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Notes on Longitarsus aphthonoides Weise 1887 and on the species groups of L. lateripunctatus (Rosenhauer 1856) and L. curtus (Allard 1860) (Coleoptera, Chrysomelidae)

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The aim of this study is to deepen the knowledge of five poorly studied Longitarsus species in order to elucidate their taxonomic position within the genus. For each of them further diagnostic characters and many new collecting localities are supplied. One new synonymy and one change of status are proposed: L. callidus Warchalowski 1967 = L. aphthonoides Weise 1887 syn. nov., and L. lateripunctatus personatus Weise 1893 is raised to valid species. The morphologically similar L. curtus (Allard 1860) and L. monticola Kutschera 1863 are also examined, providing new male and female diagnostic characters for their identification, as well as several European and Italian new collecting localities.
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Natural History Sciences. Atti Soc. it. Sci. nat. Museo civ. Stor. nat. Milano, 8 (2): 11-34, 2021 DOI: 10.4081/nhs.2021.515
Notes on Longitarsus aphthonoides Weise 1887 and on the species groups
of L. lateripunctatus (Rosenhauer 1856) and L. curtus (Allard 1860)
(Coleoptera, Chrysomelidae)
Laura Farina
Via A. Stoppani 17/A, I - 23880 Casatenovo, Italia.
E-mail: laura.farina@chrysomelidae.it
© 2021 Laura Farina
Received: 5 February 2021
Accepted for publication: 10 May 2021
Online publication: 29 October 2021
Abstract - The aim of this study is to deepen the knowledge of ve
poorly studied Longitarsus species in order to elucidate their taxonomic
position within the genus. For each of them further diagnostic charac-
ters and many new collecting localities are supplied.
One new synonymy and one change of status are proposed: L. cal-
lidus Warchalowski 1967 = L. aphthonoides Weise 1887 syn. nov., and
L. lateripunctatus personatus Weise 1893 is raised to valid species. The
morphologically similar L. curtus (Allard 1860) and L. monticola Kut-
schera 1863 are also examined, providing new male and female diag-
nostic characters for their identication, as well as several European
and Italian new collecting localities.
Key words: Alticini, European fauna, Galerucinae, Italian fauna,
Mediterranean fauna, taxonomy.
Riassunto - Note su Longitarsus aphthonoides Weise 1887 e sui
gruppi di specie L. lateripunctatus (Rosenhauer 1856) e L. curtus
(Allard 1860) (Coleoptera, Chrysomelidae).
Scopo di questo studio è approfondire la conoscenza di cinque
specie di Longitarsus scarsamente studiate al ne di chiarire la loro
posizione tassonomica all’interno del genere. Per ciascuna di esse ven-
gono forniti ulteriori caratteri anatomici comparativi e numerosi nuovi
dati geonemici.
Vengono proposti una nuova sinonimia: L. callidus Warchalowski
1967 = L. aphthonoides Weise 1887 syn. nov. e il cambio di stato di L.
lateripunctatus personatus Weise 1893 che è elevato a rango di specie.
Per L. curtus (Allard 1860) e L. monticola Kutschera 1863, specie assai
simili, vengono forniti nuovi caratteri diagnostici, sia maschili che fem-
minili, e numerose nuove località europee e italiane.
Parole chiave: Alticini, fauna europea, fauna italiana, fauna medi-
terranea, Galerucinae, tassonomia.
INTRODUCTION
Longitarsus Latreille 1829 (2: 155) (sensu Löbl &
Smetana, 2010) is the world’s largest genus of Alticini,
with about 500 species described. About 150 species are
known for the Palaearctic Region, including 93 species
for Italy only: see Biondi et al. (1995), with the addition
of L. anacardius (Allard 1866) sensu Leonardi & Sassi
(1997).
The aim of this paper is to provide new diagnostic
characters to clarify the related taxonomy of ve spe-
cies of Longitarsus, and to extend the knowledge of their
ranges by listing new collecting localities. The following
taxa are discussed:
Longitarsus aphthonoides Weise 1887
Longitarsus callidus Warchalowski 1967
Longitarsus lateripunctatus (Rosenhauer 1856)
Longitarsus personatus Weise 1893
Longitarsus curtus (Allard 1860)
Longitarsus monticola Kutschera 1863
MATERIALS AND METHODS
All observations were done using binocular micro-
scopes. To set specimens and genitalia on the label, the
author rst gets the specimen wet, after boiling in pure
water; then extracts genitalia with the aid of two pins. For
the spermatheca extraction usually it is necessary to cut
completely the abdomen of the wet specimen, separating it
from the rest of the body, and extract gently the spermath-
eca. For the aedeagus extraction, instead, the author cuts
slightly the abdomen in the middle and extracts carefully
the aedeagus. The aedeagi and spermathecae were xed
with Arabic gum and set on the same label of the speci-
men or on another label below. The spermathecae were
included in DMHF (Dimethyl-hydantoin formaldehyde)
or in entomological glue. The biometric measurements
were taken using an ocular micrometer. Drawings were
taken using an eyepiece equipped with a 20x20 reticule
and manually transferring the observed image on graph
paper. The length of elytra and prothorax were measured
keeping the specimens tilted in order to have the base and
the apex of the elytra (or the fore and the rear edge of pro-
notum) on the same plane (Farina & Leonardi, 2018). The
macrophotos were taken using a full-frame digital camera
with a 25 mm lens.
Collection localities were recorded as written on the
specimens’ labels. Countries and Italian regions were ar-
ranged in a west to east and north to south order; the lo-
calities were listed grouping them according to the region
of provenance and museum or private collections.
12
The specimens included in this study are kept in the
following institutional and private collections:
BMNH: Natural History Museum, London (BMNH-War:
A. Warchalowski collection);
CGCB: Christoph Germann collection, Basel, Switzer-
land;
DFCA: David Furth collection, Alexandria, VA, USA;
FFCJ: Frank Fritzlar collection, Jena, Germany;
FMCT: Fernando Montemurro collection, Taranto, Italy;
HNHM: Hungarian Natural History Museum, Budapest;
LFCC: Laura Farina collection, Casatenovo, Lecco, Italy;
MBCA: Maurizio Biondi collection, University of
L’Aquila, Italy;
MHNB: Naturhistorisches Museum, Basel (MHNB-Heik:
1953 F. Heikertinger collection);
MSNG: Museo Civico di Storia Naturale “G. Doria”,
Genova (MSNG-Bin: G. Binaghi collection; MSNG-
Dod: A. Dodero collection; MSNG-Man: C. Mancini
collection);
MSNM: Museo di Storia Naturale di Milano (MSNM-
Ber: M. Bergeal collection);
MSNVR: Museo Civico di Storia Naturale, Verona;
NHMW: Naturhistorisches Museum, Wien;
RPCR: Riccardo Papi collection, Ragusa, Italy;
SDEI: Senckenberg Deutsches Entomologisches Institut,
Müncheberg;
SZCM: Stefano Zoia collection, Milano, Italy;
ZMB: Museum für Naturkunde, Berlin;
ZSM: Zoologische Staatssammlung München.
Further abbreviations
Al: median lobe of aedeagus length (taken in ventral
view);
Aw: median lobe of aedeagus width (taken in ventral
view);
El: elytral length;
Ew: elytral width (taken together);
lg: lateral groove;
Lt: length of hind tibia in stretched position;
mg: median groove;
(….)m: mean value of….;
n: number of specimens in the sample;
Pl: prothorax length;
Pw: prothorax width;
(!): specimens dissected for genitalia examination;
/: line break within the label; the line break is not applied
in the species determination labels.
//: next label on the same pin.
SPECIES DISCUSSION
Longitarsus aphthonoides Weise 1887 (Figs. 1-3)
Longitarsus aphthonoides Weise, 1887: 206. Reported
locality: Chabarofka.
Heikertinger, 1930: 1326; Csiki & Heikertinger, 1940:
pars 166, 110; Lopatin, 1967: 166; Warchalowski, 1970:
106-107, 114 (Fig. 5) [designation of the Lectotype, typ.
loc.: Ussuri Gebiet, Chabarofka]; Gruev, 1980: 32; Gruev,
1981: 149; Medvedev, 1982: 114-115 (Fig. 34: 2); Gruev,
1987: 56 (Fig. 3); Medvedev & Dubeshko, 1992: 153, 216
(Fig. 30: 10); Gruev, 1994: 81; Gruev & Döberl, 1997:
148-149; Gruev & Döberl, 2005: 75; Warchalowski,
2010: 830, 831 (Fig. 4020).
= Longitarsus callidus Warchalowski, 1967: 59-60. Loc.
typ.: North Iran: Mt. Talysch (syn. nov.)
Leonardi, 1975: 207 (Figs. 27-28, 53), 212 (Fig. 57),
213-214; Fogato & Leonardi, 1980: 46, 53 (Fig. 14); Dog-
uet, 1987: 4(2): 138; Biondi, 1990a: 138; Döberl, 1994:
103, 129 (Figs. 173-175); Doguet, 1994: 224 (Fig. 93f),
228-229, 297 (Fig. 122b); Lucht, 1994: 319; Doguet &
Dufay, 1994: 306-307 (Figs. 5,7); Biondi et al., 1995: 18;
Döberl, 1995b: 53; Warchalowski, 1995a: 152 (Figs. 460-
461), 155 (Fig. 478), 224; Warchalowski, 1996: 45 (Fig. 95-
96), 48 (Fig. 113), 76-77; Rozner, 1996: 255 [“this species
is distributed from Asia Minor through Southeast Europe
to Austria. Its occurrence in Hungary is quite probably”];
Gruev & Döberl, 1997: 157-158; Geiser, 2001: 465; Brelih
et al., 2003: 96; Warchalowski, 2003: 419; Geiser, 2004:
19; Biondi, 2005: data on CD-Rom; Gruev & Döberl, 2005:
80; Čižek, 2006: 24-25; Warchalowski, 2010: 841 (Figs.
4096-4097), 842 (Fig. 4098); Özdikmen et al., 2020: 282.
Types
Lectotype (♂) and paralectotype (♀) of L. aphtho-
noides (Chabarofka, leg. Koltze, undated), selected and
studied by Warchalowski (1970: 106-107) were not re-
trieved (see below under “Notes on types search and iden-
tication”).
Two of the 5 paralectotypes (still labelled as syntypes)
of the Koltze collection, housed at SDEI, were seen as
a detailed photograph (kindly supplied by Lutz Behne)
(Fig. 2):
2 exx., labelled “Chabarofka // Weise”, undated. A sec-
ond handwritten label (by Weise, Koltze or else): “12223,
aphthonoides Ws, Chabarofka Weise determ. Type”.
Three specimens previously thought to be syntypes
of L. aphthonoides and now redened as paralectotypes
were found and examined:
1♀! labelled “Cotype” in MHNB: “Chabarowka /
aphthonoides Weise det. / Reitter don. // 1953 Coll. Heik-
ertinger // Paralectotypus / Longitarsus aphthonoides
Weise / vidit Laura Farina”.
1 ex. in HNHM labelled “Chabarowka / Longitarsus
aphthonoides Weise coll. Reitter // Paralectotypus / Lon-
gitarsus aphthonoides Weise / vidit Laura Farina”.
1 ex. in ZMB labelled “Chabarowka / coll. Koltze
// Typus // Longitarsus aphthonoides Weise det. L. N.
Medvedev, 1987 // Paralectotypus / Longitarsus aphtho-
noides Weise / vidit Laura Farina”.
The holotype of L. callidus, kept at BMNH was kindly
sent by Maxwell Barclay (Fig. 3: Holotypus; Fig. 4: ae-
deagus):
1 ♂! labelled “Talysch / Reitter // Longitarsus / cal-
lidus mihi / det. A. Warchalowski // Holotypus // BMNH /
A. Warchlowski collection”.
The study of this material (plus the many specimens
listed below) did not leave any doubt about the synonymy
proposed here:
Longitarsus aphthonoides Weise 1887 = Longitarsus
callidus Warchalowski 1967 syn. nov.
LAURA FARINA
13
Distribution
L. aphthonoides is widely distributed in the Palaearc-
tic region, west from Italy and Tunisia across Asia, east to
the Siberian Far East.
Gruev & Döberl (1997: 148, 157-158; 2005: 75, 80)
report the following regions:
Europe: Austria, Azerbaijan (Dashdatuk), Carpathian
Basin, Czechia (Moravia), France, Germany (West), Hun-
gary, Italy (North), Poland, Slovakia, Slovenia (Veržej-
Gomilica), Switzerland.
Asia: China (Heilungkiang, Fukien: Kuatun), Even-
kia, Iran (North), Kazakhstan (“Aulie Ata” = Dzhambul;
Syr Daria Valley; “Werneyi”= Alma Ata), Korea, Kurils,
Mongolia, Russia (Mid Siberia, Irkutsk District, East Si-
beria, Far East, Amurland, Maritime Province), Sakhalin,
Turkey.
North Africa: Tunisia.
Morphological remarks
L. aphthonoides belongs to the group of L. mem-
branaceus (Foudras 1860) (Leonardi, 1975: 213-214;
Doguet & Dufay, 1994: 306-307; Warchalowski, 1970:
106-107; 1995: 224); as for habitus, it is similar to L.
lycopi (Foudras 1860) (Warchalowski, 1970: 106-107;
Leonardi, 1975: 213-214).
After many years from the Weise paper dealing with
L. aphthonoides, and taking into account that the War-
chalowski description (1967: 59-60) of his L. callidus
might be considered rather short, it is useful to provide a
new description of this taxon.
Body length: ♂♂: 1.36-1.68 mm, ♀♀: 1.58-1.91 mm.
Shape elliptical, moderately narrow and on average
elongate (Figs. 1-3). Upper side (in mature individuals)
yellow to pale rusty-brownish; elytral suture lightly dark-
ened; head usually brown, darker than pronotum; upper
lip dark.
Antennae pale at least in part: in some specimens pro-
gressively darkened from 5°-6° antennomeres, at times the
last 4-5 antennomeres blackish. Legs yellow, hind femora
often dark but rarely completely pale (as yellow as body);
in some specimens, also last tarsomere and claws dark.
Frons [‘postfronssensu Heikertinger, 1913 (Fig. 5)]
and vertex [sensu Doguet, 2004 (Fig. 5a)] entirely mi-
crogranulated. Frontal tubercles distinctly separate from
frons by deep frontal furrows. Ocular furrows weak and
incomplete in terminal part as in L. lycopi.
Antennae always shorter than body. Articles ratio:
16:9:8:9:11:9:11:11:12:10:14.
Pronotum transverse [(Pw /Pl)m= 1.57-1.61], convex,
wider in middle; sides rounded, lateral calli (behind fore
corners) often showing an angular or obtuse dentiform
prole; surface clearly microgranulated and rather strong-
ly punctate; posterior angle, in intact specimens, with an
erect seta as long as last antennomere.
Elytrae bright, moderately elongated [(El/Ew)m =
1.56-1.57], slightly rounded on sides, jointly rounded api-
cally; surface less microgranulated than pronotum and
more strongly, irregularly punctate, sometimes with trac-
es of alignment. Humeral calli usually absent or weak but
rather developed on approximately 30% of the examined
specimens.
Pigidium microgranulated, covered by seta punctua-
tion, nely punctate in apical half and simply microgranu-
lated basally.
Male last sternite with a deep, oval pit (Warchalowski,
1967: 59 Fig. 3B; Leonardi, 1973: 212 Fig. 57).
First tarsomere of fore legs clearly widened in male
(Warchalowski, 1967: 59 Fig. 3A; Leonardi, 1973: 207
Fig. 43).
Median lobe of aedeagus (Figs. 4, 9-10), in lateral
view, almost straight in distal part but, sometimes, distal
2/3s weakly curved with apex shortly sinuate and apical
tip always facing upwards [a character which does not
appear in the Warchalowski drawings: both L. aphtho-
noides (Warchalowski, 1970: 114 Fig. 5) and L. callidus
(Warchalowski, 1967: 59 Fig. 3C), but appears in the
Medvedev drawing of L. aphthonoides (Medvedev, 1982:
114-115 Fig. 34: 2)]; in ventral-dorsal view its distal 1/3
tapered and rather expanded at apex with apical tip rather
short and prominent. Ventral groove moderately deep,
widened, its bottom with slight transverse wrinkles.
Basal part of spermatheca (Figs. 11-12) more or less
kidney-shaped, basal and distal part well differentiated
from each other, tip of distal part usually absent; ductus
spermathecae forming one, more or less widened, loop;
ductus insertion about in the middle of spermatheca basal
part.
Morphological data. (n=8): El = 1.05-1.26 mm;
(El)m = 1.17 mm; Ew = 0.70-0.79 mm; (Ew)m = 0.75
mm; Pl = 0.29-0.35 mm; (Pl)m = 0.31 mm; Pw = 0.47-
0.54 mm; (Pw)m = 0.50 mm; Lt = 0.44-0.54 mm; (Lt)m
= 0.50 mm; El/Ew = 1.48-1.61; (El/Ew)m = 1.56; El/Pl =
3.40-4.20; (El/Pl)m = 3.77; Pw/Pl = 1.40-1.70; (Pw/Pl)m
= 1.61; Lt/El = 0.41-0.46; (Lt/El)m = 0.43.
