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Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand
Abstract
A new cave-dwelling species of Cyrtodactylus Gray is described from Mae Hong Son Province in northern Thailand. The new species, C. erythrops sp. nov., is characterized by its moderate size (snout-vent length to at least 78 mm), relatively large, closely-spaced, flattened tubercles in 18–20 irregular rows at midbody, low number of ventral scales across midbody (28), absence of precloacal groove, presence of precloacal and femoral pores separated by a diastema, broad subcaudal plates, and dorsal pattern of dark spots and blotches. It is the fifth species of cave-dwelling Cyrtodactylus recorded from Thailand and its discovery adds to the mounting evidence that this genus exhibits unprecedented levels of localized endemism throughout tropical Southeast Asia.
... Bilateral scale counts were given as left/right. The methodology of measurements and meristic counts followed Ngo (2011) Morphological comparisons and analyses were based on specimen examination and data obtained from the literature (Hoang et al. 2007;Rösler et al. 2008;Bauer et al. 2009Bauer et al. , 2010Ngo and Grismer 2010;Nguyen et al. 2010Nguyen et al. , 2015Nguyen et al. , 2017Sumontha et al. 2010;Teynié and David 2010;Luu et al. 2011Luu et al. , 2013Luu et al. , 2016Ngo 2011;Ngo and Chan 2011;Schneider et al. 2011Schneider et al. , 2014Schneider et al. , 2020Kunya et al. 2014;Nazarov et al. 2014Nazarov et al. , 2018Nguyen et al. 2014;Pham et al. 2019). ...
... nov. differs from C. erythrops Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya, 2009 in having fewer femoral pores in males (4-10 vs 18-20), more lamellae under finger IV (18-21 vs 16) and toe IV (21-23 vs 20), and more transverse bands on the dorsum of body (8-9 vs 6-7). ...
A new species of Cyrtodactylus is described on the basis of five specimens collected from the karst formations of Zhenkang County, Yunnan Province, China. Cyrtodactylus zhenkangensis sp. nov. is recognized by having a unique combination of morphological characters, the most diagnostic being: 12–15 enlarged femoral scales on each thigh; 2–5 femoral pores on each thigh in males, 0–3 pitted scales on each thigh in females; eight or nine precloacal pores in a continuous row or separated by one poreless scale in males, 7–9 pitted scales in females; subcaudals enlarged, arranged alternately as single and double on anterior and mostly single at middle and posterior; dorsal surface of head with obvious reticulations. Phylogenetic analyses show that the new species is a member of the C. wayakonei species group and a sister taxon to a clade consisting of C. wayakonei and C. martini based on Maximum Likelihood analyses and Bayesian Inference and differs from its congeners by at least 12.0% genetic divergence in a fragment of the COI gene.
... At least 35 species of the gekkonid genus Cyrtodactylus are reported from Indochina (Smith 1935;Bauer et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Sumontha et al. 2010;Ziegler et al. 2010). Of these, twenty species are from Vietnam, most of them described only in recent years (Smith 1935;Bobrov et al. 1995;Darevsky & Szczerbak 1997;Ziegler et al. 2002;Nguyen et al. 2006;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo, 2008;Ngo & Bauer 2008;Geissler et al. 2009;Ziegler et al. 2010). ...
... Comparisons were made with material from the Zoological Collection at the University of Natural Sciences as well as original published descriptions and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Cox et al. 1998;Das & Lim 2000;Das 2006;Ziegler et al. 2002;Bauer 2002Bauer & 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Grismer et al. 2007Nguyen et al. 2006;Youmans & Grismer 2006;Heidrich et al. 2007;Hoang et al. 2007;Kraus & Allison 2006;Kraus 2007Kraus & 2008Linkem et al. 2008;Oliver et al. 2008;Oliver et al. 2009;Orlov et al. 2007;Nazarov et al. 2008;Rösler & Glaw 2008;Rösler et al. 2007;Ngo 2008;Ngo & Bauer 2008;Geissler et al. 2009;Welton et al. 2009;Lei & Hui 2010;Sumontha et al. 2010;Ziegler et al. 2010 Diagnosis. Cyrtodactylus yangbayensis differs from all other congeners by the following combination of characters: mean SVL of 83.5 mm (± 4.7 mm); original tail long (TL/SVL: 1.28); dark brown blotches on head; nuchal loop broken into two dark fragments or V-shaped; dorsal pattern consisting of five to seven irregular rows of narrow, dark brown bands between limb insertions; 6-8 precloacal pores in males; 5-16 enlarged scales beneath thighs; 18-23 interorbital scales on the frontal bone; 26-32 scales in a straight line between eye and nostril; 39-46 rows of ventral scales between ventrolateral folds; 20-23 irregular, longitudinal rows of weakly-keeled, conical tubercles at midbody between the lateral folds; 28-34 paravertebral tubercles between limb insertions; 18-20 subdigital lamellae on first toe; 15-17 subdigital lamellae on fourth toe; 9-10 enlarged scales on heel; 9-11 irregular bands on original tail; median row of enlarged subcaudal scales. ...
A new species of bent-toed gecko, Cyrtodactylus cucphuongensis sp. nov. is described from the karst forest of Cuc Phuong National Park, Ninh Binh province, Northern Vietnam. It differs from all other species of Indo-Chinese Cyrtodactylus by the following combination of characters: SVL 96.0 mm; absence of precloacal and femoral pores; 19 subdigital lamellae on first toe, 24 on fourth toe; large, dark spots on top of head; wide, prominent nuchal band; and five or six dark, irregular, broad bands between limb insertions
... At least 35 species of the gekkonid genus Cyrtodactylus are reported from Indochina (Smith 1935;Bauer et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Sumontha et al. 2010;Ziegler et al. 2010). Of these, twenty species are from Vietnam, most of them described only in recent years (Smith 1935;Bobrov et al. 1995;Darevsky & Szczerbak 1997;Ziegler et al. 2002;Nguyen et al. 2006;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo, 2008;Ngo & Bauer 2008;Geissler et al. 2009;Ziegler et al. 2010). ...
... Comparisons were made with material from the Zoological Collection at the University of Natural Sciences as well as original published descriptions and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Cox et al. 1998;Das & Lim 2000;Das 2006;Ziegler et al. 2002;Bauer 2002Bauer & 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Grismer et al. 2007Nguyen et al. 2006;Youmans & Grismer 2006;Heidrich et al. 2007;Hoang et al. 2007;Kraus & Allison 2006;Kraus 2007Kraus & 2008Linkem et al. 2008;Oliver et al. 2008;Oliver et al. 2009;Orlov et al. 2007;Nazarov et al. 2008;Rösler & Glaw 2008;Rösler et al. 2007;Ngo 2008;Ngo & Bauer 2008;Geissler et al. 2009;Welton et al. 2009;Lei & Hui 2010;Sumontha et al. 2010;Ziegler et al. 2010 Diagnosis. Cyrtodactylus yangbayensis differs from all other congeners by the following combination of characters: mean SVL of 83.5 mm (± 4.7 mm); original tail long (TL/SVL: 1.28); dark brown blotches on head; nuchal loop broken into two dark fragments or V-shaped; dorsal pattern consisting of five to seven irregular rows of narrow, dark brown bands between limb insertions; 6-8 precloacal pores in males; 5-16 enlarged scales beneath thighs; 18-23 interorbital scales on the frontal bone; 26-32 scales in a straight line between eye and nostril; 39-46 rows of ventral scales between ventrolateral folds; 20-23 irregular, longitudinal rows of weakly-keeled, conical tubercles at midbody between the lateral folds; 28-34 paravertebral tubercles between limb insertions; 18-20 subdigital lamellae on first toe; 15-17 subdigital lamellae on fourth toe; 9-10 enlarged scales on heel; 9-11 irregular bands on original tail; median row of enlarged subcaudal scales. ...
