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Notulae to the Italian alien vascular ora: 8 63
Notulae to the Italian alien vascular flora: 8
Gabriele Galasso1, Gianniantonio Domina2, Sebastiano Andreatta3,
Claudia Angiolini4, Nicola M.G. Ardenghi5, Claudio Aristarchi6, Matteo Arnoul7,
Mattia M. Azzella8, Gianluigi Bacchetta9, Fabrizio Bartolucci10, Silvia Bodino5,
Giacomo Bommartini11, Gianmaria Bonari12, Sergio Buono13, Vito Buono14,
Orazio Caldarella15, Giacomo Calvia9, Emilio Corti16, Marco D’Antraccoli17,
Rocco De Luca18, Fabrizio De Mattia19, Stefano Di Natale16,
Alessandra Di Turi6, Assunta Esposito20, Giulio Ferretti16, Tiberio Fiaschi4,
Maria C. Fogu9, LuigiForte21, Jessica Frigerio19, Leonardo Gubellini22,
Lorenzo Guzzetti23, Nicole Hofmann24, Valentina L.A. Laface25,
Gaetano Laghetti26, Andrea Lallai9, Alfonso La Rosa27, Lorenzo Lazzaro16,
Silvano Lodetti5, Michele Lonati28, Fabio Luchino29, Sara Magrini30,
Andrea Mainetti28, Michela Marignani9, GinaMaruca26, Pietro Medagli31,
Giacomo Mei24, Flavio Menini32, Valerio Mezzasalma19, Alice Misuri16,
Sara Mossini33, Michele Mugnai16, Carmelo M. Musarella25, Ginevra Nota28,
Nicola Olivieri34, Alessia Padula16, Marziano Pascale35, Federico Pasquini16,
Lorenzo Peruzzi17, Gianni Picella36, Lorenzo Pinzani17, Silvia Pirani28,
Marco Pittarello28, Lina Podda9, Simone Ravetto Enri28, Carmelo D. Rici37,
Francesco Roma-Marzio38, Rosario Romano39, Leonardo Rosati40, Filippo Scadi2,
Enrico Scarici41, Marco Scarici42, Giovanni Spampinato25, Adriano Stinca20,
Robert P. Wagensommer43, Giovanni Zanoni44, Chiara Nepi45
1Sezione di Botanica, Museo di Storia Naturale di Milano, Corso Venezia 55, 20121 Milano, Italy
2Dipartimento di Scienze Agrarie, Alimentari e Forestali (SAAF), Università di Palermo, Viale delle Scienze,
edicio 4, 90128 Palermo, Italy 3 Museo Civico di Storia Naturale di Verona, Piazza Arsenale 8, 37126
Verona, Italy 4 Dipartimento di Scienze della Vita, Università di Siena, Via P.A. Mattioli 4, 53100 Siena,
Italy 5Dipartimento di Scienze della Terra e dell’Ambiente, Università di Pavia, Via Sant’Epifanio 14, 27100
Pavia, Italy 6 Via Crocetta di Apparizione 32, 16133 Genova, Italy 7 Loc. Serre 6, 10060 Angrogna (Torino),
Italy 8Centro Ricerche Frascati, Agenzia Nazionale per le Nuove Tecnologie, l’Energia e lo Sviluppo Economico
Sostenibile (ENEA), Via E. Fermi 45, 00044 Frascati (Roma), Italy 9 Dipartimento di Scienze della Vita
e dell’Ambiente, Università di Cagliari, Viale Sant’Ignazio da Laconi 13, 09123 Cagliari, Italy 10 Centro
Ricerche Floristiche dell’Appennino (Università di Camerino – Parco Nazionale del Gran Sasso e Monti della
Laga), San Colombo, 67021 Barisciano (L’Aquila), Italy 11 Via Buggia 17, 37050 Belore (Verona), Italy
12 Department of Botany and Zoology, Masaryk University, Kotlářská 2, CZ-611 37 Brno, Czech Republic
Italian Botanist 8: 63–93 (2019)
doi: 10.3897/italianbotanist.8.48621
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Copyright Gabriele Galasso et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
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RESEARCH ARTICLE
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
64
13 Via XXV Aprile 6, 01010 Oriolo Romano (Viterbo), Italy 14 Vico Traversa 2, 70127 Bari, Italy 15 Viale
Maria Santissima Mediatrice 38, 90129 Palermo, Italy 16 Dipartimento di Biologia, Università di Firenze,
Via G. La Pira 4, 50121 Firenze, Italy 17 Dipartimento di Biologia, Università di Pisa, Via Derna 1,
56126 Pisa, Italy 18Via M. Reitano Spadafora, isolato 43, 98124 Messina, Italy 19 FEM2 Ambiente s.r.l.,
Piazza della Scienza 2, 20126 Milano, Italy 20 Dipartimento di Scienze e Tecnologie Ambientali, Biologiche
e Farmaceutiche, Università della Campania Luigi Vanvitelli, Via A. Vivaldi 43, 81100 Caserta, Italy
21Dipartimento di Biologia e Museo Orto Botanico, Università di Bari Aldo Moro, Via E. Orabona 4, 70125
Bari, Italy 22 Centro Ricerche Floristiche Marche, Provincia di Pesaro e Urbino, Via E. Barsanti 18, 61122
Pesaro (Pesaro e Urbino), Italy 23 Dipartimento di Biotecnologie e Bioscienze, Università di Milano-Bicocca,
Piazza della Scienza 2, 20126 Milano, Italy 24 Dipartimento di Scienze Agrarie, Alimentari ed Ambientali,
Università Politecnica delle Marche, Via Brecce Bianche 10, 60131 Ancona, Italy 25 Dipartimento di Agraria,
Università Mediterranea di Reggio Calabria, Feo di Vito snc, 89122 Reggio Calabria, Italy 26 Istituto di
Bioscienze e Biorisorse, Consiglio Nazionale delle Ricerche (CNR-IBBR), Via G. Amendola 165/a, 70126
Bari, Italy 27 Cooperativa Silene, Via V. D’Ondes Reggio Vito 8/a, 90127 Palermo, Italy 28 Dipartimento di
Scienze Agrarie, Forestali e Alimentari (DISAFA), Università di Torino, Largo P. Braccini 1, 10095 Grugliasco
(Torino), Italy 29 Via Torrente Allume 6/a, 98027 Roccalumera (Messina), Italy 30 Banca del Germoplasma
della Tuscia, Università della Tuscia, Largo dell’Università snc, blocco c, 01100 Viterbo, Italy 31 Dipartimento
di Scienze e Tecnologie Biologiche e Ambientali (DiSTeBA), Università del Salento, Strada Provinciale Lecce-
Monteroni 165, 73100 Lecce, Italy 32 Via D. Ricci 4, 37042 Caldiero (Verona), Italy 33 Via E. Raovati 6,
27049 Stradella (Pavia), Italy 34 Via Maestri del Lavoro 40, 64100 Teramo, Italy 35 Via della Repubblica
14, 12018 Roccavione (Cuneo), Italy 36 Via Quarnaro 16, 70121 Bari, Italy 37 Via G. Mazzini 5, 98064
Librizzi (Messina), Italy 38 Sistema Museale di Ateneo, Università di Pisa, Via L. Ghini 13, 56126 Pisa,
Italy 39 Via A. Gramsci Vicolo 1 6, 96011 Augusta (Siracusa), Italy 40 Scuola di Scienze Agrarie, Forestali
e Ambientali, Università della Basilicata, Via Ateneo Lucano 10, 85100 Potenza, Italy 41 Loc. Paglianello
1/a, 01012 Capranica (Viterbo), Italy 42 Via degli Anguillara 90, 01012 Capranica (Viterbo), Italy
43Dipartimento di Chimica, Biologia e Biotecnologie, Università di Perugia, Via del Giochetto 6, 06122
Perugia, Italy 44 Via Ambrosi 9, 37038 Soave (Verona), Italy 45 Sistema Museale di Ateneo, Università di
Firenze, Via G. La Pira 4, 50121 Firenze, Italy
Corresponding author: Gabriele Galasso (gabriele.galasso@comune.milano.it)
Academic editor: S. Biondi|Received 20 November 2019|Accepted 8 December 2019|Published 17 December2019
Citation: Galasso G, Domina G, Andreatta S, Angiolini C, Ardenghi NMG, Aristarchi C, Arnoul M, Azzella MM,
Bacchetta G, Bartolucci F, Bodino S, Bommartini G, Bonari G, Buono S, Buono V, Caldarella O, Calvia G, Corti E,
D’Antraccoli M, De Luca R, De Mattia F, Di Natale S, Di Turi A, Esposito A, Ferretti G, Fiaschi T, Fogu MC, Forte L,
Frigerio J, Gubellini L, Guzzetti L, Hofmann N, Laface VLA, Laghetti G, Lallai A, La Rosa A, Lazzaro L, Lodetti S, Lonati
M, Luchino F, Magrini S, Mainetti A, Marignani M, Maruca G, Medagli P, Mei G, Menini F, Mezzasalma V, Misuri A,
Mossini S, Mugnai M, Musarella CM, Nota G, Olivieri N, Padula A, Pascale M, Pasquini F, Peruzzi L, Picella G, Pinzani
L, Pirani S, Pittarello M, Podda L, Ravetto Enri S, Rici CD, Roma-Marzio F, Romano R, Rosati L, Scadi F, Scarici E,
Scarici M, Spampinato G, Stinca A, Wagensommer RP, Zanoni G, Nepi C (2019) Notulae to the Italian alien vascular
ora: 8. Italian Botanist 8: 63–93. https://doi.org/10.3897/italianbotanist.8.48621
Abstract
In this contribution, new data concerning the distribution of vascular ora alien to Italy are presented.
