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https://doi.org/10.59628/jast.v2No3.898 ISSN 2958-9568 JAST
Vol.2, No.3, Page 309- 326, 2024
Recent Foraminiferal genus Quinqueloculina from
Socotra Island and Hadhramout coastlines, Yemen
Mohammed Al-Wosabi1*and Wafa’a M. Al-Qadassi2
1Sana’a University, Faculty of Petroleum and Natural Resources, Earth Sciences Department, Yemen,
2Petroleum Exploration and Production Authority, Yemen.
∗Corresponding author: mo.alwosabi@su.edu.ye
Abstract
For the purpose of etablishing a database on benthic foraminifera in Yemen, two hundred and twelve recent
benthic foraminifera species have been recorded from 32 sandy samples collected from different localities in
Socotra Island during January 2017 and Hadhramout coastline, Yemen during August 2018 from a depth of 0
- 4 m, to identify benthic foraminiferal taxa to the species level. These samples were processed to extract the
foraminiferal tests using a water saline brine based on the difference between the density of the shells and the
solution. The extracted benthic foraminifera were separated according to the type of wall into three groups; the
Agglutinated, Porcellaneous and hyaline walled foraminifera. Porcellaneous tests represent the largest number
in the collected samples of the two study areas. One hundred and four species were identified. These species
belonging to twenty-eight genera, four superfamilies and eight families. Twenty-nine species of these belong to
the genus Quinqueloculina d’Orbigny which are discussed here and illustrated. The systematic classification of
the species recorded from these samples was mentioned and compared with the type species. The previous
recordings of these species, their geographical distribution regionally and globally, and their abundance in the
study samples were tracked. The samples were deposited at the Department of Earth Sciences, Faculty of
Petroleum and Natural Resources, Sana’a University.
Article Info
Keywords:
Hadhramout coastline,Quinqueloculina, Re-
cent benthic foraminifera, Socotra Island,
Yemen.
Article History:
Received: 27-February-2024,
Revised: 16-March-2024,
Accepted: 4-May-2024,
Available online: 30 June 2024.
1. INTRODUCTION
Socotra Island and Hadhramout coastlines situated at
the Arabian Sea and Indian Ocean, which are consid-
ered warm-water areas with subtropical condition and
open currents. The Arabian Sea is recognized to host
a wide range of benthic foraminiferal morphospecies [
1
].
Some studies were carried out on foraminiferal assem-
blages in the region. For example, [
2
] identified a new
forminiferal genus from the Lias of Socotra. [
3
–
6
] studied
the foraminiferal content in the Arabian Sea, coasts of
India. [
7
]. [
8
] studied the Middle Eocene-Early Miocene
larger foraminifera from Dhofar (Oman) and Socotra Is-
land (Yemen). [
9
] studied benthic foraminifera proxies in
Mukalla coastal area, Hadhramout Governorate. [
1
] stud-
ied the benthic foraminifera from Bir Ali Beach, Shabwah
Governorates. [
10
] studied an assessment of marine
pollution with some heavy metals at Hadhramout coastal
zone. The main objective of this paper is to classification
and introduce details about the Quinqueloculina species
in the beach sand of Socotra Island and the coastline of
Hadhramout Governorate.
2. STUDY AREA
Socotra Island is the largest Yemeni Island, located be-
tween the latitudes 12° 18‘ and 12° 42‘ N and longi-
tudes 53° 19‘ and 54° 33‘E [7] (Fig. 1A). Hadhramout is
the largest governorate of the Yemen in terms of area.
It occupies about 36% of the total area of Yemen. It
is restricted between the latitudes 14° 30‘ and 14° 56‘
N and longitudes 49° 07‘ and 50° 21‘ E [
8
] (Fig. 1B).
Hadhramout coastline occupies about 303.9 km of the
Yemeni coasts [
11
] and consists of a series of sandy
beaches punctuated some rocky protruding areas that
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 309
Al-Wosabi and Al-Qadassi
mostly extends into the shallow water.
3. METHODOLOGY
Fieldwork was carried out along the northern and south-
ern coastline of Socotra Island during January 2017
and Hadhramout coastline during August 2018. Thirty-
two sand samples were collected from selected sta-
tions along the coastline of the Socotra Island and
Hadhramout coastline. These samples were collected
from a depth ranging between 0 - 4 m by hand collecting
(Fig. 1A, B). Thirteen of these samples were collected
from the north and south of Socotra Island (Table 1). The
rest samples were collected from Hadhramout coastline
(Table 2). The collected samples were prepared to sepa-
rate the foraminiferal tests from sediments. According to
[
12
], this was accomplished depending on the difference
in density between saline water and the shells, which
composed of calcium carbonate. One hundred grams of
each sample was boiled to get rid of the tissue or any
unwanted material inside the shells and then they were
washed by running distilled water using a (200-mesh)
sieve to expel the mud material. Washed samples con-
taining sand and shells were immersed in a plastic basin
filled with saline solution and then stirred. As a result
of the differences in the density of the salt liquid, sand
grains and foraminiferal shells, the shells floated on the
solution surface Tests were concentrated and extracted
from the sediments and stored in a plastic vials. Tests
were sorted according to their size using sieves of dif-
ferent sizes, then picked by double zero (00) brush and
arranged in micropaleontolgical slides. This fauna was
studied and identified by using light binocular microscope.
Photos of identified foraminiferal species were taken by
12 Mega Pixel digital camera. Some of the images of the
species were not clear enough, so they were replaced by
clear images from published researches and websites
such as www.foraminifera.net/eu site.
4. SYSTEMATIC CLASSIFICATION
Identification of benthic foraminiferal species has been
completely based on the wall composition and structure,
the external morphological features including the shape
of test, arrangement of chambers, shape of chambers,
number of whorls, number of chambers per whorl,
shape of periphery, position and form of aperture, orna-
mentation of tests and the modifications of apertures.
Twenty nine species belonging to Quinqueloculina
were recorded in two study areas. Sixteen species
were recorded in Socotra Island (Table 3), while were
recorded twenty four species in Hadhramout coastline
(Table 4). The distribution of the identified species
through the samples was determined and displayed
in Table (3 and 4). All the illustrated specimens and
sediment samples utilized for this study have been
indexed and deposited in the repository of the Earth
Sciences Department, Faculty of Petroleum and Natural
Resources, Sana’a University.
Suborder: Miliolina [13]
Superfamily: Miliolacea [14]
Superfamily: Miliolacea [14]
Family: Hauerinidae [15]
Subfamily: Haueriniae [15]
Genus: Quinqueloculina [16]
Quinqueloculina agglutinans [17]
(Pl. 1, Figs. 1, 2)
1839 Quinqueloculina agglutinans D’ORBIGNY, p.
195, pl. 12, figs. 11-13.
1994 Agglutinella agglutinans (d’Orbigny); [
18
], p. 44,
pl. 70, figs. 1-9.
2001 Agglutinella agglutinans d’Orbigny; [
19
], p. 102,
pl. 11, fig. 18.
2003 Quinqueloculina agglutinans d’Orbigny; [
20
], p.
18, fig. 4.3, 4.4.
2006 Siphonaperta agglutinans d’Orbigny; [
21
], p.
155, pl. 2, fig. 12.
2012 Quinqueloculina agglutinans d’Orbigny; [
22
], p.
119.