A new species of bent-toed gecko, Cyrtodactylus yangbayensis sp. nov. is described from the monsoon evergreen forests of Truong Son Mountain Range, Khanh Hoa province. It differs from all other species of Indo-Chinese Cyrtodactylus by having a mean SVL of 83.5 mm; dark spots and blotches on the head; no continuous nuchal loop; 6-8 precloacal pores in males arranged in a chevron; five or six dark brown, irregular, dorsal bands between limb insertions; 18-20 subdigital lamellae under first toe; 15-17 subdigital lamellae under fourth toe; and a median row of enlarged subcaudal scales.
... Bilateral scale counts were given as left/right. The methodology of measurements and meristic counts followed Liu and Rao (2021a, b): Morphological comparisons were based on the original descriptions of each species (Hoang et al. 2007;Bauer et al. 2009Bauer et al. , 2010Ngo and Grismer 2010;Nguyen et al. 2010Nguyen et al. , 2015bNguyen et al. , 2017Sumontha et al. 2010;Luu et al. 2011;Ngo 2011;Ngo and Chan 2011;Kunya et al. 2014;Nazarov et al. 2014;Nguyen et al. 2014;Schneider et al. 2014Schneider et al. , 2020Le 2016;Pham et al. 2019;Liu and Rao 2021a;Liu et al. 2021;Zhang et al. 2021). ...
A new species of the Cyrtodactylus chauquangensis species group is described based on four specimens collected from the karst formations of Menglian County, Puer City, Yunnan Province, China. The new species can be separated from all other congeners by having a unique combination of morphological characters: a medium-sized body; ventrolateral folds present with interspersed small tubercles; seven precloacal pores in a continuous series in males, absent in females; enlarged femoral scales and femoral pores absent; two postcloacal tubercles on each side; and one or two rows of enlarged subcaudals. Genetically, the new species most closely related to C. wayakonei and the uncorrected sequence divergences of the ND2 gene and its flanking tRNAs between the new species and investigated congeners range from 7.2% to 18.4%.
... Prior to this study, there were 36 species of Cyrtodactylus Gray known from Thailand which is comparable to that of neighboring Myanmar (45 species), Laos (23 species), Vietnam (43 species), and Peninsular Malaysia (33 species) but far exceeds that of China (five species) and Cambodia (eight species) (Uetz et al. 2020). However, in stark contrast to the neighboring countries, the phylogenetic relationships of 44% (16) of the Thai species are unknown because most of the recent species descriptions did not include molecular data (Bauer et al. 2009(Bauer et al. , 2010Chan-ard & Makchai, 2011;Kunya et al. 2014Kunya et al. , 2015Pauwels et al. 2014aPauwels et al. ,b, 2016, thus precluding any downstream comparative analyses (see Grismer et al. 2020a). To this end, we add C. inthanon and C. doisuthep and lower that value to 38%. ...
A new species of Cyrtodactylus from Tak Province, Thailand, Cyrtodactylus amphipetraeus sp. nov., is described using an integrative taxonomic analysis based on morphology, color pattern, and the mitochondrial gene NADH dehydrogenase subunit 2 (ND2). The phylogenetic analyses place the new species within the C. sinyineensis group which was previously thought to be endemic to the Salween Basin in southern Myanmar. The phylogeny also places C. inthanon in the C. sinyneensis group which is expanded herein to also include the group’s sister species C. doisuthep. Along with C. amphipetraeus sp. nov., these are the first three species of the C. sinyineensis group to be found outside of Myanmar east of the Tenasserim Mountains. The Tenasserim Mountain region is discussed as an area of cladogeneic turnover.
... The abbreviations are as follows: / = data not available in literature; -=; characteristic not present; + = characteristic present but not uniquely determined; * = broken or regenerated tail. Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Pauwels et al. 2004Pauwels et al. , 2013Pauwels et al. , 2014Pauwels et al. , 2018Nguyen et al. 2010;Ngo & Pauwels 2010;Sumontha et al. 2010Sumontha et al. , 2012Sumontha et al. , 2014Sumontha et al. , 2015Shi & Zhao 2010;Chan-ard & Makchai 2011;David et al. 2011;Schneider et al. 2011;Grismer et al. 2012Grismer et al. , 2018Johnson et al. 2012;Kunya et al. 2014Kunya et al. , 2015Luu et al. 2014Luu et al. , 2015Luu et al. , 2016Luu et al. , 2017Nazarov et al. 2014;Panitvong et al. 2014 King 1962;Das 1993;Günther & Rösler 2002;Rösler et al. 2007;Rösler & Glaw 2008;Oliver et al. 2009;Welton et al. 2009Welton et al. , 2010Iskander et al. 2011;Kathriner et al. 2014;Riyanto et al. 2014Riyanto et al. , 2015Riyanto et al. , 1016Riyanto et al. , 2018Harvey et al. 2015;Hartmann et al. 2016;, Riyanto et al. 2018from Uetz et al. 2019). The abbreviations are as follows: / = data not available in literature; -=; characteristic not present; + = characteristic present but not uniquely determined; * = broken or regenerated tail. ...
We describe a new species of Cyrtodactylus on the basis of four specimens collected from Phu My, Binh Dinh Province, southern Vietnam. Cyrtodactylus phumyensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small (SVL up to 66.8 mm); two internasals; dorsal tubercle rows 18 or 19 at midbody; ventral scale rows 33-41; ventrolateral folds slightly developed; each thigh with 5-7 enlarged femoral scales; femoral pores absent in males and female; a series of 5-7 precloacal pores plus a pitted, enlarged precloacal scale in males; 6 pitted, enlarged precloacal scales in female; paravertebral tubercles 20-23; lamellae under toe IV 18-21; small subcaudal scales, not transversely enlarged; two irregular dark longitudinal stripes on shoulder. In phylogenetic analyses, the new species is recovered as a member of the Cyrtodactylus irregularis species group, and strongly supported as a sister taxon of C. cucdongensis from Khanh Hoa Province.
... Recent work on Cyrtodactylus cave geckos has revealed 18 species (e.g. Bauer et al. 2009;Ellis & Pauwels 2012;Sumontha, Panitvong & Deein 2010) that have been found in or near caves. Thailand also hosts the smallest bat species in the world (Craseonycteris thonglongyai), not much larger than a bumble-bee (Duangkhae 1991;Puechmaille et al. 2009) which is restricted to Kanchanaburi and neighbouring regions of Myanmar, and at least another 68 species of bat have been recorded from caves in the country. ...
... These new species discoveries are further confirmation of the highly biodiverse, and poorly inventoried, nature of southern Myanmar's forests. The genus Cyrtodactylus is a species-rich group of tropical gekkonid lizards that currently includes greater than 200 described species [32], many of which were recently discovered in Southeast Asia (e.g., [33,34,35]). These species, and closely related Cnemaspis species, are often restricted to isolated karst formations or have limited geographic distributions, which increases their vulnerability to local habitat loss or alteration [36]. ...
Myanmar's recent transition from military rule towards a more democratic government has largely ended decades of political and economic isolation. Although Myanmar remains heavily forested, increased development in recent years has been accompanied by exceptionally high rates of forest loss. In this study, we document the rapid progression of deforestation in and around the proposed Lenya National Park, which includes some of the largest remaining areas of lowland evergreen rainforest in mainland Southeast Asia. The globally unique forests in this area are rich in biodiversity and remain a critical stronghold for many threatened and endangered species, including large charismatic fauna such as tiger and Asian elephant. We also conducted a rapid assessment survey of the herpetofauna of the proposed national park, which resulted in the discovery of two new species of bent-toed geckos, genus Cyrtodactylus. We describe these new species, C. lenya sp. nov. and C. payarhtanensis sp. nov., which were found in association with karst (i.e., limestone) rock formations within mature lowland wet evergreen forest. The two species were discovered less than 35 km apart and are each known from only a single locality. Because of the isolated nature of the karst formations in the proposed Lenya National Park, these geckos likely have geographical ranges restricted to the proposed protected area and are threatened by approaching deforestation. Although lowland evergreen rainforest has vanished from most of continental Southeast Asia, Myanmar can still take decisive action to preserve one of the most biodiverse places on Earth.
