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Type material of Clausiliidae door snails from Philippe Dautzenberg in the collection of the Museum of New Zealand Te Papa Tongarewa

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A small collection of Vietnamese door snails (family Clausiliidae) was recently (re)discovered in the malacological collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ). This material previously belonged to the famed shell collection of Belgian malacologist Philippe Dautzenberg (1849-1935) and was eventually acquired by, or donated to, NMNZ in the first half of the twentieth century. The collection comprises type material of 19 taxa (species and subspecies) and possible type material of one taxon, alongside non-type material of several additional species. It is presented here in the form of an annotated and illustrated catalogue.
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Copyright © Museum of New Zealand Te Papa Tongarewa (2020)
Ty p e m a t e r i a l o f C l a u s i l i i d a e d o o r s n a i l s
from Philippe Dautzenberg in the collection
of the Museum of New Zealand Te Papa Tongarewa
Rodrigo B. Salvador1* and Abraham S.H. Breure2,3
1 Museum of New Zealand Te Papa Tongarewa, PO Box 467, Wellington 6011, New Zealand
2 Royal Belgian Institute of Natural Science, Vautierstreet 29, 1000, Brussels, Belgium
3 Department of Life Sciences, Invertebrate Division, Natural History Museum, London SW7 5BD, UK
* Corresponding author: salvador.rodrigo.b@gmail.com
ABSTRACT: A small collection of Vietnamese door snails (family Clausiliidae) was recently (re)discovered
in the malacological collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ). This
material previously belonged to the famed shell collection of Belgian malacologist Philippe Dautzenberg
(1849–1935) and was eventually acquired by, or donated to, NMNZ in the first half of the twentieth
century. The collection comprises type material of 19 taxa (species and subspecies) and possible type
material of one taxon, alongside non-type material of several additional species. It is presented here in
the form of an annotated and illustrated catalogue.
KEYWORDS: Philippe Dautzenberg, Arthur Bavay, Victor Demange, Louis Messager, Gastropoda,
collections, pulmonates, type material, snails, Vietnam.
Introduction
During ongoing work to reorganise the terrestrial and
freshwater gastropod collection of the Museum of New
Zealand Te Papa Tongarewa (NMNZ), Wellington,
New Zealand, a small but important collection of land
snails was discovered. This material belonged to the famed
shell collection of malacologist Philippe Dautzenberg and
was eventually acquired by, or donated to, NMNZ in
circumstances that remain largely unknown.
Philippe Dautzenberg was born in Brussels in 1849 and
moved to Paris in 1867 to work in the Braquenié tapestry
company. He married one of the owner’s daughters, Marie
Gabrielle Braquenié, succeeded his father-in-law Henri
Charles as director and made his fortune in the company.
This allowed him to build up a huge shell collection over
the years, which at the turn of the century was already
considered the most important private malacological
collection in the world, both in terms of the diversity of
species and the quality of specimens. When Dautzenberg
died in 1935, his collection contained about 40,000 species
and was bequeathed to the Belgian state. It is now housed in
the Royal Belgian Institute of Natural Sciences in Brussels,
together with Dautzenberg’s library and archives (Lamy
1935; Duchamps 1986, 1989).
The small Dautzenberg collection at NMNZ is composed
almost exclusively of Vietnamese door snails (family
Clausiliidae), although one species from Japan and one from
China are also present. Most material was obtained by Louis
G.M. Messager (1852–1915), who served in the French
army and was stationed for many years in the French colony
of Indochina (Breure & Páll-Gergely 2019). Further material
was collected by Victor Demange (1870–1940), a trader who
lived for many years in Hanoi (Breure & Ablett 2016). The
specimens mostly represent species and subspecies described
by Dautzenberg, in collaboration with Arthur R.J.B.
Bavay (1840–1923), a French pharmacist and malacologist
(Roth & Clover 1973). The labels of all NMNZ lots bear
the indication ‘From the type series’ or similar, so they are
potential candidates to be type material of these clausiliid
taxa. We investigate this matter in our article and provide a
list of all Dautzenberg material housed at NMNZ.
Tuhinga
31: 56–69
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Material and methods
The small Dautzenberg collection housed at NMNZ
contains 51 lots. Unfortunately, its history remains largely
uncertain. According to the internal collection registers,
the collection was received on 4 December 1935. The
register points to exactly 51 lots of Clausilia, so we can
assume that none of the original material has been lost.
About one-third of the lots had already been entered
in the NMNZ database, which means their original
labels have been discarded and substituted with new
ones (either handwritten or printed), an unfortunate
practice in NMNZ until relatively recently. However,
the original labels survived from the unregistered lots
(Fig. 1), confirming their provenance from Dautzenberg’s
collection. Furthermore, the ledger at NMNZ states that
the small collection was a donation by G.E. Mason. There
is no known biographical information available about
Mason (whose first name was likely George), other than
that he was based in the UK and that he also donated
specimens to the Naturalis Biodiversity Center (Leiden, the
Netherlands) and the Natural History Museum (London,
UK) (Mogollón Avila & Breure 2009). In any event,
given the date, Mason likely received this material from
Dautzenberg during the latter’s lifetime (Dautzenberg died
on 9 May 1935). Research at the Royal Belgian Institute
of Natural Sciences, where Dautzenberg’s collection and
archives are now housed, confirmed that this material
never reached that museum (T. Backeljau, pers. comm.,
24 December 2019).
In a few cases (e.g. NMNZ M.306040) the labels
indicate that the specimens originally belonged to Bavay’s
collection, which was reportedly dispersed after his death
(Dance 1966); it is known that Dautzenberg acquired part
of this (Breure 2015). A few other labels still indicate other
previous owners of the material, including Jules F. Mabille,
Otto F. von Möllendorff and Henry A. Pilsbry (see
below). It is uncertain whether the specimens belonged to
Dautzenberg before being acquired by Mason, but that is
likely given the uniformity of the label format.