It includes new records, conrmations, exclusions, and status changes for Italy or for Italian administra-
tive regions of taxa in the genera Bunias, Calocedrus, Calycanthus, Celosia, Clerodendrum, Convolvulus,
Crassula, Cyclamen, Datura, Dicliptera, Eragrostis, Erigeron, Gamochaeta, Gazania, Impatiens, Kolkwitzia,
Notulae to the Italian alien vascular ora: 8 65
Leucaena, Ludwigia, Medicago, Muscari, Nigella, Oenothera, Opuntia, Paulownia, Petroselinum, Phyllos-
tachys, Physalis, Pseudosasa, Quercus, Reynoutria, Roldana, Saccharum, Sedum, Semiarundinaria, Senecio,
Sisyrinchium, Solanum, Sporobolus, Tulipa, Vachellia, Verbena, and Youngia. Nomenclatural and distribu-
tion updates published elsewhere are provided as Suppl. material 1.
Keywords
Alien species, oristic data, Italy
How to contribute
e text for the new records should be submitted electronically to Chiara Nepi (chiara.
nepi@uni.it). e corresponding specimen along with its scan or photograph has to
be sent to FI Herbarium: Museo di Storia Naturale (Botanica), Sistema Museale di
Ateneo, Via G. La Pira 4, 50121 Firenze (Italy). ose texts concerning nomenclatural
novelties (typications only for accepted names), status changes, exclusions, and con-
rmations should be submitted electronically to: Gabriele Galasso (gabriele.galasso@
comune.milano.it). Each text should be within 2,000 characters (spaces included).
Floristic records
Bunias orientalis L. (Brassicaceae)
+ (NAT) EMR. – Status change from casual to naturalized alien for the ora of Emilia-
Romagna.
In Italy, this species was already cultivated in botanical gardens in the late 18th
century (e.g., in Pavia, see Anonymous 1785) and was rst recorded as a casual al-
ien in 1897 (Penzig 1897; Béguinot and Mazza 1916). Bunias orientalis now occurs
as a casual alien in most of the northern regions, with the exceptions of Piemonte
and Friuli Venezia Giulia, where it is considered naturalized (Galasso et al. 2018a).
In Emilia-Romagna, it is known as casual in the province of Ferrara (Piccoli et al.
2014). On July 2nd, 2019, a large population was discovered in the locality Casino
of the former municipality of Nibbiano (now Alta Val Tidone), province of Pia-
cenza (WGS84: 44.945631N, 9.332003E). Here, fruiting individuals form a thick
stand of 3,350 m2, with a 75% cover, on waste land colonized by Artemisia vulgaris
L., Cirsium arvense (L.) Scop., Elymus repens (L.) Gould subsp. repens, and Sambu-
cus ebulus L. More than 1,200 rosettes were counted across a mowed wheat eld
of 31,000 m2 in locality Casa Castellina (WGS84: 44.946498N, 9.330900E) and
further individuals were observed along the nearby roadsides. is species, simi-
larly to other European countries (see e.g., Clement and Foster 1994), was likely
introduced as a grain impurity. e pronounced tendency to invasiveness in these
localities needs to be monitored.
N.M.G. Ardenghi, S. Bodino, S. Lodetti
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
66
Calocedrus decurrens (Torr.) Florin (Cupressaceae)
+ (CAS) TOS: Firenze (Firenze), Parco delle Cascine, alla conuenza tra il Torrente
Mugnone e il Fiume Arno (WGS84: 43.790598N, 11.197569E), 50 m, 3 February
2019, L. Pinzani (FI). – Casual alien species new for the ora of Toscana.
Calocedrus decurrens has been already recorded as casual in Lombardia, Umbria,
and Sardegna (Galasso et al. 2018a). Some young individuals originated by seeds from
nearby cultivated plants were found in Firenze, at the Cascine Park.
L. Pinzani
Calycanthus oridus L. (Calycanthaceae)
+ (CAS) MAR: San Benedetto del Tronto (Ascoli Piceno), aiuola presso Viale delle
Palme (WGS84: 42.949722N, 13.884780E), epita su stipite di Phoenix canariensis,
ca. 8 m, 9 July 2019, N. Olivieri (FI). – Casual alien species new for the ora of Marche.
Calycanthus oridus is an ornamental species native to southeastern North America
and introduced in Italy in 1788 (Maniero 2015). In Italy, it is known as casual alien only
in Toscana (Galasso et al. 2018a). Some young individuals of the species have developed
as epiphytes on the trunk of a young Phoenix canariensis H.Wildpret, settling among
the remains of brous tissue present among the stumps of the leaf rachids. e plants
developed from seeds produced by a shrub cultivated in a owerbed at a short distance.
e area is located in a rather sheltered position due to the presence of groups of Pinus
halepensis Mill. subsp. halepensis and alignments of buildings that limit insolation and
reduce the inuence of the eastern sea winds, creating a cooler microclimate.
N. Olivieri
Celosia argentea L. (Amaranthaceae)
+ (CAS) SIC: Polizzi Generosa (Palermo), strada SS643 (WGS84: 37.839167N,
13.960000E), bordo stradale, ca. 550 m, 12 September 2005, G. Domina, S.L. Jury
(PAL); Ciminna (Palermo), strada SP33 (WGS84: 37.895676N, 13.544894E), bordi
stradali, ca. 600 m, 7 October 2018, leg. F. Scadi, det. F. Scadi, G. Domina (FI). –
Casual alien species conrmed for the ora of Sicilia.
is species was likely introduced in Italy for ornamental purposes, and the indi-
viduals found along the roadside may have originated from cultivated plants growing
nearby. is species was identied according to Iamonico (2013). In Galasso et al.
(2018a), it is reported as casual alien for many Italian administrative regions, but not
recently recorded from Sicilia.
F. Scadi, G. Domina
Clerodendrum trichotomum unb. (Lamiaceae)
+ (CAS) TOS: Borgo San Lorenzo (Firenze), fraz. Panicaglia, ex stazione ferroviaria lungo
la ferrovia Faentina (WGS84: 43.978605N, 11.407556E), alcune piante spontaneizzate ai
Notulae to the Italian alien vascular ora: 8 67
margini di un giardino in vicinanza di piante adulte coltivate, 281 m, 8 November 2018,
M. Mugnai, A. Misuri, G. Ferretti (FI). – Casual alien species new for the ora of Toscana.
is species was already recorded in Italy as naturalized, mostly in northern Italy.
Some young individuals were found at the railway station of Panicaglia, probably origi-
nating from adult fruiting plants of a neighboring garden. Given the ephemeral condi-
tion of the occurrence site, we consider this species as casual for Toscana.
M. Mugnai, A. Misuri, L. Lazzaro
Convolvulus sabatius Viv. subsp. mauritanicus (Boiss.) Murb. (Convolvulaceae)
+ (NAT) PUG: Giovinazzo (Bari), tra Giovinazzo e Santo Spirito (comune di Bari)
(WGS84: 41.179111N, 16.689172E), margine stradale, 3 m, 29 May 2019, leg. V.
Buono, det. R.P. Wagensommer (FI, BI Nos. 42141, 42142); Lecce (Lecce), presso il
cimitero (WGS84: 40.359449N, 18.165857E), aiuola incolta, 40 m, 15 June 2019, P.
Medagli (LEC). – Naturalized alien subspecies new for the ora of Puglia.
Convolvulus sabatius was rst recorded from Puglia near Giovinazzo (Bianco 1969)
and then collected in Salento (Marchiori et al. 1993), Bari and Monopoli (Perrino
et al. 2013). ese collections were all attributed to C. sabatius subsp. sabatius. Our
gatherings, from Giovinazzo and Lecce, show long spreading hairs on stems, leaves
and calyx and are, therefore, attributed to C. sabatius subsp. mauritanicus, according
to Carine and Robba (2010) and Wood et al. (2015). Consequently, we consider C.
sabatius subsp. sabatius as recorded from Puglia by mistake (Bartolucci et al. 2019).
