A CONTRIBUTION TO KNOWLEDGE OF BALKAN LEPIDOPTERA: SOME NEW AND RARE SPECIES FOR ALBANIA AND NORTH MACEDONIA (MACROLEPIDOPTERA)

Abstract

Data for 45 nocturnal Lepidoptera species collected mainly in June-August 2018 by the authors are presented. The majority of reported species are from Albania, from where six genera and 19 species are reported for the first time. Seventeen other species are reported for the second time for Albania or are confirmed for the country. Three species are new for the North Macedonia (formerly known as the yugoslav Republic of Macedonia). Two other species are reported for the second time for that country. Reported species and collecting localities are illustrated in colour, when necessary with genitalia, including everted vesicas.

Full text

242 Entomologist’s Rec. J. Var. 131 (2019)
A CONTRIBUTION TO KNOWLEDGE OF BALKAN
LEPIDOPTERA: SOME NEW AND RARE SPECIES FOR ALBANIA
AND NORTH MACEDONIA (MACROLEPIDOPTERA)
STOYAN BESHKOV 1 & ANA NAHIRNIĆ 2
National Museum of Natural History, Tsar Osvoboditel Blvd.1, 1000 Sofia, Bulgaria
stoyan.beshkov@gmail.com; ananahirnic@nmnhs.com
Abstract
Data for 45 nocturnal Lepidoptera species collected mainly in June-August 2018 by the
authors are presented. The majority of reported species are from Albania, from where
six genera and 19 species are reported for the first time. Seventeen other species are
reported for the second time for Albania or are confirmed for the country. Three species
are new for the North Macedonia (formerly known as the Yugoslav Republic of
Macedonia). Two other species are reported for the second time for that country.
Reported species and collecting localities are illustrated in colour, when necessary with
genitalia, including everted vesicas.
Keywords: Albania, North Macedonia, faunistics, Lepidoptera, Macrolepidoptera
Introduction
Survey of Lepidoptera in the southern Balkans, and especially in Albania, is
ongoing. Results of trips from September to November have been published
(Beshkov & Nahirnić, 2019) and results of a trip in May-June, with C. W. Plant,
P. Jakšić and A. King, will be published soon. A paper concerning the Pyraloidea
is also in preparation (C. W. Plant et al., in prep.).
This present paper reports the results of five collecting trips to Albania in
June, July and August 2018 by the authors, as well as some additional results
from 2016 and 2017. Additional data from an expedition during August 2018
with C. W. Plant and his colleagues will also be published separately. In 2018
we extended research to new parts of the country, mostly high mountain areas.
The Albanian Lepidoptera fauna, particularly moths, still remains poorly
explored, although in the last three years we have reported more than 120 new
macrolepidoptera species for the country. After 2019, the number of the possible
new species for the country is expected to decrease markedly, but there are large
areas without any data or with old records only. Some species, recorded in only
a few localities, may be widespread. The lack of native researchers in Albania
is unfortunate as is the fact that recent research by foreign workers remains
unpublished.
Collecting methods are described in details in Beshkov & Nahirnić (2018a,
2018b, 2019). Genitalia slides of Nola, Meganola and Eilema rungsi were
photographed by Dr. B. Zlatkov using Carl Zeiss Jena Amplival Microscope.
All material published here is deposited in the collection of Stoyan Beshkov in
the National Museum of Natural History, Sofia (NMNHS).
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 242

