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EUROPEAN RED LIST OF
TERRESTRIAL MOLLUSCS:
SNAILS, SLUGS, AND SEMI-SLUGS
B
The semislug Vitrinobrachium breve (LC) shown
here at Elfenau, close to Bern in Switzerland.
A species of forests and woodlands, it is
widespread in Central and Southern Europe.
© Beat Pfarrer
LEAST CONCERN
LC
1
European Terrestrial Molluscs
Europe has one of the most highly fragmented
landscapes of all the continents, with only a tiny
fraction of its land surface – largely the Arctic
regions – considered as wilderness. Over the last
5,000 years, much of the European landscape
has been shaped by human activity through
food, timber and fuel production. Currently within
Western Europe, more than 80% of the land
area is under some form of direct management
(European Environment Agency, 2007).
Consequently, European species are to a large
extent dependent upon semi-natural habitats
created and maintained by human activity,
particularly traditional, non-intensive forms of
land management. Over the last century, these
habitats have come under increasing pressure
from agricultural intensication, urban sprawl,
infrastructure development, land abandonment,
acidication, eutrophication and desertication.
Many species are directly affected by
overexploitation, persecution, and impacts of alien
invasive species, and climate change is set to
become an increasingly serious threat. Europe is
a vast, diverse region and the relative importance
of different threats varies widely across its
biogeographic regions and countries. This report
is one of a series of reports on the conservation
status of animals and plants in Europe, focussing
on terrestrial molluscs in the Class Gastropoda –
slugs, snails, and semi-slugs.
The Phylum Mollusca (snails, slugs, clams,
mussels, chitons, tusk shells, monoplacophorans,
polyplacophorans (chitons), Caudofoveata,
Solenogastres, squids, cuttlesh and octopuses)
contains an estimated 94,000 extant species
worldwide. This gure comprises ca. 55,000
marine gastropods, ca. 10,000 marine bivalves, at
least 25,000 terrestrial and freshwater snails and
1,300 freshwater bivalve species. The remaining
groups embrace considerably fewer numbers of
species, ca. 800 cephalopods, 1,100 chitons,
600 tusk shells and 420 Caudofoveata and
Solenogastres (currently known and estimated
numbers taken from MolluscaBase). The overall
number of described species increases each
year as more research is undertaken, especially
in under-researched regions of the world such
as the tropics. However, even within the relatively
well-studied European region, new species are
constantly being described, and the status of
many species names, as either valid species
or as synonyms of other species, needs further
taxonomic study. The actual number of extant
species of Mollusca globally is uncertain. Molluscs
can be found in almost all habitats, from the
bottom of the oceans to mountain-tops and
across tundra regions. They are very diverse, not
only in size and shape but also in their lifecycle,
lifespan and habitat. They are an important
food source for birds, sh, mammals and other
invertebrates, and in some cases for humans,
too. Together with such groups as the fungi and
saproxylic beetles, Mollusca play a key role in
soil-generation and the recycling of nutrients
in natural ecosystems. Because of their often
specic habitat requirements, they can be good
indicators of environmental quality, especially for
ancient woodlands, grasslands, wetlands and
subterranean habitats. Farmers and gardeners
usually view terrestrial molluscs as pests that
damage plants and the subsequent control of
terrestrial molluscs can cause problems from a
conservation perspective. However, these ‘pest’
species actually represent only a very small
fraction of the species that occur in the European
region. Indeed, many of the globally signicant
pest species were exported from Europe to other
parts of the world.
Terrestrial molluscs utilise a very wide range of
microhabitats – taxa have evolved to exploit these
niches, with some having very specic ecological
requirements, such as interstitial spaces within
subsoil, living on trees, found only on rock-faces
or restricted to pristine forest habitats. The
assemblage of molluscs found at a particular site
is inuenced by historical geographical isolation
and glaciation, altitude, degree and duration of sun
exposure, the quantity and duration of rainfall and
the frequency of habitat disturbance (i.e., forest
res or clear-cutting, amongst many other factors).
2
Assessment Scope
This IUCN European Red List provides an
assessment for the 2,480 species of terrestrial
mollusc species known to be present in the
European region at the start of 2019. Of these
species, 11 are considered Not Applicable
(NA) for assessment as they are not native to
the European region. The number of species
constantly changes as their taxonomy is
revised and new species are described; a
further 15-20 species have been described
for the European region since this Red List
assessment was completed. Updates can be
found on the MolluscaBase database, which
represents the taxonomic backbone used in
this project.
The data presented in this report comprises
1,233 species assessed between 2009 and
2011 (Cuttelod et al., 2011) and a further 1,261
species that were assessed between 2015
and 2018 (of which 14 were rst assessed in
the rst phase but reassessed as the result of
signicant new data or changes in taxonomic
concept). All the assessments were made
following the IUCN Red List Categories and
Criteria (IUCN, 2001; IUCN, 2012a), which is
the global standard for measuring extinction
risk. For species not endemic to the EU
Member States, the Guidelines for Application
of IUCN Red List Criteria at Regional and
National Levels (IUCN, 2003; IUCN, 2012b)
were applied.
Red List assessments were made at two
regional levels: geographical Europe (‘Pan
Europe’), and the Member States of the
European Union. In the rst stage of the
European Red List of Non-marine molluscs
(Cuttelod et al., 2011), the species were
assessed at the level of the then 27 Member
States of the EU (Croatia joined the EU in
2013), whilst in the second stage assessments
(2015-2018), the species were assessed at
the EU 28 level, with the addition of Croatia.
For Pan Europe, the scope is continent-wide,
extending from Iceland in the west to the Urals
in the east, and from Franz Josef Land in the
north to the European Macaronesian Islands
in the south. The Caucasus region is not
included.
The slug Milax verrucosus (LC) is thought to
be endemic to the Stara Planina Mountains
in Bulgaria. It occurs in mountains, in shaded
valleys, and fairly damp places, in deciduous
and mixed forests rich in litter. There are no
widespread threats and it is Least Concern.
© Ivailo Dedov
LEAST CONCERN
LC
3
Levels of Endemism in Europe
The level of endemism (92%; 2,268 species)
within the terrestrial molluscs of the European
region is much greater than in many other
taxon groups. In vertebrates, for example,
the reptiles are 52.1% endemic (73 endemic
species), mammals 26.9% (61 endemic
species), and amphibians are 76.3% endemic
(61 endemic species). There are an estimated
100,000 invertebrate species in Europe, with
six groups assessed for the European Red
List. The dragonies and damselies have a
low level of endemism (12.2%; 16 endemic
species), compared to Orthoptera (69.9%;
742 endemic species), and saproxylic beetles
(32.4%; 224 endemic species). There are
very few terrestrial mollusc species present in
Europe that are not endemic to the European
region, and most of these have ranges that do
not extend far into North Africa or Asia. Only
a few species are proven to have a holarctic
distribution pattern. As a consequence,
Europe and in particular those countries
with single-country threatened species have
a responsibility to ensure the long-term
conservation of these species.
Threat Status
Overall, 21.8% and 22.5% of the 2,469 native
species of terrestrial mollusc are considered
threatened (species assessed as Critically
Endangered, Endangered or Vulnerable) in
Europe and in the European Union Member
States (EU 27/28), respectively. These are the
mid-point values – assuming that a similar relative
proportion of the Data Decient (DD) species are
likely to be threatened – and provide the best
estimate of the proportion of threatened species
(IUCN, 2016). For Europe, the percentage of
threatened species ranges between 19.6%
(if no DD species are considered likely to be
threatened) and 29.8% (if all DD species were
to be assessed as threatened). The equivalent
gures range from 20.1% to 30.8% for the
EU27/28 Member States.
For just over one-tenth (250 species – 10.1%)
of the species in Europe there was insufcient
information to evaluate their risk of extinction and
they were assessed as Data Decient (DD). In the
EU Member States, 240 species (10.6%) were
assessed as DD, which is considerably lower
than the level of data deciency for the freshwater
molluscs (24.7%) (Cuttelod et al., 2011).
In Europe, 97 species (3.9%) have been
assessed as Critically Endangered, and of these,
23 species are considered Possibly Extinct
(CR(PE)). A further 90 species (3.6%) have been
assessed as Endangered and 296 (12.0%) as
Vulnerable (Table 1 and Figure 1a). A further 14%
(345 species) are considered Near Threatened.
There is a very slightly higher proportion of
threatened species in the EU 27/28 (4.1%
Critically Endangered (23 species are CR(PE)),
3.7% Endangered and 12.3% Vulnerable), with
12.3% Near Threatened (Table 1 and Figure 1b).
Including the Near Threatened species, 37.4%
of mollusc species present in the region of
Europe – for which sufcient data exist – can
be considered to be of elevated conservation
concern. Nearly half (47.0%, 655) of all the extant
species in the Member States of the European
Union, are of conservation concern (threatened
and Near Threatened categories).
At present, ve species are considered
to be extinct in the European region, all
previously endemic to the Member States of
the European Union, i.e. Leiostyla lamellosa,
Pseudocampylaea lowii, Vitrea storchi, Zonites
4
IUCN Red List Categories
No. of species
in Europe
(No. of endemic species)
No. species EU
Member States
(No. of endemic species)
Extinct (EX) 5 (5) 5 (5)
Extinct in the Wild (EW) 0 (0) 0 (0)
Regionally Extinct (RE) 0 (0) 0 (0)
Critically Endangered (CR) 97 (95) 93 (91)
Endangered (EN) 90 (85) 84 (77)
Vulnerable (VU) 296 (286) 278 (261)
Near Threatened (NT) 345 (333) 279 (249)
Least Concern (LC) 1,386 (1,227) 1,285 (768)
Data Decient (DD) 250 (237) 240 (204)
Total number of species 2,469 2,264
The gure (2,469) for the total of species shown in Europe excludes eleven species that are considered introduced to Europe
and are therefore Not Applicable (NA). In addition, the total number of species (2,264) shown for the EU Member States
excludes a further 206 species that are considered NA because they are not recorded for the Member States. It should be
noted that some taxa that occur in Croatia are excluded because they were assessed prior to the accession of Croatia to the
EU in 2013.
Table 1. Summary of numbers of terrestrial molluscs within each Red List Category.
The snail Rossmaessleria scherzeri
(LC) is endemic (as the subspecies
R. scherzeri scherzeri) to the Rock
of Gibraltar. Although its distribution
in Europe is highly restricted, it is
assessed as Least Concern as it occurs
within well-managed Protected Areas.
© Marco Neiber
LEAST CONCERN
LC
5
santoriniensis and Zonites siphnicus. All
ve species were small-range endemics
from islands; the rst two from Madeira and
Porto Santo, the remaining three from the
Aegean, the islands of Chios (Vitrea storchi),
Santorini (Z. santoriniensis) and Sifnos,
Sikinos and Folegandros (Z. siphnicus). The
large species from the Zonites genus live on
many Aegean islands, and often are single-
island endemics, or inhabit a small group of
neighbouring islands, as with Z. siphnicus.
Species from the Zonites genus are known to
have undergone seriously declines in recent
decades, and many of them have not been
found alive in the eld. It is not clear whether
their rapid extinction reects a natural process
of declining island faunas, whether it is
combined with climate change, or has other
causes.
The complete list of species and their Red List
status in Europe and in the EU Member States
(EU 27/28) is available as Supplementary
Material1.
Comparing mid-point values for other
European Red List species groups that have
been comprehensively assessed (Table 2) for
the European region, we nd that terrestrial
molluscs are one of the more threatened
invertebrate groups assessed for Europe
so far. Only freshwater molluscs, freshwater
shes, grasshoppers and crickets, and
amphibians have a higher percentage of
threatened species than terrestrial molluscs;
terrestrial molluscs show the highest level of
endemism (92%).
1 Supplementary Material available at:
https://portals.iucn.org/library/node/48439
Figure 1a. IUCN Red List status of
terrestrial molluscs in Europe. Figure 1b. IUCN Red List status of terrestrial
molluscs in the EU Member States.
CR
DD
EX
EN
VU
NT
LC
4.1%
10.6%
0.2%
3.7%
12.3%
12.3%
56.8%
CR
DD
EX
EN
VU
NT
LC
3.9%
10.1%
0.2%
3.6%
12.0%
14.0%
56.8%
6
Table 2. Summary of the total number of species assessed, and proportion endemic,
threatened, and of elevated conservation concern, ordered by the % Threatened.
Taxonomic group
Total species
assessed
(ex NA1)
%
Endemic
%
Threatened2
(mid-point)
%
Elevated
Conservation
Concern3
Freshwater molluscs 853 87.6 58.6 69.7
Freshwater shes 516 81.0 39.9 43.2
Grasshoppers & Crickets 1,077 68.9 28.5 43.9
Amphibians 80 76.3 24.1 40.5
Terrestrial molluscs 2,476 92.0 21.8 37.4
Lycopods & ferns 194 27.3 19.9 34.2
Reptiles 140 52.1 19.6 32.6
Mammals 227 26.9 17.6 26.6
Dragonies & damselies 131 12.2 15.1 26.2
Birds 543 6.8 12.1 17.7
Bees 1,942 20.7 9.2 21.2
Butteries 437 33.0 8.6 19.0
Notes
1 NA, Not Applicable species, usually species considered non-native to the European region. 2 Mid-point gure based on CR,
EN and VU assessments. 3 Based on Extinct (EX, EW and RE), CR, EN, VU and NT assessments.
Sources
Data extracted from the original European Red Lists: freshwater molluscs (Cuttelod et al., 2011); freshwater shes (Freyhof and
Brooks, 2011); Orthoptera (Hochkirch et al., 2016); mammals (Temple and Terry, 2007); amphibians (Temple and Cox, 2009);
reptiles (Cox and Temple, 2009); butteries and bees (van Swaay et al., 2010); dragonies (Kalkman et al., 2010); medicinal
plants (Allen et al., 2015); birds (Birdlife International, 2015); marine shes (Nieto et al., 2015); lycopods and ferns (García Criado
et al., 2017); trees (Rivers et al., 2019); updated (with recent revisions and additional taxa) by M. Bilz pers. comm. 2019).
Major Threats
A large number of species are threatened due
to their restricted range and consequential
high levels of extinction risk. The ranges of
many of these species do not lie in protected
areas and their current conservation status is
concerning.
An overview of the major threats affecting
European terrestrial molluscs is shown in
Figure 2.
Over 30% of all assessed species are
impacted by natural system modications,
including the removal of banks and hedges,
changes in re management practices,
quarrying, livestock grazing, wetland
conversion and peat extraction, changes in
land management practices, logging and
other forest management, especially of old-
growth forests.
Loss of habitats through the conversion
of land to residential and industrial use is
common on the edges of towns and villages.
Habitats at risk include forests, meadows,
fens, and marshes. Habitat loss can create
additional threats to terrestrial molluscs by
fragmenting forest habitats, reducing habitat
area and quality; larger species of terrestrial
molluscs are likely to be more at risk than
7
smaller species. Terrestrial molluscs tend
to occur in meta-populations in marshes
and fens, but land drainage can lead to
fragmentation of riverine fringing habitats and
therefore reduced opportunities for molluscs to
recolonise after ooding events.
Fire within Mediterranean zones is a natural
event, due to lightning strikes. However, in
recent decades, the frequency and severity
of res has increased. Fire is known to cause
declines in snail diversity and abundance, but
given time terrestrial molluscs can recolonise
when suitable vegetation re-emerges and
where the burn has not destroyed the soil
cover and crevices within rocky habitats. The
impact of long-term climate change on re risk
and intensity requires more research. Families
at particular risk include the Clausiliidae,
Helicidae, Geomitridae, Hygromiidae, Enidae
and Vitrinidae.
Agricultural land use has led to a decline in
snail abundance, for example, where land has
been drained or ploughed, rough grazing has
been converted to improved grasslands, or
where eld boundaries have been removed
to make large elds that are more economic
for cultivation. The conversion of heathlands,
forest and marshes/fens for agriculture use
over the last 30 years further contributes to
the loss of habitats for terrestrial molluscs.
Families at particular risk include the Helicidae,
Geomitridae, Hygromiidae, Vertiginidae and the
Succineidae.
Many slug species, which are commonly
regarded as agricultural pests, are threatened
Figure 2. Major threats to terrestrial molluscs in Europe.
Non-threatened species Threatened species
Number of species
Natural system modications
Residential & commercial development
Agriculture & aquaculture
Energy production and mining
Human intrusions and disturbance
Biological resource use
Transportation and service corridors
Climate change and sever weather
Pollution
Invasive and problematic native species
Geological events
Other threats
0100 200 300 400 500 600 700
8
by the indiscriminate use of molluscicides and
biological control measures. Some endemic
slug species do not have large ranges, and
these species are particularly vulnerable to the
use of slug controls; families at particular risk
include the Limacidae and the Arionidae.
Quarrying of limestone destroys large areas
of crags and subterranean habitats. As
many terrestrial molluscs are calciphiles,
range-restricted endemics are common in
limestone habitats, leading to rapid declines
in populations, and potentially local or global
extinctions. Families at particular risk include
the Aciculidae, Cochlostomatidae, Argnidae,
Clausiliidae, Lauriidae, Pupillidae, Chondrinidae
and Spelaeodiscidae.
The larger terrestrial molluscs, such as Helix
pomatia, Helix lucorum, Helix gulina, Helix
straminea, Helix ligata, Cornu aspersum, Otala
lactea, O. punctata, Massylaea vermiculata
and Cepaea nemoralis continue to be collected
for food within many of the Mediterranean
countries. Some endemic species, such as
the ‘Chapa’ (Iberus gualtieranus) fetch high
prices in local markets, placing increasing
pressure on these populations. There has been
little research so far on the impact of climate
change on snails and slugs, for example
through re, or through lengthening periods of
no or low rainfall. This may exceed the capacity
of the species to withstand drought periods,
and there are anecdotal observations of this
already occurring in parts of southern Europe,
for example in Greece.
Unmanaged grazing, such as shown here
by a mixed herd of goats and sheep in
the Slavyanka Mountains near the Izvora
hut in Bulgaria, is a signicant threat to
some terrestrial mollusc subpopulations.
© Fabia Knechtle
9
Population Trends
As part of the Red List process, the trend
of each species’ overall population was
assessed as either declining, stable, increasing
or unknown.
Data are presently very poor on the population
size and trend of many species of terrestrial
molluscs, largely because there are few
quantitative data collected, usually for highly
range-restricted island species. In some
countries, there are data on general declines
over 20-30 years, as distribution atlases
have been published at these intervals (e.g.
for the British Isles; Kerney, 1999). This
is an area where citizen science projects
could be launched to increase efforts in
monitoring European molluscs. However,
it has to be stressed that for most species
of invertebrates, it is extremely hard to
obtain robust data on population size, and
consequently on their long-term trends!
In Europe, 6% (138 species) of terrestrial
mollusc species are thought to be declining
in population size, whilst 42% (1,043 species)
are considered to have a stable population,
however, the trend is increasing for just 1%
of species and the trend is not known for
more than half (1,272) of all species (Figure 3).
As would be expected, the picture is worse
for threatened species, with no populations
thought to be increasing, 15% (70 species)
stable, but 20% (97 species) with declining
population trends and the trend not known for
65% (315 species).
Figure 3. Population trends of European terrestrial molluscs; all species: outer ring;
threatened species only: inner ring.
Stable
Stable
Increasing
Unknown
Unknown
Decreasing
Decreasing
42%
15%
1%
51%
65%
51%
20%
10
Protected Areas
Many species of terrestrial molluscs are
recorded from protected areas (see Figure 4),
however, this high gure is misleading, as there
are almost no protected areas designated or
managed for mollusc species. In general, only
those species listed on the Habitats Directive,
with a requirement to have designated Special
Areas of Conservation (SACs), such as the
four Vertigo species (Vertigo angustior (VU),
V. genesii (LC), V. geyeri (LC), V. moulinsiana
(VU)), Elona quimperiana (LC), and the Kerry
Slug Geomalacus maculosus (LC), are
species in protected areas with management
plans. However, there are many species of
terrestrial molluscs that are known from single
sites, especially on the islands, such as in
Macaronesia or the Aegean Sea, many of them
threatened, that do not occur within protected
areas. It is likely that the areas where these
species occur would qualify as Alliance for
Zero Extinction (AZE) sites, or should otherwise
come within the protected areas network, and
further work is required to conrm this.
Figure 4. The proportion of European terrestrial molluscs known to occur within a
protected area; all species: outer ring; threatened species only: inner ring.
No
No
Yes
Unknown
Unknown Yes
6%
10%
44%
50%
45% 45%
11
Spatial Distribution Patterns
The spatial distribution patterns of terrestrial
molluscs in Europe are shown in Figures 5 to
8. The intermediate latitudes of central Europe
clearly stand out as areas of high species
richness (Figure 5). Biodiversity hotspots are
located in mountainous areas such as the
Pyrenees, Alps, Carpathians and the Balkan
Peninsula but also on the Macaronesian
Islands. The richness of endemic species
is shown in Figure 6 and shows somewhat
similar patterns to the overall species diversity.
The Mediterranean and Macaronesian islands
have many range-restricted endemic terrestrial
snails, which can also be seen on the endemic
species richness map. The larger of the
Canary Islands, Tenerife and Gran Canaria, as
well as Madeira and Porto Santo, are densely
populated by endemic species, and typically,
each particular island has its own set of
endemic species.
The distribution of threatened species is
shown in Figure 7. Here, the pattern does not
strictly follow the simple formula “the higher
the species richness, the higher the number
of threatened taxa”. This rule may apply for
the Macaronesian islands, but certainly not
for the continent. On average, the areas of
high species richness “only” have a density of
2-3 threatened species per hexagon with rare
spot-like exceptions populated by species
of the “4-6” category. On the continent, the
highest threat level is reached on the southern
Pelopónnisos in Greece, reaching the
maximum level of 17-24 threatened species
per hexagon. This area only reaches moderate
to slightly elevated values in species richness,
endemicity and data deciency (Figure 8), so
the explanation for this exceptional issue is
likely due to elevated environmental stress in
the area, caused by re and other factors.
Figure 5. Overall species richness of European terrestrial molluscs.
12
Figure 6. Distribution of endemic European terrestrial molluscs.
Figure 7. Distribution of threatened terrestrial molluscs in Europe.
13
The distribution of Data Decient species is
shown in Figure 8. The Data Decient category
reects the limited knowledge about the
distribution, population or ecology of a species.
The distribution pattern of these species on the
continent is striking: the greatest numbers of
poorly known species occur in the western and
central arc of the Alps, and, to a lesser degree,
in southern Italy, Sicily, and the southern part
of the Balkan Peninsula. This discovery is
somewhat biased as it corresponds to the area
of the highest species richness and endemicity,
but it occurs in countries where a well-
developed scientic infrastructure is assumed
to exist. A lot of effort is needed to eliminate
the inationary creation of new taxa introduced
by the followers of the “Nouvelle École” in the
19th century. Another hotspot of DD species is
on the Canary Islands. Despite the enormous
efforts during 30 years of research by Spanish
malacologists and colleagues from abroad,
the knowledge on these islands and their rich
malacofauna is still poor.
Conservation Actions
Legislation for species
Despite the high level of threat and endemism,
just 14 of the threatened terrestrial molluscs
from Europe are listed in the Habitats Directive
annexes as requiring strict protection. Most
taxa were placed in the Directive annexes
based on data from the 1970s and have not
been reviewed in recent times, except when a
new country becomes a member state.
Protected Areas
It has been argued that many species will have
ranges that overlap the Natura 2000 network
of protected areas, which covers more than
Figure 8. Distribution of Data Decient terrestrial molluscs in Europe.
14
Fire, such as shown in this
landscape near Trun in western
Bulgaria, can completely
devastate habitats and pose
a signicant threat to range-
restricted species.
© Ivailo Dedov
18% of the EU’s land area and hence plays
a vital role in protecting species and their
habitats. It is clear, however, that in practice,
a Natura 2000 designation does not always
mean actual protection (at least in the eastern
part of Europe). Moreover, there are threatened
species in habitats that lie outside protected
areas, such as Natura 2000, Ramsar sites
and national protected areas. This needs to
be reviewed as an urgent action, to determine
whether existing protected areas could have
their boundaries extended or new sites
developed, especially for threatened single-site
(AZE) species. Habitat protection is one of the
most important conservation actions needed
for terrestrial molluscs at the national and
international levels.
Reducing damaging operations
From a nature conservation point of view,
the pivotal questions for terrestrial molluscs
are the protection of existing habitats and
the restoration and/or reconstruction of
endangered or lost habitats. This can be
seen in Figure 2 where “natural system
modications” are agged as the major threat
to terrestrial molluscs, followed by “residential &
commercial development”. Future conservation
actions have to focus on the protection of areas
rather than on individual species, as was the
practice with the species listed in Annex II of
the EU Habitats Directive. The management of
habitats for the benet of terrestrial molluscs,
where the major threats are disturbance or
destruction of the habitats, is urgently needed.
1) Logging
Logging and other forestry activities affect
many forest-dependent species in Europe. In
all European forest types, excessive logging
deeply modies or even destroys wood
habitats. This is particularly true for sensitive
species (i.e., species dependent on large
standing trees, for example, or on a diverse and
stable functioning forest ecosystem); forestry
policy has to follow sustainable management of
areas. Forest certication schemes, such as the
Forest Stewardship Council (FSC), promote the
15
responsible management of the world’s forests,
and many European and EU countries have
certied areas, and the process is supported by
the European Union Forestry Strategy. Certied
forest areas must ensure the protection of
threatened species in order to comply with the
certication requirements, and can, therefore,
play an important role in terrestrial mollusc
conservation.
2) Biodiversity-friendly forest
management
Forest management is also important, as it is
not just logging that can impact native species
of terrestrial molluscs and slugs. Dead wood
is often a favoured habitat, hence removal
of large trunks can be detrimental, and any
woodland habitat plan should ensure that the
habitats maintain a range of suitable trees of
different ages, that large trunks are allowed
to decay in situ and that clearings, and that
hedges and boundary areas retain some
understory vegetation and where possible link
across agricultural areas. Another completely
underestimated factor is the continuity in
forest habitats: decaying ground litter is
an important habitat, particularly for small
terrestrial mollusc species. For example,
species in the Vertiginidae family, and the
valloniid species Acanthinula aculeata and
Spermodea lamellata, are strongly dependent
on ground-litter continuity in forests, especially
so in Scandinavian woodlands. The role
of biological or landscape corridors for the
protection of terrestrial molluscs remains
controversial. Landsnail dispersal may be an
active process covering only short distances;
here, corridors may have a fragmenting effect.
However, many snails disperse passively
using migrating vector species (smaller or
larger mammals, fowl, human transport), so
corridors may have a positive dispersal effect
as well. Size, structure, and maintenance of
corridors are also possible factors inuencing
their suitability for conservation purposes.
Summarising, it can be said that biological
corridors are mainly designed to serve the
conservation of vertebrates, but their role for
land snail dispersal and conservation needs
further intensive study.
3) Biodiversity-friendly grassland
management
Grassland areas host many species of endemic
terrestrial molluscs. Better land management
practises, such as preventing over- and under-
grazing, reducing the density of stock to reduce
trampling, retaining boundaries such as hedges
and banks, not mowing or harvesting too
frequently, and not burning off the grass and
brush at inappropriate times of the year. For
species-rich grassland sites, the focus should be
on extensive management practices. Grassland
enrichment through the introduction of nutrients
(articial fertilisers and animal manures) results in
the reduction of plant species richness that then
affects mollusc diversity.
4) Reducing habitat destruction by
quarrying
Many terrestrial molluscs are restricted to rocky
crags and hillsides, which are often sites that
have the potential for limestone extraction.
Planning approval for quarrying usually requires
an environmental impact assessment (EIA) for
biodiversity impacts, however, these frequently
do not take account of invertebrate species,
or only look at those listed in the EU Habitats
Directive annexes. Recommended actions
include undertaking an EIA if an endemic
terrestrial mollusc is suspected to be present,
prior to quarrying of limestone resources to
ensure that these take into account biodiversity
loss, especially in regions of high terrestrial
mollusc endemism. It is possible to create areas
that are set aside as mini-reserves within the
range of a quarry region to protect the species
and serve as a refuge from where terrestrial
molluscs can recolonise once quarrying
operations cease. However, such a fragmented
landscape cannot protect subterranean or
cave-dwelling species, as their subterranean
habitat and especially its microclimate will be
severely altered by quarrying and mining; even
road construction in limestone areas is able to
damage this fragile environment.
5) Reducing habitat destruction by
drainage and water extraction
Many terrestrial molluscs are restricted to
small calcareous fens and bogs, which
16
Ilhéu de Cima: a small island southwest of Porto
Santo in the Madeira Archipelago, showing the
effect of vegetation recovery after eradication of
the feral goat population. A dense and species-
rich plant community is again found on the islet,
a vital prerequisite for terrestrial snail fauna.
© Dinarte Teixeira
are often sites identied as suitable for
agricultural improvement. Once drainage has
commenced, it is impossible for these species
to recolonise, unless it is along the margin
of rivers. Small areas of suitable habitat can
protect these species.
Wetland habitats are often fed by springs that
are identied as suitable for offtake of water
for villages and agricultural purposes. Once
offtake has commenced, it creates areas of dry
habitat and vegetation quickly changes, such
that the species sensitive to drying habitats
quickly are lost in these sites. Small areas
of suitable habitat can protect the species
provided that the water supply is maintained at
minimum levels all year around.
6) Erosion and habitat restoration
Erosion caused by a variety of factors is a
widespread phenomenon all over Europe.
However, it may become a critical issue where
erosion affects extremely small-range endemic
species. This is, for example, the case on the
Desertas, a small island group off Madeira,
and the other satellite islands, as well as
many of the small islands and island-groups
in the Aegean Sea. Trampling and massive
overgrazing of vegetation by non-native
species (goats) had led to serious erosion
across large areas on the Madeiran islands.
Consequently, mollusc populations crashed
so low that they were not recorded during
eld surveys in the 1980s and 1990s. Today,
the goat population has been eradicated,
and the vegetation is now recovering slightly;
recent surveys showed that many of the
highly threatened terrestrial mollusc species
may have survived at very low density in small
pockets, and a restoration process has begun
supported by the EU LIFE Programme.
Species conservation action
LIFE is the EU’s nancial instrument supporting
nature conservation in projects throughout the
EU and can, therefore, provide an important
tool for conservation of terrestrial molluscs
in Europe. Projects that involve a variety of
actions including habitat restoration, site
purchases, communication and awareness-
raising, protected area infrastructure and
conservation planning. Such examples are
the LIFE projects operating in the Madeiran
Archipelago.
Research on species trends,
distribution and threats
There is a clear need to stimulate and support
more research, monitoring and conservation
of terrestrial mollusc species across Europe.
Bilz et al. (2012: 69) stress that “…countries
with the least capacity [to monitor biodiversity]
tend to be custodians of the greatest quantity
of wildlife. Least is known about the status
of species where diversity is greatest”. The
opposite is true in terrestrial molluscs: as can
be derived from the distribution pattern of the
DD species, the least known species and the
highest decit in research is concentrated in
considerably wealthy countries like France,
Switzerland, Italy, and Austria.
Identication of KBA and AZE sites
for terrestrial molluscs
The IUCN European Red List can be used
to help prioritize sites (KBAs) and species
for conservation action. At present, a KBA
exercise has not commenced for terrestrial
molluscs. However, given the high proportion
of endemic and threatened species, it is
considered that terrestrial molluscs would
make ideal case studies to extend the currently
identied KBAs in Europe as a whole and for
the Mediterranean region specically (Darwall
et al., 2014; Máiz-Tomé et al., 2017). AZE
sites should also be identied for threatened
(Endangered and Critically Endangered)
single-site endemic species that meet the AZE
thresholds.