♀ (n=11): El = 1.21-1.37 mm; (El)m = 1.29 mm; Ew
= 0.77-0.91 mm; (Ew)m = 0.82 mm; Pl = 0.28-0.40 mm;
(Pl)m = 0.35 mm; Pw = 0.51-0.58 mm; (Pw)m = 0.55 mm;
Lt = 0.47-0.58 mm; (Lt)m = 0.52 mm; El/Ew = 1.48-1.66;
(El/Ew)m = 1.57; El/Pl = 3.38-4.03; (El/Pl)m = 3.69; Pw/
Pl = 1.40-1.86; (Pw/Pl)m = 1.57; Lt/El = 0.35-0.43; (Lt/
El)m = 0.40.
As mentioned above, the study of the L. callidus holo-
type has not shown any meaningful difference from the
paralectotypes of L. aphthonoides and other specimens
including topotypes. It’s worth noting that the author
could not corroborate the ♂♂ differences (between speci-
mens determined L. callidus and L. aphthonoides) in the
aedeagus shape, antennal length (which are shorter than
the body in the specimens studied), male width rst tar-
somere of fore tarsi (which is medium to strong) and the
last sternite character (with a deep, oval pit), previously
reported by Warchalowski (1967: 59, 60; 1970: 106-107,
114 Fig. 5) and Leonardi (1975: 213-214).
In conclusion, the synonymy between the two names
is here proposed.
It should be added that also Leonardi (1975: 213-214)
already reported the similarity between L. callidus War-
chalowski and L. aphthonoides Weise, as based on the
study of only a few ♀♀ specimens (including one syn-
type) of L. aphthonoides from the Frey collection (housed
at MHNB). The same author indeed noted the closeness
between the two in both external characters and the sper-
NOTES ON LONGITARSUS LATREILLE 1829
14
Figs. 1-8 – 1) L. aphthonoides. Wroclaw Wojnów. 2) L. aphthonoides. Chabarofka: paralectotypes. 3) L. callidus. Talysch Mountains:
holotype. 4) L. callidus, holotype: aedeagus. 5) L. personatus. Budapest. 6) L. lateripunctatus. Promontorio del Gargano. 7) L. monti-
cola. Monte Grigna Settentrionale. 8) L. curtus. Bisamberg. (Photos / Foto: 1, 3-8: S. Zoia; 2: L. Behne).
LAURA FARINA
15
matheca shape, not advancing the synonymy because of
the Warchalowski (1967: 59, 60; 1970: 106-107, 114 Fig.
5) claim of the several differences in the ♂♂ between
these two species.
Notes on typical locality
The exact placement of the L. aphthonoides type lo-
cality deserves a clarication.
Weise (1887: 206), in his original description, simply
reported “Chabarofka”. Warchalowski (1970: 106-107)
identied a specimen from this locality as lectotype and
correctly placed “Chabarowka” in the “Ussuri-Gebiet” (=
Ussuri region).
Today, the place can be identied with Khabarovsk
(also spelled Chabarovsk or Khabarovka; Chabarofka in
German), a town and an administrative territory of the
Russian Far East located at the conuence of Amur and
Ussuri rivers.
Notes on ecology
In Italy L. aphthonoides was observed in May-June
1973, close to Rovasenda (Vercelli, Piedmont), on Lysi-
machia vulgaris L. (Primulaceae), but probably this was
only a refuge plant (Leonardi, 1975: 213-214, reported
as L. callidus). The host plant could be Stachys L. spp. or
Teucrium L. spp. (Lamiaceae), similarly to other species
related to L. membranaceus.
L. aphthonoides was also observed on Sanguisorba of-
cinalis L. (Rosaceae) (Warchalowski, 1996: 76-77) and
on Onobrychis viciifolia Scopoli (= Onobrychis sativa
Lamarck) (Fabaceae) (Özdikmen et al., 2020: 282).
In the Rovasenda area, Lysimachia vulgaris as well as
the (supposed) host plants (Stachys spp., Teucrium spp.,
Sanguisorba ofcinalis) are rather common and wide-
spread but, unfortunately, this species could not be ob-
served any more in the several surveys carried out by the
author in the years 2018, 2019 and 2020.
Notes on types search and identification
Warchalowski (1970: 106-107) established, as lec-
totype (♂) and paralectotype (♀) of L. aphthonoides,
two specimens of “DEI” collection in Eberswalde (now
SDEI in Müncheberg), from Chabarofka, leg. Koltze,
undated.
Unfortunately, it was impossible to examine these
specimens. Indeed, they are missing from the type series
still present in SDEI, they are missing in the collection
Warchalowski housed at the BMNH and they were not
found also in the Reitter collection (housed both at MH-
NB and HNHM).
At ZMB, two specimens were found, bearing a red la-
bel “Typus”, written by a curator (Bernd Jaeger in verbis);
one of these specimens (♀) not dissected and labelled
“Chabarowka / Koltze // det. Medvedev 1987”, is prob-
ably a syntype of Weise, now a paralectotype. The other
specimen, a male, not dissected, is labelled Irkutsk and
collected by Koltze (but it cannot be considered among
the type material).
Some topotypical specimens were also found in the
MSNM-Ber, permitting to illustrate the median lobe of
the aedeagus of one of them (Fig. 9).
Materials studied
As mentioned, many specimens were studied, from
several Western and Eastern Palaearctic localities (as de-
tailed below), which were determined with either one or
the other name but all certainly belonging to this same
species.
Italy: Piemonte: 2 exx. (1♂! 1♀!) Rovasenda /
13.V.1973 / leg. Leonardi // Longitarsus callidus det. C.
Leonardi (LFCC); 3 exx. Rovasenda / 17.VI.1973 / leg.
Leonardi // Longitarsus callidus det. C. Leonardi (LFCC);
1 ex. (1♂!) Rovasenda / 17.VI.1973 / leg. Leonardi // Lon-
gitarsus callidus Warchalowski det. C. Leonardi, 1975
(BMNH); 2 exx. (1♂! 1♀!) Rovasenda / Novara VI.1959
/ brughiera / Leonardi (MSNM); 1 ex. Piemonte (VC)
/ Rovasenda / 13.V.1973 // Longitarsus callidus det. C.
Leonardi (MSNM); 66 exx. (12♂♂! 6 ♀♀!) Rovasenda /
17.VI.1973 / leg. Leonardi // Longitarsus callidus det. C.
Leonardi (MSNM); 1 ex. (1♀!) Rovasenda / 18.VI.1976
/ leg. Leonardi // Longitarsus callidus det. C. Leonardi
(MSNM); 1 ex. (1♀!) Rovasenda / 5.VI.1977 / leg. Pe-
sarini // Longitarsus callidus det. C. Leonardi (MSNM); 2
exx. (1♀!) Rovasenda / 5.VI.1977 / leg. Fogato // Longi-
tarsus callidus det. Leonardi (MSNM); 1 ex. (1♂!) Rov-
asenda / 17.VI.1973 / leg. C. Leonardi // Longitarsus cal-
lidus Warchalowski det. Leonardi, 1975 (ZSM);
Friuli Venezia Giulia: 1 ex. (1♂!) Doberdò / 15.X.1933
/ Springer (MSNM); 1 ex. Friuli V.G. (Trieste) / dint. Gro-
pada / 15.IV.1996 / leg. S. Zoia // Longitarsus callidus
det. L. Farina (SZCM);
Poland: 1 ex. Poland / Wroclaw Wojnów / 17°06’E-
51°05’N / 18.VIII.1995 / leg. M. Döberl // Longitarsus
callidus det. M. Döberl, 1995a (BMNH); 1 ex. (1♂!)
Polonia sept. / Puszcza Piska Szeroki Bór / 17.XI.1958
/ leg. A. Szujecki // Longitarsus callidus mihi / det.
A. Warchalowski (BMNH-War); 1 ex. Silesia minor /
Wojnów ad Wroclaw / 130 m 15.X.1992 / leg. A. War-
chalowski // Longitarsus callidus mihi / det. A. War-
chalowski (BMNH-War); 1 ex. Silesia minor / Wojnów
ad Wroclaw /13.IX.2008 / leg. A. Warchalowski // Lon-
gitarsus callidus mihi / det. A. Warchalowski (BMNH-
War); 1 ex. (1♂!) Poland, Silesia / Wroclaw-Wojnów /
24.X.1975 / L. Borowiec // Longitarsus callidus det. L.
Borowiec (MSNM); 1 ex. Poland, Silesia / Wroclaw-
Wojnów / 1.V.1977 / L. Borowiec // Longitarsus callidus
det. L. Borowiec (MSNM); 2 exx. (1♂! 1♀!) Poland,
Silesia / Wroclaw-Wojnów / 24.IX.1982 / L. Borowiec //
Longitarsus callidus det. L. Borowiec (MSNM); 2 exx.
(1♂!) Poland, Silesia / Wroclaw-Wojnów / 3.IX.1994 /
L. Borowiec // Longitarsus callidus det. L. Borowiec
(MSNM); 16 exx. (1♂!) Wroclaw Wojnów / 5.X.1982 /
Warchalowski (MSNM-Ber); 6 exx. (3♂♂! 3♀♀!) Polen
Silesia minor / Umg. Wroclaw Oderwiesen / 24.VII.1994
/ leg. U. Heinig // an Wiesenkopf (Sanguisorba sp.) //
Longitarsus callidus det. M. Döberl, 1995a (MSNM-
Ber) (Fig. 10); 1 ex. (1♂!) Poland / Wroclaw Wojnów
/ 17°06’E-51°05’N / 18.VIII.1995 / leg. M. Döberl //
Longitarsus callidus det. M. Döberl, 1995a (MSNM-
Ber) (Fig. 1);
Austria: 1 ex. (1♂!) Nussdorf / (S. Peter) 10.V.1936 /
Springer // Longitarsus callidus Warch. det. C. Leonardi
(MSNM); 1 ex. (1♂!) Moor von Gois / bei Salzburg / H.
NOTES ON LONGITARSUS LATREILLE 1829
16
Frieb // Longitarsus callidus det. Bergeal (MSNM-Ber);
1 ex. (1♂!) Achau NÖ / 28.VIII.1960 / Hausmauer // 1953
coll. Heikertinger // Longitarsus callidus det. C. Leonardi
(ZSM);
Slovenia: 1 ex. Slovenia / Verzej-Gomilica / 16°12’
E- 46°34’N / 19.IV.1996 Döberl // Longitarsus callidus
det. M. Döberl (MHNB); 1 ex. Valvolciana / Stara Gora
17.V.1943 / Springer (MSNM); 3 exx. (1♂! 2♀♀!) Slov-
enia Verzej-Gomilica / 16°12’E-46°34’N / 19.IV.1996
Döberl // Longitarsus callidus det. M. Döberl (MSNM-
Ber) (Fig. 12);
Russia: 1 ex. (1♂!) Derbent // cetera data nulla //
Longitarsus callidus mihi / det. A. Warchalowski (BM-
NH-War); 1 ex. Ussuria / lacus Chanka / 12.VI.1908 /
leg. Aleks. Czerski // coll. Warchalowski // Longitarsus /
aphthonoides Weise det. A. Warchalowski (BMNH-War);
1 ex. Ins. Sachalin / Konuma / 30.V.1927 / leg. Tamanuki
// coll. Warchalowski // Longitarsus aphthonoides Weise
det. A. Warchalowski (BMNH-War); 1 ex. Chabarowka /
Longitarsus aphthonoides Weise coll. Reitter // Paralec-
totypus / Longitarsus aphthonoides Weise / vidit Laura
Farina (HNHM); 1 ex. (1♀!) Chabarowka / aphthonoides
Weise det. Type / Reitter don. // Cotype // Paralectoty-
pus / Longitarsus aphthonoides Weise / vidit Laura Fa-
rina (MHNB-Heik); 1 ex. (1♀!) Baical / Obert. Transt. //
Longitarsus aphthonoides Weise (MHNB-Heik); 2 exx.
(2♂♂!) Saghalien // Longitarsus aphthonoides det. B.
Gruev (MSNM); 1 ex. (1♂!) Vladivostok / 8.VII.1927
/ B. Belov // Longitarsus aphthonoides det. B. Gruev
(MSNM-Ber); 6 exx. (2 ♂♂! 4♀♀!) Amur country /
Chabarovsk / Primorskaja / 7.VIII.1990 // leg. D. Boukal
(MSNM-Ber) (Figs. 9, 11); 2 exx. Chabarofka // Weise
// 12223 // Longitarsus aphthonoides Ws. det. J. Weise
Type (SDEI) (Fig. 2); 1 ex. Chabarowka / coll. Koltze
// Typus // Longitarsus aphthonoides Weise det. L. N.
Medvedev, 1987 // Paralectotypus / Longitarsus aphtho-
noides Weise / vidit Laura Farina (ZMB); 1 ex. Irkutsk /
coll. Koltze // Typus (ZMB); 1 ex. (1♂!) Krasnoyarsky
region / Lensk transt. / 5, 6 Station / meadow near the
forest / 15/1 / 18.VI.1975 / Guselnikov leg. (ZSM); 1 ex.
(1♂!) USSR / Evenkia / Tura / 28.VIII.1982 // leg. L.N.
Medvedev // Longitarsus aphthonoides Weise / det. B.
Gruev (ZSM);
Asia: 1 ex. (1♂!) Talysch / Reitter // Longitarsus cal-
lidus mihi / det. A. Warchalowski // Holotypus (BMNH-
War) (Fig. 3); 1 ex. (1♀!) Aulie Ata / Syr Daria (MSNM);
1 ex. N. China / P.M. Hammond / B.M. 1967-215 //
Heilungkiang / Erlungshan / 29.V.1966 / P.M. Hammond
// Longitarsus aphthonoides Weise det. B. Gruev (BM-
NH); 1 ex. (1♀!) N. China / P.M. Hammond / B.M. 1967-
215 // Heilungkiang / Erlungshan / 29.V.1966 / P.M. Ham-
mond // Longitarsus aphthonoides det. B. Gruev (ZSM);
1 ex. (1♀!) N. China / P.M. Hammond / B.M. 1967-215 //
Longitarsus aphthonoides Weise det. B. Gruev (ZSM); 1
ex. (1♀!) Korea prov. / Hamgjŏng-namdo / leg. M. Mroc-
zkowski et A. Riedel // ad lac. Čang-džin-ho / 9.VI.1965
// coll. Warchalowski // Longitarsus aphthonoides Weise
det. A. Warchalowski (BMNH-War); 1 ex. Korea Mt.
Pektusan / 2-6 km N Samzi-yan hotel wood / 18.VII.1977
// No. 372 netting in grasses / Dely & Draskovits // Lon-
gitarsus aphthonoides Weise det. B. Gruev (HNHM); 1
ex. Korea / Ryanggang Prov. / Samjlyon 1000 m / No.
976 / 3.VI.1985 / A. Vojnits & L. Zombori // Longitarsus
aphthonoides Weise det. B. Gruev (HNHM); 1 ex. Ko-
rea Mt. Pektusan / environs Sam-Zi-Yan / Hotel Lake-
Shore 19.VII.1977 // No. 376 Netting in grasses / Dely
& Draskovits // Longitarsus aphthonoides Weise det. B.
Gruev (ZSM);
North Africa: 1 ex. (1♂!) Tunis // 1953 Coll. Heik-
ertinger // Longitarsus callidus det. C. Leonardi, 1975
(ZSM).
Dissected specimens: 40 ♂♂; 27 ♀♀.
Figs. 9-12 – L. aphthonoides. 9-10) Aedeagus: median lobe / lobo mediano; a) ventral view / vista ventrale; b) lateral view / vista later-
rale. 11-12) Spermatheca. Localities / Località: 9) Chabarovsk; 10) Wroclaw; 11) Chabarovsk; 12) Verzej-Gomilica. Here and in the
ensuing groups of gures, the relative elytral lengths are shown to the top right of each drawing. / Qui e nei seguenti gruppi di gure,
le lunghezze elitrali relative sono indicate accanto a ciascun disegno in alto a destra.
LAURA FARINA
17
Longitarsus personatus Weise 1893 stat. nov. (Fig. 5)
Longitarsus lateripunctatus var. personatus Weise,
1893: 964.
Kolbe, 1920: 402-403, 404 [mut. personata]; Heik-
ertinger, 1930: 1328; Porta, 1934: 357; Normand, 1937:
133; Csiki & Heikertinger, 1940: pars 166, 134; Mohr,
1962: 62 (Figs. 32a, 32b), 63 (Fig. 32c), 98; Mohr, 1966:
228; Warchalowski, 1978: 49, 50 (Figs. 243-245); Furth,
1980: 101; Gruev & Tomov, 1986: 221-222 (Fig. 244);
Gruev, 1987: 58 (Fig. 21) [the author, in the text, wrote
about lateripunctatus but in Fig. 21 represented a sper-
matheca of personatus]; Warchalowski, 2003: 413 (forma
personatus).
= Longitarsus lateripunctatus ssp. personatus Weise
sensu Doguet, 1989: 172. Loc. typ.: Austria: Wien (as in
Gruev & Döberl, 1997:182).
Heikertinger, 1912: 194 [“Unterart personatus. In
Deutschland nicht gef. Die Normalform lateripunctatus
(biguttatus Foudras) nur in Mittelmeergebiete”]; Müller,
1949-1953: 524, 536; Kaszab, 1962: 311 (Fig. 132 I-J),
314; Gruev, 1979: 131; Biondi, 1990a: 142-143; Biondi,
1990c: 346; Gruev et al., 1993: 102; Döberl, 1994: 107,
135 (Figs. 266-268); Doguet, 1994: 220-221, 260 (Fig.
