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The present paper describes 16 new species and one new genus from French Guiana and numerous taxonomic changes are proposed to the Phasmatodea of French Guiana. Fifty-six new combinations, twenty-six new synonymies, two lectotypes and four neotypes are designated.

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... Mantis xanthomela was finally removed from the synonymy of Prexaspes lateralis by Otte & Brock (2005), who revalidated the species as Stratocles xanthomela. Recently, both species were placed in new combinations by Conle et al. (2020): Prexaspes lateralis was transferred to Paraphasma, which was corroborated by the phylogenetic hypothesis of Chiquetto-Machado & Cancello (2021), and Stratocles xanthomela was transferred to Parastratocles Redtenbacher, 1906. Here we solve one last taxonomic problem regarding these species. ...
... Phasma linearis Stoll, 1813: 77 [name for the species "Le Double-Epine Brun" of Stoll (1787: 24, fig. 27)]; Westwood, 1859: 147 [= Necroscia umbretta (Gray, 1835]; Kirby, 1904a: 415 [= Prexaspes umbretta (Gray, 1835]; Redtenbacher, 1906: 116 [= Paraphasma maculatum (Gray, 1835]; Otte & Brock, 2005: 251 [= Paraphasma maculatum (Gray, 1835]; Conle et al., 2011: 33 [= Paraphasma maculatum (Gray, 1835]; Conle et al., 2020: 126 [= Paraphasma maculatum (Gray, 1835]. Phasma lineare, Gray, 1835: 26;De Haan, 1842: 123. ...
... 77)]; Otte & Brock, 2005: 251 [= Paraphasma maculatum (Gray, 1835]; Conle et al., 2011: 33 [= Paraphasma maculatum (Gray, 1835]; Conle et al., 2020: 126 [= Paraphasma maculatum (Gray, 1835]. Unspecified type(s): ♀, Ambon Island [probably lost; locality probably wrong]. ...
Article
Paraphasma Redtenbacher, 1906 is a genus of fully-winged stick insects occurring in central and northern South America. We carried out a morphology-based taxonomic revision of this genus with emphasis on the phallic organ, a structure that has been poorly explored for taxonomic purposes in Phasmatodea. Additionally, pairwise genetic distances between mitochondrial COI gene sequences were calculated for ten Paraphasma specimens representing six species. We recognize nine valid species in the genus plus one nomen dubium, Paraphasma fasciatum Gray, 1835. We redescribe Paraphasma and the species previously assigned to it, describe Paraphasma indistinctum Chiquetto-Machado sp. nov., Paraphasma sooretama Chiquetto-Machado sp. nov. and Paraphasma spinicauda Chiquetto-Machado sp. nov., and provide a key to the species in the genus. The male of Paraphasma minus Redtenbacher, 1906 is described for the first time, as well as the eggs of six species. We transfer Paraphasma amabile Redtenbacher, 1906 to Pseudophasma Kirby, 1896 (comb. nov.) and synonymize Pseudophasma xanthotaenidium Günther, 1930 under this species (syn. nov.). In addition, Phasma perspicillaris Stoll, 1813 is removed from the synonymy of Paraphasma laterale (Fabricius, 1775) and synonymized under Parastratocles xanthomela (Olivier, 1792) (syn. nov.). The examination of the phallic organ was essential for species delimitation, as most species of Paraphasma are very similar in the external morphology of both sexes. The analysis of the COI sequences supported the species delimitation, resulting in remarkably lower pairwise distances between conspecific individuals (p-distance ≤ 2.0%) than between different species (p-distance 6.9–17.5%). We hope that this paper will stimulate further studies exploring the taxonomic and phylogenetic potential of the internal male genitalia of stick insects.
... Neotropical stick insects are generally poorly studied: many species are known only from their original descriptions and relatively few have been covered in recent taxonomic works (see, e.g., Gutiérrez-Valencia et al. 2017;Conle et al. 2020;Crispino et al. 2020;Madeira-Ott et al. 2020;Chiquetto-Machado & Cancello 2021;Ghirotto 2021;Chiquetto-Machado et al. 2022). Compared to the fauna of other continents, Neotropical phasmids are underrepresented in phylogenetic analyses, reflecting the lack of basic scientific work concerning them (Robertson et al. 2018;Simon et al. 2019). ...
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Two species of stick insect with a distinctive morphology, Candovia evoneobertii (Zompro & Adis, 2001) and Echetlus fulgens Zompro, 2004, were considered to be native to Australia and introduced into Brazil. However, Heteronemia dubia (Caudell, 1904) and Heteronemia fragilis (Brunner von Wattenwyl, 1907), both described more than a hundred years ago from South America, exhibit striking similarities with the two purportedly introduced species and are found to be conspecific with C. evoneobertii. Careful analysis of the literature and specimens revealed that these species belong to the Neotropical tribe Diapheromerini (Diapheromeridae) and represent a new genus, Arumatia Ghirotto gen. nov. We therefore propose Arumatia fulgens (Zompro, 2004) gen. et comb. nov. and Arumatia dubia (Caudell, 1904) gen. et comb. nov. We further redescribe A. dubia (Caudell, 1904) gen. et comb. nov. based on several specimens and synonymize Heteronemia fragilis syn. nov. and Candovia evoneobertii syn. nov. under it. Additionally, five new Brazilian species are described: Arumatia diamante Ghirotto gen. et sp. nov. from Abaíra, Bahia; Arumatia aramatia Ghirotto gen. et sp. nov. from Porto Nacional, Tocantins; Arumatia motenata Ghirotto gen. et sp. nov. from Serra do Cipó, Minas Gerais; Arumatia crassicercata Ghirotto, Crispino & Engelking gen. et sp. nov. from Alto Paraíso de Goiás, Goiás; and Arumatia anyami Ghirotto, Crispino & Neves gen. et sp. nov. from Costa Marques, Rondônia. Species of Arumatia gen. nov. occur mostly in the Cerrado domain, and represent the first Diapheromeridae recorded in this area. Most species are known exclusively from females with only A. aramatia gen. et sp. nov. and A. motenata gen. et sp. nov. known from both sexes. Adult and egg morphology are described and illustrated in detail for all species, as well as the nymph stages for A. dubia. Biological observations are presented, including parthenogeny in A. dubia and one of the few detailed accounts of sexual behaviour in Euphasmatodea (for A. motenata gen. et sp. nov.). Finally, a species of Diapheromerini described in error from Brazil, Diapheromera armata Piza, 1973, is synonymized under the North American Megaphasma denticrus (Stål, 1875) (syn. nov.).