Since several of these lots proved to contain type
material, we present below an annotated and illustrated
catalogue of all taxa present in this collection. The species
are listed alphabetically according to their specific epithet,
followed by information on their original description, type
material, type locality, current taxonomic status (Nordsieck
2002a,b, 2007, 2011; Grego & Szekeres 2011; Schileyko
2011; MolluscaBase 2020), a transcript of the label, and
further remarks where pertinent. A few specimens proved
not to be types; they are listed and briefly discussed in
the Appendix to provide a comprehensive overview of
Dautzenberg’s material in the NMNZ collection. Material
of several species was listed in Fischer-Piette (1950) as the
‘holotype’, which according to Article 74.6 (International
Commission on Zoological Nomenclature 1999) must be
interpreted as the lectotype designation.
Fig. 1 Examples of original labels from the Dautzenberg material. A, Clausilia acrostoma Bavay & Dautzenberg, 1909,
NMNZ M.306049; B, C. lemyrei Bavay & Dautzenberg, 1899, from Bavay’s collection, NMNZ M.306040; C,
example of an unpublished manuscript name, NMNZ M.329559.
Dautzenberg’s snail types in Te Papa’s collection
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Systematics
babeensis Bavay & Dautzenberg,
1899, Clausilia (Fig. 2B)
Clausilia babeensis
Bavay & Dautzenberg, 1899b: 278, pl.
12, fig. 3.
type mate ri al: paralectotypes NMNZ M.202247
(2 shells, Messager leg.).
labe l : (not original)
‘Clausilia babeensis
B. et D. Near Lake
Ba Be, Tonkin. Coll. Messager. Ex Dautzenberg Coll. From
the type series.
type lo cali ty: ‘dans un ilot de l’un des lacs Ba-Bé’.
Vietnam, on a small island on Lake Ba B.
current taxonomy: Accepted as Synprosphyma babeensis
(Bavay & Dautzenberg, 1899).
remarks: The label (not original) indicates only ‘near Lake
Ba Be’ as the locality, and does not mention an island or islet.
Fischer-Piette (1950: 167) listed one specimen (MNHN-
IM-2000-2479) as the ‘holotype’, which is the lectotype
designation. There are further paralectotypes in RBINS
MT.2520 (9).
backanensis Bavay & Dautzenberg,
1899, Clausilia (Fig. 2C)
Clausilia backanensis Bavay & Dautzenberg, 1899b: 281, pl.
12, fig. 5.
TYPE MATERIAL: paralectotypes NMNZ M.202239 (6 shells,
Messager leg.).
LABEL: (not original) Clausilia backanensis Bav. et Dautz.
Bac Kan, Tonkin. Coll. Messager. Ex Coll. Dautzenberg.
From the type series.’
TYPE LOCALITY: ‘Les environs de Bac-Kan’. Vietnam, Bc
Kn.
CURRENT TAXONOMY: Junior synonym of Phaedusa lypra
(Mabille, 1887).
REMARKS: Fischer-Piette (1950: 167) listed one specimen
(MNHN-IM-2000-2406) as the ‘holotype’, which is the
lectotype designation.
callistoma Bavay & Dautzenberg,
1899, Clausilia (Fig. 2D)
Clausilia callistoma Bavay & Dautzenberg, 1899b: 41, pl. 2,
figs 3–3a.
TYPE MATERIAL: paralectotypes NMNZ M.202223 (2
shells, Messager leg.).
LABEL: (not original) Clausilia callistoma Bav. et Dautz.
That Khe, Tong Kin. Coll. Lt.Col. Messager. Ex Coll. Ph
Dautzenberg. From the type series.’
TYPE LOCALITY: ‘De Lang-Son à That-Khé’. Vietnam,
between Lng Sn and Tht Khê.
CURRENT TAXONOMY: Accepted as Selenophaedusa callistoma
(Bavay & Dautzenberg, 1899).
REMARKS: Fischer-Piette (1950: 166) listed one specimen
(MNHN-IM-2000-2409) as the ‘holotype’, which is the
lectotype designation.
cristata Bavay & Dautzenberg,
1899, Clausilia dorri (Fig. 2E)
Clausilia (Pseudonenia) Dorri var.. cristata Bavay &
Dautzenberg, 1899b: 283.
TYPE MATERIAL: possible paralectotypes NMNZ M.202227
(6 shells, Messager leg.).
LABEL: (not original) Clausilia dorri Bav. et Dautz. var.
cristata Bav. et Dautz. Upper Tong Kin. Ex Coll. Ph.
Dautzenberg. From the type series.’
TYPE LOCALITY: ‘le voisinage des lacs Ba-Bé’. Vietnam,
Lake Ba B.
CURRENT TAXONOMY: Accepted as Tropidauchenia bavayi
(Lindholm, 1924), a substitute name for the preoccupied
cristata.
REMARKS: The label, although not the original, gives only
‘Upper Tongkin’ as the collection locality, which is to the
south of Lake Ba B. Bavay & Dautzenberg (1899b: 285)
stated that specimens of all four varieties they described came
from the vicinities of that lake. As such, the status of the
present specimens as types remains uncertain. Fischer-Piette
(1950: 167) listed a specimen (MNHN-IM-2000-2445) as a
‘type’ of the variety, which is here considered the lectotype
designation. There is further type material in RBINS
MT.2472 (25 paralectotypes).
INSTITUTIONAL ACRONY MS : ANSP, Academy of Natural Sciences, Drexel University (Philadelphia, USA); KIU, Kyoto
University Museum (󱮕󱁳󰥁󲀌; Kyoto, Japan); MNHN, Musée national d’Histoire naturelle (Paris, France);
NMNZ, Museum of New Zealand Te Papa Tongarewa (Wellington, New Zealand); RBINS, Royal Belgian Institute of
Natural Sciences (Brussels, Belgium).