105bis b), 297 (Fig. 122g); Lucht, 1994: 319; Döberl,
1995b: 55-56; Warchalowski, 1995a: 144 (Fig. 384), 146
(Figs. 400-401), 254-256 (Figs. 954, 956); Warchalowski,
1996: 36 (Fig. 19, L. lateripunctatus), 38 (Figs. 35, 36, L.
lateripunctatus), 137-138, 141 (Fig. 589), 142 (Fig. 591);
Vig, 1996: 125; Rozner, 1996: 256; Strejcek, 1996: 572;
Gruev & Tomov, 1998: 93; Gruev & Döberl, 1997:182;
Herger & Kamke, 1998: 123 [“neu für Kanton Schaff-
hausen und NordSchweiz”]; Callot, 1999: 43; Gruev, 2002:
27; Vig, 2002: 122; Brelih et al., 2003: 114; Geiser, 2004:
19; Gruev & Döberl, 2005: 90; Čižek, 2006: 32 (Fig. 102)-
33; Warchalowski, 2010: 864 (Figs. 4289-4290), 865 (Fig.
4292); Baviera & Biondi, 2015: 19; Fritzlar, 2017: 80.
= Longitarsus lateripunctatus (Rosenhauer 1856)
Furth, 1980: 101 [partim?]
The comparative study of a wide sample (MSNM,
MSNG, MHNB, HNHM and private collections) permit-
ted to come to the conclusion that L. personatus is neither
just a wingless form of L. lateripunctatus, as Weise sup-
posed (1893: 964) and as Warchalowski (1978: 49-50) and
Gruev & Tomov (1986: 221-222) conrmed, nor a subspe-
cies, as put forward by Heikertinger (1912: 194), Müller
(1949-1953: 524, 536), Kaszab (1962: 311, 314), Doguet
(1989: 172) and Warchalowski (1996: 137), but a valid
species. This taxonomic proposal is based on the signi-
cant morphological differences between the two species
and considers the partial overlap of their ranges (Fig. 31).
Comparative description
An accurate morphological description of both spe-
cies can be found in Doguet (1989: 172; 1994: 220-221,
260 Figs. 105 bis a, b) and Warchalowski (1995: 254-256;
1996: 137-138, 142 Figs. 590-591), which also report col-
our differences, regrettably not always a stable character.
In addition, Doguet (1989: 172; 1994: 220-221, 297 Figs.
122 f-g) observed differences in the shape of the sper-
matheca (ductus spermathecae forms 2 or 3 loops in L.
lateripunctatus and shows no loops in L. personatus).
The comparative analysis of the morphology on a wider
sample permitted to add further discriminant characters
between the two species (Table 1).
Distribution
L. personatus distribution is mainly eastern Euro-
Mediterranean.
Gruev & Döberl (1997: 182; 2005: 90) report the fol-
lowing regions:
Europe: Austria, Bosnia-Herzegovina, Bulgaria, Car-
pathian Basin, Croatia, Czechia, France (Alsace), Germa-
ny, Greece (Crete), Hungary, Italy (North and East), Mol-
davia, Montenegro, Rumania, Serbia, Slovakia, Slovenia,
Switzerland, Ukraine.
Asia: Caucasus, Cyprus, Greece (Rhodos), Israel,
Caucasic Russia (Krasnodar District), Turkey.
In previous studies (Müller, 1949-1953: 524, 536;
Doguet, 1989: 172; 1994: 220-221; Warchalowski,
1995a: 254-256; 1996: 137-138, 141; 2010: 865) where
L. lateripunctatus and L. personatus were considered as
geographical races of the same species, L. lateripunctatus
was reported to include the western Mediterranean area,
while L. personatus range would cover the northern and
eastern part of it. However, as it appears from the materi-
als studied (see below), the two species are clearly para-
patric (Fig. 31), with overlapping ranges, which mainly
include peninsular Italy and Sicily.
It is worth reporting that, at least in two instances, the
two species were found together in the same locality:
San Pier Niceto, Messina, Sicilia:
L. lateripunctatus: 1 ex. San Pier Niceto (Leonardi in
verbis)
L. personatus: 1 ex. (1♀!) (MSNM) labelled “Sicilia
Messina Peloritani / San Pier Niceto 2, 350 m / 19.VI.2011
/ leg. C. Baviera”
Lazio, Riano:
L. lateripunctatus: 1 ex. (1♂!) (MBCA) labelled
“Lazio Roma / Riano Flaminio / 1.IV.1979 M. Biondi”
L. personatus: 1 ex. (1♀!) (MSNM) labelled “Lazio
Riano / VII.1937 Garavaglia”
In other countries too, specimens of the two species
were occasionally found in the same locality. For exam-
ple, in Greece (Corfu):
L. lateripunctatus: 1 ex. (1♀!) (MHNB-Heik) labelled
“Corfù / Valle di Ropa”, undated; 2 exx. (1♀!; 1♂!) (HN-
HM, the ♀ coll. Dr. J. Fodor) labelled “Corfù / Paganetti”,
undated;
L. personatus: 2 exx. (1♀!; 1♂!) (MHNB-Heik) la-
belled “Corfù / Paganetti”, undated; 1 ex. (1♂!) (HNHM)
labelled “Corfù / Paganetti”, undated.
Materials studied
France: 1 ex. Gallia / Reitter (MSNM); 2 exx. Bois
de Sermersheim / 9.IV.1999 M. Callot / Pulmonaria
(MSNM-Ber);
Italy: Friuli Venezia Giulia: 1 ex. (1♂!) Trieste-Carso
/ Divaccia / 14.VI.2008 (MSNM) (Fig. 28); 1 ex. (1♀!)
Trieste-Carso / Divaccia / 14.VI.2008 (MSNM);
NOTES ON LONGITARSUS LATREILLE 1829
18
Liguria: 1 ex. Fontanegli / Bavari Genova / A. Bal-
iani (MBCA); 1 ex. S. Ilario Ligure / 28.III.1915 / G.
Mantero // Longitarsus lateripunctatus det. A. Dodero
(MSNG); 2 exx. (1♂! 1♀!) Fontanegli / Bavari Geno-
va / IX.1925 / A. Baliani // Longitarsus lateripuncta-
tus Rosenhauer teste M. Biondi (MSNG); 1 ex. (1♀!)
Genova / S. Ilario Ligure / 4.VII.1944 / leg. Binaghi
(MSNG) (Fig. 22); 1 ex. Liguria (GE) Genova / febbraio
1884 / A. Dodero (MSNG-Bin); 7 exx. (1♂!) Genova
/ S. Mario 4.VII.1944 // su Commelina communis G.
Binaghi (MSNG-Bin); 2 exx. (1♀! 1♂!) Liguria (GE)
Genova Sturla 1.IV.1945 / Binaghi // su Symphytum bul-
bosum (MSNG-Bin); 3 exx. (1♂!) Liguria (GE) Genova
/ Febbraio 1884 / A. Dodero (MSNG-Dod); 1 ex. Liguria
(GE) Genova / Aprile 1893 / A. Dodero (MSNG-Dod);
2 exx. (2♂♂!) Liguria (SP) Ameglia / Liguria orientale
/ 14.IX.1913 / C. Mancini (MSNG-Man); 9 exx. Ligu-
ria (GE) / Lavagna / 10.XII.1972 / S. Zoia (SZCM); 1
ex. Finale Ligure (SV) loc. Perti / greto Torrente Pora /
10.IV.1994 / S. Zoia (SZCM);
Toscana: 1 ex. (1♂!) Palude Peretola / Firenze
27.XI.1936 / A. Martelli (MSNM) (Fig. 27);
Tab. 1 - Taxonomically discriminant characters of L. lateripunctatus and L. personatus. / Caratteri tassonomici
discriminanti di L. lateripunctatus e L. personatus.
Character L. lateripunctatus (Rosenhauer 1856) L. personatus Weise 1893
Elytrae Sutural stripe of elytrae almost parallel and
usually narrow, in some specimens wider in
middle (Fig. 6; also: Doguet, 1989: 172 Fig.
1; Warchalowski, 1996: 142 Fig. 590; Doguet,
1994: 260 Fig. 105bis a), never joining lateral
black spots, except in one specimen from
Morocco (prov. Taza in MBCA) which is
considered belonging to L. lateripunctatus for
the shape of the aedeagus.
Sutural stripe of elytrae widened before
middle, sometimes joining lateral black
spots (Fig. 5; also: Doguet, 1989: 172 Fig. 2;
Warchalowski, 1996: 142 Fig. 591; Doguet,
1994: 260 Fig. 105bis b).
Uncommonly pale specimens can be found,
with sutural stripe narrower in material
studied from Cyprus: 4 exx. (4♀♀!) “Larnaka,
Tochni”, “Lemessos, Pachna”, “Lemessos,
Gerovasa”, “Larnaka, Lefkara” (FFCJ) and 1
ex. (1♂!) “Akamas, Drouseia” (MSNM-Ber).
Spermatheca Ductus spermathecae forms 2 or 3 loops
(Figs. 13-17; also: Doguet, 1989: 172 Fig.
3; 1994: 297 Fig. 122f). Distal part of
spermatheca usually wide and rounded at
apex.
Ductus spermathecae more or less long and
without loops (Figs. 18-22; also: Doguet,
1989: 172 Fig. 4; 1994: 297 Fig. 122g). In
some specimens, ductus forms a slight ripple
(Figs. 18 and 21). Distal part of spermatheca
usually narrow and apically folded.
Median lobe
of aedeagus
Median lobe of aedeagus (Figs. 23-26) on
average less elongate than in L. personatus:
(n=12) Al = 0.652-0.732 mm; (Al)m = 0.696
mm; Aw = 0.112-0.128 mm; (Aw)m = 0.122
mm; (Al/Aw) = 4.993-6.536; (Al/Aw)m =
5.732.
In ventral view comparatively straight or basal
2/3s weakly tapered, not broadened at apex
(only in a few specimens, weakly broadened);
distal 2/3s usually bent in lateral view.
Specimens from North Africa show a longer
aedeagus in ventral view (Fig. 24a) basal
2/3s tapered and weakly wider at apex, but in
lateral view bent as usual (Fig. 24b).
Median lobe of aedeagus (Figs. 27-30) more
elongate than in L. lateripunctatus:
(n=12) Al = 0.699-0.792 mm; (Al)m = 0.747
mm; Aw = 0.096-0.128 mm; (Aw)m = 0.115
mm; (Al/Aw) = 5.974-7.375; (Al/Aw)m =
6.539.
In ventral view basal 2/3s weakly tapered
and more or less broadened at apex. In
lateral view distal 2/3s appear comparatively
straight.
Wings Usually hind wings well-developed, with
humeral calli evident (75% of examined
exx.). Almost all specimens with hind
wings reduced come from islands (where
winged specimens are also present): Corfù
(1 ex.), Isola del Giglio (3 exx.), Sardegna
(3 exx.), Isola di Capraia (1 ex.), Sicilia (4
exx.). Among the specimens studied, two,
with reduced wings, come from continental
localities (Portugal: “São Martinho” and
“Serra de São Mamede”).
Usually hind wings reduced, with humeral
calli absent (73% of examined exx.).
LAURA FARINA
19
Marche: 1 ex. Macerata / 1960 / leg. S. Battoni (MSN-
VR);
Lazio: 2 exx. (1♂!) Lazio Roma / dintorni Maccarese
/ 25.II.1979 M. Biondi (MBCA); 1 ex. Lazio / Fregene /
24.IV.1978 / leg. M. Biondi (MBCA); 1 ex. (1♀!) Lazio
Riano / VII.1937 Garavaglia (MSNM) (Fig. 21);
Campania: 1 ex. Napoli / Italia // lateripunctatus det.
F. Heikertinger (MHNB-Heik);
Puglia: 1 ex. (1♀!) Puglia (TA) / Fiume Lato /
12.III.2011 / leg. F. Montemurro (LFCC) (Fig. 20);
Calabria: 4 exx. (2♀♀! 1♂!) Calabria (RC) / Sta
Eufemia / Paganetti // ex Orig. Samig. J. Breit Wien
(MHNB); 1 ex. (1♂!) Calabria (CZ) Sambiase / Calabria
/ V.1920 / C. Minozzi (MSNG-Man);
Sicilia: 1 ex. (1♀!) Sicilia Messina Peloritani / San
Pier Niceto 2, 350 m / 19.VI.2011 / leg. C. Baviera // N
38°07’35.6 E 15°23’12.4 // Longitarus lateripunctatus
personatus W. det. C. Leonardi (MSNM) (Fig. 19); 1 ex.
(1♂!) Palermo / Oreto / VI.1961 // Longitarsus lateripunc-
tatus (Rosh.) det. A. Warchalowski (MSNVR);
Czechia: 1 ex. Moravia meridionale / Palava ras 169 /
J. Macek 1992 (MSNM-Ber);
Austria: 3 exx. (3♀♀!) Hainburg / A. i. Mader // Longi-
tarsus lateripunctatus Rosh. det. A. Warchalowski, 1960-
62 (HNHM); 2 exx. (2♂♂!) Austria / Krems a. Don. / I.
Bachinger (MHNB); 1 ex. Bisamberg // ex Orig. Samig.
J. Breit Wien (MHNB); 1 ex. Mödling / Austr. Breit // ex
Orig. Samig. J. Breit Wien (MHNB); 1 ex. (1♀!) Baden-
Wien / Helenental // Helen. Bad // personatus mit Schul-
terteule (MHNB-Heik); 1 ex. (1♂!) Wien-Bisamberg / H.
Schweiger (MHNB-Heik); 1 ex. Bisamberg / 2.II.1907 //
Bisamberg A.i. / Heikertinger // Longitarsus lateripuncta-
tus (MHNB-Heik); 1 ex. (1♀!) Bisamberg / Heikertinger
/ 13.VI.1911/ Scheuch // lateripunctatus det. F. Heik-
ertinger (MHNB-Heik); 1 ex. Austria / Krems a. Don. /
I. Bachinger // Longitarsus lateripunctatus det. F. Heik-
ertinger // coll. A. Simonis dupleta coll. Winkler (MSNG);
5 exx. Vienne / Autriche (MSNG-Dod); 4 exx. (3♂♂!)
Austria / Krems a. Don. / J. Bachinger (MSNM) (Figs.
29, 30); 1 ex. Bisamberg / A. Otto (MSNM); 1 ex. (1♀!)
Austria inferiore / Bisamberg / Th. v. Wanka (MSNM); 2
exx. Wien (MSNM); 4 exx. (1♂!) Laxenburg / A. Otto //
ex. Museo A. Carret 1908 / Longitarsus lateripunctatus
personatus det. S. Doguet, 1984 (MSNM-Ber);
Slovenia: 1 ex. Pirano Is. (MHNB-Heik); 1 ex. (1♀!)
SLO Mota / Hrastje / 25.IV.2002 / leg. B. Drovenik // Lon-
gitarsus lateripunctatus personatus det. M. Döberl, 2003
(MSNM-Ber); 1 ex. SLO / Bunčani / 25.IV.2002 / leg. B.
Drovenik // Longitarsus lateripunctatus personatus det.
M. Döberl, 2003 (MSNM-Ber);
Hungary: 2 exx. Siόfok / Lichtneckert // Longitarsus
lateripunctatus personatus Weise det. Z. Kaszab, 1962
(HNHM); 1 ex. BPST Ungbg. / Budai hegys // coll. Dr.
J. Fodor // Longitarsus lateripunctatus personatus Weise
det. B. Gruev (HNHM); 2 exx. Bpst környéke / Budai he-
gys / coll. H. Diener // Longitarsus lateripunctatus per-
sonatus Weise det. Z. Kaszab, 1962 (HNHM) (Fig. 5); 2
exx. (2♂♂!) Hung. 26.IV.1931 / Budakeszi erdö / coll. H.
Diener // Longitarsus lateripunctatus personatus Weise
det. Z. Kaszab, 1962 (HNHM); 1 ex. Hu. Pest m. Monor
/ 7.VI.1936 / leg. Dr. J. Fodor // Longitarsus lateripunc-
tatus personatus Weise B. Gruev (HNHM); 1 ex. (1♂!)
Figs. 13-22 – Spermatheca. 13-17) L. lateripunctatus. 18-22) L. personatus. Localities / Località: 13) Laigueglia; 14) Talamone;
15) Circummarpiccolo; 16) Monti Peloritani; 17) Corfù, Valle di Ropa; 18) Corfù; 19) Messina, San Pier Niceto; 20) Fiume Lato;
21) Riano; 22) Sant’Ilario Ligure.
NOTES ON LONGITARSUS LATREILLE 1829
20
Dp. Kara-hegy / 16.V.1950 / Révy D. / coll. Dr. Révy
D. // Longitarsus lateripunctatus personatus Weise det.
Z. Kaszab, 1962 (HNHM); 1 ex. Mecsek, Pécs / Misina
É. / 26.V.1954 leg. Kaszab // Longitarsus lateripuncta-
tus personatus Weise det. Z. Kaszab, 1962 (HNHM); 1
ex. Hu. Vértes h. / Fány Völgy // 2.VIII.1961 / Hámoriné
// Longitarsus lateripunctatus personatus Weise det. B.
