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Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera



Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera (Aarvik et al. 2017) are presented. 44 species new to the Nordic-Baltic area as well as numerous new country records are added. Two misspellings and 14 other mistakes of various types are corrected. The genus name Tuta Kieffer & Jørgensen, 1910 is recombined with the species name absoluta Meyrick, 1917 as Tuta absoluta (Meyrick, 1917) comb. rev.
Additions and corrections to the Nordic-Baltic Checklist of
Aarvik, L., Bengtsson, B.Å., Elven, H., Ivinskis, P., Jürivete, U., Karsholt, O., Mutanen, M. &
Savenkov, N. 2021. Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera.
Norwegian Journal of Entomology 68, 1–14.
Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera (Aarvik et al. 2017) are
presented. 44 species new to the Nordic-Baltic area as well as numerous new country records are
added. Two misspellings and 14 other mistakes of various types are corrected. The genus name Tuta
Kieer & Jørgensen, 1910 is recombined with the species name absoluta Meyrick, 1917 as Tuta
absoluta (Meyrick, 1917) comb. rev.
Key words: Lepidoptera, checklist, Nordic-Baltic, additions, corrections.
Leif Aarvik, Natural History Museum, University of Oslo, P.O. Box 1172 Blindern, NO-0318 Oslo,
Norway. E-mail:
Bengt Å. Bengtsson, Lokegatan 3, S-386 93 Färjestaden, Sweden.
Hallvard Elven, Natural History Museum, University of Oslo, P.O. Box 1172 Blindern, NO-0318
Oslo, Norway. E-mail:
Povilas Ivinskis, St. Nature Research Centre, Akademijos 2, LT-08412 Vilnius-21, Lithuania.
Urmas Jürivete, Estonian Lepidopterologists’ Society, Moora umb. 8, EE-11625 Tallinn, Estonia.
Ole Karsholt, Zoologisk Museum, Natural History Museum of Denmark, University of Copenhagen,
Universitetsparken 15, DK-2100 Copenhagen Ø, Denmark. E-mail:
Marko Mutanen, Department of Genetics and Physiology, P.O. Box 3000, FI-90014 University of
Oulu, Finland. E-mail: marko.mutanen@oulu.
Nikolay Savenkov, Latvian Museum of Natural History, K. Barona st. 4, LV-1050 Riga, Latvia.
© Norwegian Journal of Entomology. 18 June 2021
The authors present faunistic and taxonomic
updates to the Nordic-Baltic Checklist of
Lepidoptera (Aarvik et al. 2017). In the past three
years, numerous publications on Lepidoptera have
appeared which have added to the knowledge
of the fauna present in the area. The studies on
phylogeny using molecular methods are continuing
and recently this has aected particularly the
classication of Gelechioidea (Wang & Li
2020) and Geometridae (Murillo-Ramos et al.
2019, Müller et al. 2019, Sihvonen et al. 2020).
New monographs covering Europe or the whole
Palaearctic region give an extended perspective at
the genus level and inevitably lead to changes. An
example is the monograph on the noctuid genus
Agrochola Hübner, 1821 and allies (Ronkay et
al. 2017). At the national level numerous new
records have been made. This reects the interest
in Lepidoptera in each of the countries, but also to
some extent a warmer climate.
It has recently been suggested (Wiemers
et al. 2018) that Linnaeus’ Fauna Svecica was
published in 1760, not 1761. However, it seems
that Linnaeus in 1760 was only circulating a proof
version of Linnaeus (1761) and not a publication.
The suggested change would aect the year of
description for a number of species. Until more
evidence eventually appears, we consider the year
of publication to be 1761.
Material and methods
New country records are based on publications
referred to in each case. In cases where there is
no reference to a publication, the information
was gathered by the authors from the dierent
countries. A new version of the checklist in the
form of an excel sheet is available on the website
of the Norwegian Entomological Society: http://
The new country records are the responsibility
of the authors representing their own country.
New species for the list is the responsibility of the
contributor from the country where the species
was discovered, but has also been accepted
by the rest of the authors. Some of the records
have been published in entomological journals,
some have been presented on the internet and
others are presented for the rst time here. In
case of unpublished records, they are given
with the permission of the discoverer/collector.
New species for Finland are recorded on the
website «Laji» which is provided by The Finnish
Biodiversity Information Facility (FinBIF):
https://laji./en. The word ‘omitted’ is used in
cases when a record was omitted by mistake in
the checklist (Aarvik et al. 2017). The number
before the name of the taxon is the same as the
one used in the checklist. A new taxon takes
the number of the taxon preceding it, adding a
letter after the number, as in the example: 4178a.
Spodoptera cilium Guenée, 1852 D (Hviid 2018).
This means that Spodoptera cilium is a species
new to Denmark as well as the whole region, and
that it should be inserted after Spodoptera exigua,
which has number 4178.
New country records
0020. Eriocrania salopiella (Stainton, 1854) Lt
(Ivinskis et al. 2018).
0047. Stigmella freyella (Heyden, 1858) S
(Bengtsson 2020).
0051. Stigmella sakhalinella Puplesis, 1984 F.
0129. Trifurcula immundella (Zeller, 1839) N
(Aarvik et al. 2021), Lt (Ivinskis et al. 2018).
0136. Etainia louisella (Sircom, 1849) S
(Bengtsson 2019).
0185. Incurvaria masculella (Denis & Schier-
müller, 1775) Lt (Ivinskis & Rimšaite 2017).
0216. Nemophora minimella (Denis & Schier-
müller, 1775) F, E.
0220. Adela reaumurella (Linnaeus, 1758) F.
0245. Coptotriche marginea (Haworth, 1828) N
(Aarvik et al. 2019).
0308. Apterona helicoidella (Vallot, 1827) D
(Buhl et al. 2018).
0368. Trichophaga scandinaviella Zagulajev,
1960 E.
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
0370. Elatobia fuliginosella (Lienig & Zeller,
1846) Lt (Ivinskis & Rimšaite 2017).
0382. Tinea svenssoni Opheim, 1966 Lt.
0473. Caloptilia della (Reutti, 1853) F, Lv, Lt
(Ivinskis et al. 2020).
0536. Phyllonorycter emberizaepenella (Bouché,
1834) I (Ólafsson 2018).
0556. Phyllonorycter viminetorum (Stainton,
1854) E.
0557. Phyllonorycter connexella (Zeller, 1846) N
(Aarvik et al. 2019).
0595. Euhyponomeutoides ribesiella (Joannis,
1900) Lt (Ivinskis et al. 2020).
0604. Swammerdamia passerella (Zetterstedt,
1839) Lt (Ivinskis et al. 2018).
0626. Argyresthia trifasciata (Staudinger, 1871)
0673. Digitivalva reticulella (Hübner, 1796) Lt
(Paulavičiūtė et al. 2017).
0691. Ypsolopha mucronella (Scopoli, 1763) F.
0734. Leucoptera laburnella (Stainton, 1851) F.
0737. Leucoptera lathyrifoliella (Stainton, 1866)
D (omitted).
0768. Choreutis diana (Hübner, 1822) Lt (Ivinskis
et al. 2019).
