Available via license: CC BY-NC-ND 4.0
Content may be subject to copyright.
Ecologica Montenegrina, 58, 2022, 42-49
https://zoobank.org/urn:lsid:zoobank.org:pub:19BB0564-4274-4D98-BA6A-4CB7C9ADB4FE
Four new species of the genera Dryophthoroides Roelofs, 1879 and
Nephius Pascoe, 1885 (Coleoptera, Curculionidae) from Oriental and
Papuan Regions
ANDREI A. LEGALOV1,2,3*
1 Institute of Systematics and Ecology of Animals, SB RAS, Frunze Street 11, Novosibirsk, 630091, Russia.
2 Altai State University, Lenina Street 61, Barnaul, 656049, Russia.
3 Tomsk State University, Lenina Prospekt 36, 634050 Tomsk, Russia.
* Corresponding author. E-mail: fossilweevils@gmail.com, https://orcid.org/0000-0001-7347-8169
Received 9 October 2022 │ Accepted by V. Pešić: 25 October 2022 │ Published online 26 October 2022.
Abstract
Four new species of the genera Dryophthoroides Roelofs, 1879 and Nephius Pascoe, 1885 from the tribe
Stromboscerini of the subfamily Dryophthorinae are described and illustrated. Dryophthoroides telnovi Legalov, sp.
nov. from New Guinea (Star Mountains) is similar to D. seftoni (Zimmermann, 1944) bit differs in the shorter and
thicker rostrum and narrower pronotum. It is distinguish from D. beccarii (Pascoe, 1885) in the smaller body size,
thicker rostrum, coarser punctured pronotum, not subparallel elytra and narrower elytral interstriae. Dryophthoroides
insularis Legalov, sp. nov. from North Moluccas (Bacan Islands) differs from D. telnovi Legalov, sp. nov. in the wider,
laterally weakly rounded pronotum with a more finely punctured pronotal disc, the stronger widened interstria five at
apex, the smaller punctures in elytral striae, the wider elytral interstriae and the longer and thinner rostrum. Nephius
continentalis Legalov, sp. nov. from Laos (Hua Phan Prov.) differs from N. kalimantanensis Legalov, 2022 in the
thicker and shorter rostrum, the pronotum lacking the sharp preapical constriction, the elytra with the greatest width
postmedium, the medially not depressed lateral sides of the pronotum, and the obsolete humeri. Nephius philippinensis
Legalov, sp. nov. from the Philippines (Mindanao Is., Bukidnon Prov.) differs from N. mindanaoensis Legalov, 2020 in
the sparsely punctate pronotum with a median longitudinal groove and lacking four premedian pilose ridges, the elytral
interstriae with weak pilose ridges, and the less curved rostrum.
Key words: Biodiversity, Curculionoidea, Dryophthorinae, Stromboscerini, Laos, Philippines, Indonesia.
Introduction
The tribe Stromboscerini belongs to subfamily Dryophthorinae of the family Curculionidae. The tribe
includes thirteen extant and two fossil genera (Grebennikov 2018; Legalov 2016; Legalov et al. 2019;
Nazarenko and Perkovsky 2009). Almost all modern species are distributed in the Oriental Region, and only
the genus Stromboscerus Schoenherr, 1838 in Madagascar and an undescribed form are listed for Uganda
Ecologica Montenegrina 58: 42-49 (2022)
This journal is available online at: www.biotaxa.org/em
https://dx.doi.org/10.37828/em.2022.58.4
LEGALOV
Ecologica Montenegrina, 58, 2022, 42-49 43
(Grebennikov 2018). This work continues the study of the tribe Stromboscerini by Legalov (2019, 2020b,
2021a, 2021b, 2021c, 2021d, 2022a, 2022b).
In this paper, the new species of the genera Dryophthoroides Roelofs, 1879 and Nephius Pascoe,
1885 from Laos, the Philippines and Indonesia are described.