59
Fig. 2 Clausiliidae types from the NMNZ collection; scale bars = 5 mm. A,
paralectotype of Clausilia porphyrostoma
Bavay & Dautzenberg, 1909, NMNZ M.202225;
B, p
aralectotype of
C. babeensis Bavay & Dautzenberg, 1899,
NMNZ M.202247; C, paralectotype of C. backanensis Bavay & Dautzenberg, 1899, NMNZ M.202239; D,
paralectotype of C. callistoma
Bavay & Dautzenberg, 1899, NMNZ M.202223;
E, p
ossible paralectotype of
C. dorri
cristata
Bavay & Dautzenberg, 1899, and C. bavayi Lindholm, 1924,
NMNZ M.202227;
F, s
yntype
of
C. duella
Mabille, 1887,
NMNZ M.306046;
G, p
aralectotype of
C.
falcifera
Bavay & Dautzenberg, 1899,
NMNZ M.306039;
H, paralectotype of C. fistulata Bavay & Dautzenberg, 1909, NMNZ M.202224; I, syntype of
Garnieria goliath H. Rolle, 1910, NMNZ M.202228.
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duella Mabille, 1887, Clausilia (Fig. 2F)
Clausilia duella Mabille, 1887a: 16.
TYPE SPECIMEN: syntype NMNZ M.306046 (1 shell, ex
Mabille colln.).
LABEL: (original) ‘DUELLA Mabille. Tonkin. Ex coll. Dr.
Ph. Dautzenberg, and Dr. Mabille. [From the Type series.]’
TYPE LOCALITY: Vietnam, Tonkin.
CURRENT TAXONOMY: Accepted as Dautzenbergiella duella
(Mabille, 1887).
REMARKS: One syntype, MNHN-IM-2000-2446, listed in
Fischer-Piette (1950: 167) as a figured specimen by Bavay
& Dautzenberg (1899a: pl. 1, fig. 7).
falcifera Bavay & Dautzenberg,
1899, Clausilia (Fig. 2G)
Clausilia falcifera Bavay & Dautzenberg, 1899b: p. 290, pl.
12, fig. 10.
TYPE MATERIAL: paralectotypes NMNZ M.306039 (2 shells,
Messager leg.).
LABEL: (original) ‘FALCIFEAR [sic] Bav. and Dautz.
Bac Kan, Tongking. Coll. Messager. Ex Coll. Dr. Ph.
Dautzenberg. [From the type series.]’
TYPE LOCALITY: ‘Entre Bac-Kan et That-Khé’. Vietnam,
between Bc Kn and Tht Khê.
CURRENT TAXONOMY: Junior synonym of Selenophaedusa
billeti (H. Fischer, 1898).
REMARKS: The label (original) mentions only Bc Kn as
the locality. Fischer-Piette (1950: 167) listed one specimen
(MNHN-IM-2000-2468) as the ‘holotype’, which is the
lectotype designation.
fistulata Bavay & Dautzenberg,
1909, Clausilia (Fig. 2H)
Clausilia fistulata Bavay & Dautzenberg, 1909: 92, pl. 2,
figs 15–16.
TYPE MATERIAL: paralectotypes NMNZ M.202224 (4 shells,
Messager leg.).
LABEL: absent; only recent electronic database information
available.
TYPE LOCALITY: ‘Pac-Kha, Muong-Bo et Gia-Phu’.
Vietnam, Bc Hà, Mng and Gia Phú.
CURRENT TAXONOMY: Accepted as Synprosphyma fistulata
(Bavay & Dautzenberg, 1909).
REMARKS: Fischer-Piette (1950: 174) listed one specimen
(MNHN-IM-2000-2471) as the ‘holotype’, which is the
lectotype designation.
goliath H. Rolle, 1910, Garnieria (Fig. 2I)
Garnieria goliath H. Rolle, 1910: 135, textfig.
TYPE MATERIAL: syntypes NMNZ M.202228 (3 shells,
Demange leg. 1910, ex H. Rolle colln.).
LABEL: (not original) Clausilia goliath Rolle. Phu-Ly
Phongy, Tonkin. Coll. Demmge [sic], 1910. Ex Coll. H.
Rolle. From the type series.’
TYPE LOCALITY: ‘Tongking, inter Phu-ly et Ké-So’.
Vietnam, between Ph Lý and KS.
CURRENT TAXONOMY: Junior synonym of Megalauchenia
proctostoma (Mabille, 1889).
REMARKS: The label (not original) gives only Ph Lý as the
locality, while the species description presents a broader
type locality (from Ph Lý to KS). Nor does the label
mention Dautzenberg, although given the remaining
material, it can be surmised the specimens came from his
collection as well.
grangeri Bavay & Dautzenberg,
1899, Clausilia (Fig. 3A)
Clausilia Grangeri Bavay & Dautzenberg, 1899a: 46, pl. 3,
figs 8–8b.
TYPE MATERIAL: syntypes NMNZ M.202242 (3 shells and 1
fragmentary aperture, Messager leg.).
LABEL: (not original) Clausilia grangeri B. et D. That Khe,
Tonkin. Coll. Messager. Ex Coll. Dautzenberg. From the
type series.’
TYPE LOCALITY: ‘Entre Lang-Son et That-Khé’. Vietnam,
between Lng Sn and Tht Khê.
CURRENT TAXONOMY: Accepted as Siphonophaedusa grangeri
(Bavay & Dautzenberg, 1899).
REMARKS: Fischer-Piette (1950: 167) listed two syntypes
(MNHN-IM-2000-2417).
61
Fig. 3 Clausiliidae types from the NMNZ collection (cont.); scale bars = 5 mm. A, syntype of Clausilia grangeri Bavay &
Dautzenberg, 1899, NMNZ M.202242; B, paralectotype of C. inanis Bavay & Dautzenberg, 1909, NMNZ M.306045;
C, paralectotype of C. lemyrei Bavay & Dautzenberg, 1899, NMNZ M.306040; D, paralectotype of C. mairei Bavay &
Dautzenberg, 1909, NMNZ M.202237; E, paralectotype of C. mairei Bavay & Dautzenberg, 1909, NMNZ M.306044
(in scale with specimen in Fig. 3D); F, paralectotype of C. margaritifera Bavay & Dautzenberg, 1909, NMNZ M.202243;
G, paralectotype of C. messageri Bavay & Dautzenberg, 1899, NMNZ M.202245; H, paralectotype of C. moirati Bavay &
Dautzenberg, 1909, NMNZ M.202229; I, syntype of Phaedusa gisota platyloma Möllendorff, 1901, NMNZ M.306043.