Gruev (HNHM); 1 ex. Hung. Békés m. Sarkad / Remetei-
erdő 90 m // Festuco pseudovinae / Quercetum // egyelés
/ 24.V.1982 / leg. Adám // Longitarsus lateripunctatus
Rosenhauer ssp. personatus Weise (HNHM); 1 ex. (1♀!)
Hung. Pest m. / Tahitόfaln / Cseresznyés-völgy // fuhálόzás
/ 27.V.1984 / leg. O. Merkl // Quercetum petrae-cerris /
300 m // Longitarsus lateripunctatus personatus Weise
det. B. Gruev (HNHM); 1 ex. (1♂!) Hung. Szabolcs – Sz.
M. / Bátorliget / Bátorligeti-láp // 29.IX.1990 / leg. Papp.
J. // Longitarsus lateripunctatus personatus Weise det. O.
Merkl 1991 (HNHM); 3 exx. (1♂!) Fünfkirchen [=Pécs]
/ Hungar. // lateripunctatus det. F. Heikertinger (MHNB-
Heik); 2 exx. Fünfkirchen / Hungar. // Longitarsus / v.
personatus // v. personatus / ?Shinam // cotypus (MHNB-
Heik); 1 ex. (1♀!) Ungarn (MSNM);
Croatia: 1 ex. Dalmatia c. / Ins. Brazza / Milna V.1936
/ Stöcklein (MHNB);
Bosnia-Herzegovina: 1 ex. (1♀) Apfelbeck / Sara-
jevo // Longitarsus lateripunctatus coll. Apfelbeck // L.
lateripunctatus Rosh. det. A. Warchalowski, 1962 (HN-
HM);
Serbia: 1 ex. (1♀!) Slavonia Ruma / Schwieger //
Longitarsus lateripunctatus Rosenhauer det. M. Biondi,
1985 (MSNG);
Rumania: 1 ex. Herkulsfürdő / coll. Mihόk // Longi-
tarsus lateripunctatus personatus Weise det. Z. Kaszab,
1962 (HNHM); 1 ex. Herkulsfürdő / 1880 Pável // Lon-
gitarsus lateripunctatus personatus Weise det. Z. Kaszab,
1962 (HNHM); 1 ex. Herkulesbad / Sd Ung. Breit // ex
Orig. Samig. J. Breit Wien (MHNB);
Greece: 1 ex. (1♂!) Greece / Peloponnes SE / Ka-
lamata oberhalb // Kastania N36°52’19’’ E22°18’50’’ 850
m // 1.X.2014 leg. C. Germann // Longitarsus personatus
Weise det. L. Farina, 2019 (CGCB); 3 exx. Kreta Rethim-
non d. 08 / Eleftherna Archea El. / 300 m NN / 14.4.2019
Fritzlar leg. // Longitarsus lateripunctatus det. F. Fritzlar,
2019 (FFCJ); 1 ex. Morea / Hagios Wlastis Breuske //
Longitarsus lateripunctatus Rosenhauer coll. Reitter
(HNHM); 1 ex. (1♂!) Corfù / Paganetti // lateripunctatus
det. K. Lopatin (HNHM); 1 ex. Peloponneso / Tripoli din-
torni Dahra / 4.VIII.1980 / leg. P. Audisio e M.C. Aneelié
(MBCA); 1 ex. (1♂!) Isola di Kos / Asklipiion ume /
23.III.1989 P. Audisio (MBCA); 1 ex. Karp. M. / Motrul
Sec. (MHNB); 1 ex. Volo / Thessalien // Moctarski // lat-
eripunctatus (MHNB-Heik); 2 exx. (1♀!) Corfù / Pagan-
etti // lateripunctatus det. F. Heikertinger (MHNB-Heik)
(Fig. 18); 1 ex. (1♀!) Vardarebene / Macedonia / A. Shatz-
mayr (MSNG-Dod); 1 ex. (1♀!) Rodi / 21.II.1931 / Torre
Tasso (MSNM); 1 ex. (1♂!) Kriti-Hanjà (44) / Aghio-
pandes / Vryses 6 km NW / 28.V.2006 Liberti (MSNM);
1 ex. Grèce Laconie Aeropoli / Neo Itilio 15.IV.1997 /
J-M Maldes & A. Madocq leg. (MSNM-Ber); 1 ex. (1♀!)
Crète (Rethimnon) / Agios Silas / 1.V.1999 / M. Bergeal
leg. (MSNM-Ber); 1 ex. (1♀!) GR. Peloponnese Arka-
dia / Eleniko 9.IV.2000 / Bords de Kouraikos / B. et M.
Bergeal leg. (MSNM-Ber); 1 ex. GR. Peloponnese Mavi
/ Kokkala 12.IV.2000 / B. et M. Bergeal leg. (MSNM-
Ber); 1 ex. (1♀!) GR. Peloponnese Ahaia / Mega Spileo
15.IV.2000 / Bords de Kouraikos / B. & M. Bergeal leg.
(MSNM-Ber); 1 ex. GR. env. Xanthi / Bordes de rivière /
22.V.2001 / B. et M. Bergeal leg. (MSNM-Ber);
Cyprus: 1 ex. (1♀!) Cyprus W 38 / Lemessos 35 km
WNW / Gerovasa S 500 m / 10.IV.2006 Fritzlar leg. //
Longitarsus lateripunctatus det. F. Fritzlar, 2007 (FFCJ);
3 exx. (1♀!) Cyprus S 63 / Lemessos 20 km NW / Pach-
na E 800 m / 3. IV.2009 Fritzlar leg. // Longitarsus lat-
eripunctatus det. F. Fritzlar, 2009 (FFCJ); 2 exx. (1♂!)
Cyprus S 67 / Lemessos 30 km NW / Omodos W 1000
m / 6. IV.2009 Fritzlar leg. // Longitarsus lateripuncta-
tus det. F. Fritzlar, 2009 (FFCJ); 1 ex. (1♀!) CY Prov. /
Larnaka 71a / Tochni NE Macchia / ehem. Friedhof 100
m / 12.III.2012 Fritzlar leg. // Longitarsus lateripunctatus
det. F. Fritzlar, 2012 (FFCJ); 3 exx. (1♀!) CY Prov. Lar-
naka 72 / Lefkara 4 km WSW Vavla 350 m / 13.III.2012
Fritzlar leg. // Longitarsus lateripunctatus det. F. Fritzlar,
2012 (FFCJ); 1 ex. CY Prov. Larnaka 76 / Choirokoitia 2
km NW / Loulas Farm 100 m / 17.III.2012 Fritzlar leg. //
Longitarsus lateripunctatus det. F. Fritzlar, 2012 (FFCJ);
1 ex. CY Prov. Paphos 111 / Polis 2 km SSE / Steni 200
m NN / 34°59’58»N 32°28’26»E / 23.III.2018 Fritzlar
leg. // Longitarsus lateripunctatus det. F. Fritzlar, 2018
(FFCJ); 1 ex. CY Prov. Paphos 104 / Polis 16 km SE / Fy-
ti 650 m / 34°55’46»N 32°32’47»E / 24.III.2018 Fritzlar
leg. // Longitarsus lateripunctatus det. F. Fritzlar, 2018
(FFCJ); 1 ex. CY Prov. Paphos 109a / Polis 9km W Neo
Chorio / Pissouromoutti 350m / 35°01’13»N 32°20’01»E
/ 27.III.2018 Fritzlar leg. // Longitarsus lateripunctatus
det. F. Fritzlar, 2018 (FFCJ); 1 ex. CY Prov. Paphos 110
/ Polis 7km NW Akamas / Aphrodite-Bad Kliff 20m /
35°03’28»N 32°20’47»E / 28.III.2018 Fritzlar leg. // Lon-
gitarsus lateripunctatus det. F. Fritzlar, 2018 (FFCJ); 1 ex.
Mt. Armenien / Cypern / lateripunctatus det. Heikerting-
er / personatus mit Schulterteule (MHNB-Heik); 2 exx.
(2♂♂!) Chypre (Akamas) / Drouseia 500 m / N3456919
E03224917 / B. et M. Bergeal leg. (MSNM-Ber);
Turkey: 1 ex. (1♀!) Turcia d. Antalya 7 / Kiris b. Ke-
mer 5 km w / Forellenhof 350 m / N:36°36.27’ E:30°28.54
/ 30.III.2000 Fritzlar leg. // Longitarsus lateripunctatus
personatus det. F. Fritzlar, 2000 (FFCJ); 1 ex. Turcia d.
Antalya 15 / Kiris b. Kemer 7 km w / Gedelme Dorf 750
m / N:36°37.04’ E:30°26.29 / 31.III.2000 Fritzlar leg.
// Longitarsus lateripunctatus personatus det. F. Frit-
zlar, 2000 (FFCJ); 1 ex. Türkei d. Antalya 02 / Alanya
20 km E 50 m NN, Mahmutlar E, Karicak / N:36°27‘28“
E:032°07‘42“ / 9.IV.2001 Fritzlar leg. // Longitarsus la-
teripunctatus personatus det. F. Fritzlar, 2002 (FFCJ); 1
ex. Türkei d. Antalya 10 / Alanya 15 km ENE 200 m NN /
Kestel N Gärten / N:36°31‘53“ E:032°05‘53“ / 13.IV.2001
Fritzlar leg. // Longitarsus lateripunctatus personatus
det. F. Fritzlar, 2002 (FFCJ); 1 ex. Türkei d. Antalya 13 /
Alanya 40 km E, 200 m NN, Demirtaş 5 km N Wegrand
/ N:36°29‘43“ E:032°16‘01“ / 15.IV.2001 Fritzlar leg. //
Longitarsus lateripunctatus personatus det. F. Fritzlar,
2002 (FFCJ); 2 exx. Turcia d. Antalya 15 / Kiris b. Kemer
7 km w / Gedelme Dorf 750 m / N:36°37.04’ E:30°26.29
LAURA FARINA
21
/ 31.III.2000 Fritzlar leg. // Longitarsus lateripunctatus
personatus det. F. Fritzlar, 2000 (LFCC); 1 ex. (1♂!)
Turchia vil. Izmir / 5 km N di Bergama m. 500 / 3.V.1991
/ Audiso-Biondi-Bologna (MBCA);
Syria: 1 ex. Syria Prov. Homs / 25 km NE of Homs
/ 475 m E34°53.285’ N36°56.066’ / 4.V.2006 / leg. Gy
Rozner // Longitarsus lateripunctatus personatus Weise,
1893 det. J. Bezdék, 2015 (HNHM);
Israel: 2 exx. (1♂! 1♀!) Israel Upper Galilee / Nach-
al Amud / 26.X.1977 / leg. D. G. Furth // on Symphytum
palaestinum // macropterous // Longitarsus lateripuncta-
tus Rosenhauer det. D.G. Furth, 1977 (MSNM); 1 ex. N.
Amud // micropterous // Upper Galilee // on Symphytum
palaestinum / 21.III.1974 / leg. D. Furth // Longitarsus lat-
eripunctatus Rosenhauer det. D. G. Furth, 1975 (MSNVR);
North Africa: 1 ex. (1♀!) Algier / Reitter Leder //
Longitarsus lateripunctatus Rosenhauer coll. Reitter
(HNHM); 1 ex. Algier / Reitter // Longitarsus lateripunc-
tatus coll. Reitter (HNHM).
Dissected specimens: 33 ♂♂; 33 ♀♀.
The author reports also these data from Israel (DFCA)
taken from the photos of specimens kindly sent by David
Furth. On the labels there is interesting information about
host plants. These specimens were not studied by the au-
thor, but they seem to have the characteristics (typical
coloring) of L. personatus:
1 ex. Israel / N. Amud / 17.X.1972 / leg. D.G. Furth
/ on Symphytum palaestinum; 1 ex. Israel: Upper Jordan
Valley / Kare Deshe / 1.XII.1972 / leg. D.G. Furth; 4 exx.
Israel / N. Amud / 8.V.1973 / leg. D.G. Furth / on Symphy-
tum palaestinum; 4 exx. Israel / N. Amud / 26.VI.1973
Figs. 23-30 – Aedeagus: median lobe / lobo mediano: a) ventral view / vista ventrale; b) lateral view / vista laterale. 23-26) L.
lateripunctatus. 27-30) L. personatus. Localities / Località: 23) Algeciras; 24) Tunisi; 25) Giglio island; 26) Marina di Modica;
27) Peretola; 28, Trieste; 29-30) Krems.
NOTES ON LONGITARSUS LATREILLE 1829
22
/ leg. D.G. Furth / on Symphytum palaestinum; 1 ex. Is-
rael: Upper Jordan Valley / Betecha / 27.IV.1974 / leg.
D.G. Furth; 1 ex. Israel: Lower Galilee / Merom 1100 m /
29.IV.1974 / leg. D.G. Furth; 1 ex. Israel: Carmel Ridge /
N. Oren / 30.IV.1974 / leg. D.G. Furth / on Salvia hierosol-
ymitana; 1 ex. Israel / Golan Hts. / Qusbiye / 31.I.1978 /
leg. D.G. Furth; 5 exx. Israel / N. Amud / 23.II.1978 / leg.
D.G. Furth / on Symphytum palaestinum; 3 exx. Israel:
Northern Coastal Plain / Nahal Keziv / 4.IV.1978 / leg.
D.G. Furth / on Symphytum palaestinum; 2 exx. Israel:
Golan Hts. / Merom HaGolan / 27.IV.1978 / leg. D.G.
Furth; 10 exx. Israel / N. Amud / 30.IV.1978 / leg. D.G.
Furth / on Symphytum palaestinum; 3 exx. Israel: Carmel
/ Nahal Oren / 4.V.1978 / leg. D.G. Furth; 2 exx. Israel /
N. Amud / 7.V.1978 / leg. D.G. Furth; 1 ex. Israel: Judean
Foothills / Shoresh / 20.V.1978 / leg. D.G. Furth; 1 ex.
Israel: Mt. Hermon / Nimrod castle 800 m / 14.VI.1978 /
leg. D.G. Furth; 2 exx. Israel: Upper Jordan Valley / Ein
Gev / 15.I.1979 / leg. D.G. Furth; 5 exx. Israel: Samaria
/ Ein Sinya / 15.I.1979 / leg. D.G. Furth / on Symphytum
palaestinum; 1 ex. Israel: Upper Galilee / Nahal Betzet /
17.I.1979 / leg. D.G. Furth / on Symphytum palaestinum);
1 ex. Israel: Upper Galilee / Elon / 17.I.1979 / leg. D.G.
Furth / on Cynoglossum creticum; 1 ex. Israel: Upper Gal-
ilee / Banias / 22.II.1979 / leg. D.G. Furth; 1 ex. Israel:
Carmel Ridge / N. Oren / 3.III.1979 / leg. D.G. Furth / on
Cynoglossum creticum; 1 ex. Israel: Upper Galilee / Ha
Tanur / 6.V.1979 / leg. D.G. Furth; 3 exx. Israel: Upper
Galilee / Banias / 23.V.1979 / leg. D.G. Furth / on Symphy-
tum palaestinum; 8 exx. Israel: Carmel Ridge / N. Oren /
2.I.1980 / leg. D.G. Furth / on Echium judaeum; 2 exx.
Israel: Lower Galilee / Ha Solelim / 2.I.1980 / leg. D.G.
Furth / on Cynoglossum sp. // Longitarsus lateripunctatus
det. D.G. Furth, 1980; 1 ex. Israel: Lower Galilee / Naza-
reth / 2.I.1980 / leg. D.G. Furth / on Echium glomeratum
// Longitarsus lateripunctatus det. D.G. Furth, 1980; 1 ex.
Israel: Upper Jordan Valley / Tiberias / 10.I.1980 / leg.
D.G. Furth on Echium judaeum; 1 ex. Israel: Carmel Ridge
/ N. Oren / 4.V.1980 / / leg. D.G. Furth // Longitarsus lat-
eripunctatus det. D.G. Furth, 1980; 1 ex. Israel / Carmel /
6.XI.1980 / leg. A. Freidberg; 2 exx. Israel: Upper Galilee
/ Banias / 2.II.1981 / leg. D.G. Furth / on Cynoglossum
sp.; 6 exx. Israel: Carmel Ridge / N. Oren / 4.II.1981 /
leg. D.G. Furth / on Echium judaeum // Longitarsus la-
teripunctatus det. D.G. Furth, 1981; 1 ex. Israel: Carmel
/ N. Oren / 20.I.1984 / leg. D.G. Furth / on Cynoglos-
sum creticum; 1 ex. Israel: Carmel / N. Oren / 20.I.1984 /
leg. D.G. Furth / on Echium judaeum; 1 ex. Israel: Yizrael
valley / Tel Qashish / 20.I.1984 / leg. D.G. Furth; 3 exx.
Israel: Lower Galilee / Ha Solelim, oak forest / 22.I.1984
/ leg. D.G. Furth / on Cynoglossum creticum.
Longitarsus lateripunctatus (Rosenhauer 1856) (Fig. 6)
Teinodactyla lateripunctata Rosenhauer, 1856: 339.
Loc. typ.: Spagna: Andalusia (Algeciras).
