Taxonomic notes on longhorned beetles with the descriptions of several new taxa (Coleoptera, Cerambycidae)

Abstract

Many newly published taxonomy modifications are discussed. Several taxonomy news is proposed.

Full text

Humanity space International almanac VOL. 13, No 1, 2024: 21-38
21
http://zoobank.org/urn:lsid:zoobank.org:pub:E5021D95-8238-4F33-A9F6-0BBE908A2D20
DOI: 10.24412/2226-0773-2024-13-1-21-38
EDN: VDXIRF
Taxonomic notes on longhorned beetles with the descriptions of
several new taxa (Coleoptera, Cerambycidae)
M.A. Lazarev
Free Economic Society of Russia, Department of Scientifics Conferences and
All-Russian Projects
Tverskaya str., 22a, Moscow, 125009, Russia
e-mail: humanityspace@gmail.com, cerambycidae@bk.ru
Key words: Coleoptera, Cerambycidae, taxonomy, zoogeography, new genus, new
subgenus, new synonyms, new rank, another subfamily added.
Abstract: Many newly published taxonomy modifications are discussed. Several
taxonomy news is proposed.
Introduction.
Many taxonomic publications on Cerambycidae are regularly
appear each year without special comments by colleagues. I propose
here alternative positions on several questionable cases. Several new
synonyms cannot be accepted. Several new names were in fact
synonyms. Certain taxonomy acts were not acceptable. Rank of six
species names was downgraded to subspecies. A new genus and a
new subgenus were proposed, as well as many new synonyms.
One old subfamily name was restored, as well as one subspecies
name. A wrong geographical record was corrected, as well as some
other wrongly published data. Rank of one old name was restored.
1. A new unacceptable tribal system of Lepturinae was proposed by
Zamoroka (2022b): Cariliini (Carilia, Acmaeops, Gaurotes,
Paragaurotes, Dinoptera, Gnathacmaeops, Cortodera); Pidoniini
(Pidonia, Fallacea), Evodiini (Evodinus, Brachyta). Lepturini
includes 6 Palaearctic genera (Anoplodera, Nivellia, Leptura,
Anastrangalia, Grammoptera, Strangalia); Stenocorini -
2 (Stenocorus, Anisorus), Rhamnusiini - 3 (Rhamnusium, Akimerus,
Enoploderes), Rhagiini - 2 (Pachyta, Rhagium).
Cortodera cannot be in one tribe with all genera around
Dinoptera; Rhamnusium, Akimerus, and Enoploderes are totally

M.A. Lazarev
22
different on larval and imaginal characters, so until better decisions it
is necessary conserve traditional system of Palaearctic Lepturinae
with 7 tribes: Xylosteini, Encyclopini, Oxymirini, Enoploderini,
Rhamnusiini, Rhagiini, Lepturini.
2. Evodinellus borealis (Gyllenhal, 1827) = Pidonia petrovi
Danilevsky, 2023a, syn. nov. on the base of the type series study.
3. Megarhagium Reitter, 1913 and Hagrium Villiers, 1978 must be
accepted as valid genera names as in Villier (1978).
4. Very different genera Brachysomida Casey, 1913 and
Pseudogaurotina Plavilstshikov, 1958 were inadequately published
by Zamoroka (2022b) as subgenera of one genus, and impossible
combination was proposed: “Brachysomida (Pseudogaurotina)
excellens”.
5. According to Zamoroka (2022b), “Evodinus borealis does not
belong to the separate genus Evodinellus Winkler, 1929” or
Evodinus LeConte, 1850 = Evodinellus Plavilstshikov, 1915. But
American Evodinus differs from Evodinellus borealis (Gyllenhal,
1827) by the position of antennal insertions, which are situated in
Evodinellus borealis in front of anterior eye margins, while in
Evodinus antennal insertions disposed behind the line connecting
anterior eye margins.
6. The names of very different genera: Acmaeops LeConte, 1850 and
Euracmaeops Danilevsky, 2014 were wrongly published by
Zamoroka (2022b) as synonyms.
7. Wrong synonyms were accepted by Zamoroka (2022a):
“Cortodera flavimana (Waltl, 1838) = C. moldovana
Danilevsky, 1995”. In fact, C. moldovana has no connection with
C. flavimana, but close to C. tibialis (Marseul, 1876), and especially
to C. tibialis rossica Danilevsky, 2001b. No evidences of the
presence of С. flavimana and C. moldovana in Ukraine exist.

