ArticlePDF Available

DATA ON 3 NEW AND 6 RARE GELECHIIDAE MOTH SPECIES (LEPIDOPTERA: GELECHIIDAE) FOR LITHUANIA BRIGITA PAULAVIČIŪTĖ

Authors:
  • Kaunas T.Ivanauskas zoological museum, Lithuania, Kaunas

Abstract

This article presents data on 3 new (Metzneria aestivella, Syncopacma larseniella, Acompsia schmidtiellus) and 6 rare for the Lithuanian fauna species of Gelechiidae. Species were collected from 9 administrative districts and municipalities of Lithuania.
68 BULLETIN OF THE LITHUANIAN ENTOMOLOGICAL SOCIETY. Volume 3 (31)
DATA ON 3 NEW AND 6 RARE GELECHIIDAE MOTH SPECIES
(LEPIDOPTERA: GELECHIIDAE) FOR LITHUANIA
BRIGITA PAULAVIČIŪTĖ
Kaunas T. Ivanauskas Museum of Zoology, Laisvės al. 106, LT-44253, Kaunas, Lithuania.
E-mail: brigita.paulaviciute@zoomuziejus.lt
Introduction
The Gelechiidae (Lepidoptera: Gelechioidea) include species that are pests of
agriculture and forestry as well as species of biological and ecological interest. It is one
of the largest families of microlepidoptera and includes more than 4600 described
species in the world referred to about 500 genera in four subfamilies (Hodges, 1999).
About 1500 species occur in the Palearctic Region (Piskunov, 1990), and about 690
species occur in the Nearctic Region, although only 30% of the species in the latter are
estimated to be described (Hodges, 1983, 1999). A large number of species are also
present in the Oriental, Neotropical, Afrotropical, and Australian Regions (Becker, 1984;
Common, 1990; Heppner, 1991; Nielsen & Common 1991; Park, 1992). Gelechiidae is
one of the least known families of micromoths in Europe, with onlymore than 600
species (Huemer & Karsholt, 1999). The last catalogue of Lithuanian Lepidoptera
includes 163 species of this family (Ivinskis & Rimšaitė, 2018).
This article presents data on 3 new (Metzneria aestivella, Syncopacma larseniella,
Acompsia schmidtiellus) and 6 rare for the Lithuanian fauna species of Gelechiidae.
Species were collected from 9 administrative districts and municipalities of Lithuania.
Material and Methods
The material was collected in different parts of Lithuania: in Kazlų Rūda and Neringa
municipalities, Jonava, Kaunas, Tauragė, Varėna and Vilnius administrative districts.
Insects were collected using an entomological net and light trapping at night (160W–
500W DRL type bulb lamp was used). The species were collected by Brigita
Paulavičiūtė (B.P.), Giedrius Švitra (G.Š.), Vaida Paznekaitė (V.P.) and Vytautas
Inokaitis (V.I.). The material was identified by Brigita Paulavičiūtė.
The geographical coordinates were measured using a Garmin eTrex GPS receiver.
The material is deposited in the collection of the Kaunas T. Ivanauskas Museum of
Zoology. Species were identified using the following sources: Huemer & Karsholt
(1999, 2010, 2013), Elsner et al. (1999). Rarity categories have been established
according to the checklist of Lepidoptera of Lithuania (Ivinskis, 2004; Ivinskis &
Rimšaitė, 2018).
LIETUVOS ENTOMOLOGŲ DRAUGIJOS DARBAI. 3 (31) tomas 69
List of localities
Localit
y
Administra
t
ive district Coordinates (LAT, LONG)
Bradeliškės Vilnius district 54.82452, 24.94517
Braziūkai Kaunas district 54.907783, 23.474718
Jovariškės Trakai district 54.65241, 24.87764
Juodkrantė Neringa mun. 55.545957, 21.119798
Katra Varėna district 53.992694, 24.586032
Lokėnėlių Miškas f. Jonava district 55.09546, 24.37448
Lomankos Miškas f. Kaunas district 54.963343, 23.769996
Runkių Miškas f. Kazlų Rūda mun. 54.812257, 23.517778
Viešvilė Nat. R. Tauragė district 55.179949, 22.461193
List of species
GELECHIIDAE
*Metzneria aestivella (Zeller, 1839)
Runkių Miškas f., 01 07 2016, 1♂ (V.I.).
