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407
ISSN 0013-8738, Entomological Review, 2019, Vol. 99, No. 3, pp. 407–415. © Pleiades Publishing, Inc., 2019.
Russian Text © The Author(s), 2019, published in Zoologicheskii Zhurnal, 2019, Vol. 98, No. 1, pp. 28–36.
The carpenter moths (Lepidoptera, Cossidae) of Cen-
tral and South America have been scarcely investigated;
in particular, the genital structures are not studied for
most of the genera. In recent years, the fi rst steps have
been taken by a number of authors to revise Cossidae of
the Neotropical fauna. Some genera were re-described
(Yakovlev, 2014b; Penco et al., 2016; Yakovlev et al.,
2016); the subfamily Cossulinae of Central America
was studied in detail (Davis et al., 2008); the catalog of
the fauna of Argentina (Penco and Yakovlev, 2015) was
compiled.
The genus Brypoctia Schoorl, 1990 was established
for Xyleutes strigifer Dyar, 1910, based on the structure
of the appendages of the head and the thoracic sclerites
(Schoorl, 1990). Schoorl reported for it the following
characters: “male antenna bipectinate for approx. 0.4
times its length, labial palpi slender and approx. 0.5
times length of eye-diameter, second segment of labial
palpi apically slightly thickened, tegula ventro-posteri-
orly bluntly pointed.” The other characters noted by him
are too fi ne details of the thorax structure to be examined
in the course of routine faunistic and taxonomic studies
aimed at identifying the material (Yakovlev, 2014a).
The genus Brypoctia was re-described by us (Pencо
et al., 2016) based on the characters of the male geni-
talia traditionally used in Lepidoptera systematics. As
the result of the study of the Neotropical representatives
of the subfamily Zeuzerinae, new species of the genus
Brypoctia were found. Descriptions of these species are
given below, the status of some taxa is clarifi ed, and a
new synonymy and new combinations are established.
MATERIALS AND METHODS
Material from the following depositories was exam-
ined: the Natural History Museum, London (BMNH),
Museum Witt, Munich (MWM) [it would be transferred
later to the Museum of the Bavarian State Collection
of Zoology, Munich (ZSSM)], the Carnegie Museum
of Natural History, Pittsburgh (CMNH), the United
States National Museum of Natural History, Washington
(USNM).
Review of the Genus Brypoctia Schoorl, 1990 (Lepidoptera,
Cossidae) with Descriptions of Five New Species
from Central and South America
R. V. Yakovleva, b*, F. Pencoc, and T. Wittd
aAltai State University, Barnaul, 656049 Russia
bTomsk State University, Tomsk, 634050 Russia
cUniversidad Maimonides, Buenos Aires, 1405BDB Argentina
dWitt Museum, Munich, D-80796 Germany
*е-mail: yakovlev_asu@mail.ru
Received October 30, 2017
Revised February 12, 2018
Accepted June 5, 2018
Abstract—An illustrated catalogue of the genus Brypoctia Schoorl (Lepidoptera, Cossidae), which includes
11 species, is presented. A new synonym, Brypoctia Schoorl, 1990 = Aramos Schoorl, 1990, syn. n., and fi ve new
combinations are established: Brypoctia aeetes (Druce, 1901), comb. n., Brypoctia desdemona (Dyar et Schaus,
1937), comb. n., Brypoctia itys (Druce, 1911), comb. n., Brypoctia ramosa (Schaus, 1892), comb. n., and Brypoctia
ramuscula (Dyar, 1906), comb. n. Five new species are described: Brypoctia greifensteini sp. n. from Venezuela,
B. eqaqo sp. n. from Peru, B. kurupi sp. n., B. monai sp. n. from Brazil, and B. itzamna sp. n. from Guatemala.
DOI: 10.1134/S0013873819030138
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
YAKOVLEV et al.
408
The constant slides of the genitalia were made using
the standard method: mounted in euparal on slides and
colored with eosin.
The drawings were processed using CorelDraw soft-
ware.