18
On the ground island conservation of snails through LIFE
Since 2010, three European Union LIFE
projects (LIFE Ilhéus do Porto Santo, LIFE
Maciço Montanhoso, and LIFE Recover
Natura) have contributed to land snail
conservation in the Madeira Archipelago,
focusing on species listed on the EU
Habitats Directive. Many of these species
occur in areas where intensive habitat
degradation has occurred in the form of
res, erosion, human activities, or the impact
of invasive species (e.g. mice, rabbits, the
Argentine Ant, or invasive plants).
Through each project, complete species
inventories were produced, and accurate
species distributions and populations
estimates produced for the target species.
The research also identied two new
species of Discula snails, and rediscovered
‘Lazarus’ species (e.g. Geomitra coronula
and Discula lyelliana) that had previously
been considered extinct.
Conservation actions such as ex situ
reproduction for population reinforcement
(e.g. Atlantica calathoides), the
establishment of new subpopulations
of priority species based on habitat
modelling (e.g. Wollastonaria turricula),
and the introduction of articial shelters for
species protection, have contributed to the
conservation of the target species. Indirect
measures have played an important role in
the restoration of the ecosystem equilibrium,
resulting in habitat recovery and enhancing
the lifecycle of the species. Interventions
have included habitat recovery through the
reduction or complete removal of invasive
plants (e.g. Nicotiana glauca and Phalaris
sp.) and animals (e.g. rabbits), which
hugely impact the species habitat, and the
elimination or mitigation of the impact of
predators such as mice and Argentine Ant
for example.
One of the most important outputs of the
LIFE projects has been the implementation
of monitoring schemes focused on the
Habitats Directive species. For these
species, a capture-mark-recapture
monitoring scheme is ongoing, along
with climate monitoring within each
subpopulation through data loggers, and
this work provides important information
about the species, such as lifecycle and
ecology data. In addition, ten-year species
action plans were produced, encompassing
the necessary measures, conservation
actions and plans to ensure a good
conservation status for the EU Habitats
Directive species.
Dinarte Teixeira
Further reading
IFCN. (2016). Final Report LIFE Ilhéus do Porto Santo (LIFE09/NAT/PT/000041). Instituto das Florestas e
Conservação da Natureza, Funchal.
IFCN. (2017a). Mid-term report Recover Natura (LIFE12/NAT/PT/000195). Instituto das Florestas e Conservação
da Natureza, Funchal.
IFCN. (2017b). Layman Report Recovery and conservation of species and habitats of the Madeiran Central
Mountainous Massif (LIFE11/NAT/PT/000327). Instituto das Florestas e Conservação da Natureza, Funchal.
IFCN. (2018). Final Report Maciço Montanhoso Central (LIFE11/NAT/PT/000327). Instituto das Florestas e
Conservação da Natureza, Funchal.
Alves, F., Rodrigues, J., Montes, R., Menezes, D., Oliveira, P. and Silva, V. (2015). Islets of Porto Santo: a treasure
to be preserved. Serviço do Parque Natural da Madeira (SPNM), Funchal.
SPNM. (2015). Layman’s Report do Projeto ‘Travar a perda da Biodiversidade Europeia através da recuperação
de habitats e espécies dos Ilhéus do Porto Santo e área marinha envolvente’ (LIFE09 NAT/PT/000041).
Instituto das Florestas e Conservação da Natureza, Funchal.
19
Key Recommendations
Policy
The IUCN European Red List of
terrestrial molluscs should be used to
inform nature and biodiversity policies to
improve the status of threatened species
and should be revised at regular intervals
of ten years. Individual species should be
reassessed whenever new data become
available, for example on their distribution
and novel threats, or as a result of
taxonomic changes and the description
of new species.
Measures should be promoted at the
EU-level to ensure the conservation of
key habitats, for example, limestone
habitats above and below ground,
fenland areas, ancient woodlands,
ancient banks and hedges and intact
forest ecosystems.
The Common Agricultural Policy should
promote the appropriate management
and protection of wood pasture habitats
and chalk grasslands across Europe.
Council of Europe Recommendations
No R (88) 112 on ancient natural and
semi-natural woodlands and R (88) 103
on the protection and management of
saproxylic organisms and their biotopes
2 Available at: https://rm.coe.int/090000168090a6bc
3 Available at: https://rm.coe.int/090000168090a6ba
Wollastonaria turricula (VU)
is an example of an extreme
small-range snail species,
endemic to the Ilhéu de Cima,
Madeira.
© Dinarte Teixeira
VULNERABLE
VU
20
of the Council of Europe to the Member
States should be fully implemented.
Measures should be put in place to
prevent illegal logging and ensure control
of wood collecting.
Guidance should be developed on
best practices for terrestrial mollusc
conservation in Natura 2000 sites.
Threatened molluscs should be specied
as species of conservation interest when
they exist in Natura 2000 sites or other
national/local nature reserves.
Species and habitat
conservation
Conservation strategies for European
terrestrial molluscs identied to be of
elevated conservation concern should be
developed and implemented, especially
were they are under-represented within
protected areas (Trochet and Schmeller,
2013).
Habitat fragmentation should be reduced
by creating ecological networks and
corridors.
The best habitat management practices
for European terrestrial molluscs should
be broadly adopted and relevant
stakeholders and land managers should
be made more aware of the available
sources of information.
Terrestrial mollusc inventories in Natura
2000 sites and other protected areas
should be made to identify priority
species in order to develop strategies for
their protection.
Veteran trees should be preserved
throughout Europe, in forests,
pastureland, orchards, and urban areas.
Identify those species that are known
from single sites, especially those that
are phyletically distinct as these are
candidate species for priority actions in
terms of designating KBAs and AZEs.
EU states should review that EIA
processes include evaluation of
invertebrates, especially terrestrial
molluscs, prior to quarrying of limestone
resources to ensure that these take into
account biodiversity loss, especially
in regions of high terrestrial mollusc
endemism.
Environmental Impact Assessments
for large dam projects should take into
account biodiversity loss, in subterranean
habitats especially in limestone regions
that have high levels of endemism.
Review impacts of wildres on the loss
of Mediterranean grassland species and
instigate better habitat management
practices for recovery of European
terrestrial molluscs in regions of frequent
res.
Research
Specic research on species that have
not been recently recorded in Europe or
have been assessed as Data Decient
should be conducted to clarify their
status.
The effects of poorly understood threats
(e.g., climate change) on terrestrial
molluscs need further study.
Effective monitoring tools and improved
research efforts on terrestrial mollusc
species should be developed and
promoted, particularly in the Natura
2000 network, in order to understand
population trends and the impacts of
implemented actions.
Further research is needed to identify
and protect old growth habitats in the
cultural landscapes of Europe.
21
Acknowledgments
This project would not have been possible without the enthusiastic cooperation of a large number
of people throughout Europe and further aeld who have helped and supported this project in
many different ways over the last ten years. Many of them shared their knowledge of autecology,
distribution, abundance and threats to terrestrial molluscs in their region of specialism. Others
provided access to the data held within their private collections. Some contributed by reviewing
the species accounts once they had been compiled. Others helped by correcting the species
distribution maps. In order to avoid lengthy lists, we here simply record people in alphabetical
order:
Abreu, C., Albrecht, C., Aldridge, D., Alonso, M.R., Araujo, R., Arconada, B., Badia Boher, J.A.,
Barker, G., Bichain, J.-M., Bilandzija, H., Bodon, M., Bouchet, P., Charrier, M., Cianfanelli, S.,
Cioboiu, O., Coles, B., Colville, B., Deli, T., Erőss, Z.P., Falniowski, A., Georgiev, D., Giotis, L.,
Gloer, P., Glogger, F.K., Haase, M., Hallgass, A., Hauffe, T., Jalzic, B., Lajtner, J., Jochum, A.,
Kebapcı, Ü., Killeen, I., Menez, A., Moorkens, E., Mylonas, M., Niederhöfer, H.-J., Paparisto, A.,
Pešić, V., Rađa, B., Radea, K., Ramos, M., Regnier, C., Ripken, T., Schembri, P.J., Schneppat,
U.E., Schreiber, K., Sinaco, R., Sket, B., Solymos, P., Subai, P., Subai, P., Szekeres, M., Tomovic,
J., Van Damme, D., van Goethem, J., Vavrova, L., Vinarksi, M. and Wiese, V.
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23
The designation of geographical entities in this brochure, and the presentation of the material, do not imply the
expression of any opinion whatsoever on the part of IUCN concerning the legal status of any country, territory,
or area, or of its authorities, or concerning the delimitation of its frontiers or boundaries.
The views expressed in this brochure do not necessarily reect those of IUCN. This brochure has been
prepared by IUCN as a deliverable of the LIFE European Red Lists project (LIFE14 PRE/BE/000001).
Project Title: Establishing a European Red List of Bryophytes, Pteridophytes, Saproxylic Beetles, Terrestrial
Molluscs and Vascular Plants (LIFE European Red Lists; LIFE14 PRE/BE/000001).
Project duration: May 2015 to September 2019. Project total costs: 1,166,667 EUR.
Contribution of the LIFE Programme: 700,000 EUR.
Co-nancers of the project: National Parks and Wildlife Service, Republic of Ireland; Ministry of Economic
Affairs, Department of Nature & Biodiversity (Ministerie van Economische Zaken, Directie Natuur &
Biodiversiteit), the Netherlands; Council of Europe; Ofce fédéral de l‘Environnement, Switzerland; Swedish
Environmental Protection Agency (Naturvardsverket), Sweden; British Entomological Society, United Kingdom;
Ministry of Sustainable Development and Infrastructure, Government of the Grand-Duché of Luxembourg.
The LIFE Programme (https://ec.europa.eu/easme/en/life) is the EU’s nancial instrument supporting
environmental, nature conservation and climate action projects throughout the EU. The general objective
of LIFE is to contribute to the implementation, updating and development of EU environmental, nature
conservation and climate policy and legislation by co-nancing projects with European added value.
The report is available online at http://ec.europa.eu/environment/nature/conservation/species/redlist
and https://www.iucnredlist.org/regions/europe
Published by: IUCN, Cambridge (UK) and Brussels (Belgium).
Copyright: 2019 IUCN, International Union for Conservation of Nature and Natural Resources. Licensed to
the European Union under conditions. Reproduction of this brochure for educational or other non-commercial
purposes is authorised without prior written permission from the copyright holder provided the source is fully
acknowledged. Reproduction of this brochure for resale or other commercial purposes is prohibited without
prior written permission of the copyright holder.
Citation: Neubert, E., Seddon, M.B., Allen, D.J., Arrébola, J., Backeljau, T., Balashov, I., Bank, R., Cameron,
R., de Frias Martins, A.M., De Mattia, W., Dedov, I., Duda, M., Falkner, G., Falkner, M., Fehér, Z., Gargominy,
O., Georgiev, D., Giusti, F., Gómez Moliner, B.J., Groh, K., Ibáñez, M., Kappes, H., Manganelli, G., Martínez-
Ortí, A., Nardi, G., Neiber, M.T., Páll-Gergely, B., Parmakelis, A., Prié, V., Reischütz, A., Reischütz, P.L.,
Rowson, B., Rüetschi, J., Slapnik, R., Son, M., Štamol, V., Teixeira, D., Triantis, K., Vardinoyannis, K., von
Proschwitz, T. and Walther, F. (2019). European Red List of Terrestrial Molluscs. IUCN: Cambridge, UK and
Brussels, Belgium. https://portals.iucn.org/library/node/48439
Design and layout: Imre Sebestyén jr. / UNITgraphics.com
All photographs used in this publication remain the property of the original copyright holder (see individual
captions for details). Photographs should not be reproduced or used in other contexts without written
permission from the copyright holder.
INTERNATIONAL UNION
FOR CONSERVATION OF NATURE
EUROPEAN REGIONAL OFFICE
64 Boulevard Louis Schmidt
1040 Brussels
Belgium
Tel: +32 2 739 0317
Fax: +32 2 732 9499
www.iucn.org/regions/europe
Alinda wagneri (LC) – is
endemic to the western parts
of the Stara Planina Mountains
in Bulgaria and Serbia. It is
assessed as Least Concern.
© Ivailo Dedov
LEAST CONCERN
LC
Neubert, E., Seddon, M.B., Allen, D.J., Arrébola, J., Backeljau, T., Balashov, I., Bank, R., Cameron, R., de Frias Martins, A.M., De Mattia, W., Dedov, I., Duda, M., Falkner, G., Falkner, M., Fehér, Z., Gargominy, O., Georgiev,
D., Giusti, F., Gómez Moliner, B.J., Groh, K., Ibáñez, M., Kappes, H., Manganelli, G., Martínez-Ortí, A., Nardi, G., Neiber, M.T., Páll-Gergely, B., Parmakelis, A., Prié, V., Reischütz, A., Reischütz, P.L., Rowson, B., Rüetschi,
J., Slapnik, R., Son, M., Štamol, V., Teixeira, D., Triantis, K., Vardinoyannis, K., von Proschwitz, T. and Walther, F. 2019.
Supplementary Material to the European Red List of terrestrial molluscs
. Cambridge, UK: IUCN.
Available at: https://portals.iucn.org/library/node/48439
1
Supplementary Material to the European Red List of terrestrial molluscs
This document provides the Red List Category and Criteria for the European terrestrial mollusc species assessed on the IUCN European Red List. Data were compiled as part of
the LIFE project, ‘Establishing a European Red List of Bryophytes, Pteridophytes, Saproxylic Beetles, Terrestrial Molluscs and Vascular Plants (LIFE European Red Lists; LIFE14
PRE/BE/000001)’. Includes data compiled through an earlier stage of the European Red List: Cuttelod, A., Seddon, M. and Neubert, E. 2011.
European Red List of Non-marine
Molluscs
. Luxembourg: Publications Office of the European Union.
An * indicates that a species was assessed for the EU 27 Member States, i.e., prior to the accession of Croatia in 2013.
IUCN Red List status of European terrestrial mollusc species
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Aciculidae
Acicula beneckei
Yes
NT
Yes*
NT
Aciculidae
Acicula benoiti
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Acicula corcyrensis
Yes
LC
Yes*
LC
Aciculidae
Acicula disjuncta
Yes
NT
No*
NT
Aciculidae
Acicula douctouyrensis
Yes
DD
Yes*
DD
Aciculidae
Acicula fusca
Yes
LC
Yes*
LC
Aciculidae
Acicula hausdorfi
Yes
NT
Yes*
NT
Aciculidae
Acicula lineata
Yes
LC
No
LC
Aciculidae
Acicula lineolata
Yes
LC
No
LC
Aciculidae
Acicula miaphene
Yes
NT
No
NA
Aciculidae
Acicula multilineata
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Acicula norrisi
Yes
VU
B1ab(iii)+2ab(iii); D2
Yes
VU
B1ab(iii)+2ab(iii); D2
Aciculidae
Acicula parcelineata
No
LC
No
LC
Aciculidae
Acicula szigethyannae
Yes
LC
No
LC
Aciculidae
Acicula vezzanii
Yes
NT
Yes*
NT
Aciculidae
Menkia dewinteri
Yes
DD
Yes*
DD
Aciculidae
Menkia horsti
Yes
NT
Yes*
NT
2
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Aciculidae
Menkia rolani
Yes
DD
Yes*
DD
Aciculidae
Platyla banatica
Yes
LC
No
LC
Aciculidae
Platyla callostoma
Yes
LC
Yes*
LC
Aciculidae
Platyla ceraunorum
Yes
DD
No
NA
Aciculidae
Platyla corpulenta
Yes
LC
No
NA
Aciculidae
Platyla cryptomena
Yes
LC
Yes*
LC
Aciculidae
Platyla curtii
Yes
LC
No
LC
Aciculidae
Platyla dupuyi
Yes
LC
Yes*
LC
Aciculidae
Platyla elisabethae
Yes NT No* NA
Aciculidae
Platyla feheri
Yes
NT
No
NA
Aciculidae
Platyla foliniana
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Platyla gracilis
Yes
LC
No
LC
Aciculidae
Platyla hedionda
Yes
DD
Yes
DD
Aciculidae
Platyla jankowskiana
Yes
VU
D2
No*
NA
Aciculidae
Platyla lusitanica
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Platyla maasseni
Yes
VU
D2
No*
NA
Aciculidae
Platyla microspira
Yes
LC
Yes*
LC
Aciculidae
Platyla minutissima
Yes
NT
Yes*
NT
Aciculidae
Platyla orthostoma
Yes
NT
Yes*
NT
Aciculidae
Platyla peloponnesica
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Platyla perpusilla
Yes
LC
No
LC
Aciculidae
Platyla pezzolii
Yes
NT
Yes*
NT
Aciculidae
Platyla pinteri
Yes
LC
No
LC
Aciculidae
Platyla polita
Yes
LC
No
LC
Aciculidae
Platyla procax
Yes
VU
D2
No*
NA
Aciculidae
Platyla sardoa
Yes
NT
Yes*
NT
Aciculidae
Platyla similis
Yes
LC
No
LC
Aciculidae
Platyla stussineri
Yes
LC
Yes*
LC
Aciculidae
Platyla subdiaphana
Yes
NT
Yes*
NT
Aciculidae
Platyla talentii
Yes
NT
Yes*
NT
Aciculidae
Platyla wilhelmi
Yes
LC
No*
NA
Aciculidae
Renea berica
Yes
LC
Yes
LC
3
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Aciculidae
Renea bourguignatiana
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Aciculidae
Renea elegantissima
Yes
DD
Yes*
DD
Aciculidae
Renea gentilei
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Renea gormonti
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Renea kobelti
Yes
NT
No*
NA
Aciculidae
Renea moutonii
Yes
NT
Yes*
NT
Aciculidae
Renea paillona
Yes
VU
D2
Yes*
VU
D2
Aciculidae
Renea spectabilis
Yes
LC
No
LC
Aciculidae
Renea veneta
Yes LC Yes* LC
Agriolimacidae
Deroceras adolphi
Yes
DD
Yes
DD
Agriolimacidae
Deroceras agreste
No
LC
No
LC
Agriolimacidae
Deroceras altimirai
Yes
LC
No
LC
Agriolimacidae
Deroceras astypalaeensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras attemsi
Yes
DD
Yes
DD
Agriolimacidae
Deroceras berytensis
No
LC
No
LC
Agriolimacidae
Deroceras bisacchianum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras bistimulatum
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras boeoticum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras boettgeri
Yes
DD
Yes
DD
Agriolimacidae
Deroceras bulgaricum
Yes
DD
Yes
DD
Agriolimacidae
Deroceras bureschi
Yes
LC
Yes
LC
Agriolimacidae
Deroceras caucasicum
No
LC
No
NA
Agriolimacidae
Deroceras cazioti
Yes
NT
Yes
NT
Agriolimacidae
Deroceras chevallieri
Yes
DD
Yes
DD
Agriolimacidae
Deroceras chrysorroyatissensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras corsicum
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras cycladicum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras dallaii
Yes
DD
Yes
DD
Agriolimacidae
Deroceras dewinteri
Yes
VU
D2
Yes
DD
Agriolimacidae
Deroceras ercinae
Yes
VU
B1ab(iii)
Yes
VU
B1ab(iii)
Agriolimacidae
Deroceras famagustensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras gavdosensis
Yes
VU
D2
Yes
VU
D2
4
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Agriolimacidae
Deroceras giustianum
Yes
DD
Yes
DD
Agriolimacidae
Deroceras golcheri
Yes
LC
Yes
LC
Agriolimacidae
Deroceras gorgonium
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras halieos
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras ikaria
Yes
DD
Yes
DD
Agriolimacidae
Deroceras invadens
Yes
LC
No
LC
Agriolimacidae
Deroceras johannae
Yes
DD
Yes
DD
Agriolimacidae
Deroceras karnaniensis
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras kasium
Yes VU D2 Yes VU D2
Agriolimacidae
Deroceras keanense
Yes
LC
Yes
LC
Agriolimacidae
Deroceras klemmi
Yes
LC
Yes
LC
Agriolimacidae
Deroceras korthionensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras kythirensis
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras laeve
No
LC
No
LC
Agriolimacidae
Deroceras lasithionensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras levisarcobelum
Yes
LC
No
LC
Agriolimacidae
Deroceras lombricoides
Yes
LC
Yes
LC
Agriolimacidae
Deroceras lothari
Yes
LC
Yes
LC
Agriolimacidae
Deroceras maasseni
Yes
NT
No
NA
Agriolimacidae
Deroceras malkini
No
LC
No
LC
Agriolimacidae
Deroceras melinum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras minoicum
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras moldavicum
Yes
LC
No
LC
Agriolimacidae
Deroceras neuteboomi
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras nitidum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras nyphoni
Yes
VU
D2
Yes
VU
D2
Agriolimacidae
Deroceras occidentale
Yes
LC
No
LC
Agriolimacidae
Deroceras oertzeni
Yes
LC
Yes
LC
Agriolimacidae
Deroceras pageti
Yes
LC
Yes
LC
Agriolimacidae
Deroceras panormitanum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras parium
Yes
LC
Yes
LC
Agriolimacidae
Deroceras parnasium
Yes
LC
Yes
LC
5
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Agriolimacidae
Deroceras planarioides
Yes
LC
Yes
LC
Agriolimacidae
Deroceras ponsonbyi
No
DD
No
DD
Agriolimacidae
Deroceras praecox
Yes
LC
Yes
LC
Agriolimacidae
Deroceras pseudopanormitanum
Yes
LC
Yes
LC
Agriolimacidae
Deroceras rethimnonensis
Yes
LC
Yes
LC
Agriolimacidae
Deroceras reticulatum
No
LC
No
LC
Agriolimacidae
Deroceras rhodensis
Yes
DD
Yes
DD
Agriolimacidae
Deroceras roblesi
Yes
LC
Yes
LC
Agriolimacidae
Deroceras rodnae
Yes LC No LC
Agriolimacidae
Deroceras samium
Yes
LC
Yes
LC
Agriolimacidae
Deroceras sardum
Yes
DD
Yes
DD
Agriolimacidae
Deroceras seriphium
Yes
LC
Yes
LC
Agriolimacidae
Deroceras sturanyi
Yes
LC
No
LC
Agriolimacidae
Deroceras tarracense
Yes
NT
Yes
NT
Agriolimacidae
Deroceras tauricum
Yes
LC
No
NA
Agriolimacidae
Deroceras thersites
Yes
LC
Yes
LC
Agriolimacidae
Deroceras turcicum
No
LC
No
LC
Agriolimacidae
Deroceras vascoanum
Yes
NT
Yes
NT
Agriolimacidae
Deroceras zilchi
Yes
DD
Yes
DD
Agriolimacidae
Furcopenis circularis
Yes
NT
Yes
NT
Agriolimacidae
Furcopenis darioi
Yes
VU
B1ab(iii)+2ab(iii)
Yes
VU
B1ab(iii)+2ab(iii)
Agriolimacidae
Furcopenis gallaeciensis
Yes
NT
Yes
NT
Agriolimacidae
Furcopenis geresiensis
Yes
NT
Yes
NT
Agriolimacidae
Krynickillus melanocephalus
No
LC
No
NA
Agriolimacidae
Krynickillus urbanskii
No
LC
No
LC
Agriolimacidae
Mesolimax brauni
No
LC
No
LC
Argnidae
Agardhiella armata
Yes
LC
No
LC
Argnidae
Agardhiella banatica
Yes
NT
Yes*
NT
Argnidae
Agardhiella biarmata
Yes
LC
No*
NA
Argnidae
Agardhiella caesa
Yes
LC
Yes*
LC
Argnidae
Agardhiella crassilabris
Yes
NT
Yes*
NT
Argnidae
Agardhiella dabovici
Yes
NT
No*
NA
6
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Argnidae
Agardhiella domokosi
Yes
LC
Yes*
LC
Argnidae
Agardhiella extravaganta
Yes
DD
No*
NA
Argnidae
Agardhiella formosa
Yes
LC
No*
NA
Argnidae
Agardhiella grossui
Yes
NT
Yes*
NT
Argnidae
Agardhiella incerta
Yes
LC
No
LC
Argnidae
Agardhiella lamellata
Yes
LC
Yes*
LC
Argnidae
Agardhiella langaleta
Yes
NT
Yes*
NT
Argnidae
Agardhiella macrodonta
Yes
LC
No
LC
Argnidae
Agardhiella mista
Yes NT Yes NT
Argnidae
Agardhiella parreyssii
Yes
LC
Yes*
LC
Argnidae
Agardhiella pirotana
Yes
NT
No*
NA
Argnidae
Agardhiella reinhardti
Yes
NT
Yes*
NT
Argnidae
Agardhiella serbica
Yes
NT
No*
NA
Argnidae
Agardhiella skipetarica
Yes
LC
No
LC
Argnidae
Agardhiella stenostoma
Yes
LC
No*
NA
Argnidae
Agardhiella truncatella
Yes
LC
No
LC
Argnidae
Agardhiella tunde
Yes
VU
D2
Yes*
VU
D2
Argnidae
Agardhiella zoltanorum
Yes
LC
No*
NA
Argnidae
Argna bielzi
Yes
LC
No
LC
Argnidae
Argna biplicata
Yes
LC
Yes*
LC
Argnidae
Argna bourguignatiana
Yes
VU
D2
Yes*
VU
D2
Argnidae
Argna ferrari
Yes
LC
No
LC
Argnidae
Argna szekeresi
Yes
DD
No
NA
Argnidae
Argna thracica
Yes
LC
Yes*
LC
Argnidae
Argna valsabina
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Argnidae
Speleodentorcula beroni
Yes
VU
D2
Yes*
VU
D2
Arionidae
Arion anthracius
Yes
LC
Yes
LC
Arionidae
Arion ater
Yes
LC
No
LC
Arionidae
Arion atripunctatus
Yes
DD
Yes
DD
Arionidae
Arion baeticus
Yes
NT
Yes
NT
Arionidae
Arion circumscriptus
Yes
LC
No
LC
Arionidae
Arion distinctus
Yes
LC
No
LC
7
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Arionidae
Arion euthymeanus
Yes
DD
Yes
DD
Arionidae
Arion fagophilus
Yes
LC
No
LC
Arionidae
Arion fasciatus
Yes
LC
No
LC
Arionidae
Arion flagellus
Yes
LC
Yes
LC
Arionidae
Arion franciscoloi
Yes
LC
Yes
LC
Arionidae
Arion fuligineus
Yes
LC
Yes
DD
Arionidae
Arion fuscus
Yes
LC
No
LC
Arionidae
Arion gilvus
Yes
LC
Yes
LC
Arionidae
Arion hispanicus