M.A. Lazarev
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8. Cornumutila quadrivittata (Gebler, 1830) was wrongly recorded
for Ukraine by Zamoroka (2018, 2022a). C. quadrivittata is widely
distributed in Siberia (Lazarev, 2009) and does not penetrate to
Europe. A single record for Moscow Region was very doubtful.
Only C. lineata (Letzner, 1844) is distributed in Europe.
9. Pedostrangalia revestita corsica Vartanis, 2024b, stat. nov. was
described from France (Corsica) as a species on the bases of red
anterior legs in combination with black other legs. No other features
are observed that distinguish the new taxon from P. r. revestita.
10. The reality of two different subgenera of
Stictoleptura [Paracorymbia sensu Miroshnikov, 2021, 2016]
subgen. Batesiata Miroshnikov, 1998 and S. subgen. Pyrrholeptura
Lazarev, 2016 was not accepted by Miroshnikov (2016, 2021).
I insist on three fundamental differences: Pyrrholeptura is
characterized by red elytral color, shallow male abdominal
emargination and dense elytral punctation. The difference in color
between two subgenera can't be denied, but Miroshnikov shows
several examples of different taxa where elytral color is not
important. Such reasoning is irrelevant. He shows male abdominal
emargination in both species, but the shape of those structures in his
photos is distinctly different, as well as elytral punctation in his
photos of “P. (Batesiata) tesserula” and “P. (B.) pyrrha”.
So, Stictoleptura (Batesiata Miroshnikov, 1998) and
S. (Pyrrholeptura Lazarev, 2016) are good valid names.
11. Stictoleptura (Maculileptura Danilevsky, 2014) was not accepted
by Miroshnikov (2021) who did not see the differences of this taxon
from Paracorymbia (s. str.) Miroshnikov, 1998. But that valid name
was established by Danilevsky (2014) instead of Paracorymbia
(s. str.) «group maculicornis» Miroshnikov (1998). All characters of
the group were published by Miroshnikov (1998: 594): Last
abdominal male sternite with shallow, narrow, short, but well distinct
impression, slightly emarginated apically; hind male tibiae not
curved, with two apical spines; elytra yellowish, monochrome or
with dark apex and lateral margin; antennal joints with light bases or
with light rings.