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inanis Bavay & Dautzenberg, 1909,
Clausilia (Fig. 3B)
Clausilia inanis Bavay & Dautzenberg, 1909: 103, pl. 3, figs
15–16.
TYPE MATERIAL: paralectotypes NMNZ M.306045 (3 shells,
Messager leg.).
LABEL: (original) ‘INANIS B. and D. Pac Kha, Tongking.
Coll. Messager. Dr. Ph. Dautzenberg. [From the Type
series.]’
TYPE LOCALITY: ‘Pac-Kha’. Vietnam, Bc Hà.
CURRENT TAXONOMY: Accepted as Phaedusa inanis (Bavay
& Dautzenberg, 1909).
REMARKS: Fischer-Piette (1950: 174) listed one specimen
(MNHN-IM-2000-2494) as the ‘holotype’, which is the
lectotype designation.
lemyrei Bavay & Dautzenberg, 1899,
Clausilia (Fig. 3C)
Clausilia lemyrei Bavay & Dautzenberg, 1899b: 275, pl. 12,
fig. 1.
TYPE MATERIAL: paralectotypes NMNZ M.306040 (2 shells,
ex A.R.J.B. Bavay colln.).
LABEL: (original) ‘LEMYREI Bav. and Dautz. Tonkin.
Coll. Ex coll. Dr. A. Bavay. [From the Type series.]’
TYPE LOCALITY: Not specifically given, but title refers to
‘Indo-Chine’.
CURRENT TAXONOMY: Accepted as Hemiphaedusa lemyrei
(Bavay & Dautzenberg, 1899).
REMARKS: The label (original) gives Tonkin as the locality.
Fischer-Piette (1950: 167) listed one specimen (MNHN-
IM-2000-2500, from ‘That-Khé’) as the ‘holotype’, which
is the lectotype designation.
mairei Bavay & Dautzenberg, 1909, Clausilia
(Figs 3D and 3E)
Clausilia Mairei Bavay & Dautzenberg, 1909: 96, pl. 3,
figs 1–2.
TYPE MATERIAL: paralectotypes NMNZ M.202237 (2 shells,
Messager leg.), NMNZ M.306044 (2 shells, Messager leg.).
LABELS: NMNZ M.202237 (not original) Clausilia mairei
Bav. et Dau. Pac Kha, Tonkin. Coll. Messager. Ex Coll. Ph.
Dautzenberg. From the type series.’; NMNZ M.306044
(original) ‘MAIREI B. and D. Lao Kay. Coll. Messager.
Dr. Ph. Dautzenberg. [From the Type series.]’.
TYPE LOCALITY: ‘province de Lao-Kay et toute du rive
droite du Fleuve rouge’. Vietnam, Lào Cai province, entire
right bank of Red River.
CURRENT TAXONOMY: Accepted as Oospira mairei (Bavay &
Dautzenberg, 1909).
REMARKS: The label (not original) of lot NMNZ M.202237
brings a more specific locality within Lào Cai province,
namely Pc Kha. This is an old name for Bc (c. 908 m,
22.53911 N, 104.29305 E). The label of the other lot,
NMNZ M.306044, is original and indicates only Lào Cai
province as the locality. It is uncertain why these lots are
separated, but the specimens from lot NMNZ M.202237
display an eroded periostracum and more slender shells
(compare Figs 3D and 3E). Fischer-Piette (1950: 171) listed
one specimen (MNHN-IM-2000-2705) as the ‘holotype’,
which is the lectotype designation.
FURTHER SPECIMENS: NMNZ M.202235 (23 shells, Vietnam,
Red River, right bank between Cam Ðng and Làng Bo,
Messager leg.). Its label (not original) bears the manuscript
name Clausilia suturalis Bav & Dautz’, which was never
published. The specimens, however, belong in Oospira mairei.
margaritifera Bavay & Dautzenberg,
1909, Clausilia (Fig. 3F)
Clausilia margaritifera Bavay & Dautzenberg, 1909: 102, pl.
3, figs 13–14.
TYPE MATERIAL: paralectotypes NMNZ M.202243 (12
shells, Messager leg.).
LABEL: (not original) Clausilia margaritifera Bav. D. Lao Kay,
Tonkin. Coll. Messager. Ex Coll. Dautzenberg. From the
type series.’
TYPE LOCALITY: ‘Muong-Hum et Nat-Son’. Vietnam,
Mng Hum; Nht Sn.
CURRENT TAXONOMY: Accepted as Margaritiphaedusa
margaritifera (Bavay & Dautzenberg, 1909).
REMARKS: The label (not original) gives only Lào Cai
province as the locality, which includes the type localities.
Fischer-Piette (1950: 174) listed one specimen (MNHN-
IM-2000-2508) as the ‘holotype’, which is the lectotype
designation.
63
messageri Bavay & Dautzenberg,
1899, Clausilia (Fig. 3G)
Clausilia (Pseudonenia) Messageri Bavay & Dautzenberg,
1899a: 47, pl. 3, figs 1–1b.
TYPE MATERIAL: paralectotypes NMNZ M.202245 (2 shells,
Messager leg.).
LABEL: (not original) Clausilia messageri B. et D. That Khe,
Tonkin. Coll. Messager. Ex Coll. Dautzenberg. From the
type series.’
TYPE LOCALITY: ‘Entre Lang-Son et That-Khé’. Vietnam,
between Lng Sn and Tht Khê.
CURRENT TAXONOMY: Accepted as Tropidauchenia orientalis
messageri (Bavay & Dautzenberg, 1899).