0797. Sparganothis pilleriana (Denis & Schier-
müller, 1775) Lv.
0973. Eupoecilia sanguisorbana (Herrich-
Schäer, 1856) E.
1001. Cochylis roseana (Haworth, 1811) S
(Bengtsson 2019).
1006. Cochylichroa atricapitana (Stephens, 1852)
(= Cochylis atricapitana) Lt (Ivinskis &
Rimšaite 2017).
1030. Apotomis lineana (Denis & Schiermüller,
1775) Lt (Ivinskis et al. 2018).
1062. Celypha woodiana (Barrett, 1882) Lt
(Ivinskis et al. 2020, Paulavičiūtė et al. 2020).
1115. Endothenia gentianaeana (Hübner, 1799)
Lt (Ivinskis et al. 2020).
1142. Ancylis comptana (Frölich, 1828) D (P.
Falck pers.comm.).
1193. Epinotia tedella (Clerck, 1759) I (E.
Ólafsson in litt.).
1195. Epinotia pusillana (Peyerimho, 1863) S
(Bengtsson 2019).
1212. Crocidosema plebejana Zeller, 1847 I
1257. Epiblema junctana (Herrich-Schäer,
1856) Lv.
1270. Notocelia rosaecolana (Doubleday, 1850) I
(E. Ólafsson in litt.).
1277. Retinia perangustana (Snellen, 1883) Lt
(Paulavičiūtė et al. 2020).
1323. Cydia conicolana (Heylaerts, 1874) F.
1330. Cydia grunertiana (Ratzeburg, 1868) Lv.
1336. Cydia leguminana (Lienig & Zeller, 1846)
N (Gustad & Aarvik in prep.)
1341. Cydia amplana (Hübner, 1799) Lv.
1354. Grapholita gemmiferana Treitschke, 1835
Lt (Ivinskis et al. 2019).
1358. Grapholita lobarzewskii (Nowicki, 1860)
1370. Pammene albuginana (Guenée, 1845) Lt
(Ivinskis & Rimšaite 2017).
1381. Pammene trauniana (Denis & Schier-
müller, 1775) S (Bengtsson 2018).
1383. Pammene aurita Razowski, 1992 Lv.
1443. Synanthedon aviventris (Staudinger, 1883)
N (Aarvik et al. 2021).
1445. Synanthedon myopaeformis (Borkhausen,
1789) F.
1476. Oegoconia deauratella (Herrich-Schäer,
1854) Lt (Ivinskis & Rimšaite 2017).
Norwegian Journal of Entomology 68, 1
14 (2021)
1486. Denisia albimaculea (Haworth, 1828) N
(Aarvik et al. 2021).
1488. Denisia luticiliella (Erscho, 1877) D
(Buhl et al. 2018).
1512. Batia internella Jäckh, 1972 F.
1520. Oecophora bractella (Linnaeus, 1758) Lv.
1649. Pancalia nodosella (Bruand, 1851) Lt
(Ivinskis & Rimšaite 2017).
1678. Aproaerema larseniella Gozmány, 1957 Lt
(Paulavičiūtė 2019).
1700. Nothris gregerseni Karsholt & Šumpich,
2015 Lt (Ivinskis & Rimšaite 2017).
1719. Dichomeris latipennella (Rebel, 1937) D
(Buhl et al. 2020).
1733. Helcystogramma albinervis (Gerasimov,
1929) F.
1803. Monochroa divisella (Douglas, 1850) E.
1866. Gelechia scotinella Herrich-Schäer, 1854
N (Aarvik et al. 2019).
1902. Scrobipalpa bryophiloides Povolný, 1966
E, D (Buhl et al. 2020).
1905. Scrobipalpa ocellatella (Boyd, 1858) S
(Bengtsson 2019).
1908. Scrobipalpa salicorniae (E. Hering, 1889)
N (Aarvik et al. 2019).
1913. Scrobipalpula tussilaginis (Stainton, 1867)
1924. Caryocolum alsinella (Zeller, 1868) N
(Aarvik et al. 2019), Lt (Ivinskis & Rimšaite
1936. Caryocolum blandelloides Karsholt, 1981
1937. Caryocolum proxima (Haworth, 1828) Lt
(Paulavičiūtė et al. 2017).
1938. Caryocolum blandulella (Tutt, 1887) Lv.
1957. Carpatolechia fugacella (Zeller, 1839) S.
2006. Elachista dispilella Zeller, 1839 Lt (Sruoga
et al. 2019).
2011. Elachista dispunctella (Duponchel, 1843)
Lt (Sruoga et al. 2019).
2041. Elachista poae Stainton, 1855 N.
2042. Elachista atricomella Stainton, 1849 F.
2052. Eachista deriventa Kaila & Mutanen, 2008
2058. Elachista bifasciella Treitschke, 1833 N
(Aarvik et al. 2019).
2081. Elachista consortella Stainton, 1851 Lv.
2091. Blastodacna hellerella (Duponchel, 1838)
2132. Coleophora amellivora Baldizzone, 1979
Lt (Ivinskis et al. 2019).
2138. Coleophora asteris Mühlig, 1864 F.
2161. Coleophora jaernaensis Björklund &
Palmqvist, 2002 E.
2188. Coleophora chalcogrammella Zeller, 1839
2207. Coleophora vulnerariae Zeller, 1839 F.
2308. Hypatopa segnella (Zeller, 1873) Lt
(Ivinskis et al. 2018).
2357. Epermenia aequidentellus (E. Hofmann,
1867) D (Buhl et al. 2018).
2390. Stenoptilia veronicae Karvonen, 1932
(eborinodactyla auct.) D (Buhl et al. 2017,
2392. S. mariaeluisae Bigot & Picard, 2002
(inopinata auct.) I (E. Ólafsson in litt.).
2419. Oxyptilus tristis (Zeller, 1841) N (Aarvik et
al. 2019).
2427. Hellinsia inulae (Zeller, 1852) S (Bengtsson
2556. Araschnia levana (Linnaeus, 1758) N
(Aarvik et al. 2019)
2589. Apatura iris (Linnaeus, 1758) N (Aarvik et
al. 2021).
2594. Lasiommata megera (Linnaeus, 1767) E.
2664. Aricia agestis (Denis & Schiermüller,
1775) Lv.
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
2729. Sciota rhenella (Zincken, 1818) S
(Bengtsson 2020).
2769. Delplanqueia inscriptella (Duponchel,
1836) F, Lt (Ivinskis et al. 2018).
2775. Nephopterix angustella (Hübner, 1796) E.
2804. Eccopisa eractella Zeller, 1848 E.
2809. Euzophera bigella (Zeller, 1848) F.
2816. Ancylosis oblitella (Zeller, 1848) N (Aarvik
et al. 2019), F, E.
2824. Phycitodes lacteella (Rothschild, 1915) Lv,
Lt (Ivinskis et al. 2018).
2838. Cadra cautella (Walker, 1863) Lt (omitted).
2853. Endotricha ammealis (Denis & Schier-
müller, 1775) E.
