Villegas, Analyn A. Cabras
38
occurrence of some Philippine tger beetles: Input to tropical biodiversity conservaton,
Manila, Philippines. ORCID: https://orcid.org/0000-0002-0980-1651
INTRODUCTION
The taxonomy of Philippine Cerambycidae (or
long-horned beetles) has recently gained
momentum through the works of both foreign
and Philippine taxonomists. Within the last
decade, several species of cerambycids have
been added to Philippine fauna with the
majority of the species from the sub-family
Lamiinae (flat-faced long-horned beetles)
(Barševskis 2015, Barševskis 2016a,
Barševskis 2018, Barševskis et al. 2020a,
Barševskis et al 2020b, Barševskis et al 2022,
Barševskis & Medina 2021, Barševskis &
Medina 2023, Medina et al., 2021a, Medina et
al 2021b, Medina et al 2022a, Vitali 2019,
Vitali & Medina 2023, Vives 2009, Vives
2012a, Vives 2012b, Vives 2013, Vives 2015).
The sub-family Lamiinae constitutes at least
70-75% of Philippine cerambycids (Roguet
2024, Tavakilian & Chevillotte 2023),
comprising roughly 1500 described species in
the Philippines, and this number is
continuously rising as more new species
and/or new genera are being added each year.
One of the most speciose groups of Lamiinae
in the Philippines is the tribe Pteropliini with
42 genera and roughly 226 species (Roguet
2024, Tavavilian & Chevillotte 2023). The
genus Callimetopus is one of the biggest
genera of the tribe in the Philippines with 44
species and 95% of the species are endemic to
the country. There are eight species listed for
Mindanao fauna, C. antonkozlovi Barsevskis,
2015, C. bilineatus Vives, 2015 both recorded
in Mt. Apo Range, C. cabrasae Barsevskis,
2018 recorded in Agusan del Sur, C. irroratus
(Newman, 1842) recently recorded in Davao
Oriental and Davao de Oro, C. lumawigi
Breuning, 1980 recorded in Mt. Apo Range,
C. siargaonus (Schultze, 1919) recently
collected in Mt. Balatukan, Gingoog City, C.
stanleyi Dela Cruz & Adorada, 2012 recorded
in Bukidnon, and C. tagalus (Heller, 1899)
recorded in Zamboanga, Surigao, and Davao
region (Roguet 2024, Tavakilian &
Chevillotte 2023) (Fig. 3A-H). Characters of
the genus and key to some species are
provided by Breuning (1962). Recent works
on the genus include a species of Callimetopus
endemic to Central Mindanao biogeographic
subregion, C. lumawigi Breuning, 1980 being
assessed by the IUCN as an endangered
species (Bowers et al., 2023, IUCN 2023-1) as
well as ecologic notes of the same species
being documented (Medina et al. 2022b).
Moreover, several catalogs of the genus
Callimetopus have been provided (Breuning
1962, Barsevskis 2016b, 2018).
The Eastern Mindanao Biodiversity Corridor
(EMBC) is an interesting group of mountain
ranges that covers the mountain areas of
Governor Generoso including the Mt.
Hamiguitan Range Wildlife Sanctuary
(MHRWS), the Mt. Mayo in Mati-Tarragona,
Mt. Candalaga-Kampalili Complex in Davao
de Oro and Davao Oriental, and the mountain
areas in Agusan del Sur, Agusan del Norte,
Surigao del Sur, Surigao del Norte, and
Dinagat Island. The EMBC is a group of
mountain ranges that line the eastern seaboard
of Mindanao and is one of the most
biologically rich areas in the country. Several
species of cerambycids and other beetle
groups have been discovered in this area
(Medina 2023, Medina et al., 2023a,b,c,
2024). Recently, a new species of
Callimetopus has been collected at the high
elevation range of Mt. Candalaga in
Maragusan, Davao de Oro. This new species
is the first Callimetopus described in the said
mountain range and a new species addition to
the fauna of EMBC.
MATERIAL AND METHODS
The specimens were collected using a beating
sheet method at an elevation of 1600 m.a.s.l at
the Northern slope of Mt. Candalaga Range in
Maragusan Davao de Oro. All specimens
were stored in 70% ethanol, labeled, and later
Callimetopus Dagtumanus Sp. Nov., a New Species of Pteropliini Thomson, 1860 (Coleoptera: Cerambycidae:
Lamiinae) from Mt. Candalaga Mountain Range in Maragusan Davao De Oro Philippines
39
examined at Davao Oriental State University
(DOrSU) Invertebrate Research Center, City
of Mati, Philippines. A gratuitous permit from
the Department of Environment and Natural
Ressources (DENR) XI was also secured
before collection.
