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Review of the Spiders, Pseudoscorpions and Scorpions in the Region of Plovdiv, S Bulgaria (Arachnida: Araneae, Pseudoscorpiones & Scorpiones)

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Abstract and Figures

Although Plovdiv is the second largest city in Bulgaria and on its territory, there are many natural habitats of high conservation importance, there is still almost no data on the species composition of the arachnid fauna. Our study is the first attempt to summarize available published and to provide original data about three orders of arachnids. The following number of the investigated taxa of arachnids was established: spiders: 312 species from 33 families, pseudoscorpions: 4 species from 4 families and one scorpion species (from Euscorpiidae family). New for the region of Plovdiv are 280 species, thirteen spider species are newly recorded for the country and five of them are first records from the Balkans as well. Lathys spasskyi seems to be a first record from Europe and Zelotes harmeron is the first published record from the European mainland. The high total number of species presented here, as well as the many new country records, show that the studies of arachnids in cities have been unjustifiably neglected.
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Bull. Nat. Hist. Mus. Plovdiv
http://rnhm.org/ en/
Regional Natural History Museum - Plovdiv
University of Plovdiv Publishing House
Mollov I., D. Georgiev, O. Todorov (Eds.)
Faun istic d ive rsity o f th e c ity o f Plo v d iv (Bulg aria),
Vo lum e 2 - Ve rte b rat e s & In v e rte b rat e s
Bulletin of the Natural History Museum - Plovdiv
Bu ll. Nat. His t. M us . P lo v d iv , 2022, Sup p le m e n t 2: 83- 110
Review of the Spiders, Pseudoscorpions and Scorpions in the
Region of Plovdiv, S Bulgaria (Arachnida: Araneae,
Pseudoscorpiones & Scorpiones)
Maria Naumova1*, Vasil Genchev2
1 - Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences,
1 Tsar Osvoboditel Blvd.,1000Sofia, BULGARIA
2 - Bulgarian Society for the Protection of Birds,Plovdiv, BULGARIA
*Corresponding author:munny@abv.bg
Abstract. Although Plovdiv is the second largest city in Bulgaria and on its territory, there
are many natural habitats of high conservation importance, there is still almost no data on
the species composition of the arachnid fauna. Our study is the first attempt to summarize
available published and to provide original data about three orders of arachnids. The
following number of the investigated taxa of arachnids was established: spiders: 312species
from 33 families, pseudoscorpions: 4 species from 4 families and one scorpion species
(from Euscorpiidae family). New for the region of Plovdiv are 280 species, thirteen spider
species are newly recorded for the country and five of them are first records from the
Balkans as well. Lath y s sp as s kyi seems to be a first record from Europe and Z e lo te s
h arm e ro n is the first published record from the European mainland. The high total
number of species presented here, as well as the many new country records, show that the
studies of arachnids in cities have been unjustifiably neglected.
Key words: Balkans, Bulgaria, new country record, taxonomy, Thrace, Upper Thracian
lowland.
Introduction
Plovdiv is the second largest city in
Bulgaria and is located in the western part of
the Upper Thracian lowland, on the both
banks of the Maritsa River. The outskirts of
Plovdiv consist mainly of open agricultural
landscapes, pastures, orchards, villages,
warehouses and service areas. In the city and
its surroundings there are many parks,
protected areas and Natura 2000 sites with
high conservation values. This provides a
variety of habitats, which implies a rich fauna.
No any study of the arachnids in the
Plovdiv region has been published to date.
The only historical species reports (21
spiders & 2 pseudoscorpions) can be found
in the papers of Drensky (1913, 1936),
Jurinic & Drensky (1917), Shikrenov (1961),
Dimitrov (1993) and Petrov (1997) and
recently (12 more spiders as part of this
research) in Naumova (2019), Geci &
Naumova (2021), Naumova & Deltshev
(2021) and Naumova et al. (2021). So, to date
only 35 species (33 spiders and 2
pseudoscorpions) were known from the
region of Plovdiv.
The aim of this study is to summarize and
provide all available data about the presence of
three orders of arachnids in Plovdiv city and its
closest surrounds and to present original
photographs of the spiders species Afraf lac illa
e p ib le m o id e s (Chyzer, 1891), At yp us m uralis
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
84
Bertkau, 1890, Dy sd e ra o se ll ai Alicata, 1973,
Erig o ne de n to sa O. Pickard-Cambridge, 1894,
Ero ko re an a Paik, 1967, Ip a te r re n us (L. Koch,
1879), Lathys s p asskyi Andreeva & Tystshenko,
1969, Prit ha p arv a Legittimo, Simeon, Di
Pompeo & Kulczycki, 2017, Te g e n aria
f aniap o llin is Brignoli, 1978, Trac h e las m in o r O.
Pickard-Cambridge, 1872, Ze lo t e s e u g e ne i
Kovblyuk, 2009 and Z. h arm e ro n Levy, 2009 to
facilitate their identification.
Material and Methods
All specimens of arachnids were
collected by hand, by sweeping grass or with
pitfall traps (with formaldehyde) from 1927
to 2021 but mainly in the last four years.
Specimens were preserved in 70-95% ethanol
and deposited (indicated in Table 1) in the
Institute of Biodiversity and Ecosystem
Researches (IBER) and The National
Museum of Natural History (NMNH) at the
Bulgarian Academy of Sciences.
The digital images were taken with a
Canon EOS 1100D digital camera attached
to Carl Zeiss Amplival microscope and
processed using Photoshop CS6 software.
The nomenclature follows the World Spider
Catalog (2021), Harvey (2013) and Kovařík
et al. (2020). All taxa are listed alphabetically.
The collectors were presented by their
initials as follows: D. Georgiev (DG), G.
Glushkov (GG), I. Delev (ID), S. Indzhov, I.
Mollov (IM), M. Naumova (MN), O.
Todorov (OT), P. Drensky (PD), P. Mitov
(PM), S. Popova (SP), T. Teofilova (TT), V.
Genchev (VG).
Other abbreviations used in the paper
include: - male; - female; hc - hand
collected; j -juvenile/ s; L - locality; Pl -
Plovdiv city; RNHM - Regional Natural
History Museum; sa - subadult, Sur -
surrounds.
The collecting sites (alphabetically) are
presented in Table 1 and mapped on Fig. 1.
The data in Table 1 includes the localities
with numbers 1 to 31, in the city of Plovdiv,
followed by sites with numbers 32 to 53 from
the nearest villages (1-10 km from the town)
and related data (UTM-code, exact or
approximate coordinates (decimal), elevation
(m a.s.l.), collecting method, collectors and
depository). All records from Plovdiv,
without exact location were tentatively added
to emblematic region Stariya Grad place (The
Old town) with number 29a and approximate
coordinates.
Table 1. The localities where the arachnids have been collected inthe Plovdiv region.
N
Locality
coordinates
Alt.
Method
coll./Author
deposit
City of PLOVDIV
1
Bratska Mogila place
N 42.1435, E 24.7178
168
sweeping
DG
IBER
2
Bunardzhik hill
N 42.1447, E 24.7394
190-236
soil traps, hc
IM, MN, PD, VG
IBER
3
Central, I
N 42.1482, E 24.7301
170
hc
VG
IBER
4
Central, II
N 42.1497, E 24.7247
163
hc
VG
IBER
5
Central, Maritsa River
N 42.1527, E 24.7230
161
hc
VG
IBER
6
Central, RNHM
N 42.1490, E 24.7430
173
hc
VG
IBER
7
Central, Sity center
N 42.1514, E 24.7415
166
hc
GG
IBER
8
Danov Halm (Sahat Tepe) hill (The
Clocktower)
N 42.1463, E 24.7469
185-210
hc, soil traps
CD, IM, MN, SP
NMNH,
IBER
9
Dzhendem Tepe (Mladezhki Halm) hill
N 42.1498, E 24.7496
168
soil traps
IM
IBER
10
Gagarin ward
N 42.1628, E 24.7544
167
hc
MN
IBER
11
Grebnata baza Place
N 42.1495, E 42.1495
hc
MN, VG
IBER
12
Han Asparuh ward
N 42.1412, E 24.7912
163
hc
MN
IBER
13
Hisar Kapiya place
N 42.1486, E 24.7541
177
hc
PD
NMNHS
14
Izgerv ward
N 42.1485, E 24.7842
159
hc
MN
IBER
15
Iztochna Promishlena Zona ward
N 42.1521, E 24.7730
158
hc
MN
IBER
16
Kamenitsa 1 ward
N 42.1395, E 24.7559
159
hc
Naumova et al.
