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A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus

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Abstract

We describe a new gekkonid lizard from Batang Padang, Perak province, Malaysia, based on a single specimen collected almost 100 years ago. Cyrtodactylus stresemanni sp. nov. apparently differs from all other species in the genus by large tubercles on the ventral side of the tail, suggesting an isolated position within the genus. A literature survey of meristic and mensural data of all described Cyrtodactylus species revealed further diagnostic characters to distinguish C. stresemanni from all other species.

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... The lizard genus Cyrtodactylus is the most speciose group of gekkonids to date (Kluge 2001) and the remarkable frequency at which new species are being added to this group shows no signs of leveling off (i.e., Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Grismer 2005, Grismer & Leong 2005Hayden et al. 2008;Hoang et al. 2007;Heidrich et al. 2007;Nazarov et al. 2008;Ngo & Bauer 2008;Nguyen et al. 2006;Orlov et al. 2007;Rösler & Glaw 2008;Rösler et al. 2007;Youmans & Grismer 2006;Ziegler et al. 2002). Cyrtodactylus are generally forest-dwelling lizards noted for their slender, inflected digits, long limbs and associated terrestrial to scansorial habits. ...
... These characteristics are likely to have contributed to their widespread radiation from tropical South Asia, Indochina, the Philippines, and through the Indo-Australian Archipelago to as far east as the Solomon Islands (Bauer and Henle, 1994). There are at least 21 species of Cyrtodactylus currently recognized from the Sunda Shelf region of Southeast Asia collectively ranging from the Isthmus of Kra, Thailand (sensu Hughes et al. 2003) southward to Borneo, Sumatra, Java, and their satellite islands (Manthey & Grossman, 1997;Grismer, 2005;Rösler & Glaw 2008;Youmans & Grismer, 2006). Ten of these species occur in Peninsular (= West) Malaysia (Grismer 2008b) and in southern Peninsular Malaysia, Grismer and Leong (2005) and Grismer et al. (2007) noted that at least four of these, C. consobrinus, C. quadrivirgatus, C. semenanjungensis, and C. sworderi, are lowland inhabitants likely occurring in sympatry (Wood et al. 2008). ...
... Some of the information on character states and their distribution in other species was obtained from Das andLim (2000), De Rooij (1915), Dring (1979), Grismer (2008a), Hikida (1990), Inger andKing (1961), Manthey and Grossman (1997), Pauwels et al. (2004), Rösler and Glaw (2008), Smith (1930), andTaylor (1963). Additional specimens examined are listed in the appendix. ...
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Three new species of the gekkonid genus Cyrtodactylus are described from Peninsular Malaysia; C. jarakensis sp. nov. from Pulau Jarak off the west coast of Perak, C. batucolus sp. nov. from Pulau Besar of the Water Islands Archipelago off the west coast of Melaka, and C. pantiensis sp. nov. from the Gunung Panti Forest Reserve, Johor. All are distinguishable from other Sunda Shelf species of Cyrtodactylus in having unique combinations of morphological and color pattern characteristics. The continued high potential for finding additional new species in unexplored regions of southern Peninsular Malaysia and its associated archipelagos is further supported with these findings.
... The lizard genus Cyrtodactylus is the largest group of gekkonids to date (Kluge 2001) and the rate at which new species are being discovered and described shows no signs of leveling off (i.e., Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Grismer 2005, Grismer & Leong 2005Hayden et al. 2008;Hoang et al. 2007;Heidrich et al. 2007;Kraus 2007;Linkem et al. 2008;Ngo & Bauer 2008;Ngyuen et al. 2006;Orlov et al. 2007;Pauwels et al. 2004;Rösler & Glaw 2008;Rösler et al. 2007;Youmans & Grismer 2006;Zeigler et al. 2002). Cyrtodactylus are terrestrial to scansorial forest-dwelling lizards noted for their slender, inflected digits, and long limbs. ...
... They collectively range from tropical South Asia, Indochina, the Philippines, and the Indo-Australian Archipelago to as far east as the Solomon Islands (Bauer & Henle 1994). There are at least 24 species of Cyrtodactylus currently recognized from the Sunda Shelf region of Southeast Asia, and these species collectively range from the Isthmus of Kra, Thailand (sensu Hughes et al. 2003) southward to Borneo, Sumatra, Java, and their satellite islands (Manthey & Grossmann 1997;Grismer 2005;Rösler & Glaw 2008;Youmans & Grismer 2006). Of these, 13 species occur in Peninsular Malaysia (Grismer 2008c) and two, C. pulchellus Gray and C. quadrivirgatus Taylor, are known from the Langkawi Archipelago off the northwest coast of Peninsular Malaysia (Fig. 1). ...