Material and methods
Type specimens are kept in the ISEA = Institute of Systematics and Ecology of Animals (Russia:
Novosibirsk) and NME = Naturkundemuseum (Germany: Erfurt).
Descriptions, body measurements, and photographs, were prepared using the Zeiss Stemi 2000-C
dissecting stereomicroscope.
The terminology of the weevil body is according to Lawrence et al. (2010).
The systematics of studied taxa are based on the works of Grebennikov (2018) and Legalov (2020a).
Systematics
Insecta: Coleoptera: Curculionidae: Dryophthorinae: Stromboscerini
Genus: Dryophthoroides Roelofs, 1879
Dryophthoroides telnovi Legalov, sp. nov.
https://zoobank.org/urn:lsid:zoobank.org:act:2413A957-AA3E-41C2-BA3D-09CCDD4BDB46
(Figs. 1A, 1E)
Type material: Holotype, female (NME), INDONESIA: New Guinea, Papua Prov., Star Mountains, 22-27
km SSE of Oksibil, 5 km around Beringin vill., 5°5’01”S, 140°43’27”E, 250-320 m, primary and secondary
lowland montane rainforest, 12-14.III.2018, D. Telnov.
Description. Female: Body black, subglabrous. Antennae, tarsi and uncus red-brown. Head nearly
conical. Rostrum subcylindrical, curved, about 0.9 times as long as pronotum, about 2.7 times as long as
wide at apex, about 2.8 times as long as wide in middle, about 2.9 times as long as wide at base, densely
punctate, with middle ventral carina. Apex of rostrum nearly smooth. Eyes large, coarsely faceted, not
protruding from contour of head, linear, narrowly separated beneath. Forehead flat, about 0.7 times as
narrow as rostrum base width, finely punctate. Temples short, punctate. Antennal scrobes directed ventrally
to base of rostrum. Antennae short, inserted before middle of rostrum, reaching pronotum. First antennomere
elongate conical, 2.4 times as long as wide at apex, nearly extending towards eyes. Second antennomere
suboval, about 1.4 times as long as wide at apex, about 0.4 times as long as and 0.7 times as narrow as first
antennomere. Third antennomere conical, equal in length and width, 0.7 times as long as and subequal in
wide to second antennomere. Fourth and fifth antennomeres wide-conical. Fourth antennomere 0.5 times as
long as wide at apex, about 0.6 times as long as and about 1.1 times as wide as third antennomere. fifth
antennomere 0.4 times as long as wide at apex, 0.9 times as long as and about 1.1 times as wide as fourth
antennomere. Antennal club obliquely truncate, apically tomentose, 1.5 times as long as wide in middle,
nearly equal to antennomeres second- fifth combined. Pronotum nearly bell-shaped, about 1.3 times as long
as wide at apex, little longer than wide in middle and at base. Disk slightly flattened, quite coarsely and
densely punctate. Sides nearly straight. Scutellum small, narrow, immersed. Elytra suboval, about 1.7 times
as long as pronotum, about 1.4 times as long as wide at base, about 1.3 times as long as wide in middle,
about 1.7 times as long as wide at apical fourth. Humeri flattened. Elytral striae distinct. Ninth stria merges
with tenth stria near metacoxa. Interstriae weak convex, with row of points, distinctly narrower than striae.