(2021)
IBER
17
Kapana ward
N 42.1503, E 24.7485
168
hc
VG
Naum o v a &G e n c he v
85
18
Kyuchuka ward
N 42.1102 , E 24.7275
168
hc
Naumova &
Deltshev (2021)
IBER
19
Lauta Park I
N 42.1352, E 24.7716
159
soil traps
ID, IM
IBER
20
Lauta Park II
N 42.1381, E 24.7824
158
hc
PM
IBER
21
Marasha ward
N 42.1507, E 24.7371
166
hc
MN
IBER
22
Maritsa river,a rivershore in Karshiaka ward
N 42.1541, E 24.7279
160
hc
MN, SP
IBER
23
Maritsa river, 4th km, suburban
N 42.1420, E 24.6990
165
soil traps
ID
IBER
24
Maritsa river, 6th km, rural
N 42.1502, E 24.6797
160
soil traps
ID
IBER
25
Maritsa river, Gerdzhika, urban
N 42.1540, E 24.7340
159
soil traps
ID
IBER
26a
Nebet Tepe hill
N 42.1508, E 24.7534
189
hc
VG
IBER
26b
Nebet Tepe hill, Maritsa River
N 42.1541, E 24.7511
161
hc
VG
IBER
26c
Nebet Tepe hill, Stariya Grad I
N 42.1493, E 24.7540
185
hc
VG
IBER
26d
Nebet Tepe hill, Stariya Grad II
N 42.1499, E 24.7540
187
hc
VG
IBER
27
North region, raspberry plantation
N 42.1763, E 24.7619
161
-
-
NMNHS
28
Promishlena Zona Izgrev ward
N 42.1479, E 24.8081
162
hc
VG
IBER
29a
Stariyat grad place I
N 42.1420, E 24.7480
165
hc
GG, PD
-
29b
Stariyat grad place II
N 42.1482, E 24.7301
-
Dimitrov (1993)
-
30
Trakia ward
N 42.1341, E24.7849
165
hc, obs.
Naumova &
Deltshev (2021)
IBER
31
Yuzhna Promishlena Zona waed
N 42.1274, E 24.7714
160
hc
-
IBER
SURROUNDS
32
Bresovitsa village
N 42.1035, E 24.6529
180
hc
PD
-
33
Katunitsa village, Chaya River
N 42.1162, E 24.8669
hc
VG
IBER
34
Kostievo village
N 42.0933, E 24.8185
165
Soil traps
TT
IBER
35
Krumovo village
N 42.0976, E 24.8149
162
hc
CD
NMNH
36
Sadovo village
N 42.1300, E 24.9233
143
hc
BP
NMNH
37
Tsaratsovo village
N 42.2109, E 24.6991
178
Soil traps
IM
IBER
38
Yagodovo village, a yard
N 42.1107, E 24.8510
hc
VG
IBER
39
Yagodovo village, cemetery
N 42.1160, E 24.8496
hc
VG
IBER
40
Yagodovo village, Chaya River I
N 42.1038, E 24.8645
hc
VG
IBER
41
Yagodovo village, Chaya River II
N 42.1140, E 24.8659
hc
VG
IBER
42
Yagodovo village, Chaya River III
N 42.1087, E 24.8656
hc
VG
IBER
43a
Yagodovo village, East I
N 42.1070, E 24.8542
161
hc
VG
IBER
43b
Yagodovo village, East II
N 42.1060, E 24.8520
hc
VG
IBER
43c
Yagodovo village, East III
N 42.1069, E 24.8557
hc
VG
IBER
44
Yagodovo village, N field I
N 42.1284, E 24.8556
152
hc
VG
IBER
45a
Yagodovo village, N field II
N 42.1312, E 24.8493
153
hc
VG
IBER
45b
Yagodovo village, N field III
N 42.1320, E 24.8429
154
hc
VG
IBER
46a
Yagodovo village, North I
N 42.1181, E 24.8532
157
hc
VG
IBER
46b
Yagodovo village, North II
N 42.1148, E 24.8526
160
hc
VG
IBER
46c
Yagodovo village, North III
N 42.1151, E 24.8516
150
hc
VG
IBER
47
Yagodovo village, North IV
N 42.1234, E 24.8527
157
hc
VG
IBER
48
Yagodovo village, North V
N 42.1260, E 24.8430
156
hc
VG
IBER
49
Yagodovo village, South I
N 42.0998, E 24.8481
164
hc
VG
IBER
50
Yagodovo village, South II
N 42.1010, E 24.8472
165
hc
VG
IBER
51
Yagodovo village, West I
N 42.1033, E 24.8378
162
hc
VG
IBER
52
Yagodovo village, West II
N 42.1128, E 24.8387
161
hc
VG
IBER
53
Yagodovo village, West III
N 42.1145, E 24.8284
158
hc
VG
IBER
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
86
Fig. 1 . Map with the localities in Plovdiv where the arachnids have been collected
(for details see Table 1).
Results
During this study the following number
of the investigated taxa of arachnids was
established: spiders: 312 species from 33
families (Agelenidae-7, Amaurobiidae-3,
Anyphaenidae-1, Araneidae-27, Atypidae-1,
Cheiracanthiidae-6, Clubionidae-2, Dictynidae-
8, Dysderidae-8, Eresidae-2, Filistatidae-2,
Gnaphosidae-30, Hahniidae-1, Linyphiidae-37,
Liocranidae-3, Lycosidae-34, Mimetidae-3,
Miturgidae-4, Oecobiidae-1, Oxyopidae-2,
Philodromidae-13, Pholcidae-4, Phrurolithidae-
3, Pisauridae-1, Salticidae-40, Scytododae-2,
Tetragnathidae-9, Theridiidae-32, Thomisidae-
19, Titanoecidae-3, Trachelidae-1, Uloboridae-1,
Zodariidae-2); pseudoscorpions: 4 species from
4 families (Cheliferidae-1, Chernetidae-1,
Chtoniidae-1, Neobisiidae-1) and one scorpions
from the family Euscorpiidae. New for the
region of Plovdiv are 280 species (marked with
an asterisk). Thirteen spider species are newly
recorded for the country (marked with 2
asterisks) and five of them are first records
from the Balkans as well (marked with 3
asterisks). Lathys s p asskyi Andreeva &
Tystshenko, 1969 seems to be a first record
from Europe and Z e lo te s har m e ro n Levy, 2009
is the first published record from the European
mainland. One speciecies from a genus
Sc y to d e s Latreille, 1804 seems to be new for
the science and will be described in separate
paper.
The records of D ys d e ra e r yt hr ina
(Walckenaer, 1802) from Plovdiv, Bunardzhika
(Drensky, 1913: 87 (sub D. c am b r idg e i),
Drensky, 1936: 12, Jurinitch & Drensky,
1917:125) is listed here as D. sp. e ry th rin a
complex. The material from Plovdiv cannot be
located (probably lost) and in accordance with
Řezáč et al., (2018) the species have been
excluded from the Bulgarian checklist of
spiders. The revision of e ry th rin a species
complex in the Balkans is in the future.
Three spider species are included in the
list below as uncertain records: Ne o n p ic tu s
Kulczyński, 1891 from the hills Bunardzhika &
Sahat Tepe (Jurinitch & Drensky, 1917: 135),
Ozy p tila rauda Simon, 1875 from Sahat Tepe
(Drensky, 1913: 101 sub O xyptila r. Simon,
1875) and Ch e irac an th ium m arg ar itae Sterghiu,
Naum o v a &G e n c he v
87
1985 (described by a single female specimen in
Romania) and reported (also by single female) from
Plovdiv (Dimitrov, 1993: 74). They have not been
reported over again and it is quite possible to be a
result of misidentifications. Since the materials of
these species cannot be located (probably lost),they
are included in the list below, with a question mark
(?) and need conformation.
Table 2. List of arachnids. Le g e n d: S - status: * - new for the Plovdiv region, ** - new country
record, *** - species new for the Balkans, # - species nov., ? - uncertain record, L - locality
(number = to the number in Table 1 and Fig. 1), j - juvenile, sa - subadult, Pl - Plovdiv city, Sur -
surrounds.