... Some of the information on character states and their distribution in other species was obtained from Das and Lim (2000), De Rooij (1915), Dring (1979), Grismer (2008a), Grismer et al. (2007, Hikida (1990), Inger and King (1961), Manthey and Grossmann (1997), Pauwels et al. (2004), Rösler and Glaw (2008), Smith (1930), and Taylor (1963). Additional specimens examined are listed in the appendix. ...
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A new species of the gekkonid lizard, Cytrodactylus macrotuberculatus sp. nov., is described from Pulau Langkawi of the Langkawi Archipelago located off the northwest coast of Peninsular Malaysia in the state of Kedah. This species had been previously recognized as C. pulchellus but is differentiated here from that species and all other Sunda Shelf Cyrtodactylus on the basis of its strong tuberculation and other scale and color pattern characteristics. This represents the second endemic species known from the Langkawi Archipelago.
... These medium-sized geckos are nocturnal and mostly live in undisturbed areas, making them very difficult to study, thus their occurrence and biology are poorly documented. Since 2000, many new species have been discovered, especially in Southeast Asian countries, namely Laos (David et al. 2004), Malaysia (Das & Lim 2000;Grismer 2005;Grismer & Ahmad 2008;Grismer & Leong 2005;Rösler & Glaw 2008;Youmans & Grismer 2006), Myanmar (Bauer 2002;2003), and Vietnam (Heidrich et al. 2007;Geissler et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Nguyen et al. 2006;Orlov et al. 2007;Quang et al. 2007;Rösler et al. 2007Ziegler et al. 2003). ...
... Comparisons were made with museum material in the collections of Chulalongkorn University Museum of Zoology (CUMZ), Thailand Natural History Museum (THNHM), Institut Royal des Sciences Naturelles de Belgique (IRSNB), and Montri Sumontha's personal collection (MS), as well as original published descriptions provided in broader fauna and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Das & Lim 2000;Bauer 2003;Bauer et al. 2002Bauer et al. , 2003Pauwels et al. 2004;Rösler & Glaw 2008) All specimens were preserved in 10% formaldehyde solution for 2 months then transferred to 80% ethanol. Etymology. ...
... We compare the new taxon to species currently known from Thailand and neighboring Laos, Myanmar and Cambodia. Table 2, modified from Rösler & Glaw (2008), presents important characters of each species in the region. Cyrtodactylus auribalteatus sp. ...
Article
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A new cave-dwelling species of the gekkonid lizard genus Cyrtodactylus Gray, C. auribalteatus sp. nov. is described on the basis of animals collected from a cave in Thung Salaeng Luang National Park, Phitsanulok Province, Thailand, bringing the number of species in this genus in Thailand to 18. The new species shares over all appearances with other cave-dwelling Cyrtodactylus spp. in having large eyes, a long snout, reduced tubercles, absence of caudal tubercles, and slender limbs for climbing. The new species is characterized by its moderate size (snout-vent length to at least 98.07 mm), weakly conical tubercles in 22–24 rows at midbody, high number of ventral scale across midbody (38–40), absence of precloacal groove, present of a single series of 6 precloacal pores and 4–5 femoral pores on each thigh separated by a diastema, broad subcaudal plates, and its dorsal pattern of three dark bands between limb insertions. It is the seventh species of cave-dwelling Cyrtodactylus, including the sandstone cave species, C. jarujini, recorded from Thailand.
... Other species assigned to this group (Grismer et al. 2010) were described only recently. Rösler and Glaw (2008) described Cyrtodactylus stresemanni on the basis of a single distinctive gecko collected 105 years ago, reportedly from Perak Province, Malaysia. Shortly thereafter, Grismer et al. (2010) described C. durio from Kedah, Malaysia, and generated a preliminary phylogenetic hypothesis for prehensile-tailed Cyrtodactylus based on morphological characters. ...
... We have shown that some distinctive characters originally suggesting a close relationship between Cyrtodactylus durio and C. stresemanni are also found in C. serratus and C. nuaulu. Although the specimen has faded with age, the dorsal pattern of the holotype of C. stresemanni remains well-defined (Rösler & Glaw 2008, their figure 7). The specimen has two pairs of elongate blotches on the flanks separated by a wide, pale vertebral stripe. ...
... Morphological similarity among Cyrtodactylus stresemanni and three Wallacea-Papuan species in our phylogeny raises questions about the origin of the holotype of C. stresemanni. Rösler and Glaw (2008) reported "Batang Padang-Tal, 800-900 m" as the type locality based on a handwritten label inside the jar with the specimen. They traced the locality to Tapah, Perak State, Peninsular Malaysia, and refer to Stresemann's own correspondence describing this locality. ...