2012 Siphonaperta agglutinans d’Orbigny; [
23
], p. 51,
fig. 13.18, 13.19.
2013 Quinqueloculina agglutinans d’Orbigny; [
24
], p.
8, pl. 2, fig. 5.
2017 Quinqueloculina agglutinans d’Orbigny; [
1
], p.
396, fig. 4.17, 4.18.
2019 Quinqueloculina agglutinans d’Orbigny; [
25
], p.
61, fig. 17.10.
2021 Quinqueloculina agglutinans d’Orbigny; [
26
], p.
102.
Material: Specimens of this species was occurred as a
frequent form in samples HDC8, HDC11, as a common
form in sample HDC2 and as a rare form in samples
FSQ10, HDC1, HDC3A, HDC4, HDC5, HDC6, HDC9B,
HDC12B, HDC13 and HDC14.
Geographic distribution: This species was originally
described from Cuba [
17
]. Later, it was recorded from
western Mexico [
27
], Japan [
28
], New Zealand [
29
],
Sahul Shelf and Timor Sea [
18
], southwestern south
China Sea, Sunda Shelf [
19
], Bermuda [
20
], Gulf of
Iskenderun, eastern Mediterranean [
21
], Indian coast
[
5
], southwestern Pacific New Caledonia [
22
], western
Mediterranean Sea [
23
], Egyptian Red Sea coast [
24
],
Delta Shelf off Myanmar, India [
30
], Bay of Bengal, In-
dia’s east coast [
25
], Yemen [
1
], [
31
], [
32
] and Venezuela
[26].
Quinqueloculina barnardi [33]
(pl. 1, Fig. 3-5)
1971 Quinqueloculina barnardi RASHEED, p. 26, 27,
pl. 2, fig. 1.
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 310
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
Figure 1. Location map of collected samples in the two study areas; A: Socotra Island, B: Hadhramout coastline (Source: Google
Earth).
1988 Quinqueloculina barnardi Rasheed; [
34
], p. 233,
pl. 4, figs. 18-20.
1997 Quinqueloculina barnardi Rasheed; [
35
], p. 265,
fig. 3.23, 3.24.
2009 Quinqueloculina barnardi Rasheed; [
36
], p. 184,
figs. 129a-f; 130a-k.
2012 Quinqueloculina barnardi Rasheed; [
22
], p.
119.
Material: Q. barnardi Rasheed was found as an abun-
dant form in samples FSQ12, FSQ19, HDC1, HDC12B
and HDC14, as a frequent form in sample FSQ23, as a
common form in samples FSQ25, HDC3A and HDC10
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 311
Al-Wosabi and Al-Qadassi
and as a rare form in samples FSQ27, FSQ28, HDC4
and HDC15.
Geographic distribution: This species was originally
described from Coral Sea, south of Papua, New Guinea
[
33
]. It was later recorded from Papuan Lagoon, New
Guinea [
34
], western Australia [
35
], Ningaloo Reef, west-
ern Australia [
36
] and southwestern Pacific, New Cale-
donia [22].
Quinqueloculina bradyana [37]
(pl. 1, Fig. 6)
1917 Quinqueloculina bradyana CUSHMAN, p. 52, pl.
18, fig. 2a-c.
1954 Quinqueloculina bradyana Cushman; [
38
], p.
332, pl. 83, fig. 26.
1995 Quinqueloculina bradyana Cushman; [
39
]], p.
83, figs. 14, 15.
2017 Quinqueloculina bradyana Cushman; [
1
], p. 369,
fig. 4.19, 4.20.
2021 Quinqueloculina bradyana Cushman; [
26
], p.
102.
Material: In the present study, this species was found as
a rare form in samples HDC13.
Geographic distribution: This species was recorded
from Marshall Islands, south Pacific [
38
], Japan [
40
],
New Zealand [
29
], the southeast Australian coast [
39
],
Bermuda [
20
], Maldives Ridge, southeastern Arabian
Sea [
6
] and New Caledonia, southwestern Pacific [
22
],
Yemen [1] and Venezuela [26].
Quinqueloeulina disparilis [16]
(pl. 1, Figs. 7-9)
1826 Quinqueloculina disparilis D’ORBIGNY, p. 302,
n. 21.
1878 Spiroloculina disparilis Terquem; [
41
], p. 55, pl.
5(10), fig. 12.
1893 Quinqueloculina disparilis d’Orbigny; [
42
], p.
212, pl. 2, figs. 55-57.
1923 Quinqueloculina disparilis d’Orbigny; [
43
], p. 47,
pl. 6, figs. 60, 61.
1929 Quinqueloculina disparilis d’Orbigny; [
44
], p. 32,
pl. 5, fig. 4.
1958 Quinqueloculina disparilis d’Orbigny; [
45
], p.
180, pl. 4, figs. 26, 27.
1974 Quinqueloculina disparilis d’Orbigny; [
46
], p.
200, fig. 55 a-g.
1980 Quinqueloeulina disparilis d’Orbigny; [
31
], p.
151, pl. 2, figs. 4-6.
1984 Quinqueloeulina disparilis d’Orbigny; [
32
], p. 48,
pl. 1, fig. 18.
1991 Quinqueloculina disparilis d’Orbigny; [
47
], p. 36,
pl. 33, figs. 1, 2.
1993 Quinqueloculina disparilis d’Orbigny; [
48
], p.
170, pl. 8, fig. 2.
1995 Quinqueloculina disparilis d’Orbigny; [
49
], p.
168, pl. 2, fig. 2.
2004 Quinqueloculina disparilis d’Orbigny; [
50
], p.73,
pl. 10, figs. 1-3.
2012 Quinqueloculina disparilis d’Orbigny; [
23
], p. 56,
fig. 15.10-15.12.
Material: This species was occurred as a frequent form
in sample FSQ29 and as a rare form in samples FSQ10
and FSQ12.
Geographic distribution: Q.disparilis was originally
described from the Gulf of Marseilles, France [
16
]. It
was later recorded from Australia [
43
], southeastern Red
Sea shores, Yemen [
31
,
32
], Mediterranean [
47
], New
Zealand [
29
], Gulf of Naples, Italy [
48
], eastern Aegean
Sea, Turkey [
50
], south eastern Australia [
51
], and west-
ern Mediterranean Sea [23].
Quinqueloculina distorqucata [38]
(Pl. 1, Figs. 10-12)
1954 Quinquelocnlina distorqucata n. sp. CUSHMAN,
p. 333, pl. 83, fig. 27.
1980 Quinquelocnlina distorqucata Cushman; [
31
], p.
151,pl. 3, figs. 4-6.
1984 Quinquelocnlina distorqucata Cushman; [
32
], p.
49, pl. 1, fig. 10.
1997 Quinquelocnlina distorqucata Cushman; [
35
], p.
265, fig. 3.25.
Material: Q. distorqucata Cushman was found as an
abundant form in samples HDC13 and HDC14, as a
frequent form in samples FSQ19, FSQ21 and HDC2, as
a common form in samples FSQ25 and HDC6 and as
a rare form in samples FSQ4, FSQ10, FSQ12, FSQ23,
HDC9B and HDC10.
Geographic distribution: This species was originally
described as a new species from Marshall Islands,
south Pacific [
38
], southeastern Red Sea shores, Yemen
[
31
,
32
], Great Barrier Reef, Australia [
52
] and western
Australia [35].
