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Checklist of checkered beetles (Coleoptera, Cleridae)
from Sakartvelo (Georgia)
Tukazban Iskandarova1, Shota Japarashvili1
1 Institute of Ecology, Ilia State University, Cholokashvili av. 3/5 Tbilisi, 0162, Georgia
http://zoobank.org/FF5432A4-011E-4336-B0DF-D8B0DA6C854B
Corresponding author: Tukazban Iskandarova (tukazban.iskandarova.1@iliauni.edu.ge)
Academic editor: Alexander Prokin ♦ Received: 4 July 2023 ♦ Accepted: 20 October 2023 ♦ Published: 20 November 2023
Abstract
A list of 18 species of checkered beetles is given, together with images of voucher specimens and a distribution map based on literature,
dry collections and databases.
Key words
CaBOL, fauna, Insecta, occurence data
Introduction
Checkered beetles (Coleoptera: Cleridae) are a member of
the superfamily Cleroidea, with about 3,500 species de-
scribed worldwide. Most often, checkered beetles are found
on owers in moist and sunny environments or under tree
barks (McNamara 1991). Checkered beetles mainly repre-
sent phytophagous animals, although there are also zooph-
agous forms known, and some species feed on animal car-
casses (e.g., Trichodes apiarius (Linnaeus, 1758) is known as
a scavenger of bee hives).
Knowledge of checkered beetle diversity in Georgia remains
poor. One of the rst lists of Caucasian checkered beetles pub-
lished by Radde (1899) reported four species for Georgia. Lat-
er on, Zaitsev (1915) updated the list for the country to 16
species. Lastly, Japoshvili et al. (2022) provided three species
of Cleridae from the Lagodekhi Protected Areas. So far, 18
species have been reported from Georgia (Tarkhnishvili et al.
2013). However, species distribution data as well as actual spe-
cies diversity are still very sporadic and incomplete.
Here we comprehensively summarize the existing data
for the checkerid beetle species found in Georgia and pro-
vide new occurrence data for some of the species.
Materials and mathods
Various published articles and online databases for beetles
in the Caucasus and Georgia were checked. e curated
databases GBIF (Global Biodiversity Information Facility)
(GBIF Secretariat 2023) and GBD (Georgian Biodiversity
Database) (Tarkhnishvili et al. 2013) were used for collect-
ing occurrences. Dry entomological collections of the Insti-
tute of Zoology of Ilia State University (ISUZI) were stud-
ied for old, unpublished material. Data used from GBIF
and CaBOL (Caucasus Barcode of Life) (https://ggbc.eu/
data) is cited in the given paper using unique IDs. Identi-
cation of CaBOL material was performed using the identi-
cation keys of Richter (1965) and Krivolutskaya (1992).
In Georgia, social media proved to be a useful tool for
collecting biodiversity data (Iankoshvili and Tarkhnishvili
2021). We used the Facebook page “Wildlife of Georgia” as
a complementary source (WGFB) as well as the internation-
al citizen science platform iNaturalist (https://inaturalist.
ca/) to retrieve the data. For several records, we were unable
to retrieve the collecting date as it was not specied in the
publication, nor did we have access to the actual material.
e taxonomy was validated using GBIF.
Caucasiana 2, 2023, 205–210 | DOI 10.3897/caucasiana.2.e109026
Copyright Iskandarova and Japarashvili. is is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use,
distribution, and reproduction in any medium, provided the original author and source are credited.
Ilia State University, Tbilisi
Iskandarova and Japarashvili: Checkered Beetles
206
e locations for the beetles recorded in older articles are
not precise, in which case the sampling locations were geo-
referenced using Google Earth Pro (v.7.3.6) with varying
accuracy (Fig. 1).
Abbreviations for collecting methods are as follows: HC
= Hand Collection; SwN = Sweeping Net. e genera and
species in the checklist are listed alphabetically. Taxonomy is
according to the Catalogue of Palaearctic Coleoptera (Löbl
and Rolcik 2007). For species with vague occurrence data,
we used “location is not dened”.
Results
Family Cleridae
Genus Clerus Geoffroy, 1762
1. Clerus mutillarius Fabricius, 1775
GEORGIA • Location is not dened (Löbl and Rolcik 2007).
Genus Denops Fischer Von Waldheim, 1829
2. Denops albofasciatus (Charpentier, 1825)
GEORGIA • Surami (Shida Kartli region); N42.02°,
E43.55°; Zaitsev (1915).
Genus Dermestoides Houlbert & Bétis, 1905
3. Dermestoides sanguinicollis (Fabricius, 1787)
GEORGIA • Location is not dened (Löbl and Rolcik 2007).