243
Entomologist’s Rec. J. Var. 131 (2019)
Collecting localities
Many of the collecting localities have been mapped, described and illustrated
already (Beshkov, 2018; Beshkov & Nahirnić, 2018a, 2018b, 2019). These are
summarised in Table 1. Map 1 indicates the location of the 2018 localities.
1.Above Theth, Shkodra County, Boga
district, Bjeshkët e Nemuna Mts
(= Prokletije Mts), Radohima Mt.,
between Qafa e Thorës Pass and Theth
Village, south-east of Shtegu
Peak, 1657m, N42.3854°: E19.7502°,
limestone rocky area above coniferous
forest (Plate 1: 1).
2.Qafa e Selitës, Malësia e Tiranës,
(Highlands of Tirana), below Maja e
Fekenit (Feken peak), west from Qafa
e Selitës Pass, 1133m, N41.3729°:
E19.9992°, slopes with Quercus,
Carpinus, Juniperus oxycedrus (Plate
2: 2).
3.Mali me Gropë, 1501m, Mali me
Gropë Mts, Central part south of
plateau, 1501m, N41.3617°: E20.0568°,
Limestone area with dolines, mountain
steppe-like grasslands with single Acer
trees (Plate 1: 2).
4.Mali me Gropë, 1405m – Mali me
Gropë Mt., above Shen Meri, 1405 m,
N41.3524°: E20.0465°, stonny slopes
with Artemisia alba, Saturea,
Juniperus, etc. (Plate 2: 1).
5. Tomori, 2379m, Mt. Tomor, Tomor peak (Abaz Aliu peak), 2379m, N40.6361°:
E20.1615°, high mountain dry grassy slopes and rocky areas (Plate 3: 1).
6. Tomori, 1739m, Mt. Tomorr, Tomorr peak (Abaz Aliu peak, below to south,
N40.6111°: E20.1885°, 1739m, mountain grassy slopes and rocky areas (Plate 3: 2).
7. Ossumi, Ossumi river canyon, Çorovodë, opposite Çerenisht village, 419m,
N40.4846°: E20.2362°, Maquis: Phyllirea, Arbutus unedo, Pistacia terebinthus,
Quercus ilex, Quercus sp., Palliurus, Cotinus coggigria, Juniperus oxycedrus,
Carpinus orientalis, etc. (Plate 4: 1).
8. Mal Çajup, Gjirokaster County, Mt. Lunxhërisë, between Mal Çajup and Erind
Village, 1015m, N40.1825°: E20.1661°, limestone mountain steppe-like grasslands
with Quercus, Carpinus, Acer trees around (Plate 4: 2).
Map 1. New in 2018 collecting localities in
Albania.
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 243

244 Entomologist’s Rec. J. Var. 131 (2019)
Inventory of species
All entries relate to Albania, unless otherwise stated to the contrary. Data are
presented chronologically.
Drepanidae Boisduval, 1828
Cilix asiatica O. Bang-Haas, 1907 Vukopoles, 25.ix.2016; Prespa Lake, 18.viii.2017,
1. In Albania known only from Shkallë (Beshkov & Nahirnić, 2018b). Second report
for Albania.
Geometridae Stephens, 1829
Proteuchloris neriaria (Herrich-Schäffer, 1852) Syri i Kaltër, 6.viii.2018, 1(Plate 5:
1). Reported for Albania from Tirana, Marikaj near Tirana, Fush-Kuqe and Kavajë
(Misja, 1977). We suppose possible misidentification because in the work by Misja
Comibaena bajularia ([Denis & Schiffermüller], 1775) is not included.
Pachycnemia tibiaria (Rambur, 1829) Vukopoles, 25.ix.2016, 1(Plate 5: 2). New
species for Albania.
Dyscia raunaria (Freyer, 1851) Boboshticë, 19.v.2017, 2 (Plate 5: 3), Gen. prep.
3./28.II.2018, S. Beshkov, genitalia; Shkallë, 29.vi.2017, Gen. prep. 1./14.II.2018,
S. Beshkov, genitalia (Plate 10: 3). Known in Albania from Mali me Gropë
(Urbahn, 1966). Second report for Albania.
Locality Number
in map Source of map
Ardenica 9 Beshkov & Nahirnić (2019)
Bistricë 3 Beshkov (2018)
Bistrice - Syri i Kaltër 15 Beshkov & Nahirnić (2018b)
Boboshticë 8 Beshkov & Nahirnić (2018b)
Butrint 30 Beshkov & Nahirnić (2018a)
Dardhë 9 Beshkov & Nahirnić (2018b)
Dishnicë 6 Beshkov & Nahirnić (2018b)
Drenovë 7 Beshkov (2018)
Ilias 8 Beshkov (2018)
Klocë 1 Beshkov (2018)
Kolonjë 13 Beshkov & Nahirnić (2018b)
Kozjak Venec - Beshkov & Micevski (2019)
Palasë 5 Beshkov (2018)
Plani i Bardhë 4 Beshkov & Nahirnić (2019)
Prespa Lake 9 Beshkov & Nahirnić (2018a)
Pustec, 862m 4 Beshkov & Nahirnić (2018b)
Skallë 1 Beshkov & Nahirnić (2018b)
Stan Karbunarë 4 Beshkov (2018)
Starë 1 Beshkov & Nahirnić (2019)
Strelcë 6 Beshkov (2018)
Syri i Kaltër 10 Beshkov & Nahirnić (2019)
Vukopoles 4 Beshkov & Nahirnić (2018a)
Zvezdë 2 Beshkov (2018)
Table 1. Key to localities mapped in earlier works.
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 244