2885. Sclerocona acutella (Eversmann, 1842) Lv.
2923. Spoladea recurvalis (Fabricius, 1775) N
(Aarvik et al. 2019), S (Bengtsson 2020), F.
2941. Cydalima perspectalis (Walker, 1859) F, E,
Lt (Paulavičiūtė & Mikalauskas 2018).
2950. Cynaeda pustulalis (Hübner, 1823) D (Buhl
et al. 2020).
3020. Pediasia luteella (Denis & Schiermüller,
1775) S (Bengtsson 2017).
3028. Agriphila latistria (Haworth, 1811) F.
3029. Agriphila aeneociliella (Eversmann, 1844)
3033. Agriphila biarmicus (Tengström, 1865) Lt
(Ivinskis et al. 2018).
3121. Idaea ochrata (Scopoli, 1763) Lv, Lt
(Švitra et al. 2017)
3124. Idaea fuscovenosa (Goeze, 1781) E.
3144. Scopula virgulata (Denis & Schiermüller,
1775) D (omitted).
3307. Pareulype berberata (Denis & Schier-
müller, 1775) E.
3361. Eupithecia abbreviata Stephens, 1831 Lt
(Švitra et al. 2018).
3387. Eupithecia insigniata (Hübner, 1790) F.
3402a. Eupithecia addictata Dietze, 1908 Lt
(Ūsaitis et al. 2019).
3458. Isturgia arenacearia (Denis & Schier-
müller, 1775) S (Palmqvist & Ryrholm 2020),
D (Larsen et al. 2020), E, Lv.
3473. Apocheima hispidaria (Denis & Schier-
müller, 1775) N (Aarvik et al. 2021).
3493. Peribatodes rhomboidaria (Denis &
Schiermüller, 1775) Lv, Lt (Švitra et al.
3548. Ennomos quercinaria (Hufnagel, 1767) F.
3623. Chlorissa cloraria (Hübner, 1813) F.
3636. Eversmannia exornata (Eversmann, 1837)
Lt (Švitra et al. 2018).
3714. Hyles euphorbiae (Linnaeus, 1758) N.
3745. Drymonia obliterata (Esper, 1785) Lt
(Kaupys 2019).
3797. Hypena lividalis (Hübner, 1790) E.
3819. Laelia coenosa (Hübner, 1808) F.
3863. Utetheisa pulchella (Linnaeus, 1758) F.
3955. Eublemma parva (Hübner, 1808) N (Aarvik
et al. 2021).
3957. Eublemma purpurina (Denis & Schier-
müller, 1775) N (Aarvik et al. 2021), S
(Palmqvist & Ryrholm 2020), D (Larsen et al.
3977. Catocala elocata (Esper, 1787) E, Lv.
4001. Chrysodeixis chalcites (Esper, 1789) F, Lv.
4067. Cucullia verbasci (Linnaeus, 1758) F, E.
4166. Bryopsis muralis (Forster, 1771)
(= Nyctobrya muralis) N, F.
4185. Caradrina kadenii Freyer, 1836 S (Palmqvist
& Ryrholm 2020), D (Larsen et al. 2019).
4187. Caradrina albina Eversmann, 1848 S
(Palmqvist 2018).
4296. Apamea remissa (Hübner, 1809) I (omitted).
4299. Apamea crenata (Hufnagel, 1766) I
4321. Resapamea vulpecula (Eversmann, 1852)
(= R. hedeni) Lv.
4328. Xylomoia graminea (Graeser, 1889) E, Lv.
4373. Conistra rubiginosa (Scopoli, 1763) E, Lv.
4384. Lithophane ornitopus (Hufnagel, 1766) N
Norwegian Journal of Entomology 68, 1
14 (2021)
(Aarvik et al. 2019).
4409. Atethmia centrago (Hübner, 1809) N
(Aarvik et al. 2021). Lv.
4423. Aporophyla lueneburgensis (Freyer, 1848)
4429. Polymixis lichenea (Hübner, 1813) N
(Aarvik et al. 2019).
4476. Polia lamuta (Herz, 1903) N (Saarenmaa
4486. Lacanobia amurensis (Staudinger, 1901)
4496. Hyssia cavernosa (Eversmann, 1842) Lv.
4507. Conisania leineri (Freyer, 1836) E.
4509. Hecatera dysodea (Denis & Schiermüller,
1775) F, E.
4536. Mythimna l-album (Linnaeus, 1767) N
(Aarvik et al. 2021), E.
4540. Leucania loreyi (Duponchel, 1827) N, S
(Palmqvist & Ryrholm 2020).
4571. Euxoa vitta (Esper, 1789) E.
4581. Agrotis bigramma (Esper, 1790) Lv.
4591. Agrotis puta (Hübner, 1803) F.
4602. Diarsia orida (F. Schmidt, 1859) F.
4626. Noctua comes Hübner, 1813 I (E. Ólafsson
in litt.).
4655. Xestia distensa (Eversmann, 1851) N
(Aarvik et al. 2019).
4697. Nola cristatula (Hübner, 1793) Lv.
New taxa to the Nordic-Baltic area
0312a. Dryadaula heindeli Gaedike & Scholz,
1998 N (Gustad & Aarvik 2017).
0324a. Triaxomera baldensis G. Petersen, 1983
Lt (Ivinskis & Rimšaite 2017).
0345a. Neurothaumasia Le Marchand, 1934.
0345b ankerella (Mann, 1867) D (P. Falck pers.
0391a. Monopis jussii Kaila, Mutanen, Huemer,
Karsholt & Autto, 2020 N, S, F (Mutanen et
al. 2020).
0439a. Micrurapteryx kollariella (Zeller, 1839) D
(Buhl et al. 2019).
0441a. Parectopa robiniella Clemens, 1863 Lv,
Lt (Ivinskis et al. 2019).
0462a. Calybites quadrisignella (Zeller, 1839)
0474a. Caloptilia honoratella (Rebel, 1914) D
(Buhl et al. 2019).
0766a. Tebenna micalis (Mann, 1857) N (Aarvik
et al. 2021). In 2014 imported to Denmark as
larvae on Helichrysum italica in several
places; a single specimen was found in a light
trap. Since it has not been recorded, and the
species is considered introduced to Denmark.
0834a. Clepsis peritana (Clemens, 1860) D (Buhl
et al. in press).
0985a. Aethes bilbaensis (Rössler, 1877) D (Buhl
et al. 2020).
1222a. Eucosma albidulana (Herrich-Schäer,
1851) S (Ohlsson 2020).
1360a. Grapholita inopinata Heinrich, 1928 F
1454a. Chamaesphecia Spuler, 1910
1454b. nigrifrons (Le Cerf, 1911) Lt (Švitra et al.
1518a. Esperia Hübner, 1825
1518b. sulphurella (Fabricius, 1775) D (Buhl et
al. 2018).
1668a. Ascalenia Wocke, 1876
1668b. viviparella Kasy 1969 Lt (Ivinskis et al.
1704a Neofaculta taigana Ponomarenko, 1998 N
(Aarvik et al. 2021), S, F.
1914a. Tuta Kieer & Jørgensen, 1910
1914b. absoluta (Meyrick, 1917) N (Aarvik et al.