REMARKS: The label (not original) gives only Tht Khê as
the locality. Fischer-Piette (1950: 167) listed one specimen
(MNHN-IM-2000-2640) as the ‘holotype’, which is the
lectotype designation.
moirati Bavay & Dautzenberg,
1909, Clausilia (Fig. 3H)
Clausilia Moirati Bavay & Dautzenberg, 1909: 100, pl. 3,
figs 10–11.
TYPE MATERIAL: paralectotype NMNZ M.202229 (1 shell,
Messager leg., ex A.R.J.B. Bavay colln.).
LABEL: (not original) Clausilia moirati Bav. et Dautz. Tong
Kin. Ex Coll. A. Bavay. From the type series.’
TYPE LOCALITY: ‘Pac-Kha, Muong-Bo et Binh-Lu’.
Vietnam, Bc Hà, Mng and Bình L.
CURRENT TAXONOMY: Accepted as Synprosphyma moirati
(Bavay & Dautzenberg, 1909).
REMARKS: Fischer-Piette (1950: 174) listed one specimen
(MNHN-IM-2000-2642) as the ‘holotype’, which is the
lectotype designation.
FURTHER SPECIMENS: NMNZ M.329559 (1 shell,
‘Banoumeoun, Upper Tonkin’, ex Rev. A.H. Cooke
colln.). The label (original) gives the manuscript name rudis
Bav. and Dautz.’ (genus Clausilia is supposedly implicit),
which was never published (Fig. 1C). The specimens,
however, belong in Synprosphyma moirati.
platyloma Mööllendorff, 1901,
Phaedusa gisota (Fig. 3I)
Phaedusa (Hemiphaedusa) gisota platyloma Möllendorff,
1901: 79.
TYPE MATERIAL: syntypes NMNZ M.306043 (2 shells, ex
Möllendorff colln.).
LABEL: (original) ‘GISOTA Bav. and Dautz. Var.
PLATYLOMA Mollff. Tongkin. Coll. Ex coll.
Dautzenberg, and von Mollendorf. [From the Type
series.]’
TYPE LOCALITY: ‘Than-moi’. Vietnam, Làng Thanh
(formerly Thanh Mi).
CURRENT TAXONOMY: Junior synonym of Notoptychia
polydona (Mabille, 1887).
REMARKS: The label (original) gives only Tonkin as the
locality.
porphyrostoma Bavay & Dautzenberg,
1909, Clausilia (Fig. 2A)
Clausilia porphyrostoma Bavay & Dautzenberg, 1909: 87, pl.
2, figs 5–6.
TYPE MATERIAL: paralectotypes NMNZ M.202225 (3 shells,
Vietnam, Bc Hà, Messager leg.).
LABEL: (not original) Clausilia porphyrostoma Bav. et Dautz.
Pac Kha, Tongkin. Coll. Lt. Col. Messager. Ex Coll. Ph.
Dautzenberg. From the type series.’
TYPE LOCALITY: ‘région de Lao-Kay’. Vietnam, Lào Cai
province.
CURRENT TAXONOMY: Accepted as Selenophaedusa
porphyrostoma (Bavay & Dautzenberg, 1909).
REMARKS: Fischer-Piette (1950: 175) listed one specimen
(MNHN-IM-2000-2661) as the ‘holotype’, which is the
lectotype designation; five further paralectotypes are in
RBINS (RBINS MT.2479).
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sykesi Bavay & Dautzenberg,
1899, Clausilia (Fig. 4A)
Clausilia Sykesi Bavay & Dautzenberg, 1899b: 293, pl. 12,
figs 12–13.
TYPE MATERIAL: paralectotypes NMNZ M.202244 (4 shells,
Messager leg.).
LABEL: (not original) Clausilia sykesi B. et D. Bac Kan,
Tonkin. Coll. Messager. Ex Coll. Dautzenberg. From the
type series.’
TYPE LOCALITY: ‘Bac-Kan’. Vietnam, Bc Kn.
CURRENT TAXONOMY: Accepted as Leptocochlea sykesi
(Bavay & Dautzenberg, 1899).
REMARKS: Fischer-Piette (1950: 167) listed one specimen
(MNHN-IM-2000-2684) as the ‘holotype’, which is the
lectotype designation.
tantilla Pilsbry, 1902, Clausilia (Fig. 4B)
Clausilia (Hemiphaedusa) tantilla Pilsbry, 1902: 655.
TYPE MATERIAL: paralectotypes NMNZ M.329558 (2 shells,
Y. Hirase leg.).
TYPE LOCALITY: ‘Goto, Uzen’. The type locality given by
Pilsbry (1902) is uncertain, as it could refer to: (1) the Gotö
Islands in Nagasaki prefecture; (2) Mt Unzen, also in Nagasaki
prefecture (near Shimabara) but not on Gotö islands; or (3)
Uzen province, an old province name in northern Japan,
now part of Yamagata prefecture. The actual locality can
be ascertained by looking into further material from Hirase
Conchological Museum, now housed at KIU. The label of
the lot of Clausilia tantilla (KIU 1730) has the locality noted
as 󲪧’, which is the Gotö Islands, Nagasaki prefecture
(S. Ishida, pers. comm., 15 January 2020).
CURRENT TAXONOMY: Accepted as Zaptyx tantilla (Pilsbry,
1902).
REMARKS: The lectotype (Baker 1963: 214) and a lot
with paralectotypes are deposited in the ANSP collection
under register numbers ANSP 83023 and ANSP 453959,
respectively. The present specimens are thus further
paralectotypes.
thatkheana Bavay & Dautzenberg,
1899, Clausilia (Fig. 4C)
Clausilia thatkheana Bavay & Dautzenberg, 1899b: 285, pl.
12, fig. 7.
TYPE MATERIAL: paralectotypes NMNZ M.202236 (2 shells,
Messager coll.).
LABEL: (not original) Clausilia thatkeana [sic] Bav. et Dautz.
That Khe, Tonkin. Coll. Messager. Ex Coll. Dautzenberg.
From the type series.’
TYPE LOCALITY: ‘Les environs de That-Khé’. Vietnam,
Tht Khê.