Genus Korynetes Herbst, 1792
4. Korynetes caeruleus (De Geer, 1775)
GEORGIA • Borjomi (Samtskhe-Javakheti region);
N41.83°, E43.38°; Radde 1899. • Manglisi (Kvemo Kartli
region); N41.70°, E44.37°; Zaitsev 1915. • 2 inds; Atskuri
(Samtskhe-Javakheti region); N41.73°, E43.16°; 11 June
1895; Zaitsev 1915. • 3 inds; Bakuriani (Samtskhe-Ja-
vakheti region); N41.74°, E43.53°; June 1910; Zaitsev
1915. • 1 ind; Telavi (Kakheti region); N41.91°, E45.47°;
April 1907; Zaitsev 1915.
Genus Necrobia Olivier,
5. Necrobia rupes (Fabricius, 1781)
GEORGIA • 8 inds; Tbilisi; N41.72°, E44.79°; Septem-
ber-October 1904; Zaitsev 1915.
6. Necrobia violacea (Linnaeus, 1758)
GEORGIA • 15 inds; Telavi (Kakheti region); N41.91°,
E45.47°; March 1907; Zaitsev 1915. • 2 inds; Tbilisi;
N41.72°, E44.79°; 12 April 1905; Zaitsev 1915. • Mtskhe-
ta (Mtskheta-Mtianeti region); N41.84°, E44.71°; 19 May
1905; Zaitsev 1915. • Tana (Ateni gorge, Shida Kartli re-
gion); N41.89°, E44.01°; Zaitsev 1915. • 1 ind; Saghamo
lake (Samtskhe-Javakheti region); N41.2941°, E43.7309°;
2006 m a.s.l.; 11 October 2021; leg. A. Seropian; Japarash-
vili et al. 2023.
Figure 1. A map of Georgia shows the localities (coordinates are given in the check-list under each of the listed species) where Cleridae
materials are collected (ESRI Satellite, ArcGIS/World_Imagery).
Caucasiana 2, 2023, 205–210 207
Genus Opetiopalpus Spinola, 1844
7. Opetiopalpus scutellaris (Panzer, 1797)
GEORGIA • Tbilisi; N41.72°, E44.79°; Zaitsev 1915.
Genus Opilo Latreille, 1802
8. Opilo domesticus (Sturm, 1837)
GEORGIA • Surami (Shida Kartli region); N42.02°,
E43.55°; Radde 1899.
9. Opilo mollis (Linnaeus, 1758)
Fig. 2A
GEORGIA • Surami (Shida Kartli region); N42.02°,
E43.55°; Zaitsev 1915. • Tana (Ateni gorge) (Shida Kar-
tli region); N41.89°, E44.01°; Zaitsev 1915. • Borjomi
(Samtskhe-Javakheti region); N41.83°, E43.38°; 8 June
1910; Zaitsev 1915. • 3 inds; Lagodekhi (Kakheti region);
N41.8715°, E46.3115°; 1351 m a.s.l.; 4-14 June 2014; Ja-
poshvili et al. 2022. • 1 ind; Lagodekhi (Kakheti region);
N41.8715°, E46.3115°; 1351 m a.s.l.; 4-14 June 2014; 14
Figure 2. A: Opilo mollis (Linnaeus, 1758), CaBOL-ID 1010268. B: Opilo taeniatus (Klug, 1842), CaBOL-ID 1010301. C: anasimus
formicarius (Linnaeus, 1758), CaBOL-ID 1009787. D: Trichodes punctatus (Fischer von Waldheim, 1829), specimen from Kiketi. E:
Trichodes quadriguttatus (Adams, 1817), specimen from Gori. Scale bars = 1 mm.
Iskandarova and Japarashvili: Checkered Beetles
208
June / 5 July 2014; Japoshvili et al. 2022. • 1 ind; Udzo (Ko-
jori, Tbilisi municipality); N41.6763°, E44.7009°; 1408 m
a.s.l.; 17 June 2021; HC; deciduous forest; CaBOL-ID:
CaBOL-1010268.
10. Opilo taeniatus (Klug, 1842)
Fig. 2B
GEORGIA • 3 inds; Tbilisi; N41.72°, E44.79°; 8 May
1903, 30 May 1905, 2 June 1906; Zaitsev 1915. • 1 ind;
Gori (Kvernaki ridge, Shida Kartli region); N41.9938°,
E44.1733°; 848 m a.s.l.; 30 May 2021; SwN; leg. A. Seropi-
an; CaBOL-ID: CaBOL-1010301. • 1 ind; Gori (Kvernaki
ridge, Shida Kartli region); N41.9938°, E44.1733°; 848 m
a.s.l.; 24 June 2021; HC; leg. N. Bulbulashvili; CaBOL-ID:
CaBOL-1010307.