S
Taxa
L (sex, date)
Pl
Sur
ARANEAE
Agelenidae
*
Ag e le n a o r ie nt alis C. L. Koch, 1837
38 (1 , 16.08.2018, 1 , 28.08.2018), 41 (2 ,
14.08.2021), 42 (1 , 4 sa, 08.09.2018)
+
Allag e le n a g rac ile n s (C. L. Koch, 1841)
11 (1 , 16.07.2021), 29a (Drensky, 1913, sub
A. s im ilis , Drensky, 1936), 33 (1 ,
25.07.2021), 44 (1 , 25.03.2020), 46b (1 ,
29.08.2020)
+
+
*
Eratig e n a ag re s tis (Walckenaer, 1802)
23 (1 , 30.06-14.07.2020), 31 (1, 22.02.1988)
+
*
Maim u na v e st ita (C. L. Koch, 1841)
2 (1 , 19.04.2017), 6 (1 ¸18.02.2022), 7 (1
¸ 02-03.2018), 8 (1 , 1 , 20.03.2018), 15 (1
, 11.11.2016), 38 (3 , 8 , 1 sa, 1 j, 16-
30.03.2018, 1 , 11-15.04.2018, 1 ,
21.2.2021)
+
+
**
Te g e n aria f an iap o llin is Brig n o li, 1978
5 (1 , 26.03.2021), 38 (1 , 1 sa,
16.06.2021, 3 , 19.06.2021, 1 , 28.10.2021, 2
, 17.02.2022)
+
+
*
Te g e n aria h as p e ri Chyzer, 1897
4 (1 , 30.06.2021), 23 (1 , 01-30.09.2020),
24 (1 , 01-30.09.2020 , 1 , 30.06-
14.07.2020), 25 (1 , 30.06-14.07.2020), 33 (2
), 38 (16.06.2019)
+
+
Te g e n aria p ar ie tina (Fourcroy, 1785)
6 (1 , 25.10.2018), 29a (1 , 25.12.2018, 1 j,
03.10.1927), 32 (Jurinic & Drensky, 1917), 38
(2 , 16-30.03.2018, 1 , 04.10.2018)
+
+
Amaurobiidae
*
Am au ro b ius e r b e ri (Keyserling, 1863)
2 (1 , 19.04.2017, 1 , 2 j, 01-04.05.2018), 6
(2 , 18.02.2022), 8 (1 , 1 , 13.02.1988, 2 ,
4, 20.03.2018), 15 (1 , 1 j, 11.11.2016), 22
(1 , 20.03.2018), 26c (2 , 27.04.2018), 38 (6
, 9 j, 16-30.03.2018, 1 , 02.04.2018, 2 , 11-
15.04.2018, 1 , 3 , 08-21.04.2019, 2 , 11-
21.05.2019, 2 , 28.10.2020, 1 , 06.11.2020)
+
+
*
Am aur o biu s f e ro x (Walckenaer, 1830)
38 (1 , 05.2018)
+
*
Am aur o biu s p allidu s L. Koch, 1868
19(1, 01-31.05.2021),20(3,3,1j,
18.04.2017)
+
Anyphaenidae
*
An yp h ae na ac c e n tu ata (Walckenaer, 1802)
33 (1 , 23.04.2022, 1 , 1 , 02.05.2022)
Araneidae
*
Ac u le pe ir a c e r o p e g ia (Walckenaer, 1802)
33(1,16.06.2022),38(1,01.07.2020,1,
03.07.2020)
+
*
Ag ale n ate a r e dii (Scopoli, 1763
38 (1 , 08-21.04.2019, 1 j , 15.04.2018)
+
*
Aran e us an g ul atus Clerck, 1757
29a (1 sa, 03.10.1927), 38 (1 , 19.08.2018),
44 (3 , 1 , 10-12.07.2018)
+
+
*
Aran e us d iad e m atu s Clerck, 1757
29a (1 , 09.11.2018, 1 , 15.11.2018, 1 ,
01.09.2019), 38 (1 , 14.08.2018, 1 ,
+
+
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
88
29.09.2018), 44 (1 , 21.10.2018, 1 ,
27.08.2021)
*
Aran ie lla c u c urb itin a (Clerck, 1757)
9 (1 , 2 , 17.06.2021), 33 (1 , 16.06.2019),
40 (1 , 1 , 04.06.2021)
+
+
*
Aran ie lla o p ist ho g rap ha (Kulczynski, 1905)
27(1, 1, 27.10.1988),38(1,28.05-
08.06.2019)
+
+
*
Arg io p e b ru e nn ic h i (Scopoli, 1772)
44 (1 , 10-12.07.2018), 48 (1 , 18.07.2018)
+
*
Arg io p e lo b ata (Pallas, 1772)
14 (1 , 18.08.2021), 28 (1 , 25.07.2018)
+
*
Ce rc id ia pr o m in e ns (Westring, 1851)
38 (1 , 11.08.2020, 1 , 30.08.2020)
+
*
Cy c lo sa c o nic a (Pallas, 1772)
35 (1 , 04.1988)
+
*
Cy c lo sa o c ulat a (Walckenaer, 1802)
38 (1 , 27-29.05.2018, 1 , 31.07.2020, 1 ,
02.08.2020, 2 , 30.08.2020)
+
*
Gib b aran e a b itu b e rc u lata (Walckenaer, 1802)
19 (1 , 26.05.2011), 40 (2 , 10-11.05.2021)
+
+
*
Gib b aran e a g ib b o s a (Walckenaer, 1802)
38 (1 , 08-21.04.2019)
+
*
Gib b aran e a u llric h i (Hahn, 1835)
38 (1 , 11-18.05.2018, 1 ,, 20.03.2019
+
*
Hyp s o s ing a p yg m ae a (Sundevall, 1831)
38 (1 , 04-14.07.2019, 1 , 16.07.2021)
+
*
Hyp s o s ing a san g uin e a (C. L. Koch, 1844)
38 (2 , 25.04-05.05.2019, 1 , 27.04.2019, 1
, 04.05.2019, 1 , 22.06.2021), 39 (1 ,
12.07.2021)
+
*
Larinio id e s ixo b o lus (Thorell, 1873)
42 (2 , 08.09.2018)
+
*
Larinio id e s p atag iat us (Clerck, 1757
44(1,25.03.2020,1,15.10.2020,1,04.09.2021)
+
*
Larinio id e s s us p ic ax (O. Pickard-Cambridge, 1876)
33 (2 , 30.05.2018)
+
*
Lev ie llu s s tro e m i (Thorell, 1870)
25 (1 , 20.05-29.06.2020),33 (1 , 25.07.2021)
+
+
*
Man g o ra ac alyp h a (Walckenaer, 1802)
1(6,03-04.06.2017),2(1,1 ¸ 01-04.05.2018),
19(1, 26.05.2011), 33(1, 25.07.2021),35(1,
04.1988), 38(1sa, 11-15.04.2018,1,04-
14.07.2019),40(1,04.06.2021)
+
+
Ne o sc o n a ad iant a (Walckenaer, 1802)
33 (1 , 18.07.2021), 38 (1 , 03-10.07.2018),
44 (Geci & Naumova, 2021)
+
Ne o sc o n a b yzan th ina (Pavesi, 1876)
33 (1 , 05.09.2022), 44 (Geci & Naumova,
2021), 45a (Geci & Naumova, 2021, 1 , 1
sa, 27.08.2021), 53 (Geci & Naumova, 2021)
+
*
Ne o sc o n a s ub f us c a (C. L. Koch, 1837)
38 (2 , 03-10.07.2018)
+
*
Nuc t e ne a um b r atic a (Clerck, 1757)
38 (1 , 30.07.2018)
+
*
Sing a nit id ula C. L. Koch, 1844
44 (1 , 24.05.2021)
+
*
Zilla d io dia (Walckenaer, 1802)
2 (1 , 01-04.05.2018), 3 (1 , 24.06.2021), 38
(1 , 25.04-05.05.2019, 2 , 23.08.2019)
+
+
Atypidae
*
Aty p us m ur alis Bertkau, 1890
43a (1 , 23.06.2021)
+
Cheiracanthiidae
*
Ch e irac an th ium e le g an s Thorell, 1875
33 (2 , 18.07.2021)
+
*
Ch e irac an th ium e rr atic u m (Walckenaer, 1802)
19 (1 , 26.05.2011), 44 (1 , 09.05.2019)
+
?