Quinqueloculina eburnean [17]
(pl. 1, Figs. 13, 14)
1839 Triloculina eburnea D’ORBIGNY, p. 180, pl. 10,
figs. 21-23.
1987 Quinqueloculina oblonga eburnea (d’Orbigny);
[52], pl. 46, figs. 3-5.
1993 Quinqueloculina eburnea (d’Orbigny); [
53
], p.
59, pl. 53, figs. 9-11; pl. 54, figs. 1-5.
2001 Quinqueloculina eburnea (d’Orbigny); [
54
], p.
128, 131, pl. 2, figs. 18, 19.
2004 Quinqueloculina eburnea (d’Orbigny); [
50
], p.
73, pl. 10, figs. 4, 5.
Material: Q. eburnea (d’Orbigny) was occurred as a
rare form in samples FSQ19, FSQ23, FSQ29, HDC1 and
HCD8.
Geographic distribution: This species was originally
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 312
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
described from Cuba [
17
]. Later, it was recorded from
northern Great Barrier Reef, Australia [
52
], the Gulf of
Aqaba, Red Sea [
53
] and eastern Aegean Sea, Turkey
[50].
Quinqueloculina erinacea [55]
(pl. 1, Figs. 15-17)
1977 Quinqueloculina erinacea MIKHALEVICH, p.
447, fig. 4.
1983 Quinqueloculina erinacea Mikhalevich; [
56
], p.
114, fig. 203a-b.
2012 Quinqueloculina erinacea Mikhalevich; [
22
], p.
122.
Material: Q. erinacea Mikhalevich was recorded as a
frequent from in samples HDC13 and HDC14 and as a
rare form in samples HDC10 and HDC12B.
Geographic distribution: This species was recorded
from northwestern Coast of Africa [
55
] and southwestern
Pacific New Caledonia [22].
Quinqueloculina flavescens in [31]
(pl. 1, Fig. 18)
1905 Quinqueloculina flavescens D’ORBIGNY, p. 68,
pl. 4, fig. 4.
1980 Quinqueloculina flavescens d’Orbigny; [
31
], p.
151, p. 5, figs. 10-12.
1984 Quinqueloculina flavescens d’Orbigny; [
32
], p.
49, p. 1, fig. 4.
2012 Quinqueloculina flavescens d’Orbigny; [7][
7
], p.
38, pl. 2, fig. 13.
Material: Specimens of this species found as an abun-
dant form in samples HDC2 and HDC14, as a frequent
form in sample HDC8, as a common form in samples
HDC12B and HDC13 and as a rare form in sample
HDC12A.
Geographic distribution: This species was originally
described from Madagascar (d’Orbigny, 1905). It was
recorded from Yemen in southeastern Red Sea shores
[31,32] and Socotra Island [7].
Quinqueloculina granulocostata [57]
(pl. 1, Figs. 19, 20)
1946 Quinqueloculina granulocostata GERMERAAD,
p. 63, pl. 7, fig. 11, pl. 8, fig. 1.
1980 Quinqueloculina granulocostata Germeraad;
[31], p. 151, pl. 1, figs. 13-15.
1984 Quinqueloculina granulocostata Germeraad;
[32], p. 49, pl. 1, fig. 21.
1987 Quinqueloculina granulocostata Germeraad;
[52], p. 87, pl. 41, fig. 3.
1988 Quinqueloculina granulocostata Germeraad;
[34], p. 233, pl. 6, figs. 8-10.
1994 Massilina granulocostata (Germeraad); [
18
], p.
47, pl. 79, figs. 1-12.
2009 Quinqueloculina granulocostata Germeraad;
[36], p. 211, figs. 150a-k, 151a-h.
2012 Quinqueloculina granulocostata Germeraad;
[22], p. 123.
2017 Quinqueloculina granulocostata Germeraad; [
1
],
p. 397, fig. 5.2, 5.3.
Material: This species was occurred as a frequent form
in sample FSQ25 and as a rare form in samples HDC13
and HDC14.
Geographic distribution: This species was originally
described from Indonesia [
57
]. It was recorded from
northern Great Barrier Reef, Australia [
52
], Japan [
58
],
Sahul Shelf and Timor Sea [
18
], the southeast Aus-
tralian coast [
39
], Papuan Lagoon, southeast coast of
New Guinea [34], east coast of India [4], Ningaloo Reef,
western Australia [
36
] and southwestern Pacific New,
Caledonia [22].
In Yemen, this species was recorded from the southeast-
ern Red Sea [
31
,
32
] and Bir Ali Beach, Arabian Sea
[1].
Quinqueloculina jugosa [59]
(pl. 2, Fig. 1)
1944 Quinqueloculina seminulum (Linné) var. jugosa
CUSHMAN, p. 13, pl. 2, fig. 5.
1980 Quinqueloculina jugosa Cushman; [
31
], p. 152,
pl. 2, figs. 1-3.
1984 Quinqueloculina jugosa Cushman; [
32
], p. 49,
pl. 1, fig. 17.
2004 Quinqueloculina jugosa Cushman; [
50
], p. 74,
pl. 10, figs. 6, 7.
2005 Quinqueloculina jugosa Cushman; [
60
], fig.
1.16.
2012 Quinqueloculina jugosa Cushman; [
22
], p. 123.
2017 Quinqueloculina jugosa Cushman; [
1
], p. 397,
fig. 5.4, 5.5.+
Material: This species was found as a frequent form in
sample FSQ29.
Geographic distribution: The present species was de-
scribed from the recent deposits of the New England
Coast [
59
], northeastern Gulf of Mexico [
61
], Gulf of
Kalloni, Greece [
60
], Turkey [
50
,
62
] and southwestern
Pacific New Caledonia [22].
In Yemen, this species recorded from the southeastern
Red Sea shores [
31
,
32
] and Bir Ali Beach, Arabian Sea,
Yemen [1].
Quinqueloculina laevigata [63]
(pl. 2, Fig. 2)
1839 Quinqueloculina laevigata D’ORBIGNY, p. 143,
pl. 3, figs. 31-33.
1974 Quinqueloculina laevigata d’Orbigny; [
64
], p.
240, pl. 2, figs. 1-5.
1976 Quinqueloculina laevigata d’Orbigny; [
65
], p.
89.
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 313
Al-Wosabi and Al-Qadassi
1980 Quinqueloculina laevigata d’Orbigny; [
31
], p.
152, pl. 5, figs. 4-6.
1984 Quinqueloculina laevigata d’Orbigny; [
32
], p. 49,
pl. 1, fig. 2.
1991 Quinqueloculina laevigata d’Orbigny; [
47
], p. 37,
pl. 33, figs. 8-11.
2003 Quinqueloculina laevigata d’Orbigny; [
20
], p. 19,
fig. 4.11.
2004 Quinqueloculina laevigata d’Orbigny; [
50
], p. 74,
pl. 10, figs. 8, 9.
2021 Quinqueloculina laevigata Deshayes; [
26
], p.
102.
Material: Specimens of this species was occurred as
an abundant form in samples HDC11 and HDC13, as
a frequent form in samples HDC2, HDC8, HDC9B and
HDC12A and as a common form in samples HDC6 and
HDC10.