M.A. Lazarev
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12. Several new synonyms are proposed: Xylotrechus (Xylotrechus
Chevrolat, 1860 = Fulvotrechus Zamoroka, 2021 =
Hieroglyphotrechus Zamoroka, 2021 = Igneotrechus Zamoroka,
2021 = syn. nn.), Xylotrechus Chevrolat, 1860 =
Spinotrechus Zamoroka, 2021, syn. nov.
In general Zamoroka’s publications were often based on
molecular data, but strong morphological argument were usually
ignored. For example, such cases as one tribe for so different genera
as Rhamnusium, Akimerus and Enoploderes, as well as joining of
Rutpela maculata (Poda von Neuhaus, 1761) and Stenurella nigra
(Linnaeus, 1758) inside one genus show complete absurdity of
his method.
13. Wrong synonyms proposed by Zamoroka (2021): Xylotrechus =
Rusticoclytus must be rejected, and valid name
Xylotrechus (Rusticoclytus Vives, 1977) generally accepted (Vives,
1977, 2000; Villiers, 1978, 1979; Demelt, 1982; Bílý & Mehl, 1989;
Marquet, 2001, 2015; Pesarini & Sabbadini, 2007; Sama, 2008; Löbl
& Smetana, 2010; Danilevsky, 2012, 2020; Alekseev & Maryutin,
2019; Stolbov et al., 2019; Trócoli, 2019; Gradinarov &
Sivilov, 2020; Sakalian et al., 2020 and others) must be preserved.
14. Xyloclytus was wrongly upgraded to genus rank by Zamoroka
(2021). Valid subgenus name Xylotrechus (Xyloclytus Reitter, 1913)
must be preserved.
15. Teratoclytus D.W. Zaitzev, 1937 cannot be moved to
Anaglyptini, as it was proposed by Zamoroka (2021) and must be
returned to Clytini (elytral bases without tubercles).
16. Humeromaculatus Özdikmen, 2011: 537 (type species Cerambyx
figuratus Scopoli, 1763) was introduced as a subgenus of
Chlorophorus. It was upgraded to genus level by Zamoroka (2021)
without sufficient reasons. Sparganophorus Zamoroka, 2021 (type
species Clytus diadema Motschulsky, 1854) was described for a
single species, which was placed by Özdikmen (2022) in
Ch. (Humeromaculatus), so Ch. (Humeromaculatus Özdikmen, 2011
= Sparganophorus Zamoroka, 2021, syn. nov.

M.A. Lazarev
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17. Perderomaculatus Özdikmen, 2011: 537 (type species
Cerambyx sartor Müller, 1766) was upgraded by Zanoroka (2021) to
genus level without sufficient reasons, the valid name must be
Chlorophorus (Perderomaculatus).
18. Chlorophorus (Viridiphorus Zamoroka, 2021, type species:
Callidium herbstii Brahm, 1790) = Chlorophorus (Brevenotatus
Özdikmen, 2022, type species: Clytus distinguendus Perroud, 1855),
syn. nov. as Ch. herbstii was included in Ch. (Brevenotatus).
19. New synonyms must be accepted:
Stenurella (Priscostenurella Özdikmen, 2013) =
Rutpela (Eduardvivesia Zamoroka, Trócoli, Shparyk & Semaniuk,
2022) = Rutpela (Nigromacularia Zamoroka, Trócoli, Shparyk &
Semaniuk, 2022), syn. nn.
20. Ropalopus hungaricus olympicus Vartanis, 2024c was described
as R. insubricus olympicus Vartanis, 2024c from Greece (Olympos
Mt., Pieria prov., 700-1000 m), and R. hungaricus creticus Vartanis,
2024c was described as a species from Crete.
21. Chlorophorus cannot be separated from Clytini, so Clytini
Mulsant, 1839 = Chlorophorini Zamoroka, 2021, syn. nov.
22. Three new combinations were incorrectly accepted by Zamoroka
et. al. (2022): Rutpela (Eduardvivesia) vaucheri (Bedel, 1900),
R. (Nigrostenurella) nigra (Linnaeus, 1758), R. (Nigromacularia)
septempunctata (Fabricius 1793). All three taxa must be left in
Stenurella as: S. (Priscostenurella) vaucheri, S. (P.) septempunctata
and S. (Nigrostenurella) nigra.
Several species were placed by Zamoroka et. al. (2022) in
wrong subgenera: Stenurella (Priscostenurella) jaegeri (Hummel,
1825), S. (P.) novercalis (Reitter, 1901), S. (s. str.) hybridula
(Reitter, 1902), S. (s. str.) approximans (Rosenhauer, 1856). Must be:
S. (Stenurelloides) jaegeri, S. (S.) novercalis, S. (Iberostenurella)
hybridula, S. (Crassostenurella) approximans.