Monochroa elongella (Heinemann, 1870)
Bradeliškės, 25 07 2015, 1♂ (G.Š.).
Eulamprotes atrella (Denis & Schiffermüller, 1775)
Jovariškės, 12 05 2018, 1♂ (G.Š.).
Bryotropha desertella (Douglas, 1850)
Katra, 12 05 2018, 2♂ (G.Š.).
Bryotropha galbanella (Zeller, 1839)
Juodkrantė, 14 07 2004, 1 spec. (B.P.); Braziūkai 30 06 2009, 1♂ (B.P.).
Bryotropha basaltinella (Zeller, 1839)
Katra, 05 08 2018, 2♀ (G.Š.).
Gelechia muscosella Zeller, 1839
Lomankos Miškas f., 30 07 2018, 1♂ (B.P.).
*Syncopacma larseniella Gozmány, 1957
Viešvilė Nat. R., 28 07 2008, 1♂ (B.P.).
*Acompsia schmidtiellus (Heyden, 1848)
Lokėnėlių Miškas f., 07 07 2017, 1♂ (V.P.).
Discussion
Gelechiidae moths are not comprehensively investigated in Lithuania. New species
are published almost every year.
Metzneria aestivella (Zeller, 1839) is spread widely from the Canary Islands
throughout Europe to Central Asia (Elsner et al., 1999). This species already is known in
many European countries: Austria, Belarus, Belgium, Bosnia Herzegovina, Bulgaria,
Croatia, Cyprus, The Czech Republic, Denmark, Estonia, Finland, France, Germany,
Greece, Hungary, Ireland, Italy, Crete, Latvia, Macedonia, Malta, Poland, Portugal,
Romania, Russia, Sardinia, Serbia, Sicily, Slovakia, Slovenia, Spain, Sweden,
Switzerland, the Netherlands (Karsholt & Nieukerken, 2013; Aarvik et al., 2017). The
moth wingspan is usually 11-14 mm (Fig. 1). Adults are very similar to M. lappella and
70 BULLETIN OF THE LITHUANIAN ENTOMOLOGICAL SOCIETY. Volume 3 (31)
M. metzneriella. M. aestivella differs with lack the black discal spot on the forewing
(Solly, 2019). Larva feeds on Carlina spp. and Tanacetum corybosum (Patočka &
Turčáni, 2005). The species is local and the adults fly during June and July (Elsner et al.,
1999).
Fig. 1. Metzneria aestivella (Zeller, 1839) (Runkių Miškas f., 01 07 2016, ♂ V.
Inokaitis).
Fig. 2. Syncopacma larseniella Gozmány,
1957 (Viešvilė Nat. R. (2), 28 07 2008, ♂
B. Paulavičiūtė).
Fig. 3. Acompsia schmidtiellus (Heyden,
1848) (Lokėnėlių Miškas f., 07 07 2017, ♂
V. Paznekaitė)
Genus Syncopacma contains several species which are virtually impossible to
distinguish except by genitalia dissection. Syncopacma larseniella Gozmány, 1957 is
mostly distributed in north-west and Central Europe: Austria, Belgium, Britain, Russia,
the Czech Republic, Denmark, France, Germany, Luxemburg, Poland, Portugal,
Romania, Slovakia, Spain, Switzerland, The Netherlands (Elsner et al., 1999; Karsholt &
Nieukerken, 2013; Baraniak et al., 2014; Aarvik et al., 2017). Moths of this species
LIETUVOS ENTOMOLOGŲ DRAUGIJOS DARBAI. 3 (31) tomas 71
show a less distinct pale fascia on the forewing underside and lack the pale spot on the
hindwing underside (Fig. 2) (Lewis, 2016). Wingspan of S. larseniella usually is 11–14
mm. Larvae feed on Lotus corniculatus in a spinning between leaves. The adults fly
during June and July (Mackay, 2019).