Article
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We redescribe Cyrtodactylus lateralis (Werner) on the basis of new specimens. Cyrtodactylus lateralis is a prehensiletailed species, known from scattered lowland to mid-elevation localities in northern Sumatra. The prehensile-tailed Cyrtodactylus are more speciose and have a wider distribution than previously thought. This group includes a mainland SE Asian clade consisting of C. elok, C. interdigitalis, and C. brevipalmatus and an insular clade containing C. durio, C. lateralis, C. nuaulu, C. serratus, C. spinosus, and C. stresemanni. However, a distinctive color pattern in the Wallacean and Papuan species and uncertainty surrounding the type locality of C. stresemanni raise unresolved questions about the inclusiveness of the insular clade. DNA sequence data supports a close relationship between C. elok and C. interdigitalis, but also reveals that C. lateralis and C. durio are not closely related to these species.
... The lizard genus Cyrtodactylus is the most speciose group of gekkonids to date (Kluge 2001) and the remarkable frequency at which new species are being added to this group shows no signs of leveling off (i.e., Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Grismer 2005, Grismer & Leong 2005Hayden et al. 2008;Hoang et al. 2007;Heidrich et al. 2007;Nazarov et al. 2008;Ngo & Bauer 2008;Nguyen et al. 2006;Orlov et al. 2007;Rösler & Glaw 2008;Rösler et al. 2007;Youmans & Grismer 2006;Ziegler et al. 2002). Cyrtodactylus are generally forest-dwelling lizards noted for their slender, inflected digits, long limbs and associated terrestrial to scansorial habits. ...
... These characteristics are likely to have contributed to their widespread radiation from tropical South Asia, Indochina, the Philippines, and through the Indo-Australian Archipelago to as far east as the Solomon Islands (Bauer and Henle, 1994). There are at least 21 species of Cyrtodactylus currently recognized from the Sunda Shelf region of Southeast Asia collectively ranging from the Isthmus of Kra, Thailand (sensu Hughes et al. 2003) southward to Borneo, Sumatra, Java, and their satellite islands (Manthey & Grossman, 1997;Grismer, 2005;Rösler & Glaw 2008;Youmans & Grismer, 2006). Ten of these species occur in Peninsular (= West) Malaysia (Grismer 2008b) and in southern Peninsular Malaysia, Grismer and Leong (2005) and Grismer et al. (2007) noted that at least four of these, C. consobrinus, C. quadrivirgatus, C. semenanjungensis, and C. sworderi, are lowland inhabitants likely occurring in sympatry (Wood et al. 2008). ...
... Some of the information on character states and their distribution in other species was obtained from Das andLim (2000), De Rooij (1915), Dring (1979), Grismer (2008a), Hikida (1990), Inger andKing (1961), Manthey and Grossman (1997), Pauwels et al. (2004), Rösler and Glaw (2008), Smith (1930), andTaylor (1963). Additional specimens examined are listed in the appendix. ...
Article
Full-text available
Three new species of the gekkonid genus Cyrtodactylus are described from Peninsular Malaysia; C. jarakensis sp. nov. from Pulau Jarak off the west coast of Perak, C. batucolus sp. nov. from Pulau Besar of the Water Islands Archipelago off the west coast of Melaka, and C. pantiensis sp. nov. from the Gunung Panti Forest Reserve, Johor. All are distinguishable from other Sunda Shelf species of Cyrtodactylus in having unique combinations of morphological and color pattern characteristics. The continued high potential for finding additional new species in unexplored regions of southern Peninsular Malaysia and its associated archipelagos is further supported with these findings.
... The lizard genus Cyrtodactylus is the largest group of gekkonids to date (Kluge 2001) and the rate at which new species are being discovered and described shows no signs of leveling off (i.e., Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Grismer 2005, Grismer & Leong 2005Hayden et al. 2008;Hoang et al. 2007;Heidrich et al. 2007;Kraus 2007;Linkem et al. 2008;Ngo & Bauer 2008;Ngyuen et al. 2006;Orlov et al. 2007;Pauwels et al. 2004;Rösler & Glaw 2008;Rösler et al. 2007;Youmans & Grismer 2006;Zeigler et al. 2002). Cyrtodactylus are terrestrial to scansorial forest-dwelling lizards noted for their slender, inflected digits, and long limbs. ...
... They collectively range from tropical South Asia, Indochina, the Philippines, and the Indo-Australian Archipelago to as far east as the Solomon Islands (Bauer & Henle 1994). There are at least 24 species of Cyrtodactylus currently recognized from the Sunda Shelf region of Southeast Asia, and these species collectively range from the Isthmus of Kra, Thailand (sensu Hughes et al. 2003) southward to Borneo, Sumatra, Java, and their satellite islands (Manthey & Grossmann 1997;Grismer 2005;Rösler & Glaw 2008;Youmans & Grismer 2006). Of these, 13 species occur in Peninsular Malaysia (Grismer 2008c) and two, C. pulchellus Gray and C. quadrivirgatus Taylor, are known from the Langkawi Archipelago off the northwest coast of Peninsular Malaysia (Fig. 1). ...