Yes DD Yes DD
Arionidae
Arion hortensis
Yes
LC
No
LC
Arionidae
Arion intermedius
Yes
LC
No
LC
Arionidae
Arion iratii
Yes
VU
D2
Yes
VU
D2
Arionidae
Arion lizarrustii
Yes
DD
Yes
DD
Arionidae
Arion luisae
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Arionidae
Arion lusitanicus
Yes
LC
Yes
LC
Arionidae
Arion magnus
Yes
DD
Yes
DD
Arionidae
Arion molinae
Yes
DD
Yes
DD
Arionidae
Arion nobrei
Yes
DD
Yes
DD
Arionidae
Arion obesoductus
Yes
LC
No
LC
Arionidae
Arion occultus
Yes
DD
Yes
DD
Arionidae
Arion owenii
Yes
LC
No
LC
Arionidae
Arion paularensis
Yes
DD
Yes
DD
Arionidae
Arion ponsi
Yes
NT
Yes
NT
Arionidae
Arion rufus
Yes
LC
No
LC
Arionidae
Arion silvaticus
Yes
LC
No
LC
Arionidae
Arion simrothi
Yes
DD
Yes
DD
Arionidae
Arion subfuscus
Yes
LC
Yes
LC
Arionidae
Arion transsylvanus
Yes
LC
Yes
LC
Arionidae
Arion urbiae
Yes
LC
Yes
LC
Arionidae
Arion vejdowskyi
Yes
DD
Yes
DD
Arionidae
Arion vulgaris
Yes
LC
No
LC
Arionidae
Arion wiktori
Yes
DD
Yes
DD
8
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Arionidae
Ariunculus isselii
Yes
LC
Yes
LC
Arionidae
Ariunculus speziae
Yes
LC
No
LC
Arionidae
Geomalacus anguiformis
Yes
NT
Yes
NT
Arionidae
Geomalacus maculosus
Yes
LC
Yes
LC
Arionidae
Geomalacus malagensis
Yes
LC
Yes
LC
Arionidae
Geomalacus oliveirae
Yes
NT
Yes
NT
Arionidae
Letourneuxia moreleti
Yes
DD
Yes
DD
Assimineidae
Assiminea avilai
Yes
DD
Yes
DD
Assimineidae
Assiminea cardonae
Yes DD Yes DD
Assimineidae
Assiminea eliae
No
LC
No
LC
Assimineidae
Assiminea gittenbergeri
No
DD
No
DD
Assimineidae
Assiminea glaubrechti
Yes
DD
Yes
DD
Assimineidae
Assiminea grayana
Yes
LC
Yes
LC
Assimineidae
Assiminea ostiorum
Yes
DD
Yes
DD
Assimineidae
Assiminea rolani
Yes
DD
Yes
DD
Assimineidae
Paludinella globularis
No
LC
No
LC
Boettgerillidae
Boettgerilla pallens
No
NA
No
NA
Bradybaenidae
Fruticicola fruticum
Yes
LC
No
LC
Charopidae
Discocharopa aperta
No
NA
No
NA
Chondrinidae
Abida ateni
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Abida attenuata
Yes
LC
Yes*
LC
Chondrinidae
Abida bigerrensis
Yes
LC
Yes*
LC
Chondrinidae
Abida cylindrica
Yes
LC
Yes*
LC
Chondrinidae
Abida gittenbergeri
Yes
NT
Yes*
NT
Chondrinidae
Abida occidentalis
Yes
LC
No
LC
Chondrinidae
Abida partioti
Yes
LC
Yes*
LC
Chondrinidae
Abida polyodon
Yes
LC
No
LC
Chondrinidae
Abida pyrenaearia
Yes
LC
Yes*
LC
Chondrinidae
Abida secale
Yes
LC
No
LC
Chondrinidae
Abida vasconica
Yes
LC
Yes*
LC
Chondrinidae
Abida vergniesiana
Yes
NT
No*
NT
Chondrinidae
Chondrina altimirai
Yes
NT
Yes*
NT
9
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Chondrinidae
Chondrina arcadica
No
LC
No
LC
Chondrinidae
Chondrina arigonis
Yes
NT
Yes*
NT
Chondrinidae
Chondrina ascendens
Yes
LC
Yes*
LC
Chondrinidae
Chondrina avenacea
Yes
LC
No
LC
Chondrinidae
Chondrina bergomensis
Yes
NT
Yes*
NT
Chondrinidae
Chondrina bigorriensis
Yes
LC
Yes*
LC
Chondrinidae
Chondrina calpica
No
LC
No
LC
Chondrinidae
Chondrina centralis
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Chondrina falkneri
Yes NT Yes* NT
Chondrinidae
Chondrina farinesii
Yes
LC
No
LC
Chondrinidae
Chondrina feneriensis
Yes
NT
Yes
NT
Chondrinidae
Chondrina gasulli
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Chondrinidae
Chondrina generosensis
Yes
LC
No
DD
Chondrinidae
Chondrina gerhardi
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Chondrina granatensis
Yes
NT
Yes*
NT
Chondrinidae
Chondrina kobelti
Yes
LC
Yes*
LC
Chondrinidae
Chondrina kobeltoides
Yes
LC
Yes*
LC
Chondrinidae
Chondrina lusitanica
Yes
LC
Yes*
LC
Chondrinidae
Chondrina maginensis
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Chondrina megacheilos
Yes
LC
No
LC
Chondrinidae
Chondrina multidentata
Yes
LC
Yes*
LC
Chondrinidae
Chondrina oligodonta
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Chondrina ripkeni
Yes
NT
Yes*
NT
Chondrinidae
Chondrina spelta
Yes
LC
No
LC
Chondrinidae
Chondrina tenuimarginata
Yes
LC
No
LC
Chondrinidae
Granaria braunii
Yes
LC
No
LC
Chondrinidae
Granaria frumentum
Yes
LC
No
LC
Chondrinidae
Granaria stabilei
Yes
LC
Yes*
LC
Chondrinidae
Granaria variabilis
Yes
DD
No*
DD
Chondrinidae
Granopupa granum
No
LC
No
LC
Chondrinidae
Rupestrella dupotetii
No
VU
D2
No*
VU
D2
Chondrinidae
Rupestrella homala
Yes
VU
D2
Yes*
VU
D2
10
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Chondrinidae
Rupestrella jaeckeli
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Rupestrella occulta
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Rupestrella philippii
No
LC
No
LC
Chondrinidae
Rupestrella rhodia
No
LC
No
LC
Chondrinidae
Rupestrella rupestris
Yes
NT
Yes*
NT
Chondrinidae
Solatopupa cianensis
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Solatopupa guidoni
Yes
DD
Yes*
DD
Chondrinidae
Solatopupa juliana
Yes
LC
Yes*
LC
Chondrinidae
Solatopupa pallida
Yes DD Yes* DD
Chondrinidae
Solatopupa psarolena
Yes
VU
D2
Yes*
VU
D2
Chondrinidae
Solatopupa similis
Yes
LC
No
LC
Clausiliidae
Agathylla abrupta
Yes
NT
Yes
NT
Clausiliidae
Agathylla biloba
Yes
LC
No
NA
Clausiliidae
Agathylla exarata
Yes
LC
No
LC
Clausiliidae
Agathylla formosa
Yes
LC
No
LC
Clausiliidae
Agathylla goldi
Yes
NT
No
NA
Clausiliidae
Agathylla lamellosa
Yes
NT
No
VU
D2
Clausiliidae
Agathylla narentana
Yes
NT
Yes
NT
Clausiliidae
Agathylla neutra
Yes
LC
No
LC
Clausiliidae
Agathylla regularis
Yes
NT
No
NA
Clausiliidae
Agathylla strigillata
Yes
NT
Yes
NT
Clausiliidae
Agathylla sulcosa
Yes
LC
No
LC
Clausiliidae
Agathylla viperina
Yes
NT
Yes
NT
Clausiliidae
Albinaria adrianae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria adriani
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria alajana
No
NT
No
NT
Clausiliidae
Albinaria amalthea
Yes
LC
Yes
LC
Clausiliidae
Albinaria arcadica
Yes
LC
Yes
LC
Clausiliidae
Albinaria argynnis
Yes
LC
Yes
LC
Clausiliidae
Albinaria ariadne
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria arthuriana
Yes
LC
Yes
LC
Clausiliidae
Albinaria avia
Yes
LC
Yes
LC
11
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Albinaria brevicollis
Yes
LC
No
LC
Clausiliidae
Albinaria broemmei
Yes
LC
Yes
LC
Clausiliidae
Albinaria butoti
Yes
LC
Yes
LC
Clausiliidae
Albinaria byzantina
Yes
LC
Yes
LC
Clausiliidae
Albinaria caerulea
No
LC
No
LC
Clausiliidae
Albinaria campylauchen
Yes
LC
Yes
LC
Clausiliidae
Albinaria candida
Yes
LC
Yes
LC
Clausiliidae
Albinaria christae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria coa
No VU D2 No VU D2
Clausiliidae
Albinaria compressa
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria confusa
Yes
NT
Yes
NT
Clausiliidae
Albinaria contaminata
Yes
LC
Yes
LC
Clausiliidae
Albinaria corrugata
Yes
LC
Yes
LC
Clausiliidae
Albinaria cretensis
Yes
LC
Yes
LC
Clausiliidae
Albinaria cristatella
Yes
LC
Yes
LC
Clausiliidae
Albinaria cytherae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria discolor
Yes
LC
Yes
LC
Clausiliidae
Albinaria eburnea
Yes
LC
Yes
LC
Clausiliidae
Albinaria edmundi
Yes
LC
Yes
LC
Clausiliidae
Albinaria evelynae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria freytagi
Yes
LC
Yes
LC
Clausiliidae
Albinaria fuchskaeufeli
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria fulvula
Yes
DD
Yes
DD
Clausiliidae
Albinaria gerolimena
Yes
LC
Yes
LC
Clausiliidae
Albinaria grayana
Yes
NT
Yes
NT
Clausiliidae
Albinaria greeni
Yes
LC
Yes
LC
Clausiliidae
Albinaria grisea
Yes
LC
Yes
LC
Clausiliidae
Albinaria haessleini
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria hians
Yes
LC
Yes
LC
Clausiliidae
Albinaria hippolyti
Yes
LC
Yes
LC
Clausiliidae
Albinaria hohorsti
Yes
LC
Yes
LC
Clausiliidae
Albinaria idaea
Yes
LC
Yes
LC
12
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Albinaria idyllica
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria ithomensis
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria jaeckeli
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria janicollis
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria janisadana
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria jonica
Yes
LC
Yes
LC
Clausiliidae
Albinaria klemmi
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria krueperi
Yes
LC
Yes
LC
Clausiliidae
Albinaria leonisorum
Yes DD Yes DD
Clausiliidae
Albinaria lerosiensis
No
LC
No
LC
Clausiliidae
Albinaria li
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria linnei
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria litoraria
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria maltzani
Yes
LC
Yes
LC
Clausiliidae
Albinaria mavromoustakisi
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Clausiliidae
Albinaria menelaus
Yes
LC
Yes
LC
Clausiliidae
Albinaria mixta
Yes
LC
Yes
LC
Clausiliidae
Albinaria moreletiana
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria munda
No
LC
No
LC
Clausiliidae
Albinaria nivea
Yes
LC
Yes
LC
Clausiliidae
Albinaria olivieri
Yes
LC
Yes
LC
Clausiliidae
Albinaria pelocarinata
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria petrosa
Yes
LC
Yes
VU
D2
Clausiliidae
Albinaria praeclara
Yes
LC
Yes
LC
Clausiliidae
Albinaria profuga
Yes
NT
Yes
NT
Clausiliidae
Albinaria proteus
Yes
LC
Yes
LC
Clausiliidae
Albinaria rebeli
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria rechingeri
Yes
CR
B2ab(iii)
Yes
CR
B2ab(iii)
Clausiliidae
Albinaria retusa
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Yes
CR
B1ab(i,iii)+2ab(i,iii)
Clausiliidae
Albinaria rollei
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria saxatilis
Yes
LC
Yes
LC
Clausiliidae
Albinaria schuchii
Yes
LC
No
LC
13
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Albinaria scopulosa
Yes
LC
No
LC
Clausiliidae
Albinaria senilis
Yes
LC
No
LC
Clausiliidae
Albinaria solicola
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria sphakiota
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria spratti
Yes
LC
Yes
LC
Clausiliidae
Albinaria sturanyi
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria subaii
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria sublamellosa
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria tenuicostata
Yes LC Yes LC
Clausiliidae
Albinaria terebra
Yes
LC
Yes
LC
Clausiliidae
Albinaria teres
Yes
LC
Yes
LC
Clausiliidae
Albinaria thiesseae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria torticollis
Yes
EN
B2ab(iii)
Yes
EN
B2ab(iii)
Clausiliidae
Albinaria troglodytes
Yes
LC
Yes
LC
Clausiliidae
Albinaria turrita
Yes
LC
Yes
LC
Clausiliidae
Albinaria ulrikae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria unicolor
Yes
NT
Yes
NT
Clausiliidae
Albinaria violacea
Yes
NT
Yes
NT
Clausiliidae
Albinaria virginea
Yes
LC
Yes
LC
Clausiliidae
Albinaria virgo
Yes
LC
Yes
LC
Clausiliidae
Albinaria voithii
Yes
LC
Yes
LC
Clausiliidae
Albinaria vrondamasa
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria wettsteini
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria wiesei
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria xanthostoma
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Albinaria zilchi
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alinda atanasovi
Yes
LC
Yes
LC
Clausiliidae
Alinda biplicata
Yes
LC
No
LC
Clausiliidae
Alinda elegantissima
Yes
NT
No
NT
Clausiliidae
Alinda fallax
Yes
LC
No
LC
Clausiliidae
Alinda jugularis
Yes
LC
Yes
LC
Clausiliidae
Alinda nordsiecki
Yes
LC
No
LC
14
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Alinda pancici
Yes
VU
D2
No
NA
Clausiliidae
Alinda serbica
Yes
LC
No
LC
Clausiliidae
Alinda stabilis
Yes
LC
No
LC
Clausiliidae
Alinda viridana
Yes
LC
Yes
LC
Clausiliidae
Alinda vratzatica
Yes
NT
Yes
NT
Clausiliidae
Alinda wagneri
Yes
LC
No
LC
Clausiliidae
Alopia alpina
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia bielzii
Yes
LC
Yes
LC
Clausiliidae
Alopia bogatensis
Yes LC Yes LC
Clausiliidae
Alopia canescens
Yes
LC
Yes
LC
Clausiliidae
Alopia glauca
Yes
LC
Yes
LC
Clausiliidae
Alopia glorifica
Yes
LC
Yes
LC
Clausiliidae
Alopia grossuana
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia hirschfelderi
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia lischkeana
Yes
LC
Yes
LC
Clausiliidae
Alopia livida
Yes
LC
Yes
LC
Clausiliidae
Alopia maciana
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia mafteiana
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia mariae
Yes
LC
Yes
LC
Clausiliidae
Alopia meschendorferi
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia monacha
Yes
LC
Yes
LC
Clausiliidae
Alopia nefasta
Yes
LC
Yes
LC
Clausiliidae
Alopia nixa
Yes
LC
Yes
LC
Clausiliidae
Alopia plumbea
Yes
LC
Yes
LC
Clausiliidae
Alopia pomatias
Yes
LC
Yes
LC
Clausiliidae
Alopia regalis
Yes
LC
Yes
LC
Clausiliidae
Alopia straminicollis
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Alopia subcosticollis
Yes
NT
Yes
NT
Clausiliidae
Alopia vicina
Yes
NT
Yes
NT
Clausiliidae
Balea heydeni
Yes
LC
No
LC
Clausiliidae
Balea kaeufeli
Yes
LC
Yes
LC
Clausiliidae
Balea nitida
Yes
NT
B2a
Yes
NT
B2a
15
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Balea perversa
Yes
LC
No
LC
Clausiliidae
Boettgeria crispa
Yes
EN
B1ab(ii,iii,iv,v)+2ab(ii,iii,iv,v)
Yes
EN
B1ab(ii,iii,iv,v)+2ab(ii,iii,iv,v)
Clausiliidae
Boettgeria deltostoma
Yes
LC
Yes
LC
Clausiliidae
Boettgeria depauperata
Yes
LC
Yes
LC
Clausiliidae
Boettgeria exigua
Yes
LC
Yes
LC
Clausiliidae
Boettgeria jensi
Yes
LC
Yes
LC
Clausiliidae
Boettgeria lowei
Yes
LC
Yes
LC
Clausiliidae
Boettgeria obesiuscula
Yes
CR
B1ab(ii,iii)+2ab(ii,iii)
Yes
CR
B1ab(ii,iii)+2ab(ii,iii)
Clausiliidae
Bofilliella subarcuata
Yes NT Yes NT
Clausiliidae
Bulgarica bulgariensis
Yes
LC
Yes
LC
Clausiliidae
Bulgarica cana
Yes
LC
No
LC
Clausiliidae
Bulgarica denticulata
No
LC
No
LC
Clausiliidae
Bulgarica erberi
No
LC
No
LC
Clausiliidae
Bulgarica fraudigera
Yes
LC
Yes
LC
Clausiliidae
Bulgarica fritillaria
Yes
LC
Yes
LC
Clausiliidae
Bulgarica hiltrudae
Yes
LC
Yes
LC
Clausiliidae
Bulgarica iniucunda
Yes
LC
Yes
LC
Clausiliidae
Bulgarica moellendorffi
Yes
VU
D2
No
NA
Clausiliidae
Bulgarica mystica
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Bulgarica pavlovici
Yes
NT
No
NA
Clausiliidae
Bulgarica pindica
Yes
NT
Yes
NT
Clausiliidae
Bulgarica pseudofraudigera
Yes
LC
Yes
LC
Clausiliidae
Bulgarica rugicollis
Yes
LC
No
LC
Clausiliidae
Bulgarica stolii
Yes
VU
D2
No
NA
Clausiliidae
Bulgarica urbanskii
Yes
NT
Yes
NT
Clausiliidae
Bulgarica varnensis
Yes
LC
Yes
LC
Clausiliidae
Bulgarica vetusta
Yes
LC
No
LC
Clausiliidae
Carinigera albicosta
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera buresi
Yes
LC
Yes
LC
Clausiliidae
Carinigera calabacensis
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera chelidromia
Yes
LC
Yes
LC
Clausiliidae
Carinigera dextrorsa
Yes
LC
Yes
LC
16
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Carinigera drenovoensis
Yes
CR
B1ab(iii)
No
NA
Clausiliidae
Carinigera eximia
Yes
LC
No
NA
Clausiliidae
Carinigera leucorhaphe
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera liebegottae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera lophauchena
Yes
EN
B1ab(i,iii)
No
EN
B1ab(i,iii)
Clausiliidae
Carinigera octava
Yes
VU
D2
No
NA
Clausiliidae
Carinigera pellucida
Yes
EN
B1ab(iii)+2ab(iii)
No
NA
Clausiliidae
Carinigera pharsalica
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Clausiliidae
Carinigera praecipua
Yes NT Yes NT
Clausiliidae
Carinigera praestans
Yes
LC
Yes
LC
Clausiliidae
Carinigera schuetti
Yes
LC
Yes
LC
Clausiliidae
Carinigera septima
Yes
VU
D2
No
NA
Clausiliidae
Carinigera stussineri
Yes
LC
Yes
LC
Clausiliidae
Carinigera superba
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera tantilla
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Carinigera thessalonica
Yes
LC
Yes
LC
Clausiliidae
Carinigera torifera
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Charpentieria calcarae
Yes
LC
Yes
LC
Clausiliidae
Charpentieria crassicostata
Yes
CR
B1ab(i,ii,iv,v); C2a(ii)
Yes
CR
B1ab(i,ii,iv,v); C2a(ii)
Clausiliidae
Charpentieria dyodon
Yes
LC
No
LC
Clausiliidae
Charpentieria eminens
Yes
EN
B1ab(iv,v)+2ab(iv,v)
Yes
EN
B1ab(iv,v)+2ab(iv,v)
Clausiliidae
Charpentieria ernae
Yes
DD
Yes
DD
Clausiliidae
Charpentieria ferrox
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Clausiliidae
Charpentieria gibbula
Yes
LC
Yes
LC
Clausiliidae
Charpentieria grohmanniana
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Clausiliidae
Charpentieria incerta
Yes
DD
Yes
DD
Clausiliidae
Charpentieria itala
Yes
LC
No
LC
Clausiliidae
Charpentieria kobeltiana
Yes
LC
Yes
LC
Clausiliidae
Charpentieria lamellata
Yes
LC
No
LC
Clausiliidae
Charpentieria leucophryna
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Clausiliidae
Charpentieria nobilis
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Clausiliidae
Charpentieria ornata
Yes
LC
Yes
LC
17
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Charpentieria paestana
Yes
LC
Yes
LC
Clausiliidae
Charpentieria pantocratoris
Yes
LC
No
LC
Clausiliidae
Charpentieria riberothi
Yes
DD
Yes
DD
Clausiliidae
Charpentieria scarificata
Yes
LC
Yes
LC
Clausiliidae
Charpentieria septemplicata
Yes
NT
Yes
NT
B1+B2
Clausiliidae
Charpentieria spezialensis
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Clausiliidae
Charpentieria splendens
Yes
LC
No
NA
Clausiliidae
Charpentieria stenzii
Yes
LC
Yes
LC
Clausiliidae
Charpentieria stigmatica
Yes LC No LC
Clausiliidae
Charpentieria tiberii
Yes
NT
Yes
NT
Clausiliidae
Charpentieria vulcanica
Yes
LC
Yes
LC
Clausiliidae
Clausilia bidentata
Yes
LC
No
LC
Clausiliidae
Clausilia brembina
Yes
LC
Yes
LC
Clausiliidae
Clausilia corynodes
Yes
LC
No
LC
Clausiliidae
Clausilia cruciata
Yes
LC
No
LC
Clausiliidae
Clausilia dubia
Yes
LC
No
LC
Clausiliidae
Clausilia pumila
Yes
LC
No
LC
Clausiliidae
Clausilia rugosa
Yes
LC
No
LC
Clausiliidae
Clausilia strobel
Yes
LC
No
LC
Clausiliidae
Clausilia whateliana
Yes
LC
Yes
LC
Clausiliidae
Cochlodina bidens
Yes
LC
Yes
LC
Clausiliidae
Cochlodina cerata
Yes
LC
No
LC
Clausiliidae
Cochlodina comensis
Yes
LC
No
LC
Clausiliidae
Cochlodina costata
Yes
LC
No
LC
Clausiliidae
Cochlodina dubiosa
Yes
LC
Yes
LC
Clausiliidae
Cochlodina fimbriata
Yes
LC
No
LC
Clausiliidae
Cochlodina inaequalis
Yes
LC
Yes
LC
Clausiliidae
Cochlodina kuesteri
Yes
LC
Yes
LC
Clausiliidae
Cochlodina laminata
No
LC
No
LC
Clausiliidae
Cochlodina liburnica
Yes
LC
Yes
LC
Clausiliidae
Cochlodina marisi
Yes
LC
No
LC
Clausiliidae
Cochlodina meisneriana
Yes
LC
Yes
LC
18
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Cochlodina orthostoma
Yes
LC
No
NT
A2
Clausiliidae
Cochlodina triloba
Yes
LC
Yes
LC
Clausiliidae
Delima albocincta
Yes
LC
Yes
LC
Clausiliidae
Delima amoena
Yes
LC
Yes
LC
Clausiliidae
Delima bilabiata
Yes
LC
No
LC
Clausiliidae
Delima binotata
Yes
LC
No
LC
Clausiliidae
Delima blanda
Yes
LC
Yes
LC
Clausiliidae
Delima edmibrani
Yes
NT
Yes
NT
Clausiliidae
Delima giselae
Yes DD Yes DD
Clausiliidae
Delima helenae
Yes
NT
Yes
NT
Clausiliidae
Delima hiltrudis
Yes
NT
Yes
NT
Clausiliidae
Delima laevissima
Yes
LC
No
LC
Clausiliidae
Delima latilabris
Yes
LC
No
LC
Clausiliidae
Delima montenegrina
Yes
LC
No
NA
Clausiliidae
Delima pachystoma
Yes
LC
No
LC
Clausiliidae
Delima pellucida
Yes
NT
Yes
NT
Clausiliidae
Delima pfeifferi
Yes
NT
Yes
VU
D2
Clausiliidae
Delima piceata
Yes
LC
Yes
LC
Clausiliidae
Delima semirugata
Yes
LC
No
LC
Clausiliidae
Delima subcylindrica
Yes
LC
Yes
LC
Clausiliidae
Delima vidovichii
Yes
LC
No
LC
Clausiliidae
Dilataria boettgeriana
Yes
LC
Yes
LC
Clausiliidae
Dilataria bosnica
Yes
NT
No
NA
Clausiliidae
Dilataria marcki
Yes
LC
Yes
LC
Clausiliidae
Dilataria pirostoma
Yes
NT
Yes
NT
Clausiliidae
Dilataria succineata
Yes
LC
Yes
LC
Clausiliidae
Dobatia goettingi
No
LC
No
LC
Clausiliidae
Elia moesta
No
LC
No
LC
Clausiliidae
Erjavecia bergeri
Yes
LC
Yes
LC
Clausiliidae
Euxina circumdata
No
LC
No
LC
Clausiliidae
Euxina persica
No
LC
No
LC
Clausiliidae
Euxina promta
No
LC
No
LC
19
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Euxinella radikae
Yes
NT
No
NA
Clausiliidae
Euxinella subaii
Yes
VU
B1ab(ii)+2ab(ii); D2
No
NA
Clausiliidae
Fusulus approximans
Yes
LC
Yes
LC
Clausiliidae
Fusulus interruptus
Yes
LC
Yes
LC
Clausiliidae
Galeata schwerzenbachii
No
LC
No
LC
Clausiliidae
Graciliaria inserta
Yes
LC
Yes
LC
Clausiliidae
Graecophaedusa sperrlei
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Herilla bosniensis
Yes
LC
No
LC
Clausiliidae
Herilla durmitoris
Yes LC No NA
Clausiliidae
Herilla illyrica
Yes
LC
No
NA
Clausiliidae
Herilla jabucica
Yes
LC
No
NA
Clausiliidae
Herilla pavlovici
Yes
LC
No
NA
Clausiliidae
Herilla ziegleri
Yes
LC
No
LC
Clausiliidae
Idyla bicristata
No
LC
No
LC
Clausiliidae
Idyla castalia
Yes
LC
Yes
LC
Clausiliidae
Idyla liebegottae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Idyla pelobsoleta
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Inchoatia haussknechti
Yes
LC
Yes
LC
Clausiliidae
Inchoatia inchoata
Yes
LC
Yes
LC
Clausiliidae
Inchoatia megdova
Yes
LC
Yes
LC
Clausiliidae
Inchoatia regina
Yes
LC
Yes
LC
Clausiliidae
Isabellaria abyssoclista
Yes
LC
Yes
LC
Clausiliidae
Isabellaria almae
Yes
LC
Yes
LC
Clausiliidae
Isabellaria bathyclista
Yes
LC
Yes
LC
Clausiliidae
Isabellaria clandestina
Yes
LC
Yes
LC
Clausiliidae
Isabellaria isabellina
Yes
LC
Yes
LC
Clausiliidae
Isabellaria lutracana
Yes
LC
Yes
LC
Clausiliidae
Isabellaria parnassia
Yes
NT
Yes
NT
Clausiliidae
Isabellaria perplana
Yes
LC
Yes
LC
Clausiliidae
Isabellaria riedeli
Yes
LC
Yes
LC
Clausiliidae
Isabellaria saxicola
Yes
LC
Yes
LC
Clausiliidae
Isabellaria sericata
Yes
LC
Yes
LC
20
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Isabellaria thermopylarum
Yes
LC
Yes
LC
Clausiliidae
Julica schmidtii
Yes
LC
Yes
LC
Clausiliidae
Laciniaria bajula
Yes
LC
Yes
LC
Clausiliidae
Laciniaria exalta
Yes
LC
Yes
LC
Clausiliidae
Laciniaria macilenta
Yes
LC
Yes
LC
Clausiliidae
Laciniaria plicata
Yes
LC
No
LC
Clausiliidae
Lampedusa imitatrix
Yes
EN
A3c; B1ab(ii,iii,v)+2ab(ii,iii,v)
Yes
EN
A3c; B1ab(ii,iii,v)+2ab(ii,iii,v)
Clausiliidae
Lampedusa lopadusae
Yes
LC
Yes
LC
Clausiliidae
Lampedusa melitensis
Yes CR B1ab(ii,iii)+2ab(ii,iii) Yes CR B1ab(ii,iii)+2ab(ii,iii)
Clausiliidae
Leucostigma candidescens
Yes
LC
Yes
LC
Clausiliidae
Macedonica brabeneci
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Macedonica frauenfeldi
Yes
LC
No
LC
Clausiliidae
Macedonica guicciardii
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Macedonica hartmuti
Yes
NT
Yes
NT
Clausiliidae
Macedonica macedonica
Yes
DD
No
DD
Clausiliidae
Macedonica marginata
Yes
LC
Yes
LC
Clausiliidae
Macedonica martae
Yes
CR
B1ab(iii)
Yes
CR
B1ab(iii)
Clausiliidae
Macedonica pindica
Yes
NT
Yes
NT
Clausiliidae
Macedonica pinteri
Yes
VU
B1ab(v)+2ab(v); D2
Yes
VU
B1ab(v)+2ab(v); D2
Clausiliidae
Macedonica pirinensis
Yes
LC
Yes
LC
Clausiliidae
Macedonica pirotana
Yes
LC
No
NA
Clausiliidae
Macedonica schatzmayri
Yes
LC
Yes
LC
Clausiliidae
Macedonica slavica
Yes
VU
D2
No
NA
Clausiliidae
Macedonica teodorae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Macedonica thasia
Yes
LC
Yes
LC
Clausiliidae
Macedonica ypsilon
Yes
LC
Yes
LC
Clausiliidae
Macedonica zilchi
Yes
VU
B1ab(v)+2ab(v); D2
Yes
VU
B1ab(v)+2ab(v); D2
Clausiliidae
Macrogastra asphaltina
Yes
LC
Yes
LC
Clausiliidae
Macrogastra attenuata
Yes
LC
No
LC
Clausiliidae
Macrogastra badia
Yes
LC
Yes
LC
Clausiliidae
Macrogastra borealis
Yes
LC
No
LC
Clausiliidae
Macrogastra densestriata
Yes
LC
No
LC
21
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Macrogastra mellae
Yes
LC
Yes
LC
Clausiliidae
Macrogastra plicatula
Yes
LC
No
LC
Clausiliidae
Macrogastra rolphii
Yes
LC
No
LC
Clausiliidae
Macrogastra tumida
Yes
LC
No
LC
Clausiliidae
Macrogastra ventricosa
Yes
LC
No
LC
Clausiliidae
Medora adensameri
Yes
NT
Yes
NT
Clausiliidae
Medora agnata
Yes
LC
Yes
LC
Clausiliidae
Medora almissana
Yes
LC
Yes
LC
Clausiliidae
Medora armata
Yes NT Yes NT
Clausiliidae
Medora contracta
Yes
LC
No
LC
Clausiliidae
Medora dalmatina
Yes
LC
No
LC
Clausiliidae
Medora eris
Yes
LC
Yes
LC
Clausiliidae
Medora hiltrudae
Yes
NT
Yes
NT
Clausiliidae
Medora italiana
Yes
LC
Yes
LC
Clausiliidae
Medora lesinensis
Yes
LC
No
NA
Clausiliidae
Medora macascarensis
Yes
LC
No
LC
Clausiliidae
Medora proxima
Yes
LC
No
NA
Clausiliidae
Medora stenostoma
Yes
NT
Yes
NT
Clausiliidae
Mentissa canalifera
Yes
LC
No
NA