RESULTS
Examination of the external characters and genital
structures of representatives of the subfamily Zeuzer-
inae (type species Zeuzera Latreille, 1804) of the New
World has revealed a number of new facts. Zeuzera ra-
mosa Schaus, 1892 (Figs. 1, 1, 2; 2, 1) (type species of
the genus Aramos Schoorl, 1990) has turned out to be
very closely related to Brypoctia strigifer (Dyar, 1910)
(Figs. 1, 3, 4; 2, 2). The male genitalia have character-
istic synapomorphies: lamellar growths of the arms of
the gnathos and a teniform process on the abdominal
margin of the valva. No signifi cant diff erences proving
the generic distinctness of Aramos were found. Thus,
Brypoctia Schoorl, 1990 = Aramos Schoorl, 1990,
syn. n. Correspondingly, the species Zeuzera ramosa,
Z. aeetes Druce, 1901, Z. ramuscula Dyar, 1906, and
Z. itys Druce, 1911 attributed by Schoorl (1990 : 126)
to Aramos should be placed in Brypoctia. In addi-
tion, Schoorl included Zeuzera masoni Schaus, 1894
(Fig. 1, 17) in the genus Aramos. Our examination of
the male genitalia of Z. masoni (Fig. 3, 4) has shown its
clear morphological diff erences from the species men-
tioned above. Zeuzera masoni has a long narrow uncus,
fi ne teniform arms of the gnathos, emarginate abdom-
inal margin of the valva, short wide lateral processes
of the juxta, long saccus, long fi ne phallus with an api-
cally situated opening of the vesica, and the vesica with
a fi ne loriform cornutus. Blanchard and Knudson (1985)
placed Z. masoni in the genus Psychonoctua Grote, 1865
(type species P. personalis Grote, 1865), but our data
do not support their opinion. The genus Psychonoctua,
redescribed by us (Yakovlev et al., 2016), has an abso-
lutely diff erent genitalia structure: a semicircular, poorly
developed saccus, long foliate lateral processes of the
juxta, and a very short and thick phallus with a strong
teniform cornutus in the vesica (Fig. 1, 18; 3, 5). Thus,
the taxonomic position of Z. masoni remains uncertain.
Xyleutes desdemona Dyar et Schaus, 1937 described
from Brazil (Dyar and Schaus, 1937) has also been in-
vestigated by us and attributed to the genus Brypoctia.
Description of New Species
Brypoctia greifensteini Yakovlev, Penco et Witt, sp. n.
(Fig. 1, 5, 6)
Material. Holotype, ♂: Venezuela, Carabobo,
Cordillera de la Costa, Bejuma, Casa Maria, 500−
1600 m, 16.VIII–4.IX.2005, leg. Th. Greifenstein
(MWM, GenPr MWM-28549). Paratypes: 4 ♂ (same
locality); 1 ♂, Venezuela, Carabobo, Cordillera de la
Costa, Bejuma, Casa Maria, 500−1600 m, 13.VIII–
2.IX.2007, leg. Th. Greifenstein (MWM); 1 ♂, Vene-
zuela, Carabobo, Cordillera de la Costa, Bejuma, Casa
Maria, 700 m, 12.VIII–1.IX.2008, leg. Th. Greifenstein
(MWM); 1 ♂, Venezuela, Est. Carabobo, Rio Bor-
burata, 675 m, 24.VIII.1943, primary forest, R. Lichy
(CMNH).
Description. Male. Length of fore wing 22 mm in
holotype, 20‒22 mm in paratypes. Antennae short,
cup-shaped, pale orange. Thorax and abdomen densely
covered with dark gray scales; tegulae and patagia
covered with pale yellow and pale orange scales. Fore
wing narrow, with pointed apex; costal, marginal, and
submarginal areas of wing diff use pale gray, without
pattern; cubital area white with undulate inconspicuous
pattern formed by fi ne gray transverse bands; anal area
pale gray with clearly visible undulate pattern formed
by dark gray transverse bands; fringe uniformly dark
gray. Hind wing gray, without pattern; fringe uniformly
dark gray.