... The Bent-toed Geckos (Cyrtodactylus Gray) constitute the most species rich reptile genus in tropical Asia, with over 200 described species (Nazarov et al. 2014;Uetz 2015). The majority of these have been described in the last decade, and many are limited to particular substrate types, such as caves and karst landscapes (e.g., Bauer et al. 2002Bauer et al. , 2009Bauer et al. , 2010Ngo 2008;Sumontha et al. 2010;Schneider et al. 2011;Ellis & Pauwels 2012;Pauwels et al. 2014a,b;Grismer et al. 2014aGrismer et al. , 2015. This has resulted in widespread microendemism and suggests that continued herpetological exploration throughout the range of the genus, but especially in south-east Asia, will continue to reveal additional species. ...
A new Bent-toed Gecko, Cyrtodactylus phetchaburiensis sp. nov. is described from the Tha Yang District of Phetchaburi Province, western Thailand. It is a medium-sized Cyrtodactylus (SVL to at least 63.2 mm), with small, mostly keeled tubercles in 20 regular longitudinal rows on dorsum; 33 scales across mid-venter between lowest rows of flank tubercles; enlarged row of femoral scales present; five precloacal pores in male, femoral pores and precloacal groove absent; 5–6 broad basal lamellae and 11 narrow distal lamellae beneath digit IV of pes; and a single median row of transversely enlarged subcaudal scales present. It has a dorsal colour pattern of large, dark, diffusely-edged markings on a fawn background and a pair of dark scapular patches. The species is a member of the Central Indochinese (Thai-Myanmar) clade of Cyrtodactylus and is most closely related to C. oldhami (Theobald), from which it differs in colour pattern.
... In many areas of Khammouane Province in central Laos, the karst mountains are isolated hills and outcrops that might have influenced Cyrtodactylus speciation (Bauer et al. 2009;Ellis & Pauwels 2012), especially within the C. phongnhakebangensis species group. For example, C. jaegeri and C. soudthichaki have been discovered at two karst localities only a few kilometres apart. ...
We describe a new species of the gekkonid genus Cyrtodactylus from Khammouane Province, central Laos based on morphological and molecular data. Morphologically, Cyrtodactylus rufford sp. Nov. differs from its congeners by a unique combination of the following characters: medium size, SVL reaching 72.5 mm; dorsal pattern with three or four light transverse bands between limb insertions; one intersupranasal; 14-16 irregular dorsal tubercle rows at midbody, weakly developed in the paravertebral region; 27-29 ventral scale rows between ventrolateral folds; 42-43 precloacal and femoral pores in a continuous row in males, enlarged femoral and precloacal scales present; 4 or 5 postcloacal tubercles on each side; dorsal tubercles present at base of tail; medial subcaudal scales enlarged. Molecular analyses show that the new species is closely related to C. khammouanensis, which was originally described from Khammouane Province.
... No Cyrtodactylus has been so far reported. Pursuing our taxonomic work and field surveys on this genus, that have already brought to light the existence of several micro-endemics in the mountains of northern Thailand (Bauer et al. 2009, 2010, Ellis & Pauwels 2012, we visited Doi Inthanon and encountered a population that is distinct from all known congeners in scalation and color pattern. This population is consequently described hereafter as a new species, Cyrtodactylus inthanon sp. ...
We describe a new forest-dwelling Cyrtodactylus from Doi Inthanon, Chiang Mai Province, northern Thailand. Cyrtodactylus inthanon sp. nov. is characterized by a maximum known SVL of 87.3 mm; 18 to 20 longitudinal rows of dorsal tubercles; a continuous series of 34 to 37 enlarged femoro-precloacal scales, including four to six pitted (female) or porebearing (male) scales on each femur separated by a diastema from five pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and three to five irregular beige dorsal bands between limb insertions. The discovery of a new reptile endemic to Doi Inthanon reinforces the high importance of this mountain in terms of biodiversity conservation.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008, Rösler & Glaw 2008, Bauer et al. 2009, Ngo & Pauwels 2010, Ziegler et al. 2010, David et al. 2011, Iskandar et al. 2011, Ngo 2011. ...
We describe a new species of the genus Cyrtodactylus based on two adult specimens from Huong Son limestone forest, Hanoi, Vietnam. Cyrtodactylus huongsonensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: medium-sized, with a maximum SVL of 89.8 mm; dorsal pattern consisting of dark nuchal loop, neck band and five in part irregular transverse body bands between limbs; two enlarged lateral chin-shields in contact with first postmental pair; dorsal tubercles present on occiput, body, forearms, hind limbs and tail base; 14–16 irregularly running dorsal tubercle rows; ventrals in 41–48 longitudinal rows at midbody; lateral skin folds present, without interspersed tubercles; six precloacal pores plus in total 17 femoral pores in males, which are separated by 8–12 poreless scales; enlarged femoral scales and precloacal scales present; three postcloacal spurs in males; subcaudal scales transversely enlarged. This is the 24 th species of Cyrtodactylus known from Vietnam.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Smith 1920Smith , 1921Smith , 1935Taylor 1963;Ulber & Grossmann 1991;Ulber 1993;Darevsky & Szczerbak 1997;Manthey & Grossmann 1997;Ziegler et al. 2002;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Ngo & Grismer 2006;Nguyen et al. 2006;Grismer et al. 2007;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Grismer & Norhayati 2008;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Bauer et al. 2009;Geissler et al. 2009). In comparison with Vietnamese congeners, Cyrtodactylus roesleri sp. ...
We describe a third cryptic species of the genus Cyrtodactylus from Phong Nha – Ke Bang National Park, Quang Binh Province, Vietnam. Cyrtodactylus roesleri sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small, with a maximum SVL of 75.3 mm; dorsal pattern consisting of dark nuchal band and 4–5 sometimes irregularly shaped dark transversal bands between limbs; ventrals in 34–40 longitudinal rows at midbody; continuous series of 20–28 precloacal and femoral pores in males; subcaudal scales transversally enlarged. It is the 15 th new reptilian taxon described as new from the karst forests of Phong Nha – Ke Bang National Park in the past decade. In addition, it is the 19 th species of Cyrtodactylus known from Vietnam and the third Cyrtodactylus occurring in sympatry in the karst forests of Phong Nha – Ke Bang National Park.
... nov. does however not show obvious affinities with other species or species complexes found in the Thai-Malay border area, cave-dwelling or not, so we have extended our comparisons to include all species known from Peninsular Malaysia, Thailand and Myanmar (see the updated lists, revisions and recent species comparisons in Ulber & Grossmann 1991;Bauer et al. 2003Bauer et al. , 2009Bauer et al. , 2010David et al. 2004;Grismer & Ahmad 2008;Mahony 2009;Chan-ard & Makchai 2011;Grismer 2011;Ellis & Pauwels 2012;Grismer et al. 2012Grismer et al. , 2013Grismer et al. , 2014aJohnson et al. 2012;Sumontha et al. 2012b;. ...