R.P. Wagensommer, V. Buono, P. Medagli
Crassula muscosa L. (Crassulaceae)
+ (CAS) MAR: San Benedetto del Tronto (Ascoli Piceno), Via C. Colombo (WGS84:
42.956186N, 13.882666E), epita su stipite di Phoenix canariensis, ca. 6 m, 9 July
2019, N. Olivieri (FI). – Casual alien species new for the ora of Marche.
Crassula muscosa is native to southern Africa and is widely cultivated as ornamen-
tal. In Italy, it is known as casual alien in Toscana, Campania, and Sicilia, while it is
considered naturalized in Liguria, Calabria, and Sardegna (Galasso et al. 2018a). Some
individuals of this species grow as epiphytes on the trunk of a Phoenix canariensis
H.Wildpret inside the city. e plants have developed among the residues of brous
tissue between the remains of the cut leaf rachids, in a partially shaded position. Indi-
viduals may have arisen via vegetative propagation from fragments of plants grown for
ornamental purposes in nearby buildings.
N. Olivieri
Cyclamen persicum Mill. (Primulaceae)
+ (CAS) LIG: Genova (Genova), lungo Via Tortona (WGS84: 44.42558N, 8.95276E),
a bordo strada, nelle crepe dell’asfalto, 81 m, 18 April 2019, A. Di Turi, C. Aristarchi
(FI, GE). – Casual alien species new for the ora of Liguria.
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
68
Cyclamen persicum is a widely cultivated plant, whose native range extends from
Algeria to the eastern Mediterranean. It is reported in Italy as a casual alien for Lom-
bardia (Ban and Galasso 2010), Sardegna (Lazzeri et al. 2015), and Lazio (Nicolella
2018). Well-developed specimens were rst recorded in 2000 in Viale G. Odino in the
centre of Genova. Recently other specimens have been found at three dierent sites,
both in the city centre (Via Fieschi, WGS84: 44.403397N, 8.935548E, 36 m) and in
more peripheral sites (Via V. Bocciardo, WGS84: 44.404441N, 8.993866E, 168 m;
and Via Tortona). All grow in the cracks of sidewalks, without any other species nearby.
One of them was in bloom when recorded (April 2019).
A. Di Turi, C. Aristarchi
Datura wrightii Regel (Solanaceae)
+ (CAS) MAR: Urbino (Pesaro e Urbino), fraz. Canavaccio, lungo il Fiume Metauro
(WGS84: 43.688780N, 12.700244E), greto uviale, ca. 175 m, 13 September 2018, L.
Gubellini, N. Hofmann (FI, PESA). – Casual alien species new for the ora of Marche.
Datura wrightii is an annual plant native to the southwestern United States and
Mexico (Verloove 2008). It is reported as casual alien in almost all regions of north-
ern and central Italy (Lombardia, Veneto, Trentino-Alto Adige, Friuli Venezia Giulia,
Liguria, Emilia-Romagna, Umbria, Lazio, Abruzzo, Campania, Puglia, and Calabria),
as naturalized alien for Toscana and Sicilia, and as invasive alien for Sardegna (Galasso
et al. 2018a). In Marche, a single individual was observed with abundant owers and
fruits in a stony bank along the Metauro River, far from gardens and urban centre. is
species has been long confused with the related D. inoxia Mill., less common in Italy,
which diers from D. wrightii for the type of indument (Verloove 2008).
L. Gubellini, N. Hofmann
Dicliptera squarrosa Nees (Acanthaceae)
+ (CAS) ITALIA (TOS): Seravezza (Lucca), fraz. Querceta, vicino all’ingresso della sta-
zione ferroviaria “Forte dei Marmi-Seravezza-Querceta”, Via Ragazzi del ‘99 (WGS84:
43.978158N, 10.198074E), margine stradale, 18 m, 23 October 2018, M. Mugnai,
L. Lazzaro, G. Ferretti (FI). – Casual alien species new for the ora of Italy (Toscana).
Dicliptera squarrosa is an ornamental plant native to South America, which pre-
sents several forms, separated mostly geographically and hardly forming discrete units
(Wasshausen and Wood 2004). is species is currently widely available for sale world-
wide and is largely used also in Italy. We found one owering individual clearly escaped
from cultivation close to the Querceta railway station. According to some authors (J.
Wood, pers. commun.), the forms cultivated in Europe should be referred to Dicliptera
suberecta (André) Bremek., currently considered as a synonym of D. squarrosa (Zuloaga
et al. 2008). Nevertheless, further studies are needed to solve this issue and we prefer to
provisionally maintain this record under D. squarrosa.
M. Mugnai, L. Lazzaro, F. Pasquini, G. Ferretti
Notulae to the Italian alien vascular ora: 8 69
Eragrostis mexicana (Hornem.) Link subsp. virescens (J.Presl) S.D.Koch &
Sánchez Vega (Poaceae)
+ (CAS) MAR: Gradara (Pesaro e Urbino), presso il cimitero comunale (WGS84:
43.942494N, 12.769445E), incolto erboso (oliveto abbandonato), ca. 330 m, 16 No-
vember 2018, L. Gubellini, N. Hofmann (FI, PESA). – Casual alien subspecies new for
the ora of Marche.
Eragrostis mexicana subsp. virescens is an alien annual grass from South America re-
corded in Europe since 1927, and in Italy since 1975 (Martini and Scholz 1998). Until
now, it was reported in Italy as naturalized alien in northern regions (Piemonte, Ligu-
ria, Lombardia, Veneto, Trentino-Alto Adige, Friuli Venezia Giulia, Emilia-Romagna)
and Calabria, and as casual alien for Valle d’Aosta, Lazio, Campania, and Puglia (Ga-
lasso et al. 2018a). A large number of individuals were detected by S. Montanari (pers.
commun.) in an uncultivated grassy eld, and the abundance of specimens suggests a
naturalization of the species, which can be conrmed by monitoring the site.
L. Gubellini, N. Hofmann
Erigeron karvinskianus DC. (Asteraceae)
+ (NAT) MAR: Ancona (Ancona), porto antico (WGS84: 43.623791N, 13.509335E),
mura in pietra del porto antico e degli scavi adiacenti, 10 m, 28 August 2018, G. Mei
(Herb. G. Mei); Sirolo (Ancona), piazzetta del belvedere e centro storico (Piazzale Ma-
rino) (WGS84: 43.522444N, 13.621030E), mura in pietra e bordure degli spazi verdi,
110 m, 12 November 2018, G. Mei (FI, ANC, Herb. G. Mei); Piobbico (Pesaro e
Urbino), vicino al ponte sul Torrente Biscubio (WGS84: 43.352610N, 12.303703E),
mura in pietra, ca. 330 m, 21 November 2018, L. Gubellini, N. Hofmann (FI, PESA).
– Naturalized alien species new for the ora of Marche.
Erigeron karvinskianus is an American perennial species native to Mexico and Gua-
temala which occurs all over western Europe, probably escaped from oriculture. To
date, it is present in almost all the Italian territory, with the exception of Valle d’Aosta,
Molise, Basilicata, and Sardegna (Galasso et al. 2018a). In all the recorded localities,
this species was also observed near road edges and in unmanaged owerbeds, mainly
colonizing the gaps in walls, where it seems to be more competitive than other spe-
cies, such as Cymbalaria muralis G.Gaertn., B.Mey. & Scherb. subsp. muralis, Linaria
vulgaris Mill. subsp. vulgaris, and Parietaria judaica L. In all the sites, this species is in
expansion, in particular in Piobbico, a site monitored since 2016.
L. Gubellini, N. Hofmann, G. Mei
+ (INV) CAM: Castellammare di Stabia (Napoli), tra il Castello Angioino e Quisisana
(WGS84: 40.687474N, 14.481252E), muro in pietra calcarea, 135 m, 24 April 2013,
A. Stinca (PORUN-Herb. Stinca); ibidem (WGS84: 40.687493N, 14.481536E), muro
in pietra calcarea, 130 m, 29 September 2018, A. Stinca (PORUN-Herb. Stinca); Ra-
vello (Salerno), Villa Rufolo (WGS84: 40.648479N, 14.613078E), muro in pietra cal-
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
70
carea, 353 m, 10 March 2014, leg. A. Stinca, M. Ravo, det. A. Stinca (PORUN-Herb.
Stinca). – Status change from naturalized to invasive alien for the ora of Campania.
Erigeron karvinskianus was reported as naturalized for Campania by Galasso et
al. (2018a). However, we found this alien plant, in dense and extensive populations,
mostly on walls of limestone and tu blocks of several sites in the Sorrento peninsula.
In these environments, it easily spreads by abundant seed production and competes
strongly with endemic species, such as Campanula fragilis Cirillo subsp. fragilis. ere-
fore, this species should be considered invasive in Campania.