Genitalia (Fig. 2, 3). Uncus relatively short, trian-
gular, gradually narrowed from base to apex; arms of
gnathos thick, apically modifi ed to form heavily scle-
rotized plates with rugose outer margins; dorsal margin
of plate semicircular, distinctly projecting; dorsal mar-
gins of arms of gnathos strongly diverging; abdominal
margins approximate; valva relatively short, its costal
margin even, outer margin oblique, abdominal margin
emarginate in middle 1/3 and bearing fi ne short (shorter
than 1/10 of valva length) teniform, weakly sclerotized
process; juxta very large, with pair of foliate lateral
processes directed dorsally; saccus large, semicircular;
phallus thick, slightly curved, as long as valva; opening
of vesica occupying complex dorsoapical position; lat-
eral surface of vesica with long teniform cornutus.
Female unknown.
Diagnosis. The new species very clearly diff ers from
all the congeners in appearance (orange antennae, con-
trasting patagia and tegulae, a distinctive wing pattern).
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
REVIEW OF THE GENUS BRYPOCTIA SCHOORL, 1990 409
Fig. 1. Adult males of Zeuzerinae (dorsal view): (1) Brypoctia ramosa (Schaus, 1892), holotype (USNM); (2) B. ramosa, Brasil, Espirito
Santo, Santa Teresa, 650 m, February, 1998, leg. Hubert Thony (MWM); (3) B. strigifer (Dyar, 1910), holotype (USNM); (4) B. strigifer,
Mexico, Oaxaca Prov., Metates, 40 km S of Valle national, 900 m, 5−13.VII.1998, leg. Salk, ex coll. R. Brechlin (MWM); (5) B. greifensteini
sp. n., holotype (MWM); (6) B. greifensteini, paratype, Venezuela, Est. Carabobo, Rio Borburata (CMNH); (7) B. eqaqo sp. n., holotype
(MWM); (8) B. kurupi sp. n., holotype (MWM); (9) B. itzamna sp. n., holotype (MWM); (10) B. monai sp. n., holotype (MWM); (11) B. aeetes
(Druce, 1901), comb. n., holotype (BMNH); (12) B. desdemona (Dyar et Schaus, 1937), comb. n., holotype (USNM); (13) B. desdemona,
Brasilien, Santa Catarina, S. Bento do Sul, Sierra Rio Natal, 850 m, Januar, 1999, leg. Hubert Thony (MWM); (14) B. itys (Druce, 1911), comb.
n., syntype (BMNH); (15) B. ramuscula (Dyar, 1906), comb. n., syntype (USNM); (16) B. ramuscula, USA, TX, Cameron Co., 25.99641°N
/ 97.56871°W, Resaca de la Palma St. Pk., subtropical Resaca woodland, 10 May, 2008, MV Sheet 1, H.L. Kons and R.J. Borth (MWM);
(17) Zeuzera masoni Schaus, 1894, USA, TX, Cameron Co., 25.99641°N / 97.56871°W, Resaca de la Palma St. Pk., subtropical Resaca
woodland, 10 May, 2008, MV Sheet 1, H.L. Kons and R.J. Borth (MWM); (18) Psychonoctua personalis Grote, 1865, Cuba, Prov. Holguin,
Sagua de Tanamo (3 km W), 250 m, La Risuena, Cuabal-veget., 27−30.V.2000, leg. Rudloff (MWM).
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
YAKOVLEV et al.
410
Fig. 2. Brypoctia spp., male genitalia (armament in front view; phallus in lateral view): (1) B. ramosa, Brasil, Espirito Santo, Santa Teresa
(GenPr Heterocera MWM: 28.548); (2) B. strigifer, Mexico, Oaxaca Prov., Metates, 40 km S Valle national (GenPr Heterocera MWM:
28.541, MWM); (3) B. greifensteini, holotype (GenPr Heterocera MWM: 28.549); (4) B. eqaqo, holotype (GenPr Heterocera MWM: 28.547);
(5) B. kurupi, holotype (GenPr Heterocera MWM: 28.546); (6) B. itzamna, holotype (GenPr Heterocera MWM: 28.545).