Genus Thanasimus Spinola, 1844
11. Tarsostenus univittatus (Rossi, 1790)
GEORGIA • Tbilisi; N41.72°, E44.79°; Zaitsev 1915.
Genus Tarsostenus Latreille, 1806
12. anasimus femoralis (Zetterstedt, 1828)
Syn. Clerus rupes Brahm, 1797
GEORGIA • 1 ind; Borjomi (Samtskhe-Javakheti region);
N41.83°, E43.38°; Zaitsev 1915. • Bakuriani (Samtskhe-Ja-
vakheti region); N41.75°, E43.53°; Zaitsev 1915. • 4 inds;
Bozali (Samtskhe-Javakheti region); N41.2714°, E43.3379°;
1866 m a.s.l.; 6 June 2022; SwN; Subalpine meadow;
leg. L-G. Japaridze; CaBOL-IDs: CaBOL-1025798,
CaBOL-1025799, CaBOL-1025800, CaBOL-1025801.
13. anasimus formicarius (Linnaeus, 1758)
Fig. 2C
GEORGIA • Lomismta Mt. (Borjomi, Samtskhe-Javakhe-
ti region); N41.87°, E43.25°; Radde 1899. • Borjomi
(Samtskhe-Javakheti region); N41.83°, E43.38°; Zaitsev
1915. • Mtskheta, (Mtskheta-Mtianeti region); N41.84°,
E44.71°; Zaitsev 1915. • 4 inds; Bakuriani (Samtskhe-Ja-
vakheti region); N41.74°, E43.53°; 19 June 1910, 25 May
1911; leg. L. Młokosiewicz; May-August 1912, 1 June
1913; leg. Kozlovsky; Zaitsev 1915. • Telavi (Kakheti re-
gion); N41.91°, E45.47°; April 1907; leg. Fursov; Zaitsev
1915. • Lagodekhi (Kakheti region); N41.87°, E46.31°; 23
May 1910; leg. L. Młokosiewicz; Zaitsev 1915. • Zeda Vedi
(previously known as Umroni) (Samegrelo-Zemo Svaneti
region); N42.89°, E42.26°; 10 July 1911; Zaitsev 1915. •
1 ind; Lagodekhi (Kakheti region); N41.8715°, E46.3115°;
1351 m a.s.l.; 5-15 May 2014; Japoshvili et al. 2022. • 1
ind; Lagodekhi (Kakheti region); N41.8525°, E46.2878°;
666 m a.s.l.; 1 May / 14 June 2014; Japoshvili et al. 2022. •
1 ind; Lagodekhi (Kakheti region); N41.8715°, E46.3115°;
1351 m a.s.l.; 15-25 May 2014; Japoshvili et al. 2022. •
1 ind; Telovani (Tbilisi); N41.8009°, E44.6807°; 888 m
a.s.l.; 2 April 2022; under the bark in deciduous forest;
HC; leg. N. Bulbulashvili; CaBOL-ID 1023282. • 1 ind;
Telovani (Tbilisi); N41.8143°, E44.6938°; 990 m a.s.l.;
16 November 2020; mixed broadleaf forest, predominant-
ly Carpinus caucasica; HC; leg. A. Seropian; CaBOL-ID:
CaBOL-1009787.
Genus Tilloidea Laporte, 1833
14. Tilloidea unifasciata (Fabricius, 1787)
GEORGIA • Borjomi (Samtskhe-Javakheti region);
N41.83°, E43.38°; Zaitsev 1915.
Genus Tillus Olivier, 1790
15. Tillus elongatus (Linnaeus, 1758)
GEORGIA • Zugdidi (Samegrelo-Zemo Svaneti region);
N42.51° E41.87°; Zaitsev 1915. • Borjomi (Samtskhe-Ja-
vakheti region); N41.83°, E43.38°; Zaitsev 1915. • El-
dari stepps (Kakheti region); N41.20°, E46.32°; Zaitsev
1915. • 1 ind; Lagodekhi (Kakheti region); N41.8715°,
E46.3115°; 1351 m a.s.l.; 4-14 June 2014; Japoshvili et
al. 2022. • 1 ind; Lagodekhi (Kakheti region); N41.8715°,
E46.3115°; 1351 m a.s.l.; 15-25 June 2014; Japoshvili et
al. 2022. • 1 ind; Lagodekhi (Kakheti region); N41.8715°,
E46.3115°; 1351 m a.s.l.; 14 June / 5 July 2014; Japoshvili
et al. 2022.