245
Entomologist’s Rec. J. Var. 131 (2019)
Plate 1. Survey areas. 1. Above Theth, 1657m; 2. Mali me Gropë, 1501m.
1
2
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 245

246 Entomologist’s Rec. J. Var. 131 (2019)
Plate 2. Survey areas. 1. Above Shen Meri, Mali me Gropë, 1405m; 2. Qafa e Selitës, 1133m.
1
2
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 246

247
Entomologist’s Rec. J. Var. 131 (2019)
Plate 3. Survey Areas. 1. Tomori summit, 2379m; 2. Tomori, below summit to the south, 1739m.
1
2
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 247

248 Entomologist’s Rec. J. Var. 131 (2019)
Plate 4. Survey Areas. 1. Ossumi River, near Çerenisht village; 2. Mal Çajup, 1015m.
1
2
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:29 Page 248

249
Entomologist’s Rec. J. Var. 131 (2019)
Protorhoe unicata (Guenée, 1858) Bistricë - Syri i Kaltër, 13.v.2017, 1, wingspan 19mm,
Gen. prep. 1./8.III.2018, S. Beshkov, genitalia on glass in euparal. Genitalia length:
3.25mm, cornutus length: 0.25mm; Ilias, 15.v.2017, 2 (Plate 5: 4), wingspan 18-
19mm, Gen. prep. 1./9.III.2018, S. Beshkov, genitalia (Plate 10: 4). Genitalia length:
3.00mm, cornutus length: 0.3mm; above Bistricë Village, 116m, 7.vi.2016, 4,
wingspan 18-20mm; Butrint, 14.v.2017, 1and 4, Gen. prep. 1./10.III.2018, S.
Beshkov, genitalia (Plate 10: 5). Protorrhoe unicata is reported as a new for Albania
by Urbahn (1966), although it is presented as a synonym of P. corollaria in Rebel &
Zerny (1931). After examination of genitalia here we confirm correct identification and
the presence of this species in Albania after the recent taxonomic revision of the genus
Protorhoe Herbulot, 1951 by Rajaei, Stadie & Hausmann (2017).
Plate 5. 1. Proteuchloris neriaria, . Syri i Kaltër; 2. Pachycnemia tibiaria, . Vukopoles; 3.
Dyscia raunaria, . Boboshticë; 4. Protorhoe unicata, . Ilias; 5. Protorhoe corollaria, . Strelcë;
6. Eupithecia carpophagata, . Mali me Gropë, 1501m.
12
34
56
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 249

250 Entomologist’s Rec. J. Var. 131 (2019)
Protorhoe corollaria(Herrich-Schäffer, 1848) Strelcë, 5.vi.2016, 1(Plate 5: 5), Gen. prep.
2./09.III.2018, S. Beshkov, genitalia (Plate 10: 6). Genitalia length: 3.65mm, cornutus
length: 0.55mm, wingspan 23mm. Protorrhoe corollaria is reported for Albania from
Rebel & Zerny (1931). After examination of genitalia here we confirm correct
identification and the presence of this species as well in Albania after the recent taxonomic
revision of the genus Protorhoe Herbulot, 1951 by Rajaei, Stadie & Hausmann (2017).
Eupithecia carpophagata Staudinger, 1871 Mali me Gropë, 1501m, 16.vii.2018, 1
(Plate 5: 6). New species for Albania.
Eupithecia gueneata Millière, 1862 Mal Çajup, 07.vi.2018, 2 (Plate 6: 1); Mali me
Gropë, 1501m, 16.vii.2018, 3 and 2. Known from Ploshtan (Rebel & Zerny,
1931) as new for the Balkan Peninsula. Second report for Albania.
Plate 6. 1. Eupithecia gueneata, . Mal Çajup; 2. Eupithecia succenturiata . Kozjak; 3. Eilema
palliatella,. Shkallë; 4. Eilema rungsi . Butrint; 5. Cucullia formosa . Kozjak; 6. Cucullia
artemisiae, . Plani i Bardhë.
12
3 4
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 250