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
2021), S, F, D (Buhl et al. in press). Placed on
the list due to numerous recent records in
1955a. Carpatolechia aenigma (Sattler, 1983) Lt
(Paulavičiūtė 2017).
2212a. Coleophora variicornis Toll, 1952 Lv.
2237a. Coleophora sirella Tabell & Mutanen,
2019 F (Tabell et al. 2019).
2332a. Scythris penicillata (Chrétien, 1900) E
2737a. Gymnancyla hornigii (Lederer, 1852) D
(Buhl et al. 2018).
2865a. Paracorsia Marion, 1959
2865b. repandalis (Denis & Schiermüller, 1775)
2906a. Udea costalis (Eversmann, 1852) Lv, Lt
2939a. Hodebertia Leraut, 2003
2939b. testalis (Fabricius, 1794) D (Buhl et al.
2978a. Eudonia angustea (Curtis, 1827) D (Buhl
et al. 2018).
3034a. Agriphila tolli (Błeszyński, 1952) F.
3185a. Xanthorhoe uralensis Choi, 2003 F.
3329a. Perizoma saxicola Tikhonov, 1994 E.
3497a. Ascotis Hübner, 1825
3497b. selenaria (Denis & Schiermüller, 1775)
S (Palmqvist & Ryrholm 2020), Lt (Ivinskis et
al. 2020).
3583a. Pungeleria Rougemont, 1903
3583b. capreolaria (Denis & Schiermüller,
1775) D (Skule & Hviid 2019).
3692a. Smerinthus caecus Ménétriés, 1857 Lv.
3773a. Spatalia Hübner, 1819
3773b. argentina (Denis & Schiermüller, 1775)
3782b. Eutelia Hübner, 1823
3782c. adulatrix (Hübner, 1813) D (Larsen et al.
3850a. Eilema caniola (Hübner, 1808) D (Larsen
4063a. Cucullia virgaureae Boisduval, 1840 F.
4068a. Phyllophila Guenée, 1852
4068b. obliterata (Rambur, 1833) F.
4178a. Spodoptera cilium Guenée, 1852 D (Hviid
4189a. Caradrina avirena Guenée, 1852 D
(Bech et al. 2017).
4387a. Lithophane leautieri (Boisduval, 1829) D
4422a. Aporophyla australis (Boisduval, 1829) D
(Larsen et al. 2019).
4531a. Mythimna alopecuri (Boisduval, 1840) D
(Larsen et al. 2019).
4629a. Noctua tertia von Mentzer, Moberg &
Fibiger, 1991 S (Palmqvist 2018).
Taxonomic changes
0029. Gazoryctra fuscoargenteus (O. Bang-
Haas, 1927) is a junior synonym. Replace with
G. uralensis (Grum-Grshimailo, 1899)
(Kallies & Farino 2018).
02590261, 02630264. According to Weidlich
& Arnscheid (2017) the parthenogenetic D.
lichenella (Linnaeus, 1761) is synonymous
with the bisexual D. fumosella (Heinemann,
1870). Robinson & Nielsen (1983)
synonymized D. lazuri (Clerck, 1759) with
D. fumosella. Consequently, the three names
lazuri, lichenella and fumosella represent
Norwegian Journal of Entomology 68, 1
14 (2021)
the same species. In addition, Weidlich &
Arnscheid (2017) synonymized D. norvegica
(Strand, 1919) with D. lichenella. These
authors overlooked the synonymy established
by Robinson & Nielsen (1983) and used the
name lazuri for a dierent Finnish species.
The latter is actually without name. Arnscheid
& Weidlich (2017) stated that it may represent a
bisexual form of D. fennicella (Suomalainen,
0346–0357. Regier et al. (2015) erected the
family Meessiidae for some genera which
until then had been placed in the subfamily
Meessiinae in Tineidae. In recent publications
on European Tineoidea the family Meessiidae
has become accepted (Budashkin & Bidzilya
2018, Gaedike 2019, Gaedike & Falck 2019).
The four genera of Meessiinae are placed in
the separate family Meessiidae before
Psychidae. However, it is still not certain if the
three genera Stenoptinea Dietz, 1905,
Karsholtia Gaedike, 1986 and Agnathosia
Amsel, 1954 really belong in Meessiidae.
0362. Haplotinea Diakono & Hinton, 1956
belongs to Myrmecozelinae (Gaedike 2019).
0361. The subfamily name Perissomasticinae
should be removed.
0664. Rhigognostis hufnagelii (Zeller, 1839) was
placed in the genus Eidophasia Stephens,
1842 (Baraniak 2020).
09921011. A molecular phylogeny of the subtribe
Cochylina was recently published (Brown
et al. 2020). These authors also indicated that
Cochylis epilinana (Duponchel, 1842)
should be placed in a genus of its own, but
it was overlooked that Longicornutia
Razowski, 1960 was available (Brown 2005,
Kovács & Kovács 2020). The reclassication
of species hitherto placed in Cochylidia
Obraztsov, 1956; Cochylis Treitschke, 1829
and Falseuncaria Obraztsov & Swatschek,
1958 is presented here:
Thyraylia Walsingham, 1897
nana (Haworth, 1811)
Cochylichroa Obraztsov & Swatschek, 1958
atricapitana (Stephens, 1852)
Pontoturania Obraztsov, 1943
posterana (Zeller, 1847)
Brevicornutia Razowski, 1960
pallidana (Zeller, 1847)
Cochylis Treitschke, 1829
roseana (Haworth, 1811)
aviciliana (Westwood, 1854)
Longicornutia Razowski, 1960
epilinana (Duponchel, 1842)
Neocochylis Razowski, 1960
hybridella (Hübner, 1813)
dubitana (Hübner, 1799)
Falseuncaria Obraztsov & Swatschek, 1958
degreyana (McLachlan, 1869)
ruciliana (Haworth, 1811)
Cochylidia Obraztsov, 1956
subroseana (Haworth, 1811)
richteriana (Fischer von Röslerstamm, 1837)
moguntiana (Rössler, 1864)
heydeniana (Herrich-Schäer, 1851)
implicitana (Wocke, 1856)
rupicola (Curtis, 1834)
0926. Acleris implexana (Walker, 1863) is
Nearctic (Gilligan et al. 2020). Replace with
A. ferrumixtana (Benander, 1934).
0987. Aethes adelaidae (Toll, 1955) raised
to species rank: (https://laji./en/taxon/
MX.5013777) F, Lv.
1471. Holcopogon bubulcellus (Staudinger,
1859) is a junior synonym and is replaced by
H. adseclella (Eversmann, 1844) (Sinev et al.
1629. Peleopodinae upgraded to family level. In
our area only the genus Carcina is represented
(Wang & Li 2020).
1635. Ethmiinae upgraded to family level and
placed close to Elachistidae (Wang & Li 2020).
1704a. Neofaculta taigana Ponomarenko, 1998 is
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
a cryptic species which has been confused with
N. infernella. It is a northern Holarctic species
and has been found in N, S and F (Aarvik et al.
1708. Anarsia lineatella Zeller, 1839 is to be
replaced by A. innoxiella Gregersen &
Karsholt, 2017 (Gregersen & Karsholt 2017).