Ch e irac an th ium m ar g aritae Sterghiu, 1985
29b (Dimitrov, 1993)
+
*
Ch e irac an th ium m ild e i L. Koch, 1864
6 (2 sa, 5 j, 18.02.2022), 29a (1 j, 03.10.1927),
38 (30.05.2018)
+
+
*
Ch e irac an th ium p e n ny i O. Pickard-Cambridge, 1873
13 (1 , 25.08.1928)
+
*
Ch eir ac anth iu m p un c to riu m (Villers, 1789)
38 (1 , 25.07.2020), 44 (2 , 1 , 10-
12.07.2018)
+
Clubionidae
*
Club io n a f ru te to rum L. Koch, 1867
24 (1 , 30.06-14.07.2020), 33 (1 ,
25.07.2021)
+
+
*
Club io n a lu te s c e n s Westring, 1851
23 (1 , 01-30.09.2020), 24 (2 , 1 , 20.05-
29.06.2020, 1 , 30.06-14.07.2020)
+
Naum o v a &G e n c he v
89
Dictynidae
Brig it te a c iv ic a (Lucas, 1850)
3 (1 , 22.05.2021), 6 (1 , 18.02.2022), 11 (1
, 17.06.2021), 13 (2 , 25.08.1928), 29a
(Drensky, 1936 sub Dic t yn a c .), 38 (1 ,
18.06.2020), 50 (2 , 26.05.2019)
+
+
*
Brig it te a v ic ina (Simon, 1873)
38 (3 , 20.06.2020)
+
*
Dic ty n a aru nd in ac ea (Linnaeus, 1758)
22 (4 sa, 20.03.2018), 33 (1 , 2 ,
04.07.2019), 38 (2 , 18-26.06.2019)
+
+
*
Dic ty n a m ajo r Menge, 1869
44 (2, 10-12.07.2018)
+
*
Dic ty n a u nc in ata Thorell, 1856
5 (1 , 26.06.2021), 27 (5 , 27.10.1988), 40 (1
, 10-11.05.2021)
+
+
***
Lathy s s p asskyi Andreeva & Tystshenko, 1969
26c (1 , 27.04.2018)
+
*
Lathy s s tig m atis ata (Menge, 1869)
35 (1 , 04.1988)
+
*
Nig m a w al c ken ae ri (Roewer, 1951)
38 (1 , 14.10.2019)
+
Dysderidae
*
Dy sd e ra c r o c ata C. L. Koch, 1838
2 (1 , 18.09.2017)
+
Dy sd e ra sp. e r yth rin a complex
29a (Drensky, 1913 sub D. c am b rid g e i,
Drensky, 1936, Jurinitch & Drensky, 1917)
+
*
Dy sd e ra lata Reuss, 1834
38 (1 , 3 , 11-15.04.2018, 1 , 28.08.2018, 1
, 16.07.2019, 1 , 1 , 15.05.2021, 1 ,
21.05.2021, 1 , 10.11.2021)
+
*
Dy sd e ra l o ng iro s tris Doblika, 1853
23 (2 , 20.05-29.06.2020, 1 , 30.06-
14.07.2020, 1 , 1 j, 01-30.09.2020), 33 (1 ,
23.04.2022, 1 , 16.06.2022)
+
***
Dy sd e ra o s e llai
26c (1 , 27.04.2018, 1 , 28.10.2018)
+
*
Harp ac te a b abo ri (Nosek, 1905)
2 (1 , 19.04.2017, 2 , 3 sa, 18.09.2017, 3
, 3 , 01-04.05.2018), 9 (1 , 19.09.2017), 23
(1 , 30.06-14.07.2020)
+
Harp ac te a le p ida (C. L. Koch, 1838)
29a (Drensky, 1913 (sub Harp ac t e s se id e lii
[sic]); Drensky, 1936)
+
*
Harp ac te a ru b ic un da (C. L. Koch, 1838)
23(1, 20.05-29.06.2020,1 ,1 ,30.06-
14.07.2020)
+
Eresidae
*
Ere su s ko llari Rossi, 1846
30 (1 , 10.10.2018)
+
Ere su s m o r av ic us Rezac, 2008
10(1, 15.07.2021), 11(Naumova&Deltshev,
2021), 18(Naumova &Deltshev,2021), 26c
(Naumova &Deltshev, 2021), 37 (1 ,12.09.2013),
38(1, 27-29.05.2018),43b(Naumova&
Deltshev,2021),50(1,26.05.2019,Naumova&
Deltshev,2021),
+
+
Filiistatidae
Prit ha p arv a Legittimo, Simeon, Di Pompeo & Kulczycki,
2017
3 (1 , 1 sa, 22.05.2021, 1 , 15.06.2021 ), 38
(Naumova & Deltshev, 2021, 1 j, 01.07.2020)
+
+
*
Filistata ins idiatrix (Forsskål, 1775)
38 (1 sa, 28.06.2022)
Gnaphosidae
*
Ap han tau lax trif as c iata (O. Pickard-Cambridge, 1872)
33 (1 , 18.07.2021), 38 (1 , 19.06.2021)
+
*
Civ ize lo t e s g rac ilis (Canestrini, 1868)
9(1,17.06.2021),38(1,16.06.2021,1,
19.06.2021)
+
+
*
Dras s o d e s c up re u s (Blackwall, 1834)
22 (2 sa, 20.03.2018)
+
*
Dras so d e s lap ido s us (Walckenaer, 1802)
11 (1 , 09.07.2021), 25 (1 , 20.05-
29.06.2020), 38 (1 , 01.05.2021, 1 ,
25.05.2021)
+
+
*
Dras s o d e s lut e sc e n s (C. L. Koch, 1839)
38 (1 , 11-15.04.2018)
+
*
Dras s o d e s p ub e sc e n s (Thorell, 1856)
25 (1 , 20.05-29.06.2020)
+
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
90
*
Dras s yllu s p rae f ic u s (L. Koch, 1866)
2(1,01-04.05.2018),9(1,17.06.2021),23(5,
2,20.05-29.06.2020,1,30.06-14.07.2020, 1
sa,01-30.09.2020), 24(1,20.05-29.06.2020),25
(1,20.05-29.06.2020),38(1,14-20.05.2020, 1
,28.05.2021),44(1,10-12.07.2018)
+
+
*
Gn ap h o sa luc if ug a (Walckenaer, 1802)
38 (1 , 05.07.2021), 43a (1 , 1 ,
27.06.2021, 1 , 29.06.2021, 1 sa,
24.09.2021)
+
*
Gn ap ho s a ru fu la (L. Koch, 1866)
34 (3 , 1 , 14.05-02.06.2017)
+
*
Hap lo dras su s bo h e m ic us Miller & Buchar, 1977
24 (1, 20.05-29.06.2020)
+
*
Hap lo d rass us sig n if e r (C. L. Koch, 1839)
25 (2 , 20.05-29.06.2020), 34 (1 , 14.05-
02.06.2017), 38 (2 , 11-15.04.2018, 2 , 08-
21.04.2019, 1 , 23.08.2019)
+
+
*
Marin aro ze lo te s m alkini (Platnick & Murphy, 1984)
23(1, 30.06-14.07.2020),34(1,14.05-
02.06.2017),38(1,03-10.07.2018, 1,
07.07.2021)
+
+
*
Mic aria alb o v itt ata (Lucas, 1846)
24(1, 30.06-14.07.2020),25(1,20.05-
29.06.2020),38(1,25-31.07.2020, 1,
18.05.2021,1 ,13.06.2020),47(1,1 ,
29.07.2019)
+
+
*
Mic aria c o arc tat a (Lucas, 1846)
11 (1 , 09.07.2021), 38 (1 , 16.07.2019)
+
+
*
Mic aria d iv e s (Lucas, 1846)
38 (1 , 05.06.2021)
+
**
Mic aria m ic ans (Blackwall, 1858)
25 (1 , 1 , 30.06-14.07.2020)
+
*
Mic aria pu lic aria (Sundevall, 1831)
38 (1 , 23.08.2019)
+
*
No m isia e xo rn ata (C. L. Koch, 1839)
26c (1 , 27.04.2018)
+
*
Sc o to p h aeu s sc u tu latus (L. Koch, 1866)
5(1,30.10.2022),6(3,18.02.2022),29a(1,
18.09.1983),33(1,06.09.2021),38(1,1,
29.09.2018,1,11-21.05.2019,1,06.11.2020,1,
16.05.2021)
+
+
*
Trac h y ze lo t e s p e de s tris (C. L. Koch, 1837)
3 (1 sa, 26.02.2021), 11 (1 , 09.07.2021), 25
(3 , 20.05-29.06.2020), 50 (1 , 26.05.2019)
+
+
**
Tu rko ze lo te s m c c o w ani (Chatzaki & Russell-Smith, 2017)
34 (4 , 1 , 14.05-02.06.2017)
+
*
Uro ze lo te s ru stic u s (L. Koch, 1872)
7 (1 , 02-03.2018), 38 (2 , 14-15.06.2020)
+
+
*
Ze lo te s ap r ic o ru m (L. Koch, 1876)
23(1, 1, 30.06-14.07.2020, 1, 01-30.09.2020)
+
*
Ze lo te s e l e c tu s (C. L. Koch, 1839)
5 (1 , 26.03.2021), 38 (1 , 19.03.2021)
+
+
*
Ze lo te s e r e b e us (Thorell, 1871)
47 (1 , 17.09.2021)
+
**
Ze lo te s e u g e n e i Kovblyuk, 2009
24(2,1,30.06-14.07.2020,1,2,01-
30.09.2020),25(1,20.05-29.06.2020,1,30.06-
14.07.2020),38(1,1,11-15.04.2018,1,
18.04.2019,1,20.06.2020,1,25.07.2020),50(1
,17.08.2019)
+
+
**
Ze lo te s h arm e ro n Levy, 2009
43c (1 , 15.07.2021)
+
*
Ze lo te s h e rm an i (Chyzer, 1897)
9(1,19.09.2017),22(1,20.03.2018), 23(4,1
,01-30.09.2020), 24(1,30.06-14.07.2020),25(5
,01-30.09.2020), 38(1,28.08.2018,1 ,
20.10.2021),39(1,19.09.2021),43a(1,
15.06.2020)
+
+
*
Ze lo te s l o ng ip e s (L. Koch, 1866)
33 (1 , 21.08.2021), 38 (1 , 29.09.2018, 2 ,
1 sa, 25.04-05.05.2019, 1 , 23.08.2019)
+
*
Ze lo te s t e nu is (L. Koch, 1866)