CURRENT TAXONOMY: Accepted as Selenophaedusa
thatkheana (Bavay & Dautzenberg, 1899).
REMARKS: Fischer-Piette (1950: 167) listed one specimen
(MNHN-IM-2000-2689) as the ‘holotype’, which is the
lectotype designation.
Fig. 4 Clausiliidae types from the NMNZ collection (cont.). A, paralectotype of Clausilia sykesi Bavay & Dautzenberg, 1899,
NMNZ M.202244; scale bar = 3 mm; B, paralectotype of C. tantilla Pilsbry, 1902, NMNZ M.329558; scale bar = 1 mm; C,
paralectotype of C. thatkheana Bavay & Dautzenberg, 1899, NMNZ M.202236; scale bar = 5 mm.
65
Concluding remarks
The present specimens from Dautzenberg’s collection
(including types of 19 taxa and possible type material of
one taxon) remained hidden for several decades within
the NMNZ collection cabinets. Not even the published
list of that institution’s molluscan type material (Marshall
1996) mentioned them. Although naturally focused on
New Zealand and the Pacific, the NMNZ collection has
a wealth of international land snails, mostly obtained in
the late nineteenth and early twentieth centuries. Even
though the purpose of this collection then was to include
representative material from all families and biogeographic
regions, there is still a wealth of information locked away
in it (e.g. Salvador 2019). We expect that further studies in
this collection (and similar collections in other museums)
will reveal more lost treasures, which need to be made
available to a broader audience through publication and
digitisation.
Acknowledgements
We are very grateful to Jean-Claude Stahl (NMNZ) for
the photographs of the specimens; Jennifer Twist, Bruce
A. Marshall and Martin Lewis (NMNZ) for their help
with old documents; Jozef Grego (Banská Bystrica) and
Miklós Szekeres (Budapest) for their help in identifying
some of the specimens bearing manuscript names; Thierry
Backeljau (RBINS) for information about the Dautzenberg
collection; So Ishida (Osaka Museum of Natural History)
and Paul Callomon (ANSP) for their help in clarifying the
type locality of Clausilia tantilla; Virginie Héros (MNHN)
for help in interpreting the type designations of Fischer-
Piette; and the two anonymous reviewers whose comments
and suggestions greatly helped us to improve this article.
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Bavay, A.R.J.B. and Dautzenberg, P. (1899a). Description
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Breure, A.S.H. and Ablett, J.D. (2016). The ‘Demange
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Conchyliologica 36: 1–9.
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Marginellidae described by Bavay, 1903–1922. The
Veliger 16(2): 207–215.
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Argentinian gastropods in the collection of the
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67
Dautzenbergiella duella (Mabille, 1887)
Clausilia duella Mabille, 1887a: 16.
SPECIMEN: NMNZ M.330021 (1 shell, Vietnam, Tonkin,
Messager leg.).
REMARKS: Despite the presence of type material of this
species in the NMNZ Dautzenberg collection (see above),
the present specimen lacks the note ‘From the Type
Series’.
Formosana pacifica (Gredler, 1884)
Clausilia (Phaedusa) pacifica Gredler, 1884: 274, pl. 19, fig. 7.
SPECIMENS: NMNZ M.330023 (2 shells, China, Hubei).
Grandinenia ardouiniana (Heude, 1885)
Clausilia ardouiniana Heude, 1885: 118, pl. 31, fig. 1.
SPECIMENS: NMNZ M.306041 (1 shell, Vietnam, HLong
Bay, Vathelet leg., ex Heude colln.), NMNZ M.306047
(2 shells, Vietnam, Tonkin Gulf, ‘Tung Nam’, Demange
leg.), NMNZ M.306048 (1 shell, Vietnam, Tonkin Gulf,
Demange leg.).
REMARKS: There is indication on the label (original) that
the specimen NMNZ M.306041 belonged to Heude’s
collection. However, neither the locality (H Long Bay, as
‘Along Bay’) nor the collector (Vathelet) given on the label
match those on the original species description ‘Ad rupes
sinus Tonquini’ and ‘Ardouin’, respectively (Heude 1885).
As such, there is little evidence to consider this specimen
a type. Johnson (1973) pointed out that paratypes of this
species are housed in the Smithsonian National Museum of
Natural History (Washington, DC, USA) and Museum of
Comparative Zoology (Cambridge, USA), but offered no
information about the holotype. Lots NMNZ M.306047
and NMNZ M.306048, however, did not belong to Heude’s
collection. Instead, they have only indications of belonging
to Dautzenberg collection and bear, respectively, the name
ardouiniana Heude Var. minor Bav. And Dautz.’ (likely
referring to ardouiniana var. minor Dautzenberg & Fischer,
1905) and the unpublished manuscript name ardouiniana
Heude Var. tenuis Bav. And Dautz.’. These specimens,
however, belong in the nominate Grandinenia ardouiniana.
Liparophaedusa auregani
(Bavay & Dautzenberg, 1904)
Clausilia Auregani Bavay & Dautzenberg, 1904: 213, pl. 9,
figs 9–11.
SPECIMEN: NMNZ M.202241 (1 shell, Messager leg.).
REMARKS: Bavay & Dautzenberg (1904) introduced this
taxon as a replacement name for Clausilia fargesiana Mabille,
1887 [non Heude, 1885]. The taxon is thus based on the
material that Mabille had at hand (collected by Balansa)
for the description. Fischer-Piette (1950: 171) listed one
specimen (MNHN-IM-2000-2414) as the ‘holotype’;
however, it was the specimen figured by Bavay &
Dautzenberg (1904). The present specimen therefore has
no type status.
Megalauchenia proctostoma (Mabille, 1889)
Clausilia proctostoma Mabille, 1889: 13.
SPECIMEN: NMNZ M.306042 (1 shell, Vietnam, Tan Phong
Island, Demange leg.).