We describe a new Cyrtodactylus, C. wangkulangkulae, from Manang District, Satun Province, southern Thailand, based on an adult female having a snout-vent length of 73.9 mm, a long tail (1.34 times snout-vent length), a complex blotched and banded dorsal pattern with four irregularly shaped dark bands between limbs insertions, bluish-grey iris, discontinuous series of enlarged poreless femoral and precloacal scales, 10 regularly arranged dorsal tubercle rows, 38 ventral scale rows between ventrolateral skin folds, transversely enlarged subcaudal plates, and no precloacal groove.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Smith 1920Smith , 1921Smith , 1935Ulber & Grossmann 1991;Darevsky & Szczerbak 1997;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009David et al. 2004David et al. , 2011Pauwels et al. 2004Pauwels et al. , 2013Nguyen S.N. et al. 2006Nguyen S.N. et al. , 2013Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Mahony 2009;Ngo & Chan 2010;Ngo & Pauwels 2010;Ngo et al. , 2011Nguyen T.Q. et al. 2010;Sumontha et al. 2010;Schneider et al. 2011;Grismer et al. 2012;Nazarov et al. 2012;Ziegler et al. 2010Ziegler et al. , 2013Luu et al. 2014;. ...
We describe a new species of the genus Cyrtodactylus from southern Vietnam, based on morphological and molecular datasets. Cyrtodactylus cucdongensis sp. nov. is described on the basis of seven specimens collected from Cuc Dong Cape, Khanh Hoa Province. The new species can be distinguished from the remaining bent-toed geckos by a combination of the following characters: maximum SVL 65.9 mm; 16–19 dorsal tubercle rows; 41–44 ventral scales at midbody; 5 or 6 precloacal pores in males, 4–6 pitted precloacal scales in females; no femoral pores; 6–13 enlarged precloacal scales; 5–9 enlarged femoral scales; no transversally enlarged subcaudals; dorsal pattern consisting of irregular dark bands. This is the 33rd species of Cyrtodactylus known from Vietnam.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Rösler & Glaw 2008;Bauer et al. 2009Bauer et al. , 2010Ngo & 36), C. pulchellus Gray 1827 (33-39), C. redimiculus King 1962 (5-8+8-9), C. seribuatensis Youmans & Grismer 2006 (40-44), C. surin Chan-ard & Makchai, 2011 (34), C. tamaiensis Mahony 2009 (40), C. trilatofasciatus Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012 (41-46), C. variegatus (Blyth 1861) (32) and C. wetariensis (Dunn 1927) (11-12+12-13). ...
We describe the third new species of the genus Cyrtodactylus from Ba Den Mountain, Tay Ninh Province, southern Vietnam based on a type series consisting of eleven specimens. Cyrtodactylus thuongae sp. nov. is distinguished from the remaining Indochinese Bent-toed Geckos by a combination of the following morphological characters: size medium, with a maximum SVL of 77.6 mm; dorsal pattern consisting of blotches between limbs and on occiput and a sometimes medially opened nuchal loop; dorsal tubercles in 16–18 irregular rows; ventrals in 29–44 longitudinal rows at midbody; lateral skin folds present, lacking tubercles; femoral pore series separated from precloacal pore series in males, 0–3 on each side; precloacal pores in males absent or only one; enlarged femoral scales and precloacal scales present, separated by a series of 15–16 smaller scales; postcloacal spurs two or three; subcaudal scales slightly enlarged. This is the 32nd described Cyrtodactylus species from Vietnam.
... nov. with all congeneric species known from Cambodia, China, Laos, Myanmar, Peninsular Malaysia, Thailand and Vietnam (see latest descriptions and intraspecific comparisons in Bauer et al. 2009Bauer et al. , 2010Geissler et al. 2009;Ngo and Chan 2010;Ngo and Pauwels 2010;Nguyen et al. 2010;Chan-ard and Makchai 2011;David et al. 2011;Grismer 2008Grismer , 2011Grismer et al. 2012;Johnson et al. 2012;Ngo and Grismer 2012;Sumontha et al. 2010Sumontha et al. , 2012Pauwels et al. 2013;Ziegler et al. 2013). Cyrtodactylus khelangensis sp. ...
We describe Cyrtodactylus khelangensis sp. nov. from a limestone cave in Pratu Pha, Mae Mo District, Lampang Province, northern Thailand. It is characterized by a maximal known SVL of 95.3 mm; 16-20 longitudinal rows of dorsal tubercles; a continuous series of 37-40 enlarged femoroprecloacal scales, including six-seven pitted or pore-bearing scales (males) or one or two pitted scales (females) on each femur separated by a diastema from 2-6 pore-bearing precloacal scales (males and females); no precloacal groove nor depression; transversely enlarged subcaudal scales; and four irregular brown dorsal bands between nuchal loop and hind limb insertions.
... Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008;Rösler & Glaw 2008;Bauer et al. 2009;Ngo & Pauwels 2010;Sumontha et al. 2010;Ziegler et al. 2010;David et al. 2011;Iskandar et al. 2011;Luu et al. 2011;Ngo 2011;Ngo & Chan 2011;Schneider et al. 2011;Nazarov et al. 2012;Ngo & Grismer 2012). ...
We describe a new species of the genus Cyrtodactylus based on five adult specimens from Dai Lanh Cape, Tuy Hoa District, Phu Yen Province, southern Vietnam. Cyrtodactylus kingsadai sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: maximum SVL of 94 mm; dorsal pattern consisting of a dark nuchal loop, continuous or partly interrupted neck band and four in part irregulartransverse body bands between limbs; internasal single; dorsal tubercles in 17–23 irregular transverse rows; ventrals in 39–46 longitudinal rows at midbody; lateral skin folds present, without interspersed tubercles; precloacal pores 7–9 plus in total 3–7 femoral pores in males (1-4 femoral pores on each side) with precloacal and femoral pore series separated from each other by 7–9 poreless scales; enlarged femoral scales and precloacal scales present; postcloacal spurs three;subcaudal scales transversely enlarged. This is the 29th species of Cyrtodactylus known from Vietnam.
... Comparisons were made with material housed in the Zoological Collection of the University of Natural Sciences, Hochiminh City, as well as with original published descriptions, data, and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Brown & Parker 1973;Ulber & Grossmann 1991;Ulber 1993;Darevsky & Szczerbak 1997;Das 1997Das , 2006Das & Lim 2000;Ziegler et al. 2002Ziegler et al. , 2010Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Batuwita & Bahir 2005;Grismer 2005;Grismer & Leong 2005;Kraus & Allison 2006;Nguyen et al. 2006;Youmans & Grismer 2006;Grismer et al. 2007Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Grismer & Ahmad 2008;Hayden et al. 2008;Kraus 2007Kraus , 2008Rösler et al. 2007Linkem et al. 2008;Nazarov et al. 2008;Rösler & Glaw 2008;Ngo 2008;Ngo & Bauer 2008;Oliver et al. 2008Oliver et al. , 2009Geissler et al. 2009;Mahony 2009;Welton et al. 2009Welton et al. & 2010 Paratypes. UNS 0527, subadult female, was collected by Ngo Van Tri between 19h00 and 20h00 on 26 April 2010 at the entrance of Lomyen Cave. ...
A new cave-dwelling bent-toed gecko, Cyrtodactylus lomyenensis sp. nov. is described from a karst forest in Gnommalath District, Khammouane Province, southern Laos. It differs from all other species of Indochinese-Thai Cyrtodactylus in the following combination of characters: maximum SVL of at least 71.2 mm; head dorsum yellowish with irregular brown blotches; presence of a brown nuchal loop reaching the posterior edge of the orbit; four narrow yellowish-cream transversal bands with irregular anterior and posterior black edges on a brown background between limb insertions; no precloacal groove; 39-40 precloacal-femoral pores in males, arranged in a continuous row; females with 32 precloacal-femoral pores in a continuous row, smaller than those of males; five postanal tubercles on each side; 16-18 subdigital lamellae on first toe; 19-23 subdigital lamellae on 4th toe; no tubercles on tail dorsum; and a median row of enlarged subcaudal scales.