Morphological characters were observed
under Leica MZ 12.5 stereomicroscope.
Habitus images were taken using Canon EOS
6D digital camera equipped with an MP-E
65mm macro lens mounted in StackShot
macro rail automated with Helicon Remote
version 4.3.0.w. All images were stacked
using Helicon Focus version 8.1.1 and
processed using a licensed Photoshop CS6
Portable software version.
Measurements of the various body parts as
follows: LB = length of body from head to
apices of clothed elytra; WH = maximum
width across head from the outer margin of a
gena to that of another; LP = length of
pronotum from base to apex along midline;
WP = maximum width across pronotum; LE =
length of elytra from level of basal margins to
apices of clothed elytra; WEH = width of
elytra at humeri.
“/” separates different lines on a label; “//”
separates different labels. All measurements
are given in millimeters (mm).
Comparative material and specimens used in
this study are deposited in the following
collections:
MMCP Milton Medina Collections, Tagum
City Philippines.
PNM Philippine National Museum,
Ermita, Philippines.
Taxonomy
Callimetopus dagtumanus Medina sp. nov.
(Fig. 1, Fig. 2)
HOLOTYPE male: PHILIPPINES –
Mindanao, Davao de Oro / Maragusan / Mt.
Candalaga iii.2023, 1500 m.a.s.l, G. Obrial, D.
Agbas leg. / MMCP, printed on red card.
PARATYPE female: same label as holotype,
MMCP. PARATYPE female: Philippines –
Mindanao, Davao de Oro / Maragusan / Mt.
Candalaga 6-12.vi.2024, 1600 m.a.s.l,
M.Medina leg. / MMCP, printed on red card.
Holotype will be deposited at the PNM.
Other materials examined: Callimetopus
irroratus (Newman, 1842) 2 males, 1 female:
Philippines – Mindanao, Davao Oriental,
Cateel, Aliwagwag Falls, i.2023, M.Medina
leg. MMCP.
Description. Dimensions Holotype: LB: 14.5
mm. WH: 3.5 mm. LP: 3.0 mm. WP: 4.0 mm.
LE: 10.0 mm. WEH: 5.0 mm. Paratype: LB:
19.0 mm. WH: 4.0 mm. 3.5 LP: mm. WP: 5.0
mm. LE: 13.0 mm. WEH: 6.5 mm.
Figure 1. Callimetopus dagtumanus sp. nov.
holotype male, habitus: A. Dorsal view, B.
Ventral view, C. Frons, D. Lateral view.
Scale = 3 mm. Photo: Milton Norman D.
Medina.
Milton Norman D. Medina, Daven Jayson D. Agbas, Graden G. Obrial, Jhonnel P. Villegas, Analyn A. Cabras
40
Figure 2. Genitalia, holotype male: A-B: whole system, A. ventral view, B. lateral view, C.
aedeagus, ventral view, D-E: tegmen, D. ventral view, E. Lateral view, F-G: VIII-X Tergites:
F. Dorsal view, G. Ventral view. Scale = 2 mm. Photo: Milton Norman D. Medina.
Adult male. Teguments head, prothorax,
elytra, abdomen, legs dull black. Eyes,
mandible, clypeus dark brownish to matte
black. Antennae scape and pedicel matte
black, antennomeres III to XI matte black with
a shade of metallic blue.
Head wider than long; frons, genae, and
clypeus densely covered with whitish
recumbent pubescence; apical margin
undulate towards the middle. Frons coarsely
punctate, finer towards vertex. Genae slightly
convex densely covered with whitish
recumbent pubescence. Lower eyelobe wider
than long, upper eyelobe semi elliptic, with
supra-orbital erect whitish setae near antennal
support. Base of clypeus lined with long erect
whitish setae; labrum with coarse punctures
towards apex covered with whitish and
yellowish recumbent setae.
Antennae: Antennae reaching apical third or
abdominal ventrite III. Scape, pedicel,
antennomere I, and basal half of antennomere
II are densely covered with long whitish
recumbent pubescence, and the rest of the
antennomeres are covered with finer whitish
pubescence, with few long semi-erect setae
mainly on the underside. Scape robust
reaching apical margin of pronotum, covered
with deep punctures; pedicel short, bell-
shaped, with few punctures. Antennomere III
longer than antennomere IV, antennomeres III
sparsely covered with fine punctures, almost
unnoticeable. Antennomeres V and VI are of
the same length, antennomeres VII to XI much
shorter than antennomere V; base of
antennomeres VI to XI glabrous; tip of
antennomere XI lined with long erect whitish
setae.