1784. Apodia bifractella (Duponchel, 1843) is to
be replaced by Apodia martinii Petry, 1911.
The former has a more southern distribution in
Europe, but the ranges of the two species are
not known in detail (Huemer & Karsholt 2020).
1812. Monochroa parvulata (Gozmány, 1957)
was transferred to the genus Pragmatodes
Walsingham, 1908 (Huemer & Karsholt 2020).
1813. Eulamprotes Bradley, 1971 is a synonym
of Oxypteryx Rebel, 1911 (Bidzilya et al.
2019, Huemer & Karsholt 2020). Its
constituent species are transferred to Oxypteryx.
1914a. Corro Chang & Metz (2021) recently
transferred Tuta absoluta to Phthorimaea,
based on a phylogenetic analysis of 22
morphological characters of eight species. We
agree that T. absoluta is more closely related to
Phthorimaea operculella than to Tuta
atriplicella Kieer & Jörgensen, 1910.
However, their male genitalia are distinctly
dierent – much more than between many
genera of Gnorimoschemini. The latter
may be oversplit, and the generic structure
of Gnorimoschemini should be analysed
based on a larger number of taxa from dierent
genera and with a character set that includes
both morphological and molecular characters.
T. absoluta is a serious pest on tomatoes
etc., and the literature on it, since it was
introduced to the Old World, runs to
thousands of publications. Most of these are
on applied entomology, and researchers in that
eld are notoriously unhappy about changes in
nomenclature. Seen in the light of the
uncertainty of the generic placement of T.
absoluta, which is also admitted by Corro
Cheng & Metz (op. cit.: 51) who write: «Our
results support a need for generic re-
classication of the taxon absoluta under
its original genus, Phthorimaea», we nd
it premature to change its generic name. We
are of the opinion that for the time the stability
of nomenclature is best served by not changing
the generic combination, and we therefore
recombine it as Tuta absoluta (Meyrick),
comb. rev.
2053. Elachista zernyi Hartig, 1941 is a junior
synonym of E. stelviella Amsel 1932 (Kaila
2392. DNA barcoding shows that Danish
specimens identied as Stenoptilia inopinata
Bigot & Picard, 2002 is conspecic with
barcoded specimens of S. mariaeluisae Bigot
& Picard, 2002 from Iceland.
2548–2553. A revised classication of Argynnis
Fabricius, 1807 aiming at reconciling the
systems used in Europe and North America
was recently published (De Moya et al. 2017).
The new system was adopted in the new
European buttery checklist (Wiemers et
al. 2018). This results in the following updated
classication of the Nordic-Baltic species:
Argynnis Fabricius, 1807
paphia (Linnaeus, 1758)
laodice (Pallas, 1771)
Fabriciana Reuss, 1920
niobe (Linnaeus, 1758)
adippe (Denis & Schiermüller, 1775)
Speyeria Scudder, 1872
aglaja (Linnaeus, 1758)
2650. Scolitantides vicrama (Moore, 1865)
is transferred to Pseudophilotes Beuret, 1958
(Wiemers et al. 2018).
2654. Phengaris Doherty, 1891 has been given
priority over Maculinea Van Eecke, 1915
(ICZN 2017). This means that the nomenclature
used in our checklist can continue.
Norwegian Journal of Entomology 68, 1
14 (2021)
2763. Khorassania Amsel, 1951 is a synonym
of 2734 Pempelia Hübner, 1825. The species
Khorassania compositella is combined with
Pempelia as Pempelia compositella
(Treitschke, 1835) (Slamka 2019).
2765. Moitrelia Leraut, 2001 is a synonym of
Uncinus Amsel, 1951. The species Moitrelia
obductella is combined with Uncinus as
Uncinus obductella (Zeller, 1839) (Slamka
2836. Ephestia unicolorella Staudinger, 1881 is to
be replaced by E. woodiella Richards &
Thomson, 1932. The former is a southern
species known from Turkey and Syria (Leraut
2014, Buhl et al. 2020).
2843. Pyralis regalis (Denis & Schiermüller,
1775) is to be replaced by P. cardinalis Kaila,
Huemer, Mutanen, Tyllinen & Wikström, 2020
which was recently separated from the former
(Wikström et al. 2020).
3101–3609. The classication of Geometridae
has recently undergone numerous changes,
and a system which includes tribes is emerging.
The new system is presented in the 6th volume
of Geometrid Moths of Europe (Müller et al.
2019). Some additions and modications can
be found in Brehm et al. (2019), Murillo-
Ramos et al. (2019) and Sihvonen et al. (2020).
3604. Elophos Boisduval, 1840 is replaced by
Yezognophos Matsumura, 1927 (Müller et al.
3606. Glacies Millière, 1874 is a synonym of
Psodos Treitschke, 1825 (Müller et al. 2019).
3960a. Euclidia mi (Clerck, 1759) is placed in the
genus Callistege Hübner, 1823. Callistege
was upgraded from status as subgenus of
Euclidia Ochsenheimer, 1816 to status as a
separate genus by Lafontaine & Schmidt
(2010). Treating the two genera as synonyms
would also mean having to synonymize the
genus Mocis Hübner, 1823 (Zahiri et al. 2012:
g. 6).
4070. Protodeltote Ueda, 1984 was upgraded from
status as subgenus of Deltote Reichenbach,
1817 to full generic status by Schmidt et
al. (2018). This results in the combination
Protodeltote pygarga (Hufnagel, 1766).
4165. Nyctobrya Boursin, 1957 is replaced by
Bryopsis Boursin 1970 (Vargas-Rodríguez et
al. 2020).
4321. Resapamea hedeni (Graeser, 1888) is
a junior synonym. Replace with R. vulpecula
(Eversmann, 1852) (Kononenko 2005).
4345. Some of the genera proposed by Beck
(1992) were treated inconsistently in the
checklist. With some doubt we accept the
diagnosis given by Beck (1992) for Fissipunctia
as nomenclaturally valid. The publication by
Beck is part of the 1991 volume of the journal
Atalanta. It was, however, printed in 1992.
4360–4370. The monographic treatment of the
Eurasian and North African Agrochola Hübner,
1821 generic complex (L. Ronkay et al. 2017)
has led to a revised classication of the group:
Agrochola Hübner, 1821
lychnidis (Denis & Schiermüller, 1775)
Anchoscelis Guenée, 1839
Omphaloscelis Hampson, 1906 syn.
nitida (Denis & Schiermüller, 1775)
lunosa (Haworth, 1809)
litura (Linnaeus, 1758)
helvola (Linnaeus, 1758)
Leptologia L.B. Prout, 1901
lota (Clerck, 1759)
macilenta (Hübner, 1809)
Sunira Franclemont, 1950
circellaris (Hufnagel, 1766)
Propenistra Berio, 1980
laevis (Hübner, 1803)
4468. We consider the diagnosis given by Beck
(1992) as valid. Thus the genus should be cited
as Coranarta Beck, 1992.