... Three nights of prospections near Camp Caïman in the Kaw Mountains resulted in the capture of additional Paraprisopus specimens. Among them was an adult female of Paraprisopus apterus Conle, Hennemann, Bellanger, Lelong, Jourdan & Valero, 2020, a species until then only described from one male (holotype) and one juvenile female (paratype). This specimen was kept in breeding in order to obtain eggs. ...
Article
The egg and the adult female of Paraprisopus apterus Conle, Hennemann, Bellanger, Lelong, Jourdan & Valero, 2020, from French Guiana, are described here for the first time. Ant-mimicking morphology and behaviour of post-hatch Paraprisopus nymphs are reported and discussed.
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The genus Pterinoxylus Serville, 1838 is redescribed and revised at the species level. It is distributed throughout most of Central America, the northern half of South America and also has one species on the Lesser Antilles. Two new species are described from Costa Rica: P. cocoense n. sp. from both sexes and the eggs and P. speciosus n. sp. from both sexes. The female of P. perarmatus (Redtenbacher, 1908) is described and illustrated for the first time, as are the eggs of the type-species P. eucnemis Serville, 1838 and P. perarmatus (Redtenbacher, 1908).
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Four new species of the genus Phantasca Redtenbacher, 1906 (Phantasca bulbosa n. sp., P. arlequina n. sp., P. kawensis n. sp. and P. margaritae n. sp.) from French Guiana are described and illustrated. The female of P. femorata Hennemann, Conle, Bellanger, Lelong & Jourdan, 2018, is re-described due to subsequent discovery of an adult specimen, and the previously unknown egg is described for the first time. The male of P. ruboligata Hennemann et al., 2018, is re-described with the holotype illustrated to rule out confusion with the male of P. margaritae n. sp. A barcoding study based on COI gene of eight species of Phantasca is presented and supports their inclusion within the genus.
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Asymmetry is a common phenom among invertebrates. A consequence of this phenom is the rise of chiral forms (or enantiomorphs), which are structures that are mirror images of each other. The asymmetry can be divided into two: directional asymmetry, when only one of the enantiomorphs exists in nature, and antisymmetry, when both chiral forms exist in nature in equal frequencies, a much rarer phenom than the prior. In this paper, I describe the unknown male and redescribe the female of Isagoras paxillus (Westwood) and a new species of Isagoras Stål from the Brazilian Atlantic Forest, Isagoras sobrali sp. nov. (based on males from Rio Grande do Norte: Natal). Furthermore, I also describe the first case of antisymmetry in Phasmatodea in these two species and discuss the relevance of this finding within a morphological, behavioral, and systematic context.
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A detailed listing of this important collection, with notes on Brunner von Wattenwyl and Redtenbacher. 72 pages, paperback A5. OUT OF PRINT but can be downloaded here: https://www.zobodat.at/pdf/kat-nhmw_13_0003-0072.pdf
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The South American genus Phantasca Redtenbacher, 1906 (Phasmatodea: Diapheromeridae: Diapheromaerinae) is re-diagnosed and revised at the species level. The precedingly unknown eggs are described for the first time. The genus Pterolibethra Günther, 1940 (type species: P. heteronemia Günther, 1940) is re-synonymised, with Phantasca (syn. nov.) and consequently the two species originally contained, P. heteronemia Günther, 1940 and P. poeciloptera Günther, 1940, are transferred to Phantasca (comb. rev.). P. laeta Conle, Hennemann & Gutierréz, 2011 is not congeneric and is transferred to the genus Jeremiodes Hennemann & Conle, 2007 (Cladomorphinae: Cladomorphini; comb. nov.). Two species are removed from Bacteria Berthold, 1827 and transferred to Phantasca; these are B. quadrilobata Chopard, 1911 and B. montana Redtenbacher, 1906 (comb. nov.). Six new species are described: P. adiposa sp. nov., P. amabile sp. nov., P. femorata sp. nov., P. guianensis sp. nov., P. nigrolineata sp. nov. and P. ruboligata sp. nov. The male and egg of P. quadrilobata (Chopard, 1911) are described and illustrated for the first time. The genus now contains 13 species that are distributed throughout the northern half of South America. A key as well as detailed descriptions and illustrations are presented for all known species.