Precoxal portion of prosternum long, slightly shorter than procoxal cavity. Postcoxal portion of prosternum
short, 0.5 times as long as precoxal portion. Procoxal cavities contiguous. Metaventrite weakly convex,
punctate, about 1.2 times as long as metacoxal cavity. Abdomen convex, densely punctate. First and second
ventrites fused, equal in length. First ventrite about 0.7 times as long as metacoxal cavity. Third and fourth
ventrites subequal in length. Third ventrite about 0.6 times as long as second ventrite. Fifth ventrite 2.5 times
as long as fourth ventrite. Procoxae large, spherical. Femora weakly thickened and flattened, lacking teeth,
densely punctate. Femora nearly straight, with lateral carinae, with large uncus and two groups of setae at
FOUR NEW SPECIES OF DRYOPHTHOROIDES AND NEPHIUS
44
apex. Tarsi long. First - third tarsomeres conical, with erect setae ventrally. Second tarsomere shorter than
first tarsomere. Third tarsomere slightly narrower than first tarsomere. Fifth tarsomere elongate. Tarsal claws
free, divergent. Total body length (without rostrum) 4.1 mm. Length of rostrum 1.1 mm.
Differential diagnosis. The new species is similar to D. seftoni (Zimmermann, 1944) from Papua
New Guinea bit differs in the shorter and thicker rostrum and the narrower pronotum. It can be distinguished
from D. beccarii (Pascoe, 1885) from Indonesian New Guinea in the smaller body size, the thicker rostrum,
the coarser punctured pronotum, the not subparallel elytra and the narrower elytral interstriae.
Etymology. Patronymic. In honour of Dmitry Telnov (Natural History Museum, London, United
Kingdom), who studies the biodiversity of New Guinea beetles.
Localisation. New Guinea: Star Mountains.
Dryophthoroides insularis Legalov, sp. nov.
https://zoobank.org/urn:lsid:zoobank.org:act:89C86230-FA2B-4B1C-9E88-C9304AAC3C57
(Figs. 1B, 1C, 1E, 1F)
Type material: Holotype, male (ISEA), INDONESIA: N Moluccas, Bacan Isl., SE slopes of Mt. Sibela, 5
km SE of Makian vill., 500-750 m, 2-12.V.2008, St. Jakl.
Description. Male: Body black, glabrous. Antennae, tarsi and uncus red-brown. Head nearly conical.
Rostrum subcylindrical, weakly curved, about 0.8 times as long as pronotum, 2.4 times as long as wide at
apex, about 2.5 times as long as wide in middle, about 2.9 times as long as wide at base, densely and quite
coarsely punctate, with middle ventral carina. Rostral apex smooth. Eyes large, coarsely faceted, not
protruding from contour of head, linear, narrowly separated beneath. Forehead flattened, about 0.6 times as
narrow as rostrum base width, punctate. Temples short, punctate. Antennal scrobes directed ventrally to base
of rostrum. Antennae short, inserted near middle of rostrum, reaching pronotum. First antennomere elongate
conical, about 4.3 times as long as wide at apex, nearly reaching eyes. Second antennomere suboval, about
1.4 times as long as wide at apex, about 0.3 times as long as and 0.9 times as narrow as first antennomere.
Third antennomere conical, equal in length and width, 0.6 times as long as and about 0.9 times as narrow as
second antennomere. Fourth and fifth antennomeres wide-conical. Fourth antennomere 0.5 times as long as
wide at apex, about 0.7 times as long as and about 1.3 times as wide as third antennomere. Fifth antennomere
about 0.4 times as long as wide at apex, subequal in length and about 1.1 times as wide as fourth
antennomere. Antennal club obliquely truncate, apically tomentose, 1.7 times as long as wide in middle,
equal to second-fifth antennomeres combined. Pronotum nearly bell-shaped, about 1.6 times as long as wide
at apex, equal to wide in middle and slightly longer than wide at base. Disk weakly flattened, quite coarsely
and densely punctate. Sides weakly curved. Scutellum small, narrow, immersed. Elytra suboval, 2.0 times as
long as pronotum, 1.5 times as long as wide at base and in middle, about 2.6 times as long as wide at apical
fourth. Humeri flattened. Elytral striae distinct. Ninth striae striae merge with striae 10 near metacoxa.
Interstriae weak convex, with row of points, distinctly wider than striae. Interstriae 5 widened at apex.