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
REVIEW OF THE GENUS BRYPOCTIA SCHOORL, 1990 411
Etymology. The new species is named after Thomas
Johann Greifenstein (Streitdorf, Germany) who col-
lected largest part of the type series.
Note. The paratype from CMNH is supplied with
a red label inscribed with black ink by H. Clench
“Holo-Typus / Caroxyleutes gen. n. / lichyi / H. Clench;”
however, both these taxa (genus and species) were not
described.
Brypoctia eqaqo Yakovlev, Penco et Witt, sp. n.
(Fig. 1, 7)
Material. Holotype, ♂: Peru, Prov. Cusco, Rio
Unebamba, Koribeni, 700 m, 1−5.XIII.1994, 700 m, leg.
Hacz and Juhasz (GenPr Heterocera MWM: 28.547).
Description. Male. Length of fore wing 20 mm.
Antennae pale brown; head and anterior part of thorax
covered with black scales; thorax and abdomen densely
covered with pale gray scales. Fore wing elongate,
narrow, with dark brown costal margin and with pale
gray wing fi eld sparsely covered with dark gray trans-
verse strokes. Hind wing white, without pattern.
Genitalia (Fig. 2, 4). Uncus small, triangular, nar-
rowed from base to apex; tegumen medium-sized; arms
of gnathos modifi ed into wide longitudinal plates with
rugose outer margin, their margins relatively even,
closely approximate along entire length; valva lanceo-
late, with costal margin weakly curved, with abdominal
margin sharply emarginate (at border between proximal
and middle thirds) and bearing digitate, heavily sclero-
tized process (about 1/5 as long as valva); juxta large,
with pair of lateral foliate processes directed dorsally;
saccus large, semicircular; phallus very large, slightly
longer than valva, with sharply thickened distal end;
opening of vesica occupying complex dorsoapical po-
sition; vesica with deep wrinkles laterally, with long
teniform cornutus.
Female unknown.
Diagnosis. This species resembles B. ramosa in
appearance but clearly diff ers in the structure of the gen-
italia:
—arms of gnathos strongly approximate (in B. ra-
mosa, dorsal margins approximate and abdominal
margins sharply diverging),
—saccus less strongly developed.
Etymology. The new species is named after Eqaqo
[in Kechua], the god of abundance and prosperity in
Inca mythology.
Brypoctia kurupi Yakovlev, Penco et Witt, sp. n.
(Fig. 1, 8)
Material. Holotype, ♂: “Brasilien, Esp. Santo
Leopoldina, Dorf Tirol, 700 m, 40°50′W/24°55′S,
08−20.XII.1996, leg. H. Thony” (GenPr Heterocera
MWM: 28.546).
Description. Male. Length of fore wing 16 mm.
Antennae pale brown; head and anterior part of thorax
covered with black scales; thorax and abdomen densely
covered with brown scales. Fore wing narrow, elongate,
pale gray with inconspicuous dark pattern (brown costal
margin, pale gray inconspicuous undulate pattern over
entire wing fi eld, fi ne brown strokes in cubital area).
Hind wing pale gray, without pattern.
Genitalia (Fig. 2, 5). Uncus small, triangular, with
pointed apex; tegumen medium-sized; arms of gnathos
modifi ed into wide plates with rugose outer margin; ab-
dominal margin of plate semicircular, clearly visible;
outer margins of plates diverging, nearly parallel to each
other; valva lanceolate, sharply narrowed from middle
third to apex, with abdominal margin distinctly emar-
ginate at border between proximal and middle thirds
and bearing digitate, heavily sclerotized process (about
1/5 as long as valva); juxta large, with a pair of lateral
foliate processes directed dorsally; saccus large, semi-
circular; phallus very large, slightly longer than valva,
with sharply thickened distal end; opening of vesica oc-
cupying complex dorsoapical position; vesica with deep
wrinkles on lateral surface, with long teniform cornutus.