Callimetopus Dagtumanus Sp. Nov., a New Species of Pteropliini Thomson, 1860 (Coleoptera: Cerambycidae:
Lamiinae) from Mt. Candalaga Mountain Range in Maragusan Davao De Oro Philippines
41
Prothorax. Pronotum wider than long; basal
margin wavy forming a spine at each lateral
end; apical margin semi-concave. Pronotal
disc slightly raised and flattened at the middle,
densely covered with whitish recumbent
pubescence and deep punctures. Propleuron
covered with deep punctures, with blunt spine
near apical margin, densely convered with
brownish recumbent pubescence. Proventral
process ovate slightly raised, sparsely covered
with brownish recumbent pubescence.
Elytra. Elyra oblong or somewhat shield-
shaped, twice longer than wide, widest at
humeri. Margin of elytral base undulate,
slightly raised at humeri. Surface of elytra
covered with deep puncturations, finer
towards suture and near apex. Two-thirds of
elytra densely covered with very fine whitish
pubescence, apical third covered with longer
whitish and yellowish recumbent pubescence.
Basal half of elytral margin covered with
relatively thick brownish pubescence with
apical end slightly protruding towards elytral
disc; apical third of elytral margin densely
lined with long erect black setae towards apex.
Scutellum matte black, bell-shaped, covered
with whitish pubescence.
Legs. Trochanters matte black, densely cove-
red with recumbent whitish pubescence; pro-
trochanter raised, mesotrochanter slightly
raised, metatrochanter not raised. Femors
densely covered with whitish recumbent
pubescence. Tibia matte black with a shade of
metallic blue, covered with whitish pube-
scence, apical half dorsal side densely covered
with long semi-erect black setae. Tarsi
metallic blue, densely covered with whitish
pubescence. Claws matte black, glabrous.
Mesepimeron, metanepisternum, metaven-
trite, mesepisternum, and lateral sides of
abdominal ventrites densely covered with
thick brownish recumbent pubescence. Me-
sosternum, median plane of metaventrite, and
abdominal ventrites densely covered with
whitish pubescence; abdominal ventrite I
broader than ventrites II, III, IV. Pygidium
dark brown, thickly covered with long
brownish and black setae.
Genitalia. Whole system of genitalia is
generally light brownish, flat at dorsal side,
slightly recurved when viewed laterally.
Aedeagus transparent at base, darker towards
apex; apex lanceolate, and slightly recurved.
Tegmen base in peculiar mucronate shape,
body expanded at middle, parameres long
oblong shaped, apex lined with long soft
yellowish setae (Fig. 2A-B). Endophalus
shorter than the whole system of genitalia
(Fig. 2C). Base of paramere slightly raised
towards the middle (Fig. 2D-E). Tergites VIII,
IX, X (Fig. 2F-G).
Adult female. There is no sexual dimorphism
between male and female, except for size
difference, female is bigger than the male
specimen.
Differential diagnosis.
The new species closely resembles C.
irroratus but differs in the following
characters: frons, genae, and clypeus of C.
dagtumanus sp. nov. is densely covered with
whitish recumbent pubescence (vs. brownish
recumbent pubescence in C. irroratus). Apical
margin of the head of the new species is
undulated towards the middle (vs. apical
margin of the head of C. irroratus flat).
Proventral process of C. dagtumanus sp. nov.
is ovate, slightly raised, sparsely covered with
brownish recumbent pubescence (vs.
proventral process of C. irroratus bell-shaped,
slightly concave, densely covered with
brownish pubescence). Underside of the body
in C. dagtumanus sp. nov. is covered with
whitish pubescence (vs. abdominal ventrites
of C. irroratus dark brown to light brown,
covered entirely with brownish recumbent
pubescence) Elytral apex of C. dagtumanus is
obtuse or normal (vs. emarginate with pointed
suture in C. irroratus). The mucronate shape
(vs. lanceolate shape in C. irroratus) of the
base of tegmen is peculiar amongst its
Callimetopus congeners in the Philippines.