A new cave-dwelling species of the gekkonid lizard genus Cyrtodactylus Gray, C. auribalteatus sp. nov. is described on the basis of animals collected from a cave in Thung Salaeng Luang National Park, Phitsanulok Province, Thailand, bringing the number of species in this genus in Thailand to 18. The new species shares over all appearances with other cave-dwelling Cyrtodactylus spp. in having large eyes, a long snout, reduced tubercles, absence of caudal tubercles, and slender limbs for climbing. The new species is characterized by its moderate size (snout-vent length to at least 98.07 mm), weakly conical tubercles in 22–24 rows at midbody, high number of ventral scale across midbody (38–40), absence of precloacal groove, present of a single series of 6 precloacal pores and 4–5 femoral pores on each thigh separated by a diastema, broad subcaudal plates, and its dorsal pattern of three dark bands between limb insertions. It is the seventh species of cave-dwelling Cyrtodactylus, including the sandstone cave species, C. jarujini, recorded from Thailand.
A new species of Bent-toed Gecko, Cyrtodactylus gunungsenyumensis sp. nov. of the sworderi complex, is described from Hutan Lipur Gunung Senyum, Pahang, Peninsular Malaysia and is differentiated from all other species in the sworderi complex by having a unique combination of characters including a maximum SVL of 74.7 mm; low, rounded, weakly keeled, body tubercles; 34-40 paravertebral tubercles; weak ventrolateral body fold lacking tubercles; 38-41 ventral scales; an abrupt transition between the posterior and ventral femoral scales; 20-23 subdigital lamellae on the fourth toe; enlarged femoral scales; no femoral or precloacal pores; no precloacal groove; wide caudal bands; and an evenly banded dorsal pattern. Cyrtodactylus gunungsenyumensis sp. nov. is a scansorial, karst forest-adapted specialist endemic to the karst ecosystem surrounding Gunung Senyum and occurs on the vertical walls of the limestone towers as well as the branches, trunks, and leaves of the vegetation in the associated karst forest. Cyrtodactylus gunungsenyumensis sp. nov. is the seventh species of karst forest-adapted Cyrtodactylus and the sixteenth endemic species of karst ecosystem reptile discovered in Peninsular Malaysia in the last seven years from only 12 different karst forests. This is a clear indication that many species remain to be discovered in the approximately 558 isolated karst ecosystems in Peninsular Malaysia not yet surveyed. These data continue to underscore the importance of karst ecosystems as reservoirs of biodiversity and microendemism and that they constitute an important component of Peninsular Malaysia's natural heritage and should be protected from the quarrying interests of foreign industrial companies.
We describe a new species of the gekkonid genus Cyrtodactylus from northern Laos, based on four specimens fro Luang Nam Tha Province. Cyrtodactylus wayakonei sp. n. is distinguished from the remaining bent-toed-geckos by the combination of the following characters: snout-vent length up to 90 mm; 7-8 supralabials; 9-10 infralabials; rostrum bearing smooth scales; 34-45 scales between fifth supralabials; dorsal tubercles smooth or slightly keeled; 17-19 dorsal tubercle rows; ventrolateral folds slightly developed, with large tubercles; 31-35 ventrals; 85-98 scales around midbody; 151-163 scales between mental and cloacal slit; 6-8 precloacal pores in both sexes; precloacal groove lacking; enlarged femoral scales absent; femoral pores lacking; subcaudals somewhat enlarged, broadened; tail tubercles flat, smooth; head and dorsum with blotched to reticulated pattern.
A new cave-dwelling species of the gekkonid lizard genus Cyrtodactylus Gray, C. auribalteatus sp. nov. is described on the basis of animals collected from a cave in Thung Salaeng Luang National Park, Phitsanulok Province, Thailand, bringing the number of species in this genus in Thailand to 18. The new species shares over all appearances with other cave-dwelling Cyrtodactylus spp. in having large eyes, a long snout, reduced tubercles, absence of caudal tubercles, and slender limbs for climbing. The new species is characterized by its moderate size (snout-vent length to at least 98.07 mm), weakly conical tubercles in 22-24 rows at midbody, high number of ventral scale across midbody (38-40), absence of precloacal groove, present of a single series of 6 precloacal pores and 4-5 femoral pores on each thigh separated by a diastema, broad subcaudal plates, and its dorsal pattern of three dark bands between limb insertions. It is the seventh species of cave-dwelling Cyrtodactylus, including the sandstone cave species, C. jarujini, recorded from Thailand.
We describe a new species of the gekkonid genus Cyrtodactylus from northwestern Laos, based on a recent collection from Vientiane Province. Cyrtodactylus pageli sp. n. is distinguished from the remaining bent-toed-geckos by the combination of the following characters: snout-vent length up to 81.8 mm; 9-12 supralabials; 9 infralabials; 9-14 rows of dorsal tubercles; no distinct lateral body fold; 41-46 midbody ventrals; precloacal pores present in both sexes (males 4, females 5 or 6); no precloacal groove; femoral scales not distinctly enlarged; no femoral pores; subdigital lamellae under the fourth toe 19-26; subcaudals transversally enlarged; dark nuchal band, followed by a transversal row of dark neck blotches (rarely forming an additional transversal neck band) in-between nuchal band and first body band; 5 in part irregular dark dorsal bands between limb insertions plus transversal band between hind limbs; tail with banded pattern.
We describe a new species of the gekkonid genus Cyrtodactylus on the basis of two specimens collected from limestone forests of Luang Prabang Province, northern Laos. Morphologically, the new species is distinguishable from its congeners by a combination of the following diagnostic characters: maximum SVL 86.1 mm; supralabials 9 or 10; infralabials 7–9; dorsal tubercles in 15 or 16 rows at midbody; ventral scale rows 34–36 at midbody; precloacal groove absent; femoral scales not distinctly enlarged; precloacal pores absent in females (unknown in males); subdigital lamellae under the fourth finger 18 or 19, under the fourth toe 18–20; subcaudals not transversally enlarged; dorsal bands white, 4 or 5 between limb insertions plus another one between hind limbs; tail banded. Based on molecular analyses, the new species is clustered in the same clade with C. wayakonei and two other species from Luang Prabang and Houaphan provinces.
We describe a new cave-dwelling Bent-toed Gecko, Cyrtodactylus kunyai (Squamata: Gekkonidae) from Suan Hin Pha Ngam, Nong Hin District, Loei Province, northeastern Thailand. It is characterized by a maximal known snout-vent length of 87.9 mm, a banded dorsal pattern with a medially interrupted nuchal loop and four of five brown bands between nuchal loop and hind limb insertions and three bicolored band interspaces between limb insertions, a dark orangeish iris, a continuous series of enlarged femoro-precloacal scales with 5-6 femoral pores on each side separated by a diastema from 3 precloacal pores in males (no pores in females), 19 irregularly arranged dorsal longitudinal tubercle rows at midbody, 34 ventral scale rows between ventrolateral skin folds, transversely enlarged subcaudal plates, and no precloacal groove.
We describe a new forest-dwelling Cyrtodactylus (Gekkonidae) from Doi Suthep, Chiang Mai Province, northern Thailand. Cyrtodactylus doisuthep sp. nov. is characterized by a maximal known SVL of 90.5 mm; 19 or 20 longitudinal rows of dorsal tubercles; a continuous series of 34 or 35 enlarged femoro-precloacal scales, including six or seven pitted scales on each femur (male and females) separated by a diastema from six pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and six or seven irregular thin beige dorsal bands between limb insertions.
We describe a new species of the genus Cyrtodactylus based on two adult specimens from Khammouane Province, Laos. Cyrtodactylus jaegeri sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: a moderately sized Cyrtodactylus with a maximum SVL reaching 68.5 mm; dorsal pattern consisting of a dark nuchal loop and four narrow brown body bands between limb insertions; dorsal tubercles in 15-17 irregular rows; ventrals in 31-32 longitudinal rows at midbody; lateral skin folds present with interspersed tubercles; precloacal-femoral pores 44 in the male, in a continuous series; enlarged femoral scales and precloacal scales present; postcloacal tubercles 3-6; subcaudals transversely enlarged. Cyrtodactylus jaegeri sp. nov. is the ninth species of Cyrtodactylus known from Laos.