Female unknown.
Diagnosis. The new species resembles B. ramosa and
B. eqaqo in appearance but diff ers in a number of ex-
ternal and inner morphological characters:
—pattern on fore wing substantially reduced,
—valva lanceolate, sharply narrowed from middle
third to apex (valva distinctly wider in B. ramosa),
—apex of uncus pointed (rounded in B. eqaqo),
—abdominal margin of arm of valve semicircular,
clearly visible (cuneiform in B. ramosa and B. eqaqo).
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
YAKOVLEV et al.
412
Etymology. The name of the species is derived from
“Kurupi,” the name of the god of fertility in Guarani
mythology.
Brypoctia itzamna Yakovlev, Penco et Witt, sp. n.
(Fig. 1, 9)
Material. Holotype, ♂: Guatemala, Peten Yaxha
national Park, lowland rainforest, 17°04′29″N /
89°23′85″W, 11−13.VII.2002, 300 m, leg. Jan-Peter
Rudloff (GenPr Heterocera MWM: 28.545). Paratype,
1 ♂, same locality (MWM).
Description. Male. Length of fore wing 21 mm in
holotype, 20 mm in paratype. Antennae pale brown;
head and anterior part of thorax covered with black
scales; thorax and abdomen densely covered with brown
scales. Fore wing long, narrow, with distinct pattern.
Costal margin dark brown; median area with wide pale
brown fi eld extending from base to apex of wing and
underlined ventrally with dark brown stripe; cubital area
(in median part of wing) with characteristic drop-shaped
dark brown stroke; cubital and anal areas with relatively
sparse gray transverse strokes against pale gray back-
ground. Hind wing white, without pattern.
Genitalia (Fig. 2, 6). Uncus triangular, with nipplelike
attenuate apex; tegumen medium-sized; arms of gnathos
modifi ed into relatively narrow plates with rugose outer
margin; abdominal margin of plate cuneiform projecting;
outer margins of plates diverging, nearly parallel to each
other; valva relatively short, wide, with nearly even
costal margin, with abdominal margin distinctly emar-
ginate at border between proximal and middle thirds and
bearing digitate, heavily sclerotized process (about 1/4
as long as valva); juxta large, with pair of lateral foliate
processes directed dorsally; saccus large, semicircular;
phallus very large, slightly longer than valva; opening of
vesica occupying complex dorsoapical position; lateral
surface of vesica with lobiform wide, relatively weakly
sclerotized cornutus.
Female unknown.
Diagnosis. The new species is most closely related
to B. ramuscula in appearance but diff ers in a number
of characters:
—apex of uncus nipple-like attenuate (uncus trian-
gular in B. ramuscula),
—valva wide (narrower in B. ramuscula),
—saccus large (smaller in B. ramuscula).
Etymology. The species is named after Itsamna, the
god of the sky in Maya mythology.
Brypoctia monai Yakovlev, Penco et Witt, sp. n.
(Fig. 1, 10)
Material. Holotype, ♂: Brasil, Espirito Santo, Santa
Leopoldina, Birricas, 700 m, January 1998, leg. Hubert
Thony (GenPr Heterocera MWM: 28.543).
Description. Male. Length of fore wing 16 mm.
Antennae, thorax, and abdomen dark gray. Fore wing
elongate, narrow, with vague pattern: wide brown fi eld
extending from costal margin to cubital area (over entire
wing fi eld), bearing drop-shaped spot in middle third;
cubital area gray, with distinct pattern formed by brown
transverse strokes. Hind wing gray, with inconspicuous
striate pattern at wing apex.