REMARKS: Mabille (1889) described this taxon on the basis of
material collected by Balansa in ‘Tonkin’ (syntypes MNHN-
IM-2000-2662, MNHN-IM-2000-2663; Fischer-Piette
1950). The present material, therefore, does not belong
to the type series. Furthermore, the locality of the present
specimen is outside the species’ known distribution.
Notoptychia polydona (Mabille, 1887)
Clausilia polydona Mabille, 1887b: 114.
SPECIMENS: NMNZ M.330018 (2 shells, Vietnam, Tonkin,
ex H.C. Fulton colln.).
Oospira javana (L. Pfeiffer, 1841)
Clausilia javana L. Pfeiffer, 1841: 49.
SPECIMENS: NMNZ M.330022 (2 shells, Java, ex H.C.
Fulton colln.).
Appendix
The following specimens belong to the Dautzenberg collection in the NMNZ, but do not represent type material (despite
some of them having ‘From the type series’ written on their labels). They are pre ented below in a similar, albeit more
simplified, manner to the main text, arranged alphabetically by genus name.
Dautzenberg’s snail types in Te Papa’s collection
Tuhinga
,
Number 31 (2020)
68
Oospira vanbuensis
(Bavay & Dautzenberg, 1899)
Clausilia vanbuensis Bavay & Dautzenberg, 1899a: 38, pl. 2,
figs 1–1b.
SPECIMENS: NMNZ M.202238 (2 shells, Vietnam, Vanbu,
Messager leg.).
REMARKS: The original description mentioned R. Bavay as
the collector (Bavay & Dautzenberg 1899a: 39). The present
specimens, however, were collected by Messager, as were
most others. As such, the present specimens are from the
type locality (route from Sn La to ‘Van-Bu’) but probably
from a later date and, thus, are unlikely to be types.
Phaedusa bocki (Sykes, 1895)
Clausilia (Pseudonenia) bocki Sykes, 1895: 263, fig. 1.
SPECIMENS: NMNZ M.329560 (2 shells, China, SE
Yunnan, Carl Bock leg., ex Hugh C. Fulton colln.).
REMARKS: The label (original) mentions only that the
specimens belonged to Fulton’s collection, and does not
list Sykes. Furthermore, even though Sykes (1895) named
his new species after Bock, he does not mention him
specifically as the collector. Finally, the type locality given
by Sykes (1895) is more specific: ‘Mang-hao, or Mon-hao,
S.E. Yunnan’. As such, the present specimens are not likely
to be type material.
Phaedusa inanis (Bavay & Dautzenberg, 1909)
Clausilia inanis Bavay & Dautzenberg, 1909: 103, pl. 3, figs
15–16.
SPECIMEN: NMNZ M.330019 (1 shell, Vietnam, Lào Cai).
REMARKS: Despite the presence of type material of this
species in the NMNZ Dautzenberg collection (see above),
the present specimen lacks the note ‘From the Type series’.
Phaedusa paviei (Morlet, 1892)
Clausilia Paviei Morlet, 1892: 319.
SPECIMENS: NMNZ M.202226 (5 shells, Vietnam, Mng
Hum, Messager coll.); NMNZ M.202240 (3 shells,
Vietnam, Tht Khê, Messager leg.); NMNZ M.202248
(8 shells, Vietnam, Mng Hum, Messager leg.), NMNZ
M.330017 (1 shell, Vietnam, MSn, ex A. Bavay colln.).
REMARKS: The labels (both not original) of lot NMNZ
M.202240 and NMNZ M.202248, respectively, give the
manuscript names Clausilia grangeri pallida and Clausilia
tonkiniana Bav & Dautz’, which were never published. The
specimens, however, belong in Phaedusa paviei.
Selenophaedusa billeti (H. Fischer, 1898)
Clausilia billeti H. Fischer, 1898: 324, pl. 18, figs 28–31.
SPECIMENS: NMNZ M.330015 (2 shells, Vietnam, Tonkin,
ex H.C. Fulton colln.).
Synprosphyma acrostoma
(Bavay & Dautzenberg, 1909)
Clausilia acrostoma Bavay & Dautzenberg, 1909: 90, pl. 2,
figs 13–14.
SPECIMENS: NMNZ M.306049 (2 shells, Vietnam, Bc Hà,
Messager leg.).
REMARKS: The specimens are not from the type locality (Gia
Phú) and as such they are not type, despite the label bearing
a ‘From the Type series’ note. Fischer-Piette (1950: 174)
mentioned one specimen (MNHN-IM-2000-2401) as the
‘holotype’, which is the lectotype designation.
Synprosphyma suilla
(Bavay & Dautzenberg, 1909)
Clausilia suilla Bavay & Dautzenberg, 1909: 88, pl. 2, figs
7–8.
SPECIMENS: NMNZ M.306051 (5 shells, Vietnam, Trnh
Tng, Messager leg.), NMNZ M.330016 (1 shell,
Vietnam, Cam Ðng, ex A.H. Cooke colln.).
REMARKS: The specimens from lot NMNZ M.306051 are
not from the type localities (Mng and Bình L),
and are instead from
Trnh Tng, c. 200 km north of
Mg and c. 50 km north of Bình L. As such, they
do not represent type material. Fischer-Piette (1950: 174)
mentioned one specimen (MNHN-IM-2000-2701) as the
‘holotype’, which is the lectotype designation; five further
paralectotypes are in RBINS (RBINS MT.2476).
Tropidauchenia bavayi (Lindholm, 1924)
Clausilia bavayi Lindholm, 1924: 57.
SPECIMENS: NMNZ M.202232 (2 shells, Vietnam, Ch
B Messager leg.), NMNZ M.330011 (2 shells, Vietnam,
region of PhLý and KS, Tonkin, ex H. Rolle colln.),
NMNZ M.330012 (1 shell, Vietnam, mountains of Lake
Ba B, ex A.H. Cooke colln.), NMNZ M.330013 (4 shells,
Vietnam, mountains of Lake Ba B, ex A.H. Cooke colln.).