A. Stinca, A. Esposito
Gamochaeta pensylvanica (Willd.) Cabrera (Asteraceae)
+ (NAT) CAL: Reggio Calabria (Reggio Calabria), Catona (WGS84: 38.185074N,
15.637955E), marciapiedi, 2 m, 12 June 2019, leg. V.L.A. Laface, det. V.L.A. Laface,
C.M. Musarella, G. Spampinato (FI, REGGIO); Villa San Giovanni (Reggio Calabria),
Viale U. Zanotti Bianco (WGS84: 38.214703N, 15.637015E), marciapiedi, 8 m, 26
June 2019, V.L.A. Laface (REGGIO); Reggio Calabria (Reggio Calabria), Cittadel-
la Universitaria (WGS84: 38.121004N, 15.662473E), aiuola, 80 m, 26 June 2019,
V.L.A. Laface, C.M. Musarella, G. Spampinato (REGGIO); Campo Calabro (Reggio
Calabria), Musalà (WGS84: 38.214607N, 15.672014E), bordo strada, 154 m, 30
June 2019, V.L.A. Laface (REGGIO); Reggio Calabria (Reggio Calabria), Gallico Ma-
rina (WGS84: 38.169410N, 15.651119E), marciapiedi, 16 m, 10 July 2019, V.L.A.
Laface (REGGIO). – Naturalized alien species new for the ora of Calabria.
Gamochaeta pensylvanica is native to North America. In Italy, its rst record by
Moraldo and La Valva (1989) for Campania, was erroneously attributed by these au-
thors to G. purpurea (L.) Cabrera (Soldano 2000) and then recorded in the same region
by Stinca et al. (2016, 2018). e origin of the introduction of this species in Italy is
uncertain. Probably, G. pensylvanica arrived in Italy through the importation of pot-
ting soil used in plant nurseries. Currently, according to Galasso et al. (2018a), G. pen-
sylvanica is a naturalized alien species in Campania, Piemonte, Lombardia, Emilia-Ro-
magna, and Sicilia, whereas it is casual in Toscana, Lazio, and Puglia. In Calabria, this
species was observed for the rst time in 2008 in locality Catona (Reggio Calabria).
C.M. Musarella, V.L.A. Laface, G. Spampinato
Gazania linearis (unb.) Druce (Asteraceae)
+ (CAS) CAL: San Calogero (Vibo Valentia), Via L. Pirandello (WGS84: 38.576041N,
16.023808E), bordo strada, 256 m, 9 June 2019, C.M. Musarella (FI, REGGIO). –
Casual alien species new for the ora of Calabria.
Gazania linearis has its native range in South Africa and Lesotho. Since it has been
cultivated as an ornamental plant since the 19th century, it has become an invasive
plant in several regions of the world (Hassler 2019). In Italy, according to Galasso et
Notulae to the Italian alien vascular ora: 8 71
al. (2018a), this species is a casual alien to Toscana, Molise, and Puglia, whereas it is
doubtfully recorded for Sardegna.
C.M. Musarella, V.L.A. Laface, G. Spampinato
Impatiens parviora DC. (Balsaminaceae)
+ (INV) TOS: Abetone Cutigliano (Pistoia), fra Pianosinatico e Cecchetto in loc. Ser-
rabosco, versante SE di Poggio del Romito (WGS84: 44.125410N, 10.717628E),
bosco misto di conifere di impianto con residui di faggeta, 900 m, 21 July 2019, F.
Roma-Marzio, M. D’Antraccoli, L. Peruzzi (PI No. 025544). – Status change from
naturalized to invasive alien for the ora of Toscana.
Impatiens parviora is native to central and eastern Asia and represents one of the
most widespread aliens in central Europe, being the only alien plant widespread in Euro-
pean forests (Godefroid and Koedam 2010; Hejda 2012). In Italy, this species is reported
as naturalized in Friuli Venezia Giulia, Emilia-Romagna, Liguria, Toscana, and Lazio,
and as invasive in Valle d’Aosta, Piemonte, Lombardia, Trentino-Alto Adige, and Veneto
(Galasso et al. 2018a). During a eld survey conducted in the Tuscan Apennines, we no-
ticed a large population of this species. e plants are particularly dense, totally covering
the herbaceous layer in shady sites and showing a preference for dry, acidic and nutrient-
poor soil conditions, as also highlighted by Godefroid and Koedam (2010). Accordingly,
we retain the status of invasive species as more appropriate for I. parviora in Toscana.
F. Roma-Marzio, M. D’Antraccoli, L. Peruzzi
Kolkwitzia amabilis Graebn. (Linnaeaceae)
+ (CAS) PIE: Avigliana (Torino), piazzola di sosta lungo la via che porta alla Sacra di San
Michele (WGS84: 45.065342N, 7.375007E), margine stradale, ca. 400 m, 8 May 2019,
M. Arnoul, M. Lonati (FI, TO). – Casual alien species new for the ora of Piemonte.
is ornamental species is native to China. It is reported in Italy as casual only for
Lombardia (Galasso et al. 2018a).
M. Arnoul, M. Lonati
Leucaena leucocephala (Lam.) de Wit subsp. glabrata (Rose) Zárate (Fabaceae)
+ (CAS) SAR: Sestu (Cagliari), loc. Su Moriscau, presso la strada provinciale (WGS84:
39.165891N; 9.044471E), aree incolte, 26 m, 11 July 2019, A. Lallai (FI, CAG);
Monserrato (Cagliari) (WGS84: 39.150639N, 9.084440E), bordo strada, 15 m, 13
July 2019, L. Podda (CAG). – Casual alien subspecies new for the ora of Sardegna.
is subspecies is native to central America and southern Mexico, and it was intro-
duced in many countries for several purposes, sometimes becoming invasive (Hughes
1998a, 1998b). In Italy, it has been reported as naturalized in Sicilia (Raimondo and
Domina 2007; Pignatti et al. 2017; Galasso et al. 2018a). In Sardegna, it has been ob-
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
72
served since 2006 in the industrial area of Sestu, where some plants are growing not far
from the cultivated parental plants. Some saplings and young trees have also been observed
in the surroundings of Monserrato, in fallow land and roadsides close to Via C. Cabras.
A. Lallai, L. Podda, G. Bacchetta
Ludwigia hexapetala (Hook. & Arn.) Zardini, H.Y.Gu & P.H.Raven (Onagraceae)
+ (NAT) LAZ: Bracciano (Roma), fraz. Vigna di Valle, Museo Storico dell’Aeronautica
Militare, sul Lago di Bracciano (WGS84: 42.085342N, 12.218902E), sulla spiaggia
e sulla riva del lago, 162 m, 12 June 2019, S. Buono (FI). – Naturalized alien species
new for the ora of Lazio.
Ludwigia hexapetala is a herbaceous perennial plant native to central and South
America; its habitat includes lakeshores, ponds, ditches, and streams. e large tolerance
of this species to the variations of hydrological and climatic conditions, as well as the
strong ability to colonize both beaches and swamps, make it a noxious invader of aquatic
ecosystems in North America and in Europe, where it is reported (as included in L.
grandiora (Michx.) Greuter & Burdet) in the list of invasive alien species of Union
concern (Regulation (EU) n. 1143/2014). It was recorded for Italy by Galasso (2007),
based on specimens collected in Lombardia and Veneto and, later, as invasive for Emilia-
Romagna (Alessandrini et al. 2017). is species is already established around the coasts of
Bracciano Lake, where large populations with hundreds of plants regularly develop owers
and fruits. Nowadays, it occurs with dense populations on about 2 km of the coast near
Vigna di Valle, together with other aliens, such as Amorpha fruticosa L., Datura wrightii
Regel, Eclipta prostrata (L.) L., Oenothera glazioviana Micheli, Physalis peruviana L., Salvia
hispanica L. (see also Galasso et al. 2018b, 2018c, 2019). Moreover, it is widespread near
Trevignano Romano (Roma), loc. Pantane, where it was wrongly reported as L. peploides
(Kunth) P.H.Raven subsp. montevidensis (Spreng.) P.H.Raven (Azzella and Iberite 2010).
Some individuals can be observed on the east coast of the lake (Lungolago di Polline).
S. Buono, M.M. Azzella, S. Magrini
Medicago ×varia Martyn (Fabaceae)
+ (CAS) TOS: Greve in Chianti (Firenze), Monte San Michele, Valico del Morellino
(WGS84: 43.550176N, 11.398473E), lungo strada, ca. 10 individui, 748 m, 27 June
2018, T. Fiaschi (FI). – Casual alien nothospecies new for the ora of Toscana.
is nothospecies (Medicago falcata L. subsp. falcata × M. sativa L.) is likely much
more widespread in Italy than currently recorded (Galasso et al. 2018a).
G. Bonari, T. Fiaschi, C. Angiolini
Muscari armeniacum Leichtlin ex Baker (Asparagaceae)
+ (CAS) PIE: Verbania (Verbano Cusio Ossola), fraz. Pallanza, Via Prossano, a
lato dell’ingresso dei Giardini Botanici di Villa Taranto (WGS84: 45.926149N,
Notulae to the Italian alien vascular ora: 8 73
8.565000E), muro di cinta, crepa nel cemento, 206 m, N, 22 April 2019, N.M.G.