Genitalia (Fig. 3, 1). Uncus triangular, relatively
narrow; tegumen medium-sized; arms of gnathos
modifi ed into wide plates with rugose outer margin; ab-
dominal margin of plate cuneiformly projecting, dorsal
margins closely approximate, and abdominal margins
diverging; valva lanceolate, with strongly narrowed
apex, weakly curved costal margin, and with abdominal
margin distinctly emarginate in middle third and bearing
short, weakly sclerotized teniform process (about 1/10
as long as valva); juxta large, with a pair of lateral foliate
processes directed dorsally; saccus large, semicircular;
phallus very large, slightly longer than valva; opening of
vesica occupying complex dorsoapical position; lateral
surface of vesica with lobiform wide, relatively weakly
sclerotized cornutus.
Female unknown.
Diagnosis. The new species is most closely related
to B. desdemona but diff ers in a number of characters:
—pattern on fore wing vague (much brighter, more
contrasting in B. desdemona),
—abdominal margin of valva with more distinct
emargination (in B. desdemona, emargination incon-
spicuous),
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
REVIEW OF THE GENUS BRYPOCTIA SCHOORL, 1990 413
Fig. 3. Zeuzerinae, male genitalia (armament in front view; phallus in lateral view): (1) B. monai, holotype (GenPr Heterocera MWM:
28.543); (2) B. desdemona, Brasilien, Santa Catarina, S. Bento do Sul, Sierra Rio Natal (GenPr Heterocera MWM: 28.542); (3) B. ramuscula,
USA, TX, Cameron Co., 25.99641°N / 97.56871°W, Resaca de la Palma St. Pk. (Gen Pr Heterocera MWM: 28.633); (4) Zeuzera masoni
(Gen Pr Heterocera MWM: 28.629); (5) Psychonoctua personalis, Cuba, Prov. Holguin, Sagua de Tanamo (3 km W), 250 m, La Risuena,
Cuabal-veget. (GenPr Heterocera MWM: 28.559).
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
YAKOVLEV et al.
414
—lamellar growths of arms of gnathos rather long
(much shorter in B. desdemona).
Etymology. The species is named after Moñái, the
god of the open fi elds in Guarani myphology.
Catalog of the Genus Brypoctia Schoorl, 1990
Brypoctia Schoorl, 1990 : 162
Type species (by monotypy) Xyleutes strigifer Dyar,
1910.
Brypoctia aeetes (Druce, 1901), comb. n. (Fig. 1, 11)
Zeuzera aeetes Druce, 1901, Ann. Mag. Nat. Hist. VII
(7 : 436). Type locality: Colombia, Bonda. Type mate-
rial: holotype (BMNH). Distribution: Colombia.
Brypoctia desdemona (Dyar et Schaus, 1937),
comb. n. (Figs. 1, 12‒13; 2, 2)
Xyleutes desdemona Dyar et Schaus, 1940, Macro-
lepidoptera of the World 6 (2) : 1266. Type locality:
Espiritu Santo (Brazil). Type material: holotype
(USNM). Distribution: Brazil. Material examined.
1 ♂, Brasilien, Santa Catarina, S. Bento do Sul, Sierra
Rio Natal, 850 m, Januar, 1999, leg. Hubert Thony
(GenPr Heterocera MWM: 28.542); 1 ♂, same data,
December, 1999 (MWM); 4 ♂, Brasil, Espirito Santa,
Santa Leopoldina, Birrincas, 700 m, January, 1998,
leg. Huber Thony (MWM); 1 ♂, Brasil, Morro Reuter,
600 m, Faz. Padre Etemo, 26.XIII.1999, leg. A. Moser
(MWM).
Brypoctia eqaqo Yakovlev, Penco et Witt, sp. n.
Type locality: Peru, Prov. Cusco, Rio Unebamba, Ko-
ribeni. Type material: holotype (MWM). Distribution:
Peru, Prov. Cuzco.
Brypoctia greifensteini Yakovlev, Penco et Witt, sp. n.
Type locality: Venezuela, Carabobo, Cordillera de
la Costa, Bejuma, Casa Maria. Type material: holotype
(MWM). Distribution: the north of Venezuela (Cara-
bobo Prov.).