11 (1 , 09.07.2021), 38 (1 , 1 , 27-
29.05.2018, 3 , 28.05-08.06.2019, 1 , 18-
26.06.2019, 1 , 16.07.2019, 1 , 14-
20.05.2020, 2 , 17.07.2020, 1 , 31.05.2021, 1
, 1 , 19.06.2021)
+
+
Hahniidae
*
Hahn ia nav a (Blackwall, 1841)
38 (1 , 25.05.2021)
+
Linyphiidae
Naum o v a &G e n c he v
91
*
Ab ac o p ro e c e s s altu um (L. Koch, 1872)
23 (1 , 20.05-29.06.2020)
+
*
Ac artau c he n ius s c ur rilis (O. Pickard-Cambridge, 1873)
2 (1 , 18.09.2017), 8 (1 , 17.09.2017), 35 (1
, 04.1988), 38 (1 , 26-30.04.2018)
+
+
**
Ag y ne t a p un c tat a Wunderlich, 1995
34 (1 , 14.05-02.06.2017)
+
*
Ag y ne t a ru re s tris (C. L. Koch, 1836)
8(1,13.02.1988),12(1,11.11.2016),31(1,
22.02.1988),35(1,04.1988),38(1,1 ,
23.08.2019,1 ,10.02.2022), 40(1, 1,
04.06.2021)
+
+
*
Can arip han te s n an us (Kulczyński, 1898)
9 (1 , 17.06.2021)
+
*
Ce n tro m e ru s sy lv atic u s (Blackwall, 1841)
19 (1 , 01-31.05.2021)
+
*
Ce ratin e lla b re v ip e s (Westring, 1851)
24 (1 , 30.06-14.07.2020)
+
*
Cre s m ato n e ta m u tin e ns is (Canestrini, 1868)
25 (1 , 01-30.09.2020), 38 (1 sa, 21-
26.10.2018, 1 , 08-21.04.2019)
+
+
*
Dic y m b ium n ig ru m (Blackwall, 1834)
38 (1 , 13.06.2020)
+
*
Dip lo st yla c o nc o lo r (Wider, 1834)
19(1, 01-31.05.2021),20(1,18.04.2017),22(1
,20.03.2018),23(2,01-30.09.2020), 24(2, 1
,30.06-14.07.2020,1 , 01-30.09.2020),25(6,9
,30.06-14.07.2020),38(1, 04-14.07.2019)
+
+
*
Erig o n e d e n tip alp is (Wider, 1834)
38 (1 , 25.04-05.05.2019)
+
***
Erig o ne d e nto s a O. Pickard-Cambridge, 1894
33 (1 , 18.07.2021)
+
*
Fro n tin e llin a f ru te to ru m (C. L. Koch, 1835)
2 (1 , 3 , 01-04.05.2018), 38 (2 , 11-
18.05.2018, 1 , 15.08.2018, 1 , 05.06.2021)
+
+
*
Gn ath o n ariu m d e n tatu m (Wider, 1834)
33 (1 , 16.07.2022)
*
Go n g ylid iu m r uf ip e s (Linnaeus, 1758)
24 (2 , 30.06-14.07.2020, 2 , 01-30.09.2020)
+
*
Im p ro p han te s d ec o lo r (Westring, 1861)
11 (1 , 19.04.2017)
+
*
Ip a t e rre n us (L. Koch, 1879)
38 (1 , 16-30.03.2018, 1 , 22.04.2019)
+
*
Liny ph ia trian g ular is (Clerck, 1757)
38 (1 , 09.11.2018), 44 (1 , 20.10.2020)
+
*
Mic ro c t e no n y x s ub it ane u s (O. Pickard-Cambridge, 1875)
38 (2 , 11-15.04.2018, 1 , 03.07.2018)
+
Mic ro lin yp h ia im p ig ra (O. Pickard-Cambridge, 1871)
38 (Naumova et al., 2021)
+
*
Mic ro lin yp h ia p u silla (Sundevall, 1830
38 (1 , 03-10.07.2018, 1 , 08-21.04.2019, 1
, 11-21.05.2019, 1 , 18-26.06.2019, 1 , 04-
14.07.2019, 1 , 1 sa, 10-11.08.2020)
+
*
Ne rie n e c lat hr ata (Sundevall, 1830)
5(1,21.04.2021),23(1,30.06-14.07.2020),24
(1,30.06-14.07.2020),25(1,30.06-14.07.2020)
38(1, 16-30.03.2018,2 ,10-11.08.2020)
+
+
*
Ne rie n e f ur tiv a (O. Pickard-Cambridge, 1871)
38 (1 , 25.05.2020, 1 , 25-31.07.2020)
+
*
O ed o th o rax ap ic atu s (Blackwall, 1850)
33 (1 , 16.06.2019)
+
*
O st e arius m e lan o p yg iu s (O. P.-Cambridge, 1879)
38 (1 , 1 j, 16-30.03.2018, 1 , 03-10.07.2018)
+
*
Pe le c o p sis e lo n g ata (Wider, 1834)
2 (1 , 18.09.2017), 15 (1 , 11.11.2016)
+
*
Prin e rig o n e v ag an s (Audouin, 1826)
31(1, 1, 22.02.1988),33(1,12.09.2021),38
(1,13.06.2020), 40(1, 10-11.05.2021,2,
27.05.2021)
+
+
*
Sint ula re tro v e rs us (O. P.-Cambridge, 1875)
2(4,2sa,18.09.2017),38(1, 3, 21-
26.10.2018)
+
+
*
Ste m o n y p han te s lin e atu s (Linnaeus, 1758)
47 (1 , 07.10.2022)
*
Sye d ra g rac ilis (Menge, 1869)
19 (1 , 1 sa, 01-31.05.2021)
+
*
Te n uip h ant e s te n e b ric o la (Wider, 1834)
2 (2 , 09.06.2021)
+
*
Te n uip h ant e s te n uis (Blackwall, 1852
9(1,19.09.2017,1,4 ,1sa,17.06.2021),23
(2,30.06-14.07.2020),33(1,04.07.2019),38(1
,11-15.04.2018,2 ,25.04-05.05.2019,2, 11-
21.05.2019,3 ,1 sa,1sa,23.08.2019, 1,
15.05.2021)
+
+
*
Tric h o n c o ide s p isc at o r (Simon, 1884)
11(1, 17.06.2021), 31(1, 22.02.1988),34(1,
14.05-02.06.2017),35(1,14.11.2021),38(1,21-
26.10.2018,1 , 26.07.2021), 46c (1 ,25.05.2018)
+
+
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
92
*
Tric h o n c us af f inis Kulczynski, 1894
37 (1 , 12.09.2013)
+
*
Tric h o n c us h ac km an i Millidge, 1955
11 (1 , 09.07.2021), 38 (1 , 01.07.2021)
+
+
*
Tric h o n c us s axic o la (O. Pickard-Cambridge, 1861)
38 (1 , 15.05.2021)
Walc ke nae ria c ap ito (Westring, 1861)
29a (Drensky, 1913 sub Walc ken ria [sic] c.,
Drensky, 1936)
+
Liocranidae
*
Ag ro e c a c up re a Menge, 1873
8 (1 , 17.09.2017, 1 , 20.03.2018), 25 (2 ,
20.05-29.06.2020, 2 , 3 , 01-30.09.2020), 38
(1 , 11-15.04.2018, 1 , 09.04.2019, 2 ,
22.04.2019, 1 , 15.05.2021, 1 , 08.06.2021)
+
+
***
Lio c rano e c a sp ass kyi Ponomarev, 2007
34 (36 , 4 , 14.05-02.06.2017)
+
Me sio t e lus t e nu iss im us (L. Koch, 1866)
29a (DRENSKY, 1913, 1936), 38 (1 , 16-
30.03.2018, 1 , 11-15.04.2018, 1 , 26-
30.04.2018, 1 , 23.08.2019, 1 , 27.02.2021, 1
, 04.08.2021), 44 (1 , 23.08.2019)
+
+
Lycosidae
*
Alo p e co s a alb o f asc iata (Brullé, 1832)
2(1,25.05.2018),33(1,27.05.2021),38(1,
28.05.2019)
+
+
*
Alo p e c o s a c un e ata (Clerck, 1757)
38 (1 , 15.12.2018)
+
*
Alo p e co s a f arin o sa (Herman, 1879)
29a (1 , 14.03.2021), 38 (1 , 28.04.2019), 51
(2 , 30.03.2021)
+
+
*
Alo p e c o s a pu lv e ru le nt a (Clerck, 1757)
25 (18 , 1 , 20.05-29.06.2020)
+
*
Alo p e co s a t aen io p us (Kulczynski, 1895)
25 (1 , 2 , 01-30.09.2020), 38 (1 ,
18.03.2019)
+
+
*
Arc t o sa c in e re a (Fabricius, 1777)
24 (1 , 20.05-29.06.2020), 33 (1 ,
09.04.2022, 1 , 23.04.2022)
+
*
Arc to s a le o p ard us (Sundevall, 1833)
5 (1 , 11.05.2021), 40 (1 , 1 sa, 10-
11.05.2021, 2 , 27.05.2021)
+
+
*
Arc t o sa v arian a C. L. Koch, 1847
38 (1 , 13.06.2021)
+
*
Aulo n ia alb im ana (Walckenaer, 1805)
4 (1 sa, 05.03.2021), 19 (2 , 1 sa, 01-
31.05.2021), 22 (1 j,20.03.2018), 23 (2 ,
20.05-29.06.2020), 24 (6 , 20.05-29.06.2020),
25 (33 , 2 , 20.05-29.06.2020), 38 (2 ,
11.05-21.05.2019, 1 , 28.05-08.06.2019, 1 ,
14-20.05.2020)
+
+
Ge o ly c o sa v ultu o sa (C. L. Koch, 1838)
2 (1 , 14.05.2021), 29a (1 , 1 sa,
03.10.1927, 1 , 1 sa, 04.10.1927), 32
(Jurinich & Drensky, 1917 sub T ro c ho s a
in fe r nalis ), 38 (1 , 16-30.03.2018, 1 , 11-
18.05.2018), 39 (1 , 03.10.2018), 47 (1 ,
23.10.2020)
+
+
*
Ho g n a rad iata (Latreille, 1817)
1 (1 sa, 03-04.06.2017), 29a (1 ,
20.08.2019), 43a (1 , 05.07.2021), 44 (1 , 10-
12.07.2018)
+
+
Lyc o s a p r ae g rand is C. L. Koch, 1836
32 (Drensky, 1915 sub Tare n tu la p .)