251
Entomologist’s Rec. J. Var. 131 (2019)
Eupithecia succenturiata (Linnaeus, 1758) North Macedonia, Kozjak, Venec,
N41.8922, E021.2304, 1046m, 07.vii.2018, 1(Plate 6: 2). New species for R. N.
Macedonia.
Sphingidae Latreille, 1802
Proserpinus proserpina (Pallas, 1772) Dishnicë, 25.vi.2017, 4 specimens, collected 1.
In Albania known from a single locality only: Oroshi (Rebel, 1913, Rebel & Zerny,
1931, Eichler & Friese, 1965). Second report for Albania.
Hyles vespertilio (Esper, [1780]) Drenovë, 7.vii. 2016, 1; Ibid, 24.vi.2017, 1; Mali me
Gropë, 1405m, 17.vii.2018. Known in Albania from Shkodra and Oroshi (Rebel, 1913,
Rebel & Zerny, 1931, Eichler & Friese, 1965) and Quarrishtë (Librazhd) (Misja, 1978).
Plate 7. 1. Cucullia dracunculi, . Pletvar; 2. Helivictoria victorina, . Kolonjë; 3. Caradrina
suscianja, . Dishnicë; 4. Caradrina wullschlegeli,. Dardhë; 5. Olivenebula subsericata, .
Shkallë; 6. Archanara dissoluta,. Prespa Lake.
1 2
3 4
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 251

252 Entomologist’s Rec. J. Var. 131 (2019)
Erebidae Leach, [1815]
Eilema costalis (Zeller, 1847) (= morosina (Herrich-Schäffer, [1847]) Ossumi,
9.viii.2018, 3. Known from Kula e Lumёs (Rebel & Zerny, 1931). Second report
for Albania.
Eilema palliatella (Scopoli, 1763) (= unita ([Denis & Schiffermüller], 1775; = palleola
Hübner, [1808]) Known from Kolgecaj [North Albanian Alps] (Rebel & Zerny, 1931).
New data: Shkallё, 29.vi.2017, 1(Plate 6: 3), Gen. prep. 1./01.V.2018, S. Beshkov,
genitalia (Plate 11: 1). Second locality for Albania.
Eilema rungsi (Toulgoët, 1960) Butrint, 28.vi.2017, 2 (Plate 6: 4), Gen. prep.
1./04.IV.2018 S. Beshkov, genitalia with semi-everted vesica on glass in euparal
(Plate 11: 2); Ibid, 22.x.2017, 1. New species for Albania.
Plate 8. 1. Globia sparganii, . Prespa lake; 2. Apamea rubrirena, . Above Theth; 3.
Apterogenum ypsillon, . Shkallë; 4. Hadena luteocincta,. Drenovë; 5. Hadena luteocincta, .
Drenovë; 6. Hadena drenowskii, . Tomori, 1739m.
1 2
34
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 252

253
Entomologist’s Rec. J. Var. 131 (2019)
Honeyania ragusana (Freyer, 1844) Borshi (Heinicke, 1965). New data: Ilias, 140m,
15.v.2017, 1; Bistricë - Syri i Kaltër, 13.v.2017, 1; Butrint, 14.v.2017, 1. Second
report for Albania.
Noctuidae Latreille, 1809
Phyllophila obliterata (Rambur, 1833) Stan Karbunarë, 11.viii.2016, 1, Gen prep
1./16.IV.2019, S. Beshkov (Plate 11: 3). Known from: Tirana Botanical Garden; 6 km
E of Tirana towards Elbasan (Beshkov & Misja, 1995), Vukopoles near Poliçan
(Nahirnić & Beshkov, 2017). The report of Beshkov & Nahirnić (2019) for Ardenica
concerns another species: Condica viscosa (Freyer, [1831]).
Plate 9. 1. Hadena tephroleuca, . Above Theth; 2. Mythimna anderreggii pseudocomma .
Above Theth; 3. Mythimna alopecuri, . Pletvar; 4. Dichagyris candelisequa, . Mali me Gropë,
1501m; 5. Chersotis anatolica, . Tomori, 2379m; 6. Standfussiana lucernea illyrica, . Dishnicë.
1 2
3 4
56
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 253

254 Entomologist’s Rec. J. Var. 131 (2019)
Cucullia formosa Rogenhofer, 1860 North Macedonia, Kozjak Venec, 1070m,
N41.8908°, E21.2239°, 18.viii.2018, 4 specimens (Plate 6: 5). Second report for R.
North Macedonaia, previously known only from Ohrid (Thurner, 1964).
Cucullia artemisiae (Hufnagel, 1766) Plani i Bardhë, 20.vii.2018, 2 and 2 (Plate
7: 4). New species for Albania.
Cucullia dracunculi (Hübner, [1813]) North Macedonia, Prilep Region, near Pletvar
Pass, 963m, N41.3702, E21.6704, 5.viii.2018, 2 (Plate 6: 6). New species for
North Macedonia; Albania, Plani i Bardhë, 20.vii.2018, 2 and 6, genitalia
checked. Reported recently as a new for the Balkan Peninsula from Albania, Shkallë
(Beshkov & Nahirnić, 2019). Second report for Albania.
Plate 10. 1. Meganola kolbi, . Ilias; 2. Nola subchlamidula, . Palasë; 3. Dyscia raunaria, 
genitalia. Shkallë; 4. Protorhoe unicata, genitalia. Ilias; 5. Protorhoe unicata, genitalia.
Butrint; 6. Protorhoe corollaria, genitalia. Strelcë.
12
3 4
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 254