Precoxal portion of prosternum long, equal to procoxal cavity. Postcoxal portion of prosternum short, about
0.3 times as long as precoxal portion. Procoxal cavities contiguous. Metaventrite weakly convex, punctate,
about 1.8 times as long as metacoxal cavity. Abdomen convex, finely punctate. First and second ventrites
fused. First ventrite about 0.6 times as long as metacoxal cavity. Second ventrite about 1.4 times as long as
first ventrites. Third and fourth ventrites equal in length. Third ventrite 0.5 times as long as second ventrite.
Fifth ventrite 1.4 times as long as fourth ventrite. Procoxae spherical. Femora weakly thickened, densely
punctate. Femora nearly straight, with lateral carinae, uncinate. Tarsi quite long. First - third tarsomeres
conical, with erect setae ventrally. Second tarsomere shorter than first tarsomere. Third tarsomere slightly
longer than first tarsomere. Fifth tarsomere elongate. Tarsal claws free, divergent. Total body length (without
rostrum) 4.1 mm. Length of rostrum 1.0 mm.
Differential diagnosis. The new species differs from D. telnovi Legalov, sp. nov. in the wider,
laterally weakly rounded pronotum with the more finely punctured pronotal disc, the stronger widened
interstria 5 at apex, the smaller punctures in elytral striae, the wider elytral interstriae and the longer and
thinner rostrum.
Etymology. From the Latin ‘insular’.
Localisation. North Moluccas, Bacan Islands.
LEGALOV
Ecologica Montenegrina, 58, 2022, 42-49 45
Figure 1. Dryophthoroides spp., holotypes: A – D. telnovi sp. nov., habitus, dorsal view; B – D. insularis sp. nov.,
habitus, dorsal view; C – D. insularis sp. nov., aedeagus, dorsal view; D – D. telnovi sp. nov., habitus, lateral view; E –
D. insularis sp. nov., habitus, lateral view; F – D. insularis sp. nov., aedeagus, lateral view. Scale bar = 1.0 mm for A,
B, D, El 0.5 mm for C, F.
FOUR NEW SPECIES OF DRYOPHTHOROIDES AND NEPHIUS
46
Genus: Nephius Pascoe, 1885
Nephius continentalis Legalov, sp. nov.
https://zoobank.org/urn:lsid:zoobank.org:act:FC85F2DF-CC4A-40CF-B7DB-6A056AFE65DA
(Figs. 2A, 2C)
Type material: Holotype, female (ISEA), LAOS: NE Laos, Hua Phan Prov., Phu Phan Mountains, 20°12′N,
103°59′E, 1200-1600 m, 10-22.V.2011, St. Jakl.
Description. Female: Body black, with matted pubescence. Antennae, apex of tibiae and tarsi
brownish. Head subconical. Rostrum long, equal in length to pronotum, 4.8 times as long as wide at apex and
at midlength, 3.6 times as long as wide at base, evenly curved, densely punctate. Apex of rostrum punctate,
without pubescence. Eyes large, coarsely faceted, not protruding from contour of head, sublinear, contiguous
ventrally. Forehead flattened, about 1.1 times as long as rostrum base width, glabrous. Antennal scrobes
directed ventrally to base of rostrum. Antennae inserted beyond of rostrum middle. Scape long, 4.8 times as
long as wide at apex, not reaching eye. Funicle sex-segmented. Second antennomere suboval, about 1.8
times as long as wide at apex, about 0.2 times as long as and 0.7 times as narrow as scape. Third
antennomere conical, 1.7 times as long as wide at apex, 0.9 times as long as and about 0.9 times as narrow as
second antennomere. Fourth-seventh antennomeres wide-conical. Third and fourth antennomeres subequal in
width. Fourth antennomere 0.6 times as long as wide at apex, about 0.3 times as long as third antennomere.