+
*
Pard o s a ag r e stis (Westring, 1861)
24 (9 , 5 , 30.06-14.07.2020), 31 (1 ,
22.02.1988), 33 (2 , 16.06.2019, 2 ,
18.07.2021), 34 (1 , 1 , 14.05-02.06.2017),
38 (1 , 18-26.06.2019), 43a (1 , 29.06.2021),
45b (3 , 09.05.2019), 51 (2 , 30.03.2021)
+
+
*
Pard o sa ag ric o la (Thorell, 1856)
24 (1 , 20.05-29.06.2020, 7 , 2 sa, 01-
30.09.2020), 29a (2 , 5 , 05.09.1988)
+
*
Pard o s a alac ris (C. L. Koch, 1833)
23 (1 , 1 , 20.05-29.06.2020)
+
*
Pard o s a am e n tat a (Clerck, 1757)
25 (2 , 30.06-14.07.2020), 33 (1 ,
18.07.2021), 38 (2 , 3 , 13.06.2020), 40 (2
, 2 , 10-11.05.2021, 1 , 1 , 27.05.2021)
+
+
*
Pard o sa ato m aria (C. L. Koch, 1847)
33 (1 , 18.07.2021, 1 , 12.09.2021)
+
Naum o v a &G e n c he v
93
*
Pard o sa h o rte ns is (Thorell, 1872)
5 (2 , 21.04.2021), 23 (3 , 8 , 20.05-
29.06.2020, 1 , 30.06-14.07.2020), 24 (2 ,
20.05-29.06.2020, 5 , 30.06-14.07.2020), 33 (2
, 04.07.2019, 3 , 18.07.2021), 34 (2 ,
14.05-02.06.2017), 38 (1 , 2 sa, 5 sa, 16-
30.03.2018, 1 , 03.04.2018, 1 , 1 , 11-
15.04.2018, 1 , 11-18.05.2018, 1 ,
23.08.2019, 1 sa, 1 sa, 08.11.2020, 1 , 1 ,
12-13.04.2021, 1 , 19.06.2021), 40 (2 ,
04.06.2021), 44 (1 , 30.05.2021), 51 (2 ,
30.03.2021)
+
+
*
Pard o s a lu g ub ris (Walckenaer, 1802)
19 (36 , 6 , 1 sa, 01-31.05.2021), 25 (1 ,
20.05-29.06.2020), 33 (2 , 16.06.2019), 38 (1
, 1 , 11-15.04.2018), 40 (1 , 10-
11.05.2021)
+
+
*
Pard o sa m o nt ic o la (Clerck, 1757)
45b (2 , 2 sa, 09.05.2019)
+
*
Pard o sa n eb ulo s a (Thorell, 1872)
11 (1 , 09.07.2021), 33 (1 , 16.06.2019, 1 ,
04.07.2019, 1 , 14.06.2021)
+
+
*
Pard o sa p alud ic o la (Clerck, 1757)
22(2, 2, 1sa,20.03.2018),51(1,
30.03.2021)
+
+
*
Pard o sa p ro xima (C. L. Koch, 1847)
1 (3 , 03-04.06.2017), 22 (1 , 20.03.2018),
24 (8 , 9 , 30.06-14.07.2020), 25 (4 , 5 ,
30.06-14.07.2020), 38 (1 , 18-26.06.2019, 1 ,
01.07.2020), 50 (1 , 27.05.2019)
+
+
*
Pard o sa p ullata (Clerck, 1757)
40 (1 , 27.05.2021)
+
*
Pirat a p iratic u s (Clerck, 1757)
40 (2 , 1 , 10-11.05.2021, 2 , 27.05.2021)
+
*
Pirat a te n u itars is Simon, 1876
40 (1 , 27.05.2021)
+
*
Pirat ula hy g ro p h ila (Thorell, 1872)
23 (16 , 20.05-29.06.2020), 40 (,3 , 2 , 10-
11.05.2021, 1 , 2 , 27.05.2021)
+
+
*
Piratula latitan s (Blackwall, 1841)
23 (9 , 20.05-29.06.2020), 40 (2 , 3 , 10-
11.05.2021, 2 , 1 , 27.05.2021)
+
+
*
Tro c h o s a h isp an ic a Simon, 1870
23(32,1sa,20.05-29.06.2020,2,01-
30.09.2020),24(1,20.05-29.06.2020,3 ,1 ,
01-30.09.2020),25(26, 4, 20.05-29.06.2020),38
(1,11-15.04.2018,1 ,11-18.05.2018, 1,
14.04.2021)
+
+
*
Tro c h o sa ro b us ta (Simon, 1876)
34 (1 , 3 , 14.05-02.06.2017)
+
Tro c h o sa ru ric o la (De Geer, 1778)
9 (1 , 19.09.2017), 23 (2 , 1 , 20.05-
29.06.2020, 2 , 30.06-14.07.2020), 24 (2 ,
30.06-14.07.2020, 1 , 01-30.09.2020), 25 (1 ,
3, 20.05-29.06.2020, 1 , 9 , 30.06-
14.07.2020, 5 , 2 , 01-30.09.2020), 32
(Jurinich & Drensky, 1917, Drensky, 1936), 38
(2 , 1 , 16-30.03.2018)
+
+
Tro c h o sa te rric o la Thorell, 1856
11 (1 , 19.04.2017), 32 (Jurinich & Drensky,
1917, Drensky, 1936), 40 (1 , 10-11.05.2021)
+
+
*
Xer o lyc o s a m in iata (C. L. Koch, 1834)
24 (1 , 1 , 30.06-14.07.2020), 29a (1 , 1
, ), 38 (1 , 25.04-05.05.2019, 2 , 11-
21.05.2019, 1 , 14-20.05.2020, 1 , 25-
31.07.2020), 44 (1 , 30.05.2021), 45b (3 ,
09.05.2019)
+
+
Xer o lyc o s a n e m o ralis (Westring, 1861)
29a (Drensky, 1936), 32 (Drensky, 1936). 38 (1
, 25.05.2021)
+
+
Mimetidae
*
Ero aphana (Walckenaer, 1802)
38 (1 , 01.06.2019, 1 , 28.05-08.06.2019)
+
***
Ero ko re an a Paik, 1967
33 , (2 , 16.07.2022)
*
Ero tu b e rc ul ata (De Geer, 1778)
8 (1 , 17.09.2017)
+
Miturgidae
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
94
*
Zo r a ar m illata Simon, 1878
38 (1 , 23.08.2019), 51 (1 , 23.05.2021)
+
*
Zo r a m an ic ata Simon, 1878
23 (1 , 20.05-29.06.2020), 38 (1 ,
21.05.2019, 1 , 30.05.2021)
+
+
*
Zo ra n em o r alis (Blackwall, 1861)
25 (1 , 01-30.09.2020)
+
*
Zo r a p aralle l a Simon, 1878
38 (1 , 13.04.2021)
+
Oecobiidae
*
O ec o b ius m ac u latu s Simon, 1870
4 (1 , 30.06.2021), 26b (1 , 05.06.2020), 26c
(1 , 19.07.2021), 38 (1 , 11.06.2019, 2 ,
18-20.06.2020, 1 , 02.08.2020, 1 ,
19.06.2021)
+
Oxyopidae
*