Geographic distribution: Q.laevigata was described
from the Holocene of the Canary Islands and from the
Tertiary of Paris [
63
]. It was recorded from the west-
ern Mexico [
27
], western north America [
66
], Greece
[
64
], southern California [
65
], Japan [
28
], Mediterranean
Sea [
47
], New Zealand [
29
], Bermuda [
20
], Turkey [
50
],
Gulf of Iskenderun, eastern Mediterranean [
21
], north
Portugal [67] and Venezuela [26].
In Yemen, Q. laevigata occurs from the southeastern
Red Sea shores [31,32].
Quinquelocnlina lamarckiana [17]
(Pl. 2, Figs. 3, 4)
1839 Quinqueloculina lamarckiana D’ORBIGNY, p.
189, pl. 11, figs. 14, 15.
1921 Quinqueloculina lamarckiana d’Orbigny; [
68
], p.
419, pl. 87, figs. 2, 3.
1953 Quinqueloculina lamarckiana d’Orbigny; [
69
], p.
177, pl. 2J, fig. 3.
1956 Quinqueloculina lamarckiana d’Orbigny; [
70
], p.
17, pl. 2, fig. 10.
1961 Quinqueloculina lamarckiana d’Orbigny; [
71
], p.
16, pl. 1, fig. 10.
1969 Quinqueloculina lamarckiana d’Orbigny; [
72
], p.
401.
1976 Quinqueloculina lamarckiana d’Orbigny; [
65
], p.
63.
1980 QuinquelocuLina lamarckiana d’Orbigny; [
31
], p.
152, pl. 4, figs. 13-15.
1984 QuinquelocuLina lamarckiana d’Orbigny; [
32
], p.
50, pl. 1, fig. 23.
1992 Quinqueloculina lamarckiana d’Orbigny; [
40
], p.
67, pl. 7, figs.7a, b.
2004 Quinquelocıılina lamarckiana d’Orbigny; [
50
], p.
76, pl. 10, figs. 10, 11.
2020 Quıinquelocıılina lamarckiana d’Orbigny; [
73
], p.
96, pl. III, fig. 1.
2022 Quinquelocıılina lamarckiana d’Orbigny;.[
74
], p.
67, fig. 15K–M.
Material: Specimens of this species was occurred as a
frequent form in samples HDC13 and HDC14 and as a
rare form in samples HDC2 and HDC3A.
Geographic distribution: This cosmopolitan species
was described from the recent deposits Cuba and Ja-
maica [
17
]. It was later recorded from the several parts
of the world such as California [
65
,
69
,
71
], west In-
dia [
70
], western Mexico [
27
], the Atlantic continental
shelf of U.S.A. [
72
], Japan [
40
], New Zealand [
29
], south
eastern Australia [
51
], Bermuda [
20
], Maldives Ridge,
southeastern Arabian Sea [
6
], north Portugal [
67
], Egypt
[
75
], Turkey [
50
], Yemen [
1
], [
31
,
32
], southeast coast of
India, Tamil Nadu [73] and Mauritanian shelf [74].
Quinqueloculina limbata [16]
(pl. 2, Figs. 5-7)
1826 Quinqueloculina limbata D’ORBIGNY, p. 302,
no. 20.
1923 Miliolina limbata (d’Orbigny); [
43
], p. 45, pl. 6,
fig. 51.
1991 Quinqueloculina limbata d’Orbigny; [
47
], p. 37,
pl. 34, figs. 1-5.
1994 Quinqueloculina limbata d’Orbigny; [
18
], p. 49,
pl. 78, figs. 10-12.
2002 Quinqueloculina limbata d’Orbigny; [
76
], p. 63,
pl. 1, figs. 16, 17.
2004 Quinqueloculina limbata d’Orbigny; [
50
], p. 77,
pl. 10, fig. 12.
2011 Quinqueloculina limbata (d’Orbigny); [
77
], p.
357, fig. 7.12, 7.13.
2016 Quinqueloculina cf. Q.limbata (d’Orbigny); [
78
],
p. 158, fig. 6.3.
2018 Quinqueloculina limbata (d’Orbigny); [
79
], p.
295, pl. 2, figs. 23, 24.
Material: In the study area, this species was found as an
abundant form in sample HDC13, as a frequent form in
samples HDC2, HDC9B, HDC10, HDC12A and HDC14,
as a common form in samples HDC6 and as a rare form
in samples FSQ10, FSQ27, FSQ28, HDC1, HDC8 and
HDC11.
Geographic distribution: This species was recorded
from Sahul Shelf and Timor Sea [
18
], Mediterranean [
47
],
Turkey [
50
], Gulf of Iskenderun, eastern Mediterranean
[
21
], Saudi Arabia [
77
–
79
], Egypt [
75
] and southwestern
Pacific New Caledonia [22].
Quinqueloculina mosharrafai [80]
(Pl. 2, Figs. 8, 9)
1949 QuinquelocuLina mosharrafai SAID, p. 10, pl. 1,
fig. 23.
1956 QuinquelocuLina cf. mosharrafai Said; [
70
]], p.
17, pl. 2, fig. 11.
1980 QuinquelocuLina cf. mosharrafai Said; [
31
], p.
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 314
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
153, pl. 1, figs. 16-18.
1984 QuinquelocuLina cf. mosharrafai Said; [
32
], p.
50, pl. 1, fig. 14.
2013 Quinqueloculina mosharrafai Said; [
75
], p. 294,
pl. 1, fig. 17.
2013Quinqueloculina mosharrafi Said; [
24
], p. 10, pl.
2, fig. 6.
2016 Quinqueloculina mosharrafai Said; [
78
], p. 156,
fig. 5.26.
Material: Q. mosharrafai Said was found as a common
form in sample FSQ23 and as a rare form in samples
FSQ10 and FSQ25.
Geographic distribution: This form was described from
northern Red Sea [
80
]. It was later recorded from west-
ern India [
70
], southeastern Red Sea shores, Yemen
[31,32], Egypt [75] and Saudi Arabia [78].
Quinqueloculina neostraitula [81]
(pl. 2, Figs. 10, 11)
1932 Quinqueloculina striatula Cushman; [
82
], p. 27,
pl. 7, figs. 3, 4.
1950 Quinqueloculina neostraitula THALMANN, p.
45.
1950 Quinqueloculina straitula Cushman; [
83
], p. 5,
pl.1, fig. 9.
1954 Quinqueloculina neostriatula Thalmann; [
38
], p.
333, pl. 83, fig. 28.
1987 Quinqueloculina neostriatula Thalmann; [
52
], p.
91, pl. 43, figs. 1-6.
1988 Quinqueloculina neostriatula Thalmann; [
34
], p.
234, pl. 6, figs. 22-25.
1980 Qumqueloculina neostraitula Thalmann; [
31
],p.
154, pl. 1, figs. 1-3.
1984 Quinqueloculina neostraitula Thalmann; [
32
], p.
51, pl. 1, fig. 25.
2009 Quinqueloculina neostriatula Thalmann; [
36
], p.
225, figs 162, 163.
2011 Varidentella neostriatula (Thalmann); [
77
], p.
357, figs. 7.18, 7.19.
2012 Quinqueloculina neostriatula Thalmann [Vari-
dentella neostriatula]; [22], p. 124.
Material: Specimens of this species was occurred as
a frequent form in sample HDC13 and as a rare form in
samples HCD1 and HDC14.