Fifth and sixth antennomeres subequal. Fifth antennomere about 0.6 times as long as wide at apex, about 1.3
times as long as and about 1.1 times wide as Fourth antennomere. Seventh antennomere 0.7 times as long as
wide, 1.2 times as long as and about 1.1 times as wide as sixth antennomere. Antennal club compact, about
1.7 times as long as wide, 0.6 times as long as second-seventh antennomeres combined, with subconical
tomentose apex. Pronotum campanulate, 1.7 times as long as wide at apex, about 1.1 times as long as wide at
midlength and at pronotal base, without sharp preapical constriction. Pronotal disk weakly convex dorsally,
sparsely punctate, without median carina. Intervals between points larger than their diameter. Sides of the not
depressed in middle. Maximum width before apex and at base. Base of pronotum 0.9 times as narrow as base
of elytra. Scutellum small, suboval. Elytra subparallel, 1.9 times as long as wide at base, 1.7 times as long as
wide at midlength, 3.5 times as long as wide at apical fourth, 2.0 times as long as pronotum. Humeri
completely flattened. Greatest width of elytra behind middle. Elytral striae distinct. Stria 9 short, fused with
stria 10 at level of metacoxae. Interstriae convex, wide, slightly wider than striae, with pilose pustules.
Prosternum punctate, with distinct postocular lobes. Precoxal portion of prosternum equal in length to
procoxal cavity. Postcoxal portion of prosternum 0.5 times as long as precoxal portion. Procoxal cavities
contiguous. Mesocoxal cavities narrowly separated. Metaventrite 1.7 times as long as length of metacoxa,
weakly convex, sparsely punctate. Abdomen weakly convex ventrally, punctate. First ventrites about 0.9
times as long as length of metacoxa. Second ventrite 1.2 times as long as First ventrites. Third ventrite about
0.8 times as long as second ventrite. Fourth ventrite equal in length to third ventrite. Fifth ventrite convex,
1.8 times as long as fourth ventrite. Procoxae subconical. Mesocoxae spherical, narrowly separated.
Metacoxae transverse. Femora slightly thickened, without tooth. Tibiae weakly curved, with large uncus.
Tarsi long. First - third tarsomeres conical, with erect setae ventrally. Fifth tarsomere elongate. Tarsal claws
free, divergent. Total body length (without rostrum) 4.6 mm. Length of rostrum 1.4 mm.
Differential diagnosis. The new species differs from N. kalimantanensis Legalov, 2022 from
Kalimantan (Indonesia) in the thicker and shorter rostrum, the pronotum lacking sharp preapical constriction,
the elytra with maximum width postmedium, the lateral sides of the pronotum lacking median depression,
and the completely flattened humeri.
Etymology. From the Latin ‘continentalis’ (continental).
Localisation. Laos, Hua Phan Prov.
Nephius philippinensis Legalov, sp. nov.
https://zoobank.org/urn:lsid:zoobank.org:act:EF98C8E9-C666-4C5F-A1BD-9517CADF7E6E
(Fig. 2B)
Type material: Holotype, female (ISEA), PHILIPPINES: Mindanao, Bukidnon, Kalatungan, IX.2014.
Paratype, female (ISEA), Bukidnon, Panamokan, XI.2016.
LEGALOV
Ecologica Montenegrina, 58, 2022, 42-49 47
Figure 2. Nephius spp., holotypes, habitus: A – N. continentalis sp. nov., dorsal view; B – N. philippinensis sp. nov.,
dorsal view; C – N. continentalis sp. nov., lateral view. Scale bar = 1.0 mm.