Clausiliidae
Mentissa gracilicosta
Yes
LC
No
NA
Clausiliidae
Mentissa velutina
Yes
LC
No
NA
Clausiliidae
Mentissella rebeli
Yes
LC
Yes
LC
Clausiliidae
Micridyla pinteri
Yes
LC
Yes
LC
Clausiliidae
Micropontica caucasica
No
NA
No
NA
Clausiliidae
Montenegrina apfelbecki
Yes
NT
No
NA
Clausiliidae
Montenegrina attemsi
Yes
NT
No
NA
Clausiliidae
Montenegrina cattaroensis
Yes
LC
No
NA
Clausiliidae
Montenegrina chiasma
Yes
NT
No
NA
Clausiliidae
Montenegrina dofleini
Yes
LC
No
LC
Clausiliidae
Montenegrina drimmeri
Yes
NT
No
NA
Clausiliidae
Montenegrina fuchsi
Yes
LC
No
LC
Clausiliidae
Montenegrina grammica
Yes
NT
No
NT
B1a+2a; D2
22
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Montenegrina haringae
Yes
NT
No
NA
Clausiliidae
Montenegrina helvola
Yes
LC
No
NA
Clausiliidae
Montenegrina hiltrudae
Yes
LC
No
LC
Clausiliidae
Montenegrina janinensis
Yes
LC
Yes
LC
Clausiliidae
Montenegrina laxa
Yes
LC
No
LC
Clausiliidae
Montenegrina lillae
Yes
NT
No
NA
Clausiliidae
Montenegrina minuscula
Yes
NT
No
NA
Clausiliidae
Montenegrina nana
Yes
LC
No
NA
Clausiliidae
Montenegrina okolensis
Yes NT No NA
Clausiliidae
Montenegrina perstriata
Yes
LC
No
NA
Clausiliidae
Montenegrina prokletiana
Yes
NT
No
NA
Clausiliidae
Montenegrina rugilabris
Yes
LC
No
LC
Clausiliidae
Montenegrina skipetarica
Yes
LC
No
LC
Clausiliidae
Montenegrina soosi
Yes
LC
No
NA
Clausiliidae
Montenegrina sporadica
Yes
NT
No
NA
Clausiliidae
Montenegrina stankovici
Yes
VU
No
NA
Clausiliidae
Montenegrina sturanyana
Yes
NT
No
NA
Clausiliidae
Montenegrina subcristata
Yes
LC
No
NA
Clausiliidae
Montenegrina timeae
Yes
NT
No
NA
Clausiliidae
Montenegrina tomorosi
Yes
LC
No
NA
Clausiliidae
Montenegrina zilchi
Yes
NT
Yes
NT
Clausiliidae
Muticaria macrostoma
Yes
LC
Yes
LC
Clausiliidae
Muticaria neuteboomi
Yes
LC
Yes
LC
Clausiliidae
Muticaria syracusana
Yes
LC
Yes
LC
Clausiliidae
Neniatlanta pauli
Yes
LC
Yes
LC
Clausiliidae
Papillifera papillaris
No
LC
No
LC
Clausiliidae
Papillifera solida
Yes
LC
Yes
LC
Clausiliidae
Protoherilla baleiformis
Yes
LC
No
NA
Clausiliidae
Protoherilla mirabilis
Yes
VU
D2
No
NA
Clausiliidae
Protoherilla pseudofallax
Yes
LC
No
NA
Clausiliidae
Pseudofusulus varians
Yes
LC
Yes
LC
Clausiliidae
Ruthenica filograna
Yes
LC
No
LC
23
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Clausiliidae
Ruthenica gallinae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Sciocochlea collasi
Yes
NT
Yes
NT
Clausiliidae
Sciocochlea cryptica
Yes
LC
No
LC
Clausiliidae
Sciocochlea llogaraensis
Yes
LC
No
NA
Clausiliidae
Sciocochlea nordsiecki
Yes
NT
No
NT
Clausiliidae
Serrulina serrulata
No
LC
No
LC
Clausiliidae
Strigilodelima conspersa
Yes
LC
No
LC
Clausiliidae
Strigilodelima pentheri
Yes
NT
No
NA
Clausiliidae
Triloba sandrii
Yes LC No NA
Clausiliidae
Triloba thaumasia
Yes
LC
No
LC
Clausiliidae
Tsoukatosia arabatzis
Yes
NT
Yes
NT
Clausiliidae
Tsoukatosia argolica
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Tsoukatosia christinae
Yes
LC
Yes
LC
Clausiliidae
Tsoukatosia evauemgei
Yes
CR(PE)
B2ab(iii,iv); D
Yes
CR(PE)
B2ab(iii,iv); D
Clausiliidae
Tsoukatosia liae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Tsoukatosia nicoleae
Yes
VU
D2
Yes
VU
D2
Clausiliidae
Tsoukatosia pallgergelyi
Yes
NT
Yes
NT
Clausiliidae
Tsoukatosia subaii
Yes
NT
Yes
NT
Clausiliidae
Vallatia vallata
Yes
LC
Yes
LC
Clausiliidae
Vestia elata
Yes
LC
No
LC
Clausiliidae
Vestia gulo
Yes
LC
No
LC
Clausiliidae
Vestia ranojevici
Yes
LC
No
LC
Clausiliidae
Vestia roschitzi
Yes
LC
No
LC
Clausiliidae
Vestia turgida
Yes
LC
No
LC
Cochlicellidae
Cochlicella acuta
No
LC
No
LC
Cochlicellidae
Cochlicella barbara
No
LC
No
LC
Cochlicellidae
Cochlicella conoidea
No
LC
No
LC
Cochlicellidae
Monilearia arguineguinensis
Yes
CR
B2ab(ii,iii)
Yes*
CR
B2ab(ii,iii)
Cochlicellidae
Monilearia caementitia
Yes
LC
Yes*
LC
Cochlicellidae
Monilearia granostriata
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Cochlicellidae
Monilearia inops
Yes
DD
Yes*
DD
Cochlicellidae
Monilearia loweana
Yes
LC
Yes*
LC
24
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Cochlicellidae
Monilearia monilifera
Yes
LC
Yes*
LC
Cochlicellidae
Monilearia montigena
Yes
NT
Yes*
NT
Cochlicellidae
Monilearia multipunctata
Yes
NT
Yes*
NT
Cochlicellidae
Monilearia oleacea
Yes
LC
Yes*
LC
Cochlicellidae
Monilearia persimilis
Yes
LC
Yes*
LC
Cochlicellidae
Monilearia phalerata
Yes
NT
Yes*
NT
Cochlicellidae
Monilearia praeposita
Yes
DD
Yes*
DD
Cochlicellidae
Monilearia pulverulenta
Yes
CR(PE)
B1ab(iii)+2ab(iii)
Yes*
CR(PE)
B1ab(iii)+2ab(iii)
Cochlicellidae
Monilearia tubaeformis
Yes NT Yes* NT
Cochlicellidae
Monilearia tumulorum
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Cochlicellidae
Monilearia umbicula
Yes
LC
Yes*
LC
Cochlicellidae
Monilearia watsoniana
Yes
DD
Yes*
DD
Cochlicellidae
Monilearia woodwardia
Yes
LC
Yes*
LC
Cochlicellidae
Obelus despreauxii
Yes
VU
D2
Yes*
VU
D2
Cochlicellidae
Obelus discogranulatus
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Cochlicellidae
Obelus mirandae
Yes
LC
Yes*
LC
Cochlicellidae
Obelus moderatus
Yes
LC
Yes*
LC
Cochlicellidae
Obelus moratus
Yes
VU
D2
Yes*
VU
D2
Cochlicellidae
Ripkeniella petrophila
Yes
NT
Yes*
NT
Cochlicopidae
Azeca goodalli
Yes
LC
Yes
LC
Cochlicopidae
Cochlicopa lubrica
No
LC
No
LC
Cochlicopidae
Cochlicopa lubricella
No
LC
No
LC
Cochlicopidae
Cochlicopa nitens
No
LC
No
LC
Cochlicopidae
Cryptazeca elongata
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Cochlicopidae
Cryptazeca monodonta
Yes
EN
B2ab(iii)
Yes
EN
B2ab(iii)
Cochlicopidae
Cryptazeca spelaea
Yes
NT
Yes
NT
Cochlicopidae
Cryptazeca subcylindrica
Yes
LC
Yes
LC
Cochlicopidae
Hypnophila bisacchii
Yes
NT
Yes
NT
Cochlicopidae
Hypnophila boissii
Yes
LC
No
LC
Cochlicopidae
Hypnophila cyclothyra
Yes
LC
Yes
LC
Cochlicopidae
Hypnophila cylindrica
Yes
DD
Yes
DD
Cochlicopidae
Hypnophila dohrni
Yes
LC
Yes
LC
25
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Cochlicopidae
Hypnophila emiliana
Yes
VU
D2
Yes
VU
D2
Cochlicopidae
Hypnophila girottii
Yes
VU
D2
Yes
VU
D2
Cochlicopidae
Hypnophila incerta
Yes
NT
Yes
NT
Cochlicopidae
Hypnophila malagana
No
NT
No
NT
Cochlicopidae
Hypnophila polita
Yes
LC
No
LC
Cochlicopidae
Hypnophila pupaeformis
Yes
LC
No
LC
Cochlicopidae
Hypnophila remyi
Yes
LC
Yes
LC
Cochlicopidae
Hypnophila zacynthia
Yes
LC
Yes
LC
Craspedopomatidae
Craspedopoma costata
Yes LC Yes LC
Craspedopomatidae
Craspedopoma hespericum
Yes
LC
Yes
LC
Cyclophoridae
Pholeoteras euthrix
Yes
LC
No
LC
Diplommatinidae
Cochlostoma achaicum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma acutum
Yes
VU
D2
Yes*
VU
D2
Diplommatinidae
Cochlostoma adamii
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma affine
Yes
VU
D2
Yes*
VU
D2
Diplommatinidae
Cochlostoma alleryanum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma anomphale
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma auritum
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma braueri
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma canestrinii
Yes
VU
D2
Yes*
VU
D2
Diplommatinidae
Cochlostoma cinerascens
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma cretense
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma crosseanum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma dalmatinum
Yes
VU
D2
No*
NA
Diplommatinidae
Cochlostoma elegans
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma erika
Yes
VU
D2
No*
NA
Diplommatinidae
Cochlostoma euboicum
Yes
NT
Yes*
NT
Diplommatinidae
Cochlostoma fuchsi
Yes
VU
D2
No*
NA
Diplommatinidae
Cochlostoma georgi
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma gracile
Yes
LC
No
LC
Diplommatinidae
Cochlostoma hellenicum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma henricae
Yes
LC
Yes*
LC
26
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Diplommatinidae
Cochlostoma hoyeri
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma intermedium
Yes
DD
Yes*
DD
Diplommatinidae
Cochlostoma kleciaki
Yes
DD
No*
DD
Diplommatinidae
Cochlostoma macei
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma montanum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma mostarensis
Yes
NT
No*
NA
Diplommatinidae
Cochlostoma nanum
Yes
LC
No
LC
Diplommatinidae
Cochlostoma pageti
Yes
NT
Yes*
NT
Diplommatinidae
Cochlostoma paladilhianum
Yes VU D2 Yes* VU D2
Diplommatinidae
Cochlostoma parnonis
Yes
NT
Yes*
NT
Diplommatinidae
Cochlostoma patulum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma philippianum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma pinteri
Yes
NT
No*
NA
Diplommatinidae
Cochlostoma porroi
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma roseoli
Yes
LC
No*
NA
Diplommatinidae
Cochlostoma sardoum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma scalarinum
Yes
LC
No
LC
Diplommatinidae
Cochlostoma septemspirale
Yes
LC
No
LC
Diplommatinidae
Cochlostoma simrothi
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma stossichi
Yes
NT
No*
NA
Diplommatinidae
Cochlostoma subalpinum
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma tergestinum
Yes
LC
No
LC
Diplommatinidae
Cochlostoma tessellatum
Yes
LC
No
LC
Diplommatinidae
Cochlostoma villae
Yes
LC
Yes*
LC
Diplommatinidae
Cochlostoma waldemari
Yes
LC
No
LC
Diplommatinidae
Cochlostoma westerlundi
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella aprica
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella asturica
Yes
NT
Yes*
NT
Diplommatinidae
Obscurella bicostulata
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella conica
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella gigas
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella hidalgoi
Yes
LC
Yes*
LC
27
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Diplommatinidae
Obscurella martorelli
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella nouleti
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella oscitans
Yes
LC
Yes*
LC
Diplommatinidae
Obscurella partioti
Yes
LC
Yes*
LC
Diplommatinidae
Striolata striolata
Yes
LC
Yes*
LC
Discidae
Atlantica anagaensis
Yes
VU
D2
Yes
VU
D2
Discidae
Atlantica calathoides
Yes
CR
B2ab(iii,v)
Yes
CR
B2ab(iii,v)
Discidae
Atlantica engonata
Yes
CR(PE)
D
Yes
CR(PE)
D
Discidae
Atlantica ganoda
Yes LC Yes LC
Discidae
Atlantica gomerensis
Yes
CR(PE)
D
Yes
CR(PE)
D
Discidae
Atlantica guerinianus
Yes
CR(PE)
B1ab(iii,v)
Yes
CR(PE)
B1ab(iii,v)
Discidae
Atlantica kompsa
Yes
NT
Yes
NT
Discidae
Atlantica laurisilvae
Yes
VU
D2
Yes
VU
D2
Discidae
Atlantica putrescens
Yes
CR(PE)
D
Yes
CR(PE)
D1
Discidae
Atlantica retexta
Yes
DD
Yes
DD
Discidae
Atlantica rupivaga
Yes
VU
D2
Yes
VU
D2
Discidae
Atlantica saproxylophaga
Yes
LC
Yes
LC
Discidae
Atlantica scutula
Yes
LC
Yes
LC
Discidae
Atlantica textilis
Yes
DD
Yes
DD
Discidae
Discus perspectivus
Yes
LC
No
LC
Discidae
Discus rotundatus
Yes
LC
No
LC
Discidae
Discus ruderatus
No
LC
No
LC
Discidae
Keraea deflorata
Yes
DD
Yes
DD
Ellobiidae
Carychium biondii
Yes
LC
Yes
LC
Ellobiidae
Carychium hellenicum
Yes
LC
No
LC
Ellobiidae
Carychium ibazoricum
Yes
LC
Yes
LC
Ellobiidae
Carychium mariae
Yes
LC
Yes
LC
Ellobiidae
Carychium minimum
No
LC
No
LC
Ellobiidae
Carychium paganettii
Yes
LC
Yes
LC
Ellobiidae
Carychium tridentatum
No
LC
No
LC
Ellobiidae
Leucophytia bidentata
Yes
LC
Yes
LC
Ellobiidae
Myosotella denticulata
Yes
LC
Yes
LC
28
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Ellobiidae
Myosotella myosotis
Yes
LC
Yes
LC
Ellobiidae
Ovatella aequalis
Yes
LC
Yes
LC
Ellobiidae
Ovatella firminii
No
LC
No
LC
Ellobiidae
Ovatella vulcani
Yes
LC
Yes
LC
Ellobiidae
Pedipes pedipes
No
LC
No
LC
Ellobiidae
Pseudomelampus exiguus
No
LC
No
LC
Ellobiidae
Zospeum allegrettii
Yes
VU
D2
Yes
VU
D2
Ellobiidae
Zospeum alpestre
Yes
LC
Yes
LC
Ellobiidae
Zospeum amoenum
Yes LC No LC
Ellobiidae
Zospeum bellesi
Yes
LC
Yes
LC
Ellobiidae
Zospeum biscaiense
Yes
NT
Yes
NT
Ellobiidae
Zospeum cariadeghense
Yes
DD
Yes
DD
Ellobiidae
Zospeum exiguum
Yes
VU
D2
Yes
VU
D2
Ellobiidae
Zospeum frauenfeldi
Yes
LC
Yes
LC
Ellobiidae
Zospeum galvagnii
Yes
DD
Yes
DD
Ellobiidae
Zospeum globosum
Yes
LC
Yes
LC
Ellobiidae
Zospeum isselianum
Yes
LC
No
LC
Ellobiidae
Zospeum kupitzense
Yes
NT
Yes
NT
Ellobiidae
Zospeum kusceri
Yes
LC
Yes
LC
Ellobiidae
Zospeum lautum
Yes
LC
Yes
LC
Ellobiidae
Zospeum obesum
Yes
LC
Yes
LC
Ellobiidae
Zospeum pretneri
Yes
LC
Yes
LC
Ellobiidae
Zospeum schaufussi
Yes
DD
Yes
DD
Ellobiidae
Zospeum spelaeum
Yes
LC
Yes
LC
Ellobiidae
Zospeum suarezi
Yes
NT
Yes
NT
Ellobiidae
Zospeum subobesum
Yes
LC
No
LC
Ellobiidae
Zospeum tholussum
Yes
NT
Yes
NT
Ellobiidae
Zospeum trebicianum
Yes
NT
Yes
NT
Ellobiidae
Zospeum turriculatum
Yes
DD
Yes
DD
Ellobiidae
Zospeum vasconicum
Yes
NT
Yes
NT
Ellobiidae
Zospeum zaldivarae
Yes
NT
Yes
NT
Elonidae
Elona quimperiana
Yes
LC
Yes
LC
29
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Elonidae
Norelona pyrenaica
Yes
LC
No
LC
Enidae
Brephulopsis bidens
Yes
LC
No*
NA
Enidae
Brephulopsis cylindrica
Yes
LC
No*
NA
Enidae
Brephulopsis subulata
Yes
EN
B1ab(iii)+2ab(iii)
No
NA
Enidae
Buliminus carneus
Yes
LC
No
LC
Enidae
Chondrula beieri
Yes
LC
Yes*
LC
Enidae
Chondrula bergeri
Yes
DD
Yes*
DD
Enidae
Chondrula consentanea
Yes
DD
No*
NA
Enidae
Chondrula lugorensis
Yes NT No* NA
Enidae
Chondrula macedonica
Yes
LC
No
LC
Enidae
Chondrula microtragus
Yes
LC
No
LC
Enidae
Chondrula munita
Yes
DD
Yes*
DD
Enidae
Chondrula peloponnesica
Yes
LC
Yes*
LC
Enidae
Chondrula pindica
Yes
LC
Yes*
LC
Enidae
Chondrula quinquedentata
Yes
LC
No
LC
Enidae
Chondrula tricuspidata
Yes
LC
No
LC
Enidae
Chondrula tridens
No
NT
No*
NT
Enidae
Chondrula werneri
Yes
LC
No
VU
D2
Enidae
Chondrus tournefortianus
Yes
LC
No
LC
Enidae
Chondrus zebra
Yes
LC
No
LC
Enidae
Ena concolor
Yes
LC
No*
NA
Enidae
Ena elongata
Yes
DD
Yes*
DD
Enidae
Ena montana
Yes
LC
No
LC
Enidae
Ena monticola
Yes
LC
Yes*
LC
Enidae
Ena subtilis
Yes
LC
No
LC
Enidae
Eubrephulus bicallosa
Yes
LC
No
LC
Enidae
Euchondrus ledereri
Yes
LC
No
LC
Enidae
Euchondrus limbodentatus
Yes
NT
Yes*
NT
Enidae
Euchondrus nucifragus
Yes
LC
Yes*
LC
Enidae
Euchondrus parreyssi
Yes
LC
Yes*
LC
Enidae
Jaminia loewii
Yes
LC
No
LC
Enidae
Jaminia quadridens
Yes
LC
No
LC
30
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Enidae
Jaminia thiesseanus
Yes
LC
Yes*
LC
Enidae
Mastus abundans
Yes
LC
Yes*
LC
Enidae
Mastus alpicolus
Yes
LC
Yes*
LC
Enidae
Mastus amenazada
Yes
VU
D2
Yes*
VU
D2
Enidae
Mastus athensis
Yes
LC
No
LC
Enidae
Mastus bielzi
Yes
LC
No
LC
Enidae
Mastus butoti
Yes
LC
Yes*
LC
Enidae
Mastus carneolus
Yes
LC
No
LC
Enidae
Mastus claudia
Yes VU B1ab(iii)+2ab(iii); D2 Yes* VU B1ab(iii)+2ab(iii); D2
Enidae
Mastus cretensis
Yes
LC
Yes*
LC
Enidae
Mastus dirphicus
Yes
LC
Yes*
LC
Enidae
Mastus emarginatus
Yes
NT
Yes*
NT
Enidae
Mastus etuberculatus
Yes
LC
No
LC
Enidae
Mastus gittenbergeri
Yes
LC
Yes*
LC
Enidae
Mastus giuricus
Yes
LC
Yes*
LC
Enidae
Mastus grandis
Yes
LC
No
LC
Enidae
Mastus hemmeni
Yes
LC
Yes*
LC
Enidae
Mastus ierapetrana
Yes
LC
Yes*
LC
Enidae
Mastus itanosensis
Yes
LC
Yes*
LC
Enidae
Mastus olivaceus
Yes
LC
Yes*
LC
Enidae
Mastus procax
Yes
LC
Yes*
LC
Enidae
Mastus pupa
No
LC
No
LC
Enidae
Mastus pusio
Yes
LC
Yes*
LC
Enidae
Mastus riedeli
Yes
LC
Yes*
LC
Enidae
Mastus rossmaessleri
Yes
LC
No
LC
Enidae
Mastus sitiensis
Yes
LC
Yes*
LC
Enidae
Mastus sphakiota
Yes
LC
Yes*
LC
Enidae
Mastus subaii
Yes
LC
Yes*
LC
Enidae
Mastus transsylvanicus
Yes
LC
Yes*
LC
Enidae
Mastus turgidus
Yes
LC
Yes*
LC
Enidae
Mastus venerabilis
Yes
LC
Yes*
LC
Enidae
Mastus violacea
Yes
LC
Yes*
LC
31
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Enidae
Meijeriella canaliculata
No
LC
No
LC
Enidae
Merdigera obscura
No
LC
No
LC
Enidae
Multidentula lamellifera
Yes
LC
No
LC
Enidae
Multidentula ovularis
Yes
LC
No
LC
Enidae
Multidentula squalina
No
DD
No
NA
Enidae
Multidentula stylus
Yes
LC
Yes*
LC
Enidae
Napaeopsis cefalonica
Yes
LC
No
LC
Enidae
Napaeopsis mennoi
Yes
LC
Yes*
LC
Enidae
Napaeopsis merditanus
Yes LC No* NA
Enidae
Napaeopsis minima
Yes
LC
Yes*
LC
Enidae
Napaeopsis ossica
Yes
LC
Yes*
LC
Enidae
Napaeus alabastrinus
Yes
LC
Yes*
LC
Enidae
Napaeus alucensis
Yes
LC
Yes
LC
Enidae
Napaeus anaga
Yes
DD
Yes*
DD
Enidae
Napaeus arinagaensis
Yes
CR(PE)
B2ab(iii)
Yes
CR(PE)
B2ab(iii)
Enidae
Napaeus atlanticus
Yes
LC
Yes*
LC
Enidae
Napaeus avaloensis
Yes
NT
Yes*
NT
Enidae
Napaeus badiosus
Yes
LC
Yes*
LC
Enidae
Napaeus baeticatus
Yes
LC
Yes*
LC
Enidae
Napaeus bajamarensis
Yes
LC
Yes*
LC
Enidae
Napaeus barquini
Yes
LC
Yes*
LC
Enidae
Napaeus bechi
Yes
LC
Yes*
LC
Enidae
Napaeus beguirae
Yes
NT
Yes*
NT
Enidae
Napaeus bertheloti
Yes
LC
Yes*
LC
Enidae
Napaeus boucheti
Yes
VU
D2
Yes*
VU
D2
Enidae
Napaeus chrysaloides
Yes
DD
Yes*
DD
Enidae
Napaeus consecoanus
Yes
NT
Yes*
NT
Enidae
Napaeus delibutus
Yes
LC
Yes*
LC
Enidae
Napaeus delicatus
Yes
NT
Yes
NT
Enidae
Napaeus doliolum
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Enidae
Napaeus dolorosae
Yes
CR
D
Yes
NT
Enidae
Napaeus elegans
Yes
VU
B2ab(iii)
Yes*
VU
B2ab(iii)
32
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Enidae
Napaeus encaustus
Yes
LC
Yes*
LC
Enidae
Napaeus esbeltus
Yes
VU
B2ab(iii)
Yes*
VU
B2ab(iii)
Enidae
Napaeus estherae
Yes
LC
Yes
LC
Enidae
Napaeus exilis
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Enidae
Napaeus flavoterminatus
Yes
DD
Yes*
DD
Enidae
Napaeus gomerensis
Yes
LC
Yes
LC
Enidae
Napaeus grohi
Yes
DD
Yes
DD
Enidae
Napaeus gruereanus
Yes
DD
Yes*
DD
Enidae
Napaeus hartungi
Yes LC Yes* LC
Enidae
Napaeus helvolus
Yes
LC
Yes*
LC
Enidae
Napaeus huttereri
Yes
NT
Yes*
NT
Enidae
Napaeus indifferens
Yes
DD
Yes*
DD
Enidae
Napaeus inflatiusculus
Yes
DD
Yes*
DD
Enidae
Napaeus interpunctatus
Yes
DD
Yes*
DD
Enidae
Napaeus isletae
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Enidae
Napaeus josei
Yes
LC
Yes
LC
Enidae
Napaeus lichenicola
Yes
VU
D2
Yes*
VU
D2
Enidae
Napaeus lowei
Yes
DD
Yes*
DD
Enidae
Napaeus maculatus
Yes
NT
Yes*
NT
Enidae
Napaeus maffioteanus
Yes
DD
Yes*
DD
Enidae
Napaeus magnus
Yes
LC
Yes
LC
Enidae
Napaeus minimus
Yes
VU
D2
Yes
VU
D2
Enidae
Napaeus moquinianus
Yes
LC
Yes*
LC
Enidae
Napaeus moroi
Yes
NT
Yes
NT
Enidae
Napaeus myosotis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Enidae
Napaeus nanodes
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Enidae
Napaeus obesatus
Yes
LC
Yes*
LC
Enidae
Napaeus ocellatus
Yes
DD
Yes*
DD
Enidae
Napaeus orientalis
Yes
NT
Yes*
NT
Enidae
Napaeus ornamentatus
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Enidae
Napaeus osoriensis
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Enidae
Napaeus palmaensis
Yes
DD
Yes*
DD
33
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Enidae
Napaeus procerus
Yes
NT
Yes*
NT
Enidae
Napaeus propinquus
Yes
DD
Yes*
DD
Enidae
Napaeus pruninus
Yes
LC
Yes*
LC
Enidae
Napaeus pygmaeus
Yes
LC
Yes*
LC
Enidae
Napaeus roccellicola
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Enidae
Napaeus rufobrunneus
Yes
LC
Yes*
LC
Enidae
Napaeus rupicola
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Enidae
Napaeus savinosa
Yes
DD
Yes*
DD
Enidae
Napaeus servus
Yes DD Yes* DD
Enidae
Napaeus severus
Yes
DD
Yes*
DD
Enidae
Napaeus subgracilior
Yes
DD
Yes*
DD
Enidae
Napaeus subsimplex
Yes
DD
Yes*
DD
Enidae
Napaeus tabidus
Yes
LC
Yes*
LC
Enidae
Napaeus tafadaensis
Yes
NT
Yes*
NT
Enidae
Napaeus tagamichensis
Yes
VU
D2
Yes*
VU
D2
Enidae
Napaeus taguluchensis
Yes
VU
D2
Yes*
VU
D2
Enidae
Napaeus tenoensis
Yes
LC
Yes*
LC
Enidae
Napaeus teobaldoi
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Enidae
Napaeus texturatus
Yes
DD
Yes*
DD
Enidae
Napaeus torilensis
Yes
NT
Yes
NT
Enidae
Napaeus tremulans
Yes
LC
Yes*
LC
Enidae
Napaeus validoi
Yes
NT
Yes
NT
Enidae
Napaeus variatus
Yes
LC
Yes*
LC
Enidae
Napaeus venegueraensis
Yes
LC
Yes
LC
Enidae
Napaeus voggenreiteri
Yes
NT
Yes*
NT
Enidae
Napaeus vulgaris
Yes
LC
Yes*
LC
Enidae
Paramastus cyprius
Yes
LC
Yes*
LC
Enidae
Peristoma merduenianum
Yes
VU
B1ab(iii)+2ab(iii)
No
NA
Enidae
Peristoma rupestre
Yes
LC
No
LC
Enidae
Pseudochondrula seductilis
No
VU
A2b
No*
VU
A2ab
Enidae
Rhabdoena cosensis
No
LC
No
LC
Enidae
Rhabdoena mirifica
Yes
NT
Yes*
NT
34
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Enidae
Rhabdoena stokesi
Yes
NT
Yes*
NT
Enidae
Rhabdoena zasiensis
Yes
NT
Yes*
NT
Enidae
Thoanteus ferrarii
Yes
LC
No*
NA
Enidae
Thoanteus gibber
Yes
LC
No*
NA
Enidae
Turanena carpathia
Yes
NT
Yes*
NT
Enidae
Turanena hemmeni
No
NT
No*
NT
Enidae
Turanena katerinae
Yes
NT
Yes*
NT
Enidae
Zebrina detrita
No
LC
No
LC
Enidae
Zebrina fasciolata
Yes LC No LC
Enidae
Zebrina kindermanni
No
LC
No
LC
Enidae
Zebrina varnensis
Yes
LC
Yes*
LC
Euconulidae
Euconulus callopisticus
Yes
DD
Yes
DD
Euconulidae
Euconulus fulvus
No
LC
No
LC
Euconulidae
Euconulus praticola
Yes
DD
No
DD
Euconulidae
Euconulus trochiformis
Yes
LC
No
LC
Ferussaciidae
Amphorella cimensis
Yes
NT
Yes
NT
Ferussaciidae
Amphorella gracilis
Yes
LC
Yes
LC
Ferussaciidae
Amphorella hypselia
Yes
VU
D2
Yes
VU
D2
Ferussaciidae
Amphorella intermedia
Yes
DD
Yes
DD
Ferussaciidae
Amphorella iridescens
Yes
VU
D2
Yes
VU
D2
Ferussaciidae
Amphorella melampoides
Yes
LC
Yes
LC
Ferussaciidae
Amphorella mitriformis
Yes
LC
Yes
LC
Ferussaciidae
Amphorella oryza
Yes
LC
Yes
LC
Ferussaciidae
Amphorella tornatellina
Yes
LC
Yes
LC
Ferussaciidae
Amphorella triticea
Yes
LC
Yes
LC
Ferussaciidae
Amphorella tuberculata
Yes
EN
B1ab(iv,v)+2ab(iv,v)
Yes
EN
B1ab(iv,v)+2ab(iv,v)
Ferussaciidae
Calaxis cypria
Yes
LC
Yes
LC
Ferussaciidae
Cecilioides acicula
No
LC
No
LC
Ferussaciidae
Cecilioides actoniana
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides barbozae
Yes
LC
Yes
LC
Ferussaciidae
Cecilioides bourguignatiana
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides clessini
Yes
NT
Yes
NT
35
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Ferussaciidae
Cecilioides connollyi
Yes
VU
D2
Yes
VU
D2
Ferussaciidae
Cecilioides dalmatina
Yes
DD
No
DD
Ferussaciidae
Cecilioides eucharista
No
NA
No
NA
Ferussaciidae
Cecilioides eulima
Yes
CR
B1ab(iii)+2ab(iii); D
Yes
CR
B1ab(iii)+2ab(iii); D
Ferussaciidae
Cecilioides jeskalovicensis
Yes
CR
B2ab(iii)
Yes
CR
B2ab(iii)
Ferussaciidae
Cecilioides petitiana
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides poupillieri
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides raddei
No
DD
No
NA
Ferussaciidae
Cecilioides raphidia
No LC No LC
Ferussaciidae
Cecilioides rizzeana
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides spelaea
Yes
DD
No
DD
Ferussaciidae
Cecilioides stephaniana
Yes
DD
Yes
DD
Ferussaciidae
Cecilioides tumulorum
No
LC
No
LC
Ferussaciidae
Cecilioides veneta
No
LC
No
LC
Ferussaciidae
Coilostele acus
Yes
DD
Yes
DD
Ferussaciidae
Cylichnidia ovuliformis
Yes
VU
D2
Yes
VU
D2
Ferussaciidae
Ferussacia attenuata
Yes
LC
Yes
LC
Ferussaciidae
Ferussacia carnea
No
NA
No
NA
Ferussaciidae
Ferussacia folliculum
Yes
LC
Yes
LC
Ferussaciidae
Ferussacia fritschi
Yes
LC
Yes
LC
Ferussaciidae
Ferussacia lanzarotensis
Yes
LC
Yes
LC
Ferussaciidae
Ferussacia submajor
Yes
NT
Yes
NT
Ferussaciidae
Ferussacia tumidula
Yes
DD
Yes
DD
Ferussaciidae
Ferussacia valida
Yes
NT
Yes
NT
Ferussaciidae
Ferussacia vitrea
Yes
LC
Yes
LC
Ferussaciidae
Hohenwartiana aradasiana
Yes
DD
Yes
DD
Ferussaciidae
Hohenwartiana disparata
Yes
LC
Yes
LC
Ferussaciidae
Hohenwartiana hohenwarti
Yes
LC
Yes
LC
Ferussaciidae
Pyrgella leacociana
Yes
LC
Yes
LC
Ferussaciidae
Sculptiferussacia clausiliaeformis
Yes
NT
Yes
NT
Gastrodontidae
Janulus bifrons
Yes
LC
Yes
LC
Gastrodontidae
Janulus pompylius
Yes
CR(PE)
D
Yes
CR(PE)
D
36
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Gastrodontidae
Janulus stephanophorus
Yes
VU
B2ab(iii)
Yes
VU
B2ab(iii)
Gastrodontidae
Janulus traviesus
Yes
NT
Yes
NT
Gastrodontidae
Striatura milium
No
DD
No
DD
Gastrodontidae
Zonitoides arboreus
No
NA
No
NA
Gastrodontidae
Zonitoides excavatus
Yes
LC
Yes
LC
Gastrodontidae
Zonitoides jaccetanicus
Yes
DD
Yes
DD
Gastrodontidae
Zonitoides nitidus
No
LC
No
LC
Helicidae
Allognathus graellsianus
Yes
LC
Yes*
LC
Helicidae
Allognathus hispanicus
Yes LC Yes* LC
Helicidae
Allognathus pyrenaicus
Yes
NT
Yes*
NT
Helicidae
Arianta aethiops
Yes
LC
No
LC
Helicidae
Arianta arbustorum
Yes
LC
No
LC
Helicidae
Arianta chamaeleon
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Helicidae