Geographic distribution: The present species was de-
scribed from the recent deposits of Fiji Islands, Pacific
Ocean [82] as Q. straitula. This form was also recorded
as Q. neostriatula (Thalmann) from northern Red Sea
[
83
], Marshall Islands, south Pacific [
38
], southeastern
Red Sea shores, Yemen [
31
,
32
], northern Great Barrier
Reef, Australia [
52
], Japan [
40
], Papuan Lagoon, south-
east coast of New Guinea [
34
], western Australia [
35
],
Ningaloo Reef, western Australia [
36
] and Saudi Arabia
[77].
Quinqueloculina parkeri [84]
(Pl. 2, Fig. 12)
1881 Miliolina parkeri BRADY, p. 46. 1917 Quinque-
loculina parkeri (Brady); [37], p. 50, pl. 15, fig. 3.
1932 Quinqueloculina parkeri (Brady); [
82
], p. 25, pl.
6, figs. 3, 4.
1957 Quinqueloculina parkeri (Brady); [
85
], p. 286, pl.
85, figs. 13a-14b.
1980 Quinqueloculina parkeri (Brady); [
31
], p. 154, pl.
1, figs. 10-12.
1984 Quinqueloculina parkeri (Brady); [
32
], p. 51, pl.
1, figs. 24.
2009 Quinqueloculina parkeri (Brady); [
5
], p. 25, im-
age 3.
2009 Quinqueloculina parkeri (Brady); [
6
], p. 40, pl.
8, fig. 6.
2012 Quinqueloculina parkeri (Brady); [
7
], p. 38, pl.
2, figs. 16, 17.
2012 Quinqueloculina parkeri (Brady); [22], p. 12.
2014 Quinqueloculina parkeri (Brady); [
30
], p. 131,
pl. 9, fig. 13a-c.
2017 Quinqueloculina parkeri (Brady); [
1
],p. 400,
figs. 18, 19.
2019 Quinqueloculina parkeri (Brady); [
25
], p. 63, fig.
18.7.
2021 Quinqueloculina parkeri (Brady); [26], p. 102.
Material: Specimens of this species was found a fre-
quent form in samples HDC2, HDC3A and HDC6, as a
common form in sample HDC14 and as a rare form in
sample FSQ10.
Geographic distribution: Q. parkeri (Brady) was
recorded in different regions such as Hawaii Islands
[
84
], northern Red Sea [
83
], Marshall Islands, south Pa-
cific [
38
], southern Thailand [
86
], east coast of India [
4
],
Japan [
40
], western Australia [
87
], southwestern Pacific,
New Caledonia [
22
], Delta Shelf off Myanmar, India [
30
],
Bay of Bengal, India’s east coast [
25
] and Venezuela
[26].
In Yemen: This species was recorded from the south-
eastern Red Sea shores [
31
,
32
], Socotra Island [
7
] and
Bir Ali Beach, Arabian Sea [1].
Quinqueloculina philippinensis [68]
(pl. 2, Fig. 13)
1917 Quinqueloculina reticulata [
37
], p. 55, pl. 16, fig.
1
1921 Quinqueloculina kerimbatica (Heron-Allen & Ear-
land) var. philippinensis CUSHMAN, p.438, pl. 89,
figs. 2-3, text-fig. 34.
1941 Quinqueloculina pseudoreticulata Parr; [
88
], p.
305.
1993 Pseudotriloculina philippinensis Hottinger et al.;
[53], p. 55-56, pl. 47, figs. 1-7.
1994 Quinqueloculina pseudoreticulata Parr; [
89
], p.
25, pl. 9, figs. 2-3.
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 315
Al-Wosabi and Al-Qadassi
1994 Quinqueloculina philippinensis Cushman; [
18
],
p. 50, pl. 81, figs. 1-10.
2007 Quinqueloculina pseudoreticulata Parr; [
90
], p.
18, pl. 1, fig. 8a-b.
2012 Quinqueloculina pseudoreticulata Parr; [
22
], p.
125.
2014 Quinqueloculina philippinensis Cushman; [
30
],
p. 131, pl. 9, fig. 15a-c.
2018 Quinqueloculina philippinensis Cushman; [
91
],
p. 133, pl. 4.1, figs. 40, 41.
2019 Quinqueloculina philippinensis Cushman; [
25
],
p. 65, fig. 18.9-18.11.
Material: Specimens of this species was occurred as a
rare form in sample FSQ10.
Geographic distribution: This species was recorded
the Gulf of Aqaba, Red Sea [
53
], Sahul Shelf and Timor
Sea [
18
], southwestern south China Sea, Sunda Shelf
[
19
], Ningaloo Reef, western Australia [
36
], Papua, New
Guinea [
92
], southwestern Pacific, New Caledonia [
22
],
Delta Shelf off Myanmar, India [
30
], western Australian
shelf [91] and Bay of Bengal, India’s east coast [25].
Quinqueloculina plicosa [93]
(pl. 2, Fig. 14)
1856 Quinqueloculina plicosa COSTA, p. 322, pl. 25,
fig. 2, 5, 7.
1980 Quinqueloculina plicosa Costa; [
31
], p. 154, pl.
1, figs. 7-9.
1984 Quinqueloculina plicosa Costa; [
32
], p. 51, pl. 1,
fig. 16.
2000 Quinqueloculina plicosa Costa; [
94
], p. 168, fig.
2.
2017 Quinqueloculina plicosa Costa; [
1
], p. 400, fig.
5.20-5.22.
Material: This species was occurred as a rare form in
sample HDC12B.
Geographic distribution: This species was recorded
from Italy [93], Qatar [94] and Yemen [1,31,32].
Quinqueloculina poeyana [17]
(Pl. 2, Figs. 15, 16)
1839 Quinqueloculina poeyana D’ORBIGNY, p. 191,
pl. 11, figs. 25-27.
1980 Quinqueloculina poeyana d’Orbigny; [
31
], p.
154, pl. 2, figs. 7-9.
1984 Quinqueloculina poeyana d’Orbigny; [
32
], p. 51,
pl. 1, fig. 20.
2010 Quinqueloculina poeyana d’Orbigny; [
95
], p. 6,
pl. 1, fig. 5.
2011 Quinqueloculina poeyana d’Orbigny; [
77
], p.
357, fig. 7.14, 7.15.
2012 Quinqueloculina poeyana d’Orbigny; [
7
], p. 28,
pl. 2, figs. 14, 15.
2015 Quinqueloculina poeyana d’Orbigny; [
96
], p.
141, fig. 4.h.
2017 Quinqueloculina poeyana d’Orbigny; [
1
], p. 400,
fig. 5.23-5.25.
2022 Quinqueloculina poeyana d’Orbigny; [
97
], p.
270,fig. 4.7.
Material: This species was found as a frequent form in
sample HDC11 and as a rare form in samples FSQ12,
HDC4 and HDC8.
Geographic distribution: Q. poeyana d’Orbigny previ-
ously recorded from the recent deposits of Cuba [
17
]. It
was later recorded from California [
71
], northeastern Gulf
of Mexico [
61
], western Mexico [
27
], the Atlantic conti-
nental shelf of U.S.A. [
72
], Brazil [
98
], New Zealand [
29
],
South eastern Australia [
51
], western Australia [
87
,
99
],
north Basrah-Iraq [
95
], north Portugal [
67
], Farasan Is-
land, southern Red Sea, Saudi Arabia [
77
], Oman Coast
[96] and Venezuela [26].