255
Entomologist’s Rec. J. Var. 131 (2019)
Helivictoria victorina (Sodoffsky, 1849) Kolonjë, 27.VI.2017, 1(Plate 7: 2). New
genus and a new species for Albania.
Condica viscosa (Freyer, [1831]) Stan Karbunarë, 11.viii.2016, 11 and 6, Gen
prep 1./19.IV.2019 S. Beshkov (Plate 11: 4: ); Ardenica, 4 and 1. In Beshkov &
Nahirnić (2019), the specimens mentioned and illustrated as Phyllophila obliterata
are Condica viscosa. New genus and a new species for Albania.
Cryphia algae (Fabricius, 1775) Known in Albania at a single locality only: Tirana
(Rebel & Zerny, 1931, Heinicke, 1965). The locality Galičica reported by Rebel &
Zerny (1931), following the report of Drenowsky (1930) is in North Macedonia, not
Plate 11. 1. Eilema paliatella, genitalia with everted vesica. Shkallë; 2. Eilema rungsi, 
genitalia with semi-everted vesica. Butrint; 3. Phyllophila obliterata, genitalia with semi-everted
vesica. Stan Karbunarë; 4. Condica viscosa, genitalia with everted vesica. Stan Karbunarë; 5.
Caradrina suscianja, genitalia with everted vesica. Dardhë; 6. Caradrina willschlaegeli, 
genitalia with everted vesica. Dardhë.
1 2
3 4
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:30 Page 255

256 Entomologist’s Rec. J. Var. 131 (2019)
in Albania. New data: Shkallë, 29.vi.2017, 3, genitalia checked and 23.viii.2017.
Second report for Albania.
Caradrina suscianja (von Mentzer, 1981) Zvezdë, 06.vii.2016, 1, genitalia with
evertred vesica checked and 1; Drenovë, 07.vii.2016, 1; Ibid, 24.vi.2017, 6;
Boboshticë, 08.vii.2016, 1; Dishnicë, 25.vi.2017, 1(Plate 7: 3), Gen. prep.
1./14.III.2018, S. Beshkov, genitalia; Dardhë, 26.vi.2017, 4, Gen. preps 1,
2./13.III.2018, S. Beshkov, genitalia with everted vesica on glass in euparal (Plate
11: 5) and 4. New species for Albania.
Caradrina wullschlegeli wullschlegeli Püngeler, 1903 (= wullschlegeli schwingenschussi
Boursin, 1936) Previously in Albania known only from Mali me Gropё, Bizё near
Plate 12. 1. Hadena luteocincta, genitalia. Drenovë; 2. Hadena luteocincta, genitalia with
everted vesica. Drenovë; 3. Mythimna alopecuri, clasper mechanism. Pustec; 4. Mythimna sicula,
clasper mechanism. Pustec; 5. Standfussiana lucernea illyrica, genitalia with everted vesica.
Shkallë; 6. Meganola kolbi, genitalia. Ilias.
1 2
3 4
5 6
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:31 Page 256

257
Entomologist’s Rec. J. Var. 131 (2019)
Shёngjergji (Heinicke, 1965). New data: Boboshticë, 08.vii.2016, 1; Shkallë,
29.vi.2017, 1; Dishnicë, 25.vi.2017, 4, Gen. prep. 2./14.III.2018, S. Beshkov,
genitalia with everted vesica on glass in euparal and 1; Dardhë, 26.vi.2017, 4
(Plate 7: 4), Gen. prep. 3./14.III.2018, S. Beshkov, genitalia (Plate 11: 6). Second
report for Albania.
Olivenebula subsericata (Herrich-Schäffer, 1861) Shkallë, 23.viii.2017, 3 (Plate 7:
5). New genus and a new species for Albania.
Archanara dissoluta (Treitschke, 1825) Prespa Lake, 10.vii.2018, 1(Plate 7: 6). New
genus and a new species for Albania
Globia sparganii (Esper, [1790]) Prespa Lake, 10.vii.2018, 1(Plate 8: 1). New genus
and a new species for Albania
Apamea rubrirena (Treitschke, 1825) above Theth, 19.vii.2019, 1(Plate 8: 2). New
species for Albania.
Apterogenum ypsillon ([Denis & Schiffermüller], 1775) Shkallë, 08.vi.2018, 1(Plate
8: 3). New genus and a new species for Albania.
Hadena luteocincta (Rambur, 1834) Drenovë, 7.vii.2016, 1(Plate 8: 4), Gen. prep.
1./29.I.2018, S. Beshkov,genitalia (Plate 12: 1), apophyses posterior with ovipositor
lobes = 6.5mm; Ibid, 24.vi.2017, 1(Plate 8: 5), Gen. prep. 1./31.I.2018, S. Beshkov,
genitalia with everted vesica (Plate 12: 2); Qafa e Selitës, 15.vii.2018, 1. New
species for Albania.
Plate 13. 1. Meganola strigula, genitalia. Pustec; 2. Nola chlamitulalis, genitalia. Bistricë -
Syri i Kaltër; 3. Nola subchlamidula, genitalia. Palasë.
1 2
3
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:31 Page 257