Ardenghi, S. Mossini (FI). – Casual alien species new for the ora of Piemonte.
Muscari armeniacum, usually grown for ornamental purposes, is known as a
casual alien in dierent regions of northern and central Italy, except Piemonte (Ga-
lasso et al. 2018a). A single individual was found within the crack of a wall near
the entrance of the Botanical Gardens of Villa Taranto, where this species is widely
cultivated in owerbeds.
N.M.G. Ardenghi, S. Mossini
Nigella sativa L. (Ranunculaceae)
+ (CAS) PUG: Bari (Bari), Lungomare N. Sauro (WGS84: 41.122055N, 16.878997E),
fessure di marciapiede, 1 m, 22 May 2019, leg. G. Picella, det. L. Forte, R.P. Wagensommer
(BI No. 42139); ibidem, 17 June 2019, leg. L. Forte, det. L. Forte, R.P. Wagensommer
(FI, BI No. 42140). – Casual alien species new for the ora of Puglia.
Nigella sativa grows in many countries of the temperate regions, where it is cul-
tivated for its aromatic seeds (Zohary 1983). In Italy, it was already cultivated in An-
cient Rome (Arrigoni and Viegi 2011), and it is currently reported as a casual alien in
Sardegna, extinct in Piemonte, and not recently recorded for Friuli Venezia Giulia and
Toscana (Galasso et al. 2018a). In the latter region, Arcangeli (1882) already indicated
its occurrence in Casentino as doubtful. No recent information about cultivation of
this species in Puglia is available.
L. Forte, R.P. Wagensommer, G. Picella
Oenothera speciosa Nutt. (Onagraceae)
+ (NAT) MAR: Fano (Pesaro e Urbino), loc. Rosciano (WGS84: 43.822978N,
12.995544E), margini stradali e campi, ca. 28 m, 2 June 2017, L. Gubellini (FI,
PESA). – Naturalized alien species conrmed for the ora of Marche.
Oenothera speciosa is a showy perennial alien introduced as ornamental, native to
prairies in the United States of America (Missouri and Nebraska) and northern Mexico
(Wager et al. 2007; Keener et al. 2019). In Italy, this species is reported as casual alien
for Lombardia, Veneto, Toscana, and as naturalized for Emilia-Romagna (Galasso et al.
2018a). For Marche, the occurrence of a Oenothera with pink owers near Senigallia
was reported by G. Mazzueri (pers. commun.). e same data was later veried and
recorded by Montanari and Marconi (2010), but no precise locality information was
provided. In Rosciano, several specimens have been observed for some years along
roadsides and uncultivated areas, where they are slowly spreading.
L. Gubellini, N. Hofmann
+ (CAS) SIC: Augusta (Siracusa), centro urbano, villetta comunale tra Via A. Gramsci e
Via Papa Giovanni XXIII (WGS84: 37.247270N, 15.221336E), 25 May 2019, leg. R.
Romano, det. N.M.G. Ardenghi (FI). – Casual alien species new for the ora of Sicilia.
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
74
Many individuals grow in an abandoned urban garden, probably introduced a few
years ago for ornamental purposes in a small owerbed. Currently, O. speciosa displays
an 80% cover of the owerbed and is expanding in the surrounding areas.
R. Romano, O. Caldarella, A. La Rosa, F. Luchino, N.M.G. Ardenghi
Opuntia scheeri F.A.C.Weber (Cactaceae)
+ (CAS) UMB: Castiglione del Lago (Perugia), loc. Badiaccia, vicino alla sponda
del Lago Trasimeno, lungo la strada SR71, presso l’ingresso di un’abitazione privata
(WGS84: 43.168985N, 12.014299E), coltivata e spontaneizzata lungo un fosso, 261
m, 9 October 2018, M. Mugnai, S. Di Natale, G. Ferretti (FI). – Casual alien species
new for the ora of Umbria.
+ (CAS) TOS: Sesto Fiorentino (Firenze), fraz. Montorsoli, nei pressi della ex stazione
ferroviaria lungo la ferrovia Faentina (WGS84: 43.836202N, 11.284043E), pianta
spontaneizzata sulla scarpata al margine stradale, 265 m, 7 February 2019, M. Mugnai
(FI). – Casual alien species new for the ora of Toscana.
Opuntia scheeri is a species native to Mexico, often cultivated as an ornamental
plant. It was recorded for the rst time in Italy in 1994 (Guiggi 2008), and currently
occurs in several regions of northern Italy (Piemonte, Lombardia, Trentino-Alto Adige,
Veneto, Emilia-Romagna: Galasso et al. 2018a). Both the records reported here refer
to individuals growing close to inhabited areas and derived most likely from cultivated
plants. e record from Umbria refers to several well-established plants, probably orig-
inated by vegetative means from individuals cultivated nearby. e Tuscan occurrence,
instead, consists of a single established individual.
M. Mugnai, E. Corti, S. Di Natale
Paulownia tomentosa (unb.) Steud. (Paulowniaceae)
+ (CAS) TOS: Vaglia (Firenze), fraz. Fontebuona, ex stazione ferroviaria lungo la fer-
rovia Faentina (WGS84: 43.880886N, 11.289335E), numerose piante spontaneizzate
nei pressi dei marciapiedi e degli edici della stazione, 332 m, 8 November 2018, M.
Mugnai, A. Misuri, G. Ferretti (FI). – Casual alien species new for the ora of Toscana.
Paulownia tomentosa is an ornamental plant native to China and introduced to Eu-
rope. It is usually cultivated in parks and gardens, but it is also used for timber production
thanks to its fast growth and high-quality wood. e size of plantations in Italy has been
increasing rapidly since 1989 (Mezzalira and Colonna 2002). is species occasionally
escapes cultivation and becomes invasive, growing rapidly in disturbed areas. It is consid-
ered as invasive in the USA, and a potentially invasive species in Europe and South Amer-
ica, where it has been introduced (CABI 2019). We observed an abundant population at
the Fontebuona railway station, close to a large cultivated plant. e population consists
of numerous individuals of various ages, deriving from both seeds and root suckers. Re-
cently (May 8th, 2019) this species was detected in another site, on the right bank of the
Arno River in loc. Riscaggio (Reggello, Firenze, WGS84: 43.7249776N, 11.4662411E).
A. Misuri, L. Pinzani, G. Ferretti
Notulae to the Italian alien vascular ora: 8 75
Petroselinum crispum (Mill.) Fuss (Apiaceae)
+ (NAT) ITALIA (SAR). Status change from casual to naturalized alien for the ora
of Italy (Sardegna).
In Italy, Petroselinum crispum is reported for most of the regions (Galasso et al.
2018a). Although an agronomic study on populations naturalized in Trentino-Alto
Adige was published recently (Fusani et al. 2016), it is considered as casual alien at
national level. We detected numerous plants inhabiting steep and shady calcarenitic
clis at Capo Sant’Elia (Cagliari, Sardegna). is population displays a well-structured
partition in age classes, with seedlings, juveniles, and fruiting individuals that suggest
the establishment of a naturalized population. Interestingly, the presence in this area
of the phyto-toponym “su perdusemini”, clearly referring to parsley, and used at least
from the 18th century to name a tower probably built during the 16th century, suggests
that naturalized populations may be present in this area since a long time. However, P.
crispum was not previously recorded in the accurate ora of Capo Sant’Elia compiled
by Martinoli (1950). In this context, it must be pointed out that the origin of this
widely cultivated plant has not yet been identied with certainty, though it possibly
originates in the eastern or central Mediterranean region (Agyare et al. 2017; Pignatti
et al. 2018). It is noteworthy that Linnaeus (1753) stated its wild habitat to be Sarde-
gna, close to springs.
M.C. Fogu, M. Marignani, L. Rosati
Phyllostachys viridiglaucescens (Carrière) Rivière & C.Rivière (Poaceae)
– VDA. – Alien species to be excluded from the ora of Valle d’Aosta.
Phyllostachys viridiglaucescens in Valle d’Aosta was recorded for two locali-
ties (Mainetti and Ban 2018). Surveys in 2018 [Champdepraz (Aosta), terrazza-
menti abbandonati a ca. 300 m dalla fraz. Chef-Lieu (WGS84: 45.68873546N,
7.65795915E), terrazzamenti abbandonati, ca. 540 m, 7 October 2018, A. Mainetti,
S. Ravetto Enri, V.Mezzasalma (FI); Arnad (Aosta), boscaglia a lato della strada SS26
sul conne con il comune di Hône (WGS84: 45.624517N, 7.736778E), boscaglia
ripariale, ca. 350 m, 7 October 2018, A. Mainetti, S. Ravetto Enri, V. Mezzasalma
(FI)] revealed short oblique internodes at the base of the culms for both the locali-
ties. is is a distinctive feature of P. aurea Carrière ex Rivière & C.Rivière (Tison
and de Foucault 2014), a species already reported from Valle d’Aosta (Galasso et al.