In Yemen, this species was recorded from the southeast-
ern Red Sea shores [
31
,
32
], Socotra Island [
7
], Bir Ali
Beach, Arabian Sea [
1
] and Al-Bawadi Island, southern
Red Sea [97].
Quinqueloculina polygona [17]
(pl. 2, Fig. 17)
1839 Quinqueloculina polygona D’ORBIGNY, p. 198,
pl. 12, figs. 21-23.
1921 Quinqueloculina polygona d’Orbigny; [
68
], p. 66,
pl. 16, figs. 3, 4.
1929 Quinqueloculina polygona d’Orbigny; [
44
], p. 28,
pl. 3, figs 5a-c.
1932 Quinqueloculina polygona d’Orbigny; [
82
], p. 25,
pl. 6, fig. 6.
1973 Quinqueloculina polygona d’Orbigny; [
100
], p.
406, pl. 6, fig. 19, 20.
1980 Quinqueloculina polygona d’Orbigny; [
31
], p.
155, pl. 4, figs. 4-6.
2012 Quinqueloculina polygona d’Orbigny; [
22
], p.
125.
2021 Quinqueloculina polygona d’Orbigny; [
26
], p.
103.
Material: Specimens of this species was occurred as a
rare form in samples HDC14 and HDC15.
Geographic distribution: The present species was de-
scribed from the recent deposits of Cuba and Jamaica
[
17
]. It was later recorded from Puerto Rico [
100
], south-
eastern Red Sea shores, Yemen [
31
,
32
], New Zealand
[
29
], Japan [
40
], southwestern Pacific, New Caledonia
[22], and Venezuela [26].
Quinqueloculina pseudoreticulata [88]
(pl. 2, Figs. 18-20)
1941 Quinqueloculina pseudoreticulata PARR, p. 305.
1994 Quinqueloculina philippinensis Cushman; [
18
],
p. 50, pl. 81, figs. 1-10.
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 316
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
2012 Quinqueloculina pseudoreticulata Parr [Quin-
queloculina variolata d’Orbigny; Quinqueloculina
philippinensis Cushman]; [22], p. 125.
2018 Quinqueloculina pseudoreticulata Parr; [
79
], p.
295, pl. 2, fig. 22.
Material: This species was found as a common form in
sample FSQ19 and as a rare form in samples FSQ21,
FSQ10 and HDC14.
Geographic distribution: This species was recorded
from Sahul Shelf and Timor Sea [
18
], Ningaloo Reef,
western Australia [
36
], Indian coast [
5
], southwestern
Pacific New Caledonia [22] and Saudi Arabia [79].
Quinqueloculina rariformis [101]
(pl. 3, Fig. 1)
1981 Quinqueloculina?rariformis MCCULLOCH, p.
51, pl. 15, fig. 12a-b.
2012 Quinqueloculina rariformis McCulloch; [
22
], p.
125.
Material: This species was recorded as a common form
in sample HDC13 and as a frequent form in sample
HDC14.
Geographic distribution: This species was recorded
from southwestern Pacific New Caledonia [22].
Quinqueloculina seminulum [102]
(pl. 3, Figs. 2-4)
1758 Serpula seminulum LINNÉ, p. 786, pl. 2, fig.
1a-c.
1956 Quinqueloculina seminulum Linné; [
70
], p. 17,
pl. 2, fig. 9.
1966 Quinqueloculina seminulum Linné; [
103
], p. 68,
pl. 7, figs. 5-7.
1969 Quinqueloculina seminulum Linné; [
72
], p. 416.
1973 Quinqueloculina seminulum Linné; [
100
], p.
391.
1976 Quinqueloculina seminulum Linné; [65], p. 89.
1980 Quinqueloculina seminula (Linné); [
31
],p. 155,
pl. 5, figs. 13-15.
1984 Quinqueloculina seminulum (Linné) [
32
], p. 54,
pl. 1, fig. 5.
2003 Quinqueloculina seminulum (Linné); [
104
], p.
17, pl. 4, figs. 11, 12.
2007 Quinqueloculina seminulum (Linné); [
90
], p. 18,
pl.1, fig. 9 a-b.
2010 Quinqueloculina seminula (Linné); [
105
], p. 101,
pl. 7, fig. 8a-c.
2012 Quinqueloculina seminula (Linné); [
23
], p. 57,
fig. 15.30, 15.31.
2012 Quinqueloculina seminula (Linné); [
22
], p. 126.
2019 Quinqueloculina seminulum (Linné); [
25
],p. 65,
fig. 18.13, 18.14.
2022 Quinqueloculina seminulum(Linné); [
97
], p. 270,
fig. 4.6a, b.
Material: This species was occurred as a frequent form
in sample HDC12B, as a common form in samples
HDC12A and HDC13 and as a rare form in samples
HDC8 and HDC15.
Geographic distribution: This cosmopolitan species
was originally described from the recent deposits of the
Adriatic Sea [
102
]. It was later recorded from the numer-
ous localities such as Belgium [
106
], India [
70
], western
Mexico [
27
], Denmark [
103
], the Atlantic continental shelf
of U.S.A. [
72
], New foundland [
107
], Puerto Rico [
100
],
southern California [
65
], Japan [
40
], New Zealand [
29
],
south eastern Australia [
51
], Marmara Sea [
108
], the
Hebridean shelf, west of Scotland [
104
], France [
109
],
north Portugal [
67
], southwestern Pacific, New Caledo-
nia [
22
], western Mediterranean Sea [
23
], south Africa
coast line [
110
], Saudi Arabia [
78
], Bay of Bengal, India’s
east coast [25], Venezuela [26] and Yemen [31,32,97].
Quinqueloculina sulcata [16]
(pl. 3, Fig. 5)
1826 Quinqueloculina sulcata D’ORBIGNY, p. 301,
no. 17.
1900 Quinqueloculina sulcata Fornasini, [
111
], p. 364,
fig. 9.
1932 Quinqueloculina sulcata d’Orbigny; [
82
], p. 28,
pl. 7, figs. 5-8.
1954 Quinqueloculina sulcata d’Orbigny;. [
38
], p. 334,
pl. 84, figs. 1, 2.
1994 Quinqueloculina sulcata d’ Orbigny; [18][
18
], p.
50, pl. 82, figs. 1-6.
2007 Quinqueloculina sulcata d’Orbigny; [
90
], p. 18,
pl. 1, fig. 10a-b.
2012 Quinqueloculina cf. Q. sulcata d’Orbigny; [
22
],
p. 127.
2019 Quinqueloculina sulcata Fornasini; [
25
], p. 65,
fig. 18.19, 18.20.
Material: This species was found as a rare form in sam-
ples HDC6, HDC13 and HDC14.
Geographic distribution: This species was recorded
from Marshall Islands, south Pacific [
38
], Saurashtra
coast, Gujarat [
90
], southwestern Pacific, New Caledonia
[22] and Bay of Bengal, India’s east coast [25].
Quinqueloculina trigonula [41]
(Pl. 3, Fig. 6)
I876 Quinquelocnlina trigonula TERQUEM, 84, pl. 12,
fig. 4.
1975 Quinquelocnlina trigonula Terquem; [
112
], p.172.
pl, figs. 10-15.
1980 Quinquelocnlina trigonula Terquem; [
31
],p. 157,
pl. 3, figs. 1-3.
1984 Quinquelocnlina trigonula Terquem; [
32
], p. 55,
pl. 1, fig. 9.