M.A. Lazarev
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23. Oxypleurus nodieri Mulsant, 1839 was placed by Zamoroka
(2022a) in Atimiini without sufficient reasons. The species belongs
to Saphanini as it is generally accepted.
24. Wrong synonyms were accepted by Zamoroka (2022a):
“Tetropium fuscum (Fabricius, 1787) = T. tauricum
Shapovalov, 2007”. The holotype of T. tauricum strongly differs
from many hundreds of known T. fuscum. Up to now only very
peculiar holotype of T. tauricum is known. So, rather probably it was
just a teratic specimen of local species - T. castaneum (Linnaeus,
1758). According to Plavilstshikov (1940), T. fuscum was not
collected in Crimea. T. castaneum only was recorded for Crimean
fauna by Zagaikevich (1991: 153). In fact T. fuscum is absent in
Crimea. It was only recorded by Bartenev (2009) with question mark
on the base of WEB European Cerambycidae list of 2000 by
M.L. Danilevsky.
25. Anoplistes balcanicus Sláma, 2010 described from Bulgaria was
wrongly published as a subspecies of A. halodendri (Pallas, 1773) by
Danilevsky (2020) on the bases of the records by Muraj (1960) of
Purpuricenus ephippium for Albania. Asias ephippium was also
recorded for Bulgaria by Angelov (1995) and for Rumania by Panin,
Săvulescu (1961). But in fact, Anoplistes balcanicus Sláma, 2010
does not similar to A. halodendri because of short antenna and legs,
peculiar elytral design. So, Anoplistes balcanicus Sláma, 2010 must
be accepted as a valid species name, as it was originally introduced.
The presence of Anoplistes halodendri in Balcan area rests
rather doubtful.
26. Plagionotus detritus caucasicola Plavilstshikov, 1940 =
P. d. grecus Vartanis, 2023, syn. nov.
Plagionotus detritus caucasicola Plavilstshikov, 1940 was
described with two taxonomy rank in the original publication: as
“m.” [morpha] and as “форма” [forma]. The later makes the name
available, and it was generally accepted (Özdikmen & Turgut, 2009;
Löbl & Smetana, 2010; Danilevsky, 2010; Özdikmen, 2014; Vitali,
2016; Rapuzzi & Sama, 2018; Lazarev, 2019; Danilevsky, 2020;
Vartanis, 2023 and others).

M.A. Lazarev
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Vartanis (2023) accepted P. d. caucasicola Plavilstshikov,
1940 as a valid name, but proposed no distinguishing characters. In
fact, the Caucasian populations are just identic to Greece
populations. Populations from Peloponnesus were described by
Vartanis (2023) and I’ve got good series from Northern Greece
(Thessaly): 7 males, 8 females from Ossa Mt. and 1 male, 2 females
from near Kalabaka (Meteora). Greece specimens are also rather pale
as specimens from Caucasian populations, have same wide yellow
pronotum anteriorly, wide yellow elytral stripes, and posterior elytral
stripes are partly or totally conjugated.
Acronyms of collections:
MD - collection of M. Danilevsky (Moscow)
ML - collection of M. Lazarev (Moscow)
SM - collection of S. Murzin (Moscow)
ZMM - collection of Zoological Museum of the Moscow State University
Plagionotus detritus caucasicola Plavilstshikov, 1940
Figs. 1-3
Type locality. Russia, Republic of Adygea, Maykop environs (on the
bases of lectotype designation).
Description. Dark areas of pronotum and elytra rather light-brown;
elytra with widened transverse yellow stripes behind middle, dark
stripes in between very narrow; anterior pronotal yellow area very
wide, often joined with central transverse pronotal stripe; narrow
yellow transverse stripe behind elytral bases more or less reduced or
totally absent; body length: 14.3-17.5 mm.
P. d. detritus (Linnaeus, 1758) is characterized by dark-
brown ground color of elytra and pronotum; light central pronotal
stripe usually well developed; yellow elytral stripes usually very
narrow; body length: 10-19 mm (Plavilstshikov, 1940).
Material. Lectotype (Figs 1-2) designated by Danilevsky (2009),
published by Lazarev (2019), male (length: 14.0 mm; width:
4.2 mm) with 4 labels: 1) “Cauc. occ. bor. / Maikop / 25.V.[1]928”;
2) “ex coll. Shaposhnikov”; 3) [red] “LECTOTYPUS / Plagionotus
detritus / forma CAUCASICOLA / Plavilstshikov, 1940 /
M. Danilevsky des., 2008”; 4) [pink] “Зоомузей МГУ (Москва,