258 Entomologist’s Rec. J. Var. 131 (2019)
Hadena drenowskii drenowskii (Rebel, 1930) Tomori, 1739m, 13.vii.2018, 1and 1
(Plate 8: 6); Mali me Gropë, 1501m, 16.vii.2018, 3 and 9; Mali me Gropë,
1405m, 17.vii.2018, 2. In some internet sources for the range of H. drenowskii
drenowskii Albania is mentioned but without any further data and proofs. New species
for Albania.
Hadena tephroleuca tephroleuca (Boisduval, 1833) Above Theth, 19.vii.2018, 1(Plate
9: 1). In Albania known only from Gram Dibar as Hadena tephroleuca asiatica (F.
Wagner, 1931) (Beshkov & Misja, 1995). According to Beshkov & Junnilainen (2016)
the population of Albania belongs to nominotypical H. t. tephroleuca (Bsdv.). Second
report for Albania.
Mythimna anderreggii pseudocomma Rebel & Zerny, 1931 Above Theth, 19.vii.2018,
2 (Plate 9: 2); Tomori, 2379m, 14.vii.2018, 3. Till now this taxon in Albania
was known only from its type locality: Pashtriq [Beshtriku] (T.L. of ssp.
pseudocomma) (Rebel & Zerny, 1931, Heinicke, 1965). Second report for Albania.
Mythimna alopecuri (Boisduval, 1840) North Macedonia, Galičica Mts, between Dvata
Javora and Bulgarska Čuka Top, 1587m, N40.9908°; E20.8578°, 17.viii.2013, 4
and 1; Pelister Mt.-Prespa lake, between Slivnica Village and Sveta Bogorodica
Monastery, 1135m, N40.9684°; E21.0917°, 7.viii.2016, 2; Prilep Region, near
Pletvar Pass, 963m, N41.3702°, E21.6704°, 5.viii.2018, 2 (Plate 9: 3); Albania,
Pustec, 862m, 19.viii.2017, 2. In Albania known only from Klocë (Beshkov &
Nahirnić, 2018a). New species for North Macedonia and second report for Albania.
Although Mythimna alopecuri can be split by appearance from Mythimna sicula
(Treitschke, 1835) (= scirpi Duponchel, 1836), the easier way for correct identification
is examination of clasper mechanism. Differences are shown on Plate 12: 3, 4.
Albocosta musiva (Hübner, [1803]) North Macedonia, Bitola Region, Pelister Mt.,
“Široka”, 1955m, N41.0047°; E21.1686°, 06.viii.2016, 1. Second locality for North
Macedonia, recently reported as a new for the country from Galičica Mts (Beshkov,
2014).
Dichagyris candelisequa ([Denis & Schiffermüller], 1775) Mal Çajup, 7.viii.2018, 1;
Mali me Gropë, 1501m, 16.vii.2018, 1(Plate 9: 4). New species for Albania.
Euxoa conspicua (Hübner, [1827]) (= agricola Boisduval, 1829) Tomori, 2379m
14.vii.2018, 1and 2. In Albania known only from a single locality: Quarrishta
(Beshkov, Misja & Abadjiev, 1996). Second locality for Albania.
Rhyacia nyctymerides stavroitiacus Toulechkoff, 1951 Tomori, 2379m 14.vii.2018,
abundant, collected more than 40 specimens. In Albania known only from a single 
from Boboshticë (Beshkov, 2018). Second report for Albania.
Chersotis anatolica (Draudt, 1936) Tomori, 2379m 14.vii.2018, 2 (Plate 9: 5). New
species for Albania.
Standfussiana lucernea illyrica (Rebel & Zerny, 1931) (= lucernea bureschiTuleschkow,
1932) Previously known from Beshtriku; Djalica e Lumёs [Mali i Gjalicёs], Korab
(Rebel & Zerny, 1931, Heinicke, 1965). New data: Dishnicë, 25.vi.2017, 4 (Plate
9: 6), genitalia checked and 1; Shkallë, 29.vi.2017, 1, Gen. prep. 2./02.II.2018,
S. Beshkov, genitalia with everted vesica on glass in euparal (Plate 12: 5); Tomori,
1739, 13.vii.2018, 3; Tomori, 2379m 14.vii.2018, abundant; Mali me Gropë,
1501m, 16.vii.2018, 2; Mali me Gropë, 1405m, 17.vii.2018.
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:31 Page 258