Allard, 1860: 116-117; Allard, 1866: 133, 241-242
[Thyamis]; Portevin, 1934: 304; Kutschera, 1863: 292,
300-302 [Longitarsus]; Oertzen, 1866: 290; Weise, 1893:
964; Kolbe, 1920: 402-403; Heikertinger, 1930: 1328; Por-
ta, 1934: 357; Normand, 1937: 133; Csiki & Heikerting-
er, 1940: pars 166, 134; Müller, 1949-1953: 524, 536;
Mohr, 1966: 228 (Fig. 51: 34); Warchalowski, 1978: 49,
50 (Figs. 243-245); Furth, 1980: 101 [possibly all Israeli
specimens belong to L. personatus]; Doguet, 1989: 172;
Biondi, 1990b: 168; Biondi, 1990c: 346; Borges, 1990:
114; Gruev, 1990: 327; Doguet, 1994: 220-221, 224 (Fig.
93c), 297 (Fig. 122f); Lucht, 1994: 319; Warchalowski,
1995a: 254-256 (Figs. 954-955); Warchalowski, 1996:
137-138, 141 (Fig. 589), 142 (Fig. 590); Gruev & Döberl,
1997: 181; Warchalowski, 2003: 413; Gruev & Döberl,
2005: 90; Baviera & Biondi, 2015: 19.
= Longitarsus azoricus Israelson, 1991: 1-4 [syno-
myzed by Gruev & Döberl, 1997: 181]
= Haltica signata Reiche et de Saulcy, 1858: 49 [syno-
myzed by Allard, 1860: 116]
= Teinodactyla bigutatta Foudras, 1860: 172 [syno-
myzed by Allard, 1860: 116]
Distribution
L. lateripunctatus is distributed primarily in the west-
ern Mediterranean area.
Gruev & Döberl (1997: 181; 2005: 90) report the fol-
lowing regions:
Europe: Croatia (Dalmatia: Is. Vis), France (South;
incl. Corsica), Greece (incl. Ionian Islands), Italy (South;
incl. Sardinia and Sicily), Malta, Portugal, Spain (incl.
Baleares).
Asia: Cyprus, Greece (Rhodos), Israel.
North Africa: Algeria, Morocco, Tunisia.
Atlantic islands: Azores (Portugal).
Materials studied
Portugal: 1 ex. S. Martinho / Portugal // ex Orig.
Samig. J. Breit Wien (MHNB); 1 ex. Port Setubal / Palma
25.IV.1996 / M. Bergeal leg. (MSNM-Ber); 1 ex. Port
Algarve / Serra de Monchique / Silves 26.IV.1996 / M.
Bergeal leg. (MSNM-Ber); 1 ex. (1♂!) Port Alto Alentejo
/ Serra de Sao Mamede / Alegrete 28.IV.1996 / M. Bergeal
leg. (MSNM-Ber); 1 ex. Port Beira Baxa / Serra da Estrela
/ Corvilha 29.IV.1996 / M. Bergeal leg. (MSNM-Ber);
Spain: 9 exx. (1♂!) Spagna prov. Cadiz / dint. San
Roque / Valle F. Guadarranque / m. 100-300 10.V.1979 /
leg. M. Biondi // Longitarsus lateripunctatus Rosenhauer
det. M. Biondi (MBCA); 2 exx. (1♂!) Algeciras / An-
dalusièn / prof. G. Strobl // lateripunctatus det. F. Heik-
ertinger (MHNB-Heik) (Fig. 23); 1 ex. (1♀!) Süd-Spanien
/ Ur. Fr. Ohaner / vend. 26.IV.1915 // Algeciras (MSNM);
1 ex. P. Chanta 5 km / westlich (teich) / 17.IV.1995 leg.
Stüben / Sammlung Dieter Silde (MSNM-Ber); 10 exx.
(7♀♀! 1♂!) E. Pr. Cadix / Los Barrios / 21.IV.1995 / M.
Bergeal leg. (MSNM-Ber); 2 exx. E. Pr. Malaga / Ronda
18.IV.1996 / Al Monte Corto / M. Bergeal leg. (MSNM-
Ber); 1 ex. E Andalusien Conil / Cadix 18.II.2007 Mes-
sutat // det. M. Bergeal, 2006 (MSNM-Ber);
France: 1 ex. (1♀!) Corsica / Étang de Biguglia /
16.IV.1927 b. Furiani // Longitarsus lateripunctatus
det. F. Heikertinger (MHNB); 4 exx. Le Cannet / AM /
22.IV.1951 / P. Bonadona (MSNM-Ber);
Italy: Liguria: 1 ex. (1♂!) Liguria (SV) Sassello / 5 km
N (Ponte Erro) / 44°31.3’N 8°28.2’E / 300 m 5.VI.2017
leg. Liberti (LFCC); 1 ex. Liguria (IM) / San Remo / leg.
LAURA FARINA
23
Schneider // lateripunctatus det. F. Heikertinger (MH-
NB-Heik); 1 ex. (1♀!) Liguria / P.M. Fusan // Longitar-
sus lateripunctatus teste Dodero (MSNG); 1 ex. (1♀!)
Laigueglia / Liguria occidentale Solari (MSNM) (Fig.
13); 1 ex. (1♀!) Alassio / Liguria occidentale / 1943 //
22.V.1943 (MSNM); 1 ex. (1♀!) Liguria occidentale
/ Laigueglia / 6.VI.1961 Liberti (MSNM); 1 ex. Finale
Ligure (SV) / Altipiano di S. Bernardino / 16.VI.1987 /
S. Zoia (SZCM);
Toscana: 1 ex. (1♀!) Is. Giglio / IV.1901 / G. Doria
// Longitarsus lateripunctatus Rosenhauer teste Springer
(MSNG); 1 ex. Is. Giglio / V.1901 / G. Doria // Longitar-
sus lateripunctatus Rosenhauer teste Springer (MSNG); 1
ex. (1♂!) Toscana (GR) / Is. Giglio / XI.1902 / G. Doria
// Longitarsus lateripunctatus Rosenhauer teste Springer
(MSNG) (Fig. 25); 1 ex. Toscana (LI) / Isola Capraia / To-
scana / S. Rocco / VI.1930 C. Mancini // Longitarsus la-
teripunctatus Rosenhauer teste Springer (MSNG); 2 exx.
Toscana (GR) / Is. Giglio / G. Doria (MSNG-Dod); 1 ex.
(1♀!) Is. Giglio / V.1901 / G. Doria (MSNM); 1 ex. (1♀!)
Ric. zone umide della Toscana / Osa-La Selva / Talamone
/ (GR) / Bordoni 5.VI.2001 // Longitarsus lateripuncta-
tus s.str. det. C. Leonardi, 2003 (MSNM) (Fig. 14); 1 ex.
Riserva zone umide della Toscana / Osa (GR) / Bordoni
29.V.2002 // Longitarsus lateripunctatus s.str. det. C. Leo-
nardi, 2003 (MSNM);
Lazio: 2 exx. Lazio Torvaianica / 4.VI.1978 / leg.
M. Biondi (MBCA); 9 exx. Roma / Nuova Florida /
4.VII.1978 leg. M. Biondi (MBCA); 1 ex. Lazio / Tor-
vaianica / 15.VII.1978 / leg. M. Biondi (MBCA); 1 ex.
Lazio Roma / Settecamini / m. 100 25.X.1978 / leg. Italo
Mercati (MBCA); 1 ex. Lazio Roma / Riano Flaminio /
1.IV.1979 M. Biondi (MBCA); 2 exx. Lazio Roma / Nuo-
va Florida / 16.IV.1979 / leg. M. Biondi (MBCA); 3 exx.
Roma / Nuova Florida / 9.IV.1986 / leg. M. Biondi // su
Cerinthe sp. (MBCA); 1 ex. Roma Marcigliana / stazione
ENEL di Bufalotta / 11.V.1999 Colonnelli (MBCA);
Sardegna: 1 ex. (1♀!) Sardegna (SS) / Ploaghe 4 Km
N / 40°41.9’N 8°43.5’E / 500 m 18.V.2017 leg. G. Liberti
(LFCC); 1 ex. Sardegna 12.X.1984 / Asinara M. Scomu-
nica pendici / M. Biondi (MBCA); 2 exx. Sardegna Is. San
Pietro / Bacino acquedotto / 11.V.1988 Osella (MBCA);
1 ex. Sardegna 12.V.1988 / Loc. Sa Scrocca Manna / S.
Antioco M. Biondi (MBCA); 1 ex. (1♀!) Sardegna (OR)
Oristano / Sardinia (MHNB); 2 exx. (1♂! 1♀!) Sardegna
(CA) Sardegna / V. 1873 / Cagliari / R. Gestro // Longi-
tarsus lateripunctatus det. Dodero (MSNG); 1 ex. (1♂!)
Sardegna (OR) Asuni / Sardegna / Dr. A. Krausse / coll.
A. Dodero (MSNG-Bin); 1 ex. Sardegna (CA) Cagliari /
Marzo / Lostia (MSNG-Bin); 1 ex. (1♀!) Sardegna (SU)
Domusnovas / Sardegna / 26.V.1892 A. Dodero (MSNG-
Bin); 5 exx. (2♂♂!) Sardegna (SU) Carloforte / Sardegna
Fig. 31 – European and Mediterranean distribution of L. lateripunctatus and L. personatus. / Distribuzione europea e meditterranea di L. lat-
eripunctatus e L. personatus. Blank map modied from / Cartina modicata da: https://d-maps.com/ carte.php?num_car=3127&lang=it
NOTES ON LONGITARSUS LATREILLE 1829
24
/ 2.V.1912 A. Dodero (MSNG-Bin); 1 ex. Cagliari Cor.
/ 2.VI.1901 A. Dodero (MSNG-Dod); 9 exx. (1♂!) Sar-
degna / Flumini / 3-8.V.1971 / leg. C. Leonardi (MSNM);
4 exx. (3♂♂!) Sardegna Ruinas / 10-13.V.1972 / leg. C.
Leonardi (MSNM); 2 exx. (1♂! 1♀!) Sardegna / Riola
/ 12.V.1972 / leg. C. Leonardi (MSNM); 12 exx. (3♀♀!
1♂!) Sardegna (SU) / Sadali / 17.V.1972 / leg. C. Le-
onardi (MSNM); 6 exx. (1♂! 1♀!) Sardegna / Ortueri /
19.V.1972 / leg. C. Leonardi (MSNM); 2 exx. Sardegna
/ Alghero / 23.V.1973 / leg. C. Leonardi (MSNM); 1 ex.
Iglesias Lago Corsi / (Cagliari) m 250 ca / 20.IV.1997 C.
Giusto (SZCM);
Basilicata: 1 ex. (1♂!) Basilicata / str. Irsina-Grassano
/ su F. Bradano (MT) 150 m / 22.IX.1996 leg. Angelini
(MSNM-Ber);
Puglia: 3 exx. (3♂♂!) Puglia Promontorio del Garga-
no W Vieste / N41°53’07’’E16°07’54’ 23 m / Olivenhain
/ 2.IV.2016 / leg. C. Germann // Longitarsus lateripunc-
tatus Rosenhauer det. L. Farina, 2020 (CGCB) (Fig. 6); 2
exx. Puglia (TA) / Bosco di San Basilio / 12.V.2018 / leg.
F. Montemurro (FMCT); 5 exx. Puglia (TA) / Bosco di
San Basilio / 27.V.2018 / leg. F. Montemurro (FMCT); 1
ex. (1♀!) Puglia (TA) / Circummarpiccolo / 26.V.2012 /
leg. F. Montemurro (LFCC) (Fig. 15); 1 ex. (1♂!) Puglia
/ Murge Tarantine / San Basilio 25.VI.2011 / leg. C. Le-
onardi (MSNM);
Calabria: 1 ex. Calabria (CZ) / Marina di Nicotera /
12.XI.1989 / Duna costiera / leg. M. Bologna & C. Ma-
rangoni (MBCA); 2 exx. (1♂!) Calabria / Vibo Valentia /
22.IV.1990 / leg. C. Leonardi (MSNM); 5 exx. (1♂! 1♀!)
Calabria / Capo Vaticano / 22.IV.1990 / leg. C. Leonardi
(MSNM); 2 exx. (1♂) Calabria / fra Ioppolo e Nicotera
23.IV.1990 / leg. C. Leonardi (MSNM); 2 exx. Calabria
/ Samo / 25.IV.1990 / leg. C. Leonardi (MSNM); 1 ex.
Calabria (RC) / Samo / 25.IV.1990 / leg. C. Leonardi
(MSNM); 3 exx. Calabria / Cirò Marina / 29.IV.1990 /
leg. C. Leonardi (MSNM);
Sicilia: 5 exx (1♂!) Sicilia (SR) Pachino / 13-17.V.1908
/ A. Dodero (MSNG-Dod); 1 ex. (1♂!) Sicilia (ME)
Messina / Sicilia / F. Vitale (MSNG-Man); 1 ex. (1♀!)
Messina / Peloritani / 8.?II.1916 / Schatzmayr (MSNM)
(Fig. 16); 5 exx. (1♂!) Sicilia / Grotta Monello (SR) /
18.VI.1991 / leg. Sabella // Longitarsus lateripunctatus
det. C. Leonardi (MSNM); 5 exx. (1♀!) Sicilia / Grotta
Monello (SR) / 28.VIII.1991 / leg. Sabella // Longitar-
sus lateripunctatus det. C. Leonardi (MSNM); 45 exx.
(1♂! 1♀!) Sicilia / Kamarina 6.V.1992 / leg. C. Leonardi
(MSNM); 1 ex. Sicilia 13.V.1992 / foce Fiume Belice /
leg. C. Leonardi (MSNM); 1 ex. (1♂!) Sicilia Siracusa
Lentini / su piante di agrumi / 17.V.2000 / leg. L.S.A. //
Longitarsus lateripunctatus (Rosenh.) det. C. Leonardi,
2004 (MSNM); 19 exx. (1♂!) Sicilia 23.IV.2004 / Ma-
rina di Modica / leg. C. Leonardi (MSNM); 1 ex. Sicilia
23.IV.2004 / Pozzallo-Marina di Modica / leg. C. Leon-
ardi (MSNM) (Fig. 26); 1 ex. Favignana / M.S. Caterina
250 m / 37°55.7N 12°18.8E / 12.V.2008 Liberti (MSNM);
1 ex. (1♂!) Favignana / M.S. Caterina 250 m / 37°55.7N
12°18.8E / 18.V.2008 Liberti (MSNM); 1 ex. Fiumara To-
no (Messina) // Sicilia / VI.1959 // Longitarsus lateripunc-
tatus (Rosh.) det. A. Warchalowski (MSNVR); 1 ex. Cava
d’Ispica (RG) / 3-13.VI.2014 leg. R. Papi (RPCR);
Malta: 1 ex. (1♂!) Malte / Mellieha 1.VI.2004 / Rolf
Rober (MSNM-Ber);
Montenegro: 1 ex. (1♀!) Sutorina / leg. Paganetti //
lateripunctatus det. F. Heikertinger (MHNB-Heik);
Greece: 1 ex. (1♀!) Peloponnes S-Pylos / Mesochori /
env. 36.880430, 21.713645 25 m / 27.IX.2016 leg. C. Ger-
mann (CGCB); 1 ex. (1♂!) Corfù / Paganetti // Longitar-
sus lateripunctatus Rosenhauer det. K. Lopatin (HNHM);
1 ex. Corfù / Paganetti // coll. Dr. J. Fodor // Longitarsus
lateripunctatus Rosenhauer det. B. Gruev (HNHM); 1 ex.
Creta 9.IV.1989 / Foci del Petrès / P. Audisio (MBCA);
1 ex. (1♀!) Corfù / V. d. Ropa // ex Orig. Samig. J. Breit
Wien (MHNB) (Fig. 17); 1 ex. (1♀!) Kephallenia / leg.
Paganetti // ex Orig. Samig. J. Breit Wien (MHNB); 1 ex.
Corfù / leg. Paganetti // lateripunctatus Rosenhauer det. F.
Heikertinger (MHNB-Heik);
North Africa: 1 ex. Algier / Reitter Leder / coll. F.
Speiser // Longitarsus lateripunctatus det. Warchalowski
1960 (HNHM); 1 ex. Algerie m. 1000 / P.N. d’Akfadou /
for. Des Beni Ghorbi / 13.IV.1988 Colonnelli (MBCA); 1
ex. Algerie m. 600 / wil. Tizi Ouzou / Col de Chergoui /
16.IV.1988 Colonnelli (MBCA); 7 exx. (2♂♂ 1♀!) Alge-
ria // leg. Peyerimhoff // lateripunctatus Rosenhauer det.
Heikertinger (MHNB-Heik); 1 ex. St. Charles/ Algérie /
A. Thery (MHNB-Heik); 1 ex. (1♂!) Les Glacières / Bli-
da Jordan (MHNB-Heik); 3 exx. (1♂!) Mustapha / Alge-
ria (MSNG-Man); 5 exx. Sellaoua Announa / 29.III.1985
Algerie / M. Bergeal leg. (MSNM-Ber); 1 ex. (1♀!) Dl.
Mahouna / Guelma (Algerie) / M. Bergeal 4.IV.1985
(MSNM-Ber); 1 ex. (1♀!) Oued Mafraq / Annaba (Alge-
rie) / M. Bergeal leg. 6.IV.1985 (MSNM-Ber);
1 ex. Tunis 1904 / Ujbeiyi // Longitarsus lateripuncta-
tus det. Warchalowski 1962 (HNHM); 1 ex. (1♀!) Tunis /
Hamm. Lif. // Nahtsaum erweitert Schultereck // Longi-
tarsus lateripunctatus det. F. Heikertinger (MHNB-Heik);
1 ex. (1♂!) Tunisi dint. / 1881 G. e L. Doria // Longitarsus
lateripunctatus teste Dodero (MSNG) (Fig. 24);
1 ex. (1♂!) Marocco prov. Taza / Cascate di Ras-el-
Oued / 15.V.1979 m. 900 ca / leg. M. Biondi (MBCA).