Arianta hessei
Yes
NT
Yes*
NT
Helicidae
Arianta schmidtii
Yes
NT
Yes*
NT
Helicidae
Arianta stenzii
Yes
NT
Yes*
NT
Helicidae
Arianta xatartii
Yes
VU
D2
Yes*
VU
D2
Helicidae
Assyriella bellardii
Yes
DD
Yes*
DD
Helicidae
Assyriella rechingeri
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Helicidae
Cantareus apertus
Yes
LC
No
LC
Helicidae
Cattania ardica
Yes
NT
Yes*
NT
Helicidae
Cattania haberhaueri
Yes
LC
No
LC
Helicidae
Cattania ista
Yes
DD
Yes
DD
Helicidae
Cattania kattingeri
Yes
LC
Yes*
LC
Helicidae
Cattania maranajensis
Yes
NT
No*
NA
Helicidae
Cattania petrovici
Yes
NT
No*
NA
Helicidae
Cattania rumelica
Yes
LC
No
LC
Helicidae
Cattania trizona
Yes
LC
No
LC
Helicidae
Causa holosericea
Yes
LC
No
LC
Helicidae
Cepaea hortensis
Yes
LC
No
LC
Helicidae
Cepaea nemoralis
Yes
LC
No
LC
Helicidae
Cepaea sylvatica
Yes
LC
No
LC
37
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Cepaea vindobonensis
Yes
LC
No
LC
Helicidae
Chilostoma achates
Yes
NT
Yes*
NT
Helicidae
Chilostoma acrotricha
Yes
NT
Yes*
NT
Helicidae
Chilostoma adelozona
Yes
VU
B1ab(iv)
No*
VU
B1ab(iv)
Helicidae
Chilostoma albanograeca
Yes
LC
No
LC
Helicidae
Chilostoma ambrosi
Yes
NT
Yes*
NT
Helicidae
Chilostoma amorgia
Yes
LC
Yes*
LC
Helicidae
Chilostoma apfelbecki
Yes
LC
No*
NA
Helicidae
Chilostoma arcadica
Yes LC Yes* LC
Helicidae
Chilostoma argentellei
Yes
LC
Yes*
LC
Helicidae
Chilostoma bacchica
Yes
LC
Yes*
LC
Helicidae
Chilostoma brenskei
Yes
LC
Yes*
LC
Helicidae
Chilostoma byshekensis
Yes
LC
No
LC
Helicidae
Chilostoma choristochila
Yes
LC
Yes*
LC
Helicidae
Chilostoma cingulatum
Yes
DD
No*
DD
Helicidae
Chilostoma comythophora
Yes
LC
Yes*
LC
Helicidae
Chilostoma conemenosi
Yes
LC
Yes*
LC
Helicidae
Chilostoma crinita
Yes
NT
No*
NA
Helicidae
Chilostoma crombezi
Yes
CR
B1ab(iii)
Yes*
CR
B1ab(iii)
Helicidae
Chilostoma cyclolabris
Yes
LC
Yes*
LC
Helicidae
Chilostoma denudata
Yes
NT
No*
NA
Helicidae
Chilostoma desmoulinsii
Yes
LC
No
LC
Helicidae
Chilostoma dochii
Yes
LC
No*
NA
Helicidae
Chilostoma dunjana
Yes
LC
No*
NA
Helicidae
Chilostoma edlaueri
Yes
DD
No*
NA
Helicidae
Chilostoma eliaca
Yes
LC
Yes*
LC
Helicidae
Chilostoma euboeae
Yes
NT
Yes*
NT
Helicidae
Chilostoma faueri
Yes
LC
Yes*
LC
Helicidae
Chilostoma fontenillii
Yes
LC
Yes*
LC
Helicidae
Chilostoma frigidum
Yes
LC
Yes*
LC
Helicidae
Chilostoma fuchsi
Yes
NT
No*
NA
Helicidae
Chilostoma fuchsiana
Yes
LC
Yes*
LC
38
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Chilostoma glaciale
Yes
LC
No
LC
Helicidae
Chilostoma harpya
Yes
CR
B1ab(ii,iii,v)+2ab(ii,iii,v)
No*
NA
Helicidae
Chilostoma heldreichi
Yes
LC
Yes*
LC
Helicidae
Chilostoma hemonica
Yes
LC
Yes*
LC
Helicidae
Chilostoma hirta
Yes
LC
No*
NA
Helicidae
Chilostoma hoffmanni
Yes
LC
No*
NA
Helicidae
Chilostoma hymetti
Yes
LC
Yes*
LC
Helicidae
Chilostoma insolita
Yes
LC
No*
NA
Helicidae
Chilostoma intermedium
Yes LC No LC
Helicidae
Chilostoma istriana
Yes
DD
No*
NA
Helicidae
Chilostoma kleciachi
Yes
LC
No*
NA
Helicidae
Chilostoma kollari
Yes
LC
No*
NA
Helicidae
Chilostoma krueperi
Yes
LC
Yes*
LC
Helicidae
Chilostoma lefeburiana
Yes
LC
No
LC
Helicidae
Chilostoma melpomene
Yes
LC
Yes*
LC
Helicidae
Chilostoma millieri
Yes
LC
Yes*
LC
Helicidae
Chilostoma moellendorffi
Yes
LC
No*
NA
Helicidae
Chilostoma nympha
Yes
LC
Yes*
LC
Helicidae
Chilostoma pelia
Yes
LC
Yes*
LC
Helicidae
Chilostoma pentheri
Yes
NT
No*
NA
Helicidae
Chilostoma phocaea
Yes
LC
No
LC
Helicidae
Chilostoma pieperi
Yes
LC
Yes*
LC
Helicidae
Chilostoma planospira
Yes
DD
Yes*
DD
Helicidae
Chilostoma polinskii
Yes
LC
Yes*
LC
Helicidae
Chilostoma polyhymnia
Yes
LC
Yes*
LC
Helicidae
Chilostoma posthuma
Yes
LC
Yes*
LC
Helicidae
Chilostoma pouzolzi
Yes
LC
No*
NA
Helicidae
Chilostoma serbica
Yes
LC
No*
NA
Helicidae
Chilostoma setigera
Yes
LC
No*
NA
Helicidae
Chilostoma setosa
Yes
LC
No*
NA
Helicidae
Chilostoma sphaeriostoma
Yes
LC
Yes*
LC
Helicidae
Chilostoma squamatinum
Yes
LC
No
LC
39
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Chilostoma stenomphala
Yes
LC
No*
NA
Helicidae
Chilostoma subaii
Yes
LC
Yes*
LC
Helicidae
Chilostoma subzonata
Yes
LC
Yes*
LC
Helicidae
Chilostoma sztolcmani
Yes
NT
Yes*
NT
Helicidae
Chilostoma valkanovi
Yes
LC
Yes*
LC
Helicidae
Chilostoma zebiana
Yes
NT
No*
NA
Helicidae
Chilostoma ziegleri
Yes
NT
Yes*
NT
Helicidae
Chilostoma zonatum
Yes
LC
No
LC
Helicidae
Codringtonia codringtonii
Yes EN B1ab(i,ii,iii,iv) Yes* EN B1ab(i,ii,iii,iv)
Helicidae
Codringtonia elisabethae
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia eucineta
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia gittenbergeri
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia helenae
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia intusplicata
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia neocrassa
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
No*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Codringtonia parnassia
Yes
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Yes*
VU
B1ab(i,ii,iv)+2ab(i,ii,iv)
Helicidae
Cornu aspersum
No
LC
No
LC
Helicidae
Cornu mazzullii
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Helicidae
Cylindrus obtusus
Yes
LC
Yes*
LC
Helicidae
Drobacia banatica
Yes
DD
No*
DD
Helicidae
Drobacia maeotica
Yes
DD
Yes*
DD
Helicidae
Eobania vermiculata
No
LC
No
LC
Helicidae
Faustina cingulella
Yes
LC
Yes*
LC
Helicidae
Faustina faustina
Yes
LC
No
LC
Helicidae
Faustina illyrica
Yes
LC
No
LC
Helicidae
Helicigona lapicida
Yes
LC
No
LC
Helicidae
Helix albescens
Yes
LC
No
LC
Helicidae
Helix anatolica
Yes
DD
No*
DD
Helicidae
Helix asemnis
Yes
LC
No
LC
Helicidae
Helix borealis
Yes
DD
Yes*
DD
Helicidae
Helix cincta
Yes
LC
No
LC
Helicidae
Helix delpretiana
Yes
DD
Yes*
DD
40
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Helix dormitoris
Yes
LC
No*
NA
Helicidae
Helix figulina
Yes
LC
No
LC
Helicidae
Helix godetiana
Yes
EN
B2ab(i,ii,iii,iv,v)
Yes*
EN
B2ab(i,ii,iii,iv,v)
Helicidae
Helix ligata
Yes
DD
Yes*
DD
Helicidae
Helix lucorum
No
LC
No
LC
Helicidae
Helix lutescens
Yes
LC
No
LC
Helicidae
Helix melanostoma
No
LC
No
LC
Helicidae
Helix nucula
No
NT
No*
NT
Helicidae
Helix philibinensis
Yes LC No LC
Helicidae
Helix pomacella
Yes
LC
No
LC
Helicidae
Helix pomatia
Yes
LC
No
LC
Helicidae
Helix pronuba
No
LC
No
LC
Helicidae
Helix secernenda
Yes
LC
No
LC
Helicidae
Helix straminea
Yes
LC
Yes
LC
Helicidae
Helix thessalica
Yes
LC
No
LC
Helicidae
Helix valentini
Yes
EN
B1ab(i,ii,iii)+2ab(i,ii,iii)
No*
EN
B1ab(i,ii,iii)+2ab(i,ii,iii)
Helicidae
Helix vladica
Yes
LC
No*
NA
Helicidae
Hemicycla berkeleii
Yes
DD
Yes*
DD
Helicidae
Hemicycla bethencourtiana
Yes
NT
Yes*
NT
Helicidae
Hemicycla bidentalis
Yes
LC
Yes*
LC
Helicidae
Hemicycla consobrina
Yes
LC
Yes*
LC
Helicidae
Hemicycla diegoi
Yes
NT
Yes
NT
Helicidae
Hemicycla distensa
Yes
DD
Yes*
DD
Helicidae
Hemicycla efferata
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Helicidae
Hemicycla ethelema
Yes
DD
Yes*
DD
Helicidae
Hemicycla eurythyra
Yes
VU
D2
Yes*
VU
D2
Helicidae
Hemicycla flavistoma
Yes
LC
Yes*
LC
Helicidae
Hemicycla fritschi
Yes
DD
Yes*
DD
Helicidae
Hemicycla fuenterroquensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Helicidae
Hemicycla fulgida
Yes
LC
Yes*
LC
Helicidae
Hemicycla gaudryi
Yes
DD
Yes*
DD
Helicidae
Hemicycla glasiana
Yes
LC
Yes*
LC
41
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Hemicycla glyceia
Yes
NT
Yes*
NT
Helicidae
Hemicycla gomerensis
Yes
DD
Yes*
DD
Helicidae
Hemicycla granomalleata
Yes
DD
Yes*
DD
Helicidae
Hemicycla guamartemes
Yes
DD
Yes*
DD
Helicidae
Hemicycla hedybia
Yes
DD
Yes*
DD
Helicidae
Hemicycla incisogranulata
Yes
NT
Yes*
NT
Helicidae
Hemicycla inutilis
Yes
VU
B2ab(iii)
Yes*
VU
B2ab(iii)
Helicidae
Hemicycla invernicata
Yes
LC
Yes*
LC
Helicidae
Hemicycla laurijona
Yes NT Yes* NT
Helicidae
Hemicycla mascaensis
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Helicidae
Hemicycla maugeana
Yes
DD
Yes*
DD
Helicidae
Hemicycla melchori
Yes
LC
Yes*
LC
Helicidae
Hemicycla modesta
Yes
CR(PE)
D
Yes*
CR(PE)
D
Helicidae
Hemicycla paeteliana
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Helicidae
Hemicycla paivanopsis
Yes
DD
Yes*
DD
Helicidae
Hemicycla perraudierei
Yes
LC
Yes*
LC
Helicidae
Hemicycla planorbella
Yes
DD
Yes*
DD
Helicidae
Hemicycla plicaria
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Helicidae
Hemicycla pouchadan
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Helicidae
Hemicycla pouchet
Yes
VU
B2ab(ii)
Yes*
VU
B2ab(ii)
Helicidae
Hemicycla psathyra
Yes
DD
Yes*
DD
Helicidae
Hemicycla quadricincta
Yes
NT
Yes*
NT
Helicidae
Hemicycla saponacea
Yes
NT
Yes*
NT
Helicidae
Hemicycla sarcostoma
Yes
LC
Yes*
LC
Helicidae
Hemicycla saulcyi
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Helicidae
Hemicycla vermiplicata
Yes
DD
Yes*
DD
Helicidae
Iberus alonensis
Yes
NT
Yes*
NT
Helicidae
Iberus angustatus
Yes
LC
Yes*
LC
Helicidae
Iberus campesinus
Yes
VU
B1ab(iii)
Yes*
VU
B1ab(iii)
Helicidae
Iberus carthaginiensis
Yes
NT
Yes*
NT
Helicidae
Iberus gualtieranus
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes*
EN
B1ab(iii,v)+2ab(iii,v)
Helicidae
Iberus guiraoanus
Yes
LC
Yes*
LC
42
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Iberus marmoratus
Yes
LC
Yes*
LC
Helicidae
Iberus ortizi
Yes
VU
D2
Yes*
VU
D2
Helicidae
Idiomela subplicata
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Helicidae
Isognomostoma isognomostomos
Yes
LC
No
LC
Helicidae
Josephinella vikosensis
Yes
DD
Yes*
DD
Helicidae
Lampadia webbiana
Yes
EN
B1ab(ii,iii)+2ab(ii,iii)
Yes*
EN
B1ab(ii,iii)+2ab(ii,iii)
Helicidae
Levantina spiriplana
Yes
LC
No
LC
Helicidae
Liburnica grisea
Yes
NT
No*
NA
Helicidae
Liburnica kulmakana
Yes NT No* NA
Helicidae
Liburnica skipetarica
Yes
LC
No*
NA
Helicidae
Macularia niciensis
Yes
LC
Yes*
LC
Helicidae
Macularia saintivesi
Yes
VU
D2
Yes*
VU
D2
Helicidae
Marmorana fuscolabiata
Yes
DD
Yes*
DD
Helicidae
Marmorana globularis
Yes
LC
Yes*
LC
Helicidae
Marmorana muralis
Yes
LC
Yes*
LC
Helicidae
Marmorana nebrodensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Helicidae
Marmorana platychela
Yes
DD
Yes*
DD
Helicidae
Marmorana saxetana
Yes
NT
Yes*
NT
Helicidae
Marmorana scabriuscula
Yes
DD
Yes*
DD
Helicidae
Marmorana serpentina
Yes
LC
Yes*
LC
Helicidae
Marmorana signata
Yes
NT
Yes*
NT
Helicidae
Otala lactea
No
LC
No
LC
Helicidae
Otala punctata
No
LC
No
LC
Helicidae
Pseudotachea litturata
No
NT
No*
NT
Helicidae
Pseudotachea splendida
Yes
LC
No
LC
Helicidae
Rossmaessleria scherzeri
No
LC
No
LC
Helicidae
Superba reischuetzi
Yes
NT
Yes*
NT
Helicidae
Tacheocampylaea acropachia
Yes
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Yes*
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Helicidae
Tacheocampylaea carotii
Yes
VU
B1ab(iii,v)+2ab(iii,v)
Yes*
VU
B1ab(iii,v)+2ab(iii,v)
Helicidae
Tacheocampylaea cyrniaca
Yes
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Yes*
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Helicidae
Tacheocampylaea raspailii
Yes
VU
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Yes*
VU
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Helicidae
Tacheocampylaea romagnolii
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Yes*
CR
B1ab(iii,v)+2ab(iii,v)
43
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicidae
Tacheocampylaea tacheoides
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes*
EN
B1ab(iii,v)+2ab(iii,v)
Helicidae
Theba andalusica
Yes
DD
Yes*
DD
Helicidae
Theba arinagae
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Helicidae
Theba clausoinflata
Yes
NT
Yes*
NT
Helicidae
Theba geminata
Yes
DD
Yes*
DD
Helicidae
Theba grasseti
Yes
EN
B1ab(ii,iii)+2ab(ii,iii)
Yes*
EN
B1ab(ii,iii)+2ab(ii,iii)
Helicidae
Theba impugnata
Yes
VU
B1ab(iii)
Yes*
VU
B1ab(iii)
Helicidae
Theba macandrewiana
Yes
NT
Yes
NT
Helicidae
Theba pisana
No LC No LC
Helicidae
Theba subdentata
No
VU
D2
No*
VU
D2
Helicidae
Tyrrhenaria ceratina
Yes
CR
B1ab(i,ii,iii)+2ab(i,ii,iii)
Yes*
CR
B1ab(i,ii,iii)+2ab(i,ii,iii)
Helicidae
Tyrrheniberus ridens
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Helicidae
Tyrrheniberus sardonius
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Helicidae
Tyrrheniberus villicus
Yes
VU
D2
Yes*
VU
D2
Helicidae
Vidovicia coerulans
Yes
VU
A2b
No*
NA
Helicodiscidae
Helicodiscus parallelus
No
NA
No
NA
Helicodiscidae
Lucilla scintilla
No
DD
No
DD
Helicodiscidae
Lucilla singleyana
No
NA
No
NA
Helicodontidae
Atenia quadrasi
Yes
LC
Yes*
LC
Helicodontidae
Drepanostoma nautiliforme
Yes
NT
No*
DD
Helicodontidae
Falkneria camerani
Yes
VU
D2
Yes*
VU
D2
Helicodontidae
Helicodonta angigyra
Yes
LC
No
LC
Helicodontidae
Helicodonta langhofferi
Yes
DD
No*
NA
Helicodontidae
Helicodonta obvoluta
Yes
LC
No
LC
Helicodontidae
Helicodonta wilhelminae
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Helicodontidae
Lindholmiola barbata
Yes
LC
Yes*
LC
Helicodontidae
Lindholmiola corcyrensis
Yes
LC
No
LC
Helicodontidae
Lindholmiola girva
Yes
LC
No
LC
Helicodontidae
Lindholmiola lens
Yes
LC
No
LC
Helicodontidae
Lindholmiola pirinensis
Yes
LC
No
LC
Helicodontidae
Lindholmiola regisborisi
Yes
LC
Yes*
LC
Helicodontidae
Lindholmiola reischuetzi
Yes
LC
Yes*
LC
44
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Helicodontidae
Lindholmiola spectabilis
Yes
LC
Yes*
LC
Helicodontidae
Soosia diodonta
Yes
NT
No*
NT
Hydrocenidae
Hydrocena cattaroensis
Yes
LC
No
LC
Hydrocenidae
Hydrocena gutta
Yes
LC
Yes
LC
Hygromiidae
Actinella actinophora
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Actinella anaglyptica
Yes
NT
Yes*
NT
Hygromiidae
Actinella arcta
Yes
LC
Yes*
LC
Hygromiidae
Actinella armitageana
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Actinella arridens
Yes CR B1ab(iii) Yes* CR B1ab(iii)
Hygromiidae
Actinella carinofausta
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Hygromiidae
Actinella fausta
Yes
LC
Yes*
LC
Hygromiidae
Actinella giramica
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Actinella laciniosa
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Actinella lentiginosa
Yes
LC
Yes*
LC
Hygromiidae
Actinella littorinella
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Actinella nitidiuscula
Yes
LC
Yes*
LC
Hygromiidae
Actinella obserata
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Actinella robusta
Yes
DD
Yes*
DD
Hygromiidae
Ashfordia granulata
Yes
LC
Yes*
LC
Hygromiidae
Canariella berthelotii
Yes
LC
Yes*
LC
Hygromiidae
Canariella bimbachensis
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Canariella discobolus
Yes
NT
Yes*
NT
Hygromiidae
Canariella eutropis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Canariella falkneri
Yes
LC
Yes*
LC
Hygromiidae
Canariella fortunata
Yes
VU
B1ab(ii,iii)+2ab(ii,iii)
Yes*
VU
B1ab(ii,iii)+2ab(ii,iii)
Hygromiidae
Canariella giustii
Yes
LC
Yes*
LC
Hygromiidae
Canariella gomerae
Yes
LC
Yes*
LC
Hygromiidae
Canariella hispidula
Yes
VU
B1ab(i,ii,iii)+2ab(i,ii,iii)
Yes*
VU
B1ab(i,ii,iii)+2ab(i,ii,iii)
Hygromiidae
Canariella huttereri
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Canariella jandiaensis
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Canariella lanosa
Yes
LC
Yes*
LC
Hygromiidae
Canariella leprosa
Yes
VU
B1ab(iii)
Yes*
VU
B1ab(iii)
45
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Canariella multigranosa
Yes
LC
Yes*
LC
Hygromiidae
Canariella planaria
Yes
LC
Yes*
LC
Hygromiidae
Canariella plutonia
Yes
LC
Yes*
LC
Hygromiidae
Canariella pontelirae
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Canariella pthonera
Yes
VU
B1ab(iii)
Yes*
VU
B1ab(iii)
Hygromiidae
Canariella ronceroi
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Hygromiidae
Canariella squamata
Yes
NT
Yes*
NT
Hygromiidae
Canariella subhispidula
Yes
LC
Yes*
LC
Hygromiidae
Canariella tenuicostulata
Yes LC Yes* LC
Hygromiidae
Canariella tillieri
Yes
NT
Yes*
NT
Hygromiidae
Candidula arganica
Yes
LC
Yes*
LC
Hygromiidae
Candidula belemensis
Yes
LC
Yes*
LC
Hygromiidae
Candidula camporroblensis
Yes
LC
Yes*
LC
Hygromiidae
Candidula castriota
Yes
NT
No*
NA
Hygromiidae
Candidula cavannae
Yes
NT
Yes*
NT
Hygromiidae
Candidula codia
Yes
LC
Yes*
LC
Hygromiidae
Candidula corbellai
Yes
LC
Yes
LC
Hygromiidae
Candidula coudensis
Yes
NT
Yes
NT
Hygromiidae
Candidula fiorii
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Candidula gigaxii
Yes
LC
Yes*
LC
Hygromiidae
Candidula grovesiana
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes*
EN
B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Candidula intersecta
Yes
LC
Yes*
LC
Hygromiidae
Candidula lernaea
Yes
LC
Yes*
LC
Hygromiidae
Candidula najerensis
Yes
LC
Yes*
LC
Hygromiidae
Candidula olisippensis
Yes
LC
Yes*
LC
Hygromiidae
Candidula rhabdotoides
Yes
LC
No
LC
Hygromiidae
Candidula rocandioi
Yes
LC
Yes*
LC
Hygromiidae
Candidula setubalensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Candidula spadae
Yes
VU
B1ab(iii)
Yes*
VU
B1ab(iii)
Hygromiidae
Candidula syrensis
Yes
LC
Yes*
LC
Hygromiidae
Candidula ultima
Yes
NT
Yes*
NT
Hygromiidae
Candidula unifasciata
Yes
LC
No
LC
46
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Caseolus abjectus
Yes
LC
Yes*
LC
Hygromiidae
Caseolus baixoensis
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Caseolus calculus
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Caseolus calvus
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Caseolus commixtus
Yes
LC
Yes*
LC
Hygromiidae
Caseolus consors
Yes
LC
Yes*
LC
Hygromiidae
Caseolus hartungi
Yes
LC
Yes*
LC
Hygromiidae
Caseolus innominatus
Yes
LC
Yes*
LC
Hygromiidae
Caseolus leptostictus
Yes VU B2ab(iii); D2 Yes* VU B2ab(iii); D2
Hygromiidae
Caseolus punctulatus
Yes
LC
Yes*
LC
Hygromiidae
Caseolus setulosus
Yes
LC
Yes*
LC
Hygromiidae
Caseolus subcalliferus
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Cernuella aginnica
Yes
LC
Yes*
LC
Hygromiidae
Cernuella amanda
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Cernuella aradasii
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Cernuella caruanae
No
LC
No
LC
Hygromiidae
Cernuella cisalpina
Yes
LC
No
LC
Hygromiidae
Cernuella hydrutina
Yes
NT
Yes*
NT
Hygromiidae
Cernuella lampedusae
Yes
NT
Yes*
NT
Hygromiidae
Cernuella neglecta
Yes
LC
No
LC
Hygromiidae
Cernuella rugosa
Yes
EN
B1ab(i,ii,iii,iv)+2ab(i,ii,iii,iv)
Yes*
EN
B1ab(i,ii,iii,iv)+2ab(i,ii,iii,iv)
Hygromiidae
Cernuella virgata
No
LC
No
LC
Hygromiidae
Cernuellopsis ghisottii
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Ciliella ciliata
Yes
LC
No
LC
Hygromiidae
Ciliellopsis oglasae
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Cryptosaccus asturiensis
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Cyrnotheba corsica
Yes
LC
Yes*
LC
Hygromiidae
Discula attrita
Yes
NT
Yes*
NT
Hygromiidae
Discula bulverii
Yes
CR
B2ab(iii,v)
Yes*
CR
B2ab(iii,v)
Hygromiidae
Discula calcigena
Yes
LC
Yes*
LC
Hygromiidae
Discula cheiranthicola
Yes
NT
Yes*
NT
Hygromiidae
Discula lyelliana
Yes
CR(PE)
B1ab(iii)
Yes*
CR(PE)
B1ab(iii)
47
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Discula polymorpha
Yes
LC
Yes*
LC
Hygromiidae
Discula pulvinata
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
Hygromiidae
Discula rotula
Yes
LC
Yes*
LC
Hygromiidae
Discula tabellata
Yes
CR
B1ab(iii)
Yes*
CR
B1ab(iii)
Hygromiidae
Discula tectiformis
Yes
EN
B1ab(iii)
Yes*
EN
B1ab(iii)
Hygromiidae
Discula testudinalis
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Hygromiidae
Discula tetrica
Yes
CR(PE)
B1ab(iii)
Yes*
CR(PE)
B1ab(iii)
Hygromiidae
Disculella compar
Yes
NT
Yes*
NT
Hygromiidae
Disculella madeirensis
Yes LC Yes* LC
Hygromiidae
Disculella spirulina
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Euomphalia strigella
Yes
LC
No
LC
Hygromiidae
Ganula gadirana
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Ganula lanuginosa
Yes
LC
Yes*
LC
Hygromiidae
Geomitra coronula
Yes
CR
B1ab(iii,v)+2ab(iii,v); D
Yes
CR
B1ab(iii,v)+2ab(iii,v); D
Hygromiidae
Geomitra delphinuloides
Yes
CR(PE)
D
Yes*
CR(PE)
D
Hygromiidae
Geomitra grabhami
Yes
CR
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Geomitra moniziana
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Geomitra tiarella
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Geomitra watsoni
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes
EN
B1ab(iii,v)+2ab(iii))
Hygromiidae
Helicella bolenensis
Yes
DD
Yes*
DD
Hygromiidae
Helicella cistorum
Yes
LC
Yes*
LC
Hygromiidae
Helicella iberica
Yes
LC
Yes*
LC
Hygromiidae
Helicella itala
Yes
LC
No
LC
Hygromiidae
Helicella juglans
Yes
LC
Yes*
LC
Hygromiidae
Helicella nubigena
Yes
DD
No*
DD
Hygromiidae
Helicella ordunensis
Yes
LC
Yes*
LC
Hygromiidae
Helicella orzai
Yes
NT
Yes*
NT
Hygromiidae
Helicella sabulivaga
Yes
DD
Yes*
DD
Hygromiidae
Helicella stiparum
Yes
EN
B1ab(iii)
Yes*
EN
B1ab(iii)
Hygromiidae
Helicella striatitala
Yes
LC
Yes*
LC
Hygromiidae
Helicella valdeona
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Helicopsis arenosa
Yes
DD
No*
DD
48
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Helicopsis cereoflava
Yes
LC
Yes*
LC
Hygromiidae
Helicopsis cypriola
Yes
LC
Yes*
LC
Hygromiidae
Helicopsis filimargo
Yes
LC
No*
NA
Hygromiidae
Helicopsis gittenbergeri
Yes
LC
Yes*
LC
Hygromiidae
Helicopsis instabilis
Yes
LC
No
LC
Hygromiidae
Helicopsis retowskii
Yes
LC
No*
NA
Hygromiidae
Helicopsis striata
Yes
LC
No
LC
Hygromiidae
Helicotricha carusoi
Yes
LC
Yes*
LC
Hygromiidae
Heterostoma desertae
Yes DD Yes* DD
Hygromiidae
Heterostoma duplex
Yes
DD
Yes*
DD
Hygromiidae
Heterostoma pauperculum
Yes
LC
Yes*
LC
Hygromiidae
Hiltrudia kusmici
Yes
LC
No*
NA
Hygromiidae
Hiltrudia mathildae
Yes
LC
No*
NA
Hygromiidae
Hygromia cinctella
Yes
LC
No
LC
Hygromiidae
Hygromia limbata
Yes
LC
No
LC
Hygromiidae
Hygromia odeca
Yes
VU
B1ab(iii,iv)
Yes*
VU
B1ab(iii,iv)
Hygromiidae
Hygromia tassyi
Yes
VU
D2
No*
VU
D2
Hygromiidae
Hystricella bicarinata
Yes
NT
Yes*
NT
Hygromiidae
Hystricella echinulata
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Hygromiidae
Hystricella leacockiana
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Hystricella oxytropis
Yes
NT
Yes*
NT
Hygromiidae
Hystricella turricula
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Hystricella vermetiformis
Yes
CR(PE)
D
Yes
CR(PE)
D
Hygromiidae
Ichnusomunda sacchii
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Hygromiidae
Ichnusomunda usticensis
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Ichnusotricha berninii
Yes
LC
Yes*
LC
Hygromiidae
Kovacsia kovacsi
Yes
LC
Yes*
LC
Hygromiidae
Lemniscia galeata
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Hygromiidae
Lemniscia michaudi
Yes
NT
Yes*
NT
Hygromiidae
Leptaxis azorica
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis caldeirarum
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Leptaxis drouetiana
Yes
LC
Yes*
LC
49
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Leptaxis furva
Yes
VU
B2ab(iii)
Yes*
VU
B2ab(iii)
Hygromiidae
Leptaxis groviana
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis membranacea
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis minor
Yes
EN
B1ab(iii)
Yes*
EN
B1ab(iii)
Hygromiidae
Leptaxis nivosa
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis sanctaemariae
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis simia
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis terceirana
Yes
LC
Yes*
LC
Hygromiidae
Leptaxis wollastoni
Yes EN B1ab(iii,v)+2ab(iii,v) Yes* EN B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Lozekia deubeli
Yes
LC
Yes*
LC
Hygromiidae
Lozekia transsilvanica
Yes
LC
Yes*
LC
Hygromiidae
Mengoana jeschaui
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola andrius
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola berytensis
Yes
LC
No
LC
Hygromiidae
Metafruticicola coartatus
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola crassicosta
Yes
VU
D2
Yes
VU
D2
Hygromiidae
Metafruticicola dictaeus
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola naxianus
Yes
DD
Yes*
DD
Hygromiidae
Metafruticicola nicosianus
Yes
DD
Yes*
DD
Hygromiidae
Metafruticicola noverca
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola occidentalis
Yes
LC
No
LC
Hygromiidae
Metafruticicola pellitus
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola pieperi