Species of bent-toed gecko (Cyrtodactylus) in Vietnam have been described at a rate of nearly four species per year since 2007 mostly based on morphological data. A tool that guides species delimitation will accelerate the rate of documentation, and at a time when the recognition of species greatly benefits conservation. We use DNA barcoding using COI (550 bp) to re-examine the levels of genetic divergence and taxonomic status of 21 described species of Vietnamese bent-toed geckos. Tree-based analyses resolve all sampled species and identify potential undescribed taxa. Kimura 2-parameter genetic distances between the described species average 21.0±4.2% and range from 4.3% to 28.7%. Further, our analyses discover two potentially new species from Vietnam, two from Laos and one from China. Herein we describe the new species Cyrtodactylus puhuensis sp. nov. from Vietnam on the basis of both genetics and morphology. Genetically, it differs from the remaining species by an average K2P distance of 24.0±1.8%. Morphologically, the new species is diagnosed by its medium-size (snout-vent length 79.24 mm and tail length 82.59 mm, for the single known individual), in having a series of moderately enlarged transverse subcaudals and a series of moderately enlarged femoral scales that extend from precloacal scales, in possessing femoral scales without pores, with males having five precloacal pores, and in exhibiting 8 supralabials, 10 infralabials, 23 narrow subdigital lamellae on its fourth toe, and 36 transverse ventrals.
A new species of Cyrtodactylus is described from the Truong Son (Annamite mountain range) of Quang Binh Province in central Vietnam. It is characterized by 3–5 transversal dorsal bands between the limb insertions, a neckband reaching the posterior margin of eye, 19–20 irregular longitudinal rows of dorsal tubercles, forelimbs with tubercles, 47–50 longitudinal rows of ventrals at midbody, 9–11 precloacal pores in an angular series in males and 16–27 enlarged precloacal scales in both sexes, 20–23 lamellae below the 4th toe, a segmented tail, and lacking of a distinct ventrolateral fold, a precloacal groove and femoral pores, as well as transversally enlarged subcaudal plates. The new species is known only from the karst forests of Phong Nha - Ke Bang National Park. Preliminary data on its natural history are provided.
Three new species of the gekkonid genus Cyrtodactylus are described from Peninsular Malaysia; C. jarakensis sp. nov. from Pulau Jarak off the west coast of Perak, C. batucolus sp. nov. from Pulau Besar of the Water Islands Archipelago off the west coast of Melaka, and C. pantiensis sp. nov. from the Gunung Panti Forest Reserve, Johor. All are distinguishable from other Sunda Shelf species of Cyrtodactylus in having unique combinations of morphological and color pattern characteristics. The continued high potential for finding additional new species in unexplored regions of southern Peninsular Malaysia and its associated archipelagos is further supported with these findings.
A new species of the gekkonid lizard, Cytrodactylus macrotuberculatus sp. nov., is described from Pulau Langkawi of the Langkawi Archipelago located off the northwest coast of Peninsular Malaysia in the state of Kedah. This species had been previously recognized as C. pulchellus but is differentiated here from that species and all other Sunda Shelf Cyrtodactylus on the basis of its strong tuberculation and other scale and color pattern characteristics. This represents the second endemic species known from the Langkawi Archipelago.
A new species of large Cyrtodactylus is described from lowland rainforest on Batanta Island in the Raja Ampat Archipelago, Papua Barat Province, Indonesian New Guinea. The new species can be distinguished from all other Melanesian Cyrtodactylus by the combination of large size (over 110 mm SVL), very robust build, presence of enlarged ventral tubercles below the lateral fold and around the angle of the lower jaw only, and dorsal colouration consisting of three to four irregular dark greyish-brown blotches. It is the second species of Cyrtodactylus known with certainty only from the Raja Ampat Islands. The morphology of the new species places it within the C. loriae group and suggests that it is closely related to Cyrtodactylus irianjayaensis.
A new species of Cyrtodactylus is described from Pulau Tioman, an island off the east coast of Peninsular Malaysia. The new species can be differentiated from congeners from south-east Asia in showing the following combination of characters: medium-sized Cyrtodactylus (SVL to 83.2 mm); rostral partially divided by rostral groove, contacted posteriorly by two nostrils and two semi-circular supranasals; pectoral and abdominal scales smooth, rounded, semi-circular, imbricate; no preanal groove, a distinct preanal depression; sharp boundary between small scales on posterior surface of thighs and larger ones on ventral surface of thighs; tail without lateral denticles or tubercles forming whorls or segments; supralabials (to midorbit position) 8-11; infralabials 9-11; midventral scale rows at belly to lowest row of tubercles 36-40; lamellae under toe IV 20-22; preanofemoral pores 19; and dorsal pattern comprises four pale yellow transverse bands, each narrower than the intervening pale brown areas, edges with dark brown, and a pale yellow nuchal loop joining posterior edges of eyes.
A new species of bent-toed gecko, Cyrtodactylus adleri, is described, based on seven examples from the island of Great Nicobar, India. Although reported as conspecific with C. rubidus, it can be differentiated from the Andamanese species in possessing dark spots (vs. dark bands) on the dorsum; 48-50 imbricate (vs. 38-42 juxtaposed) midventrals; six (vs. four) postnasals; two (vs. three) postcloacal tubercles; and presence of a preanal groove. In the lack of a preanal groove, imbricate ventrals, and dark spots on dorsum, the new species appears close to the poorly known Sumatran endemic, C. Interalis, from which is may be differentiated in its small body size (SVL 53.5-64.7 mm in four adult females, 61.8-68.5 mm in three adult males), dark spots on dorsum, and 48-50 midventrals.
This paper describes a new species of the genus Cyrtodactylus. Cyrtodactylus chauquangensis sp. nov. occurs in the karst of Quy Hop District, Nghe An Province (North Central Vietnam). It can be distinguished from all other Cyrtodactylus species occurring in Vietnam by a combination of characters: the number of pre-anal pores, the presence or absence of femoral pores or larger femoral scales, scales across the abdomen in the middle of the body, interorbital scales, snout-vent length, proportions of the body and color pattern.
A new species of Cyrtodactylus of Gekkonidae family, assigned to the Cyrtodactylus irregularis complex on the basis of twelve specimens collected from the Chu Yang Sin National Park, Krong Bong District, Dac Lac Prov-ince, central Vietnam is described. Forms of C. irregularis complex inhabiting an extensive territory, covering the southern part of the Annamite Mountains and Central Highlands, their distribution and variation are dis-cussed; a detailed description of the type specimens of C. irregularis complex and a description of a new species are given.
This paper describes a new species of the genus Cyrtodactylus. Cyrtodactylus caovansungi sp. nov. occuring in the Ninh Thuan Province (South Vietnam). It can be distinguished from all other Cyrtodactylus species distributed in Vietnam by a male with well developed nine V-shaped preanal pores. There are 8 enlarged femoral scales on each side; however femoral pores are present only on three of them, located closer to the knee bend. Roundish-triangular dorsal tubercles with well keels are placed at 16 – 18 not regular longitudinal rows.
A new species of Cyrtodactylus is described from Sulawesi and the adjacent island of Kabaena, Indonesia. This new species is the largest Cyrtodactylus known from Sulawesi and differs from all other congeners by the absence of precloacal and femoral pores, enlarged femoral scales or pubic groove; medium to large size (up to 113.6 mm snout-vent length) and dark purple dorsum with irregular dark bands and yellow spots. Only two other species of Cyrtodactylus are currently described from Sulawesi and surrounding islands. Current taxonomy of the genus does not accurately reflect the diversity of Cyrtodactylus in Sulawesi. This species is one of several taxa new to science currently being described from the region.