Genus Trichodes Herbst, 1792
16. Trichodes apiarius (Linnaeus, 1758)
GEORGIA • Borjomi (Samtskhe-Javakheti region);
N41.83°, E43.38°; Zaitsev 1915. • Bakuriani (Samtskhe-Ja-
vakheti region); N41.7510°, E43.5299°; Zaitsev 1915.
• Tbilisi; N41.72°, E44.79°; June 1912; Zaitsev 1915.
• Gvirgvina Mt. (Samtskhe-Javakheti region); N41.82°,
E43.57°; 20 July 1896; leg. Satunin; Zaitsev 1915. • 2
ind; Bakuriani (Samtskhe-Javakheti region); N41.74°,
E43.53°; June 1910, July 1911; leg. Satunin; Zaitsev 1915.
• Pasanauri (Mtskheta-Mtianeti region); N42.35°, E44.69°;
July 1913; leg. Satunin; Zaitsev 1915. • Mtskheta, Ksani
(Mtskheta-Mtianeti region); N41.88°, E44.57°; June 1913;
leg. Satunin; Zaitsev 1915. • Borjomi, Lomismta Mt.
(Samtskhe-Javakheti region); N41.87°, E43.25°; August
1896; leg. Fursov; Zaitsev 1915. • Tana (Ateni gorge, Shida
Kartli region); N41.89°, E44.01°; July 1907; leg. Fursov;
Zaitsev 1915. • Teliani (Shida Kartli region); N41.95°,
44.28°; July 1907; leg. Fursov; Zaitsev 1915. • Lagodekhi
(Kakheti region); N41.87°, E46.31°; June 1905; leg. L.
Młokosiewicz; Zaitsev 1915. • Chikauri (Imereti region);
N42.30°, E43.31°; July 1911; Zaitsev 1915. • Zeda Vedi
(previously known as Umroni) (Samegrelo-Zemo Svaneti
region); N42.89°, E42.26°; July 1911; Zaitsev 1915. • 1
ind; Shilda (Kakheti region); N42.0246°, E45.7422°; 591
m a.s.l.; 28 July 2015; leg. G. Ianqoshvili; Tarkhnishvili et
al. 2013. • 1 ind; Mtskheta (Mtskheta-Mtianeti region);
N41.84°, E44.71°; 2 August 2020; leg. G. Nemsadze;
Tarkhnishvili et al. 2013. • Aspindza (Samtskhe-Javakhe-
ti region); N41.5778°, E43.2499°; 18 July 2016; person-
Caucasiana 2, 2023, 205–210 209
al observation by Vasikoo Beridze; WGFB. • Tashiskari
(Samtskhe-Javakheti region); N41.9536°, E43.5010°; 10
July 2015; WGFB. • Tashiskari (Samtskhe-Javakheti re-
gion); N41.9536°, E43.5010°; 10 July 2015; personal
observation by Zauri Khachidze; WGFB. • Abastumani
(Samtskhe-Javakheti region); N41.75804°, E42.8296°;
personal observation by Tatia Adamia; WGFB. • Tbilisi;
N41.6958°, E44.7403°; 13 June 2017; personal observa-
tion by Lamia Strix; WGFB. • Sartichala (Gardabani, Kve-
mo Kartli); N41.7071°, E45.1652°; 15 June 2023; personal
observation by Nika Nika; WGFB. • 1 ind; Lentekhi-Tsa-
na road (0.7 km N of Tsana; Racha-Lechkhumi and Kve-
mo Svaneti region) N42.8887°, E43.1437°; 1770 m a.s.l.;
24 July 2018; Alpine meadows; leg. B. Rulik; ZFMK-
TIS-8003896 (Zoological Research Museum Alexander
Koenig). • 1 ind; Borjomi (Samtskhe-Javakheti region);
N41.8430°, E43.2730°; 1894 m a.s.l.; 22 August 2021; leg.