M.A. Lazarev
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РОССИЯ) / № ZMMU Col 00180 / Zool. Mus. Mosq. Univ. /
(Mosquae, ROSSIA) / ex coll. N. N. Plavilstshikov” - ZMM;
Paralectotype (Fig. 3): 1 female, Maykop, 5.6.1935 - ZMM.
Additional materials. 1 male, Krasnodar Reg., L’vovskaya,
16.6.1966 - ZMM; 1 female, Ekaterinodar, 9.7.1914, Lyutkovsky -
ZMM; 1 male, 1 female, Stavropol, V. Lutshnik - ZMM; 1 male,
Armenia, Dilizhan, 5000’, 26.7.1934, N. Plavilstshikov - ZMM;
1 female, Transcauc., Kars - ZMM; 1 male, Krasnodar,
Novoprokhladnoe, 13.6.1956-1959 - MD; 1 female, Sochi,
Lazarevskoe, 4.8.1983, A. Koval - MD; 1 male,
Krasnodar, Ubinskoe, 10.5.1976, M. Kravchenko - MD; 1 male,
Krasnodar, Ubinskoe, 2.5.1991, A. Abramov - MD; 1 male,
Krasnodar, Ubinskoe, 5.8.1976, Belov - ML; 1 male, Krasnodar,
Novoprokhladnoe, 25.5.1959 - MD;. 3 males (Fig. 4),
4 females (Fig. 5), Ossa Mt. (East), VII.2001, P. Tauzin - ML;
4 males, 4 females, with the same label - SM; 1 male, 2 females, 1-2
km N Kalabaka, Meteora, VI, 1981, M. Sláma - ML.
Distribution. Russia, North Caucasus, Georgia, Armenia,
Azerbaijan, Iran, Turkey; Greece (Thessaly, Peloponnesus); the
records for Syria (Plavilstshikov, 1940; Danilevsky, 2020; Vartanis,
2023) were most probably connected with P. detritus
africaeseptentrionalis Tippmann, 1952.
27. Dorcadion fulvum erythropterum Fischer von Waldheim, 1823 =
Dorcadion fulvum opillicum Zamoroka, 2019, syn. nov. Big
available series of the species (including series from “Opillya” -
geographic region of the Podolian Upland in Lvov Oblast, Ivano-
Frankovsk Oblast and Ternopol Oblast in western Ukraine) show a
great degree of geographical variability masking local forms.
28. Dorcadion fulvum heracles Vartanis, 2024a, stat. nov. was
described from Greece (Olympus Mt., Pieria prov.) as a species on
the bases of black first antennal joint and black anterior legs.
No other features are observed that distinguish the new taxon from
D. f. fulvum (Scopoli, 1763).