2018a). Furthermore, the identity of this plant was conrmed by a DNA nger-
printing (RAPD) analysis performed by FEM2-Environment Company (spin-o of
the University of Milano-Bicocca) within the BambApp Project (BambApp 2019)
(Dipartimento di Scienze Agrarie, Forestali e Alimentari, Università di Torino), us-
ing samples from a private botanical collection (T. Froese: Cravanzana, Cuneo, Italy)
veried by us as reference base. Consequently, P. viridiglaucescens should be excluded
from the ora of Valle d’Aosta.
A. Mainetti, S. Ravetto Enri, V. Mezzasalma
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
76
Phyllostachys viridis (R.A.Young) McClure (Poaceae)
+ (NAT) PIE: Arona (Novara), fraz. Montrigiasco, zona Cascina Motto (WGS84:
45.77122N, 8.51728E), bosco a Robinia pseudoacacia dominante, ca. 425 m, 15 January
2018, M. Pittarello, A. Mainetti, F. De Mattia (FI). – Status change from casual to natu-
ralized alien for the ora of Italy; naturalized alien species new for the ora of Piemonte.
+ (NAT) VDA: Antey-Saint-André (Aosta), loc. Filey, nei pressi della riva orograca
sinistra del Torrente Marmore (WGS84: 45.81154804N, 7.58866366E), prato da
sfalcio, ca. 1030 m, 21 July 2018, M. Lonati, S. Pirani, J. Frigerio (FI). – Naturalized
alien species new for the ora of Valle d’Aosta.
According to Galasso et al. (2018a), Phyllostachys viridis was previously reported in Ita-
ly only for Lombardia. Its identity was conrmed by a DNA ngerprinting (RAPD) anal-
ysis performed by FEM2-Environment Company (spin-o of the University of Milano-
Bicocca) within the BambApp Project (BambApp 2019), using samples from a private
botanical collection (T. Froese: Cravanzana, Cuneo, Italy) veried by us as reference base.
ese populations originated from agamic propagation of nearby cultivated plants.
M. Pittarello, A. Mainetti, F. De Mattia, M. Lonati, S. Pirani, J. Frigerio
Physalis angulata L. (Solanaceae)
+ (CAS) UMB: Otricoli (Terni), area archeologica Utriculum, riva idrograca sinistra
del Fiume Tevere (WSG84: 42.408889N, 12.458889E), coltivo, 42 m, 30 July 2015,
E. Scarici, M. Scarici (FI). – Casual alien species new for the ora of Umbria.
Physalis angulata is a tropical American species that it is occasionaly cultivated for
its edible fruits (Hawkes 1972). It is reported in Italy only in Lombardia, Veneto, and
Lazio (Galasso et al. 2018a). Many individuals grow in cultivated areas along the river.
E. Scarici, M. Scarici
Pseudosasa japonica (Siebold & Zucc. ex Steud.) Makino ex Nakai (Poaceae)
+ (NAT) VDA: Châtillon (Aosta), traversa di Strada Chemin de Barat che porta alla
stazione di servizio autostradale (direzione Aosta) (WGS84: 45.747689N, 7.618339E),
prato da sfalcio, ca. 490 m, 13 October 2018, S. Ravetto Enri, M. Lonati, L. Guzzetti
(FI). – Naturalized alien species new for the ora of Valle d’Aosta.
In Italy, Pseudosasa japonica was reported for all northern regions with the exception of
Liguria and Valle d’Aosta (Galasso et al. 2018a). Single branches per node and palmfont-
like leaves clearly permitted to identify the species (Li et al. 2006; Tison and de Foucault
2014). In addition, the identity was conrmed by a DNA ngerprinting (RAPD) analysis
conducted by FEM2-Environment Company (spin-o of the University of Milano-
Bicocca) within the BambApp Project (BambApp 2019), using samples from a private
botanical collection (T. Froese: Cravanzana, Cuneo, Italy) veried by us as reference base.
e recorded population originated from agamic propagation of nearby cultivated plants.
S. Ravetto Enri, M. Lonati, L. Guzzetti
Notulae to the Italian alien vascular ora: 8 77
Quercus rubra L. (Fagaceae)
+ (CAS) SAR: Villagrande Strisaili (Nuoro), Monte Idolo (WGS84: 39.940833N,
9.488056E), graniti, 881 m, 2 June 2019, G. Bacchetta (FI, CAG). – Casual alien spe-
cies new for the ora of Sardegna.
e red oak is an American taxon, which was imported in Europe starting from the
17th century (Magni Diaz 2004), and in Italy from 1803 (Maniero 2015). In Sardegna,
it was introduced in reforestations and for ornamental purposes (Veri and Bruno 1974;
Arrigoni 2006). In recent years, numerous trees and saplings were found on the eastern
side of the Gennargentu Massif (Monte Idolo), all growing close to reforestations with
red oak and other alien trees.
G. Bacchetta, G. Calvia, L. Podda
Reynoutria bohemica Chrtek & Chrtková (Polygonaceae)
+ (NAT) MAR: Urbino (Pesaro e Urbino), lungo la strada SS73bis (WGS84:
43.730503N, 12.635836E), scarpata stradale, ca. 410 m, 16 November 2018, N. Hof-
mann (FI, PESA). – Naturalized alien species new for the ora of Marche.
Reynoutria bohemica is of hybrid origin between the alien species R. japonica Houtt.
and R. sachalinensis (F.Schmidt) Nakai, and it has been recognized and described only
at the end of the last century in the Czech Republic (Chrtek and Chrtková 1983). Like
other congener species, R. bohemica colonizes ruderal environments, roadsides and
waterways, and forms dense stands that shade and crowd out all other plants, thereby
reducing the biodiversity of invaded plant communities and damaging habitats beyond
repair (Padula et al. 2008). In Italy, it has been reported, so far, for Valle d’Aosta, Pie-
monte, Lombardia, Veneto, and Toscana as invasive alien, for Friuli Venezia Giulia and
Emilia-Romagna as naturalized alien, and for Trentino-Alto Adige, Liguria as casual
alien (Galasso et al. 2018a). In the Urbino site, which represents the rst record for
Marche, a large number of individuals has been monitored for several years, and a
considerable increase of the population was observed. For this reason, containment
measures should be taken.
L. Gubellini, N. Hofmann
Roldana petasitis (Sims) H.Rob. & Brettell (Asteraceae)
+ (CAS) SIC: Librizzi (Messina), Via A. Cullura (WGS84: 38.096521N,
14.957978E), su scarpata stradale alberata con suolo profondo, 8 April 2019, C.D.
Rici (FI). – Casual alien species conrmed for the ora of Sicilia.
Roldana petasitis is native to central America (Jerey 1986). According to Galasso
et al. (2018a), this species is naturalized in Liguria, while in Lazio, Puglia, and Ba-
silicata it is considered as a casual alien. Although Fiori (1927) reported this taxon as
growing wild in Sicilia, Giardina et al. (2007) excluded it from this region. A few indi-
viduals of dierent age were found in Librizzi, growing along the roadside with other
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
78
nitrophilous species typical of urban areas. e population, monitored since 2013, is
particularly resilient, despite the continuous cuts made during ordinary maintenance
of public owerbeds. In Sicilia, this species occurs also in Siracusa, at Latomia dei Cap-
puccini, in a limestone quarry (R. Genovese, pers. commun.).
C.D. Rici, A. La Rosa, O. Caldarella, F. Luchino
Saccharum biorum Forssk. (Poaceae)
+ (NAT) PUG: Melendugno (Lecce) lungo la Circonvallazione di Melendugno
(WGS84: 40.270194N, 18.343408E), bordo strada, 35 m, 22 August 2019, G.
Laghetti, G. Maruca, C.M. Musarella (REGGIO). – Naturalized alien species con-
rmed for the ora of Puglia.
For Italy, Saccharum biorum was known, until now, only in Sicilia and Sardegna,
whereas it was not, until recently, recorded in Puglia (Galasso et al. 2018a). A popu-
lation was found also in Puglia, between a road and an abandoned eld, covering a
surface of about 20 m2. Due to its extension and to the number of the owering stems,
we can consider this species as naturalized in this locality.
C.M. Musarella, G. Maruca, G. Laghetti
Sedum palmeri S.Watson (Crassulaceae)
+ (CAS) PIE: Verbania (Verbano Cusio Ossola), fraz. Pallanza, Viale G. Azari (WGS84:
45.928311N, 8.552684E), marciapiede, 214 m, 22 April 2019, N.M.G. Ardenghi, S.
Mossini (FI). – Casual alien species new for the ora of Piemonte.
Sedum palmeri, commonly cultivated as an ornamental pot plant, has been record-
ed from many northern Italian regions, except Piemonte (Galasso et al. 2018a). Some
individuals were discovered growing within the cracks of a sidewalk. is species may
be more widespread across the region, especially in urban areas.