... Some of the information on character states and their distribution in other species was obtained from Das and Lim (2000), De Rooij (1915), Dring (1979), Grismer (2008a), Grismer et al. (2007, Hikida (1990), Inger and King (1961), Manthey and Grossmann (1997), Pauwels et al. (2004), Rösler and Glaw (2008), Smith (1930), and Taylor (1963). Additional specimens examined are listed in the appendix. ...
Article
Full-text available
A new species of the gekkonid lizard, Cytrodactylus macrotuberculatus sp. nov., is described from Pulau Langkawi of the Langkawi Archipelago located off the northwest coast of Peninsular Malaysia in the state of Kedah. This species had been previously recognized as C. pulchellus but is differentiated here from that species and all other Sunda Shelf Cyrtodactylus on the basis of its strong tuberculation and other scale and color pattern characteristics. This represents the second endemic species known from the Langkawi Archipelago.
... Its widespread radiation comprises tropical South Asia, Indochina, the Philippines, the Indo-Australian Archipelago, and the Solomon Islands in the East (Bauer & Henle 1994). New species are regularly being described (e.g., Bauer 2002, Günther & Rösler 2002, Pauwels et al. 2004, Batuwita & Bahir 2005, Grismer 2005, Grismer & Leong 2005, Grismer & Norhayati 2008, Kraus & Allison 2006, Youmans & Grismer 2006, Rösler & Glaw 2008, Ngo & Chan 2010, Shi & Zhao 2010, Welton et al. 2009, Ziegler et al. 2010. One of the regions of greatest discovery of new Cyrtodactylus has been Vietnam. ...
... The size and thus the number of midbody ventrals in Cyrtodactylus varies between 22-82 (C. variegatus versus C. caovansungi, see Rösler & Glaw 2008). Cyrtodactylus wayakonei sp. ...
... n. differs from the representatives of the subgenus Geckoella by a larger SVL (90 versus 52-80 mm) and by the presence of precloacal pores in both sexes from C. (Deraniyagala, 1945). Cyrtodactylus (Geckoella) triedrus (Günther, 1864) differs from the new species in having 3-4 precloacal and 3-4 femoral pores (Das 1996, Rösler & Glaw 2008. ...
Article
Full-text available
We describe a new species of the gekkonid genus Cyrtodactylus from northern Laos, based on four specimens fro Luang Nam Tha Province. Cyrtodactylus wayakonei sp. n. is distinguished from the remaining bent-toed-geckos by the combination of the following characters: snout-vent length up to 90 mm; 7-8 supralabials; 9-10 infralabials; rostrum bearing smooth scales; 34-45 scales between fifth supralabials; dorsal tubercles smooth or slightly keeled; 17-19 dorsal tubercle rows; ventrolateral folds slightly developed, with large tubercles; 31-35 ventrals; 85-98 scales around midbody; 151-163 scales between mental and cloacal slit; 6-8 precloacal pores in both sexes; precloacal groove lacking; enlarged femoral scales absent; femoral pores lacking; subcaudals somewhat enlarged, broadened; tail tubercles flat, smooth; head and dorsum with blotched to reticulated pattern.
... These medium-sized geckos are nocturnal and mostly live in undisturbed areas, making them very difficult to study, thus their occurrence and biology are poorly documented. Since 2000, many new species have been discovered, especially in Southeast Asian countries, namely Laos (David et al. 2004), Malaysia (Das & Lim 2000;Grismer 2005;Grismer & Ahmad 2008;Grismer & Leong 2005;Rösler & Glaw 2008;Youmans & Grismer 2006), Myanmar (Bauer 2002;2003), and Vietnam (Heidrich et al. 2007;Geissler et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Nguyen et al. 2006;Orlov et al. 2007;Quang et al. 2007;Rösler et al. 2007Ziegler et al. 2003). ...
... Comparisons were made with museum material in the collections of Chulalongkorn University Museum of Zoology (CUMZ), Thailand Natural History Museum (THNHM), Institut Royal des Sciences Naturelles de Belgique (IRSNB), and Montri Sumontha's personal collection (MS), as well as original published descriptions provided in broader fauna and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Das & Lim 2000;Bauer 2003;Bauer et al. 2002Bauer et al. , 2003Pauwels et al. 2004;Rösler & Glaw 2008) All specimens were preserved in 10% formaldehyde solution for 2 months then transferred to 80% ethanol. Etymology. ...