Acompsia schmidtiellus (Heyden, 1848) is found locally in central, eastern and
southern Europe: Austria, Belgium, Britain, the Czech Republic, Denmark, France,
Germany, Italy, Luxemburg, Portugal, Russia, Sardinia, Slovakia, Spain, Switzerland,
the Netherlands. In the east, the range extends to Ukraine (Karsholt & Nieukerken, 2013;
Aarvik et al., 2017). Wingspan of the moths is usually 14–16 mm (Fig. 3). Forewing is
light orange-brown, mottled with some black scales; two or three black spots are
distributed as follows: one distinct in the cell, one indistinct (sometimes missing) above
it, and one rather indistinct (rarely missing) at the end of the cell; a small patch of black
scales at tornus; an indistinct light fascia from outside tornus to costa; termen
emarginated below apex, with a fine, black line; cilia concolorous with forewing;
hindwing grey, with yellow cilia. (Karsholt & Riedl, 1996). Larva feeds on Origanum
vulgare, folding a leaf and spinning it together, only leaving a small entrance in each
end, through which it rapidly disappears if disturbed. Adults fly during June and late
August (Huemer & Karsholt, 2002).
Acknowledgements
I would like to thank Mr. Kazimieras Martinaitis (Kaunas T. Ivanauskas Museum of
Zoology) for photos.
References
Aarvik L., Bengtsson B. A., Elven H., Ivinskis P., Jürivete U., Karsholt O., Mutanen M.,
Savenkov N. 2017. Nordic-Baltic Checklist of Lepidoptera. Norwegian Journal of
Entomology, Supplement 3: 1–236.
Baraniak E., Walczak U., Karsholt O. 2014. Materiały do znajomości Gelechiidae
(Lepidoptera) Wielkopolskiego Parku Narodowego. Contributions to the knowledge
of Gelechiidae (Lepidoptera) of the Wielkopolski National Park. Wiadomości
Entomologiczne 33 (1): 21–31.
Becker V. O. 1984. Gelechiidae. In: Heppner, J. B. (ed.) Atlas of Neotropical Lepidoptera.
Checklist: Part 1. Boston, 44–53.
Common I. F. B. 1990. Moths of Australia. Melbourne, 585pp.
Elsner G., Huemer P., Tokar Z. 1999. Die Palpenmotten (Lepidoptera, Gelechiidae)
Mitteleuropas. Bratislava.
Heppner J. B. 1991. Faunal regions and the diversity of Lepidoptera. Tropical
Lepidoptera 2 (Supplement 1): 1–85.
Hodges R. W. 1983. Gelechioidea. In: Hodges R. W., Dominick T., Davis D. R.,
Ferguson D. C., Franclemont J. G., Munroe E. G., Powell J. A. (eds.) Check List of the
Lepidoptera of America North of Mexico. London, 11–25.
Hodges R. W. 1999. Gelechioidea, Gelechiidae (part), Gelechiinae (Part-Chionodes). In:
Hodges R. W., Davis D. R., Dominick T., Ferguson D. C., Munroe E. G., Powell J. A.
(eds.) Moths of America North of Mexico. Washington, 1–5.
72 BULLETIN OF THE LITHUANIAN ENTOMOLOGICAL SOCIETY. Volume 3 (31)
Huemer P., Karsholt O. 1999. Gelechiidae I (Gelechiinae: Teleiodini, Gelechiini). In:
Huemer P., Karsholt O., Lyneborg L. (eds.). Microlepidoptera of Europe (Vol. 3).
Brill, Netherlands.
Huemer P., Karsholt O. 2002. A review of the genus Acompsia Hübner, 1825, with
description of new species (Gelechiidae). Nota Lepidopterologica 25 (2/3): 109–151.
Huemer P., Karsholt O. 2010. Gelechiidae II (Gelechiinae: Gnorimoschemini). In:
Huemer P, Karsholt O, Nuss M. (eds.) Microlepidoptra of Europe (Vol. 6). Stenstrup.
Ivinskis P. 2004. Lepidoptera of Lithuania. Annotated catalogue. Vilnius.
Ivinskis P., Rimšaitė J. 2018. Check-list of the Lithuanian Lepidoptera. Vilnius.
Karsholt O., Nieukerken E. J. 2013 Lepidoptera, Moths. Fauna Europaea version
2017.06, https://fauna-eu.org (Accessed October 05, 2018).
Karsholt O., Riedl T. 1996. Gelechiidae. In: Karsholt O., Razowski J. (eds) The
Lepidoptera of Europe. Stenstrup, 103–122.
Mackay A. 2019. UK moths. Available from https://ukmoths.org.uk (Accessed 20
October, 2019).
Lewis Ch. 2016. Available from https://britishlepidoptera.weebly.com (Accessed 20
October, 2019).