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
4486. Lacanobia amurensis (Staudinger, 1901).
For this species the name Lacanobia aliena
(Hübner, 1809) has been used in European
literature. It was pointed out by Poole
(1989) that Noctua aliena Hübner, 1809 (now
in Lacanobia) is a primary homonym of
Noctua aliena Hübner, 1808 (a synonym of
Eriopygodes imbecilla (Fabricius, 1794)). In
the Danish catalogue (Karsholt & Stadel
Nielsen 1998) the name Lacanobia amurensis
(Staudinger, 1901) was adopted as a
replacement name. This was a nomenclatural
act in accordance with the 3rd edition of
the ICZN Code of 1985 which was in use
at that time. Had it been after 1999 when the
4th edition of the ICZN Code was published,
'reversed precedence' could have been used
to give Noctua aliena Hübner, 1809 precedence
over N. aliena Hübner, 1808. With Lacanobia
amurensis (Staudinger, 1901) now in prevailing
usage, reversal of precedence is not permitted.
Consequently, Lacanobia aliena (Hübner, 1809)
cannot be used as the valid name for this species.
4474. Polia conspicua (A. Bang-Haas, 1912)
is a junior synonym. Replace with P. vesperugo
Eversmann, 1856 (Varga et al. 2017).
0536. Phyllonorycter emberizaepenella. Spelling
of species name.
1339. Cydia splendana. Add parenthesis before
Hübner, 1799.
1480. Deuterogonia pudorina (Wocke, 1857).
Year of description.
1538. Agnoea elsae. Add parenthesis before and
after Svensson, 1982.
1683. Aproaerema vinella Bankes, 1898. Remove
2467. Carcharodus alceae (Esper, 1780). Year of
2512. Pontia chloridice (Hübner, 1813). Year of
2522. Colias croceus (Georoy, 1785). Author.
2546. Brenthis daphne (Denis & Schiermüller,
1775). Author and year.
2566. Nymphalis xanthomelas (Denis & Schier-
müller, 1775). Author and year.
2630. Chazara briseis. Spelling of species name
2666. Aricia nicias (Meigen, 1829). Year of
2857. Uresiphita gilvata (Fabricius, 1794). Delete
I’ (E. Ólafsson in litt.).
3396. Eupithecia goossensiata. The reference in
the comment on page 166 should be (Hausmann
et al. 2011) (not Hausmann et al. 2013).
3654. Lasiocampa quercus (Linnaeus, 1758).
Delete ‘I’ (E. Ólafsson in litt.).
4267. Longalatedes Beck, 1992. Year of
Acknowledgements. The authors wish to thank all persons
who have shared information about new records. In particular
we thank Thomas Pape, Copenhagen, for giving advice on
the nomenclatural problem concerning Lacanobia amurensis,
and Erling Ólafsson, Reykjavik, Iceland for information
on additions to the Lepidoptera fauna of Iceland. Per Falck,
Neksø, Denmark, informed about new species for Denmark in
2020. We are indebted to the referees for careful checking of
the manuscript, which resulted in numerous improvements.
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Accepted: 21 March 2021
Aarvik et al.: Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera
... It is native to North America, but was accidentally introduced to Italy, where it was first found in Milano in 1970 [7]. The moth has now been recorded in a number of European countries, ranging from Spain to Latvia [8][9][10][11][12][13][14][15][16][17][18][19][20][21]. [7]. ...
... [7]. The moth has now been recorded in a number of European countries, ranging from Spain to Latvia [8][9][10][11][12][13][14][15][16][17][18][19][20][21]. On average, the spread of the pest occurs at a speed of about 100 km per year [22]. ...
... Localities for P. robiniella were gathered from GBIF [31,32], records found in the literature (see https:// [13][14][15][16][17][18][19][20][21]), and our personal field surveys. As many uncertainties are associated with SDM projections, particularly when it comes to building a SDM for a species expanding its home range in a new area [33], we used for the analysis only records of European localities. ...
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The spread and outbreaks of phytophagous pests are often associated with global warming. In addition to economic interest, these species may be of interest in terms of biological indication of climate changes. In this context, we considered the locust digitate leafminer Parectopa robiniella Clemens, 1863 (Lepidoptera: Gracillariidae). This phytophage was first discovered in Europe in 1970 near Milano in Italy. Since then, it has been spreading across the continent. In Ukraine, it was recorded for the first time in 2003. In 2020–2021, we found areas of massive leaf damage caused by the black locust (Robinia pseudoacacia) in locations on Trukhaniv Island in Kyiv and some places in the Kyiv administrative region. Using 1041 georeferenced records of P. robiniella across Europe and a Bayesian additive regression trees algorithm (BART), we modeled the distribution of the moth. Predictors of current climate (WorldClim v.2, CliMond v.1.2 and ENVIREM) and a black locust habitat suitability raster were employed. Sets of SDMs built for P. robiniella with and without the habitat suitability raster for the host tree performed equally well. Amongst the factors that determine the niche of the locust digitate leafminer, most important are temperature-related conditions assumed to facilitate the spread and naturalization of the pest. In Ukraine, the appearance of the moth has coincided with increasing mean annual temperatures. Particularly favorable for the species are areas in the west and south-west of the country, and Transcarpathia. In the near future, the moth could reach locations in Nordic countries, Estonia, the British Isles, Black Sea coastal areas in Turkey, further into Russia, etc.
... The first appearance of the citrus flower moth (Prays citri (Millière, 1873) (Ivinskis, Rimšaitė, 2018). All three species are widely distributed in Northern Europe and develop on naturally growing plants (Aarvik et al., 2017). The species of the genus Prays is not easily distinguished, e.g., P. ruficeps can be separated from P. fraxinella only by DNA barcodes (Aarvik et al., 2017). ...
... All three species are widely distributed in Northern Europe and develop on naturally growing plants (Aarvik et al., 2017). The species of the genus Prays is not easily distinguished, e.g., P. ruficeps can be separated from P. fraxinella only by DNA barcodes (Aarvik et al., 2017). Two of them, the olive moth P. oleae (Bernard, 1788) and the citrus flower moth P. citri (Millière, 1873) can cause substantial economic losses in agriculture in the Mediterranean Basin (Carter, 1984). ...
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The citrus flower moth (Prays citri (Millière, 1873)) feeding on calamondin (Citrus × macrocarpa) in greenhouses in Lithuania were found on 20 April 2021. Based on a comprehensive analysis of morphological features of the moths, they were identified as Prays citri (Millière, 1873) (Lepidoptera: Praydidae). A small population of moths successfully survived in the unheated greenhouse till November. This species is one of the most harmful pests of citrus plants and can cause substantial economic losses in the Mediterranean basin and some tropical areas of the world. However, it has been detected in some central and even northern European countries in the last two decades. The article provides information about the first appearance of P. citri in Lithuania. Additionally, the paper includes a detailed and originally illustrated morphological description based on the collected specimens and a brief review of species distribution, biology, and the possible risk of this pest spreading in the country.
... The moths were attracted at light on a rainy night during the passage of a thunderstorm front between 11:00 P.M. and midnight. (Kozhantshikov 1950;Sugi 1982;Dubatolov et al. 1995;Kononenko et al. 1998;Zolotarenko & Dubatolov 2000;Ahola & Silvonen 2005;Kononenko 2005Kononenko , 2010Nupponen & Fibiger 2006;Matov et al. 2008;Fibiger et al. 2009;Volynkin 2012;Aarvik et al 2017). ...