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Citation: Büscher TH, Gorb SN (2017) Subdivision of the neotropical Prisopodinae Brunner von Wattenwyl, 1893 based on features of tarsal attachment pads (Insecta, Phasmatodea). ZooKeys 645: 1–11. https://doi. Abstract The euplantulae of species from all five genera of the Prisopodinae Brunner von Wattenwyl, 1893 were examined using scanning electron microscopy with the aim to reveal the significance of attachment pads regarding their phylogenetic relationships. The split into the conventional two sister groups is supported by the two-lobed structure of the euplantulae with a smooth surface in the Prisopodini and a nubby surface microstructure in the Paraprisopodini. The two lineages are well distinguishable by this feature, as well as by the shape of the euplantulae themselves. The functional importance of the attachment pad surface features is discussed.
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The anareolate New World subfamily Cladomorphinae Bradley & Galil, 1977 is reviewed and keys to the six tribes currently included are presented; these are: Cladomorphini Bradley & Galil, 1977, Cladoxerini Karny, 1923, Cranidiini Günther, 1953, Pterinoxylini n. trib., Hesperophasmatini Bradley & Galil, 1977 and Haplopodini Günther, 1953 rev. stat.. New diagnoses are presented for all these tribes and possible relationships within Cladomorphinae are discusssed. Morphology of the genitalia and egg-structures indicate Cladomorphinae as presently treated to be polyphyletic. Two subordinate groups are recognized within present Cladomorphinae, which differ considerably in numerous morphological characters of the insects and eggs. The first group and here regarded as Cladomorphinae sensu stricto is formed by the mostly South American Cladomorphini + Cranidiini + Cladoxerini, while the second group is formed by the predominantly Caribbean Hesperophasmatini + Pterinoxylini n. trib. + Haplopodini. Members of the first group (= Cladomorphini sensu stricto) share the dorsally carinate basitarsus in which the two dorsal carinae are melted with another, increasingly elongated gonapophyses VIII of females which are noticeably longer than gonapophyses IX and lamellate as well as strongly displaced medioventral carina of the profemora. Cranidiini + Cladomorphini share the strongly elongated and filiform gonapophyses VIII and presence of gonoplacs in the females, specialized poculum of males and presence of a median line in the eggs. Cranidiini differs from all other tribes of Cladomorphinae by the entirely unarmed legs of both sexes, distinctly broadened and leaf-like body and prominent longitudinal keel of the mesosternum of females, prominently enlarged poculum and spinulose phallus of males as well as the conspicuous narrowing of the posteromedian gap of the internal micropylar plate of the eggs and noticeably separated median line. Cladomorphini is characteristic for the specialized vomer and poculum of males and distinct opercular structures of the eggs. Certain representatives of Cladomorphini indicate relationships to the "Phanocles-group" of Diapheromerinae: Diapheromerini, hence Cladomorphini as presently treated may be paraphyletic. The exclusively South American Cladoxerini (= Baculini n. syn.) differs from the other two tribes of Cladomorphinae sensu stricto by the distinctly serrate profemora of both sexes and conspicuously shortened antennae of females, which consist of less than 30 segments and are much shorter than the profemora in females. Genital morphology, such as the elongated gonapophyses VIII and presence of gonoplacs in females, as well as the lamellate medioventral carina of the profemora indicate close relation to Cladomorphini. Cranidiini appears to be the sister-taxon of Cladomorphini + Cladoxerini. The tribe Baculini Günther, 1953 is synonymised with Cladoxerini (n. syn.), on the basis that the type-genera of both tribes are congeneric, with Baculum Saussure, 1861 being a junior synonym of Cladoxerus St. Fargeau & Audinet-Serville, 1827 (n. syn.). The genus Tersomia Kirby, 1904 is removed from Hesperophasmatini and transferred to Cladoxerini. Wattenwylia Toledo Piza, 1938 is removed from Pachymorphinae: Gratidiini and transferred to Cladoxerini. A detailed new diagnosis is presented for Cranidiini along with a detailed differentiation and the tribe is shown to be monotypical, only containing its type-genus Cranidium Westwood, 1843. All Caribbean genera subsequently added to Cranidiini are removed and transferred to Haplopodini rev. stat.. The three tribes Hesperophasmatini + Pterinoxylini n. trib. + Haplopodini rev. stat. are closely related and might form a monophyletic clade within Cladomorphinae sensu lato. They differ from Cladomorphinae sensu stricto by the short gonapophyses VIII and reduced gonoplacs of females, unspecialized poculum of males and lack of a micropylar line in the eggs. Haplopodini Günther, 1953 is re-established (rev. stat.) and comprises almost exclusively Caribbean genera previously placed in Hesperophasmatini by Bradley & Galil (1977) or Cranidiini by Zompro, (2004). Aploploides Rehn & Hebard, 1938, Diapherodes Gray, 1835, Haplopus Burmeister, 1838 and Paracranidium Brock, 1998 were misplaced in Cranidiini and are transferred to Haplopodini. On the basis of numerous morphological characters of the insects and eggs Hesperophasmatini is removed from Pseudophasmatidae: Xerosomatinae and re-transferred to its previous position in the subfamily