Yes
VU
D2
Yes
VU
D2
Hygromiidae
Metafruticicola redtenbacheri
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola sublecta
Yes
LC
Yes*
LC
Hygromiidae
Metafruticicola zonella
Yes
LC
Yes*
LC
Hygromiidae
Microxeromagna lowei
No
LC
No
LC
Hygromiidae
Monacha aniliensis
Yes
LC
Yes*
LC
Hygromiidae
Monacha atacis
Yes
LC
Yes*
LC
Hygromiidae
Monacha auturica
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Monacha cantiana
Yes
LC
No
LC
Hygromiidae
Monacha carascaloides
Yes
LC
No
LC
50
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Monacha cartusiana
Yes
LC
No
LC
Hygromiidae
Monacha cemenelea
Yes
LC
Yes*
LC
Hygromiidae
Monacha claustralis
Yes
LC
No
LC
Hygromiidae
Monacha consona
Yes
NT
Yes*
NT
Hygromiidae
Monacha cretica
Yes
LC
Yes*
LC
Hygromiidae
Monacha dirphica
Yes
DD
Yes*
DD
Hygromiidae
Monacha dofleini
Yes
LC
No*
NA
Hygromiidae
Monacha emigrata
Yes
LC
No
LC
Hygromiidae
Monacha euboeica
Yes DD Yes* DD
Hygromiidae
Monacha frequens
Yes
LC
No
LC
Hygromiidae
Monacha fruticola
Yes
LC
No*
NA
Hygromiidae
Monacha gregaria
Yes
NT
Yes*
NT
Hygromiidae
Monacha haussknechti
Yes
LC
Yes*
LC
Hygromiidae
Monacha lamalouensis
Yes
DD
Yes*
DD
Hygromiidae
Monacha liebegottae
Yes
LC
No
LC
Hygromiidae
Monacha maasseni
Yes
LC
Yes*
LC
Hygromiidae
Monacha messenica
Yes
LC
Yes*
LC
Hygromiidae
Monacha microtricha
Yes
LC
No*
NA
Hygromiidae
Monacha ocellata
Yes
LC
No
LC
Hygromiidae
Monacha orsinii
Yes
NT
Yes*
NT
Hygromiidae
Monacha oshanovae
Yes
NT
Yes*
NT
Hygromiidae
Monacha ovularis
Yes
NT
No*
NT
Hygromiidae
Monacha parumcincta
Yes
LC
No
LC
Hygromiidae
Monacha pseudorothii
Yes
LC
Yes*
LC
Hygromiidae
Monacha rizzae
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Monacha rothii
Yes
LC
Yes*
LC
Hygromiidae
Monacha ruffoi
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Monacha solidior
Yes
LC
No
LC
Hygromiidae
Monacha syriaca
No
LC
No
LC
Hygromiidae
Monacha venusta
Yes
NT
No*
NT
Hygromiidae
Monachoides bacescui
Yes
LC
Yes*
LC
Hygromiidae
Monachoides fallax
Yes
VU
D2
No*
NA
51
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Monachoides incarnatus
Yes
LC
No
LC
Hygromiidae
Monachoides kosovoensis
Yes
NT
No*
NA
Hygromiidae
Monachoides taraensis
Yes
LC
No*
NA
Hygromiidae
Monachoides vicinus
Yes
LC
No
LC
Hygromiidae
Montserratina becasis
Yes
CR(PE)
B2ab(iii)
Yes*
CR(PE)
B2ab(iii)
Hygromiidae
Montserratina bofilliana
Yes
NT
Yes*
NT
Hygromiidae
Montserratina martorelli
Yes
LC
Yes*
LC
Hygromiidae
Moreletina horripila
Yes
LC
Yes*
LC
Hygromiidae
Moreletina obruta
Yes VU D2 Yes* VU D2
Hygromiidae
Moreletina vespertina
Yes
LC
Yes*
LC
Hygromiidae
Nienhuisiella antonellae
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Perforatella bidentata
Yes
LC
No
LC
Hygromiidae
Perforatella dibothrion
Yes
LC
No
LC
Hygromiidae
Petasina bakowskii
Yes
LC
No
LC
Hygromiidae
Petasina bielzi
Yes
LC
No
LC
Hygromiidae
Petasina edentula
Yes
LC
No
LC
Hygromiidae
Petasina filicina
Yes
LC
No
LC
Hygromiidae
Petasina leucozona
Yes
LC
Yes*
LC
Hygromiidae
Petasina lurida
Yes
LC
No
LC
Hygromiidae
Petasina subtecta
Yes
NT
Yes*
NT
Hygromiidae
Petasina unidentata
Yes
LC
No
LC
Hygromiidae
Plentuisa vendia
Yes
NT
Yes*
NT
Hygromiidae
Polloneria contermina
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Hygromiidae
Ponentina martigena
Yes
LC
Yes*
LC
Hygromiidae
Ponentina ponentina
Yes
LC
Yes*
LC
Hygromiidae
Ponentina revelata
Yes
DD
Yes*
DD
Hygromiidae
Ponentina subvirescens
Yes
LC
Yes*
LC
Hygromiidae
Portugala inchoata
Yes
LC
Yes*
LC
Hygromiidae
Pseudocampylaea lowii
Yes
EX
Yes*
EX
Hygromiidae
Pseudocampylaea portosanctana
Yes
LC
Yes*
LC
Hygromiidae
Pseudotrichia rubiginosa
Yes
LC
No
LC
Hygromiidae
Pseudoxerophila bathytera
Yes
LC
Yes*
LC
52
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Pseudoxerophila confusa
Yes
LC
Yes*
LC
Hygromiidae
Pseudoxerophila oertzeni
Yes
LC
Yes*
LC
Hygromiidae
Pyrenaearia cantabrica
Yes
LC
Yes*
LC
Hygromiidae
Pyrenaearia carascalensis
Yes
LC
No
LC
Hygromiidae
Pyrenaearia carascalopsis
Yes
NT
Yes*
NT
Hygromiidae
Pyrenaearia cotiellae
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Pyrenaearia daanidentata
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Pyrenaearia molae
Yes
CR
B1ab(iii)+2ab(iii)
Yes*
CR
B1ab(iii)+2ab(iii)
Hygromiidae
Pyrenaearia navasi
Yes VU D2 Yes* VU D2
Hygromiidae
Pyrenaearia organiaca
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes*
EN
B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Pyrenaearia parva
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Pyrenaearia velascoi
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Schileykiella bodoni
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Schileykiella parlatoris
Yes
NT
Yes*
NT
Hygromiidae
Schileykiella reinae
Yes
NT
Yes*
NT
Hygromiidae
Semifruticicola costulata
Yes
LC
No*
NA
Hygromiidae
Serratorotula acarinata
Yes
CR
B1ab(iii)+2ab(iii); D
Yes
CR
B1ab(iii)+2ab(iii); D
Hygromiidae
Serratorotula coronata
Yes
EN
B1ab(ii,iii)+2ab(ii,iii)
Yes*
EN
B1ab(ii,iii)+2ab(ii,iii)
Hygromiidae
Spirorbula depauperata
Yes
LC
Yes*
LC
Hygromiidae
Spirorbula latens
Yes
LC
Yes*
LC
Hygromiidae
Spirorbula obtecta
Yes
LC
Yes*
LC
Hygromiidae
Spirorbula squalida
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Thyrreniellina josephi
Yes
NT
Yes*
NT
Hygromiidae
Trochoidea carinatoglobosa
Yes
DD
Yes*
DD
Hygromiidae
Trochoidea caroni
Yes
LC
Yes*
LC
Hygromiidae
Trochoidea cumiae
Yes
NT
Yes*
NT
Hygromiidae
Trochoidea elegans
Yes
LC
Yes*
LC
Hygromiidae
Trochoidea liebetruti
Yes
NT
Yes*
NT
Hygromiidae
Trochoidea pyramidata
No
LC
No
LC
Hygromiidae
Trochoidea tarentina
Yes
DD
Yes*
DD
Hygromiidae
Trochoidea trochoides
No
LC
No
LC
Hygromiidae
Trochulus alpicola
Yes
DD
No*
NA
53
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Trochulus ataxiacus
Yes
NT
Yes*
NT
Hygromiidae
Trochulus biconicus
Yes
EN
B1ab(ii,iii,iv)
No*
NA
Hygromiidae
Trochulus caelatus
Yes
NT
No*
NA
Hygromiidae
Trochulus clandestinus
Yes
LC
No
LC
Hygromiidae
Trochulus coelomphala
Yes
DD
Yes*
DD
Hygromiidae
Trochulus erjaveci
Yes
LC
No
LC
Hygromiidae
Trochulus graminicola
Yes
NT
Yes*
NT
Hygromiidae
Trochulus hispidus
Yes
LC
No
LC
Hygromiidae
Trochulus lubomirskii
Yes LC No LC
Hygromiidae
Trochulus montanus
Yes
DD
No*
DD
Hygromiidae
Trochulus oreinos
Yes
NT
Yes*
NT
Hygromiidae
Trochulus phorochaetia
Yes
LC
Yes*
LC
Hygromiidae
Trochulus piccardi
Yes
DD
No*
NA
Hygromiidae
Trochulus plebeius
Yes
DD
No*
DD
Hygromiidae
Trochulus sericeus
Yes
LC
No
LC
Hygromiidae
Trochulus striolatus
Yes
LC
No
LC
Hygromiidae
Trochulus suberecta
Yes
DD
No*
DD
Hygromiidae
Trochulus villosulus
Yes
LC
No
LC
Hygromiidae
Trochulus villosus
Yes
LC
No
LC
Hygromiidae
Trochulus waldemari
Yes
LC
No*
NA
Hygromiidae
Urticicola glabellus
Yes
LC
Yes*
LC
Hygromiidae
Urticicola isaricus
Yes
DD
Yes*
DD
Hygromiidae
Urticicola mounierensis
Yes
NT
Yes*
NT
Hygromiidae
Urticicola moutonii
Yes
DD
Yes*
DD
Hygromiidae
Urticicola suberinus
Yes
DD
Yes*
DD
Hygromiidae
Urticicola umbrosus
Yes
LC
No
LC
Hygromiidae
Wollastonaria jessicae
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Wollastonaria klausgrohi
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Yes
CR
B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Xerocampylaea zelebori
Yes
LC
No
VU
D2
Hygromiidae
Xerocrassa amphiconus
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa barceloi
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa cardonae
Yes
VU
D2
Yes*
VU
D2
54
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Xerocrassa caroli
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa cisternasi
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa claudia
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa claudiconus
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa cobosi
Yes
DD
Yes*
DD
Hygromiidae
Xerocrassa cretica
Yes
LC
No
LC
Hygromiidae
Xerocrassa derogata
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa ebusitana
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa edmundi
Yes EN B1ab(iii)+2ab(iii) Yes* EN B1ab(iii)+2ab(iii)
Hygromiidae
Xerocrassa ferreri
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa franciscoi
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa frater
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa geyeri
Yes
DD
No*
DD
Hygromiidae
Xerocrassa grabusana
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa grata
Yes
DD
Yes*
DD
Hygromiidae
Xerocrassa heraklea
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa homeyeri
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa jimenensis
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa kydonia
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa lasithiensis
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa meda
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa mesostena
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa molinae
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa montserratensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Xerocrassa moraguesi
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes*
EN
B1ab(iii,v)+2ab(iii,v)
Hygromiidae
Xerocrassa nicosiana
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa nyeli
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa penchinati
Yes
LC
No*
LC
Hygromiidae
Xerocrassa poecilodoma
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa prietoi
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa ripacurcica
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa rithymna
Yes
LC
Yes*
LC
55
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Xerocrassa roblesi
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Xerocrassa siderensis
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa siphnica
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa subrogata
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa subvariegata
Yes
LC
Yes*
LC
Hygromiidae
Xerocrassa turolensis
Yes
NT
Yes*
NT
Hygromiidae
Xerocrassa zaharensis
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Xerolenta depulsa
Yes
NT
Yes*
NT
Hygromiidae
Xerolenta macedonica
Yes LC No LC
Hygromiidae
Xerolenta obvia
No
LC
No
LC
Hygromiidae
Xerolenta razlogi
Yes
DD
Yes*
DD
Hygromiidae
Xerolenta spiruloides
Yes
LC
Yes*
LC
Hygromiidae
Xerolenta thasia
Yes
LC
Yes*
LC
Hygromiidae
Xeromunda alticola
Yes
LC
Yes*
LC
Hygromiidae
Xeromunda candiota
Yes
LC
No
LC
Hygromiidae
Xeromunda durieui
No
NT
No*
NT
Hygromiidae
Xeromunda peloponnesia
Yes
LC
Yes*
LC
Hygromiidae
Xeromunda thessalica
Yes
LC
Yes*
LC
Hygromiidae
Xeromunda vulgarissima
Yes
LC
No
LC
Hygromiidae
Xeropicta akrotirica
Yes
LC
Yes*
LC
Hygromiidae
Xeropicta derbentina
No
LC
No
LC
Hygromiidae
Xeropicta krynickii
No
LC
No
LC
Hygromiidae
Xeropicta ledereri
Yes
LC
Yes*
LC
Hygromiidae
Xerosecta adolfi
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Xerosecta arigonis
Yes
LC
No
LC
Hygromiidae
Xerosecta cespitum
Yes
LC
Yes*
LC
Hygromiidae
Xerosecta dohrni
Yes
DD
Yes*
DD
Hygromiidae
Xerosecta explanata
No
EN
B2ab(ii,iii,iv)
No*
EN
B2ab(ii,iii,iv)
Hygromiidae
Xerosecta giustii
Yes
CR
B1ac(iv)+2ac(iv)
Yes*
CR
B1ac(iv)+2ac(iv)
Hygromiidae
Xerosecta hillyeriana
Yes
DD
Yes*
DD
Hygromiidae
Xerosecta introducta
Yes
LC
Yes*
LC
Hygromiidae
Xerosecta promissa
Yes
LC
Yes*
LC
56
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Hygromiidae
Xerosecta reboudiana
No
LC
No
LC
Hygromiidae
Xerosecta terverii
Yes
DD
Yes*
DD
Hygromiidae
Xerotricha adoptata
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha apicina
No
LC
No
LC
Hygromiidae
Xerotricha bierzona
Yes
VU
D2
Yes*
VU
D2
Hygromiidae
Xerotricha conspurcata
No
LC
No
LC
Hygromiidae
Xerotricha corderoi
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha crispolanata
Yes
DD
Yes*
DD
Hygromiidae
Xerotricha garachicoensis
Yes CR(PE) B1ab(iii)+2ab(iii); D Yes CR(PE) B1ab(iii)+2ab(iii); D
Hygromiidae
Xerotricha gasulli
Yes
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Yes*
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Hygromiidae
Xerotricha gonzalezi
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha huidobroi
Yes
NT
Yes*
NT
Hygromiidae
Xerotricha jamuzensis
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha lancerottensis
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha madritensis
Yes
LC
No
LC
Hygromiidae
Xerotricha mariae
Yes
DD
Yes*
DD
Hygromiidae
Xerotricha nodosostriata
Yes
DD
Yes*
DD
Hygromiidae
Xerotricha nubivaga
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha orbignii
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha pavida
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Hygromiidae
Xerotricha renei
Yes
DD
Yes*
DD
Hygromiidae
Xerotricha silosensis
Yes
LC
Yes*
LC
Hygromiidae
Xerotricha vatonniana
No
VU
B1ab(iii)+2ab(iii); D2
No*
VU
B1ab(iii)+2ab(iii); D2
Hygromiidae
Xerotricha zaratei
Yes
NT
Yes*
NT
Hygromiidae
Xerotricha zujarensis
Yes
DD
Yes*
DD
Hygromiidae
Zenobiella subrufescens
No
LC
Yes*
LC
Lauriidae
Hemilauria limnaeana
Yes
LC
Yes*
LC
Lauriidae
Lauria fanalensis
Yes
LC
Yes*
LC
Lauriidae
Lauria fasciolata
Yes
LC
Yes*
LC
Lauriidae
Leiostyla abbreviata
Yes
CR(PE)
B2ab(iii)
Yes*
CR(PE)
B2ab(iii)
Lauriidae
Leiostyla arborea
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla calathiscus
Yes
NT
Yes*
NT
57
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Lauriidae
Leiostyla cassida
Yes
CR(PE)
B2ab(iii)
Yes*
CR(PE)
B2ab(iii)
Lauriidae
Leiostyla cassidula
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Lauriidae
Leiostyla castanea
Yes
DD
Yes*
DD
Lauriidae
Leiostyla cheilogona
Yes
LC
Yes*
LC
Lauriidae
Leiostyla colvillei
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla concinna
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Lauriidae
Leiostyla corneocostata
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla degenerata
Yes
NT
Yes*
NT
Lauriidae
Leiostyla falknerorum
Yes EN B2ab(iii) Yes* EN B2ab(iii)
Lauriidae
Leiostyla ferraria
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla filicum
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla fusca
Yes
LC
Yes*
LC
Lauriidae
Leiostyla fuscidula
Yes
DD
Yes*
DD
Lauriidae
Leiostyla gibba
Yes
CR(PE)
B1ab(iii)
Yes*
CR(PE)
B1ab(iii)
Lauriidae
Leiostyla heterodon
Yes
VU
D2
Yes*
VU
D2
Lauriidae
Leiostyla irrigua
Yes
LC
Yes*
LC
Lauriidae
Leiostyla lamellosa
Yes
EX
Yes*
EX
Lauriidae
Leiostyla laurinea
Yes
VU
B1ab(iii); D2
Yes*
VU
B1ab(iii); D2
Lauriidae
Leiostyla loweana
Yes
LC
Yes*
LC
Lauriidae
Leiostyla macilenta
Yes
VU
B2ab(iii)
Yes*
VU
B2ab(iii)
Lauriidae
Leiostyla millegrana
Yes
LC
Yes*
LC
Lauriidae
Leiostyla monticola
Yes
LC
Yes*
LC
Lauriidae
Leiostyla recta
Yes
LC
Yes*
LC
Lauriidae
Leiostyla relevata
Yes
NT
Yes*
NT
Lauriidae
Leiostyla rugulosa
Yes
LC
Yes*
LC
Lauriidae
Leiostyla simulator
Yes
CR(PE)
D
Yes*
CR(PE)
D
Lauriidae
Leiostyla sphinctostoma
Yes
LC
Yes*
LC
Lauriidae
Leiostyla taeniata
Yes
DD
Yes*
DD
Lauriidae
Leiostyla tesselata
Yes
DD
Yes*
DD
Lauriidae
Leiostyla vermiculosa
Yes
DD
Yes*
DD
Lauriidae
Leiostyla vincta
Yes
LC
Yes*
LC
Limacidae
Bielzia coerulans
Yes
LC
No
LC
58
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Limacidae
Gigantomilax benjaminus
Yes
LC
Yes
LC
Limacidae
Gigantomilax majoricensis
Yes
LC
Yes
LC
Limacidae
Lehmannia brunneri
Yes
LC
No
LC
Limacidae
Lehmannia horezia
Yes
LC
Yes
LC
Limacidae
Lehmannia islandica
Yes
DD
No
DD
Limacidae
Lehmannia janetscheki
Yes
DD
Yes
DD
Limacidae
Lehmannia jaroslaviae
Yes
LC
No
LC
Limacidae
Lehmannia macroflagellata
Yes
LC
No
LC
Limacidae
Lehmannia marginata
No LC No LC
Limacidae
Lehmannia medioflagellata
Yes
VU
D2
Yes
VU
D2
Limacidae
Lehmannia melitensis
Yes
LC
Yes
LC
Limacidae
Lehmannia nyctelia
Yes
LC
No
LC
Limacidae
Lehmannia requienii
Yes
NT
Yes
NT
Limacidae
Lehmannia rupicola
Yes
LC
No
LC
Limacidae
Lehmannia szigethyae
Yes
LC
No
LC
Limacidae
Lehmannia valentiana
Yes
LC
No
LC
Limacidae
Lehmannia vrancensis
Yes
VU
D2
Yes
VU
D2
Limacidae
Limacus flavus
Yes
LC
No
LC
Limacidae
Limacus maculatus
No
LC
No
LC
Limacidae
Limax aeolianus
Yes
NT
Yes
NT
Limacidae
Limax albipes
Yes
LC
Yes
LC
Limacidae
Limax bielzii
Yes
LC
Yes
LC
Limacidae
Limax brandstetteri
Yes
DD
Yes
DD
Limacidae
Limax canapicianus
Yes
DD
Yes
DD
Limacidae
Limax cephalonicus
Yes
LC
No
LC
Limacidae
Limax ciminensis
Yes
DD
Yes
DD
Limacidae
Limax cinereoniger
Yes
LC
No
LC
Limacidae
Limax conemenosi
Yes
LC
No
LC
Limacidae
Limax corsicus
Yes
LC
Yes
LC
Limacidae
Limax dacampi
Yes
LC
No
LC
Limacidae
Limax dobrogicus
Yes
VU
D2
Yes
VU
D2
Limacidae
Limax erythrus
Yes
LC
Yes
LC
59
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Limacidae
Limax gerhardti
Yes
DD
Yes
DD
Limacidae
Limax giovannellae
Yes
DD
Yes
DD
Limacidae
Limax graecus
Yes
LC
No
LC
Limacidae
Limax granosus
Yes
DD
Yes
DD
Limacidae
Limax hemmeni
Yes
VU
D2
Yes
VU
D2
Limacidae
Limax ianninii
Yes
LC
Yes
LC
Limacidae
Limax lachensis
Yes
NT
Yes
NT
Limacidae
Limax maximus
Yes
LC
No
LC
Limacidae
Limax millipunctatus
Yes DD Yes DD
Limacidae
Limax pironae
Yes
DD
Yes
DD
Limacidae
Limax polipunctatus
Yes
DD
Yes
DD
Limacidae
Limax punctulatus
Yes
DD
Yes
DD
Limacidae
Limax redii
Yes
LC
No
LC
Limacidae
Limax sarnensis
Yes
LC
No
LC
Limacidae
Limax squamosus
Yes
DD
Yes
DD
Limacidae
Limax strobeli
Yes
DD
Yes
DD
Limacidae
Limax subalpinus
Yes
LC
Yes
LC
Limacidae
Limax veronensis
Yes
LC
Yes
LC
Limacidae
Limax vizzavonensis
Yes
NT
Yes
NT
Limacidae
Limax wohlberedti
Yes
LC
No
NA
Limacidae
Limax wolterstorffi
Yes
LC
Yes
LC
Limacidae
Malacolimax kostalii
Yes
DD
Yes
DD
Limacidae
Malacolimax mrazeki
Yes
LC
No
LC
Limacidae
Malacolimax tenellus
Yes
LC
No
LC
Limacidae
Malacolimax wiktori
Yes
EN
B2ab(iii,v)
Yes
EN
B2ab(iii,v)
Megalomastomatida
Cochlostoma mariannae
Yes
LC
Yes
LC
Megalomastomatida
Cochlostoma scalariniforme
Yes
DD
No
DD
Megalomastomatida
Cochlostoma sturanii
Yes
LC
No
LC
Megalomastomatida
Obscurella crassilabrum
Yes
LC
Yes
LC
Megalomastomatida
Obscurella obscura
Yes
LC
Yes
LC
Milacidae
Milax aegaeicus
Yes
LC
Yes
LC
Milacidae
Milax altenai
No
LC
No
LC
60
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Milacidae
Milax cyprius
Yes
LC
Yes
LC
Milacidae
Milax gagates
No
LC
No
LC
Milacidae
Milax nigricans
No
LC
No
LC
Milacidae
Milax ochraceus
Yes
DD
Yes
DD
Milacidae
Milax parvulus
Yes
LC
Yes
LC
Milacidae
Milax verrucosus
Yes
LC
Yes
LC
Milacidae
Tandonia albanica
Yes
LC
No
NA
Milacidae
Tandonia bolensis
Yes
VU
D2
Yes
VU
D2
Milacidae
Tandonia bosnensis
Yes DD No NA
Milacidae
Tandonia budapestensis
Yes
LC
No
LC
Milacidae
Tandonia cavicola
Yes
LC
No
LC
Milacidae
Tandonia cretica
Yes
LC
No
LC
Milacidae
Tandonia cristata
No
LC
No
LC
Milacidae
Tandonia croatica
Yes
DD
Yes
DD
Milacidae
Tandonia dalmatina
Yes
DD
Yes
DD
Milacidae
Tandonia fejervaryi
Yes
LC
Yes
LC
Milacidae
Tandonia jablanacensis
Yes
DD
Yes
DD
Milacidae
Tandonia kaleniczenkoi
No
NT
No
NT
Milacidae
Tandonia kusceri
Yes
LC
No
LC
Milacidae
Tandonia lagostana
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Milacidae
Tandonia macedonica
Yes
NT
No
NA
Milacidae
Tandonia marinellii
Yes
NT
Yes
NT
Milacidae
Tandonia melanica
Yes
LC
Yes
LC
Milacidae
Tandonia nigra
Yes
LC
No
LC
Milacidae
Tandonia pageti
No
VU
D2
No
VU
D2
Milacidae
Tandonia pinteri
Yes
LC
Yes
LC
Milacidae
Tandonia piriniana
Yes
NT
Yes
NT
Milacidae
Tandonia rara
Yes
LC
Yes
LC
Milacidae
Tandonia reuleauxi
Yes
LC
No
LC
Milacidae
Tandonia robici
Yes
LC
No
LC
Milacidae
Tandonia rustica
Yes
LC
No
LC
Milacidae
Tandonia sapkarevi
Yes
DD
Yes
DD
61
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Milacidae
Tandonia serbica
Yes
LC
No
LC
Milacidae
Tandonia simrothi
Yes
LC
Yes
LC
Milacidae
Tandonia sowerbyi
Yes
LC
No
LC
Milacidae
Tandonia totevi
Yes
LC
Yes
LC
Oleacinidae
Poiretia compressa
Yes
LC
No
LC
Oleacinidae
Poiretia cornea
Yes
LC
No
LC
Oleacinidae
Poiretia delesserti
Yes
LC
No
LC
Oleacinidae
Poiretia dilatata
Yes
LC
Yes
LC
Oleacinidae
Sardopoiretia emanueli
Yes NT Yes NT
Onchidiidae
Onchidella celtica
Yes
LC
Yes
LC
Orculidae
Odontocyclas kokeilii
Yes
LC
No
LC
Orculidae
Orcula austriaca
Yes
LC
Yes*
LC
Orculidae
Orcula conica
Yes
LC
No
LC
Orculidae
Orcula dobrogica
Yes
DD
Yes*
DD
Orculidae
Orcula dolium
Yes
LC
No
LC
Orculidae
Orcula fuchsi
Yes
CR
B1ab(iii,v)
Yes*
CR
B1ab(iii,v)
Orculidae
Orcula gularis
Yes
LC
Yes*
LC
Orculidae
Orcula jetschini
Yes
LC
Yes*
LC
Orculidae
Orcula pseudodolium
Yes
NT
Yes*
NT
Orculidae
Orcula restituta
Yes
LC
Yes*
LC
Orculidae
Orcula schmidtii
Yes
LC
No
LC
Orculidae
Orcula spoliata
Yes
DD
Yes*
DD
Orculidae
Orcula tolminensis
Yes
LC
Yes*
LC
Orculidae
Orcula wagneri
Yes
NT
No*
NT
Orculidae
Orcula zilchi
No
VU
D2
No*
VU
D2
Orculidae
Orculella aragonica
Yes
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Yes
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Orculidae
Orculella astirakiensis
Yes
LC
Yes*
LC
Orculidae
Orculella bulgarica
No
LC
No
LC
Orculidae
Orculella creantirudis
Yes
LC
Yes*
LC
Orculidae
Orculella creticostata
Yes
LC
Yes*
LC
Orculidae
Orculella cretilasithi
Yes
LC
Yes*
LC
Orculidae
Orculella cretimaxima
Yes
LC
Yes*
LC
62
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Orculidae
Orculella cretiminuta
Yes
LC
Yes*
LC
Orculidae
Orculella cretioreina
Yes
LC
Yes*
LC
Orculidae
Orculella critica
Yes
LC
No
LC
Orculidae
Orculella diensis
Yes
LC
Yes*
LC
Orculidae
Orculella exaggerata
Yes
LC
Yes*
LC
Orculidae
Orculella fodela
Yes
LC
Yes*
LC
Orculidae
Orculella franciscoi
Yes
LC
Yes*
LC
Orculidae
Orculella ignorata
Yes
LC
No
LC
Orculidae
Orculella scalaris
Yes LC Yes* LC
Orculidae
Orculella sirianocoriensis
Yes
LC
No
LC
Orculidae
Orculella templorum
Yes
NT
Yes*
NT
Orculidae
Pagodulina austeniana
Yes
DD
No*
DD
Orculidae
Pagodulina epirotes
Yes
LC
Yes*
LC
Orculidae
Pagodulina hauseri
Yes
LC
Yes*
LC
Orculidae
Pagodulina kaeufeli
Yes
LC
No*
NA
Orculidae
Pagodulina klemmi
Yes
LC
Yes*
LC
Orculidae
Pagodulina pagodula
Yes
LC
No
LC
Orculidae
Pagodulina sparsa
Yes
LC
No
LC
Orculidae
Pagodulina subdola
Yes
LC
No
LC
Orculidae
Pagodulina tschapecki
Yes
DD
Yes*
DD
Orculidae
Sphyradium doliolum
No
LC
No
LC
Orculidae
Walklea rossmaessleri
Yes
LC
Yes*
LC
Otinidae
Otina ovata
Yes
LC
Yes
LC
Oxychilidae
Aegopinella cisalpina
Yes
LC
Yes
LC
Oxychilidae
Aegopinella epipedostoma
Yes
LC
No
LC
Oxychilidae
Aegopinella forcarti
Yes
LC
Yes
LC
Oxychilidae
Aegopinella graziadei
Yes
LC
Yes
LC
Oxychilidae
Aegopinella minor
No
LC
No
LC
Oxychilidae
Aegopinella nitens
Yes
LC
No
LC
Oxychilidae
Aegopinella nitidula
Yes
LC
No
LC
Oxychilidae
Aegopinella pura
No
LC
No
LC
Oxychilidae
Aegopinella ressmanni
Yes
LC
Yes
LC
63
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Oxychilidae
Carpathica bielawskii
Yes
LC
Yes
LC
Oxychilidae
Carpathica boettgeri
Yes
LC
No
NA
Oxychilidae
Carpathica calophana
Yes
LC
No
LC
Oxychilidae
Carpathica cretica
Yes
LC
Yes
LC
Oxychilidae
Carpathica denticulata
Yes
DD
Yes
DD
Oxychilidae
Carpathica elisabethae
Yes
VU
D2
Yes
VU
D2
Oxychilidae
Carpathica insularis
Yes
LC
Yes
LC
Oxychilidae
Carpathica langi
Yes
LC
No
LC
Oxychilidae
Carpathica stussineri
Yes LC No LC
Oxychilidae
Carpathica wirthi
No
DD
No
DD
Oxychilidae
Cellariopsis deubeli
Yes
LC
No
LC
Oxychilidae
Cibinia nana
Yes
VU
D2
Yes
VU
D2
Oxychilidae
Cibinia transsilvanica
Yes
DD
Yes
DD
Oxychilidae
Daudebardia brevipes
No
LC
No
LC
Oxychilidae
Daudebardia dacica
Yes
LC
Yes
LC
Oxychilidae
Daudebardia parvula
Yes
DD
Yes
DD
Oxychilidae
Daudebardia ponorica
Yes
DD
Yes
DD
Oxychilidae
Daudebardia rufa
No
LC
No
LC
Oxychilidae
Daudebardia wiktori
Yes
LC
Yes
LC
Oxychilidae
Eopolita protensa
No
LC
No
LC
Oxychilidae
Mediterranea adamii
Yes
LC
No
LC
Oxychilidae
Mediterranea amaltheae
Yes
CR
B2ab(iii)
Yes
CR
B2ab(iii)
Oxychilidae
Mediterranea depressa
Yes
LC
No
LC
Oxychilidae
Mediterranea hydatina
No
LC
No
LC
Oxychilidae
Mediterranea inopinata
Yes
LC
No
LC
Oxychilidae
Mediterranea ionica
Yes
LC
Yes
LC
Oxychilidae
Mediterranea juliae
Yes
LC
No
LC
Oxychilidae
Mediterranea mariensis
Yes
DD
Yes
DD
Oxychilidae
Mediterranea montivaga
Yes
LC
Yes
LC
Oxychilidae
Mediterranea mylonasi
Yes
LC
Yes
LC
Oxychilidae
Mediterranea pieperi
Yes
VU
D2
Yes
VU
D2
Oxychilidae
Mediterranea planorbis
Yes
LC
No
LC
64
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Oxychilidae
Mediterranea planospiroides
Yes
NT
No
NA
Oxychilidae