M.A. Lazarev
29
29. The new wrong synonyms were published by Zamoroka (2022a):
without analises of corresponding materials and with false statement:
“ranges of some of them completely overlap”. “Dorcadion
cinerarium cinerarium (Fabricius, 1787) = D. c. macropoides
Plavilstshikov, 1932 = D. c. zubovi Lazarev, 2011”,
“D. c. panticapaeum Plavilstshikov, 1951 = D. c. bartenevi
Lazarev, 2011 = D. c. skrylniki Lazarev, 2011 = D. c. azovense
Lazarev, 2011 = D. c. gorodinskii Danilevsky, 1996 = D. c. demidovi
Danilevsky, 2013 = D. c. mosyakini Danilevsky, 2021“; “Dorcadion
equestre (Laxmann, 1770) = D. e. vadimi Danilevsky, 2021”;
“Dorcadion holosericeum Krynicki, 1832 = D. h. ustinovi
Danilevsky, 2021”. Most probably Zamoroka did not know such
specimens, so, valid names must be preserved: D. c. macropoides
Plavilstshikov, 1932; D. c. zubovi Lazarev, 2011; D. c. bartenevi
Lazarev, 2011; D. c. skrylniki Lazarev, 2011; D. c. azovense
Lazarev, 2011; D. c. gorodinskii Danilevsky, 1996; D. c. demidovi
Danilevsky, 2013; D. c. mosyakini Danilevsky, 2021; D. e. vadimi
Danilevsky, 2021; D. h. ustinovi Danilevsky, 2021.
30. The wrong synonyms Falsomesosella truncatipennis Pic, 1944 =
F. taibaishana Lazarev, 2021 published by Lin, Weigel & Ge (2021),
can’t be accepted, as holotype of F. truncatipennis (depicted by Lin
et al., 2021) from Zhejiang (see Lin, 2015) is distinctly wider with
more elongated prothorax, besides its type area is strongly distant.
So, F. taibaishana Lazarev, 2021 is a valid name.
31. Quasimesosella ussuriensis was recorded by Danilevsky (2023b:
334) for the south of Khabarovsk Region (Listvyanaya River), but
according to personal communication by N. Anisimov (November,
2023), that record must be connected with Listvenichnaya River
from Malyi Khingan in Jewish Autonomous Republic. The record of
that species for Duchin must be connected with Dichun River south-
eastwards Radde (Jewish Autonomous Republic).
32. Paratetrops gen. nov.
Type species. Tetrops warnckei Holzschuh, 1977.
Description. Body densely covered with long thick black erect setae;
antennae extremely sick, with apical joints about as long as wide;

M.A. Lazarev
30
pronotum with smooth central stripe; elytra with rough big
punctation. A single species distributed in south Turkey is known -
Paratetrops warnckei (Holzschuh, 1977), new rank.
Etymology. close to Tetrops. Gender masculine.
33. Tetrops peterkai Scořepa, 2020 is downgraded to subspecies
rank: Tetrops praeustus peterkai Scořepa, 2020, stat. nov.;
type locality: Czech Republic, Moravia, Horní Pole environs.
According to the original description the species is distributed
in Czech Republic, Slovakia, Austria, Germany.
34. Tetrops praetermitus Sláma, 2020a is downgraded to subspecies
rank: Tetrops praeustus praetermitus Sláma, 2020, stat. nov.; type
locality. Bohemia, Lásenice. According to Sláma (2020b) the taxon
is distributed in South Bohemia only.
35. Phytoecia (Musaria) rubropunctata (Goeze 1777) was wrongly
recorded for Ukraine by Zamoroka (2022a). It is a West European
species absent in Ukraine; it does not penetrate east of Germany (Bense,
1995; Sama, 2003), but was recently recorded for Spain, France and
Italy only (Löbl & Smetana, 2010; Danilevsky, 2020). Old wrong
published records could be based on specimens of Ph. (M.) argus
(Frölich, 1793) or Ph. (M.) faldermanni (Faldermann, 1837).
36. A new subgenus Phytoecia (Danilevskia subgen. nov., type
species: Saperda molybdaena Dalman, 1817) is proposed for
4 species: Ph. (Danilevskia) molybdaena (Dalman, 1817),
Ph. (Danilevskia) uncinata (W. Redtenbacher, 1842),
Ph. (Danilevskia) tenuilinea Fairmaire, 1876 and Ph. (Danilevskia)
badenkoi Danilevsky, 1988. The new taxon is characterized by the
absence of the dense solid elytral scaly cover consisting of small
scales or very short setae. Such cover is typical for
Ph. (Opsilia Mulsant, 1862). Besides all Ph. (Danilevskia) has
unicuspid mandibulae and eyes with joined dorsal and ventral lobes.
New subgenus name is feminine. The name is dedicated to Mikhail
Leontievich Danilevsky - a specialist on Palaearctic Cerambycidae
and my constant colleague.