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Trichosea ludifica (Linnaeus, 1758) is reported from Kazakhstan (East Kazakhstan Region, West Altai Mountains) for the first time. The species habitat in Kazakhstan is briefly characterized and illustrated. A list of noctuid moths occurring syntopically with T. ludifica is also provided.
... For consistency, the genus-level taxonomy of Aarvik et al. [17] was followed supplemented by that of Waring & Townsend [18]. Between them, these two references cover the entire region of Northern Europe, as described under the United Nations geoscheme (https://unstats.un. ...
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Ecological and life-history data on the Northern European macromoth (Lepidoptera: Macroheterocera) fauna is widely available and ideal for use in answering phylogeny-based research questions: for example, in comparative biology. However, phylogenetic information for such studies lags behind. Here, as a synthesis of all currently available phylogenetic information on the group, we produce a supertree of 114 Northern European macromoth genera (in four superfamilies, with Geometroidea considered separately), providing the most complete phylogenetic picture of this fauna available to date. In doing so, we assess those parts of the phylogeny that are well resolved and those that are uncertain. Furthermore, we identify those genera for which phylogenetic information is currently too poor to include in such a supertree, or entirely absent, as targets for future work. As an aid to studies involving these genera, we provide information on their likely positions within the macromoth tree. With phylogenies playing an ever more important role in the field, this supertree should be useful in informing future ecological and evolutionary studies.
... Lepidoptera families are arranged according to Aarvik et al. (2017). Sequence and nomenclature of families Erebidae and Noctuidae follow Fibiger et al. (2011) with subsequent changes incorporated from recent taxonomic revisions. ...
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Paracossulus thrips (Hübner, 1818) and Lignyoptera fumidaria (Hübner, 1825), both species listed on Annex II of the Council Directive 92/43/EEC, are reported for the first time for Serbia from two distinct localities on Vidlič Mt. Some new localities from the edge of the range of both these species are reported from Bulgaria. Previous reports of these species in the Balkan Peninsula are summarized and comments are given on the habitats, threats and conservation requirements of both species. Serbian vernacular names for both species are proposed.
... As there is just one signum in the bursa copulatrix of N. estonica (Figures 49-51), it appears undeniable that this species cannot be conspecific with N. holsatica. This view is further supported by the report by Aarvik et al. (2017) that the DNA barcodes of N. holsatica and N. aerugula are 'almost identical', whereas there is a large (in average 0.042) genetic distance between the DNA barcodes of N. aerugula and N. estonica. ...
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Nola estonica Õunap sp. nov. (Lepidoptera, Nolidae, Nolinae) is described based on type material from Estonia. The lectotype is designated for Glaphyra atomosa Bremer, 1861, which is reinstated from a subspecies of Nola aerugula (Hübner, [1793]) to a full species: Nola atomosa (Bremer, 1861) stat. rev. The status of these three taxa as separate species is supported by the results of phylogenetic analysis of DNA barcodes, as well as external and genital morphology of adult specimens. Two new synonyms are established as follows: Nola atomosa (Bremer, 1861) = Nola candidalis Staudinger, 1892 syn. nov. and Nola shin Inoue, 1982 syn. nov. N. estonica occurs sympatrically with N. aerugula in Estonia, and with N. atomosa in South Korea and easternmost Russia. While the available data suggest a disjunct distribution of N. estonica (eastern Europe and the temperate Far East), it appears highly possible that the species has a wide transpalaearctic distribution.
... Whereas R. tomentosum is becoming increasingly rare towards the South, with only a few relict populations in peat bogs of Central Europe north of the Alps, Myrica gale is completely absent from the southern part of Central Europe. Following its host-plants L. ledi is widely distributed from northern Europe to the Far East (Aarvik et al. 2017;Baryshnikova 2019), with an increasingly patchy distribution pattern in northern Central Europe (Spitzer et al. 1996), and isolated populations in southern Czech Republic and northernmost Austria (Lower Austria) (Laštůvka and Liška 2011;Huemer 2013). The newly detected mountainous population of L. ledi from Switzerland is highly isolated at a distance of about 440 km from the closest confirmed records in the northern part of Lower Austria (Huemer 2013). ...
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Lyonetia ledi Wocke, 1859 (Lyonetiidae), was hitherto considered as a boreal species with a circumpolar distribution pattern and relict populations in isolated peat bogs north-east of the Alps (Austria, Czech Republic, Germany). In Europe it is known as a leaf-miner on Rhododendron tomentosum Stokes ex Harmaja (Ericaceae) as the primary host-plant and also Myrica gale L. (Myricaceae). The first record of L. ledi from the Swiss Alps on Rhododendron ferrugineum L., the famous Alpenrose, indicates an ancient host-plant switch during postglacial periods when R. tomentosum and R. ferrugineum shared habitat in the prealps. Conspecificity with northern populations is supported by the adult morphology and supplementing DNA barcodes (mtDNA COI gene). L. ledi is the first obligatory leaf-mining species on R. ferrugineum . Details of the life-history and habitat are described and figured. The record finally substantiates the probability of an autochthonous population in Carinthia (Austria), from where the species was recently published as new to the Alps.
... Funnene som presenteres her er basert på materiale innsamlet eller sjekket av forfatterne. Taksonomi og nomenklatur følger den nordisk-baltiske sjekklista (Aarvik et al. 2017 Med et vingespenn på 6-8 mm., er dette en stor art til dvergmøll å vaere. Den er helt uten tegninger, og varierer i fargen fra grågul til gråbrun, men enda lysere eller mørkere individer forekommer. ...
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Insekt-Nytt 46 (1) 2021 New species of lepidoptera discovered in Norway. Denne oppdateringen inneholder i hovedsak funn fra 2019 og 2020. De fleste nye arter som har dukket opp i løpet av de to sesongene er enten migranter eller arter som er i ekspansjon mot nord. To av artene er resultat av endret taksonomi, det vil si artssplittinger.
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A male specimen of Phyllonorycter schreberella (Fabricius, 1781) (Lepidoptera: Gracillariidae) was found in western Norway in Vestland at Lærdal: Husum in June 2021. The circumstances of the record, the species’ biology, geographical range, identification and status are outlined. Photos of the moth and the locality are given.
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Insekt-Nytt 46 (1) 2021 New species of lepidoptera discovered in Norway. Denne oppdateringen inneholder i hovedsak funn fra 2019 og 2020. De fleste nye arter som har dukket opp i løpet av de to sesongene er enten migranter eller arter som er i ekspansjon mot nord. To av artene er resultat av endret taksonomi, det vil si artssplittinger.
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This report represents data on 2 new (Celypha woodiana and Retinia perangustana) and 42 rare for the Lithuanian fauna Tortricidae species. Species were collected from 11 administrative districts of Lithuania.