Cladomorphinae sensu lato. A detailed newdiagnosis of Hesperophasmatini is presented, but is only provisional since the true diversity is as yet only fractionally known. The lack of a gula distinguishes Hesperophasmatini from all other tribes. The genus Laciphorus Redtenbacher, 1908 is removed from Hesperophasmatini and transferred to Diapheromeridae: Diapheromerinae: Diapheromerini. The new tribe Pterinoxylini n. trib. is established to contain only the type-genus Pterinoxylus Audinet-Serville, 1838. It is closely related and perhaps the sister taxon of Hesperophasmatini, with which it shares the presence of rough sensory areas on the probasisternum and profurcasternum. It differs from Hesperophasmatini and Haplopodini by the presence of a tympanal region (= stridulatory organ) in the alae of females and the alveolar eggs, which possess peripheral opercular and polar structures. Haplopodini is likely to be the sister group of Pterinoxylini n. trib. + Hesperophasmatini. The tribe Haplopodini rev. stat. is revised at the species level and comprises eight almost exclusively Caribbean genera, four of which are newly described. All eight genera now contained in Haplopodini are described in detail, differentiated from their closest relatives and their relationships and systematic position within Haplopodini are discussed. Keys and maps showing their distributions are presented along with a discussion of the distributional patterns. Detailed descriptions, differential diagnoses, synonymic listings, illustrations, material listings and measurements are given of all 26 currently known species and subspecies of Haplopodini. Four new genera are described within Haplopodini. The monotypical Apteroplopus n. gen. (type-species: Dyme grosse-tuberculata Brunner v. Wattenwyl, 1907) from Honduras is the only taxon of the tribe represented in Central America. It is only known from the male which differs from all other genera by being entirely apterous. Cephaloplopus n. gen. (type-species: Cephaloplopus pulchellus n. sp.) and Parhaplopus n. gen. (type-species: Haplopus cubensis Saussure, 1868) occur only on Hispaniola and Cuba. Both are closely related to Haplopus Burmeister, 1838 but in addition to having noticeably different eggs, both genera differ from Haplopus in several morphological characters. The monotypical Venupherodes n. gen. (type-species: Platycrana venustula Audinet-Serville, 1838) is endemic to Cuba, and in females being apterous resembles the second exclusively Cuban genus Aploploides Rehn & Hebard, 1938. It however differs from all other members of Haplopodini by the laterally expanded mesonotum of females, which overlaps the mesopleurae, as well as the morphology of the eggs. Two species-groups are recognized within Diapherodes Gray, 1835. The gigantea species-group comprises the species from the Lesser Antilles, which are: D. angulata (Fabricius, 1793), Diapherodes dominicae (Rehn & Hebard, 1938), D. gigantea gigantea (Gmélin, 1789), D. gigantea saintluciae n. ssp. and Diapherodes martinicensis Lelong & Langlois, 2005. The three species of the jamaicensis species-group, which are D. achalus (Rehn, 1904), D. jamaicensis (Drury, 1773) and D. laevicollis Redtenbacher, 1906, are restricted to the two Greater Antillean islands Jamaica and Puerto Rico. Haplopus Burmeister, 1838 is the most widely distributed genus being represented on all islands of the Greater Antilles except Jamaica, and also in the Virgin Islands, Bahamas, Florida Keys, Dry Tortugas and as far southwest as the Cayman Islands and Swan Islands. Nine new species and one new subspecies are described: Cephaloplopus alope n. sp. and Haplopus sobrinus n. sp. from Cuba, Cephaloplopus euchlorus n. sp., Cephaloplopus laetus n. sp., Cephaloplopus pulchellus n. sp., Haplopus brachypterus n. sp., Haplopus intermedius n. sp. and Parhaplopus navarroi n. sp. from Hispaniola, Haplopus woodruffi n. sp. from Cayman Brac (Cayman Islands) and Diapherodes gigantea saintluciae n. ssp. from Saint Lucia. Seven of these are described from both sexes but Cephaloplopus alope n. sp. and Haplopus sobrinus n. sp. are only known from the females and Cephaloplopus laetus n. sp. only from the males. The previously unknown males of Diapherodes angulata (Fabricius, 1793), Diapherodes laevicollis Redtenbacher, 1908, Haplopus bicuspidatus de Haan, 1842 and Parhaplopus cubensis (Saussure, 1868) as well as the previously unknown female of Parhaplopus evadne (Westwood, 1859) n. comb. are described and illustrated for the first time. Descriptions and illustrations of the eggs of eleven species are presented: Cephaloplopus euchlorus n. sp., Cephaloplopus pulchellus n. sp., Diapherodes achalus (Rehn, 1904), Diapherodes dominicae (Rehn & Hebard, 1938), Diapherodes gigantea gigantea (Gmélin, 1789), Diapherodes martinicensis Lelong & Langlois, 2005, Diapherodes jamaicensis (Drury, 1773), Haplopus bicuspidatus de Haan, 1842, Haplopus micropterus St. Fargeau & Audinet-Serville, 1825, Parhaplopus navarroi n. sp. and Venupherodes venustula (Audinet-Seville, 1838) n. comb.. Type specimens of the newly described taxa are deposited in the collections of ANSP, NHMUK, IIBZ, FSCA, MCZC, MNHN and USNM. Six species are transferred to other genera (n. comb.): Bacteria grossetuberculata (Brunner v. Wattenwyl, 1907) to Apteroplopus n. gen.; Haplopus cubensis Saussure, 1868 and Haplopus evadne Westwood, 1859 to Parhaplopus n. gen.; Diapherodes venustula (Audinet-Serville, 1838) to Venupherodes n. gen.; Haplopus jamaicensis (Drury, 1773) and Haplopus achalus Rehn, 1904 to Diapherodes Gray, 1835. Mantis angulata Fabricius, 1793 and Diapherodes gigantea dominicae Rehn & Hebard, 1938 are removed from synonymy with D. gigantea (Gmélin, 1789) and shown to be valid species (n. stat.). Fifteen new synonymies are revealed amongst the species studied: Diapherodes longiscapha Redtenbacher, 1908 = Diapherodes achalus (Rehn, 1904) n. syn.; Haplopus grayi Kaup, 1871 = Diapherodes angulata (Fabricius, 1793) n. syn.; Diapherodes glabricollis Gray, 1835 = Diapherodes jamaicensis (Drury, 1773) n. syn.; Diapherodes pulverulentus Gray, 1835 = Diapherodes jamaicensis (Drury, 1773) n. syn.; Diapherodes christopheri Westwood, 1859 = Diapherodes jamaicensis (Drury, 1773) n. syn.; Haplopus murinus Redtenbacher, 1908 = Diapherodes jamaicensis (Drury, 1773) n. syn.; Haplopus bituberculatum de Haan, 1842 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn.; Haplopus cythereus Westwood, 1859 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn.; Haplopus ligiolus Redtenbacher, 1908 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn.; Haplopus ligia Westwood, 1859 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn.; Haplopus mayeri Caudell, 1905 = Haplopus scabricollis (Gray, 1835) n. syn.; Aplopus similis Rehn, 1904 = Haplopus scabricollis Gray, 1835 n. syn.; Diapherodes spinipes Gray, 1835 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn.; Haplopus obtusus Redtenbacher, 1908 = Haplopus micropterus (St. Fargeau & Audinet-Serville, 1825) n. syn. and Haplopus juvenis Redtenbacher, 1908 = Venupherodes venustula (Audinet-Serville, 1838) n. syn.. The previously presumed lost holotype of Cyprocrana microptera St. Fargeau & Audinet-Serville, 1825 (= Phasma angulata Stoll, 1813), was traced in the collection of RMNH. The designation of a neotype has become necessary for Mantis angulata Fabricius, 1793 (in MNCN) and Platycrana venustula Audinet-Serville, 1838 (in MNHU). Lectotypes are designated for eight species: Diapherodes longiscapha Redtenbacher, 1908; Diapherodes scabricollis Gray, 1835; Haplopus christopheri Westwood, 1859; Haplopus cytherea Westwood, 1859; Haplopus juvenis Redtenbacher, 1908; Haplopus ligiolus Redtenbacher, 1908; Haplopus ligia Westwood, 1859 and Haplopus murinus Redtenbacher, 1908.
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L'association ASPER a été missionnée par le Parc Amazonien de Guyane pour réaliser l'inventaire des Phasmatodea de Saül, village isolé au centre de la Guyane. Le matériel et la méthode utilisés sont détaillés et les espèces observées sont listées. Au total ce sont 32 espèces de 21 genres différents qui ont été inventoriées. Abstract. – Contribution to the inventory of Phasmatodea of Saül, French Guiana. The organization ASPER has been engaged by the Parc Amazonien de Guyane to make the inventory of Phasmatodea of Saül, isolated village in the center of French Guiana. Material and method are detailed. A total of 32 species were observed, belonging to 21 different genera.
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Une liste des Phasmatodea de Trinidad a pu être établie à la suite d'une mission d'inventaire réalisée en 2010. Elle montre que cette île abrite au moins sept espèces appartenant à cet ordre. Deux nouvelles espèces sont décrites, dont une d'un nouveau genre : Apteroxylus chaguaramalensis n. gen., n. sp., et Clonistria caputaurata n. sp. La femelle de Caribbiopheromera trinitatis est redécrite et illustrée suite à la découverte de nouvelles formes. Les adultes et l'oeuf de Paraphanocles keratosqueleton sont redécrits suite à la découverte de variations intraspécifiques, et une nouvelle synonymie est établie avec Bacteria mutica. Quelques informations sur la biologie sont données et des clés d'identification des adultes des deux sexes et des oeufs sont proposées. Abstract. – Contribution to the inventory and biology of the Phasmatodea of Trinidad. A list of the Phasmatodea of Trinidad has been established following an inventory made in 2010. This island houses at least seven species of this order. Two new species are described, one of them belongs to a new genus: Apteroxylus chaguaramalensis n. gen., n. sp., and Clonistria caputaurata n. sp. The female of Caribbiopheromera trinitatis is redescribed and illustrated after the discovery of new forms. Adults and egg of Paraphanocles keratosqueleton are redescribed after the discovery of intraspecific variations, and a new synonymy is established with Bacteria mutica. Some information about biology is given and identification keys for adults of both sexes and eggs are proposed.
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The type material of Phasmatodea deposited in Brazilian museums and institutions is listed for the first time. New synonyms are proposed: Phibalosoma paulense Toledo Piza, 1938, Phibalosoma rochai Toledo Piza, 1938, Bacteria tuberculata Toledo Piza, 1938 and Bacteria tuberculata var. argentina Toledo Piza, 1938 are junior synonyms of Cladomorphus phyllinus (Gray, 1835). Nineteen new combinations are established.O material-tipo de Phasmatodea depositado em museus e instituições brasileiras é listado pela primeira vez. Novas sinonímias são propostas: Phibalosoma paulense Toledo Piza, 1938, Phibalosoma rochai Toledo Piza, 1938, Bacteria tuberculata Toledo Piza, 1938 e Bacteria tuberculata var. argentina Toledo Piza, 1938 são sinônimos júnior de Cladomorphus phyllinus (Gray, 1835). Dezenove novas combinações são estabelecidas.