Dissected specimens: 40 ♂♂; 38 ♀♀.
Longitarsus monticola Kutschera 1863 (Fig. 7) and
Longitarsus curtus (Allard 1860) (Fig. 8)
L. monticola was originally described on a few speci-
mens from Styria. Kutschera (1863: 294) reported ap-
terism and colour differences against L. curtus. Later
on, most authors considered monticola as a variety or a
subspecies of curtus. First Doguet (1984: 302-303; 1994:
289-293) in careful studies of both species reported sig-
nicant differences in the habitus and shape of genitalia.
Döberl (1994: 110; 1995: 40-41) considered these two
taxa as valid species reporting diagnostic characters based
on a substantial sample.
Comparative distribution
According to Döberl (1995: 40-41), L. curtus is wide-
spread in Europe, while L. monticola is mainly distibuted
in the Alpine Region. According to Doguet (1994: 289-
293), L. curtus seemingly shows a more western Euro-
pean distribution than L. monticola. Later, a map of the
LAURA FARINA
25
Palaearctic distribution of both species was reported by
Warchalowski (1996: 118 Fig. 571).
Although several papers provided more information
about their Italian distribution (Leonardi, 1973: 35; Bi-
ondi, 1988: 135; 2005; D’Alessandro & Biondi, 2005: 91,
96, 104, 108; D’Alessandro & Biondi, 2007: 465, 471;
Baviera & Biondi, 2015: 21; Angelini, 2020: 196), in the
author’s opinion, the knowledge of their ranges in Italy
is still incomplete, also due to the possible confusion be-
tween these similar species.
As a result of the study of a large sample, it has been
possible to provide both new diagnostic characters and
new distributional data and to attempt the production of
Italian distribution maps for both species (Fig. 44). As can
be seen, L. curtus is widespread in Northern Italy and L.
monticola is occurring almost everywhere in the Penin-
sula, showing a preference for mountain areas.
All the specimens in MBCA, from Central and South-
ern Italy, are L. monticola. The other specimens (reported
as L. curtus) indicated in the Checkmap (Biondi, 2005)
from Apennines and Southern Italy (and not studied in the
present work) in the author’s opinion, are also attributable
to L. monticola.
Comparative description
A morphological description of both species can be
found in Doguet, 1984: 302-303; 1994: 289-293; Döberl,
1995a: 40-41 and Warchalowski, 1995b: 107-110; 1996:
87-89, 157-158. It is worth noting that these species in-
clude both forms with complete wings and forms with re-
duced wings. For discriminant characters see Table 2.
Longitarsus monticola Kutschera 1863
Longitarsus monticola Kutschera, 1863: 294; VIII,
1864: 44-46; pag. Sp.: 22, 259. Locality indicated: Aus-
tria: Styria [= Steiermark] (Maria Zell) [type lost].
Allard, 1866: 136, 204-205 [Thyamis]; Redtenbacher,
1874: 506 [Longitarsus]; Döberl, 1994: 110, 139 (Figs.
317-319); Doguet, 1994: 272 (Fig. 110d), 292-293, 300
(Fig. 124h) [in the description of the species the author
wrote “Remarque: longtemps considerée comme syn-
onyme de L. curtus, cette espèce a été récemment réha-
bilitée (Döberl, op. cit.)”. In the references Doguet put the
paper of Döberl with this indication: Longitarsus monti-
cola Kutschera, 1863 spec. propria (sous presse)”]; Lucht,
1994: 319; Döberl, 1995a: 40-41 [denition of the Neo-
type, ♀, NHMW, typ. loc. Kärnten];berl, 1995b: 57 (“In
die Sammlungen gewöhnlich als curtusaufgenommen”);
Warchalowski, 1995a: 180 (Figs. 724-725), 183 (Fig. 749),
229-231 (Fig. 936); Warchalowski, 1995b: 107-110; Ro-
zner, 1996: 255; Warchalowski, 1996: 76 (Figs. 359, 360),
79 (Fig. 384), 157-158; Geiser, 1997: 68; Gruev & Döberl,
1997: 191-192; Fritzlar, 1998: 203, 210; Gruev & Tomov,
1998: 95; Fritzlar, 1999: 68, 75; Döberl et al., 2000: 70;
Bezděk & Zúber, 2001: 149 (“L. monticola resurrected by
Döberl”); Geiser, 2001: 465; Kapp, 2001: 63, 507; Vig,
2002: 125; Brelih et al., 2003: 100-101; Fritzlar, 2003: 130;
Warchalowski, 2003: 445; Geiser, 2004: 19; Biondi, 2005:
data on CD-Rom; D’Alessandro & Biondi, 2005: 91, 96,
104, 108; Fritzlar, 2005: 232, 238-239; Gruev & Döberl,
2005: 95; Čižek, 2006: 26, 27 (Fig. 78); D’Alessandro &
Biondi, 2007: 465, 471; Fritzlar, 2009: 195, 207; Baviera &
Biondi, 2015: 21; Fritzlar, 2017: 81; Angelini, 2020: 196.
Tab. 2 - Taxonomically discriminant characters of L. monticola and L. curtus. / Caratteri tassonomici
discriminanti in L. monticola e L. curtus.
Character L. monticola Kutschera 1863 L. curtus (Allard 1860)
Median lobe of aedeagus Median lobe of aedeagus (Figs. 32-36) in
ventral-dorsal view with straight (slighty
sinuous) sides, narrowed apically. Next
to median groove (mg, Fig. 33a) a weak
and short lateral groove (lg, Fig. 33a),
sometimes replaced by longitudinal
wrinkles (Fig. 35a). In lateral view distal
2/3s clearly bent, apically shortly sinuate,
apical tip usually bent upwards.
Median lobe of aedeagus (Figs. 37-39) in
ventral-dorsal view with straight sides, less
narrowed apically. Both margins sometimes
with weak wrinkles, without grooves.
In lateral view almost straight, in a few
specimens distal 2/3s slightly bent, but at
apex straight or almost so.
Spermatheca Basal part of spermatheca narrow and
gradually widened apically (Figs. 40-41;
also: Doguet, 1984: 302-303; Döberl,
1995: 40-41; Warchalowski, 1995a: 107-
110). Ductus insertion in basal part located
slightly forwards (but a variable character).
Basal part of spermatheca wider in the
middle, sometimes sub-cylindric (Figs. 42-
43; also: Doguet, 1984: 302-303; Döberl,
1995: 40-41; Warchalowski, 1995a: 107-
110). Ductus insertion in basal part often
located slightly backwards (but a variable
character).
Wings Hind wings reduced, humeral calli not
or weakly developed in most examined
specimens (82%).
Hind wings usually well-developed,
humeral calli evident in most examined
specimens (62%). In the sample, among the
winged specimens, females are the majority
(67%) while among the reduced wings
specimens, males prevail (68%).
NOTES ON LONGITARSUS LATREILLE 1829
26
Figs. 32-43 – 32-39) Aedeagus: median lobe / lobo mediano: a) ventral view / vista ventrale; b) lateral view / vista laterale. 32-36) L.
monticola. 37-39) L. curtus. 40-43) Spermatheca. 40-41) L. monticola. 42-43) L. curtus. Localities / Località: 32) Raibl; 33) Orso-
marso, Grisolia Pantanelle; 34) Costacciaro Monte Cucco; 35) Westfalen; 36) Paularo; 37) Cattaro; 38) Nevea Cregnedul; 39) Beira
Altas Manteigas; 40) Campigna; 41) Costacciaro Colle d’Orlando; 42) Nevea Cregnedul; 43) Beira Altas Manteigas. mg) median
groove / solco mediano; lg) lateral groove / solco laterale.
LAURA FARINA
27
= Longitarsus curtus, Leonardi, 1973: 35 (Fig. 62);
Gruev & Tomov, 1986: 206-207 (Fig. 225); Biondi, 1988:
135; Döberl, 1995b: 53
= L. curtus ab. monticola, Heikertinger, 1930: 1326
(forma aptera); Csiki & Heikertinger, 1940: pars 166,
119; Kaszab, 1962: 311.
= L. curtus fa. monticola, Král, 1945: 32; Mohr, 1962:
71 (Fig. 56), 88-89
= L. curtus mut. monticola, Kolbe, 1920: 402-403, 404
[referring to monticola Kolbe wrote “es ist anscheinend
eine werdende neue Art”].
= L. curtus var. monticola, Weise, 1893: 977; Heik-
ertinger, 1912: 193; Kuhnt, 1913: 885; Porta, 1934: 359;
Portevin, 1934: 303; Müller, 1949-1953: 524; Mohr, 1966:
229 (Fig. 51: 18), 230; Warchalowski, 1978: 64 (Figs.
346-347), 66 (Fig. 363), 68; Doguet, 1984: 302-303.
Distribution
L. monticola shows a preference for mountain areas.
Gruev & Döberl (1997: 191; 2005: 95) report the fol-
lowing regions:
Europe: mountain of Albania (Kelmend, Nikci/
Shkenbejt e Jarnes), Austria, Bulgaria, Carpathian Basin,
Czechia, France, Germany (South, Middle, West), Hun-
gary, Italy, Poland, Rumania (Transylvania), Slovakia,
Slovenia, Switzerland.
Asia: Caucasus, Russia, Tajikistan.
According to several papers on the Italian distribution
of these two species, L. monticola is recorded from sev-
eral localities: Raibl (Leonardi, 1973: 35 Fig. 32); Monte
Raut (Carnia), Gressoney (Aosta), Nava (Maritime Alps),
Monte Penna (Ligurian Apennines), Gargonza (Arezzo),
Campigna (Romagna), Maddalena Mountains (Potenza),
Monti Peloritani, Boschi di Malabotta (Messina), Monti
Peloritani and Nebrodi (Biondi, 1988: 135; Baviera &
Biondi, 2015: 21); Rocca di Cambio (L’Aquila) (Biondi,
2005: data on CD-Rom); Altopiano delle Rocche (Sirente-
Velino mountain area) (D’Alessandro & Biondi, 2005: 91,
96, 104, 108; D’Alessandro & Biondi, 2007: 465, 471);
Central Lucan Apennines (Angelini, 2020: 196).
Materials studied
Germany: 2 exx. (1♂! 1♀!) Bayerischer Wald / Regen-
tal b. Teisnach / 15.VIII.1987 / leg. Döberl // Longitarsus
monticola det. Döberl (MBCA); 1 ex. (1♂!) Bayerischer
Wald / Rusel +700 m / 3.IX.1980 leg. Döberl // Longitar-
sus monticola det. M. Döberl, 1981 (MSNM-Ber); 2 exx.
(1♂! 1♀!) Bayerischer Wald N. Teisnach / 30.VIII.1984
leg. Döberl (MSNM-Ber); 1 ex. (1♂!) D. Bayer. Wald /
27.VII.1988 M. Bergeal (MSNM-Ber); 1 ex. (1♀!) Au
Col des Brunner / 3.VIII.1991 / M. Bergeal // Longitarsus
monticola / det. M. Bergeal 1994 (MSNM-Ber); 2 exx.
(1♂!) Umg. Weilheim Obb. / Hardt-Wald / leg. Stein-
hausen // Longitarsus monticola det. Steinhausen (ZSM);
Italy: Piemonte: 1 ex. (1♂!) Piemonte (CN) Limone /
Piemonte / Agosto 1892 / Dodero (MHNB-Heik); 2 exx.
(2♂♂!) Piemonte / Coll. Fej (MSNG); 2 exx. (1♂! 1♀!)
Piemonte (BI) / Alpe Finestre / 24.VI.1922 / leg. F. Capra
(MSNG); 2 exx. (1♂!) Piemonte (BI) / Piedicavallo / Val
Cervo / Biella Piemonte / 10.IX.1956 F. Capra // Longitar-
sus curtus det. M. Biondi 1989 (MSNG); 1 ex. (1♀!) Pie-
monte // Longitarsus curtus det. Springer (MSNG-Bin);
9 exx. (1♂! 7♀♀!) Piemonte (BI) / Santuario di Oropa /
Biellese (Piemonte) / VII.1927 / A. Dodero (MSNG-Bin);
2 exx. (2♀♀!) Piemonte (CN) / Crissolo Baucl (MSNG-
Dod); 4 exx. (2♂♂! 2♀♀!) Piemonte (CN) / Limone / Pie-
monte / Agosto 1892 / A. Dodero (MSNG-Dod); 1 ex. (1♀!)
Piemonte (BI) / Santuario di Oropa / Biellese (Piemonte) /
VIII.1926 / A. Dodero (MSNG-Dod); 1 ex. (1♀!) Piemonte
(BI) / Santuario di Oropa / Biellese (Piemonte) / VI.1927 /
A. Dodero (MSNG-Dod); 4 exx. (2♀♀!) Piemonte (BI) /
Santuario di Oropa / Biellese (Piemonte) / VII.1927 / A. Do-
dero (MSNG-Dod); 1 ex. (1♀!) Piemonte (VC) / Val Sesia //
Longitarsus curtus All. det. Springer (MSNG-Man);
Lombardia: 16 exx. (1♂! 1♀!) Pasturo / Monte Grigna
Settentrionale versante E / Foppa del Ger 1420-1500 m /
12.IX.2013 / R. Regalin (FLCC) (Fig. 7); 1 ex. (1♀!) Antea
VII.1959 S. Pellegrino / Leonardi (MSNM); 1 ex. (1♀!)
Lombardia / Bratto / 7.VII.1973 / leg. C. Leonardi (MSNM);
Trentino Alto Adige: 1 ex. (1♂!) Trentino (TN) Moena
Val di Fassa / Trentino m. 1300 / 5.VIII.1941 C. Conci
// Longitarsus curtus det. C. Leonardi (MSNG); 1 ex.
(1♂!) Trentino (TN) Südtirol Nonstal / St. Felix 1250 m /
2.IX.1979 Döberl // Longitarsus monticola det. M. Döberl
1982 (MSNM-Ber);
Veneto: 1 ex. (1♀!) Lessini Alpen / Ital. Grigital //
Longitarsus curtus All. det. Warchalowski (HNHM); 3
exx. (1♂!) Veneto (BL) Cansiglio / Pian Osteria 2 km E /
46°03.7’N 12°25.5’E / 1052 m 4.VII.2017 Liberti (LFCC);
2 exx. (1♂!) Cadore / Voltago / m. 800 / 24.VIII.1958 /
Fig. 44 – Italian distribution of L. monticola and L. curtus. / Distribu-
zione italiana di L. monticola e L. curtus.
NOTES ON LONGITARSUS LATREILLE 1829
28
leg. Italo Mercati (MBCA); 1 ex. (1♀!) Belluno / Voltago
Agordine / m. 850 25.VIII.1959 / leg. Italo Mercati (MB-
CA); 1 ex. (1♀!) Dolomiti / Voltago / Mercati // m. 800 /
2.IX.1959 (MBCA); 1 ex. (1♂!) Belluno / Voltago Ago-
rdino / m. 850 4.VIII.1961 / leg. Italo Mercati (MBCA);
6 exx. (1♂! 3♀♀!) Dolomiti / Agordino / Colonnelli leg.
// Voltago / m. 900 / 16.VIII.1970 (MSNM); 1 ex. (1♀!)
VR Lessini / 16.V.1976 leg. Sette // Fittanze / mt. 1300
// curtus / det. C. Leonardi (MSNM); 1 ex. (1♀!) Prealpi
vicentine // Laghi / Vanzi 18.V.1980 (MSNM); 26 exx.
(2♂♂! 1♀!) Veneto (BL) / I-Dolomiti Agordino Voltago
/ VIII.1959 leg. Mercati (MSNM-Ber); 1 ex. (1♀!) VR /
Prun // 15.IX.1969 / leg. Zanetti // Long. curtus det. Mohr
(MSNVR);
Friuli Venezia Giulia: 2 exx. (1♂! 1♀!) Carnia /
Forni di Sopra / A. Gagliardi (MSNM); 2 exx. (2♂♂!)
Friuli Venezia Giulia (UD) / Raibl / VIII.1928 / Scharte /
Springer (MSNM) (Fig. 32); 5 exx. (4♂♂!) Friuli Venezia
Giulia (UD) / Paularo / VIII.1929 / A. Gagliardi (MSNM)
(Fig. 36); 2 exx. (1♂! 1♀!) Carnia VIII.1929 / Paularo
(MSNM); 1 ex. Friuli Venezia Giulia (UD) / Jul. Alp.
Raibl / Scharré / 1948 / Springer (MSNM);
Liguria: 1 ex. (1♀!) Alpi Liguri Collardente / Foresta
Gerbonte m. 1800 / 25.VIII.1991 leg. M. Biondi (MBCA);
1 ex. (1♂!) Liguria (GE) / Monte Penna / Agosto 1897 /
Dodero (MHNB-Heik); 9 exx. (1♂! 7♀♀!) Liguria (GE)
/ Monte Penna / Agosto 1897 / A. Dodero (MSNG-Bin);
7 exx. (4♀♀!) Liguria (GE) / Monte Penna / Agosto 1897
/ A. Dodero (MSNG-Dod); 4 exx. (1♂! 2♀♀!) Liguria
(GE) / S. Lorenzo di Cas. / Genova / F. Solari (MSNM);
4 exx. (4♂♂!) Liguria / Monte Penna / 3.VIII.1972 / leg.