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Monopis jussii Kaila, Mutanen, Huemer, Karsholt & Autto, sp. nov. (Lepidoptera, Tineidae) is described as a new species. It is closely related to the widespread and common M. laevigella ([Denis & Schiffermül-ler], 1775), but differs in its distinct COI DNA barcode sequences, four examined nuclear loci as well as details in forewing coloration and pattern. Most reared specimens of M. jussii have emerged from the nest remnants of the Boreal owl (Aegolius funereus (Linnaeus, 1758)), but also nests of the Ural owl (Strix uralensis Pallas, 1771) and the Great tit (Parus major Linnaeus, 1758) have been observed as suitable habitats. Based on the present knowledge, the new species has a boreo-montane distribution as it is recorded only from northern Europe and the Alps. Several extensive rearing experiments from Strix spp. nest remnants from southern Finland did not produce any M. jussii, but thousands of M. laevigella, suggesting that the species is lacking in the area or, more unlikely, that the nest of these owl species do not serve as good habitat for the new species. This unexpected species discovery highlights, once again, the usefulness of DNA barcoding in revealing the cryptic layers of biodiversity. To serve stability we select a neotype for Tinea laevigella [Denis & Schiffermüller], 1775, and discuss the complicated synonymy and nomenclature of this species. A peer-reviewed open-access journal Marko Mutanen et al. / ZooKeys 992: 157-181 (2020) 158
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In support of a comprehensive update to the checklist of the moths of North America, we attempt to determine the status of 151 species of Tortricidae present in North America that may be Holarctic, introduced, or sibling species of their European counterparts. Discovering the natural distributions of these taxa is often difficult, if not impossible, but several criteria can be applied to determine if a species that is present in both Europe and North America is natively Holarctic, introduced, or represented by different but closely related species on each continent. We use DNA barcodes (when available), morphology, host plants, and historical records (literature and museum specimens) to make these assessments and propose several taxonomic changes, as well as future areas of research. The following taxa are raised from synonymy to species status: Acleris ferrumixtana (Benander, 1934), stat. rev.; Acleris viburnana (Clemens, 1860), stat. rev.; Acleris pulverosana (Walker, 1863), stat. rev.; Acleris placidana (Robinson, 1869), stat. rev.; Lobesia spiraeae (McDunnough, 1938), stat. rev.; and Epiblema arctica Miller, 1985, stat. rev. Cydia saltitans (Westwood, 1858), stat. rev., is determined to be the valid name for the "jumping bean moth," and Phiaris glaciana (Möschler, 1860), comb. n., is placed in a new genus. We determine that the number of Holarctic species has been overestimated by at least 20% in the past, and that the overall number of introduced species in North America is unexpectedly high, with Tortricidae accounting for approximately 23-30% of the total number of Lepidoptera species introduced to North America.
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The taxonomic position of two species Plutella hufnagelii Zeller 1839 and Caunaca insulella Walsingham 1900, in the family Plutellidae: has been unclear. In current fauna studies, these species are usually placed in the genera Rhigognostis and Eidophasia respectively. In this paper, I address the taxonomic position of P. hufnagelli and C. insulella and describethe genitalia of both sexes. Analysis of genital structure confirms the correct placement of these species into the genus Eidophasia.
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The checklist of European Gelechiidae covers 865 species, belonging to 109 genera, with three species records which require confirmation. Further, it is the first checklist to include a complete coverage of proved synonyms of species and at generic level. The following taxonomic changes are introduced: Pseudosophronia con-stanti (Nel, 1998) syn. nov. A peer-reviewed open-access journal Peter Huemer & Ole Karsholt / ZooKeys 921: 65-140 (2020) 66 Dichomeris neatodes Meyrick, 1923 sp. rev.; Caryocolum horoscopa (Meyrick, 1926) stat. rev.; Ivanauskiella occitanica (Nel & Varenne, 2013) sp. rev.; Apodia martinii Petry, 1911 sp. rev.; Caulastrocecis cryptoxena (Gozmány, 1952) sp. rev. Following Article 23.9.2 ICZN we propose Caryocolum blandella (Douglas, 1852) (Gelechia) nom. protectum and Caryocolum signatella (Eversmann, 1844) (Lita) nom. oblitum.
A multigene phylogenetic study was carried out to test current, mostly morphology‐based hypotheses on Sterrhinae phylogeny with additional material included from further geographical areas and morphologically different lineages. A maximum likelihood analysis (11 molecular markers and 7665 bp) was conducted on 76 species and 41 genera using iq‐tree software. The resulting phylogenetic hypothesis is well resolved and branches have high support values. Results generally agree with earlier hypotheses at tribal levels and support the hypothesis that Sterrhinae comprises two major lineages. Based on the molecular phylogeny and extensive morphological examination, nine tribes are considered valid and the following taxonomic changes are introduced to recognize monophyletic groups: Mecoceratini Guenée, 1858 (= Ametridini Prout, 1910) is transferred from Desmobathrinae to Sterrhinae, and it is considered valid at tribal level new classification; Haemaleini Sihvonen & Brehm is described as a new tribe and deemed sister to Scopulini + Lissoblemmini; Lissoblemmini Sihvonen & Staude is described as a new tribe and sister to Scopulini; Lythriini Herbulot, 1962 is now a junior synonym of Rhodometrini Agenjo, 1952 syn.n.; and Rhodostrophiini Prout, 1935 is now a junior synonym of Cyllopodini Kirby, 1892 syn.n. In addition, 48 taxa are transferred from other geometrid subfamilies to Sterrhinae, or within Sterrhinae from one tribe to another, or they are classified into a tribe for the first time, or a new genus classification is proposed. The results demonstrate the limited explanatory power of earlier classifications, particularly at the tribal level. This is probably a result of earlier classifications being based on superficial characters and biased towards the European and North American fauna. The species richness and distribution of Sterrhinae and its constituent tribes are reviewed, showing that the globally distributed Sterrhinae are most diverse in the Neotropics (31% of global fauna). They are species‐rich in the Palaearctic (22%), Afrotropics (19%) and Indo‐Malay (16%) regions, whereas they are almost absent in Oceania (1%). In terms of the described fauna, the most species‐rich tribes are Scopulini (928 species), Sterrhini (876 species) and Cosymbiini (553 species), all of which have a cosmopolitan distribution. Mecoceratiini and Haemaleini are almost entirely Neotropical. Timandrini and Lissoblemmini, by contrast, are absent in the Neotropics. We present a revised classification of the global Sterrhinae fauna, which includes about 3000 putatively valid species, classified into nine tribes and 97 genera. Four genera are of uncertain position within Sterrhinae. Our results highlight the compelling need to include more genera from a global perspective in molecular phylogenetic studies, in order to create a stable global classification for this subfamily. This published work has been registered on ZooBank,‐06D6‐4908‐893E‐E8B124BB99B1. We analysed the phylogeny of Sterrhinae moths based on molecular dataset of 76 species and 11 genes, combined those with morphology and included the results in a global classification framework. Two new tribes are described, Mecoceratini is transferred from Desmobathrinae to Sterrhinae, and 50 other taxonomic changes are proposed. Sterrhinae are a cosmopolitan group of about 3000 species, with the highest species richness in the Neotropics and the lowest in Oceania.