259
Entomologist’s Rec. J. Var. 131 (2019)
Nolidae Bruand, 1847
Meganola kolbi Daniel, 1935 Ilias, 140m, 15.v.2017, 1(Plate 10: 1), Gen, prep.
4./18.IV.2018, S. Beshkov, genitalia (Plate 12: 6); Strelcë, 5.vi.2016, 1, genitalia
checked. New species for Albania. Female genitalia are distinctive and easily separate
this species from M. strigula.
Meganola strigula ([Denis & Schiffermüller], 1775) In Albania known from Kula e
Lumёs (Rebel & Zerny, 1931), Levan near Fieri [Koshovica Gorge] (Beshkov, 1995).
This reports may concern both M. strigula and M. kolbi. New data: Pustec, 862m,
19.viii.2017, 1, Gen. prep. 5./18.IV.2018, S. Beshkov, genitalia (Plate 13: 1).
Nola chlamitulalis (Hübner, [1813]) In Albania known from six localities (Rebel &
Zerny, 1931, Beshkov & Misja, 1995, Beshkov, 1995). New data: Klocë, 4.vii.2016;
Stan-Karbunarë, 11.viii.2016; Vukopoles, 25.ix.2016, 2, Gen. preps 2,
3./18.IV.2018, S. Beshkov, genitalia; Bistricë - Syri i Kaltër, 13.v.2017, 1, Gen.
prep. 1./18.IV.2018, S. Beshkov, genitalia (Plate 13: 2).
Nola subchlamydula Staudinger, 1871 Palasë, 6.vi.2016, 1(Plate 10: 2), Gen. prep.
2./17.IV.2018, S. Beshkov, genitalia (Plate 13: 3). New species for Albania.
Corrections to earlier published list
Phyllophila obliterata (Rambur, 1833) is reported in error from Ardenica
(Beshkov & Nahirnić, 2019). The specimens (Plate 11: 3, 4) are Condica
viscosa (Freyer, [1831]). The specimen illustrated in the same article (Plate 13:
6) as Lithophane socia (Hufnagel, 1766) from Starë is Lithophane semibrunnea
(Haworth, 1809) which is also new for Albania (genitalia checked).
Acknowledgements
The authors are thankful to Dr Boyan Zlatkov (Sofia, Bulgaria) for the
microscopic photographs of the Nola, Meganola and Eilema rungsi genitalia as
well as to Colin W. Plant (Bishops Stortford, UK) for linguistic corrections. As
members of the Macedonian Entomological Society ENTOMAK, the authors
express their sincere gratitude to Nikola Micevski (Skopije) for the membership
cards and for including their names in the collective collecting permit.
References
Beshkov, S. 1995. A contribution to the knowledge of the Lepidoptera fauna of Albania 2. Some findings
of a collecting trip in September 1993 (Lepidoptera, Macrolepidoptera). Atalanta 26(1/2): 365-399.
Beshkov, S., 2014. Contribution on the Macrolepidoptera of the Republic of Macedonia with a report of
one new genus and four new Noctuidae species for the country. Entomologist’s Rec. J. Var. 126(5):
175-185.
Beshkov, S., 2018. A Contribution to knowledge of the Balkan Lepidoptera: New and rare Erebidae and
Noctuidae for Albania, collected in 2016. Entomologist’s Rec. J. Var. 130(3): 155-172.
Beshkov, S. & Junnilainen, J., 2016. The first report of Hadena tephroleuca (Boisduval, 1833)
(Lepidoptera: Noctuidae) in Bulgaria. ZooNotes 93: 1-3.
Beshkov, S. & Micevski, N., 2019. Eupithecia gratiosata Herrich-Schäffer, 1861 and Metachrostis dardouini
(Boisduval, 1840) new for the Republic of Macedonia. Historia naturalis bulgarica 37: 1-4.
Beshkov, S. & Misja, K., 1995. A contribution to the knowledge of the Lepidoptera fauna of Albania. 1. Some
materials from the collection of K. Misja in the Natural History Museum Tirana and some results of the
collecting trip of Beshkov during 1992 (Lepidoptera, Macrolepidoptera). Atalanta 26(1/2): 345-363.
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:31 Page 259