... We compare the new taxon to species currently known from Thailand and neighboring Laos, Myanmar and Cambodia. Table 2, modified from Rösler & Glaw (2008), presents important characters of each species in the region. Cyrtodactylus auribalteatus sp. ...
Article
Full-text available
A new cave-dwelling species of the gekkonid lizard genus Cyrtodactylus Gray, C. auribalteatus sp. nov. is described on the basis of animals collected from a cave in Thung Salaeng Luang National Park, Phitsanulok Province, Thailand, bringing the number of species in this genus in Thailand to 18. The new species shares over all appearances with other cave-dwelling Cyrtodactylus spp. in having large eyes, a long snout, reduced tubercles, absence of caudal tubercles, and slender limbs for climbing. The new species is characterized by its moderate size (snout-vent length to at least 98.07 mm), weakly conical tubercles in 22-24 rows at midbody, high number of ventral scale across midbody (38-40), absence of precloacal groove, present of a single series of 6 precloacal pores and 4-5 femoral pores on each thigh separated by a diastema, broad subcaudal plates, and its dorsal pattern of three dark bands between limb insertions. It is the seventh species of cave-dwelling Cyrtodactylus, including the sandstone cave species, C. jarujini, recorded from Thailand.
... Its widespread radiation comprises tropical South Asia, Indochina, the Philippines, the Indo-Australian Archipelago, and the Solomon Islands in the East (Bauer & Henle 1994). New species are regularly being described (e.g., Bauer 2002, Günther & Rösler 2002, Pauwels et al. 2004, Batuwita & Bahir 2005, Grismer 2005, Grismer & Leong 2005, Grismer & Norhayati 2008, Kraus & Allison 2006, Youmans & Grismer 2006, Rösler & Glaw 2008, Ngo & Chan 2010, Shi & Zhao 2010, Welton et al. 2009, Ziegler et al. 2010. One of the regions of greatest discovery of new Cyrtodactylus has been Vietnam. ...
... The size and thus the number of midbody ventrals in Cyrtodactylus varies between 22-82 (C. variegatus versus C. caovansungi, see Rösler & Glaw 2008). Cyrtodactylus wayakonei sp. ...
... n. differs from the representatives of the subgenus Geckoella by a larger SVL (90 versus 52-80 mm) and by the presence of precloacal pores in both sexes from C. (Deraniyagala, 1945). Cyrtodactylus (Geckoella) triedrus (Günther, 1864) differs from the new species in having 3-4 precloacal and 3-4 femoral pores (Das 1996, Rösler & Glaw 2008. ...
Article
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We describe a new species of the gekkonid genus Cyrtodactylus from northwestern Laos, based on a recent collection from Vientiane Province. Cyrtodactylus pageli sp. n. is distinguished from the remaining bent-toed-geckos by the combination of the following characters: snout-vent length up to 81.8 mm; 9-12 supralabials; 9 infralabials; 9-14 rows of dorsal tubercles; no distinct lateral body fold; 41-46 midbody ventrals; precloacal pores present in both sexes (males 4, females 5 or 6); no precloacal groove; femoral scales not distinctly enlarged; no femoral pores; subdigital lamellae under the fourth toe 19-26; subcaudals transversally enlarged; dark nuchal band, followed by a transversal row of dark neck blotches (rarely forming an additional transversal neck band) in-between nuchal band and first body band; 5 in part irregular dark dorsal bands between limb insertions plus transversal band between hind limbs; tail with banded pattern.
... Comparisons were made with museum material in the collections of the California Academy of Sciences, San Francisco (CAS), Museum of Comparative Zoology, Cambridge, Massachusetts (MCZ), and Western Australian Museum (WAM; where the holotype is deposited). Original published descriptions and descriptions provided in broader faunal and taxonomic treatments (e.g., Annandale 1913;de Rooij 1917;Darevsky 1964;Auffenberg 1980;Bauer 2002Bauer , 2003Rösler & Glaw 2008;Linkem et al. 2008) were also consulted. ...
... nov. is at least 70% of the maximum size the species might attain. Of named species summarized by Rösler and Glaw (2008), only the following species had maximum SVLs of less than 65 mm, the maximum size of C. kimberleyensis sp. nov. ...
... so predicted: C. consobrinoides (Annandale), C. feae (Boulenger, C. annandalei Bauer, C. gansi Bauer, and C. wakeorum Bauer-all from Myanmar, C. agamensis (Bleeker) from Sumatra, C. buchardi David et al. from Laos, C. gubernatoris (Annandale) from the eastern Himalayas, C. laevigatus Darevsky from Komodo and Flores, C. malcolmsmithi (Constable) from northern India, and several species in the subgenus Geckoella occurring in peninsular India and Sri Lanka. Cyrtodactylus jellesmae (Boulenger) from Sulawesi was reported to have a maximum SVL of 63 mm by Rösler & Glaw (2008) but adults are now known to reach at least 75 mm SVL . All descriptions published since the summary of Rösler and Glaw (2008) were consulted and the only additional taxon with a mature SVL of less than 65 mm subsequently described is C. mandalayensis Mahony. ...