ÐA specimen of the gekkonid genus Cyrtodactylus Gray, 1827, collected in Xepian National Biodiversity and Conservation Area, Champasak Province, southern Laos, is here referred to a new species, Cyrtodactylus buchardi sp. nov. The holotype, a juvenile male, differs from other known species by the combination of a slender body, a barely visible lateral fold, a slender but short tail, only 12 subdigital lamellae beneath 4 th toe, 25 rows of tuberculate dorsal scales, no precloacal groove, three series of enlarged precloacal scales, no enlarged femoral scales, no distinctly enlarged subcaudal scales, 13-14 supralabials, a dorsal pattern made of five transversal series of irregular blotches, and a nuchal collar not reaching the posterior margin of the eyes. The new species is compared with other species known from the Indo-Chinese Peninsula and Thailand. It appears to be related to Cyrtodactylus angularis (Smith, 1921) and C. papilionoides Ulber & Grossmann, 1991, both from central and eastern Thailand.
The genus Cyrtodactylus Gray, 1827 has been considered to be represented in Sri Lanka by a single widely distributed species, C. fraenatus (Günther, 1864). A survey of rainforest sites in the island's wet zone shows that at least six species are present, five of them new. Cyrtodactylus cracens and C. subsolanus were recorded from the western and eastern sides respectively, of the Sinharaja World Heritage Site; C. edwardtaylori, from a refuge at Namunukula; C. ramboda, from the north-western slopes of the central hills; and C. soba from the Knuckles mountains. Cyrtodactylus fraenatus sensu stricto, restricted to the hills around Kandy, is redefined based on one of its syntypes and fresh collections.
A new cave-dwelling species of Cyrtodactylus is described from Khao Sonk in Surat Thani Province, southern peninsular Thailand. The new species is characterized by its slender body (snout-vent length to at least 80 mm) with a banded pattern, elongate tail and digits, low number (14) of rows of dorsal tubercles, absence of precloacal groove, and enlarged series of precloacal and femoral scales, all lacking pores. It is the third species of cave-dwelling Cyrtodactylus recorded from Thailand.
A new species of the gekkonid lizard genus Cyrtodactylus Gray, 1827 is described on the basis of material collected in limestone caves at Khao Wong, Rayong Province, Thailand. Cyrtodactylus sumonthai sp. nov. is characterized by a slender body, elongate tail and digits, small number (12) of rows of dorsal tubercles, and enlarged patch of precloacal scales bearing only two precloacal pores. The new species is the second tropical Asian Cyrtodactylus believed to be largely associated with cave environments and is the twelfth species of the genus thus far recorded from Thailand.
A new species of Cyrtodactylus is described from Lore-Lindu National Park, Sulawesi Island, Indonesia. It is distinguished from all other Cyrtodactylus by a unique suite of scalation characters and a distinctive color pattern. The new species is the fourth Cyrtodactylus known from the island of Sulawesi and one of two new species found in 2004. These recent discoveries suggest that the diversity of the herpetofauna in Wallacea, a poorly studied biological ''hotspot,'' may be far richer than previously thought. THE GENUS Cyrtodactylus contains 95 de-scribed species distributed throughout the Indo-Australian Archipelago westward to In-dia (Bauer and Henle, 1994). Although many species recently have been reassigned to other genera such as Tenuidactylus, Cyrtopodion, Nactus, and Geckoella (Golubev and Szczer-bak, 1985; Kluge, 1983, 1991, 1993, 2001; Macey et al., 2000; Szczerbak and Golubev, 1984, 1986), the number of species currently or formerly in this genus continues to grow. New species have been recently described from Myanmar (Bauer, 2002, 2003), Sri Lanka (Batuwita and Bahir, 2005), Malaysia (Gris-mer, 2005; Grismer and Leong, 2005; You-mans and Grismer, 2006), Thailand (Bauer et al., 2002, 2003; Pauwels et al., 2004), Vietnam (Heidrich et al., 2007; Orlov et al., 2007; Quang et al., 2007; Ziegler et al., 2002), and southern Laos (David et al., 2004). For the island of Sulawesi, Boulenger (1897) and de Rooij (1915) listed three species of Cyrtodactylus: C. fumosus, C. jellesmae, and C. marmoratus. Based on overlap in pore characters, Brongersma (1934) synonomized C. fumosus with C. marmoratus thereby reducing the number of species on the island to two. Hayden et al. (2008) recently de-scribed a third species of Cyrtodactylus from the southwestern peninsula of Sulawesi. Herein we describe a fourth species of Cyrtodactylus from Sulawesi that differs dramatically from all known congeners.
We describe a new gekkonid lizard from Batang Padang, Perak province, Malaysia, based on a single specimen collected almost 100 years ago. Cyrtodactylus stresemanni sp. nov. apparently differs from all other species in the genus by large tubercles on the ventral side of the tail, suggesting an isolated position within the genus. A literature survey of meristic and mensural data of all described Cyrtodactylus species revealed further diagnostic characters to distinguish C. stresemanni from all other species.
The Melinau limestone formation within the Gunung Mulu National Park is highly cavernous, providing an important subterranean habitat which is exploited by numerous invertebrate species and by at least 31 vertebrates, 13 bats, 3 swiftlets, 2 lizards and perhaps 2 fish use the caves as shelter and perhaps as a breeding site, but leave the cave to feed outside. At least 4 mammals and one snake make sporadic visits to the cave in order to eat cavernicoles or bat guano. At least 6 fish species may perhaps be able to sustain permanent populations in the caves; of these, Silurus furnessi may prove to be largely confined to caves. -Author
Two new species of the gekkonid lizard genus Cyrtodactylus Gray, 1827 are described on the basis of voucher specimens collected in isolated mountains separated from the Truong Son Mountain Range, southern Vietnam. Both species were collected from shallow caves, as have been many of the recently described Cyrtodactylus from across Southeast Asia. Both species, C. takouensis sp. nov. from Ta Kou Nature Reserve, Binh Thuan Province and C. huynhi sp. nov. from Chua Chan Mountain, Dong Nai Province, are characterized by a small number of enlarged femoral scales separated from the precloacal scales by a large diastema. They may be distinguished from one another and from all other congeners by differences in rows of tubercles and ventral scales, subcaudal scalation and dorsal color pattern. These new taxa bring the number of currently recognized Cyrtodactylus species in Vietnam to twelve.
A new endemic cave-dwelling species of gekkonid lizard, Cyrtodactylus hontreensis sp. nov., is described from Hon Tre Island in Kien Giang Biosphere Reserve in southwestern Vietnam and differentiated from all other species by a unique suite of the following characters: SVL of 79.7 ± 4.9 mm, three broad transverse dorsal body bands edged in white between limb insertions, very low smooth dorsal body tubercles, the absence of a precloacal groove, 7-8 precloacal pores arranged in a chevron in males, a shallow dorsal groove on the tail, absence of caudal tubercles, and various aspects of color pattern.
Two new cave-dwelling species of Cyrtodactylus from mainland southwestern Vietnam and an offshore island are differentiated from all other congeners in lacking a precloacal groove and precloacal pores, presence of enlarged femoral scales beneath thigh, and in having a color pattern consisting of four or five narrow white bands on the body dorsum and one on the tail. Cyrtodactylus grismeri sp. nov. is reddish brown, has a mean SVL of 87.6 ± 3.8 mm (n = 9), 18-22 irregular longitudinal rows of weakly-keeled tubercles at midbody between the lateral folds, 33-38 ventral scales between ventrolateral folds, 0-3 enlarged scales beneath thighs, and 20-24 subdigital lamellae under the first toe. Cyrtodactylus eisenmani sp. nov. is chocolate brown, has a mean SVL of 81.3 ± 5.0 mm (n = 5), 14 irregular longitudinal rows of weakly keeled tubercles at midbody between ventrolateral folds, 44-45 ventral scales between ventrolateral folds at midbody, 4-6 enlarged femoral scales beneath each thigh, and 22-25 subdigital lamellae under the first toe. This discovery increases the total number of Cyrtodactylus found in Vietnam to fifteen.