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A key to the anareolate stick-insect genera (Insecta: Phasmatodea) of the New World (North, Central and South America) is provided. Otocraniella flagelloantennata gen. n. sp. n. and Echetlus fulgens n. sp. are described as new. A new genus, Aplopocranidium gen. n., is erected for Bacteria waehneri (Günther, 1940). Baculum ramosum (Saussure, 1861) is redescribed. New synonyms have been traced during the works on this study. Hypocyrtus (Redtenbacher, 1908) is a subjective junior synonym of Lamponius (Stål, 1875), Steleoxiphus (Rehn, 1907) of Paraleptynia (Caudell, 1904), and both Abrachia (Kirby, 1889) and Ceratiscus (Caudell, 1904) of Baculum (Saussure, 1861). The anareolate tribe Hesperophasmatini is recognized as a member of the Pseudophasmatidae: Xerosomatinae. A new tribe, Paraleptyniini trib n., is introduced to encompass the anareolatae genera Paraleptynia (Caudell, 1904), Xiphophasma (Rehn, 1913) and Parabacillus (Caudell, 1903). A study of egg material showed these taxa to belong into Heteronemiidae.
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A taxonomic survey of the anareolate neotropical tribe Cladomorphini Brunner v. Wattenwyl, 1893 (subfamily Cladomorphinae) is provided, along with a redescription and keys to the seven genera now contained. Two genera previously only known from the females, are redescribed and revised at the species level. The monotypical Aplopocranidium Zompro, 2004 (Type-species: Bacteria waehneri Günther, 1940) was misplaced in Cranidiini Günther, 1953 and is here transferred to the tribe Cladomorphini Brunner v. Wattenwyl, 1893. It is closely related to Jeremia Redtenbacher, 1908 and Jeremiodes Hennemann & Conle, 2007, but differs from both by the prominent spines of the thorax. The previously unknown male is described and illustrated for the first time and a new extended generic description provided. The genus Jeremia Redtenbacher, 1908 (Type-species: Jeremia grossedentata Redtenbacher, 1908) is redescribed and a key provided to distinguish the two species contained (J. grossedentata Redtenbacher, 1908 and J. gymnota Günther, 1930). The previously unknown male and eggs are described and illustrated for the first time. Jeremia is closely related to Jeremiodes and Aplopocranidium, but differs by the distinct ventral teeth of the middle and hind legs.
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Khamul n. gen., a distinctive eurytomid in the subfamily Eurytominae is described from the Neotropics based upon the type species, K. erwini, n. sp. A hypothesis of its phylogenetic placement within Eurytominae is presented, and four new species are described: K. erwini, K. gothmogi, K. lanceolatus, and K. tolkeini. Diagnostic features are included to dis-tincguish this taxon from other eurytomines and a key to species presented. Its biology is unknown, but label data indi-cate walking stick eggs (Prisopus sp.; Phasmatodea: Prisopodidae) as a possible host.
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Type-species of several genera of the insect order Phasmatodea, described by Redtenbacher, are designated. The genera are: Agrostia, Anisacantha, Antongilia, Brizoides, Citrina, Eucles, Euphasma, Hypocyrtus, Mirophasma, Neophasma, Paraphasma, Paraprisopus, Perliodes
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A generic revision of the insect order Phasmatodea: The New World genera of the stick insect subfamily Diapheromerinae (Diapheromeridae) = Heteronemiinae (Heteronemiidae) sensu Bradley & Galil, 1977. The North and South American genera of the phasmatodean subfamily Diapheromerinae = Heteronemiinae sensu Bradley & Galil are revised. All genera are redescribed, type species are mentioned or designated, and synonyms listed. Two new tribes, Ocnophilini and Oreophoetini, are established, and the Libethrini are synonymized with Diapheromerini. Genitalia, eggs and important other characters of most genera are figured. Ten new genera are introduced and five new species are described. Previously unknown males, females and eggs of several species are described. The genera are arranged in groups, with separate keys to all tribes and generic-level groups, including males, females, and eggs to the extent currently known
Article
The taxonomy of Dinelytron Gray (Prisopodidae: Prisopodinae) is controversially discussed in the literature, as well as the delimitation between Prisopodini genera. The phylogenetic relationships between Prisopodini genera were never studied. Attempting to resolve these problems, Dinelytron was reviewed and a phylogeny was conducted to test the monophyly of the genus and better delimitate the limits of the tribe genera. Resulting, Prisopoides gen. nov., with four species was described. In Dinelytron, one species is redescribed and five new Brazilian species are described. Dinelytron unilineatus (Redtenbacher) comb. nov. was transferred from Damasippus Stål. New diagnosis for Damasippus and Prisopus are proposed, based on external characters and of the male genitalia. The phylogeny recovered two most parsimonious trees, both supporting the taxonomic changes proposed in the present study. Prisopodini and each of its genera are shown to be monophyletic. Novelties on the male genitalia are described and discussed, as well as their impact over Phasmatodea systematics.
Article
Isagoras aurocaudata sp. nov. is being described from two female specimens from the States of Minas Gerais, Brazil. Diagnostic characters for the new species are the yellowish compound eye with brown spots, the yellowish spot at basal third of tegmina and the yellowish abdomen segments 8–11. Furthermore, Isagoras aurocaudata sp. nov. is compared to species of Isagoras Stål and Planudes Stål.