Mediterranea polygyra
Yes
LC
Yes
LC
Oxychilidae
Mediterranea pygmaeus
Yes
VU
D2
Yes
VU
D2
Oxychilidae
Mediterranea serbica
Yes
LC
No
NA
Oxychilidae
Mediterranea wiktori
Yes
LC
No
NA
Oxychilidae
Morlina glabra
Yes
LC
No
LC
Oxychilidae
Morlina urbanskii
No
LC
No
LC
Oxychilidae
Nesovitrea hammonis
No
LC
No
LC
Oxychilidae
Nesovitrea petronella
No LC No LC
Oxychilidae
Oxychilus absoloni
Yes
DD
No
NA
Oxychilidae
Oxychilus adjaciensis
Yes
LC
Yes
LC
Oxychilidae
Oxychilus aegopinoides
Yes
EN
B1ab(i,iv)
Yes
EN
B1ab(i,iv)
Oxychilidae
Oxychilus agostinhoi
Yes
CR
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Yes
CR
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Oxychilidae
Oxychilus alicurensis
Yes
NT
Yes
NT
Oxychilidae
Oxychilus alliarius
Yes
LC
No
LC
Oxychilidae
Oxychilus altimirai
Yes
DD
Yes
DD
Oxychilidae
Oxychilus amblyopus
Yes
VU
D2
Yes
VU
D2
Oxychilidae
Oxychilus anjana
Yes
LC
Yes
LC
Oxychilidae
Oxychilus atlanticus
Yes
NT
Yes
NT
Oxychilidae
Oxychilus basajauna
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Oxychilidae
Oxychilus batalhanus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus blauneri
Yes
LC
Yes
LC
Oxychilidae
Oxychilus brincki
Yes
LC
Yes
LC
Oxychilidae
Oxychilus canini
Yes
NT
Yes
NT
Oxychilidae
Oxychilus cellarius
Yes
LC
No
LC
Oxychilidae
Oxychilus clarus
Yes
NT
No
NT
Oxychilidae
Oxychilus colliourensis
Yes
DD
Yes
DD
Oxychilidae
Oxychilus courquini
Yes
LC
Yes
LC
Oxychilidae
Oxychilus deilus
No
LC
No
LC
Oxychilidae
Oxychilus denatale
Yes
NT
Yes
NT
Oxychilidae
Oxychilus diaphanellus
Yes
LC
No
NA
Oxychilidae
Oxychilus diductus
Yes
LC
Yes
LC
65
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Oxychilidae
Oxychilus draparnaudi
Yes
LC
No
LC
Oxychilidae
Oxychilus edmundi
Yes
DD
Yes
DD
Oxychilidae
Oxychilus egadiensis
Yes
LC
Yes
LC
Oxychilidae
Oxychilus farinesianus
Yes
VU
D2
Yes
DD
Oxychilidae
Oxychilus furtadoi
Yes
LC
Yes
LC
Oxychilidae
Oxychilus fuscosus
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Oxychilidae
Oxychilus gardinii
Yes
LC
Yes
LC
Oxychilidae
Oxychilus jugoslavica
Yes
DD
No
NA
Oxychilidae
Oxychilus juvenostriatus
Yes DD Yes DD
Oxychilidae
Oxychilus lagrecai
Yes
NT
Yes
NT
Oxychilidae
Oxychilus lathyri
Yes
NT
Yes
NT
Oxychilidae
Oxychilus lentiformis
Yes
LC
Yes
LC
Oxychilidae
Oxychilus lineolatus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus maceanus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus majori
Yes
LC
Yes
LC
Oxychilidae
Oxychilus mavromoustakisi
Yes
LC
Yes
LC
Oxychilidae
Oxychilus mercadali
Yes
LC
Yes
LC
Oxychilidae
Oxychilus meridionalis
Yes
LC
Yes
LC
Oxychilidae
Oxychilus miceui
Yes
NT
Yes
NT
Oxychilidae
Oxychilus miguelinus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus minor
Yes
LC
Yes
LC
Oxychilidae
Oxychilus mortilleti
Yes
LC
Yes
LC
Oxychilidae
Oxychilus nautiliformis
Yes
NT
No
NA
Oxychilidae
Oxychilus navarricus
Yes
LC
No
LC
Oxychilidae
Oxychilus nortoni
Yes
NT
Yes
NT
Oxychilidae
Oxychilus obscuratus
Yes
NT
Yes
NT
Oxychilidae
Oxychilus oglasicola
Yes
VU
A1e
Yes
VU
A1e
Oxychilidae
Oxychilus oppressus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus ornatus
Yes
DD
Yes
DD
Oxychilidae
Oxychilus paulucciae
Yes
LC
Yes
LC
Oxychilidae
Oxychilus perspectivus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus pilula
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
Yes
EN
B1ab(i,ii,iii,v)+2ab(i,ii,iii,v)
66
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Oxychilidae
Oxychilus pityusanus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus rateranus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus requienii
Yes
LC
Yes
LC
Oxychilidae
Oxychilus scoliura
Yes
NT
Yes
NT
Oxychilidae
Oxychilus shuttleworthianus
Yes
NT
Yes
NT
Oxychilidae
Oxychilus spectabilis
Yes
LC
Yes
LC
Oxychilidae
Oxychilus superfluus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus tomlini
Yes
LC
Yes
LC
Oxychilidae
Oxychilus translucidus
No NA No NA
Oxychilidae
Oxychilus tropidophorus
Yes
LC
Yes
LC
Oxychilidae
Oxychilus uziellii
Yes
LC
Yes
LC
Oxychilidae
Oxychilus viridescens
Yes
LC
Yes
LC
Oxychilidae
Oxychilus volutella
Yes
LC
Yes
LC
Oxychilidae
Retinella circumsessa
Yes
LC
Yes
LC
Oxychilidae
Retinella giustii
Yes
LC
Yes
LC
Oxychilidae
Retinella hierroensis
Yes
LC
Yes
LC
Oxychilidae
Retinella hiulca
Yes
LC
No
LC
Oxychilidae
Retinella incerta
Yes
LC
Yes
LC
Oxychilidae
Retinella lenis
Yes
NT
Yes
NT
Oxychilidae
Retinella olivetorum
Yes
LC
Yes
LC
Oxychilidae
Retinella osoriensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Oxychilidae
Retinella pseudoaegopinella
Yes
LC
Yes
LC
Oxychilidae
Retinella rochebruni
Yes
LC
Yes
LC
Oxychilidae
Retinella stabilei
Yes
DD
Yes
DD
Oxychilidae
Schistophallus camelinus
No
LC
No
LC
Oxychilidae
Schistophallus carotii
Yes
NT
Yes
NT
Oxychilidae
Schistophallus cyprius
No
LC
No
LC
Oxychilidae
Schistophallus investigatus
No
LC
No
LC
Oxychilidae
Schistophallus kobelti
No
EN
B1ab(iii)+2ab(iii)
No
NA
Oxychilidae
Schistophallus minoicus
Yes
LC
Yes
LC
Oxychilidae
Schistophallus oscari
Yes
NT
Yes
NT
Oxychilidae
Schistophallus samius
No
LC
No
LC
67
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Oxychilidae
Schistophallus seidli
Yes
LC
Yes
LC
Oxychilidae
Schistophallus spratti
Yes
LC
Yes
LC
Papillodermatidae
Papilloderma altonagai
Yes
VU
D2
Yes
VU
D2
Parmacellidae
Cryptella alegranzae
Yes
CR
B2ab(iii,v); C2a(ii)
Yes
CR
B2ab(iii,v); C2a(ii)
Parmacellidae
Cryptella auriculata
Yes
VU
B2ab(iv,v)
Yes
VU
B2ab(iv,v)
Parmacellidae
Cryptella canariensis
Yes
NT
Yes
NT
Parmacellidae
Cryptella famarae
Yes
VU
B1ab(iv,v)+2ab(iv,v); D2
Yes
VU
B1ab(iv,v)+2ab(iv,v); D2
Parmacellidae
Cryptella susannae
Yes
VU
D2
Yes
VU
D2
Parmacellidae
Cryptella tamaranensis
Yes CR(PE) D Yes CR(PE) D
Parmacellidae
Drusia deshayesii
No
DD
No
DD
Parmacellidae
Drusia tenerifensis
Yes
EN
B2ab(iii,v)
Yes
EN
B2ab(iii,v)
Parmacellidae
Drusia valenciennii
Yes
LC
Yes
LC
Pleurodiscidae
Pleurodiscus balmei
No
LC
No
LC
Pleurodiscidae
Pleurodiscus cyprius
Yes
NT
Yes
NT
Pleurodiscidae
Pleurodiscus sudensis
Yes
LC
Yes
LC
Pomatiidae
Leonia mamillaris
No
LC
No
LC
Pomatiidae
Pomatias canariensis
Yes
DD
Yes
DD
Pomatiidae
Pomatias elegans
No
LC
No
LC
Pomatiidae
Pomatias glaucus
No
DD
No
DD
Pomatiidae
Pomatias laevigatus
Yes
DD
Yes
DD
Pomatiidae
Pomatias lanzarotensis
Yes
DD
Yes
DD
Pomatiidae
Pomatias palmensis
Yes
DD
Yes
DD
Pomatiidae
Pomatias raricosta
Yes
DD
Yes
DD
Pomatiidae
Pomatias rivularis
No
LC
No
LC
Pomatiidae
Tudorella ferruginea
Yes
LC
Yes
LC
Pomatiidae
Tudorella mauretanica
No
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
No
EN
B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)
Pomatiidae
Tudorella melitensis
Yes
LC
Yes
LC
Pomatiidae
Tudorella multisulcata
No
LC
No
LC
Pomatiidae
Tudorella panormitana
Yes
LC
Yes
LC
Pomatiidae
Tudorella sulcata
No
LC
No
LC
Pristilomatidae
Gollumia torumbilicata
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina candida
Yes
LC
No
LC
68
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Pristilomatidae
Gyralina circumlineata
Yes
LC
No
NT
Pristilomatidae
Gyralina epeirotica
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina ermonae
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina formosa
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina gjirokastrana
Yes
NT
No
NA
Pristilomatidae
Gyralina graeca
Yes
NT
Yes
NT
Pristilomatidae
Gyralina gyralinaeformis
Yes
NT
No
NA
Pristilomatidae
Gyralina hausdorfi
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina korabensis
Yes NT No NA
Pristilomatidae
Gyralina lunikense
Yes
NT
No
NA
Pristilomatidae
Gyralina mirabilis
Yes
LC
No
NA
Pristilomatidae
Gyralina mljetica
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina nautilopsis
Yes
VU
D2
No
NA
Pristilomatidae
Gyralina nopcsai
Yes
NT
No
NA
Pristilomatidae
Gyralina orientalis
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina pageti
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina rempei
Yes
LC
No
NA
Pristilomatidae
Gyralina riedeli
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina sattmanni
Yes
LC
Yes
LC
Pristilomatidae
Gyralina tarabosensis
Yes
VU
D2
No
NA
Pristilomatidae
Gyralina tsatsae
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina velasensis
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Gyralina velkovrhi
Yes
VU
D2
No
NA
Pristilomatidae
Hawaiia minuscula
No
NA
No
NA
Pristilomatidae
Lindbergia beroni
Yes
CR
B1ab(iii)+2ab(iii)
Yes
CR
B1ab(iii)+2ab(iii)
Pristilomatidae
Lindbergia garganoensis
Yes
DD
Yes
DD
Pristilomatidae
Lindbergia gittenbergeri
Yes
CR(PE)
B1ab(iii)+2ab(iii)
Yes
CR(PE)
B1ab(iii)+2ab(iii)
Pristilomatidae
Lindbergia orbicularis
Yes
LC
Yes
LC
Pristilomatidae
Lindbergia pageti
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Lindbergia parnonensis
Yes
DD
Yes
DD
Pristilomatidae
Lindbergia pinteri
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Lindbergia pseudoillyrica
Yes
LC
Yes
LC
69
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Pristilomatidae
Lindbergia spiliaenymphis
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Lindbergia stylokamarae
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Spinophallus uminskii
Yes
NT
Yes
NT
Pristilomatidae
Taurinellushka babugana
Yes
NT
No
NA
Pristilomatidae
Troglovitrea argintarui
Yes
LC
Yes
LC
Pristilomatidae
Vermetum festinans
Yes
CR
B1ab(iii,iv,v)
Yes
CR
B1ab(iii,iv,v)
Pristilomatidae
Vermetum tamadabaensis
Yes
EN
B2ab(iii)
Yes
EN
B2ab(iii)
Pristilomatidae
Vitrea angystropha
No
LC
No
LC
Pristilomatidae
Vitrea argolica
Yes LC Yes LC
Pristilomatidae
Vitrea binderi
Yes
NT
Yes
NT
Pristilomatidae
Vitrea botterii
Yes
LC
No
LC
Pristilomatidae
Vitrea bulgarica
No
LC
No
LC
Pristilomatidae
Vitrea clessini
Yes
LC
Yes
LC
Pristilomatidae
Vitrea contracta
No
LC
No
LC
Pristilomatidae
Vitrea crystallina
Yes
LC
No
LC
Pristilomatidae
Vitrea cyprina
No
LC
No
LC
Pristilomatidae
Vitrea demiobasensis
Yes
LC
Yes
LC
Pristilomatidae
Vitrea diaphana
Yes
LC
No
LC
Pristilomatidae
Vitrea ephesina
No
LC
No
LC
Pristilomatidae
Vitrea erjaveci
Yes
LC
No
LC
Pristilomatidae
Vitrea etrusca
Yes
LC
Yes
LC
Pristilomatidae
Vitrea gasulli
Yes
LC
Yes
LC
Pristilomatidae
Vitrea illyrica
Yes
LC
No
LC
Pristilomatidae
Vitrea inae
Yes
DD
Yes
DD
Pristilomatidae
Vitrea keaana
Yes
LC
Yes
LC
Pristilomatidae
Vitrea kiliasi
Yes
LC
No
NA
Pristilomatidae
Vitrea klemmi
Yes
LC
Yes
LC
Pristilomatidae
Vitrea kutschigi
Yes
LC
No
LC
Pristilomatidae
Vitrea matsakisi
Yes
LC
Yes
LC
Pristilomatidae
Vitrea megistislavras
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Vitrea meijeri
Yes
NT
B2a
No
NA
Pristilomatidae
Vitrea mikuskai
Yes
NT
No
NA
70
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Pristilomatidae
Vitrea minellii
Yes
DD
Yes
DD
Pristilomatidae
Vitrea nadejdae
Yes
EN
B1ab(iii)+2ab(iii)
No
NA
Pristilomatidae
Vitrea narbonensis
Yes
LC
Yes
LC
Pristilomatidae
Vitrea neglecta
Yes
LC
No
LC
Pristilomatidae
Vitrea olympica
Yes
NT
Yes
NT
Pristilomatidae
Vitrea ossaea
Yes
DD
Yes
DD
Pristilomatidae
Vitrea pageti
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Vitrea pieperiana
Yes
CR
B1ab(i,ii,iv)c(ii)+2ab(i,ii,iv)c(ii)
Yes
CR
B1ab(i,ii,iv)c(ii)+2ab(i,ii,iv)c(ii)
Pristilomatidae
Vitrea pinteri
Yes VU D2 Yes VU D2
Pristilomatidae
Vitrea pseudotrolli
Yes
NT
Yes
NT
Pristilomatidae
Vitrea pygmaea
No
LC
No
LC
Pristilomatidae
Vitrea riedeli
No
LC
No
LC
Pristilomatidae
Vitrea riedeliana
No
LC
No
LC
Pristilomatidae
Vitrea schneideri
Yes
NT
Yes
NT
Pristilomatidae
Vitrea schuetti
Yes
LC
Yes
LC
Pristilomatidae
Vitrea selecta
Yes
LC
No
LC
Pristilomatidae
Vitrea siveci
Yes
LC
No
LC
Pristilomatidae
Vitrea sossellai
No
LC
No
LC
Pristilomatidae
Vitrea spelaea
Yes
EN
B1ab(iii)+2ab(iii)
No
NA
Pristilomatidae
Vitrea sporadica
Yes
LC
Yes
LC
Pristilomatidae
Vitrea storchi
Yes
EX
Yes
EX
Pristilomatidae
Vitrea striata
Yes
CR(PE)
B1ab(iii)+2ab(iii)
Yes
CR(PE)
B1ab(iii)+2ab(iii)
Pristilomatidae
Vitrea sturanyi
Yes
LC
No
NA
Pristilomatidae
Vitrea subaii
Yes
LC
Yes
LC
Pristilomatidae
Vitrea subcarinata
Yes
LC
Yes
LC
Pristilomatidae
Vitrea subrimata
Yes
LC
No
LC
Pristilomatidae
Vitrea szekeresi
Yes
LC
Yes
LC
Pristilomatidae
Vitrea thasia
Yes
LC
Yes
LC
Pristilomatidae
Vitrea transsylvanica
Yes
LC
No
LC
Pristilomatidae
Vitrea trolli
Yes
DD
Yes
DD
Pristilomatidae
Vitrea ulrichi
Yes
VU
D2
Yes
VU
D2
Pristilomatidae
Vitrea vereae
Yes
LC
Yes
LC
71
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Pristilomatidae
Vitrea zakynthia
No
LC
No
LC
Pristilomatidae
Vitrea zilchi
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Punctidae
Paralaoma servilis
No
DD
No
DD
Punctidae
Punctum azoricum
Yes
LC
Yes
LC
Punctidae
Punctum pygmaeum
No
LC
No
LC
Pupillidae
Pupilla alpicola
Yes
LC
No
LC
Pupillidae
Pupilla bigranata
Yes
DD
No*
DD
Pupillidae
Pupilla muscorum
Yes
LC
No
LC
Pupillidae
Pupilla sterrii
No LC No LC
Pupillidae
Pupilla triplicata
No
LC
No
LC
Pyramidulidae
Pyramidula cephalonica
No
LC
No
LC
Pyramidulidae
Pyramidula chorismenostoma
No
LC
No
LC
Pyramidulidae
Pyramidula jaenensis
No
LC
No
LC
Pyramidulidae
Pyramidula pusilla
No
LC
No
LC
Pyramidulidae
Pyramidula rupestris
No
LC
No
LC
Pyramidulidae
Pyramidula saxatilis
Yes
LC
No
LC
Spelaeodiscidae
Spelaeodiscus albanicus
Yes
LC
No
NA
Spelaeodiscidae
Spelaeodiscus bulgaricus
Yes
VU
D2
Yes
VU
D2
Spelaeodiscidae
Spelaeodiscus dejongi
Yes
LC
No
NA
Spelaeodiscidae
Spelaeodiscus hauffeni
Yes
LC
Yes
LC
Spelaeodiscidae
Spelaeodiscus obodensis
Yes
LC
No
NA
Spelaeodiscidae
Spelaeodiscus triarius
Yes
LC
No
LC
Spelaeodiscidae
Spelaeodiscus unidentatus
Yes
LC
No
NA
Sphincterochilidae
Sphincterochila baetica
No
LC
No
LC
Sphincterochilidae
Sphincterochila candidissima
No
LC
No
LC
Sphincterochilidae
Sphincterochila cariosula
No
LC
No
LC
Sphincterochilidae
Sphincterochila insularis
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Yes
EN
B1ab(iii,v)+2ab(iii,v)
Streptaxidae
Gibbulinella dealbata
Yes
LC
Yes
LC
Streptaxidae
Gibbulinella dewinteri
Yes
LC
Yes
LC
Strobilopsidae
Klemmia magnicosta
Yes
NT
No
NA
Strobilopsidae
Klemmia sinistrorsa
Yes
NT
No
NA
Strobilopsidae
Virpazaria adrianae
Yes
LC
No
NA
72
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Strobilopsidae
Virpazaria aspectulabeatidis
Yes
NT
No
NA
Strobilopsidae
Virpazaria backhuysi
Yes
NT
No
NA
Strobilopsidae
Virpazaria deelemanorum
Yes
LC
No
NA
Strobilopsidae
Virpazaria nicoleae
Yes
NT
No
NA
Strobilopsidae
Virpazaria pageti
Yes
LC
No
LC
Strobilopsidae
Virpazaria ripkeni
Yes
LC
No
NA
Strobilopsidae
Virpazaria stojaspali
Yes
NT
No
NA
Subulinidae
Rumina decollata
No
LC
No
LC
Subulinidae
Rumina saharica
No LC No LC
Succineidae
Oxyloma dunkeri
Yes
DD
Yes
DD
Succineidae
Oxyloma elegans
No
LC
No
LC
Succineidae
Oxyloma pinteri
Yes
VU
D2
Yes
VU
D2
Succineidae
Oxyloma sarsii
No
LC
No
LC
Succineidae
Quickella arenaria
No
LC
No
LC
Succineidae
Succinea putris
No
LC
No
LC
Succineidae
Succinella oblonga
No
LC
No
LC
Testacellidae
Testacella bisulcata
No
DD
No
DD
Testacellidae
Testacella bracciai
Yes
NT
Yes
VU
B1ab(iii)+2ab(iii)
Testacellidae
Testacella gestroi
Yes
LC
Yes
LC
Testacellidae
Testacella haliotidea
Yes
LC
Yes
LC
Testacellidae
Testacella maugei
No
LC
No
LC
Testacellidae
Testacella riedeli
No
CR
B1ab(iii)+2ab(iii)
No
CR
B1ab(iii)+2ab(iii)
Testacellidae
Testacella scutulum
Yes
LC
No
LC
Trigonochlamydidae
Selenochlamys ysbryda
Yes
DD
No
NA
Trissexodontidae
Caracollina lenticula
No
LC
No
LC
Trissexodontidae
Gasullia gasulli
Yes
NT
Yes*
NT
Trissexodontidae
Gasulliella simplicula
No
LC
No
LC
Trissexodontidae
Gittenbergeria turriplana
Yes
NT
Yes*
NT
Trissexodontidae
Hatumia cobosi
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Trissexodontidae
Hatumia zapateri
Yes
NT
Yes*
NT
Trissexodontidae
Mastigophallus rangianus
Yes
NT
Yes*
NT
Trissexodontidae
Oestophora barbula
Yes
LC
Yes*
LC
73
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Trissexodontidae
Oestophora calpeana
No
EN
B1ab(iii)+2ab(iii)
No*
EN
B1ab(iii)+2ab(iii)
Trissexodontidae
Oestophora dorotheae
No
NA
No*
NA
Trissexodontidae
Oestophora ebria
Yes
NT
Yes*
NT
Trissexodontidae
Oestophora granesae
Yes
VU
D2
Yes*
VU
D2
Trissexodontidae
Oestophora lusitanica
Yes
LC
Yes*
LC
Trissexodontidae
Oestophora mariae
Yes
VU
D2
Yes*
VU
D2
Trissexodontidae
Oestophora ortizi
Yes
NT
Yes*
NT
Trissexodontidae
Oestophora prietoi
Yes
NT
Yes*
NT
Trissexodontidae
Oestophora silvae
Yes LC Yes* LC
Trissexodontidae
Oestophora tarnieri
No
LC
No
LC
Trissexodontidae
Oestophorella buvinieri
Yes
LC
Yes*
LC
Trissexodontidae
Suboestophora altamirai
Yes
VU
D2
Yes*
VU
D2
Trissexodontidae
Suboestophora boscae
Yes
LC
Yes*
LC
Trissexodontidae
Suboestophora hispanica
Yes
VU
D2
Yes*
VU
D2
Trissexodontidae
Suboestophora jeresae
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Trissexodontidae
Suboestophora tarraconensis
Yes
NT
Yes*
NT
Trissexodontidae
Trissexodon constrictus
Yes
LC
Yes*
LC
Truncatellidae
Truncatella subcylindrica
No
LC
No
LC
Valloniidae
Acanthinula aculeata
No
LC
No
LC
Valloniidae
Acanthinula azorica
Yes
LC
Yes*
LC
Valloniidae
Acanthinula spinifera
Yes
DD
Yes*
DD
Valloniidae
Gittenbergia sororcula
No
LC
No
LC
Valloniidae
Plagyrona angusta
Yes
LC
Yes
LC
Valloniidae
Plagyrona placida
No
LC
No
LC
Valloniidae
Spermodea lamellata
Yes
NT
No*
NT
Valloniidae
Spermodea monas
Yes
LC
Yes*
LC
Valloniidae
Vallonia costata
No
LC
No
LC
Valloniidae
Vallonia declivis
Yes
NT
No*
NT
Valloniidae
Vallonia enniensis
No
NT
No*
NT
Valloniidae
Vallonia excentrica
No
LC
No
LC
Valloniidae
Vallonia pulchella
No
LC
No
LC
Valloniidae
Vallonia suevica
Yes
EN
B2ab(iii)
Yes*
EN
B2ab(iii)
74
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Vertiginidae
Columella aspera
No
LC
No
LC
Vertiginidae
Columella columella
Yes
LC
No
LC
Vertiginidae
Columella edentula
Yes
LC
No
LC
Vertiginidae
Columella microspora
Yes
LC
Yes*
LC
Vertiginidae
Spelaeoconcha paganettii
Yes
LC
No
LC
Vertiginidae
Staurodon seminulum
Yes
LC
Yes*
LC
Vertiginidae
Truncatellina arcyensis
Yes
EN
B1ab(iii)+2ab(iii)
Yes*
EN
B1ab(iii)+2ab(iii)
Vertiginidae
Truncatellina atomus
Yes
DD
Yes*
DD
Vertiginidae
Truncatellina callicratis
No LC No LC
Vertiginidae
Truncatellina cameroni
Yes
LC
Yes*
LC
Vertiginidae
Truncatellina claustralis
No
LC
No
LC
Vertiginidae
Truncatellina costulata
No
LC
No
LC
Vertiginidae
Truncatellina cylindrica
No
LC
No
LC
Vertiginidae
Truncatellina laeviuscula
Yes
DD
No*
NA
Vertiginidae
Truncatellina linearis
Yes
DD
Yes*
DD
Vertiginidae
Truncatellina lussinensis
Yes
VU
D2
No*
NA
Vertiginidae
Truncatellina monodon
Yes
LC
No
LC
Vertiginidae
Truncatellina opisthodon
Yes
NT
Yes*
NT
Vertiginidae
Truncatellina purpuraria
Yes
DD
Yes*
DD
Vertiginidae
Truncatellina rothi
Yes
LC
No
LC
Vertiginidae
Truncatellina uniarmata
Yes
DD
No*
DD
Vertiginidae
Truncatellina velkovrhi
Yes
NT
No*
NA
Vertiginidae
Vertigo alpestris
No
LC
No
LC
Vertiginidae
Vertigo angustior
No
VU
A2ac+3c
No*
VU
A2ac+3c
Vertiginidae
Vertigo antivertigo
No
LC
No
LC
Vertiginidae
Vertigo arctica
Yes
NT
No*
NT
Vertiginidae
Vertigo extima
No
DD
No*
DD
Vertiginidae
Vertigo genesii
Yes
LC
No
LC
Vertiginidae
Vertigo geyeri
Yes
LC
No
LC
Vertiginidae
Vertigo heldi
Yes
EN
B2ab(iii)
No*
EN
B2ab(iii)
Vertiginidae
Vertigo lilljeborgi
Yes
NT
No*
NT
Vertiginidae
Vertigo moulinsiana
No
VU
A2ac
No*
VU
A2ac
75
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Vertiginidae
Vertigo parcedentata
Yes
VU
D2
No*
NA
Vertiginidae
Vertigo pusilla
No
LC
No
LC
Vertiginidae
Vertigo pygmaea
No
LC
No
LC
Vertiginidae
Vertigo ronnebyensis
Yes
LC
No
LC
Vertiginidae
Vertigo substriata
No
LC
No
LC
Vertiginidae
Vertigo ultimathule
Yes
NT
No*
NT
Vitrinidae
Azorivitrina brevispira
Yes
LC
Yes*
LC
Vitrinidae
Azorivitrina brumalis
Yes
LC
Yes*
LC
Vitrinidae
Azorivitrina finitima
Yes LC Yes* LC
Vitrinidae
Azorivitrina laxata
Yes
LC
Yes*
LC
Vitrinidae
Eucobresia diaphana
Yes
LC
No
LC
Vitrinidae
Eucobresia glacialis
Yes
DD
No*
DD
Vitrinidae
Eucobresia nivalis
Yes
DD
No*
DD
Vitrinidae
Eucobresia pegorarii
Yes
EN
B2ab(iv)
No*
DD
Vitrinidae
Oligolimax annularis
No
LC
No
LC
Vitrinidae
Oligolimax apatelus
Yes
NT
No*
NA
Vitrinidae
Oligolimax cephalonica
Yes
LC
Yes*
LC
Vitrinidae
Oligolimax musignani
Yes
VU
D2
Yes*
VU
D2
Vitrinidae
Phenacolimax blanci
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Vitrinidae
Phenacolimax major
Yes
NT
No*
NT
Vitrinidae
Phenacolimax stabilei
Yes
DD
Yes*
DD
Vitrinidae
Plutonia albopalliata
Yes
VU
B1ab(iii)+2ab(iii)
Yes*
VU
B1ab(iii)+2ab(iii)
Vitrinidae
Plutonia angulosa
Yes
CR
B1ab(iii,iv)+2ab(iii,iv)
Yes*
CR
B1ab(iii,iv)+2ab(iii,iv)
Vitrinidae
Plutonia atlantica
Yes
LC
Yes*
LC
Vitrinidae
Plutonia behnii
Yes
LC
Yes*
LC
Vitrinidae
Plutonia blauneri
Yes
LC
Yes*
LC
Vitrinidae
Plutonia canariensis
Yes
NT
Yes*
NT
Vitrinidae
Plutonia christinae
Yes
NT
Yes*
NT
Vitrinidae
Plutonia crassa
Yes
DD
Yes
DD
Vitrinidae
Plutonia cuticula
Yes
NT
Yes*
NT
Vitrinidae
Plutonia dianae
Yes
VU
D2
Yes*
VU
D2
Vitrinidae
Plutonia eceroensis
Yes
LC
Yes*
LC
76
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Vitrinidae
Plutonia emmersoni
Yes
NT
Yes*
NT
Vitrinidae
Plutonia falcifera
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Vitrinidae
Plutonia gomerensis
Yes
LC
Yes*
LC
Vitrinidae
Plutonia lamarckii
Yes
LC
Yes*
LC
Vitrinidae
Plutonia machadoi
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Vitrinidae
Plutonia marcida
Yes
LC
Yes*
LC
Vitrinidae
Plutonia mascaensis
Yes
NT
Yes*
NT
Vitrinidae
Plutonia media
Yes
LC
Yes*
LC
Vitrinidae
Plutonia nitida
Yes LC Yes* LC
Vitrinidae
Plutonia nogalesi
Yes
LC
Yes*
LC
Vitrinidae
Plutonia oromii
Yes
LC
Yes*
LC
Vitrinidae
Plutonia parryi
Yes
NT
Yes*
NT
Vitrinidae
Plutonia pelagica
Yes
LC
Yes*
LC
Vitrinidae
Plutonia raquelae
Yes
NT
Yes
NT
Vitrinidae
Plutonia reticulata
Yes
CR
B2ab(iii)
Yes*
CR
B2ab(iii)
Vitrinidae
Plutonia ripkeni
Yes
LC
Yes*
LC
Vitrinidae
Plutonia ruivensis
Yes
LC
Yes*
LC
Vitrinidae
Plutonia solemi
Yes
LC
Yes*
LC
Vitrinidae
Plutonia taburientensis
Yes
LC
Yes*
LC
Vitrinidae
Plutonia tamaranensis
Yes
LC
Yes*
LC
Vitrinidae
Plutonia tuberculata
Yes
NT
Yes*
NT
Vitrinidae
Sardovitrina polloneriana
Yes
VU
D2
Yes*
VU
D2
Vitrinidae
Semilimacella bonellii
Yes
LC
No
LC
Vitrinidae
Semilimacella carniolica
Yes
LC
No
LC
Vitrinidae
Semilimax carinthiacus
Yes
LC
No
LC
Vitrinidae
Semilimax kotulae
Yes
LC
No
LC
Vitrinidae
Semilimax pyrenaicus
Yes
LC
No
LC
Vitrinidae
Semilimax semilimax
Yes
LC
No
LC
Vitrinidae
Vitrina pellucida
No
LC
No
LC
Vitrinidae
Vitrinobrachium baccettii
Yes
NT
Yes*
NT
Vitrinidae
Vitrinobrachium breve
No
LC
No
LC
Vitrinidae
Vitrinobrachium tridentinum
Yes
DD
No*
DD
77
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Zonitidae
Aegopis acies
Yes
LC
No
LC
Zonitidae
Aegopis compressus
Yes
LC
Yes
LC
Zonitidae
Aegopis croaticus
Yes
LC
No
LC
Zonitidae
Aegopis gemonensis
Yes
LC
Yes
LC
Zonitidae
Aegopis italicus
Yes
LC
Yes
LC
Zonitidae
Aegopis septentrionalis
Yes
LC
No
LC
Zonitidae
Aegopis spelaeus
Yes
NT
No
VU
D2
Zonitidae
Aegopis tenerrimus
Yes
NT
No
NA
Zonitidae
Aegopis verticillus
Yes LC No LC
Zonitidae
Allaegopis amphikypellon
Yes
NT
No
NA
Zonitidae
Allaegopis corcyrensis
Yes
DD
Yes
DD
Zonitidae
Allaegopis jonicus
Yes
LC
Yes
LC
Zonitidae
Allaegopis kerketianus
Yes
NT
Yes
NT
Zonitidae
Allaegopis meridionalis
Yes
LC
Yes
LC
Zonitidae
Allaegopis skanderbegianus
Yes
LC
No
LC
Zonitidae
Allaegopis subariedeli
Yes
VU
D2
Yes
VU
D2
Zonitidae
Allaegopis transiens
Yes
LC
Yes
LC
Zonitidae
Balcanodiscus beroni
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus carinatus
Yes
LC
Yes
LC
Zonitidae
Balcanodiscus cerberus
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus danyii
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus difficilis
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus frivaldskyanus
Yes
LC
Yes
LC
Zonitidae
Balcanodiscus magnus
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus mirus
Yes
NT
Yes
NT
Zonitidae
Balcanodiscus stummerorum
Yes
NT
Yes
NT
Zonitidae
Doraegopis boeoticus
Yes
EN
B2ab(iii)
Yes
EN
B2ab(iii)
Zonitidae
Doraegopis carinatus
Yes
VU
D2
Yes
VU
D2
Zonitidae
Doraegopis euboeicus
Yes
NT
Yes
NT
Zonitidae
Doraegopis parnonicus
Yes
VU
D2
Yes
VU
D2
Zonitidae
Doraegopis subaii
Yes
VU
D2
Yes
VU
D2
Zonitidae
Meledella werneri
Yes
LC
Yes
LC
78
Family
Species
Endemic to
Europe?