User 83484 (Observation.org); GBIF ID: 3722877749. • 1
ind; Mestia (Samegrelo-Zemo Svaneti region); N43.0628°
E42.7214°; 2237 m.a.s.l.; 3 August 2021; leg. User
83484 (Observation.org); GBIF ID: 3722105898. • 2
inds; Bakuriani (Samtskhe-Javakheti region); N41.7510°
E43.5292°; 1 August 1982; leg. R. Kuresoo (TU); GBIF
ID: 3017596476, GBIF ID: 3017445492. • 1 ind; Mck-
heta (Mtskheta-Mtianeti region); N41.8412°, E44.7074°;
458 m a.s.l.; 25 August 1986; leg. W.J. Veldkamp (NMR);
GBIF ID: 2995105316. • 1 ind; Mtskheta (Mtskhe-
ta-Mtianeti region); N41.8412°, E44.7074°; 458 m a.s.l.;
21 August 1986; leg. W.J. Veldkamp (NMR); GBIF ID:
2995102323. • 1 ind; Tbilisi; N41.7317°, E44.7400°; 547
m a.s.l.; 27 November 2018; leg. pavithra1999 (iNaturalist);
GBIF ID: 2992803633. • 1 ind; Aspindza (Samtskhe-Ja-
vakheti); N41.3798°, E43.2865°; 1238 m a.s.l.; 16 June
2019; leg. F. Riegel (iNaturalist); GBIF ID: 2311394988.
• 1 ind; Baisubani (Kakheti); N41.83875°, E46.145517°;
341 m a.s.l.; 6 May 2023; leg. shota_japarashvili (iNatural-
ist); GBID ID: 4137672136. • 2 ind; Borjomi-Kharagauli
National Park, Romanov trail hut (Samtskhe-Javakheti);
N41.84055°, E43.27416°; 1901 m a.s.l.; 22 July 2006; leg.
G. Chaladze; ISUZI.
17. Trichodes punctatus Fischer von Waldheim, 1829
Fig. 2D
GEORGIA • 1 ind; Mchkheta (Jvari monastery, Mtskhe-
ta-Mtianeti region); N41.838°, E44.733°; 616 m a.s.l.; 22
May 2019; leg. M. Szewczyk; Tarkhnishvili et al. 2013. •
1 ind; Kiketi (Tbilisi municipality); N41.65°, E44.65°; 25
May 2021; leg. A. Seropian; Tarkhnishvili et al. 2013.
18. Trichodes quadriguttatus Adams, 1817
Fig. 2E
GEORGIA • Tbilisi; N41.72°, E44.79°; 12 July 1912;
Zaitsev 1915. • Mtskheta (Mtskheta-Mtianeti region);
N41.84°, E44.71°; Zaitsev 1915. • 1 ind; Gori (Kvernaki
ridge, Shida Kartli region); N42.0074°, E44.0958°; 30 May
2021; leg. A. Seropian; Tarkhnishvili et al. 2013. • 1 ind;
Lisi Lake (Tbilisi); N41.7447°, E44.7408°; 19 June 2022;
personal observation by Giorgi Makharadze; WGFB
Discussion
So far, the species-occurrence records were obtained mostly
from the central part of Georgia. Still, the surveyed area in
Georgia is very sporadic and patchy (Fig. 1). Most of the
parts of the country need to be further studied to provide
comprehensive distributional data for the members of the
Clerida family.
Two of the species given in the checklist, Clerus mutillar-
ius and Dermestoides sanguinicollis, were recorded in Geor-
gia; however, there is no exact data about their localities.
ey were included in the checklist since the Catalogue of
Palaearctic Coleoptera (Löbl and Rolcik 2007) indicates
their presence in the area. Nevertheless, further research is
needed to conrm their occurrence in Georgia.
In addition to the listed species, some others that are
known to occur in the neighborhood of Geogia are ex-
pected to be discovered here too. For instance, Trichodes
sipylus (Linnaeus, 1758) is distributed in Armenia, Azerbai-
jan, and Turkey (Zaitsev 1915; Löbl and Rolcik 2007); T.
ephippiger (Chevrolat, 1874) and T. longissimus (Abeille de
Perrin, 1881) are distributed in Turkey (Bulak et al. 2012);
T. persicus Kraatz, 1894 is distributed in Armenia, Azerbai-
jan (Zaitsev 1915; Löbl and Rolcik 2007), with the closest
occurrence recorded in Gyumri (Armenia) (Slieker et al.
2023). us, all these species are also expected to be found
in Georgia. Nevertheless, targeted research on checkered
beetles in Georgia would uncover the yet-unknown diversi-
ty and distribution patterns of this taxon.
Acknowledgements
e work undertaken in the current study was funded by
the Federal Ministry of Education and Research under
grant 01DK20014A. We express our deepest gratitude to
the anonymous reviewers for their evaluation and advice,
and to Armen Seropian and Levan Mumladze for their valu-
able comments and suggestions.
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