REMARKS: The label (not original) of lot NMNZ M.202232
indicates it is a type specimen. However, its locality
(ChB) does not match the species type locality (vicinity
of Lake Ba B; see above) and therefore it is not type
material.
69
Tropidauchenia giardi (H. Fischer, 1898)
Clausilia Giardi H. Fischer, 1898: 323, textfig. 2, pl. 18,
figs 22–27.
SPECIMENS: NMNZ M.202231 (6 shells, Vietnam, Lang Son,
Messager leg., ex H. Fischer colln.), NMNZ M.330014
(2 shells, Vietnam, Tht Khê, Messager leg., ex H. Rolle
colln.), NMNZ M.330020 (2 shells, Vietnam, Long Phú, ex
H. Rolle colln.).
REMARKS: The label (not original) states the specimens
belong to the type series and that they belonged to Fischer’s
collection. However, both the collector (Messager) and the
locality (Lang Son) noted on the label do not match the
original descriptions (Billet and Ðèo Ma Phúc respectively).
As such, the present specimens almost certainly do not
represent type material.
Tropidauchenia orientalis (Mabille, 1887)
Nenia orientalis Mabille, 1887a: 17.
SPECIMENS: NMNZ M.306050 (2 shells, Vietnam, Tonkin,
‘Tpan Mai’, ex A.R.J.B. Bavay colln.).
REMARKS: The label (original) mentions only that the
specimens belonged to Bavay’s collection and does not list
Mabille’s collection (as was the case for Dautzenbergiella
duella, above). As such, the present specimens are not likely
to be type material.
Dautzenberg’s snail types in Te Papa’s collection
... A small but important collection of land snails is part of the Mollusca collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ, Wellington, New Zealand), which has come to our attention during ongoing efforts to reorganize the museum's terrestrial and freshwater gastropod holdings (see also Salvador and Breure 2020). This material comprises thirty-nine lots of land snail shells from South Africa belonging to the family Streptaxidae, commonly known as hunter snails. ...
... Unfortunately, the original labels of Suter's specimens were discarded when Suter's collection was incorporated into the NMNZ's collection, but the information on them was transcribed. Suter kept careful notes regarding the provenance of the material and it is expected that the transcribed labels would bear the same information, as is the case for Dautzenberg's specimens at the NMNZ, for which some of the original labels remain (Salvador and Breure 2020). Similar to Dautzenberg's material, ten of Burnup's lots indicate that the specimens are paratypes. ...
... The present specimens from Burnup's collection include previously unrecognised type specimens of eight taxa. Further historical type material of terrestrial snails has also been recently re-discovered in the NMNZ collection (Salvador and Breure 2020), which although focused on New Zealand and Australasia, has a wealth of land snails from other parts of the world, a large part of which was obtained via Suter's collection or in earlier days of the then Dominion Museum. There is still much information locked away in this extralimital material (Salvador 2019), including surprising type specimens such as the present ones. ...
Article
Full-text available
A small collection containing thirty-nine lots of South African Streptaxidae land snails is housed in the collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ). This material previously belonged to British/South African malacologist Henry C. Burnup, who either donated it to, or exchanged it with New Zealand-based Swiss malacologist Henry Suter, whose land snail collection was eventually acquired by the NMNZ. The lots contain type specimens of eight taxa (species and subspecies) and are presented herein in the form of an annotated and illustrated catalogue.
... The ongoing reorganisation of the terrestrial gastropod collection has unveiled some hidden treasures, including type material (e.g. Salvador & Breure (2020); Salvador & Ablett (2020); Salvador (2021)) and specimens that add meaningful geographical data to our present knowledge (Salvador 2019). In this article, we examine the South American land snails in the NMNZ collection and present the compiled data below, excluding those from Brazil, Uruguay and Argentina, which were covered in Salvador (2019). ...
... The present specimens are paralectotypes of Nenia smithiaePilsbry, 1902 (originally spelled Nenia Smithiae) and were very likely acquired by Henry Suter directly from Henry Pilsbry, who described the species. Other lots acquired from Pilsbry are known in the NMNZ collection (e.g.Salvador & Breure (2020)). The lectotype and five other paralectotypes of N. smithiae are housed in the collection of the Academy of Natural Sciences, Drexel University, Philadelphia, USA, under register numbers ANSP 81272 and ANSP 461410, respectively. ...
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Despite its natural focus on the New Zealand region, the malacological collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ), also includes a variety of specimens from elsewhere in the world, including a fair share of South American specimens. Examination of this material reveals valuable distributional data for several species, as well as previously unreported type material. This article examines and reidentifies all South American (and Trinidadian) gastropods from the NMNZ collection, excluding those from Brazil, Uruguay and Argentina, which were analysed in a separate publication. All the information gathered is compiled and presented here as an annotated species checklist, including photographs of the most important specimens. In total, 91 species are reported. Paralectotypes of Neniops smithiae (Pilsbry, 1902) (from the private collection of zoologist Henry Suter (1841-1918)) were found in the NMNZ collection and are illustrated here.
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The malacological collection of the Museum of New Zealand Te Papa Tongarewa (NMNZ), despite naturally focusing on New Zealand species, also includes a variety of specimens from South America. Examination of this material revealed new distributional data for several species. All Brazilian, Uruguayan and Argentinian terrestrial gastropods from the NMNZ collection were examined and reidentified (no material from Paraguay was found). The information gathered was compiled and is presented in this article, and may contain significant data for malacologists working with the region's fauna. In summary, 99 species are reported, 13 of which represent new records and meaningful increments in geographical distribution, either extending their known range or filling distributional gaps. Moreover, the NMNZ collection houses the type material of six species from Brazil and Argentina described by the New Zealand malacologist Henry Suter (1841-1918) in 1900.
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Description de coquilles nouvelles de l'Indo-Chine (4e Suite)
  • A R J B Bavay
  • P Dautzenberg
Bavay, A.R.J.B. and Dautzenberg, P. (1909). Description de coquilles nouvelles de l'Indo-Chine (4e Suite). Journal de Conchyliologie 57(2): 81-105, pls 2-3.