IUCN Red
List
Category
IUCN Red List Criteria
(Europe)
Endemic to
the EU?
IUCN Red List
Category (EU)
IUCN Red List Criteria
(EU)
Zonitidae
Paraegopis albanicus
Yes
LC
No
LC
Zonitidae
Paraegopis bizonus
Yes
LC
No
NA
Zonitidae
Paraegopis mauritii
Yes
LC
No
LC
Zonitidae
Paraegopis oberwimmeri
Yes
NT
No
NA
Zonitidae
Paraegopis skipetaricus
Yes
LC
No
NA
Zonitidae
Troglaegopis mosorensis
Yes
LC
Yes
LC
Zonitidae
Zonites algirus
Yes
VU
D2
Yes
VU
D2
Zonitidae
Zonites anaphiensis
Yes
VU
D2
Yes
VU
D2
Zonitidae
Zonites astakidae
Yes CR B1ab(iii) Yes CR B1ab(iii)
Zonitidae
Zonites caricus
No
VU
D2
No
VU
D2
Zonitidae
Zonites casius
No
NT
No
NT
Zonitidae
Zonites chloroticus
No
CR
B1ab(i,iii,iv,v)c(i)
No
CR
B1ab(i,iii,iv,v)c(i)
Zonitidae
Zonites embolium
Yes
CR(PE)
B1ab(i,iii,iv,v)
Yes
CR(PE)
B1ab(i,iii,iv,v)
Zonitidae
Zonites euboeicus
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Zonitidae
Zonites festai
No
NT
No
NT
Zonitidae
Zonites graecus
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Zonitidae
Zonites invitus
Yes
CR
B1ab(iii)
Yes
CR
B1ab(iii)
Zonitidae
Zonites kobelti
Yes
EN
B1ab(iii)
Yes
EN
B1ab(iii)
Zonitidae
Zonites labiosus
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Zonitidae
Zonites messenicus
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Yes
EN
B1ab(ii,iii,v)+2ab(ii,iii,v)
Zonitidae
Zonites nautarum
Yes
CR
B1ab(i,iii)
Yes
CR
B1ab(i,iii)
Zonitidae
Zonites nikariae
Yes
VU
D2
Yes
VU
D2
Zonitidae
Zonites nisyrius
Yes
VU
D2
Yes
VU
D2
Zonitidae
Zonites oertzeni
Yes
EN
B1ab(iii)+2ab(iii)
Yes
EN
B1ab(iii)+2ab(iii)
Zonitidae
Zonites parnonensis
Yes
LC
Yes
LC
Zonitidae
Zonites pergranulatus
Yes
VU
B1ab(i,iv,v)
Yes
VU
B1ab(i,iv)
Zonitidae
Zonites rhodius
No
EN
B1ab(i,iii,iv,v)
No
EN
B1ab(i,iii,iv,v)
Zonitidae
Zonites santoriniensis
Yes
EX
Yes
EX
Zonitidae
Zonites sariae
Yes
VU
D2
Yes
VU
D2
Zonitidae
Zonites siphnicus
Yes
EX
Yes
EX
Zonitidae
Zonites smyrnensis
No
NT
No
NT
79
... En las ediciones de la IUCN Red List de 1996 y 1994, P. raricosta aparecía catalogada como como "Lower Risk/Least Concern" (bAIllIe & GRoombRIdGe, 1996) y "Rare" (GRoombRIdGe, 1994). Posteriormente, la especie ha sido catalogada como "Data Deficient" (GRoh, 2017;neubeRt et al. , 2019), debido al escaso conocimiento existente sobre el estado de sus poblaciones y su distribución geográfica. Dada su reducida área de distribución y la baja densidad de sus efectivos, se propone a la UICN la reevaluación del grado de amenaza de esta especie. ...
Article
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Resumen Se realiza una recopilación de los datos conocidos sobre Pomatias raricosta (Wollaston, 1878) y se aportan nuevas localidades en la isla de Tenerife. Además, se completa la descripción de la especie en base a sus características conquiliológicas y aspecto externo del animal, apor-tando las primeras imágenes del animal vivo. Abstract A compilation of known data on Pomatias raricosta (Wollaston, 1878) and new records from the island of Tenerife are provided. In addition, the description of the species is completed on the basis on its conchyliological characteristics and external appearance of the animal, providing the first images of the live animal.
... snails, rats, flatworms) (Holland 2009;Chiba and Cowie 2016). Particularly for European land snail species, the most significant threats are natural ecosystem modifications, residential and commercial development and agriculture (Neubert et al. 2019). ...
Article
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Land snails and the Aegean Archipelago offer an intriguing combination for studying biodiversity, biogeography and ecology. A region with high environmental and temporal heterogeneity and a tri-continental biotic influence and a group of organisms with low active dispersal abilities, high endemism, as well as the particularity to leave shells as traces of past presence, set an ideal stage for testing biodiversity patterns and exploring multisource threats, especially in the era of the ongoing biodiversity crisis. In this study, we examine Helix godetiana, a large-sized, threatened and endemic land snail of the central and south Aegean Islands. The species has been extirpated from 22 of the 32 islands where it was historically present. We identify potential drivers of its extinction, as Helix godetiana faces several threats across its current range, including competitive exclusion by Cornu aspersum, a species with continuing expansion in the Aegean and climate change disrupting its unusual breeding cycle, which occurs in late spring. Our findings shed light on potentially major, yet previously unexplored, threats on endemic molluscs of the Aegean Islands, a European biodiversity hotspot.
... Slugs and snails, comprising 80% of species diversity, are hermaphrodites, possessing both male and female reproductive organs for self-fertilization, enabling greater adaptability in challenging environments (Zając and Kramarz 2017). Terrestrial gastropods, despite being considered pests, contribute to litter decomposition and play a role in plant disease control in agroecosystems (Neubert et al. 2019). Additionally, the International Union for Conservation of Nature (IUCN) Red List https://www.iucnredlist.org ...
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Tañan VB, Sumaya NHN. 2024. Bio-inventory of terrestrial gastropod species in Northern Mindanao, Philippines. Biodiversitas 25: 3391-3402. Gastropods are the largest class in the phylum Mollusca, with an estimated 70,000 to 76,000 species. The prevalence of gastropods is undeniable evidence of their successful adaptation to various habitats, including terrestrial, freshwater, and marine environments. Slugs and snails are extremely effective in the terrestrial environment, where they are involved with several ecological activities such as nutrient recycling, decomposing waste material, and calcium enrichment in calcium-deficient habitats. However, many species (notably the non-indigenous ones) are well-known pests that damage crops, increase pesticide demand, harm natural ecosystems, reduce native biodiversity, and even carry pathogens and parasites. Northern Mindanao is one of the regions in Mindanao, Philippines, with abundant agricultural resources. In comparison to other taxa in Mindanao, gastropod species have received less attention from researchers. Thus, we collected, characterized, and delineated species of gastropods from the different agricultural areas in Northern Mindanao, Philippines. During sampling, environmental parameters were recorded, and it was established which gastropod species are associated with crops and, at their worst, can become pests to banana (Musa acuminata Colla), corn (Zea mays L.), cabbage (Brassica oleracea L.), carrots (Daucus carota L.), rice (Oryza sativa L.), eggplant (Solanum melongena L.), chayote (Sechium edule (Jacq.) Sw.), radish (Raphanus sativus L.), and many more. Morphology and morphometrics following published protocols on the collected organisms successfully characterized the gastropod species. Slug species, namely, Laevicaulis alte (Férussac, 1822), Sarasinula plebeia (P.Fischer, 1868), and Deroceras (Deroceras) laeve (O.F.Müller, 1774) and snail species, Lissachatina fulica (Férussac, 1821), Pomacea canaliculata (Lamarck, 1822), Subulina octona (Bruguière, 1789), Oxychilus (Ortizius) alliarius (J.S.Miller, 1822), and Bradybaena similaris (A.Férussac, 1822), were identified. The study reveals that exotic terrestrial gastropods are prevalent in the agricultural areas across Northern Mindanao. Consequently, additional surveys are necessary to determine the extent of their dispersal, crop preferences, pest status, and to its associated parasites within the Philippines.
... After the political transition, the explorations of terrestrial mollusks fauna started again with some expeditions organized in collaboration between Albanian and Austrian researchers (Welter-Schultes 1996; Riedel & Welter-Schultes, 1996;Dhora & Welter-Schultes, 1996;b). Until 1996, the fauna of terrestrial mollusks counted 237 species of terrestrial and freshwater mollusks for Albania (Dhora, 1994;1995;1995-96;1997a;1997b;1997-2000Dhora & Welter-Schultes, 1996). ...
... Although this chapter focuses heavily on mollusc extinction, noting that this is particularly severe on oceanic islands, it is becoming clear that much greater numbers of non-marine molluscs, though not yet extinct, are under considerable threat. For example, the European Red List of terrestrial molluscs (Neubert et al., 2019) considers 2469 species as native in Europe, but only five of these as Extinct, apparently in marked contrast to the picture painted above. However, 19.5% of the species are Critically Endangered, Endangered or Vulnerable. ...
Chapter
Since 1970, there has been an overall decline in wildlife populations in the order of 52%. Freshwater species populations have declined by 76%; species populations in Central and South America have declined by 83%; and in the Indo-Pacific by 67%. These are often not complete extinctions, but large declines in the numbers of animals in each species, as well as habitat loss. This presents us with a tremendous opportunity, before it is too late to rescue many species. This book documents the present state of wildlife on a global scale, using a taxonomic approach, and serving as a one stop place for people involved in conservation to be able to find out what is in decline, and the success stories that have occurred to bring back species from the brink of extinction - primarily due to conservation management techniques - as models for what we might achieve in the future.
... Sphyradium doliolum is not a globally threatened species (Neubert et al., 2019), although it is regionally threatened or near threatened in many European countries (Wiktor & Riedel, 2002;Beran et al., 2005;Rüetschi et al., 2012;Welter-Schultes, 2012;Balashov, 2016 a, etc). In Ukraine, this species is offi cially protected since 2021 by the Red Book of Ukraine (offi cially listed as a protected species by the Decree of the Ministry of Ecology of Ukraine from 19 January 2021, in eff ect from 12 March 2021). ...
Article
Full-text available
All existing data and most of the available materials on Sphyradium doliolum from Ukraine and Moldova are reviewed. The species is reported from 38 localities (some including several sites) on uplands east of the Carpathians, the Dnipro Upland and in the Crimean Mountains. It mostly inhabits the broadleaved forests, but in one locality was found in a forestless rocky meadow steppe at an altitude of ca. 1400 m in the Crimea. Statistical analysis of the morphometric shell characters shows a significant geographical variability between the populations of S. doliolum from the different parts of Ukraine and Moldova. The most prominent morphometric differences were found between the populations from the Crimean Mountains and from all other East European populations combined. An origin of S. doliolum in Eastern Europe is discussed. The results of the morphometric analyses support the hypothesis of the two different origins during the two separate periods for the populations of S. doliolum in the Crimean Mountains and in the East European Plain. These two groups of populations could be representing the two different lineages that perhaps could be of the two different subspecies, but their status requires additional studies involving molecular genetic methods and material from a wider area.
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A recent paper highlighted the need for increased attention towards species that have vanished from scientific literature and proposed a new nomenclature in view of needed conservation actions, based on the time interval since their disappearance. Here, we propose to implement this concept by showcasing a taxonomic perspective that is often overlooked in conservation studies. In recent decades, research on vertebrates showed that, in several cases, species considered as undescribed were, in fact, first discovered and already named a long time ago. Quite often they fell into oblivion while being considered synonyms of other taxa until they were re-evaluated in subsequent taxonomic revisions. In this context, we are proposing the concept of “long-lost synonymized” species, asking for greater attention to the discipline of taxonomy, the relevance of specimen-based taxonomy and the value of natural history collections within conservation and academic communities.
Article
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Census of the Mollusca (Gastropoda, Bivalvia) living in Carobbio Valley (Lombardy, Brescia). Since the last three years the malacological fauna of Valle del Carobbio, in the southern slope of Monte Maddalena (Municipality of Brescia, Lombardy), was studied: 53 different taxa were recorded, 50 terrestrial and 3 freshwater, 48 of which with shells and 5 slugs. The endemic Italian species/subspecies present in Valle del Carobbio are 4: Chondrina avenoides (Westerlund, 1874), Macrogastra attenuata modulata (Rossmässler, 1835), Helicodonta obvoluta dentata (Westerlund, 1876), and Chilostoma cingulatum baldense (Rossmässler, 1839); the sub-endemic ones, that is endemic of Italy but shared with neighboring foreign areas, are 5: Charpentieria itala albopustulata (De Cristofori & Jan, 1832), Aegopinella cisalpina Riedel, 1983, Aegopis gemonensis (Férussac, 1832), Limax dacampi dacampi Menegazzi, 1855 and Helicodonta angigyra (Rossmässler, 1834); the allochthonous species ascertained are 4: Paralaoma servilis (Shuttleworth, 1852), Arion vulgaris Moquin-Tandon, 1855, Helix cincta Müller, 1774 and Helix lucorum Linnaeus, 1758. Rumina decollata (Linnaeus, 1758), is not indigenous but probably transfaunated from other Italian localities. The malacological fauna present in the investigated area reflects the characteristics of the environment: many calciphilous and thermophilic species are represented, both rupicolous (e.g. Granaria frumentum illyrica (Rossmässler, 1835), Chondrina avenoides (Westerlund, 1874), Charpentieria itala albopustulata (De Cristofori & Jan, 1832), Chilostoma cingulatum baldense (Rossmässler, 1839)), and from litter and on the ground (e.g. Rumina decollata (Linnaeus, 1758), Truncatellina callicratis (Scacc hi, 1833), Lauria sempronii (Charpentier, 1837), Granopupa granum (Draparnaud, 1801), Zebrina detrita (Müller, 1774), Chondrula tridens tridens (Risso, 1826), Paralaoma servilis (Shuttleworth, 1852), Xerotrica consuprcata (Draparnaud, 1801), Candidula unifasciata unifasciata (Poiret, 1801)). In the central part of the Valle del Carobbio, near to Baldovera stream basin, where there are strips of mesophilous woodland and slightly more humid areas, instead more hygrophilous species are present (e.g. Acicula lineolata banki Boeters, Gittenberger & Subai, 1989, Cecilioides acicula (Müller, 1774), Columella edentula (Draparnaud, 1805), Vallonia costata (Müller, 1774), Zonitoides nitidus (Müller, 1774), Aegopinella cisalpina Riedel, 1983, Aegopis gemonensis (Férussac, 1832), Vitrinobrachium cf. breve (Férussac, 1821), Hygromia cinctella (Draparnaud, 1801)) together with some slugs (e.g. Tandonia rustica (Millet, 1843), Limax dacampi dacampi Menegazzi, 1855, Lehmannia marginata (Müller, 1774), Deroceras reticulatum (Müller, 1774)). The studied area is characterized by a developed karst; however, no malacological species were recorded inside the caves, except for their photic area (entrance to the cavities). In the rare water basins of the valley, represented by some pools in the upper part, 2 bivalves were also recorded: Musculium lacustre (Müller, 1774) and Pisidium casertanum (Poli, 1791). By a chorological point of view, 21 different geoelements are represented, such as the European (EUR) with 10 entities, the Southern European (SEU) with 8 entities, the Mediterranean European (EUM) with 6 entities and the Southern Alpine (ALPS) with 5 entities. The 53 malacological species surveyed in Valle del Carobbio are not globally threatened and are almost exclusively listed into the Least Concern (LC) category, according to the IUCN criteria; however, this area is subject to two potential risk elements: the anthropic pressure for the extreme proximity of Brescia, the second city of Lombardy for number of inhabitants, and the fires, which have become increasingly frequent in recent years, also due to the considerable drought recorded during the last summers.
Chapter
European ecosystems and species remain under pressure from intensive agriculture and forestry, fishing, pollution, urban sprawl, invasive species and climate change. This book provides a detailed description and critical analysis of nature conservation responses, achievements and failures, motivated by the concerning state of nature and missed biodiversity targets. It summarises Europe's nature and the impact of human activities, and then gives an overview of relevant international biodiversity treaties and the EU nature conservation policy and legislative framework. The core of the book comprises chapters written by national experts, which cover the UK and twenty-five EU Member States, providing comparative case studies from which valuable lessons are drawn. Covering wide-ranging topics such as biodiversity pressures, legislation and governance, biodiversity strategies, species protection, protected areas, habitat management, and funding, this book is of interest to a wide audience, including academics and professionals involved in nature conservation and related environmental fields.
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Arthropods' decline has been documented in global and local studies, yet they are still not sufficiently protected on a global scale. Crete (Greece), a Mediterranean hotspot is a continental island, renowned for its diverse geology, ecosystems and endemicity of flora and fauna, with many studies on all of the above disciplines dating back to the 19th century. Here we investigate the conservation status of the Cretan Arthropods using Preliminary Automated Conservation Assessments (PACA) and the overlap of Cretan Arthropod distributions with the Natura 2000 protected areas. In order to perform this analysis, we assembled occurrences of the endemic Arthropods in Crete located in the collections of the Natural History Museum of Crete together with literature data. These assessments resulted in 75% of endemic Arthropods as potentially threatened. The hotspots of endemic and threatened taxa are distributed mostly on the mountainous areas where the Natura 2000 protected areas have great coverage. Yet human activities have significant impact even in those areas, while some taxa are not sufficiently covered by Natura 2000. These findings call for countermeasures and conservation actions.
Technical Report
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The Mediterranean Basin biodiversity hotspot is known to be one of the most biologically rich and complex regions on Earth. The north-western part of the hotspot (the area that covers all of the Iberian Peninsula, southern France, most of Italy and Malta) supports important river, lake, karst and wetland systems critical for the survival of threatened and restricted range freshwater species. Through this project we have identified freshwater Key Biodiversity Areas, defined as sites of importance for the global persistence of biodiversity, within the north-western Mediterranean region of the hotspot. Four priority taxonomic groups including freshwater fishes, molluscs, odonata and aquatic plants, were selected to represent a range of trophic levels within the food webs that support wetland ecosystems. Based on the IUCN Red List published data on species conservation status and distributions 218 freshwater KBAs were confirmed by regional experts for these taxonomic groups, covering an area of 100,021 km2 (13% of the sub-region). The KBAs validated through the project support 109 globally threatened species (Critically Endangered, Endangered or Vulnerable species), 91 geographically restricted range species and demographic aggregations of four species. Of these KBAs, 11 also qualify as Alliance for Zero Extinction sites (AZE) confirming the urgency to develop and implement effective conservation actions for freshwater biodiversity in the northwestern Mediterranean countries. The spatial overlap between freshwater KBAs and existing protected areas, in particular sites within the Natura 2000 Network, and other KBAs was higher than in other Mediterranean sub-regions. Around 84% of the total area of these freshwater KBAs (83,921 km2) lies within the boundaries of pre-existing protected areas or KBAs. However, most of these existing management units have been delineated primarily for terrestrial species and often fail to cover and manage effectively the restricted range and threatened species living in freshwater habitats. The remaining 16% of the freshwater KBAs area (16,100 km2 ), lies outside of any existing protected areas or KBAs, representing priority gaps in the current network. These KBAs should be considered within future strategies for improving representation of freshwater biodiversity within the protected areas network. Specific recommendations for conservation actions are mainly focused on improving water management, especially in relation to the over-abstraction of water from springs and ground water, dam construction and pollution from agriculture and urban areas. Increased efforts are also required to control and/or eradicate invasive alien species in freshwater systems. Management of freshwater KBAs at the catchment scale is needed to: i) ensure effective species protection from both upstream and downstream threats often originating from some distance outside of KBAs, and, ii) maintain the quality, quantity and timing of water flows required to sustain freshwater ecosystems. Countries in the sub-region are recommended to implement an Integrated River Basin Management approach (IRBM or similar) to better coordinate conservation and management actions across sectors. KBAs should be identified at local and national level through a participatory process that involves the relevant stakeholders to maximise buy-in and subsequent implementation of conservation actions. As part of the process 128 potential Site Champions have been identified as individuals/organizations best placed to raise awareness of the KBAs and the issues faced with regard to biodiversity threat, and to help to implement the required actions to safeguard these globally important sites. The information presented in this report will help guide and inform policies and conservation actions for freshwater biodiversity in the region.The KBAs identified will be useful to: - inform the strategic expansion of protected area networks by governments and civil society working toward the achievement of the Aichi Biodiversity Targets (in particular Targets 11 and 12), as established by the Convention on Biological Diversity (CBD); - to inform the description or identification of sites under international conventions (such as wetlands of international importance designated under the Ramsar Convention, natural World Heritage Sites, and Ecologically and Biologically Significant Areas as described under the CBD); - inform private sector safeguard policies, environmental standards, and certification schemes; support conservation planning and priority-setting at national and regional levels; and provide local communities with opportunities for employment, recognition, economic investment and societal mobilization. The identification and management of freshwater KBAs can also provide a metric for measurement of progress towards Sustainable Development Goals 6 and 15 to target 6.5, focused on implementation of integrated water resources management at all levels; target 6.6 focused on protecting and restoring water-related ecosystems; target 15.1 focused on the conservation, restoration and sustainable use of terrestrial and inland freshwater ecosystems and their services; and target 15.5 focused on taking urgent and significant action to reduce the degradation of natural habitats, halt the loss of biodiversity and, by 2020, protect and prevent the extinction of threatened species. Finally, it must be noted that the identification and delineation of KBAs is necessarily a fluid and ongoing process responding to the provision of new information and a constantly changing environment. It is therefore expected that this current freshwater KBA dataset will continue to be refined and updated by each of the countries in the region. The work presented here represents the first steps in taking this process forwards and it provides a baseline to inform future freshwater KBA designations. All validated KBAs will soon be publicly available on the World Database of Key Biodiversity Areas website managed by BirdLife International on behalf of the KBA partnership.
Technical Report
Full-text available
Aim | The European Red List is a review of the conservation status of European species according to IUCN regional Red Listing guidelines. It identifies those species that are threatened with extinction at the regional level, so that appropriate conservation action can be taken to improve their status. This Red List publication summarises results for all described native European marine fishes. Scope | All marine fishes, including all hagfish, cartilaginous fish and bony fish species native to or naturalised in Europe before AD 1500 (a total of 1,220 species), have been assessed in this Red List. The geographic scope encompasses the Mediterranean Sea, the Black Sea, the Baltic Sea, the North Sea and/or the European part of the Atlantic Ocean (i.e., the territorial waters and the Exclusive Economic Zones (EEZs) of all European countries in the Eastern part of the Atlantic Ocean, including the EEZs of the Macaronesian islands belonging to Portugal and Spain). Status assessment | The status of all species was assessed using the IUCN Red List Categories and Criteria (IUCN 2012a), which are the world’s most widely accepted system for measuring extinction risk. All assessments followed the Guidelines for Application of IUCN Red List Criteria at Regional and National Levels (IUCN 2012b). These assessments were compiled based on the data and knowledge from a network of leading European and international experts on marine fish. The assessments were then completed and reviewed at three large workshops held in Spain, Belgium and the UK, as well as through email correspondence with relevant experts. More than 110 experts participated in the assessment and review process for European marine fishes. Assessments are available on the European Red List website and data portal: http://ec.europa.eu/environment/nature/ conservation/species/redlist and http://www.iucnredlist. org/initiatives/europe. Results | Overall, 7.5% of the total of European marine fish species that were assessed in this study are considered threatened (i.e., assessed as having an elevated risk of extinction) in European waters. A further 2.6% (26 species) are considered Near Threatened. However, for 204 species (20.6%), there was insufficient scientific information available to be able to evaluate their risk of extinction and thus they were classified as Data Deficient (DD). When more data become available, some of these species might also prove to be threatened. By comparison, of those other groups that were assessed comprehensively in Europe, 59% of freshwater molluscs, 40% of freshwater fishes, 23% of amphibians, 20% of reptiles, 17% of mammals, 16% of dragonflies, 13% of birds, 9% of butterflies and bees, 8% of aquatic plants and 2% of medicinal plants are threatened (IUCN 2011a, Nieto et al. 2014, Allen et al. 2014, BirdLife International 2015). Additional European Red Lists assessing a selection of species showed that 22% of terrestrial molluscs, 16% of crop wild relatives and 15% of saproxylic beetles are also threatened (IUCN 2011a). No other groups have yet been assessed at the European level. Looking at the population trends of European marine fish species, 8.4% (83 species) have declining populations, 21.5% (212 species) are more or less stable and 1.7% (17 species) are increasing. The population trends for 676 species (68.4%) remain unknown. Eight of the 184 species (4.3%) that are endemic to Europe (i.e., they are found nowhere else in the world) are threatened (Critically Endangered, Endangered, or Vulnerable), highlighting the responsibility that European countries have to protect the global populations of these species. Overall, the European areas with the highest diversity of species are the coast of Portugal, the Macaronesian islands and the western Mediterranean Sea. Hotspots of endemic species are found in the Mediterranean Sea, in particular along the European coast, including the Balearic, Ligurian, Tyrrhenian, Adriatic and Aegean Seas. Furthermore, the greatest concentrations of threatened species appear off the Iberian Peninsula, the Mediterranean Sea and the Canary Islands. Finally, the highest number of Data Deficient (DD) species is present near the Macaronesian islands, particularly in the Madeiran and Canarian archipelagos. The main threats to European marine fishes are overfishing, coastal development, energy production and mining, and pollution. Recommendations • Use the European and national Red Lists of marine fishes to inform revisions and implementation of relevant European legislation (including the Marine Strategy Framework Directive and the Maritime Spatial Planning Directive) to improve the status of threatened marine species, and to improve the knowledge of Data Deficient species. • Continue, and where necessary, expand multi-national fisheries conservation initiatives and management of commercially shared stocks of exploited species, particularly in the Mediterranean and Black Seas. • Take immediate measures to (i) reduce target and incidental catches of species assessed as threatened (i.e., Critically Endangered, Endangered and Vulnerable), (ii) enact measures to ensure sustainable management of species threatened by exploitation, and (iii) set and enforce science-based fishing opportunities and multi-annual management plans for all commercial species, especially those fisheries that target or affect species assessed as threatened and Near Threatened based on exploitation. • Ensure compliance with the requirements under the Common Fisheries Policy (CFP) to apply the precautionary approach by harvesting species at levels to ensure Maximum Sustainable Yield (MSY) for all EU fisheries, especially those for highly migratory species. • Ensure compliance with the requirements, under the Common Fisheries Policy (CFP) to apply the ecosystem-based management of fisheries. • Improve collection of fisheries-dependent data of commercial fisheries, in particular data on catch composition, by-catch, landings, discards, and catch per unit effort, and where not already in place, establish monitoring schemes for small scale artisanal and recreational (line and spear) fisheries. • Require fully documented fisheries and proper mapping of fisheries and fishing effort deployed though control technologies such as Vessel Monitoring System (VMS), Vessel Detection System (VDS) and/ or Automatic Identification System (AIS). • Expand fisheries-independent data collection monitoring, especially for Data Deficient species, and monitoring of threatened species that are not 6 currently sampled effectively, and ensure that such data are exchanged with relevant scientific bodies and Regional Fisheries Management Organisations (RFMO) (e.g., ICES, GFCM, NEAFC, ICCAT). Improve EU and RFMO requirements for speciesspecific reporting of catches and landings of all species, especially chondrichthyans, and for quality assurance of these data so that improved analyses of long-term trends can be undertaken. • Carry out analyses of species trends in both the Northeast Atlantic and the Mediterranean Sea. In particular in the Mediterranean Sea, although there is the ongoing MEDITS survey, there is an urgent need for the development of region-wide time-series analyses of these data. • For those taxa with threatened species and taxonomic problems, improved species identification is required in all data collection exercises (including both commercial landings as well as scientific surveys). For that purpose, trainings of species identification to fishers should be provided, in particular for sharks, skates and rays species, with a view to ensuring that these species are not confused and that juveniles in particular can be distinguished. • Ensure that all designated Marine Protected Areas (MPAs) and areas restricted to fisheries, at the national, regional (Regional Seas Conventions) and European levels (Natura 2000 network) provide adequate protection to threatened marine fishes and particularly protect critical habitats for key life cycle stages (e.g., spawning, pupping, nursery and feeding grounds). • Fully adopt and enforce fisheries management measures for designated MPAs with the view to alleviate pressure on marine fish species and on the habitats that are necessary for their conservation. • Expand area-based conservation measures to meet and exceed the globally agreed-upon Aichi Target 11 of 10% coverage of each marine region, by identifying and establishing Fish Stock Recovery areas, as per the Common Fisheries Policy provisions, to protect spawning grounds and concentration of juvenile fish for those commercial fish species assessed as Critically Endangered, Endangered, Vulnerable and Near Threatened. • Revise the list of threatened European marine fishes regularly, and whenever new data becomes available. • Conduct basic biological research for deep-sea and Data Deficient species, especially those that are or have been commercially exploited (e.g., Orange Roughy, Hoplostethus atlanticus, and the wolf-fishes, Anarhichas spp.).
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The world's biodiversity is currently in rapid decline -Europe being no exception -with as principal cause a human-mediated global change. The Natura 2000 network is an important conservation tool for Euro-pean biodiversity; it is a network of natural and semi-natural sites within Europe with high heritage values due to the exceptional flora and fauna they contain. Here, we evaluated the coverage of 300 threatened species by the Natura 2000 network, and determined potential factors influencing the designation of sites and the structure of the network within a country (social, ecological and demographic national fac-tors). Our analysis was based on a coverage ratio between the Natura 2000 sites and distribution maps of threatened European species. We showed that the distributions of a large proportion of threatened species of mammals, birds and reptiles considered in our study were highly covered (above 90%) by the current Natura 2000 network, demonstrating that the Natura 2000 network also covers species not listed in the annexes of the Nature Directives. However, our results confirm that a large proportion of threatened spe-cies (some of them listed on the European annexes), especially fishes, are currently poorly covered by the Natura 2000 network. The coverage of species likely seemed to be highly related to national demographic factors, i.e. the proportion of the national urban population. Our analysis also suggested that the designa-tion of sites depends too strongly on governmental politics, economic and cultural criteria, and interac-tions between society and the environment. A more effective process might be necessary to ensure the Natura 2000 network reaches its potential as the most important and comprehensive network of protected areas intended to halt the loss of biodiversity in Europe in the near future. A peer-reviewed open-access journal