Nielsen E. S., Common I. F. B. 1991. Lepidoptera (moths and butterflies). In: Division
of Entomology, CSIRO (ed) The Insects of Australia. Second Edition. New York,
817–915.
Park K. T. 1992. Systematics of the Subfamily Gelechiinae (Lepidoptera, Gelechiidae) in
Korea II: Tribe Teleiodini. Insecta Koreana 9: 1–33.
Piskunov V. I. 1990. Gelechiidae. In: Medvedev G. S. (ed.) Keys to the Insects of the
European Part of the USSR IV, Lepidoptera, part 1: 889–1024.[English translation]
Solly F. 2019. UK moths. Available from https://ukmoths.org.uk (Accessed 20 October,
2019).
Patočka J., Turčáni M. 2005. Lepidoptera Pupae, Central European Species. Stenstrup.
Nauji duomenys apie 3 naujas ir 6 retas Gelechiidae drugių rūšis (Lepidoptera:
Gelechiidae) Lietuvoje
BRIGITA PAULAVIČIŪTĖ
Santrauka
Straipsnyje pateikiami duomenys apie 3 naujas ir 6 retas Gelechiidae šeimos rūšis
Lietuvos teritorijoje. Metzneria aestivella, Syncopacma larseniella, Acompsia
schmidtiellus yra naujos Lietuvos faunos drugių rūšys. Straipsnyje pateikta kiekvienos
rūšies sugavimo vieta, data, individų skaičius ir rinkėjai.
Received: 24 October, 2019
Article
Full-text available
Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera (Aarvik et al. 2017) are presented. 44 species new to the Nordic-Baltic area as well as numerous new country records are added. Two misspellings and 14 other mistakes of various types are corrected. The genus name Tuta Kieffer & Jørgensen, 1910 is recombined with the species name absoluta Meyrick, 1917 as Tuta absoluta (Meyrick, 1917) comb. rev.
Article
Full-text available
Additions and corrections to the Nordic-Baltic Checklist of Lepidoptera (Aarvik et al. 2017) are presented. 44 species new to the Nordic-Baltic area as well as numerous new country records are added. Two misspellings and 14 other mistakes of various types are corrected. The genus name Tuta Kieffer & Jørgensen, 1910 is recombined with the species name absoluta Meyrick, 1917 as Tuta absoluta (Meyrick, 1917) comb. rev.
Article
Full-text available
The Palaearctic genus Acompsia is revised and two subgenera are considered: Acompsia Hübner, 1825 and Telephila Meyrick, 1923. Altogether 17 species are dealt with in detail and genitalia and adults are figured. 7 new species are described: Acompsia (A.) pyrenaella sp. n. (Spain: Pyrenees), A. (A.) ponomarenkoae sp. n. (Albania, Greece), A. (A.) schepleri sp. n. (Turkey), A. (A.) fibigeri sp. n. (Turkey), A. (A.) bidzilyai sp. n. (Russia: Transbaikalia), A. (A.) caucasella sp. n. (Russia: Caucasus) and A. (T.) syriella sp. n. (Syria). Lectotypes for A. maculosella (Stainton, 1851), A. dimorpha Petry, 1904 and A. minorella (Rebel, 1899) and a neotype for A. tripunctella ([Denis & Schiffermüller], 1775) are designated.