2017 Quinquelocnlina trigonula Terquem; [1], p. 402,
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 317
Al-Wosabi and Al-Qadassi
fig. 6.27, 6.28.
Material: This species was occurred as a frequent form
in samples HDC11 and HDC16, as a common form in
samples FSQ29 and HDC2 and as a rare form in sam-
ples HDC10 and HDC12B.
Geographic distribution: This species was described
from the northern shores of France [
41
]. In Yemen, it
was recorded from the southeastern Red Sea shores by
[31,32] and from Bir Ali Beach, Arabian Sea by [1].
Quinqueloculina tropicalis [113]
(Pl. 3, Fig. 7)
1924 Quinqueloculina tropicalis CUSHMAN, p. 63, pl.
23, figs. 9, 10.
1960 Quinqueloculina tropicalis Cushman; [
114
], p.
10, pl. 5, fig. 3.
1980 Quinqueloculina tropicalis Cushman; [
31
], p.
157, pl. 5, figs. 7-9.
1984 Quinqueloculina tropicalis Cushman; [
32
], p. 55,
pl. 1, fig. 3.
1994 Quinqueloculina tropicalis Cushman; [
18
], pp.
50-51, pl. 78, figs. 13-15.
1995 Quinqueloculina tropicalis Cushman; [
39
], p. 85,
figs. 170, 171, 174, 175.
2012 Quinqueloculina tropicalis Cushman; [
22
], p.
127.
2016 Quinqueloculina tropicalis Cushman; [
78
], p.
156, fig. 5.25.
Material: Specimens of this species was occurred as
an abundant form in sample HDC6, as a frequent form
in samples HDC1, HDC2, HDC5, HDC12A and HDC14
and as a common form in samples HDC3B, HDC10 and
HDC11.
Geographic distribution: [
113
] was described Q.
tropiealis from the recent deposits of Pago Pago Harbor.
It was recorded later from the numerous localities such
as the southeastern Red Sea shores, Yemen [
31
,
32
],
southeastern Australia [
39
,
51
], Papua, New Guinea [
92
],
Indian coast [
5
], southwestern Pacific New Caledonia
[22] and Saudi Arabia [78].
Quinqueloculina tubus [85]
(Pl. 3, Fig. 8)
1957 Quinqueloculina tubus TODD, p. 306, pl. 85,
fig. 18.
1988 Quinqueloculina tubus Todd; [
34
], p. 234, pl.
8, figs. 25-28.
1994 Quinqueloculina cuvieriana d’Orbigny; [
18
],
p. 48, pl. 78, figs. 4-6.
2009 Quinqueloculina tubus Todd; [
36
], p. 276,
figs. 198a-l, 199a-g, 200a-i.
2012 Quinqueloculina tubus Todd; [22], p. 128.
Material: Specimens of this species was found as a
rare form in samples FSQ4, FSQ10 and HDC14.
Geographic distribution: This species was recorded
from Saipan, Mariana Islands [
85
], Sahul shelf and
Timor Sea [
18
], Papuan Lagoon, southeast coast of
New Guinea [
34
], Ningaloo Reef, western Australia
[36] and southwestern Pacific, New Caledonia [22].
Quinqueloculina undulosecostata [115]
(Pl. 3, Fig. 9)
1882 Quinqueloculina undulosecostata
TERQUEM, p. 185, pl. 20, figs. 18, 19.
1956 Quinqueloculina undulosecostata Terquem;
[70], p. 17, pl. 2, fig. 8.
1980 Quinqueloculina undulosecostata Terquem;
[31], p. 157, pl. 1, figs. 4-6.
1984 Quinqueloculina undulosecostata Terquem;
[32], p. 55, pl. 1, fig. 15.
2017 Quinqueloculina undulosecostata Terquem;
[1], p. 402, fig. 7.2, 7.3.
Material: Specimens of this species was found as an
abundant form in sample HDC13 and as a rare form
in sample HDC4.
Geographic distribution: Q. undulosecostata was
described from the Eocene of Paris [
115
]. It was later
recorded from the recent deposits of the shores of
India [
70
] and the main Sea shores of Yemen [
1
,
31
,
32].
Quinqueloculina variolata [16]
(pl. 3, Figs. 10, 11)
1826 Quinqueloculina variolata D’ORBIGNY, p.
302, no. 26
1977 Quinqueloculina variolata d’Orbigny; [
116
] ,
p. 102, figs. 15-17.
1993 Quinqueloculina variolata d’Orbigny; [
48
], p.
180, pl. 8, fig. 1.
2012 Quinqueloculina pseudoreticulata Parr [Quin-
queloculina variolata d’Orbigny; Quinqueloculina
philippinensis Cushman]; [22], p. 125.
2016 Quinqueloculina variolata d’Orbigny; [
117
], p.
10, pl. 1, fig. 15.
Material: Specimens of this species was occurred as
a common form in sample FSQ25 and as a rare form
in samples FSQ12 and FSQ23.
Geographic distribution: This species was recorded
from Italy [
48
], southwestern Pacific, New Caledonia
[22] and Tunisia [117].
5. CONCLUSION
Thirty-two surface sand samples were collected along
the northern and southern coastlines of Socotra Is-
land and the Hadhramout coastline. These samples
were collected at a depth ranging between 0 – 4 m. to
identify the benthic foraminiferal species in these ar-
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 318
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
eas. In this paper, the systematic position of recorded
foraminiferal species is applied to identify twenty nine
Quinqueloculina species. Quinqueloculina genus as
mentioned above represent excellent indicators for
the nearshore conditions, and inner shelf environ-
ments such as the intertidal and turbulent zones, la-
goons, bays, reefal deposits, shallow open ocean
(warm water) and marshes and rare in bathyal and
deeper environments. These species were recorded
in previous studies in the main Sea shores of Yemen
as southeastern Red Sea shores and Socotra Island.
ACKNOWLEDGEMENTS
The author gratefully acknowledges American insti-
tute for Yemeni studies for its scholarship We are
thankful to Dr. Ahmed Al- Aydrus of Sana’a University,
for his help in collecting samples from Hadhramout
coastline.
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© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 321
Al-Wosabi and Al-Qadassi
Table 1: Coordinates of the studied stations, their measured environmental parameters and main type of substrates in Socotra Island.