N.M.G. Ardenghi, S. Mossini
+ (CAS) TOS: Figline e Incisa Valdarno (Firenze), loc. C. Torrione (WGS84:
43.6586857N, 11.4246546E), interno cipresseta, 310 m, 24 February 2019, L. Pin-
zani (FI). – Casual alien species new for the ora of Toscana.
In Italy, Sedum palmeri is recorded from Lombardia, Veneto, Friuli Venezia Giulia,
Emilia-Romagna, Liguria, Lazio, Campania, and Sardegna (Galasso et al. 2018a).
Various groups of individuals grow within a cypress wood. e main one is represented
by more than 100 individuals.
L. Pinzani
Semiarundinaria fastuosa (Lat.-Marl. ex Mitford) Makino (Poaceae)
+ (NAT) ITALIA (PIE): Borgo San Dalmazzo (Cuneo), Via Mangiacane (WGS84:
44.34109541N, 7.50195817E), canale di irrigazione e margine di seminativo, ca.
Notulae to the Italian alien vascular ora: 8 79
620m, 1 February 2018, M. Pascale, G. Nota, V. Mezzasalma (FI). – Naturalized alien
species new for the ora of Italy (Piemonte).
Semiarundinaria fastuosa is a bamboo native to Japan (south-western Honshu).
e recorded population originated from agamic propagation from a private garden
and colonized a nearby canal bank. Several branches per node, partially deciduous
culm sheaths and minute auricles allowed us to identify this species (Li et al. 2006;
Tison and de Foucault 2014). e identication was conrmed by DNA ngerprint-
ing (RAPD) analysis performed by FEM2-Environment Company (spin-o of the
University of Milano-Bicocca) within the BambApp Project (BambApp 2019), using
samples from a private botanical collection (T. Froese: Cravanzana, Cuneo, Italy) veri-
ed by us as reference base.
M. Pascale, G. Nota, V. Mezzasalma
Senecio angulatus L.f. (Asteraceae)
+ (CAS) ABR: San Vito Chietino (Chieti), loc. Marina, muro di contenimento coperto
da vegetazione presso la strada SS16 Adriatica (WGS84: 42.305208N, 14.450116E),
ca. 15m, SW, 18 May 2019, N. Olivieri (FI). – Casual alien species new for the ora
of Abruzzo.
Senecio angulatus is a succulent climbing plant native to South Africa, introduced
for ornamental purposes in southern Europe, Macaronesia, northern Africa, Califor-
nia, Chile, Australia, and New Zealand. Currently, it is naturalized in Albania (Barina
et al. 2011), Croatia (Milović et al. 2010), Iberian peninsula (Romero Buján 2007;
Pyke 2008), and Chile (Ugarte et al. 2011) and is considered one of the most invasive
species in the western Mediterranean area (Brundu et al. 1999), Mediterranean France
(Brunel and Tison 2005), Australia (Ross and Walsh 2003; Randall 2007), and New
Zealand (Bergin 2006). is species was introduced in Italy in 1875 (Maniero 2015).
It is known as a casual alien in Lazio and Calabria, while it is naturalized in Puglia,
Campania, Basilicata, Sicilia, and invasive in Liguria, Toscana, and Sardegna (Galasso
et al. 2018a). In San Vito Chietino, this species grows on a brick retaining wall, located
below the site of the Adriatic State Road, in a sunny and sheltered position, close to the
Adriatic Sea. Here the plant is established along with Arundo plinii Turra, Ficus carica
L., and Rubus ulmifolius Schott.
N. Olivieri
Senecio inaequidens DC. (Asteraceae)
+ (INV) TOS. – Status change from naturalized to invasive alien for the ora of Toscana.
Senecio inaequidens is native to South Africa. It was recorded in Europe for the rst
time in the mid-twentieth century and observed in Italy in 1947 (Carrara Pantano and
Tosco 1959; Anzalone 1976). It was reported as present throughout central and northern
Italy and has been rapidly expanding since the beginning of the 1980s (Pignatti 1982).
Now it is widespread in all Italian regions and often considered invasive (Galasso et al.
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
80
2018a). Our recent eld investigations revealed the presence of this species in all Tuscan
provinces, conrming many previous observations and adding several new occurrences
(Peruzzi et al. 2019). Consequently, this species is abundant and well distributed in
anthropized sites of Toscana, where it is spreading notwithstanding the control actions
often undertaken. Moreover, this species has been observed in some natural sites.
Accordingly, we regard the status of invasive alien as the most appropriate.
A. Misuri, G. Ferretti, M. Mugnai
Sisyrinchium rosulatum E.P.Bicknell (Iridaceae)
+ (CAS) SAR: Olbia (Sassari), Parco F. Noce, presso il canale di Via L. Galvani
(WGS84: 40.554511N, 9.295523E), prati e aiuole, 1–2 m, 25 June 2017, G. Cal-
via (FI); ibidem, Parco F. Noce, lato Via G. D’Annunzio, ai lati della pista (WGS84:
40.554352N, 9.300366E), aiuole e prati inglesi, 2 m, 25 June 2017, G. Calvia (Herb.
G. Calvia). – Casual alien species new for the ora of Sardegna.
Sisyrinchium rosulatum is a species native to North America, introduced in Europe
and other continents, and now naturalized in several countries (Nicolella and Ardenghi
2013). In Italy, this species has been reported as casual alien in Lazio (Nicolella and
Ardenghi 2013; Galasso et al. 2018a). In Sardegna, it has been observed starting to
2015 in the town of Olbia, where it grows in the Fausto Noce community park and
neighboring areas, above all in lawns but also in owerbeds and along paths. It prob-
ably arrived there thanks to seed dispersed in lawns.
G. Calvia
Solanum bonariense L. (Solanaceae)
+ (CAS) LIG: Genova (Genova), lungo Via Apparizione, nel tratto pedonale (WGS84:
44.40443N, 8.98889E), bordo strada, 42 m, 20 April 2019, A. Di Turi, C. Aristarchi
(FI, GE, GDOR). – Casual alien species new for the ora of Liguria.
Solanum bonariense is a perennial shrub native to Uruguay, northern Argentina,
and southern Brazil where it is widespread in pastures. Introduced in Europe as an
ornamental, it is nowadays recorded in Italy as a casual species for Lombardia, Lazio,
Campania, and as naturalized for Toscana and Sicilia (Galasso et al. 2018a). A well-
developed specimen, growing together with Parietaria judaica L., has been recorded
in a pedestrian street of Genova among houses surrounded by orchards and gardens.
A. Di Turi, C. Aristarchi
Solanum laciniatum Aiton (Solanaceae)
+ (NAT) ITALIA (TOS): Monte Argentario (Grosseto), lungo la strada sterrata Via
Panoramica, sopra Cala dell’Acqua Dolce (WGS84: 42.374541N, 11.185636E),
macchia, 70 m, 22 June 2019, F. Roma-Marzio, P. Liguori (FI, Herb. F. Roma-
Marzio). – Naturalized alien species conrmed for the ora of Italy and new for the
ora of Toscana.
Notulae to the Italian alien vascular ora: 8 81
Solanum laciniatum is a species native to New Zealand and Australia from south-east-
ern Australia, Victoria, and Tasmania (Simon 1981). is species belongs to Solanum subg.
Archaesolanum Bitter ex Marzell, composed of eight species occurring only in the SW-Pacif-
ic region (Poczai et al. 2011). In the Euro+Med area, S. laciniatum is recorded in Morocco,
France, Spain, Israel, and Tunisia (Valdés 2012), whereas in Italy it is doubtfully occurring
based on a record for Puglia (Beccarisi et al. 2015; Galasso et al. 2018a). is species is
similar to S. aviculare G.Forst, that mainly diers from S. laciniatum in the shape of petals
(notched in S. laciniatum and acute in S. aviculare), and in the colour of mature fruits (or-
ange-yellow in S. laciniatum and orange-red to scarlet in S. aviculare). About six big tufts,
probably originated from cultivated plants at a nearby hotel, were counted mixed with na-
tive species typical of the Mediterranean scrub. Furthermore, in the same area plants are pre-
sent since 2006, as highlighted by some photos published on the Portal to the Flora of Italy
(http://dryades.units.it/oritaly/index.php?procedure=taxon_page&tipo=all&id=11471).
F. Roma-Marzio
Sporobolus indicus (L.) R.Br. (Poaceae)
+ (CAS) MAR: Piobbico (Pesaro e Urbino), alla conuenza tra il Torrente Biscubio e il Fi-
ume Candigliano (WGS84: 43.589956N, 12.510999E), greto uviale, ca. 335 m, 10 De-
cember 2018, N. Hofmann (FI, PESA). – Casual alien species new for the ora of Marche.
is perennial grass is naturalized throughout peninsular and insular Italy, except
for Valle d’Aosta, Marche, Umbria, and Puglia (Galasso et al. 2018a). In Marche, a few
individuals grow in the grassy edge of a riverbed on alluvial sandy soil. e occurrence of
Sporobolus indicus could be due to the abundant presence in the site of migratory birds
(especially ducks), that inhabit riverbanks and contribute to the conveyance of seeds.