Article
Type specimens of 75 taxa of Phasmida have been identified in the Nationaal Natuurhistorisch Museum, Leiden. The taxa are listed alphabetically by species, with the number of specimens, sex and locality data. The material includes species described by Bragg, Bragg & Chan, Gunther, de Haan, Kirby, Lichtenstein, Olivier, and Stoll. The type of pin used was found to be particularly useful as an aid to identification of older material. There are a number of omissions and discrepancies in the published data, these are clarified. Some background information on the original publications and the described material is included.
Article
A catalogue of the type-material of the insect order Phasmatodea housed in the Museum für Naturkunde der Humboldt-Universität, Berlin, Germany, and in the collection of the Institut für Zoologie der Martin-Luther-Universität in Halle (Saale), Germany is published. The collection in Berlin contains types of more than 280 species and is especially strong in material of New Guinea, described by Bragg, Brunner v. Wattenwyl, Charpentier, Fritzsche, Gerstäcker, Günther, Hennemann, Karsch, Pictet, Redtenbacher, Rehn, Schaum, Sjöstedt, Westwood and Zompro. The catalogue includes the specimens from Baltic amber described by Pictet & Berendt (1854), housed in the Palaeontological Institute of the museum. The collection in Halle contains types of eight species, which are all described by Burmeister. A lectotype is designated for Cuniculina obnoxia Brunner v. W., 1907. New synonyms are: Cuniculina modesta Brunner v. W., 1907, of Clitumnus lobipes Brunner v. W., 1907, and Parapachymorpha quadrispinosa Hennemann, Gehler & Conle, 1995 of Clitumnus spiniger Brunner v. W., 1907. Two new African genera and species of Phasmatodea: Bacillidae: Antongilliinae, Ulugurucharax uluguruensis n. gen. n. sp. and Tuberculatocharax fritzsicki n. gen. n. sp. are described for the first time. Tuberculatocharax n. gen. is the only genus of Tuberculatocharacini n. trib., which is characterized by the reduction of the profemoral area apicalis. Beside two genera of Bacillini (Bacilloidea: Bacillidae), this is the only case known in Phasmatodea. (© 2005 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim)
Article
An attempt is made to standardize further the descriptive terminology of the phasmid egg capsule by introducing stricter definitions and standard abbreviations. In addition, the various forms of the internal micropylar plate are categorized. Eophasma Sellick is replaced by Eophasmodes nov.n. A key to 131 generic forms of these eggs is provided. Where more than one egg form is associated with a genus, a diagnosis of the subgroups is provided.
Article
The type material in the Zoologisches Museum der Universität Hamburg, Germany (ZMH) is listed. The collection contains type specimens of 113 species, mainly described by BRUNNER V. WATTENWYL, GÜNTHER and REDTENBACHER. The collection is especially strong in material from Borneo, Costa Rica, Fiji, Madagascar and Brazil. Lectotypes are designated for Creoxylus hagani REDTENBACHER, 1906, Xerosoma michaelis REDTENBACHER, 1906, Bacteria tenella REDTENBACHER, 1908 and Leprocaulus insularis verrucifer GÜNTHER, 1934
  • Redtenbacher Isagoras Brevipes
Isagoras brevipes Redtenbacher, 1906: 134 [Brazil: Espírito Santo] Isagoras brunni (Redtenbacher, 1906: 132) n. comb. [Venezuela] Isagoras cortex (Hebard, 1919: 155), pl. 22:4 n. comb. [Colombia] Isagoras crenulipes (Rehn, 1904: 100) n. comb. [Costa Rica] Isagoras dentipes Redtenbacher, 1906: 134 [Panama]
Isagoras funestus (Redtenbacher, 1906: 132) n. comb. [Colombia] Isagoras glyptomerion (Rehn, 1904): 101 n. comb
  • Hebard Isagoras Ecuadoricus
Isagoras ecuadoricus Hebard, 1933: 37 [Ecuador] Isagoras funestus (Redtenbacher, 1906: 132) n. comb. [Colombia] Isagoras glyptomerion (Rehn, 1904): 101 n. comb. [Panama, Costa Rica, Ecuador, Colombia].
Amazonas] Isagoras taeniatus (Piza, 1944: 43) n. comb. [Brazil: São Paulo] Isagoras venezuelae Rehn, 1947: 6 [Venezuela] Isagoras vignieri
  • Günther Isagoras Tacanae
Isagoras tacanae Günther, 1940: 494 [Colombia, Peru, Brazil: Amazonas] Isagoras taeniatus (Piza, 1944: 43) n. comb. [Brazil: São Paulo] Isagoras venezuelae Rehn, 1947: 6 [Venezuela] Isagoras vignieri (Redtenbacher, 1906): 144 [Panama]
Stéphane Brûlé 14.02
  • French Guiana
FRENCH GUIANA: 1 ♀: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., leg. Stéphane Brûlé 14.02.2010 [coll. OC, No. 0280-1]; 1 ♀: Französisch Guyana: Commune de Saül, Belvédère de Saül, 3°37'22"N-53°12'57"W, 326m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 26.08.2011 [coll. OC, No. 0280-2]; 1 ♀: Französisch Guyana: Commune de Saül, Aussichtspunkt Belvédère de Saül, 3°37'22"N-53°12'57"W, 326m, S.E.A.G., leg. Stéphane Brûlé 23.09.2011 [coll. OC, No. 0280-3];
1 ♂: Französisch Guyana: Commune de Roura
  • No Oc
OC, No. 0251-39];. 2 ♂♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 29.04.2017 [coll. OC, No. 0251-40, 0251-45];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 03.06.2017 [coll. OC, No. 0251-42];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg.