Item Sample No. Location Latitude N Longitude E pH S (%0) T °C Position Substrate
1 FSQ4 Deleshiah beach A 12°41‘ 09.5“ 54°07‘ 53.1“ 7.22 34.3 24.8 Northen coastline Brownish fine-med. Sands
2 FSQ10 Nojahar Beach 12°38‘ 34.3“ 53°56‘ 12.6“ 5.94 34 18.4 Northen coastline Whitish medium sands
3 FSQ12 Ras Irsel 12°33‘ 12.4“ 54°30‘ 53.4“ 8.16 34 22.5 Northen coastline Whitish medium sands
4 FSQ17 Dhihamri beach 12°40‘ 16.1“ 54°11‘ 46.5“ 8.26 33.9 12.8 Northen coastline Coarse- med. sands
5 FSQ19 Detwah beach 12°41‘ 49.5“ 53°29‘ 12.7“ 7.83 34.5 26.2 Northen coastline Fine-med sands
6 FSQ20 Detwah lagoon 12°41‘ 54.7“ 53°30‘ 08.6“ 7.76 35 24.9 Northen coastline Med.-fine sands
7 FSQ21 Qadamah beach 12°41‘ 48“ 53°39‘ 24.1“ 8.26 34.4 28.2 Northen coastline Pebbly- sandy beach
8 FSQ23 Deleshiah beach B 12°40‘ 16.6“ 54°09‘ 01.3“ 8.6 34.2 28.4 Northen Sandy beach
9 FSQ25 Mahferhen- Khaisat Nahab 12°24‘ 04.2“ 54°13‘ 31.4“ 8.98 34 28.8 Southern coastline Fine sand
10 FSQ27 Haif 12°21‘ 13.2“ 54°04‘ 39.8“ 7.83 34.1 29.8 Southern coastline Semi restricted lagoon, fine sands
11 FSQ28 Haif 12°20‘ 12“ 54°04‘ 30.4“ 8.21 34 28.5 Southern coastline Open sea, med-fine sands
12 FSQ29 Awmaq 12°20‘ 40.0“ 54°01‘ 11.3“ 8.88 34.2 28.2 Southern coastline Fine – med. sands
13 FSQ30 Stero 12°19‘ 46.7“ 53°55‘ 08.0“ 8.92 34.1 28.5 Southern coastline Fine –med.sands
Table 2: Coordinates of the studied stations, their measured environmental parameters and main type of substrates in Hadhramout coastline.
Item Sample No. Location Latitude N Longitude E pH S (%0) T °C Substrate
1 HDC1 Husayhisah 14 14‘ 49.8“ 48°52‘ 23.8“ 7.52 34.2 24.8 Rocky sand beach
2 HDC2 Zalomah 14 18‘ 3.4“ 48°56‘ 14.3“ 7.98 34.1 28.2 Rocky sand beach
3 HDC3A Burum A 14°21‘ 31.3“ 48°58‘ 56.4“ 8.1 34.2 25.3 sandy beach
4 HDC3B Burum B 14°21‘ 36.2“ 48°58‘ 56.8“ 8.5 34.2 25.5 sandy beach
5 HDC4 Sixty St. Al-Shafi’i 14°27‘ 19.8“ 49°01‘ 49.4“ 8.38 34.1 26.2 sandy beach
6 HDC5 Sixty Street 14°30‘ 57“ 49°05‘ 10“ 8.6 34.3 26.9 sandy beach
7 HDC6 Al-Mukalla 14°31‘ 44.4“ 49°06‘ 36.7“ 8.18 34.2 27.4 sandy beach
8 HDC7 Al-Mihdhar Corniche 14°30‘ 37.8“ 49°09‘ 53.3“ 8.16 34 28.5 Rocky beach
9 HDC8 Al-Selail 14°32‘ 53.8“ 49°10‘ 38.7“ 8.48 34.1 28.8 Rocky beach
10 HDC9A Rawkab A 14°34‘ 2.7“ 49°12‘55.2“ 8.33 34.1 29.8 sandy beach
11 HDC9B Rawkab B 14°34‘ 5.8“ 49°12‘ 55.4“ 8.3 34.1 28.5 sandy beach
12 HDC10 Al-Shihr 1 14°45‘ 17.7“ 49°37‘ 3.0“ 8.2 34.2 29.1 sandy beach
13 HDC11 Al-Shihr 2 Al-‘Eqah 14°45‘ 29.6“ 49°37‘ 30.6“ 8.18 34.3 28.5 sandy beach
14 HDC12A Al-Hami A 14°48‘ 12.9“ 49°48‘ 31.9“ 8.4 34.3 29.5 sandy beach
15 HDC12B Al-Hami B 14°49‘ 09.4“ 49°51‘ 41.2“ 8.3 34.3 29.1 sandy beach
16 HDC13 Al-Qarn 14°49‘ 27.3“ 50°00‘ 47.8“ 7.5 34 30.5 sandy beach
17 HDC14 Sharmah 14°50‘ 55.2“ 50°01‘35.97“ 8.23 34.2 27.8 sandy beach
18 HDC15 Qusayr 14°55‘ 55.9“ 50°20‘ 18.2“ 8.04 34.1 28.1 sandy beach
19 HDC16 Ar-Raydah Al-Sharqiah 15°01‘14.16“ 50°28‘ 7.6“ 7.26 34.2 28.9 sandy beach
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 322
Recent Foraminiferal genus Quinqueloculina from Socotra Island and Hadhramout coastlines, Yemen
Table 1. Species of Quinqueloculina genus from Socotra Island. Abbreviations: R, Rare, less than 5 organisms by taxa/sample; F,
Frequent, between 6 to 10 organisms by taxa/sample; C, Common, between 11–15 organisms by taxa/sample; A, Abundant, more
than 15 organisms by taxa/sample.
Suborder Species
FSQ4
FSQ10
FSQ12
FSQ17
FSQ19
FSQ20
FSQ21
FSQ23
FSQ25
FSQ27
FSQ28
FSQ29
FSQ30
Miliolina
Quinqueloculina agglutinans R
Quinqueloculina barnardi A A F C R R
Quinqueloeulina disparilis R R F
Quinqueloculina distorqucata R R R F F R C
Quinqueloculina eburnean R R R
Quinqueloculina granulocostata F
Quinqueloculina jugosa F
Quinqueloculina philippinensis R
Quinqueloculina limbata R R R
Quinqueloculina mosharrafai R C R
Quinqueloculina parkeri R
Quinqueloculina poeyana R
Quinqueloculina pseudoreticulata R C R
Quinqueloculina trigonula R C
Quinqueloculina tubus R R
Quinqueloculina variolata R R C
Table 2. Species of Quinqueloculina genus from Hadhramout coastline. Abbreviations: R, Rare, less than 5 organisms by
taxa/sample; F, Frequent, between 6 to 10 organisms by taxa/sample; C, Common, between 11–15 organisms by taxa/sample; A,
Abundant, more than 15 organisms by taxa/sample.
Suborder Species
HDC1
HDC2
HDC3A
HDC3B
HDC4
HDC5
HDC6
HDC7
HDC8
HDC9A
HDC9B
HDC10
HDC11
HDC12A
HDC12B
HDC13
HDC14
HDC15
HDC16
Miliolina
Quinqueloculina agglutinans R C R R R R F R R F R R R
Quinqueloculina barnardi A C R F C A A R
Quinqueloculina bradyana R
Quinqueloculina distorqucata F C R R A A
Quinqueloculina eburnean R R
Quinqueloculina erinacea R R F F
Quinqueloculina flavescens A F C R C C A
Quinqueloculina granulocostata R R
Quinqueloculina laevigata F C F F C A F C A
Quinqueloculina lamarckiana R R F
Quinqueloculina limbata R F C R F F R F A F
Quinqueloculina neostraitula R F R
Quinqueloculina parkeri F F F C
Quinqueloculina plicosa R
Quinqueloculina poeyana R R F
Quinqueloculina polygona R R
Quinqueloculina pseudoreticulata R
Quinqueloculina rariformis C F
Quinqueloculina seminula R C F C R
Quinqueloculina sulcata R R R
Quinqueloculina trigonula C R F R A F
Quinqueloculina tropicalis F F C F A C C F F
Quinqueloculina tubus R
Quinqueloculina undulosecostata R A
© 2024 JAST Sana’a University Journal of Applied Sciences and Technology 323