C. Leonardi (MSNM); 1 ex. (1♀!) Alpi Liguri (Garessio)
/ pendici Nord M. Galero / 1300 m 29-30.V.1976 / A. e L.
Briganti (SZCM);
Emilia Romagna: 1 ex. (1♀!) Romagna / Foresta di
Campigna / 18-23.VIII.2018 / leg. F. Montemurro (FM-
CT); 1 ex. (1♀!) Campigna (FC) / 23.VI.1972 / leg. Le-
onardi (MSNM) (Fig. 40); 1 ex. (1♀!) Romagna / Foresta
di Campigna / 20-23.VIII.2017 / leg. F. Montemurro
(MSNM);
Toscana: 3 exx. (1♂! 1♀!) Toscana (AR) / F. Camal-
doli / 26.VIII.2005 / Zoia (LFCC); 3 exx. (2♀♀!) Toscana
(PT) / Boscolungo Pistoiese / VII-VIII.1891 / A. Dodero
(MSNG-Dod);
Umbria: 1 ex. (1♀!) Umbria (PG) / Lippiano / Arezzo
/ Andreini // Longitarsus / curtus / det. M. Biondi, 1989
(MSNG) (on April, 15th 1927 the municipality of San-
ta Maria, with Lippiano, became Province of Perugia,
RDL 31/3/1927 n. 468, GU n. 86 del 13/4/1927); 1 ex.
(1♂!) Umbria (PG) / Lippiano Arezzo / VII-VIII.1921 /
A. Andreini // Longitarsus curtus det. M. Biondi 1989
(MSNG); 1 ex. (1♂!) Umbria (PG) Scheggia e Pascelupo
/ loc. Valdorobia / rive f. Sentiero 520 m / 3.V.2015 M.
Bocci (MSNM); 1 ex. (1♂!) Umbria (PG) Costacciaro
/ str. per Monte Cucco sentiero /n. 24 650 m 18.X.2015
/ leg. M. Bocci (MSNM) (Fig. 34); 1 ex. (1♀!) Umbria
(PG) Scheggia e Pascelupo / Perticano Fossa / Rave 400
m 19.VI.2016 / leg. F. Angelini & M. Bocci (MSNM); 1
ex. (1♀!) Umbria (PG) Costacciaro / Colle d’Orlando /
Fonte Acqua Passera 1140 m / 15.V.2016 leg. M. Bocci
(MSNM) (Fig. 41); 1 ex. (1♂!) Umbria (TR) / Italien /
Umbrien b. Orvieto c.a. 500 m NN / VIII.1997 leg. M.
Langer (MSNM-Ber);
Lazio: 1 ex. (1♀!) Colli Albani / M. Artemisio /
4.IV.1980 / P. Audisio (MBCA); 1 ex. (1♀!) Lazio (VT) /
Selva di Lamone / F.so Olpeta 9.VIV.1983 / leg. P. Audi-
sio (MBCA); 1 ex. (1♀!) Capricchia (RI) / m. 1100-1500
// M.ti della Laga / IX.1968 (MSNVR);
Abruzzo: 1 ex. (1♂!) Abruzzo Zittola (AQ) / Castel di
Sangro / 17.XI.1990 / Fiordigigli-Osella (MBCA); 3 exx.
(1♂! 1♀!) dint. N Pratola Peligna / m 340 (L’Aquila) /
8.IX.1990 / S. Zoia leg. (SZCM);
Sardegna: 1 ex. (1♂!) Sardinia / Reitter // Longitarsus
curtus coll. Reitter (HNHM) (generally indicated for Sar-
dinia on the map Fig. 44);
Campania: 1 ex. (1♀!) Campania (NA) Ottaviano /
sentiero per i Cognoli 600 m. ca. / pitfall traps castagneta
mista / UTM: 33TVF54102040 / leg. G. Nardi (MBCA);
Basilicata: 1 ex. Abriola (PZ) / leg. Angelini (MSNM);
1 ex. (1♂!) Basilicata / Pignola / Lago di Pantano (PZ)
770 m / 19.X.1996 / trapp. aceto / leg. Angelini (MSNM);
1 ex. (1♀!) Basilicata 800 m / Castelsaraceno (PZ) / Tor-
rente Cagliandrino / 28.X.2001 / leg. Angelini // Longi-
tarsus monticola Kutschera det. C. Leonardi (MSNM); 1
ex. (1♀!) Basilicata / La Maddalena (PZ) / Abriola 1400
m / 9.V.2003 Angelini // Longitarsus monticola Kutschera
det. C. Leonardi (MSNM);
Calabria: 1 ex. (1♂!) Calabria Orsomarso / Grisolia
Pantanelle (CS) / 11.IX.1993 700 m / leg. F. Montemur-
ro // Longitarsus monticola Kutschera det. C. Leonardi
(MSNM) (Fig. 33);
Sicilia: 1 ex. (1♂!) Sicilia (ME) M.ti Peloritani /
Foresta di Malabotta m. 1185 / cerreta 24.V.1981 / leg. M.
Biondi (MBCA); 1 ex. (1♀!) Sicilia (ME) M.ti Peloritani
/ Foresta di Malabotta m. 1185 / cerreta 19.VII.1981 / leg.
M. Biondi (MBCA); 1 ex. (1♂!) Sicilia (ME) M.ti Pelori-
tani / Foresta di Malabotta m. 1185 / cerreta 10.VIII.1981
/ leg. M. Biondi (MBCA); 8 exx. (4♂♂! 4♀♀!) Sicilia
(ME) M.ti Peloritani / Foresta di Malabotta m. 1185 / cer-
reta 17.IX.1981 / leg. M. Biondi (MBCA); 1 ex. (1♀!)
Sicilia (ME) M.ti Peloritani / Foresta di Malabotta m.
1215 / faggeta X.1981 / leg. M. Biondi (MBCA); 1 ex.
(1♀!) Sicilia (ME) M.ti Peloritani / Foresta di Malabotta
m. 1185 / cerreta 26.X.1981 / leg. M. Biondi (MBCA);
1 ex. (1♂!) Sicilia (ME) M.ti Nebrodi / P.lla Maulazzo
m. 1460 / faggeta 21.XI.1981 / faggeta X.1981 / leg. M.
Biondi (MBCA); 1 ex. (1♂!) Sicilia (ME) M.ti Peloritani /
Foresta di Malabotta m. 1185 / cerreta 2.III.1982 / leg. M.
Biondi (MBCA); 1 ex. (1♂!) Sicilia (ME) M.ti Peloritani
/ Foresta di Malabotta m. 1185 / cerreta 30.III.1982 / leg.
M. Biondi (MBCA);
Lithuania: 1 ex. (1♂!) Babiagura / Natterer //
30.VII.1911 (MHNB-Heik);
Poland: 2 exx. (2♀♀!) Obiźowa / P. Zaleserpyti /
18.IX.1936 // Polonia / S. Tenk Taum (MHNB-Heik);
Austria: 1 ex. (1♂!) Lunz / Schuster // monticola Kut-
schera det. Z. Kaszab (HNHM); 1 ex. (1♂!) Hesersdf. /
IX.1924 / Heikertinger Longitarsus curtus All. det. Dr. Z.
Kaszab 1962 (HNHM); 1 ex. (1♂!) Turnau / Styr. Bor. //
ex Orig. Samig. J. Breit Wien // Longitarsus curtus det.
C. Leonardi (MHNB); 1 ex. (1♂!) Neudorf Hu. / Ödenbg.
Com. // ex Orig. Samig. J. Breit Wien // Longitarsus curtus
LAURA FARINA
29
det. C. Leonardi (MHNB); 1 ex. (1♀!) Helmgeblet / Tyr
IX.1928 // curtus det. F. Heikertinger (MHNB); 1 ex. (1♂!)
Villgratnertal / Tyr VIII.1929 // curtus det. F. Heikertinger
(MHNB); 1 ex. (1♀!) Weitlanbrunn / Tyr 25.VIII.1931 //
curtus det. F. Heikertinger (MHNB); 1 ex. (1♂!) West-
falen (MHNB-Heik) (Fig. 35); 1 ex. (1♀!) Damberg / Aus-
tr. Sup. Mont. / Petz (MHNB-Heik); 1 ex. (1♂!) Lunz /
Schuster // L. viduus / don. Reitter // v. Reitter als L. viduus
erhalten! (MHNB-Heik); 2 exx. (1♂! 1♀!) Shoberstein /
Austr. Sup. / Mont. Petz (MHNB-Heik); 1 ex. (1♀!) Car-
inthia / v. Weise (MHNB-Heik); 2 exx. (2♀♀!) Maria Zell
700-1000 / 21.VIII.1908 / Heikertinger (MHNB-Heik); 1
ex. (1♀!) Neuhaus b. Gam / 22.VIII.1908 Heikertinger //
curtus det. F. Heikertinger (MHNB-Heik); 1 ex. (1♀!) Hof
Mor. / Scheerpeltz // Longitarsus curtus det. F. Heikerting-
er // coll. A. Simonis dupleta coll. Winkler (MSNG); 1
ex. (1♀!) Wien Umg / Heikertinger (MSNM); 1 ex. (1♀!)
Lautsch. / Alp Styr / Messa (MSNM); 1 ex. (1♀!) Terri-
torio del Dobratsch / A. Sazmaier (MSNM); 1 ex. (1♀!)
Austria Kürnten / Karawanken / doch-Obir 9.VIII.1983 /
leg. M. Döberl // Longitarsus curtus det. M. Döberl, 1984
(MSNM-Ber); 1 ex. (1♂!) Austria / b. Fiss c.a. 1440 m /
26.VIII.1998 leg. M. Langer (MSNM-Ber); 1 ex. (1♂!) A.
Tirol (LA) / b. Fiss c.a. 1440 m / IX.2000 leg. M. Langer
gekeschert (MSNM-Ber);
Slovenia: 1 ex. (1♀!) Waladir / Longitarsus curtus det.
F. Heikertinger // L. curtus coll. Reitter (HNHM);
Hungary: 1 ex. (1♀!) Lazsiscsinai völgy // Marmaros /
leg. Pável 1875 Longitarsus curtus All. det. Dr. Z. Kaszab
1962 (HNHM); 1 ex. (1♀!) Tátra / 16.VIII.1890 / Birό L.
// Longitarsus curtus All. det. Dr. Z. Kaszab 1962 (HN-
HM); 1 ex. (1♂!) Hu. Occ. 1951 / Velencei-hegys / Na-
dap / Meleg-hegy // X.1915 / erdőszél rostálva // legit Dr.
Kaszab // monticola Kutschera det. Z. Kaszab (HNHM);
2 exx. (1♂! 1♀!) Dobogό-kő / 10-19.VIII.1917 / Révy D.
// coll. Dr. D. Révy // monticola Kutsch. det. Z. Kaszab
(HNHM); 1 ex. (1♀!) Hung. N.P. Miskolc / Nagy-me
850 m / Lolio-Cynosuretum / fühálόzás / 23.IX.1982 / leg.
Vágό E. // Longitarsus curtus All. det. B. Gruev (HNHM);
1 ex. (1♀!) Hung. Bükki N. P. / Bükkzsére / Szarba-völgy
// Phyllitidi-Aceretum / fényre / 29.IX.1982 / leg. Vágό E.
// Longitarsus curtus All. det. B. Gruev (HNHM); 1 ex.
(1♀!) Hung. Bukki N.P. / Kisgyόr 450 m / Gyertyán-völgy
/ Querco petrae-Carpinetum // fühálόzás / 18.IX.1984 / leg.
Merkl O. // Longitarsus curtus det. B. Gruev (HNHM); 1
ex. (1♀!) Stirnhöcker / deutlich // Hungar (MHNB-Heik);
Rumania: 1 ex. (1♂!) Sugatag / Uhl // monticola Kut-
schera det. Z. Kaszab (HNHM); 1 ex. (1♀!) Transsylv.
Csik m. / hidegség / Hegyes-havas 10.IX.1918 / leg. Dr. J.
Fodor / Longitarsus curtus All. det. B. Gruev (HNHM); 1
ex. (1♀!) Transs. / Deubel // curtus v. monticola apterus
von Weise // Longitarsus curtus v. monticola Weise Coty-
pus (MHNB-Heik).
Dissected specimens: 70 ♂♂; 99 ♀♀.
Longitarsus curtus (Allard 1860)
Teinodactyla curta Allard, 1860, 8: 137, 143, 832 Loc.
typ.: France: Paris.
Kutschera, 1863: 294; 1864: 41-43; Allard, 1866: 135,
242-243 [Thyamis]; Bedel, 1892: 306; Portevin, 1934:
303; Redtenbacher, 1874: 507 [Longitarsus]; Weise,
1893: 977; Heikertinger, 1912: 193; Kuhnt, 1913: 885;
Heikertinger, 1930: 1326; Porta, 1934: 358; Csiki & Heik-
ertinger, 1940: 118; Mequignon, 1948: 80 [var. ciliifer as
in Löbl & Smetana, 2010: 524]; Müller, 1949-1953: 514
(Fig. 15), 524, 532; Kaszab, 1962: 311 [ab. Lichtneckerti,
unavailable name (ICZN 1999, Art. 45.6.2)]; Mohr, 1962:
88-89; Mohr, 1966: 230; Warchalowski, 1978: 68; Dog-
uet, 1984: 302-303 [denition of the Lectotype (typ. loc.:
Retz) and Paralectotype (typ. loc.: Paris)]; Bourdonné
& Doguet, 1986: 21; Gruev, 1990: 325; Döberl, 1994:
110, 137 (Figs. 314-316); Doguet, 1994: 272 (Fig. 110c),
289-292; 300 (Fig. 124g); Döberl, 1995a: 40-41; Döberl,
1995b: 53 (“Alle bisher überprüften “curtus”- Stücke er-
wiesen sich als monticola. Das Vorkommen in der Sch-
weiz muß neu bestätigt warden”); Warchalowski, 1995a:
180 (Figs. 722-723), 183 (Fig. 748), 229-231 (Fig. 936);
Warchalowski, 1995b: 107-110; Rozner, 1996: 255; Vig,
1996: 123; Warchalowski, 1996: 76 (Figs. 357-358), 79
(Fig. 383), 87-89, 118 (Fig. 571); Gruev & Döberl, 1997:
161-162; Vig, 1997: 195; Fritzlar, 1998: 203; Gruev &
Tomov, 1998: 90; Fritzlar, 1999: 75; Baselga & Novoa,
2000: 18; Bezděk & Zúber, 2001: 149; Geiser, 2001: 465;
Kapp, 2001: 63, 506; Vig, 2002: 119-120; Brelih et al.,
2003: 100; Fritzlar, 2003: 129; Warchalowski, 2003: 445;
Geiser, 2004: 19; Biondi, 2005: data on CD-Rom; Fritzlar,
2005: 232, 238-239; Gruev & Döberl, 2005: 82; Čižek,
2006: 25, 27 (Fig. 77); Fritzlar, 2009: 193.
= Thyamis liliputana Allard, 1866: 138, 180-181; Be-
del, 1889-1901: 191, 306; Heikertinger, 1930: 1326 [Lon-
gitarsus] [synonymized by Heikertinger, 1930: 1326]
= ?Thyamis moscovita Allard, 1866: 135, 247 [syno-
nymized by Heikertinger, 1930: 1326]
= Thyamis papaveris Allard, 1866: 133, 226-227; Be-
del, 1889-1901: 191, 306 [synonymized by Bedel, 1892:
306]
= Teinodactyla pratensis Allard, 1860: 137, 832 (prat-
ensis auct. nec Panzer) [synonymized by Allard, 1860:
832: “J’ai donné à tort le nom de Teinodactyla pratensis
à l’espèce que j’ai décrite à la page 137 84; il faut lui
substituer celui de Teinodactyla curta Allard”]
= Thyamis subquadrata Allard, 1866: 138, 246 [syno-
nymized by Döberl in Löbl & Smetana, 2010: 524]
= Thyamis vidua Allard, 1866: 139, 172-173; Leesberg
1882: 160, 171 [Longitarsus]; Bedel, 1892: 306 [syno-
nymized by Bedel, 1892: 306]
Distribution
L. curtus is widespread in Europe.
Gruev & Döberl (1997: 161-162; 2005: 82) report the
following regions:
Europe: Albania, Austria, Belgium, Bosnia-Herze-
govina, Bulgaria, Byelorussia, Carpathian Basin, Croatia
(included Adriatic islands), Czechia, Denmark, United
Kingdom, Isle of Man, Estonia, Finland, France (also
Corsica and East), Germany, Greece (Ionian Islands),
Hungary, Ireland, Italy (including Sicily), Latvia, Lithua-
nia, Luxembourg, Montenegro, Netherlands, Poland,
Rumania, Russia (incl. Karelia), Serbia, Slovakia, Slov-
enia, Spain, Sweden, ?Switzerland (Döberl, 1995b: 53),
Ukraine.
NOTES ON LONGITARSUS LATREILLE 1829
30
Asia: Afghanistan, Caucasus, Daghestan, Russia
(Transbaikalia: Buriatia), Tajikistan, Turkestan.