Brypoctia itys (Druce, 1911), comb. n. (Fig. 1, 14)
Zeuzera itys Druce, 1911, Ann. Mag. Nat. Hist. VIII
(7 : 291).
Type locality: Colombia, Jimenez; Rio Dagua. Type
material: syntypes (BMNH). Distribution: Colombia.
Brypoctia itzamna Yakovlev, Penco et Witt, sp. n.
Type locality: Guatemala, Peten Yaxha National
Park, 17°04′29″N / 89°23′85″W. Type material: holo-
type (MWM). Distribution: Guatemala.
Brypoctia kurupi Yakovlev, Penco et Witt, sp. n.
Type locality: Brasilien, Esp. Santo Leopoldina,
Dorf Tirol, 40°50′W/24°55′S. Type material: holotype
(MWM). Distribution: Brazil.
Brypoctia monai Yakovlev, Penco et Witt, sp. n.
Type locality: Brasil, Espirito Santo, Santa Leo-
poldina, Birricas. Type material: holotype (MWM).
Distribution: Brazil.
Brypoctia ramosa (Schaus, 1892), comb. n.
Zeuzera ramosa Schaus, 1892, Proc. Zool. Soc.
London, 1892 : 329. Type locality: Rio Janeiro [Brazil].
Type material: holotype (USNH). Distribution: Brazil.
Material examined: 1 ♂, Brasil, Espirito Santo, Santa
Teresa, 650 m, February, 1998, leg. Hubert Thony
(GenPr Heterocera MWM: 28.548).
Brypoctia ramuscula (Dyar, 1906), comb. n.
(Figs. 1, 15‒16; 3, 3)
Zeuzera ramuscula Dyar, 1906, Sci. Bull. Mus.
Brooklyn Inst. Arts and Sciences 1 (8), 200. Type lo-
cality: Brownsville, Texas [the USA]. Type material:
syntypes (USNM). Distribution: USA (Texas). Material
examined: 6 ♂, USA, TX, Cameron Co., 25.99641°N /
97.56871°W, Resaca de la Palma St. Pk., subtropical Re-
saca woodland, 10 May, 2008, MV Sheet 1, H.L. Kons
and R.J. Borth (Gen Pr Heterocera MWM: 28.633).
Brypoctia strigifer (Dyar, 1910)
Xyleutes strigifer Dyar, 1910, Proc. U. S. Nat. Mus.,
38 (1742), 269. Type locality: Orizaba, [Veracruz State],
Mexico [18°51′N / 97°06′W]. Type material: holotype
USNM. Distribution: Mexico, Costa Rica, Honduras,
Guatemala and Colombia (Schoorl, 1990). Material ex-
amined. 1 ♂, Mexico, Oaxaca Prov., Metates, 40 km S
Valle national, 900 m, 5−13.VII.1998, leg. Salk, ex coll.
ENTOMOLOGICAL REVIEW Vol. 99 No. 3 2019
REVIEW OF THE GENUS BRYPOCTIA SCHOORL, 1990 415
R. Brechlin (GenPrMWM: 28.541, MWM); 1 ♂, Gua-
temala, Peten: Yaxha national park, lowland rainforest,
17°04.29′N / 89°23.85′W, 300 m, 11−13.VII.2002, leg.
J.-P. Rudloff (GenPrMWM: 28.540).
ACKNOWLEDGMENTS
We express sincere gratitude to our colleagues for help
in the work on the paper: Harald Sulak (Munich, Ger-
many) and Patricia Gentili-Poole (Washington, USA)
for photographs of the specimens of Cossidae depos-
ited in USNM and CMNH. We thank also Geoff Martin
(Great Britain) and the BMNH Council of Trustees for
the permission to publish images of the type specimens
deposited in the museum.
FUNDING
The study was fi nancially supported by the Russian
Foundation for Basic Research (Grant № 18-04-00440).
REFERENCES
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soni (Schaus), New Combination (Lepidoptera: Coss-
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Texas and USA,” Proceedings of the Entomological So-
ciety of Washington 87 (2), 426‒431 (1985).
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