Article
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A diminutive new species of Cyrtodactylus is described from East Montalivet Island off the north coast of the Kimberley region of Western Australia. Cyrtodactylus kimberleyensis sp. nov. may be distinguished from all other congeners by its small size (gravid female holotype 45 mm SVL), its lack of enlarged subcaudal plates, 16–18 rows of dorsal tubercles, weakly developed ventrolateral skin fold, and dorsal pattern lacking dark transverse bands and enlarged blotches. The new species is one of the smallest in the genus and is the first Cyrtodactylus known from Western Australia. It is not closely related to the large-bodied species of the Cape York Peninsula, Queensland, but rather has affinities to small to mid-sized species occurring on Timor and in the Lesser Sundas, and thus represents a second pathway of colonization of northern Australia.
... Since 2005 21 new species, most with highly restricted distributions, have been described in Peninsular Malaysia and its offshore islands (Grismer 2005(Grismer , 2006Grismer & Leong 2005;Youmans & Grismer 2006;Grismer & Ahmad 2008;Rösler & Glaw 2008) and Vietnam (Nguyen et al. 2006;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Ngo & Bauer 2008;Ngo 2008;Nazarov et al. 2008; alone. Other relatively recent discoveries include a new Cyrtodactylus from Laos (David et al. 2004), nine from Myanmar (Bauer 2002(Bauer , 2003, and four from Thailand Pauwels et al. 2004). ...
... Although highly artificial, we have employed commonly used and easily scorable single differential characters for this purpose. Recent summaries of some of these, and other, features for large numbers of Cyrtodactylus species have been provided by Youmans & Grismer (2006), Grismer et al. (2007), Grismer & Ahmad (2008), Linkem et al. (2008), and Rösler & Glaw (2008). ...
Article
A new cave-dwelling species of Cyrtodactylus Gray is described from Mae Hong Son Province in northern Thailand. The new species, C. erythrops sp. nov., is characterized by its moderate size (snout-vent length to at least 78 mm), relatively large, closely-spaced, flattened tubercles in 18–20 irregular rows at midbody, low number of ventral scales across midbody (28), absence of precloacal groove, presence of precloacal and femoral pores separated by a diastema, broad subcaudal plates, and dorsal pattern of dark spots and blotches. It is the fifth species of cave-dwelling Cyrtodactylus recorded from Thailand and its discovery adds to the mounting evidence that this genus exhibits unprecedented levels of localized endemism throughout tropical Southeast Asia.
... Some of the information on character states and their distribution in other species was obtained from Das and Lim (2000), De Rooij (1915), Dring (1979), Grismer (2008a), Hikida (1990), Inger and King (1961), Manthey and Grossman (1997), Pauwels et al. (2004), Rösler and Glaw (2008), Smith (1930), and Taylor (1963). UKMHC refers to the Herpetological Collection of Universiti Kebangsaan Malaysia (UKM), Bangi, Selangor, Peninsular Malaysia; ZRC refers to the Zoological Reference Collection at the Raffles Museum of Biodiversity Research, National University of Singapore; the type series has been deposited at the Herpetological Collection of Universiti Kebangsaan Malayia, Bangi, Selangor and the Raffles Museum of Biodiversity Research, National University of Singapore. ...
... Additionally, Cyrtodactylus leegrismeri differs from C. condorensis and C. paradoxus in body markings (see Fig. 2A Cyrtodactylus leegrismeri and other Sunda Shelf Cyrtodactylus are presented in Table 1. For comparisons with Cyrtodactylus in other regions, see Rösler and Glaw (2008). ...
Article
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A new species of endemic Cyrtodactylus is described from Pulau Tenggol, Terengganu, off the northeastern coast of Peninsular Malaysia. It differs from other Sunda Shelf species by having a maximum SVL of 92 mm, enlarged tubercles on body and hind limbs but none on fore limbs, 27-35 ventral scale, a single row of transversely enlarged, median subcaudal scales, 18-20 subdigital lamellae on fourth toe, four precloacal pores, and a single row of enlarged, non porebearing femoral scales beneath each thigh which are not continuous with the precloacal pores.