We describe a new gecko, Cyrtodactylus murua, from Woodlark Island, off the southeastern tip of
Papua New Guinea. The new species differs from all Papuan congeners except C. louisiadensis (de
Vis) and C. aaroni Günther & Rösler in having a single row of enlarged subcaudal scales. From C.
louisiadensis, the new species is easily distinguished by its color pattern and by having fewer rows
of enlarged dorsal tubercles; from C. aaroni by its larger size and fewer dark dorsal bands. The new
species is almost certainly restricted to Woodlark Island and represents the first endemic reptile
described from that island.
I describe a new gecko, Cyrtodactylus serratus, from the Star Mts. of the western extreme of Papua New Guinea. The new species differs from all Papuan congeners in having a row of enlarged, dentate tubercles extending the length of each lateral skin fold. Further differences with the other species of Papuan Cyrtodactylus are identified and serve to highlight the uniqueness of the new species. I speculate that relationships of the new species may lie closest to C. loriae Boulenger
among named forms, but numerous additional species of Cyrtodactylus remain to be described from New Guinea and this suggestion can only be considered tentative.
As recently understood, the gecko species Cyrtodactylus louisiadensis (de Vis, 1892) ranges across eastern New Guinea and adjacent islands, the Solomon Islands, and northern Queensland. Within that portion of its range encompassed by Papua New Guinea I show that five species are currently masquerading under this appellation. Two of these occur in sympatry at the type locality for the species, Sudest Island. Because the type specimen is lost, the taxonomy of this complex has been confused, and two species are present at the type locality, I designate a neotype for C. louisiadensis so as to stabilize taxonomy within this complex. I describe the four additional species, C. epiroticus sp. nov., C. klugei sp. nov., C. robustus sp. nov., and C. tripartitus sp. nov., which differ from each other in a variety of scalational, mensural, and color-pattern features. Four of these species are confined to single islands off the southeastern peninsula of New Guinea. The fifth occupies the eastern rim of New Guinea.
A new species of Cyrtodactylus is described from lowland forest of southern Peninsular Malaysia. It differs from all other Cyrtodactylus by having a unique suite of characteristics involving coloration and scalation. This is the third species of amphiban or reptile restricted to the southern portion of the Malay Peninsula.
In 1985 Wells and Wellington resurrected from the synonymy of Hemidactylus the Genus Pnoepus of Fitzinger, 1843. As then, I still believe that Hemidactylus is a polyphyletic group (Type Species: Gecko tuberculosus Daudin, 1802 = Hemidactylus mabouia Moreau de Jonnes, 1818) and that all Asiatic members of the frenatus group should be placed in the genus Pnoepus. Additionally, a number of synonyms of frenatus were resurrected by Wells and Wellington (1985) in the belief that frenatus was a species complex in need of major revision. Although it would appear that no one has supported these opinions to date, I still hold the above views so consequently maintain my use of the genus Pnoepus for the populations present in Australia.
A new species of Crytodactylus from Pulau Aur in the Seribuat Archipelago off the southeast coast of Peninsular Malaysia is differentiated from all other Sundaland species in having a unique suite of characteristics involving tuberculation, scale counts, and color pattern. This increases the total number of endemic species in the Seribuat Archipelago to at least 18 and is the second endemic gecko known from Pulau Aur.
Notes on Cyrtodactyline Gekkos from Thailand with Descriptions of Two New Species (Reptilia: Geckonidae)From a small collection of cyrtodactyline geckos made by J. NABHITABHATA in various parts of Thailand, four species are morphologically described. Two of them turned out to be new and are consequently described as Cyrtodactylus interdigitalis sp. nov. and Cyrtodactylus jarujini sp. nov. Whereas in the former it is very obvious that the new species is cladistically related to C. brevipalmatus (SMITH) and C. elok DRING by sharing the feature of interdigital membranes, the relationships of the latter are not quite clear. From its habitus and colouration this new species is similar to C. khasiensis (JERDON) and possibly C. medogensis (ZHAO & LI), but differs considerably from these by having femoral pores. Furthermore, a sample of the poorly known C. oldhami (THEOBALD) with an unusual colour pattern and a single female specimen of C. brevipalmatus (SMITH) are described. For each species ecological data are supplied additionally.
Two new species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar
- A M Bauer
Bauer, A.M. (2002) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar. Proceedings of the
California Academy of Sciences, 53, 75-88.
Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma)
- A M Bauer
Bauer, A.M. (2003) Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of
Myanmar (Burma). Proceedings of the California Academy of Sciences, 54, 463-498.
- A M Bauer
- M Sumontha
- O S G Pauwels
Bauer, A.M., Sumontha, M. & Pauwels, O.S.G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata:
Gekkonidae) from Thailand. Zootaxa, 376, 1-18.
Assignment of currently misplaced Cnemaspis gordongekkoi Das, 1993 (Reptilia: Gekkonidae) to Cyrtodactylus Gray, 1827
- S Biswas
Biswas, S. (2007). Assignment of currently misplaced Cnemaspis gordongekkoi Das, 1993 (Reptilia: Gekkonidae) to
Cyrtodactylus Gray, 1827. Russian Journal of Herpetology, 14, 15-20.
Comments on the taxonomic status of Cyrtodactylus abrae Wells
- P J Couper
- A P Amey
- G M Shea
Couper, P.J., Amey, A.P. & Shea, G.M. (2004) Comments on the taxonomic status of Cyrtodactylus abrae Wells 2002.
Memoirs of the Queensland Museum, 49, 648.
The Caves of Thailand
- J R Dunkley
Dunkley J.R. (1995) The Caves of Thailand. Speleological Research Council, Broadway, Australia, 124 pp.
Slender toes in southern Malaysia
- L L Grismer
Grismer, L.L. (2006) Slender toes in southern Malaysia. Iguana, 13(1), 2-7.
Redescription of the gekkonid lizard Cyrtodactylus sworderi (Smith, 1925) from southern Peninsular Malaysia
- L L Grismer
- P L Wood
- Jr
- T M Youmans
Grismer, L.L., Wood, P.L., Jr. & Youmans, T.M. (2007) Redescription of the gekkonid lizard Cyrtodactylus sworderi
(Smith, 1925) from southern Peninsular Malaysia. Hamadryad, 31, 250-257.
Eine neue Art der Gattung Cyrtodactylus Gray, 1827 aus dem Westen von Neuguinea (Reptilia: Sauria: Gekkonidae)
- R Günther
- H Rösler
Günther, R. & Rösler, H. (2003) Eine neue Art der Gattung Cyrtodactylus Gray, 1827 aus dem Westen von Neuguinea
(Reptilia: Sauria: Gekkonidae). Salamandra, 38, 195-212.
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Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species
- T Hikida
Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of
three new species. Japanese Journal of Herpetology, 13, 91-107.
Eine neue großwüchsige Cyrtodactylus-Art von Neuguinea (Reptilia: Sauria: Gekkonidae)
- H Rösler
Rösler, H. (2001) Eine neue großwüchsige Cyrtodactylus-Art von Neuguinea (Reptilia: Sauria: Gekkonidae).
Zoologische Abhandlungen, Staatliches Museum für Tierkunde Dresden, 51, 61-71.
Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae)
- H Rösler
- S J Richards
- R Günther
Rösler, H., Richards, S.J. & Günther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus
Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria:
Gekkonidae). Salamandra, 43, 193-230.
A new Cyrtodactylus (Squamata: Gekkonidae) from central Vietnam
- H Rösler
- N T Vu
- Q T Nguyen
- V T Ngo
- T Ziegler
Rösler, H., Vu, N.T.,Nguyen, Q.T., Ngo, V.T. & Ziegler, T. (2008) A new Cyrtodactylus (Squamata: Gekkonidae) from
central Vietnam. Hamadryad, 33, 48-63.