Description. Female: Body black, with matted pubescence. Antennae, apex of tibiae and tarsi reddish-
brown. Head nearly conical. Mandibles not large. Rostrum long, 1.1 times as long as pronotum, 4.9-5.1 times
as long as wide at apex, 3.9-4.8 times as long as wide at midlength, 3.7-4.5 times as long as wide at base,
curved. Apex of rostrum smooth. Eyes large, not protruding from contour of head, linear, narrowly separated
FOUR NEW SPECIES OF DRYOPHTHOROIDES AND NEPHIUS
48
beneath. Forehead wide, flattened, 1.1 times as long as rostrum base width, densely punctate. Antennal
scrobes directed ventrally to base of rostrum. Antennae inserted in apical fourth of rostrum. Scape long, 4.6
times as long as wide at apex, not reaching eye. Funicle 6-segmented. Second-seventh antennomeres conical.
Second antennomere about 1.6 times as long as wide at apex, about 0.2 times as long as and about 0.6 times
as narrow as scape. Third antennomere about 1.8 times as long as wide at apex, subequal in length and about
0.9 times as long as second antennomere. Fourth antennomere about 0.8 times as long as wide at apex, about
0.4 times as long as and same width as third antennomere. Fifth antennomere about 0.7 times as long as
wide, equal in length and about 1.1 times as wide as fourth antennomere. Sixth antennomere about 0.6 times
as long as wide, about 1.2 times as long as and about 1.2 times as wide as fifth antennomere. Seventh
antennomere about 0.6 times as long as wide, equal in length and about 1.1 times as wide as sixth
antennomere. Antennal club compact, about 2.1 times as long as wide, about 0.7 times as long as second-
seventh antennomeres combined, with subconical tomentose apex. Pronotum campanulate, 1.2-1.5 times as
long as wide at apex, slightly longer than wide at midlength, about 1.1 times as long at pronotal base.
Pronotal disk weakly convex dorsally, quite sparsely punctate, with two small pilose ridges at base and with
median longitudinal groove. Intervals between points larger than their diameter. Sides nearly straight. Base
of pronotum about 0.7 times as narrow as base of elytra. Scutellum small, triangular. Elytra nearly
subparallel, 1.7-1.8 times as long as wide at base and at midlength, about 2.2 times as long as wide at apical
fourth, 2.3-2.5 times as long as pronotum. Humeri weakly convex. Elytral striae distinct. Ninth stria short,
fused with tenth stria at level of metacoxae. Interstriae weakly convex, wide, 3.0-3.5 times as wide as striae
width. Odd interstriae with row of weak and short longitudinally oriented and pilose ridges. Prosternum
punctate, with strong postocular lobes. Precoxal portion of prosternum long, equal to procoxal cavity.
Postcoxal portion of prosternum about 0.4 times as long as procoxal cavity. Procoxal cavities contiguous.
Mesocoxal cavities narrowly separated. Metaventrite 2.2 times as long as length of metacoxa, weakly
convex, sparsely punctate. Abdomen weakly convex ventrally, punctate. First ventrite 0.6 times as long as
length of metacoxa. Second ventrite 1.2 times as long as first ventrite. Third ventrite about 0.6 times as long
as second ventrite. Fourth ventrite slightly longer than third ventrite. Fifth ventrite 1.9 times as long as fourth
ventrite. Procoxae conical. Mesocoxae spherical, narrowly separated. Metacoxae transverse. Femora slightly
thickened, without tooth. Tibiae weakly curved, with large uncus. Tarsi long. First - third tarsomeres conical,
with erect setae ventrally. Fifth tarsomere elongate. Tarsal claws free, divergent. Total body length (without
rostrum) 5.1-5.8 mm. Length of rostrum 1.5-1.7 mm.
Differential diagnosis. The new species differs from N. mindanaoensis Legalov, 2020 from
Mindanao (Philippines) in the sparsely punctate pronotum with a median longitudinal groove and without
four pilose ridges before the middle, elytral interstriae with weak pilose ridges, and weaker curved rostrum.
Etymology. Named after the Philippines.
Localisation. Philippines, Mindanao Is., Bukidnon Prov.
Acknowledgements
The author thanks D. Telnov (Natural History Museum, London, United Kingdom), for the opportunity to
study material from Indonesia, and anonymous reviewers for the valuable comments that improved the
manuscript.