Lichtfang, leg. Stéphane Brûlé 02.08
  • No Oc
OC, No. 0251-44];. 1 ♂: Französisch Guyana: Commune de Saül, Aussichtspunkt Belvédère de Saül, 3°37'22" N-53°12'57" W, 326m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 02.08.2017 [coll. OC, No. 0251-41];. 1 ♂: Französisch Guyana: Communde de Montsinery, Savane Lambert, Savane littorale, N 4°53'05,09" W 52°31'04,70", S.E.A.G, Lichtfang, leg. Stéphane Brûlé 09.07.2016 [coll. OC, No. 0251-37]; 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Canopy UV light trap, leg.
Lichtfang, leg. Stéphane Brûlé 19.12.2015 [coll. OC, No. 0251-28];. 1 ♂: Französisch Guyana: Commune de Saül, Aussichtspunkt Belvédère de Saül, 3°37'22" N-53°12'57" W, 326m
  • S E A G Lichtfang
Stéphane Brûlé 25.07.2015 [coll. OC, No. 0251-26];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 17.10.2015 [coll. OC 0251-27];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 19.12.2015 [coll. OC, No. 0251-28];. 1 ♂: Französisch Guyana: Commune de Saül, Aussichtspunkt Belvédère de Saül, 3°37'22" N-53°12'57" W, 326m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 01.09.2015 [coll. OC, No. 0251-29];. 2 ♂♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 18.06.2016 [coll. OC, No. 0251-30, 0251-31];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 13.02.2016 [coll. OC, No. 0251-32];. 2 ♂♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 16.07.2016 [coll. OC, No. 0251-33, 0251-34];. 2 ♂♂♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg. Stéphane Brûlé 13.08.2016 [coll. OC, No. 0251-35, 0251-36];. 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, RN2 PK22, 4°44'56"N-52°26'28"W, 75m, S.E.A.G., Lichtfang, leg.
Automatic light trap blue, leg. Stéphane Brûlé 23.12.2017 [coll. OC, No. 0287-56]; 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, 4°44'31"N-52°25'53"W, 90m
  • Französisch Guyana
1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, 4°44'31"N-52°25'53"W, 90m, S.E.A.G., Automatic light trap blue, leg. Stéphane Brûlé 23.12.2017 [coll. OC, No. 0287-56]; 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, 4°44'31"N-52°25'53"W, 90m, S.E.A.G., Automatic light trap blue, leg. Stéphane Brûlé 18.08.2018 [coll. OC, No. 0287-57]; 1 ♂: Französisch Guyana: Commune de Roura, Montagne des Chevaux, 4°44'31"N-52°25'53"W, 90m, S.E.A.G., Automatic light trap blue, leg. Stéphane Brûlé 18.08.2018 [coll. OC, No.
Isagoras obscurum (Guérin-Méneville, 1838: 72) rev. stat. Isagoras similis n. sp. Metriophasma (Metriophasma) Uvarov
  • Isagoras Stål
Isagoras Stål, 1875 Isagoras obscurum (Guérin-Méneville, 1838: 72) rev. stat. Isagoras similis n. sp. Metriophasma (Metriophasma) Uvarov, 1940
removed from Bacteria Berthold, 1827 and transferred to Phanocloidea Zompro
  • Redtenbacher Bacteria Pallidenotata
Bacteria pallidenotata Redtenbacher, 1908: 415, removed from Bacteria Berthold, 1827 and transferred to Phanocloidea Zompro, 2001 [Valid name: Phanocloidea pallidenotata (Redtenbacher, 1908) (n. comb.) → 4.2]
removed from Perliodes Redtenbacher, 1906 and transferred to Agrostia Redtenbacher
  • Redtenbacher Perliodes Nigrogranulosus
Perliodes nigrogranulosus Redtenbacher, 1906, removed from Perliodes Redtenbacher, 1906 and transferred to Agrostia Redtenbacher, 1906 [Valid name: Agrostia nigrogranulosa (Redtenbacher, 1906) (n. comb.) → 5.3.1]
1859: 164, is removed from Dinelytron Gray, 1835 and transferred to Paraprisopus Redtenbacher
  • Westwood Dinelytron Agrion
Dinelytron agrion westwood, 1859: 164, is removed from Dinelytron Gray, 1835 and transferred to Paraprisopus Redtenbacher, 1906 [Valid name: Paraprisopus agrion (westwood, 1859) (n. comb.) → 5.2.1]
341, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher
  • Chopard Isagoras Affinis
Isagoras affinis Chopard, 1911: 341, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher, 1906 [Valid name: Tenerella affinis (Chopard, 1911) (n. comb.) → 5.3.2]
164, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher
  • Hebard Isagoras Chocoensis
Isagoras chocoensis Hebard, 1921: 164, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher, 1906 [Valid name: Tenerella chocoensis (Hebard, 1921) (n. comb.) → 5.3.2]
17, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher
  • Isagoras Schraderi Rehn
Isagoras schraderi Rehn, 1947: 17, is removed from Isagoras Chopard, 1911 and transferred to Tenerella Redtenbacher, 1906 [Valid name: Tenerella affinis (Rehn, 1947) (n. comb.) → 5.3.2]