... At least 35 species of the gekkonid genus Cyrtodactylus are reported from Indochina (Smith 1935;Bauer et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Sumontha et al. 2010;Ziegler et al. 2010). Of these, twenty species are from Vietnam, most of them described only in recent years (Smith 1935;Bobrov et al. 1995;Darevsky & Szczerbak 1997;Ziegler et al. 2002;Nguyen et al. 2006;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo, 2008;Ngo & Bauer 2008;Geissler et al. 2009;Ziegler et al. 2010). ...
... Comparisons were made with material from the Zoological Collection at the University of Natural Sciences as well as original published descriptions and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Cox et al. 1998;Das & Lim 2000;Das 2006;Ziegler et al. 2002;Bauer 2002Bauer & 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Grismer et al. 2007Nguyen et al. 2006;Youmans & Grismer 2006;Heidrich et al. 2007;Hoang et al. 2007;Kraus & Allison 2006;Kraus 2007Kraus & 2008Linkem et al. 2008;Oliver et al. 2008;Oliver et al. 2009;Orlov et al. 2007;Nazarov et al. 2008;Rösler & Glaw 2008;Rösler et al. 2007;Ngo 2008;Ngo & Bauer 2008;Geissler et al. 2009;Welton et al. 2009;Lei & Hui 2010;Sumontha et al. 2010;Ziegler et al. 2010 Diagnosis. Cyrtodactylus yangbayensis differs from all other congeners by the following combination of characters: mean SVL of 83.5 mm (± 4.7 mm); original tail long (TL/SVL: 1.28); dark brown blotches on head; nuchal loop broken into two dark fragments or V-shaped; dorsal pattern consisting of five to seven irregular rows of narrow, dark brown bands between limb insertions; 6-8 precloacal pores in males; 5-16 enlarged scales beneath thighs; 18-23 interorbital scales on the frontal bone; 26-32 scales in a straight line between eye and nostril; 39-46 rows of ventral scales between ventrolateral folds; 20-23 irregular, longitudinal rows of weakly-keeled, conical tubercles at midbody between the lateral folds; 28-34 paravertebral tubercles between limb insertions; 18-20 subdigital lamellae on first toe; 15-17 subdigital lamellae on fourth toe; 9-10 enlarged scales on heel; 9-11 irregular bands on original tail; median row of enlarged subcaudal scales. ...
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A new species of bent-toed gecko, Cyrtodactylus cucphuongensis sp. nov. is described from the karst forest of Cuc Phuong National Park, Ninh Binh province, Northern Vietnam. It differs from all other species of Indo-Chinese Cyrtodactylus by the following combination of characters: SVL 96.0 mm; absence of precloacal and femoral pores; 19 subdigital lamellae on first toe, 24 on fourth toe; large, dark spots on top of head; wide, prominent nuchal band; and five or six dark, irregular, broad bands between limb insertions
... At least 35 species of the gekkonid genus Cyrtodactylus are reported from Indochina (Smith 1935;Bauer et al. 2009;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Sumontha et al. 2010;Ziegler et al. 2010). Of these, twenty species are from Vietnam, most of them described only in recent years (Smith 1935;Bobrov et al. 1995;Darevsky & Szczerbak 1997;Ziegler et al. 2002;Nguyen et al. 2006;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo, 2008;Ngo & Bauer 2008;Geissler et al. 2009;Ziegler et al. 2010). ...
... Comparisons were made with material from the Zoological Collection at the University of Natural Sciences as well as original published descriptions and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Cox et al. 1998;Das & Lim 2000;Das 2006;Ziegler et al. 2002;Bauer 2002Bauer & 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Grismer et al. 2007Nguyen et al. 2006;Youmans & Grismer 2006;Heidrich et al. 2007;Hoang et al. 2007;Kraus & Allison 2006;Kraus 2007Kraus & 2008Linkem et al. 2008;Oliver et al. 2008;Oliver et al. 2009;Orlov et al. 2007;Nazarov et al. 2008;Rösler & Glaw 2008;Rösler et al. 2007;Ngo 2008;Ngo & Bauer 2008;Geissler et al. 2009;Welton et al. 2009;Lei & Hui 2010;Sumontha et al. 2010;Ziegler et al. 2010 Diagnosis. Cyrtodactylus yangbayensis differs from all other congeners by the following combination of characters: mean SVL of 83.5 mm (± 4.7 mm); original tail long (TL/SVL: 1.28); dark brown blotches on head; nuchal loop broken into two dark fragments or V-shaped; dorsal pattern consisting of five to seven irregular rows of narrow, dark brown bands between limb insertions; 6-8 precloacal pores in males; 5-16 enlarged scales beneath thighs; 18-23 interorbital scales on the frontal bone; 26-32 scales in a straight line between eye and nostril; 39-46 rows of ventral scales between ventrolateral folds; 20-23 irregular, longitudinal rows of weakly-keeled, conical tubercles at midbody between the lateral folds; 28-34 paravertebral tubercles between limb insertions; 18-20 subdigital lamellae on first toe; 15-17 subdigital lamellae on fourth toe; 9-10 enlarged scales on heel; 9-11 irregular bands on original tail; median row of enlarged subcaudal scales. ...