O xyo p e s h e te ro p h th alm us (Latreille, 1804)
19 (1 , 1 sa, 1 sa, 26.05.2011), 33 (1 ,
16.06.2019), 38 (1 , 14-20.05.2020), 44 (1 ,
11.07.2020), 52 (1 , 29.05.2019)
+
+
*
O xyo p e s lin e atu s Latreille, 1806
1 (1 , 4 sa, 3 sa, 03-04.06.2017), 19 (13 j,
26.05.2011), 33 (1 , 16.06.2019, 1 , 1 ,
18.07.2021), 38 (2 j, 16-30.03.2018, 2 j, 11-
15.04.2018, 1 , 1 sa, 27-29.05.2018, 1 , 03-
10.07.2018, 1 , 17.06.2020)
+
+
Philodromidae
*
Ph ilo dr o m us alb idu s Kulczyński, 1911
9 (1 , 17.06.2021)
+
*
Ph ilo dr o m us aur eo lu s (Clerck, 1757)
38 (1 , 24.03, 30.05.2019)
+
*
Ph ilo dr o m us c o llin us C. L. Koch, 1835
42 (1 , 13.06.2019)
+
*
Ph ilo dr o m us lo n gip alp is Simon, 1870
2(1,20.06.2021),11(1,09.07.2021),23(1,
30.06-14.07.2020),33(3,16.06.2019,1,
04.07.2019,2 ,2 ,18.07.2021, 2,25.07.2021,1
,21.08.2021),38(1,03-10.07.2018,1 ,1 ,
13-14.06.2019,1 ,18.06.2019,1,25.06.2019,2
,18-26.06.2019,1 ,13.06.2020,1,20.06.2020),
48(1, 21.06.2019)
+
+
*
Ph ilo d ro m us m ar m o ratu s Kulczyński, 1891
40 (1 , 04.06.2021), 44 (1 , 30.05.2021)
+
*
Ph ilo dr o m us p o ec ilu s (Thorell, 1872)
6 (1 j, 18.02.2022), 20 (1 , 18.04.2017), 38 (1
, 24.03.2019, 1 , 20.04.2018, 1 ,
22.06.2020), 40 (1 , 17.06.2020)
+
+
*
Ph ilo dr o m us ru fu s Walckenaer, 1826
38 (2 , 14-20.05.2020)
+
*
Th anat us ar e nar ius L. Koch, 1872
25 (1 , 20.05-29.06.2020), 40 (1 ,
27.05.2021),
+
+
*
Th anatus atratus Simon, 1875
25 (1 , 30.06-14.07.2020), 33 (1 ,
21.08.2021), 38 (1 , 1 , 13-15.06.2019), 39
(1 , 17.07.2021), 43a (1 , 27.06.2021), 45b (1
, 13.06.2019)
+
+
*
Th anat us f o rm ic in us (Clerck, 1757)
38 (1 , 23.06.2018)
+
*
Th anat us v u lg aris Simon, 1870
1(1,1 sa,03-04.06.2017),11(1,1,
09.07.2021)
+
*
Tib e llu s m ac e llu s Simon, 1875
1 (1 , 03-04.06.2017)
+
*
Tib e llus o b lo ng u s (Walckenaer, 1802)
6 (1 sa, 14.05.2021), 19 (7 , 8 , 10 j,
26.05.2011), 38 (1 , 11.11.2018, 1 ,
13.06.2020, 1 , , 09.05.20211 , 06.06.2021)
+
+
Pholcidae
Ho lo c n e m u s p luc h e i (Scopoli, 1763)
7 (1 , 1 j, 02-03.2018), 11 (1 , 2 j,
17.06.2021), 13 (1 , 25.08.1928), 26c (1 ,
27.04.2018), 29a (Drensky, 1913 sub H.
riv u latus , Drensky, 1936)
+
*
Ph o lc us o p ilio nid e s (Schrank, 1781)
38 (1 , 2 j, 16-30.03.2018, 1 , 21-26.10.2018)
+
*
Ph o lc us p hal ang io ide s (Fuesslin, 1775)
7 (1 , 02-03.2018), 38 (1 , 16-30.03.2018, 1
, 11-15.04.2018)
+
+
Naum o v a &G e n c he v
95
*
Spe rm o p h o ra s e no c u lata (Dugès, 1836)
26c (19.07.2021, 1 ), 38 (3 j, 16.03.-
30.04.2018, 1 , 25.04-05.05.2019, 1 j, 11-
21.05.2019), 50 (1 , 26.05.2019)
+
+
Phrurolithidae
*
Ph ru ro lith u s f e stiv u s (C. L. Koch, 1835)
25(1, 30.06-14.07.2020),34(2,14.05-
02.06.2017),38(1,8sa,11-15.04.2018,1 ,27-
29.05.2018,1 ,1 ,03-10.07.2018, 1, 08-
21.04.2019)
+
+
*
Ph rur o lithu s p ullatu s Kulczyński, 1897
38(1sa,1sa,16-30.03.2018,1,1,26-
30.04.2018,1,21-26.10.2018,1,1sa,11-
18.05.2018,1,28.05-08.06.2019,2,2,
23.08.2019)
+
*
Ph rur o lithu s szily i Herman, 1879
5 (1 , 21.04.2022), 38 (1 , 1 sa, 11-
18.05.2018)
+
+
Pisauridae
*
Pis aura m irab ilis (Clerck, 1757)
23 (1 , 20.05-29.06.2020, 1 j,01-30.09.2020),
24 (1 , 1 j, 20.05-29.06.2020), 25 (1 , 1 ,
20.05-29.06.2020), 33 (22 , 16.06.2019), 38 (1
, 1 sa, 26-30.04.2018, 1 , 11-18.05.2018)
+
+
Salticidae
*
Ae lur illus v - in sig n itu s (Clerck, 1757)
40 (1 , 29.04.2019)
+
Af raf lac illa e pib le m o id e s (Chyzer, 1891)
38 (1 , 03.10.2018, 1 , 1 , 25.03.2020, 1 ,
05.08.2020, 1 , 14.08.2020, 1 j, 25.09.2020),
47 (NAUMOVA e t al. 2021, 1 , 08.07.2019, 1
, 1 , 25.03.2020, 1 , 1 , 05.08.2020, 1 j,
25.09.2020)
+
*
Att ulu s dis tin g u e nd us (Simon, 1868)
38 (1 , 11-18.05.2018)
+
Att ulus s altat o r (O. Pickard-Cambridge, 1868)
29a (Drensky, 1913 sub Attu s s., Drensky, 1936
sub Sittic u s s.)
+
*
Ballu s c h aly be iu s (Walckenaer, 1802)
38 (1 , 08-21.04.2019, 1 , 25.04-05.05.2019,
1, 28.05-08.06.2019, 1 , 23.06.2021), 40 (1
, 27.05.2021)
+
Carrh o tu s xanth o g ram m a (Latreille, 1819)
29a (Drensky, 1913 sub P. b ic o lo r, Drensky,
1936), 38 (1 , 21.03.2019)
+
+
Ch alc o sc irt us in fim u s (Simon, 1868)
29a (Jurinitch & Drensky, 1917 sub
Ch alc o sc o it us [sic] (Callie th e ra [sic]) i.)