Book
The Gnorimoschemini (Gelechiidae: Gelechiinae) of Europe are reviewed. A total of 211 species is recognized, including two species not yet found in Europe with certainty and two others recently introduced. Descriptions and diagnoses are accompanied by colour figures of the adults, usually depicting variation, and black and white photographs of male and female genitalia characters. Notes on distribution and biology are added for every species. 15 new species are described: Gnorimoschema nupponeni sp. n. (Russia, Ukraine), Scrobipalpa skulei sp. n. (Slovenia), Scrobipalpa dorsolutea sp. n. (Russia), Scrobipalpa corleyi sp. n. (Portugal), Scrobipalpa oleksiyella sp. n. (Russia), Scrobipalpa tokari sp. n. (Croatia), Scrobipalpa heimi sp. n. (Russia), Scrobipalpa jarii sp. n. (Bulgaria), Scrobipalpa abstrusa sp. n. (Russia), Scrobipalpa hendrikseni sp. n. (Greece), Scrobipalpa deutschi sp. n. (Russia, Algeria), Scrobipalpa mixta sp. n. (Macedonia, ?Albania, Ukraine), Scrobipalpa postulatella sp. n. (Spain), Scrobipalpa halimioniella sp. n. (France), Caryocolum baischi sp. n. (Greece). The following 30 species group names are synonymized and two taxa are revised as subspecies: Lita porcella Heinemann, 1870 syn. n. of Scrobipalpa acuminatella (Sircom, 1850); Scrobipalpa sibirica Bidzilya, 2009 syn. n. of Scrobipalpa occulta (Povolný, 2002); Scrobipalpa deleta Povolný, 1981 syn. n. of Scrobipalpa clintoni Povolný, 1968; Euscrobipalpa bezengensis Povolný, 2001 syn. n. of Scrobipalpa caucasica (Povolný, 2001); Scrobipalpa submagnificella Povolný, 1977 syn. n. of Scrobipalpa heretica Povolný, 1973; Scrobipalpa glaserorum Povolný, 1977 syn. n. of Scrobipalpa bradleyi Povolný, 1971; Scrobipalpa fraterna Povolný, 1969 syn. n. of Scrobipalpa selectella (Caradja, 1920); Scrobipalpa milleri Povolný, 1977 syn. n. of Scrobipalpa niveifacies Povolný, 1977; Gnorimoschema hyoscyamivora Gerasimov, 1940 and Gnorimoschema pseudobsoletellum Povolný & Gregor, 1955 syn. n. of Scrobipalpa indignella (Staudinger, 1879); Lita atriplicella var. infumatella Fuchs, 1901 syn. n. of Scrobipalpa atriplicella (Fischer von Röslerstamm, 1841); Scrobipalpa hartigi Povolný, 1977 syn. n. of Scrobipalpa suaedella (Richardson, 1893); Scrobipalpa rezniki Piskunov, 1990 and Scrobipalpa rezniki turkmenica Piskunov, 1990 syn. n. of Scrobipalpa dagmaris Povolný, 1987; Scrobipalpa artemisiella mongolensis Povolný, 1969 syn. n. of Scrobipalpa artemisiella (Treitschke, 1833); Scrobipalpa japonica Povolný, 1977 syn. n. of Scrobipalpa rebeli (Preissecker, 1914); Gelechia eremaula Meyrick, 1891, Gelechia lyciella Walsingham, 1900, Lita desertella Rebel, 1901, Phthorimaea bertramella Lucas, 1940, Teleia leroyella Lucas 1950, Gnorimoschema reisseri Povolný & Gregor, 1955 and Gnorimoschema philolycii Hering, 1957 syn. n. of Scrobipalpa portosanctana (Stainton, 1859); Scrobipalpa trebujenae Povolný, 1977 syn. n. of Scrobipalpa salinella (Zeller, 1847); Lita caliacrae Caradja, 1932 syn. n. of Scrobipalpa salicorniae (Hering, 1889); Phthorimaea argentinae (Povolný, 1989) syn. n. of Phthorimaea operculella (Zeller, 1873); Opacopsis buvati Povolný, 1992 syn. n. of Ephysteris deserticolella (Staudinger, 1871); Doryphora praticolella Christoph, 1872 (as ssp.) and Opacopsis gallica Povolný, 1992 syn. n. of Ephysteris insulella (Heinemann, 1870); Ephysteris (Opacopsis) treskensis Povolný, 1964, Ephysteris (Opacopsis) hispanica Povolný, 1981 and Ephysteris (Opacopsis) foulonsensis Povolný, 1981 syn. n. of Ephysteris diminutella (Zeller, 1847); Ephysteris (Opacopsis) olympica ssp. monticola Povolný, 1981 syn. n. of Ephysteris olympica Povolný, 1968; Phthorimaea microcasis Meyrick, 1929, Hedma abzacella Dumont, 1932, Teleia hyoscyami Amsel, 1935, Megalocypha