M.A. Lazarev
31
37. Up to now Palaearctic Cerambycidae includes 8 subfamilies
(Parandrinae, Prioninae, Lepturinae, Necydalinae, Spondylidinae,
Apatophyseinae, Cerambycinae, Lamiinae). Now another one must
be accepted: Agapanthiinae Mulsant, 1839 (type genus Agapanthia
Audinet-Serville, 1835, monobasic), which was traditionally
included in Lamiinae.
Agapanthiinae is characterized by moderately or small body
size; usually strongly elongated; parallel sided or with sides slightly
diverging posteriorly; antennae usually long, often much longer than
body, usually 12-segmented (in Pseudocalamobius -
11-segmented); frons usually sloping backwards; prothorax always
without lateral spines or tubercles; legs short; anterior coxae
spherical; claws simple, divergent, without tooth-like appendages or
denticles; metepisternae very narrow, parallel sided. Larvae with
cylindrical “C”-like curved body, without legs; head slightly
elongated, strongly prominent; abdominal ventral ampullae
partly reduced.
Agapanthiinae differ from Lamiinae by many larval
characters: “C”-like curved body, slightly elongated head, rounded
laterally, ventral ampullae partly or totally reduced.
38. Agapanthia kindermanni Pic, 1905 must be returned to the
original subspecies rank as Agapanthia dahli kindermanni Pic, 1905.
39. Agapanthia lateralis Ganglbauer, 1884 is downgraded to subspecies
rank: Agapanthia dahli lateralis Ganglbauer, 1884, stat. nov.
40. Agapanthia mutinensium Sama & Rapuzzi, 2010 is downgraded
to subspecies rank: Agapanthia dahli mutinensium Sama & Rapuzzi,
2010, stat. nov.
41. Agapanthia pustulifera Pic, 1905 is downgraded to subspecies
rank: Agapanthia dahli pustulifera Pic, 1905, stat. nov.
42. Agapanthia salviae Holzschuh, 1975 is downgraded to subspecies
rank: Agapanthia dahli salviae Holzschuh, 1975, stat. nov.

M.A. Lazarev
32
43. Agapanthia schurmanni Sama, 1979 is downgraded to subspecies
rank: Agapanthia dahli schurmanni Sama, 1979, stat. nov.
44. Agapanthia subsimplicicornis Sama & Rapuzzi, 2010 is
downgraded to subspecies rank: Agapanthia dahli subsimplicicornis
Sama & Rapuzzi, 2010, stat. nov.
45. Zaitzev D.W. (original spelling - Zaitzev 1931, 1937) was
wrongly published several times (Löbl & Smetana, 2010;
Danilevsky, 2020) as Zaitzev D.A. The original spelling was
unacceptably changed by Zamoroka (2021) to “Zajciw”.
Acknowledgements. I am very grateful to M. Danilevsky (A.N. Severtsov
Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow,
Russia), A. Gusakov (Zoological Museum of Moscow State University,
Moscow, Russia) and S. Murzin (Moscow, Russsia) for supplying me with
material for study. My special thanks to M. Danilevsky for
permanent consultations.

M.A. Lazarev
33
Figs 1-5. Plagionotus detritus caucasicola Plavilstshikov, 1940:
1. Lectotype, male; 2. Lectotype labels; 3. Paralectotype, female, Maykop,
5.6.1935; 4. Male, Ossa Mt. (East), VII.2001, P. Tauzin;
5. Female with the same label.

M.A. Lazarev
34
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Received: 20.12.2023
Accepted: 29.01.2024