+
*
Euo p hr ys f ro n talis (Walckenaer, 1802)
9 (1 , 19.09.2017), 38 (1 , 14-20.05.2020)
+
+
*
Euo p h ry s h e rb ig rad a (Simon, 1871)
23 (1 , 30.06-14.07.2020)
+
*
Ev arc ha arc u ata (Clerck, 1757)
1 (1 , 03-04.06.2017), 33 (1 , 25.07.2021),
38 (3 , 11-15.04.2018, 1 , 25.04-05.05.2019,
1, 13.06.2020), 40 (1 , 04.06.2021)
+
+
*
He lio p h anu s aur atus C. L. Koch, 1835
33 (2 , 18.07.2021, 4 , 25.07.2021, 1 ,
12.09.2021), 38 (1 , 11-15.04.2018, 1 ,
25.04-05.05.2019, 1 , 15.09.2019, 1 ,
07.08.2021), 45b (1 , 09.05.2019)
+
*
He lio p h anu s c u pr e us (Walckenaer, 1802)
5 (2 , 21.04.2021), 38 (2 , 11-15.04.2018, 1
, 25.04-05.05.2019)
+
+
*
He lio p h anu s f lav ip e s (Hahn, 1832)
1(1,03-04.06.2017),19(2,1 ,2 sa,
26.05.2011)
+
*
He lio ph an us ko c hii Simon, 1868
2 (1 , 01-04.05.2018, 1 , 14.05.2021), 8 (1
, 3 sa, 20.03.2018), 26c (1 , 19.07.2021),
38 (1 , 27-29.05.2018, 1 , 03-10.07.2018, 1
, 15.08.2018, 1 , 11-21.05.2019, 1 ,
20.05.2021)
+
+
*
He lio p h anu s lin e iv e nt ris Simon, 1868
38 (1 , 23.06.2018)
+
*
He lio p h anu s m e lin us L. Koch, 1867
2 (1 , 30.05.2018), 38 (1 , 1 , 04.06.2020),
49 (2 , 2 , 02.06.2020)
+
+
Re v iew o f t he Spide rs, Ps eud o sc o rp io ns and Sc o rp io ns in th e Re g io n o f Plo v div , S B ulg aria...
96
*
He lio p h anu s p atag iatu s Thorell, 1875
33 (1 , 1 sa, 18.07.2021, 1 , 3 ,
25.07.2021, 1 , 17.08.2021)
+
Ic iu s ham at us (C. L. Koch, 1846)
4 (1 , 05.03.2021, 1 , 21.04.2021), 16
(Naumova et al., 2021)
+
Lep to rc h e st e s siko rs kii Prószyński, 2000
39(Naumovaetal.2021,1 sa,12.04.2020,1 ,
11.07.2020,1 ,19.06.2021,1 ,21.06.2020,1,
01.08.2020,1 , 10.07.2021), 52 (Naumova et al.,
2021)
+
*
Mac aro e ris f lav ic o m is (Simon, 1884)
38 (1 , 04.07.2019)
+
*
Mac aro e ris n id ic o le n s (Walckenaer, 1802)
38 (1 , 03.04.2019, 1 , 25.04.2019, 1 ,
11.05-21.05.2019, 1 , 28.05-08.06.2019)
+
*
Marp iss a m u sc o s a (Clerck, 1757)
29a (Drensky, 1936)
+
*
Marp is sa niv o y i (Lucas, 1846)
38 (1 , 27-29.05.2018, 1 , 14-20.05.2020)
+
Marp issa rad iata (Grube, 1859)
32 (Drensky, 1913 sub Marp tu sa r.)
+
*
Me n e m e rus se m ilim b atu s (Hahn, 1829)
2 (1 , 01-04.05.2018), 21 (1 j, 10.10.2021), 26c
(1 , 26.06.2016), 29a (1 , 1 j, 25.08.2021), 38
(4 sa, 16-30.03.2018, 1 , 11-15.04.2018)
+
+
*
Me n e m e rus tae n iatu s (L. Koch, 1867)
38(2sa,16-30.03.2018, 2, 1sa, 1, 03-
10.07.2018,1 ,25.04-05.05.2019, 1, 11.05-
21.05.2019),46a(1,28.09.2018), 52(1,
29.05.2019)
+
?
Ne o n p ic tu s Kulczyński, 1891
2 (Jurinic &Drensky,1917),8 (Jurinic &Drensky,
1917)
+
Pe lle n e s alle g rii Caporiacco, 1935
38 (Naumova et al., 2021)
+
*
Pe lle n e s b re v is (Simon, 1868)
38 (1 , 23.06.2018)
+
*
Pe lle n e s n ig ro c iliat us (Simon, 1875)
38 (1 , 08-21.04.2019)
+
*
Pe lle n e s s e riatu s (Thorell, 1875)
47 (1 , 1 , 13.5.2022)
*
Ph ilae us c h ry so p s (Poda, 1761)
49 (1 sa, 1 sa, 29.09.2018)
+
*
Ph int e lla c as trie s iana (Grube, 1861)
33(1, 30.06.2018,1 ,16.06.2019, 1,
18.07.2021)
+
*
Ph le g ra fas c iata (Hahn, 1826)
2 (1 , 01-04.05.2018), 24 (5 , 2 , 30.06-
14.07.2020), 38 (1 , 11-21.05.2019), 40 (1 ,
29.04.2019), 44 (1 , 09.05.2019)
+
+
Ps e ud e uo p h ry s o b so le ta (Simon, 1868)
2 (1 , 3 j, 18.09.2017, 1 , 01-04.05.2018, 1
, 09.06.2021), 5 (1 , 21.04.2021), 9 (2 j,
18.09.2017), 25 (1 j), 29a (Drensky, 1913, sub
Euo p hr is o ., Drensky, 1936), 38 (1 , 25.04-
05.05.2019), 47 (1 , 05.08.2020)
+
+
*
Ps e ud ic iu s e nc arp atu s (Walckenaer, 1802)
33 (1 , 18.07.2021), 38 (1 , 08.08.2019, 1 ,
20.06.2020, 1 , 28.08.2020), 40 (1 , 10-
11.05.2021)
+
*
Saitis tau ric us Kulczyński, 1904
2 (1 , 01-04.05.2018, 1 , 27.07.2018), 5 (1
, 26.03.2021), 6 (1 j, 18.02.2022), 38 (3 , 1
, 14 j, 16-30.03.2018, 2 , 11-15.04.2018, 1
, 27-29.05.2018, 1 , 26.05.2020)
+
+
*
Saltic u s u nic o lo r (Simon, 1868)
2 (1 , 09.06.2021), 5 (1 , 21.04.2021), 38 (1
, 25.05.2019, 1 , 1 , 14-20.05.2020, 1 ,
20.06.2020, 1 , 23.06.2021)
+
+
**
Talav e ra ap e rta (Miller, 1971)
38 (1 , 1 , 15.05.2021)
+
*
Talav e ra lo g u no v i Kovblyuk &Kastrygina, 2015
38 (1 , 1 , 15.05.2021)
+
Scytodidae
#
Sc yt o d e s sp .
38 (1 , 1 , 19.06.2021)
+
*
Sc yt o de s t ho rac ic a (Latreille, 1802)
2(1,2 ,4 j,18.09.2017,1 ,01-04.05.2018),6
(1j,18.02.2022),9(1,18.09.2017),23(1,20.05-
29.06.2020),25(1,1, 30.06-14.07.2020),38(1
,04.04.2018),42(1,08.09.2018)
+
+
Naum o v a &G e n c he v
97
Tetragnathidae
*
Me te llin a m e ng e i (Blackwall, 1869)
5 (1 , 21.04.2021), 38 (1 , 2 , 11-
15.04.2018)
+
+
*
Me te llin a se g m e n tat a (Clerck, 1757)
35 (1 , 04.1988)
+
*
Pac h yg n ath a c le rc ki Sundevall, 1823
5 (1 , 26.03.2021), 25 (1 , 30.06-14.07.2020),
33 (1 , 18.07.2021), 38 (1 , 16-30.03.2018),
40 (1 , 10-11.05.2021)
+
+
*
Pac h y g nat ha d e g e e ri Sundevall, 1830
23(1, 4, 20.05-29.06.2020, 3, 3,30.06-
14.07.2020,1 ,2 ,01-30.09.2020),24(1,
20.05-29.06.2020,1,1 ,30.06-14.07.2020), 25(2
,30.06-14.07.2020),31(1, 22.02.1988),38(3,
3j,21-26.10.2018,1 ,18-26.06.2019,1,04-
14.07.2019,4 ,1 ,23.08.2019, 1,30.08.2020)
+
+
*
Te tr ag nat ha e xte ns a (Linnaeus, 1758)
1 (1 , 1 , 03-04.06.2017), 33 (1 ,
17.08.2021), 38 (2 , 14-20.05.2020), 40 (1 ,
10-11.05.2021, 1 , 2 , 27.05.2021)
+
+
*
Te tr ag nat ha m o n tan a Simon, 1874
33 (1 , 18.07.2021), 38 (1 , 1 , 27-
29.05.2018), 40 (1 , 10-11.05.2021)
+
*
Te tr ag nat ha n ig rita Lendl, 1886
33 (1 , 18.07.2021, 1 , 25.07.2021)
+
*
Te tr ag nat ha o b tu sa C. L. Koch, 1837
13 (3 , 25.08.1928), 33 (1 , 18.07.2021), 38
(1 , 20.06.2020), 50 (1 , 26.05.2019)
+
+
*
Te tr ag nat ha p inic o la L. Koch, 1870
38 (1 , 28.05.2021)
+
Theridiidae
*
Asag e n a m e rid io n alis Kulczynski, 1894
38 (1 , 18.02.2019)
+
*
Asag e n a ph ale rat a (Panzer, 1801)
2 (2 , 1 sa, 18.09.2017), 9 (1