polioptera Janse, 1960 and Ephysteris aellographa Janse, 1960 syn. n. of Microlechia chretieni Turati, 1924; Lita roseella Zetterstedt, 1839 syn. n. of Cosmardia moritzella (Treitschke, 1835); Lita crepusculella Teich, 1889 syn. n. of Caryocolum viscariella (Stainton, 1855); Caryocolum emarginatum Huemer, 1988 syn. n. of Caryocolum stramentella Rebel, 1935; Phthorimaea horoscopa Meyrick, 1926 as ssp. stat. n. of Caryocolum blandella (Douglas, 1852). The following genus group names are synonymized: Hedma Dumont, 1932 and Megalocypha Janse, 1960 syn n. of Microlechia Turati, 1924. The following new combinations are introduced: Scrobipalpa karadaghi (Povolný, 2001) comb. n.; Scrobipalpa acuta (Povolný, 2001) comb. n.; Scrobipalpa alterna (Falkovitsh & Bidzilya, 2006) comb. n.; Scrobipalpa grossoides (Povolný, 2001) comb. n.; Microlechia rhamnifoliae (Amsel & Hering, 1931) comb. n.; Microlechia karsholti (Nupponen, 2010) comb. n. The following taxa are recalled from synonymy: Gnorimoschema nilsi Huemer, 1996 sp. rev.; Gnorimoschema hoefneri (Rebel, 1909) sp. rev.; Scrobipalpa divisella (Rebel, 1936) sp. rev.; Scrobipalpa salicorniae (Hering, 1889) sp. rev.; Scrobipalpa spegulariella (Chrétien, 1910) sp. rev.; Scrobipalpa thymelaeae (Amsel, 1939) sp. rev.; Scrobipalpa halymella (Millière, 1864) sp. rev.; Klimeschiopsis terroris (Hartig, 1938) sp. rev.; Microlechia Turati, 1924 gen. rev. The specific status of following taxa is revised: Gnorimoschema alaskense Povolný, 1967 stat. rev.; Scrobipalpa stabilis Povolný, 1977 stat. n.; Ephysteris inustella gredosensis (Rebel, 1935) stat. rev.; Ephysteris insulella praticolella (Christoph, 1872) stat. n.; Lutilabria lutilabrella olympica Huemer, 1993 stat. n.; Caryocolum blandella horoscopa (Meyrick, 1926) stat. n. Finally the synonymy of Gnorimoschema antiquum Povolný, 1966 syn. rev. and the generic combinations of Scrobipalpa lagodes (Meyrick, 1926) comb. rev. and Scrobipalpa ergasima (Meyrick, 1916) comb. rev. are revised. Lectotypes are designated for the following taxa: Phthorimaea calaritanella Amsel, 1952; Gelechia portosanctana Stainton, 1859; Gelechia lyciella Walsingham, 1900; Gelechia salinella Zeller, 1847; Teleia hyoscyamella Rebel, 1912; Oecophora moritzella Treitschke, 1835; Lita roseella Zetterstedt, 1839.
Book
The book presents description and identification keys of more than 2000 Lepidopteran species living in Europe.
Metzneria aestivella (Zeller, 1839) is spread widely from the Canary Islands throughout Europe to Central Asia
  • Elsner
Discussion Gelechiidae moths are not comprehensively investigated in Lithuania. New species are published almost every year. Metzneria aestivella (Zeller, 1839) is spread widely from the Canary Islands throughout Europe to Central Asia (Elsner et al., 1999). This species already is known in many European countries: Austria, Belarus, Belgium, Bosnia Herzegovina, Bulgaria, Croatia, Cyprus, The Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Crete, Latvia, Macedonia, Malta, Poland, Portugal, Romania, Russia, Sardinia, Serbia, Sicily, Slovakia, Slovenia, Spain, Sweden, Switzerland, the Netherlands (Karsholt & Nieukerken, 2013; Aarvik et al., 2017). The moth wingspan is usually 11-14 mm (Fig. 1). Adults are very similar to M. lappella and References
Die Palpenmotten (Lepidoptera, Gelechiidae) Mitteleuropas
  • G Elsner
  • P Huemer
  • Z Tokar
Elsner G., Huemer P., Tokar Z. 1999. Die Palpenmotten (Lepidoptera, Gelechiidae) Mitteleuropas. Bratislava.
Gelechiinae (Part-Chionodes)
  • R W Hodges
Hodges R. W. 1999. Gelechioidea, Gelechiidae (part), Gelechiinae (Part-Chionodes). In: Hodges R. W., Davis D. R., Dominick T., Ferguson D. C., Munroe E. G., Powell J. A. (eds.) Moths of America North of Mexico. Washington, 1-5.