References
Grebennikov, V.V. (2018) Dryophthorinae weevils (Coleoptera: Curculionidae) of the forest floor in
Southeast Asia: illustrated overview of nominal Stromboscerini genera. Zootaxa, 4418 (2), 121–135.
https://doi.org/10.11646/zootaxa.4418.2.2
Lawrence, J.F., Beutel, R.G., Leschen, R.A.B., Slipinsky, S.A. (2010). Chapter 2. Glossary of Morphological
Terms. Handbook of Zoology. Arthropoda: Insecta. Tb. 40: Coleoptera (Beetles). Vol. 2:
Morphology and Systematic (Elateroidea, Bostrichformia, Cucujiformia partim), P. 9–20.
Legalov, A.A. (2016) Two new genera and four new species of fossil weevils (Coleoptera: Curculionoidea)
in Baltic amber. Entomologica Fennica, 27 (2), 57–69. https://doi.org/10.33338/ef.59124
LEGALOV
Ecologica Montenegrina, 58, 2022, 42-49 49
Legalov, A.A. (2019) A new species of the genus Dryophthoroides Roelofs, 1879 (Coleoptera:
Curculionidae) from Philippines. Far Eastern Entomologist, 375, 20–24.
https://doi.org/10.25221/fee.375.4
Legalov, A.A. (2020a) Annotated key to weevils of the world. Part 4. Subfamilies Erirhininae,
Dryophthorinae and Cossoninae (Curculionidae). Ukrainian Journal of Ecology, 10 (2), 319–331.
https://doi.org/10.15421/2020_104
Legalov, A.A. (2020b) A new species of the genus Nephius Pascoe, 1885 (Coleoptera, Curculionidae) from
Philippines. Baltic Journal of Coleopterology, 20 (1), 15–18.
Legalov, A.A. (2021a) A new species of the genus Tasactes Faust, 1894 (Coleoptera, Curculionidae) from
Nepal. Ecologica Montenegrina, 40, 128–132. https://doi.org/10.37828/em.2021.40.11
Legalov, A.A. (2021b) A new species of the genus Allaeotes Pascoe, 1885 (Coleoptera, Curculionidae) from
Philippines. Baltic Journal of Coleopterology, 21 (1), 25–29.
Legalov, A.A. (2021c) A new species of the genus Nephius Pascoe, 1885 (Coleoptera, Curculionidae) from
Indonesia. Ecologica Montenegrina, 47, 33–36. http://dx.doi.org/10.37828/em.2021.47.5
Legalov, A.A. (2021d) A new species of the genus Dexipeus Pascoe, 1885 (Coleoptera, Curculionidae) from
the Philippines. Baltic Journal of Coleopterology, 21 (2), 175–180.
Legalov, A.A. (2022a) Orthosinus laosensis Legalov, a new species of the tribe Strombocerini (Coleoptera:
Curculionidae) from North-eastern Laos. Ecologica Montenegrina, 51, 15–18.
https://dx.doi.org/10.37828/em.2022.51.2
Legalov, A.A. (2022b) A new species of the genus Synommatus Wollaston, 1873 (Coleoptera:
Curculionidae) from the Philippines. Far Eastern Entomologist, 462, 1–7.
https://doi.org/10.25221/fee.462.1
Legalov, A.A., Nazarenko, V.Yu., Perkovsky, E.E. (2019) New weevils (Coleoptera: Curculionidae) from
Rovno amber. Paleontological Journal, 53 (10), 1045–1059.
https://doi.org/10.1134/S0031030119100101
Nazarenko, V.Y., Perkovsky, E.E. (2009) A new genus and species of Dryophthorid weevils (Coleoptera,
Dryophthoridae: Stromboscerinae) from the Rovno amber. Paleontological Journal, 43, 1097–1100.
https://doi.org/10.1134/s003103010909010