Etymology. The new species name ‘dagtu-
manus’ is from the Filipino-Bisayan term
Milton Norman D. Medina, Daven Jayson D. Agbas, Graden G. Obrial, Jhonnel P. Villegas, Analyn A. Cabras
42
‘dagtum’ meaning pitch black, referring to the
general matte black coloration of the species.
Notes on the species habitat
The new species was discovered on the
northern edge of Mount Candalaga in the
municipality of Maragusan, Davao De Oro,
along Langgawisan-Bahi Road at around 1600
m.a.s.l (Fig. 4). The locals referred to this area
as Almaciga forest since it is densely covered
with towering Agathis philippinensis Warb
(Araucariaceae), a Philippine native tree spe-
cies that is a source of resin that the locals used
as fuel. Generally, the area is characterized as
a high-elevation montane ecosystem compo-
sed of old-growth secondary forests with
relatively high humidity, and the understorey
is composed of several species of ferns,
shrubs, and flowering plants.
At present, the immediate threat to the species'
habitat is its conversion to agricultural lands.
With this, several fragments of forest clearing
are visible allowing the entrance of invasive
and exotic species and making the land area to
landslide. The newly cemented road from
Maragusan to Langawisan-Bahi also promotes
the entrance of human habitation which may
put pressure on forests for higher demand for
food, shelter, and other socio-economic
activities. The newly cemented road also poses
an easy access amongst poachers which also
contributes to the decline of species
population in the wild.
Figure 3. Callimetopus species in Mindanao Island: A. C. dagtumanus sp. nov. holotype male,
B. C. irroratus (Newman, 1842), C. C. lumawigi Breuning, 1980, D. C. siargaonus (Schultze,
1919), E. C. bilineatus Vives, 2015, F. C. cabrasae Barsevskis, 2018, G. C. stanleyi Dela Cruz &
Adorada, 2012, H. C. tagalus (Heller, 1899). Images D-H credit to A. Barsevskis. Scale = 3 mm.
Callimetopus Dagtumanus Sp. Nov., a New Species of Pteropliini Thomson, 1860 (Coleoptera: Cerambycidae:
Lamiinae) from Mt. Candalaga Mountain Range in Maragusan Davao De Oro Philippines
43
Figure 4. Species habitat in Mt. Candalaga Range, Maragusan, Davao de Oro, Philippines.
Photo: G. Obrial, 2024.
Distribution. Philippines: Mindanao (Davao
de Oro, Maragusan, Mt. Candalaga).
ACKNOWLEDGEMENTS
Our gratitude to DOrSU president Roy Ponce
for the enabling support to the Coleoptera
research team. To Vanessa Asaias and Mark
Binocal of the Center for Futures Thinking and
Regenerative Development (CFTRD) of
Davao Oriental State University for the seed
grant. To Jose Letchoncito (the DENR PAMO
of MHRWS) for processing our Gratuitous
Permit (GP) at DENR XI. To our local guides
and colleagues who helped us during the
expedition in Mt. Candalaga: Dexter Patalita,
Jessa Patalita, Eleuterio Avergonzado, Jenelyn
Japitan, Nestor “Tatay Bobong” Badilla, John
Mark Badilla, Jovil Mamparo, Rinato
Concepcion, Neshan Badilla, Erick Pescadero,
and Alexander Anichtchenko (Daugavpils
University, Latvia). Thanks to two anonymous
reviewers.
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of the genus Eczemotellus Heller from the
Philippines (Coleoptera, Cerambycidae).
Lambillionea CXXIII 1: 73–77.
Vives E. 2009. Cerambycidos nuevos o
interesantes de Filipinas (Part II)
(Coleoptera, Cerambycidae). Les Cahiers
Magellanes NS 88: 1–25.
Vives E. 2012b. New or interesting
Cerambycidae from the Philippines (Part
VI) (Coleoptera, Cerambycidae). Les
Cahiers Magellanes NS 7: 43–46.
Vives E. 2012a. New or interesting Ceram-
bycidae from the Philippines (Part V)
(Coleoptera, Cerambycidae). Les Cahiers
Magellanes NS 7: 1–70.
Vives E. 2013. New or interesting Ceram-
bycidae from the Philippines (Part VII)
(Coleoptera, Cerambycidae). Les Cahiers
Magellanes NS 11: 62–75.
Vives E. 2015. New or interesting Ceram-
bycidae from the Philippines (Coleoptera,
Cerambycidae, Lamiinae) (Part XII).
Boletín de la Sociedad Entomológica
Aragonesa 56: 49–60.
Received:24.04.2024.
Accepted: 02.08.2024.