L. Gubellini, N. Hofmann
Sporobolus vaginiorus (Torr. ex A.Gray) Alph.Wood (Poaceae)
+ (NAT) TOS: Fiesole (Firenze), fraz. Caldine, stazione ferroviaria di Caldine-Fiesole
(WGS84: 43.830543N, 11.308060E), marciapiedi lungo il binario, 169 m, 8 Novem-
ber 2018, M. Mugnai, A. Misuri, G. Ferretti (FI). – Naturalized alien species new for
the ora of Toscana.
Sporobolus vaginiorus is a North American species already present in most of
northern regions of Italy (Galasso et al. 2018a). e population reported here dis-
played several mature fruiting individuals spanning alongside the sidewalks of the
Caldine-Fiesole railway station and in the surrounding areas.
M. Mugnai, S. Di Natale, A. Padula
Tulipa clusiana Redouté (Liliaceae)
+ (NAT) VEN: Soave (Verona), alla ‘Colombara’ (WGS84: 45.44168063N, 11.24953327E),
boschetto termolo, 168 m, 2 March 2019, G. Bommartini, G. Zanoni, F. Menini (VER
No. FDC7708). – Status change from casual to naturalized alien for the ora of Veneto.
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
82
Tulipa clusiana is native to Syria and Persia, in the Middle East (Ban and Galasso
2010), and is recorded as a casual alien in several central-northern Italian regions, and
as naturalized in Piemonte, Lombardia, and Marche (Galasso et al. 2018a). In Veneto,
there was only one conrmed report by Busnardo (2000) in Bassano del Grappa
(Vicenza). For the Verona province, there is only a historical sample collected by Goiran
(1897, 1900, VER) and a recent indication of occasional presence in Custoza (F. Prosser,
pers. commun.). In the locality reported here, the population consists of thousands of
seedlings, which grow both within a thermophilic grove formed by dierent species,
such as Dioscorea communis (L.) Caddick & Wilkin, Fraxinus ornus L. subsp. ornus,
Ligustrum vulgare L., Quercus pubescens Willd. subsp. pubescens, Robinia pseudoacacia
L., Rubus ulmifolius Schott, and Sambucus nigra L., and inside olive groves. is
species was found in two small woods about 250 meters apart, and more on two other
adjacent banks. Other localities have been found on the slopes of Monte Tenda, just
above the medieval castle of Soave (WGS84: 45.44145545N, 11.24924856E, 95 m),
more than 2 km away from the above-mentioned sites. e total area occupied, albeit
discontinuously, by T. clusiana is over 10,000 m2 and hosts thousands of individuals.
G. Bommartini, G. Zanoni, F. Menini, S. Andreatta
Vachellia farnesiana (L.) Wight & Arn. (Fabaceae)
+ (CAS) CAL: Bova Marina (Reggio Calabria), loc. Vena (WGS84: 37.937774N,
15.911936E), scarpata bordo strada, 44 m, 27 April 2019, leg. V.L.A. Laface, det. V.L.A.
Laface, C.M. Musarella, G. Spampinato (FI, REGGIO); Reggio Calabria (Reggio Calabria),
Gallico, loc. Pietre della Zita (WGS84: 38.161215N, 15.663414E), scarpata bordo stra-
da, 47 m, 9 October 2019, V.L.A. Laface (REGGIO); Brancaleone (Reggio Calabria), loc.
Fiumarella (WGS84: 37.982290N, 16.089573E), bordo strada, 35 m, 28 October 2019,
V.L.A. Laface (REGGIO). – Casual alien species new for the ora of Calabria.
e native range of Vachellia farnesiana is considered to be the New World (New
1984), and in particular North America (Gilman and Watson 1993). However, its exact
origin is nowadays debated (Luken and ieret 1996; Roskov 2006). In Europe, it oc-
curs in France, Italy, and Spain (Roskov 2006). Currently, according to Galasso et al.
(2018a), it is a casual alien in Sicilia and Sardegna. In this new Calabrian locality, we ob-
served several seedlings near the mature plants. is is the rst record for peninsular Italy.
C.M. Musarella, V.L.A. Laface, G. Spampinato
Verbena bonariensis L. (Verbenaceae)
+ (CAS) FVG: Gorizia (Gorizia), Borgo Castello, sulle mura del castello subito dopo
Porta Leopoldina (WGS84: 45.942638N, 13.628783E), su mura di arenaria, 100 m,
25 April 2019, F. Roma-Marzio, P. Liguori (FI, Herb. F. Roma-Marzio). – Casual alien
species new for the ora of Friuli Venezia Giulia.
Verbena bonariensis is native to South America (southern Brazil, Uruguay, Para-
guay, northern Argentina) and has been introduced in many countries of Africa, Asia,
Notulae to the Italian alien vascular ora: 8 83
Australia, and Europe and in the USA (Munir 2002; Nesom 2010). In Italy, it is re-
ported as naturalized alien in Liguria and as casual in Lombardia, Trentino-Alto Adige,
Emilia-Romagna, Toscana, Umbria, and Lazio (Galasso et al. 2018a). About ve plants
were found on the ancient walls, probably as a result of escaped cultivated plants.
Specimens were identied using the key reported by Nesom (2010).
F. Roma-Marzio
Youngia japonica (L.) DC. subsp. japonica (Asteraceae)
+ (CAS) SIC: Messina (Messina), Rodia, loc. Contrada Marmora, presso il complesso
residenziale Baia Verde (WGS84: 38.267442N, 15.478063E), fessure nella
pavimentazione del marciapiede e interstizi tra marciapiede e muro, 14 February 2019,
F. Luchino (FI). – Casual alien species new for the ora of Sicilia.
According to Shi and Kilian (2011), the Sicilian populations of Youngia japonica
belong to the autonymic subspecies, native probably to China and naturalized in warm
areas of all continents (Galasso et al. 2016). e single Italian record of this species
in Genova (Liguria) is very recent (Galasso et al. 2016). We found approximately 30
individuals growing inside sidewalk cracks and in shady micro-soil located at the base
of the walls. In the same area, the herbaceous vegetation consists mainly of several
ruderal species linked to anthropic environments. Y. japonica has been observed as
alien also in north-eastern Sicilia (A. Crisafulli and R.M. Picone, pers. commun.),
namely in Messina along urban roads (Via F. Bisazza), in the owerbeds and lawns of
the Comando Arma dei Carabinieri (near Villa Mazzini) and in Milazzo (Messina) at
C.da Scaccia in an uncultivated wet habitat.
F. Luchino, O. Caldarella, A. La Rosa, R. De Luca
Nomenclatural and distribution updates from other literature sources
Nomenclatural, status, and distribution updates according to Saccardo (1909), Viegi
et al. (1974), Ricciardi and Anzalone (1988), Greuter et al. (1989), Prosser et al.
(2009), Castellano and Spadaro (2011), Licitra and Napoli (2011), Schaefer and
Renner (2011), Himmelreich et al. (2012), Pasta (2012), Sebastian et al. (2012),
Coulot and Rabaute (2013), Gestri and Peruzzi (2016), Pasta et al. (2016), iede
(2017), Ardenghi (2018), Carta et al. (2018), Gallo et al. (2018), Rich et al. (2018),
Trejo-Torres et al. (2018), Antonietti and Dellavedova (2019), Ardenghi (2019),
Badalamenti (2019), Benetti (2019), Berselli et al. (2019), Buccheri et al. (2019),
Compton et al. (2019), Del Guacchio et al. (2019), Galasso (2019), Gallo (2019),
Gariboldi and Frezzini (2019), Groom (2019), Marchetti (2019), Musarella (2019),
Musarella et al. (2019), Pascale and Pellegrino (2019), Paton et al. (2019), Picco et
al. (2019), Prosser et al. (2019), Sarmati et al. (2019), Stinca and Mei (2019), Stinca
et al. (2019), Verloove et al. (2019), and corrections to Galasso et al. (2018a) are
provided in Suppl material 1.
G. Galasso, F. Bartolucci
Gabriele Galasso et al. / Italian Botanist 8: 63–93 (2019)
84
Acknowledgements
We gratefully acknowledge Enrico Ban, Maurizio Bovio, and Filippo Prosser, who
provided distribution, nomenclatural and taxonomic suggestions.
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Supplementary material 1
Supplementary data
Authors: Gabriele Galasso, Fabrizio Bartolucci
Data type: species data
Explanation note: 1. Nomenclatural updates; 2. Note updates; 3. Distribution up-
dates; 4. Synonyms, misapplied or included names.
Copyright notice: is dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). e Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/italianbotanist.8.48621.suppl1
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