Materials studied
Portugal: 3 exx. (2♀♀! 1♂!) Portugal / Beira Altas
Manteigas / Gletschertal / Zêzere 1300 m / N40°19’58’’
W7°34’14’’/ Betula-Hangwald / 26.IV.2014 / leg. C. Ger-
mann // Longitarsus curtus Allard det. L. Farina, 2018
(CGCB) (Figs. 39, 43); 1 ex. (1♀!) Portugal / Beira Alta
Manteigas / Gletschertal / Zêzere 1300 m / N40°19’58’’
W7°34’14’’ / Betula-Hangwald / 26.IV.2014 leg. C. Ger-
mann (LFCC);
Spain: 3 exx. (2 ♂♂!) Spagna Pirenei / sopra Salardù
1700-1900 m / 10.VIII.1986 / leg. M. Biondi (MBCA);
2 exx. (2♂♂!) Spagna (Avila) 1400 m / Navacepeda
de Tormes / 4.V.1989 /M. Biondi P. Audisio (MBCA);
1 ex. (1♀!) Spagna (Salamanca) / Candelario 1200 m /
10.V.1989 / M. Biondi P. Audisio (MBCA); 1 ex. (1♂!)
España-Castilla / Navacepeda dint. / 26.VI.1991 / C.
Giusto leg. (SZCM);
France: 1 ex. (1♀!) Gallia / coll. Lichtneckert // Lon-
gitarsus curtus All. det. B. Gruev (HNHM); 1 ex. (1♀!)
Gallia / Pyr. Or. / Le Vernet / 6.VIII.1904 / A. Dodero
(MSNG-Dod); 1 ex. (1♀!) Vichy (Gallia) / 9.VII.1934 /
Tasso / Schatzm. (MSNM); 1 ex. (1♂!) Francia06 / 1353
m 9.VII.2010 / Le Bourguet 3 km S (pont de la Serra)
/ 44°11.875N 006°57.167E / Leonardi (MSNM); 1 ex.
(1♂!) Riom // Longitarsus curtus det. M. Bergeal 1994
(MSNM-Ber); 12 exx. (6♀♀!) F-03 / Chemilly 26.V.1987
/ M. Bergeal // Longitarsus curtus det. M. Bergeal 1994
(MSNM-Ber); 1 ex. (1♂!) Meyrueis / 19.VIII.1987 /
M. Bergeal (MSNM-Ber); 4 exx. (2♀♀! 2♂♂!) F-48 /
Meyrueis / 20.VIII.1988 / M. Bergeal // Longitarsus cur-
tus det. M. Bergeal, 1994 (MSNM-Ber);
Sweden: 1 ex. (1♂!) S. Sk. Hovdala / 26.XI.1983 / G.
Gillerfors (MSNM) (specimen with an anomalous dark
brown coloring on elytrae); 2 exx. (2♂♂!) Gotland Sun-
dre / 25.VIII.1961 / E. Wirén // Longitarsus curtus det. M.
Bergeal, 1996 (MSNM-Ber);
Finland: 1 ex. (1♀!) Fennia / Kb. Hammaslaakari / P.
Kontkanen // 22.VIII.1928 // curtus det. F. Heikertinger
(MHNB-Heik);
Germany: 1 ex. (1♀!) Bayern / Stadt / curtus / Fld.
Stark / Weitlänf. Pkt. (MHNB-Heik);
Switzerland: 2 exx. (1♂! 1♀!) Zermatt / Wallis (MH-
NB);
Italy: Trentino Alto Adige: 1 ex. (1♀!) Tirol / Seis. //
curtus det. Heikertinger // coll. A. Simonis dupleta coll.
Winkler (MSNG); 2 exx. (2♀♀!) Trentino (TN) Levico /
Ag.-Ott. 1884 / C. Doria (MSNG); 1 ex. Alto Adige (BZ)
/ Pusteria Brunico / 26.VII.1934 / G. Binaghi (MSNG); 1
ex. (1♀!) Alto Adige (BZ) Renon mt. 1100 / 14.IX.1968 /
Mercati (MSNM);
Friuli Venezia Giulia: 1 ex. (1♂!) Jul. Alp. / Nevea
Cregnedul / 1800 m 18.IX.1949 / Springer (MSNM) (Fig.
38); 1 ex. (1♂!) Jul. Alp. / Nevea Cregnedul / 7.X.1949 /
1700 m Springer (MSNM); 1 ex. (1♂!) Jul. Alp. / Nevea
Cregnedul Springer // 1900 m 7.X.1949 (MSNM); 1 ex.
(1♂!) Jul. Alp. / Nevea Cregnedul / VII.1950 / Springer
(MSNM); 1 ex. (1♀!) Jul. Alp. / Nevea Cregnedul / VI-
II.1950 Springer (MSNM); 5 exx. (2♂♂! 3♀♀!) Friuli
Venezia Giulia (UD) Jul. Alp. / Nevea / Cregnedul /
10.IX.1950 / Springer (MSNM) (Fig. 42); 1 ex. (1♀!)
Jul. Alp. / Nevea Cregnedul / 17.IX.1950 / Springer
(MSNM);
Liguria: 1 ex. (1♀!) Alpi Liguri / Viozene 1745 m /
IX.1942 // ex. collez. A. Porta (MSNM);
Poland: 1 ex. (1♂!) Krakau / Natterer // 6.III.1907 (MH-
NB-Heik); 1 ex. (1♀!) Krakau / Natterer // 23.VII.1907
(MHNB-Heik); 1 ex. (1♀!) Polonia / Lukawica / Prov. Lis
Ro. / 6.VIII.1926 (MSNM);
Czechia: 1 ex. (1♀!) Erzgebirge Bohemia // ex. coll.
Dr. I. Peregi (HNHM); 1 ex. (1♂!) Friedberg / Bohem.
Wald / Skalitzky (MHNB-Heik); 1 ex. (1♀!) Moravia /
Seitendorf / Zanchtel / 21.VIII.1915 (MHNB-Heik); 1
ex. (1♀!) Sobèšice / rexocanb / Czechoslov. (ZSM); 1
ex. Slov. 4.IX.1948 / L’adová u Dobš / leg. Josef Král //
Czechoslov. // Longitarsus curtus det. B. Gruev (ZSM);
Austria: 1 ex. (1♀!) St. Radegund / Dr. Beszédes
/ Longitarsus curtus det. B. Gruev // coll. Dr. J. Fodor
(HNHM); 2 exx. (1♀!) Boehm-Wald // ex Orig. Samig.
J. Breit Wien // Longitarsus curtus det. C. Leonardi (MH-
NB); 1 ex. (1♀!) Puntigam-Au / Styr 19.IX.1932 / curtus
det. P. Heymes (MHNB); 1 ex. (1♀!) Bohemia Barzdorf /
coll. Dr. Tölg (MHNB-Heik); 1 ex. (1♀!) Bohemia / Ele-
onorenh / Natterer (MHNB-Heik); 2 exx. (1♀!) Bohemia
/ Barzdorf / August / Coll. Dr. Tölg // curtus / Halschild
/ dünkel (MHNB-Heik); 1 ex. (1♀!) Bohemia / Barzdorf
/ Coll. Dr. Tölg // curtus / Halschild / dünkel (MHNB-
Heik); 1 ex. Wien / Umg A. Winkler (MHNB-Heik); 1
ex. (1♂!) Carinthia / Weise (MHNB-Heik); 1 ex. (1♂!)
Stiria bor. / Dr. Krauss / curtus Stirnhöcka (MHNB-Heik);
1 ex. Otto 1893 / Ferlach (MHNB-Heik); 1 ex. Bisamberg
A. i. / 14.IX.1907 / Heikertinger (MHNB-Heik) (Fig. 8);
1 ex. A. Baden / Natterer // 15.IX.1907 (MHNB-Heik);
2 exx. (2♂♂!) Taffatal bei Horn A.i. 1908 / Heikerting-
er (MHNB-Heik); 3 exx. (2♂♂!) Taffatal b. Horn A.i. /
1.IX.1908 / Heikertinger // Longitarsus curtus det. Heik-
ertinger (MHNB-Heik); 1 ex. (1♂!) Föhrenwald / b. Egg-
enburg / 6.IX.1908 / Heikertinger (MHNB-Heik); 1 ex.
(1♀!) Wien erstein / 20.IX.1908 / Heikertinger // curtus
det. F. Heikertinger (MHNB-Heik); 1 ex. (1♀!) Franken-
fels / VII.1913 / Longitarsus curtus det. Heikertinger //
coll. A. Simonis dupleta coll. Winkler (MSNG); 2 exx.
(1♂!) Territorio del Dobratsch / A. Sazmaier (MSNM); 1
ex. (1♂!) St. Johann / Herb. / Bezirk. Hartberg / Tierpark
400 / 20.VI.1995 / leg. E. Holzer (MSNM-Ber); 1 ex.
(1♂!) STMK. Arzberg / Bez. Weiz Grösser / Raabklamm
500 / 28.IX.1997 / leg. E. Holzer (MSNM-Ber);
Slovenia: 1 ex. Paskau / Reitter // coll. Reitter // Longi-
tarsus curtus All. det. A. Warchalowski, 1960 (HNHM); 1
ex. (1♂!) Laibach / 17.IX.1931 / Dr. Staudacher (MSNM);
1 ex. (1♂!) Julisch. Alp. / Raibl VIII.1948 / leg. Springer
// Longitarsus curtus det. C. Leonardi (ZSM);
Hungary: 1 ex. (1♂!) Siόfok / Lichtneckert // mon-
ticola Kutschera det. Z. Kaszab (HNHM); 1 ex. (1♂!)
Hungaria / Marostordam / Ehik & Loksa // Gödemester-
háza 750 m / 1-15.VIII.1943 // monticola Kutschera det.
Z. Kaszab (HNHM); 1 ex. (1♂!) Vértes h. Hu. / Fáni
völgy / rostálva 11.IV.1961 / Endrődy / Hungary (ZSM);
1 ex. (1♂!) Hu. Vértes h. / Fáni völgy // 2-4.VIII.1961
/ Hámoriné (ZSM); 1 ex. Hung. Bukki N.P. / Szilvás-
LAURA FARINA
31
váradi / Rakmava 600 m / Sambucetum ebuli / egyelés /
12.VII.1983 / leg. Ádám (ZSM); 1 ex. Hung. Bükki N.P.
/ Felsótárkány Tav. -kó 900 m // Tilio-Fraxinetum // tala-
jcsapda / 4.X.1983 / leg. Ádám (ZSM); 1 ex. (1♂!) Hung.
Bükki N.P. / Bükkszemtkeveszt / Pénzpatak-tetö 600 m //
ablakcsapda / 20.IX.1984 leg. Merkl O. // Querco petrae-
Carpinetum (ZSM);
Croazia: 1 ex. (1♀!) Pakrac / Apfelbeck / monticola
Kutschera det. Z. Kaszab (HNHM);
Montenegro: 2 exx. Krivosije / Paganetti // Longitar-
sus luridus Scop. det. A. Warchalowski, 1960 (HNHM);
1 ex. (1♂!) Cattaro / Dalmat. // Cattaro / VI-VII.1906
(MHNB-Heik) (Fig. 37);
Albania: 1 ex. (1♀!) Albania / Ipek Akad Balk. Exp. /
Csiki 1917 (HNHM); 1 ex. (1♀!) Albania / Ipek / VI.1927
/ Akad Balk. Exp. (HNHM);
Greece: 1 ex. (1♀!) Grecia nom. Trikala / M.ti Vout-
sikaki dint. / Moskhoton tra M.te Khatzi e M. te Avgo m.
1000-1400 / 23.V.1982 M. Biondi (MBCA);
Russia: 2 exx. (1♂! 1♀!) Russia Belgorod /
15.VII.1957 / leg. L.M. Medvedev (MSNM-Ber); 1 ex.
(1♂!) Ud.S.S.R. / Streleckaja / step. 1.IX.1970 / leg.
B. Gruev // Longitarsus curtus det. B. Gruev, 1972
(ZSM);
Asia: 19 exx. (1♂! 3♀♀!) Armenia / Gegamskij / Mts
Hosrov res. / 1-16.VI.1987 / V. Karasjov (MSNM).
Dissected specimens: 44 ♂♂; 50 ♀♀.
Acknowledgments
I am very grateful to the following friends and col-
leagues: Carlo Leonardi (Museo Civico di Storia Natu-
rale di Milano), for his support, suggestions and criticism
during the draft of the text and for his precious check of
my drawings; Gianfranco Liberti (Uboldo) for his im-
portant critical review of the manuscript, his suggestions
and help in improving my English text; David Furth (De-
partment of Entomology, Smithsonian Institution, Wash-
ington, DC), for his critical review of my manuscript,
his interesting suggestions, the sending of L. personatus
specimen photographs from Israel and his careful check
of the English text; Stefano Zoia (Milano) who, with his
usual competence and availability, made the photographs;
Lutz Behne (Senckenberg Deutsches Entomologisches
Institut, Müncheberg), for his kind permission to publish
the photograph of the paralectotypes of L. aphthonoides;
Maurizio Biondi (Università degli Studi dell’Aquila), for
his suggestions and for the loan of very interesting speci-
mens; Claudio Pagliarin (Museo di Storia Naturale di Mi-
lano) who, with his patience and graphic skills, helped me
to improve the drawings and the distribution maps; Maria
Chiara Sibille (Ente di gestione delle aree protette del Ti-
cino e del Lago Maggiore) for her assistance in the eld;
Rita Mascolo and Enrico Muzio (Museo di Storia Natu-
rale di Milano) for their help in digging out overlooked
references hidden on the library shelves.
I also wish to thank the following people for the loan
of specimens and for their friendly, professional help:
Martin Albrecht (Schüpfen), Michael Balke (Zoologische
Staatssammlung, München), Ditta Amran Balke (Zo-
ologische Staatssammlung, München), Maxwell Barclay
(Natural History Museum, London), Mauro Daccordi
(Museo Civico di Storia Naturale, Verona), Frank Fritzlar
(Jena), Michael Geiser (Natural History Museum, Lon-
don), Christoph Germann (Naturistorische Museum, Ba-
sel), Pascale Hutter (Basel), Bernd Jaeger (Museum für
Naturkunde, Berlin), Leonardo Latella (Museo Civico di
Storia Naturale, Verona), Otto Merkl (Hungarian Natu-
ral History Museum, Budapest), Fernando Montemurro
(Taranto), Roberto Poggi (Museo Civico di Storia Natu-
rale “Giacomo Doria”, Genova), Fabrizio Rigato (Museo
di Storia Naturale di Milano).
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d’Europe et des bordes de la Mer Méditerranée. An-
nales de la Société Entomologique de France, Paris,
sér. 3, 8: 39-144, 369-418, 539-578, 785-834.
Allard, E., 1866 – Monographie des Alticides. L’Abeille,
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The paper shows how faunistic research was initiated and supported by Manfred Döberl. For the leaf beetle fauna of Slovenia new records are listed and interpreted. Records concern 116 leaf beetle species from region Gorenjsko und from region Štajersko. Observations on habitats and host plants are added for the species Cryptocephalus albolineatus, Cryptocephalus scapularis, Sclerophaedon carniolicus, Sclerophaedon orbicularis, Aphthona juliana, Longitarsus rectilineatus, Longitarsus fulgens, Neocrepidodera cyanescens, Chaetocnema semicoerulea, Argopus ahrensi and Cassida alpina. Cryptocephalus saliceti, Cryptocephalus transiens, Cryptocephalus transcaucasicus, Chrysochus asclepiadeus and Oreina liturata are listed as species of special interest. Additionally listed are records from flea beetles determined by Manfred Döberl (Chaetocnema compressa, Chaetocnema subcoerulea, Longitarsus languidus, Longitarsus longiseta and Longitarsus lateripunctatus personatus) as well as records on flea beetles collected by B. Kofler, Škovja Loca (among them Aphthona stussineri, Longitarsus monticola and Longitarsus pinguis). Kurzzusammenfassung: Die Arbeit ist ein Beispiel für faunistische Forschung an Blattkäfern, die von Manfred Döberl angeregt und unterstützt worden ist. Zur Blattkäferfauna Sloweniens werden Funde faunistisch interessanter Arten vorgestellt. Von 116 Blattkäfer-Arten der Julischen Alpen (Region Oberkrain, Gorenjsko) und des Bacher-Gebirges (Pohorie) (Region Steiermark, Štajersko) werden Funddaten mitgeteilt. Für Cryptocephalus albolineatus, Cryptocephalus scapularis, Cryptocephalus saliceti, Cryptocephalus transiens, Cryptocephalus transcaucasicus, Chrysochus asclepiadeus, Oreina liturata, Sclerophaedon carniolicus, Sclerophaedon orbicularis, Aphthona juliana, Longitarsus rectilineatus, Longitarsus fulgens, Neocrepidodera cyanescens, Chaetocnema semicoerulea, Argopus ahrensi und Cassida alpina werden Lebensräume und Wirtspflanzen beschrieben. Cryptocephalus saliceti, Cryptocephalus transiens, Cryptocephalus transcaucasicus, Chrysochus asclepiadeus und Oreina liturata sind weitere faunistisch interessante Arten. Ergänzend werden von Manfred Döberl bestimmte Funde (Chaetocnema compressa, Chaetocnema subcoerulea, Longitarsus languidus, Longitarsus longiseta und Longitarsus lateripunctatus personatus) und bemerkenswerte Nachweise von Flohkäfern (leg. B. Kofler, Škovja Loca) veröffentlicht (darunter Aphthona stussineri, Longitarsus monticola und Longitarsus pinguis).
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