260 Entomologist’s Rec. J. Var. 131 (2019)
Beshkov, S., Misja, K. & Abadjiev, S., 1996. A Contribution to the knowledge of the Lepidoptera fauna
of Albania. 3. Some results of the collecting in April 1994 and some materials from the collection of
the Natural History Museum of Tirana (Lepidoptera, Macrolepidoptera, Heterocera). Atalanta
27(3/4): 623-648.
Beshkov, S. & Nahirnić, A., 2018a. New and rare Noctuoidea species for Albania (Lepidoptera: Nolidae,
Erebidae and Noctuidae). Atalanta 49(1-4): 159-170.
Beshkov, S. & Nahirnić, A., 2018b. A contribution to knowledge of the Balkan Lepidoptera: new and
rare moths for Albania, collected in 2017. Entomologist’s Rec. J. Var., 130: 215-232.
Beshkov, S. & Nahirnić, A., 2019. A contribution to knowledge of the Balkan Lepidoptera: species
collected in the autumn of 2018 in Albania (macrolepidoptera with some crambidae). Entomologist’s
Rec. J. Var., 132: 66-102.
Drenowsky, Al., 1930. Beitrag zur Lepidopterenfauna S.W. Macedoniens. Zeitschrift Bulg. Akad.
Wissensch. 42: 120-177.
Eichler, F. & Friese. G., (1965): Ergebnisse der Albanien-Expedition 1961 des Deutschen
Entomologischen Institutes. 32. Beitrag. Lepidoptera: Sphingidae. Beitrage zur Entomologie 15(5-6):
633-640.
Heinicke, W., 1965. Ergebniss der Albanien-Expedition1961 des Deutschen Entomologischen Institutes.
31. Beitrag. Lepidoptera: Noctuidae. Beiträge zur Entomologie 15 (5/6): 501–632.
Misja, K., 1978. Donnees sur l’etude des papillons du groupe Heteroneura de l’Albanie. - Buletini i
Shkencave tё Natyrёs 3: 65-73. Universiteti i Tiranёs, Fakulteti i Shkencave tё Natyrёs. (in
Albanian). Tirana.
Misja, K., 1977. Donnees preliminaires sur l’etude des papillons nocturnes de la Famille Geometridae.
Buletini i Shkencave të Natyrës 2: 81-91. Universiteti i Tiranës, Fakulteti i Shkencave të Natyrës. (in
Albanian).
Nahirnić, A. &Beshkov, S., 2017. Penestoglossa dardoinella (Millière, 1863) (Lepidoptera: Psychidae)
recorded for the first time in Albania. ZooNotes 101: 1-3.
Rajaei, H., Stadie, D. & A. Hausmann, 2017. Taxonomic revision of the genus Protorhoe Herbulot, 1951
(Lepidoptera, Geometridae, Larentiinae), new taxonomic changes and description of two new species.
Zootaxa 4282 (2): 269-291.
Rebel, H. & Zerny, H., 1931. Die Lepidopterenfauna Albaniens. Denkschriften der Akademie der
Wissenschaften, Math. Nat. Kl. Wien 103: 37-161.
Rebel, H., 1913. Studien über die Lepidopterenfauna der Balkanländer, Teil III. Sammelerrgebnisse aus
Montenegro, Albanien, Mazedonien und Trazien. Ann., k.k. naturhist. Hofmus., Wien 27: 281-334.
Thurner, J., 1964. Die Lepidopterenfauna Jugoslavisch Mazedoniens. 1. Rhopalocera, Grypocera und
Noctuidae. Prirodonaucen muzej Skopje, Posebno izdanie Nr. 1., 159 pp.
Urbahn, E., 1966. Ergebnisse der Albanien-Expedition 1961 des Deutschen Entomologischen Institutes.
51. Beitrag. Lepidoptera: Geometridae. Beitrage zur Entomologie 16(3-4): 407-446.
Ent Rec 131(5).qxp_Layout 1 26/09/2019 15:31 Page 260