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A new species of bent-toed gecko, Cyrtodactylus yangbayensis sp. nov. is described from the monsoon evergreen forests of Truong Son Mountain Range, Khanh Hoa province. It differs from all other species of Indo-Chinese Cyrtodactylus by having a mean SVL of 83.5 mm; dark spots and blotches on the head; no continuous nuchal loop; 6-8 precloacal pores in males arranged in a chevron; five or six dark brown, irregular, dorsal bands between limb insertions; 18-20 subdigital lamellae under first toe; 15-17 subdigital lamellae under fourth toe; and a median row of enlarged subcaudal scales.
... The genus Cyrtodactylus is the most diverse gekkonid genus with 196 species (Uetz 2014) and recent studies on the genus have improved our ability to define species boundaries (e.g., Rösler & Glaw 2008, Grismer et al. 2012b, as the pace of herpetological fieldwork in Southeast Asia has accelerated (e.g., Bauer 2002, 2003Das 2004-Borneo;Grismer et al. 2011-Indochina and Malaysia;Kaiser et al. 2011-Timor;Linkem et al. 2008Setiadi et al. 2011-Indonesia). Cyrtodactylus is a geographically wide-ranging genus with species occupying diverse ecological niches. ...
... Cyrtodactylus is a geographically wide-ranging genus with species occupying diverse ecological niches. Species of Cyrtodactylus are found in mainland Asia, as far northwest as northern India, Nepal, and Tibet, and members of the genus are more-or-less continuously distributed eastwards throughout Southeast Asia and the Indo-Australian Archipelago, reaching the Philippines, New Guinea, northern Australia, and the Solomon Islands (Rösler & Glaw 2008;Uetz 2014;Wood et al. 2012). In view of this broad distribution, the relative paucity of known forms from the ecologically diverse Lesser Sunda Archipelago (Fig. 1), currently limited to C. darmandvillei (Weber, 1890), C. gordongekkoi (Das, 1993), C. laevigatus Darevsky, 1964 and its subspecies C. l. laevigatus and C. l. uniformis Auffenberg, 1980, and C. wetariensis (Dunn, 1927), is intriguing. ...
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We describe a new species of bent-toed gecko from a single specimen initially collected in 1924 by Malcolm Smith on Timor Island in the Lesser Sunda Archipelago of Indonesia. Cyrtodactylus celatus sp. nov. is distinguished from all other congeners by the following combination of characters: small adult size; without spinose tubercles on the ventrolateral body fold and along the lateral margin of the tail; 16 longitudinal rows of tubercles at midbody; 42 ventral scales between the ventrolateral folds at midbody; no transversely enlarged, median subcaudal scales; 17 subdigital lamellae (seven basal + ten distal) under the fourth toe; no abrupt transition between postfemoral and ventral femoral scale series. The specimen is the earliest confirmed record of the genus Cyrtodactylus for Timor, and it is the first putatively endemic gecko species described from this island.
... The condition of precloacal and femoral scales and pores in males has traditionally been used to distinguish members of the genus Cyrtodactylus from one another (e.g., Smith, 1935;Darevsky & Szcerbak, 1997;Bauer, 2002Bauer, , 2003Bauer et al., 2003). Cyrtodactylus grismeri and Cyrtodactylus eisenmani differ from all but 10 species (two of which are variable in this trait) of Cyrtodactylus in lacking both precloacal and femoral pores in adult males (see Rösler & Glaw, 2008). The banded dorsal pattern of these two species distinguishes them from all of these except C. badenensis Nguyen et al.,C. ...
... Comparison among species of Cyrtodactylus that lack precloacal and femoral pores. Question marks indicate data unavailable in the literature (seeRösler & Glaw, 2008). ...
Article
Two new cave-dwelling species of Cyrtodactylus from mainland southwestern Vietnam and an offshore island are differ-entiated from all other congeners in lacking a precloacal groove and precloacal pores, presence of enlarged femoral scales beneath thigh, and in having a color pattern consisting of four or five narrow white bands on the body dorsum and one on the tail. Cyrtodactylus grismeri sp. nov. is reddish brown, has a mean SVL of 87.6 ± 3.8 mm (n = 9), 18–22 irreg-ular longitudinal rows of weakly-keeled tubercles at midbody between the lateral folds, 33–38 ventral scales between ventrolateral folds, 0–3 enlarged scales beneath thighs, and 20–24 subdigital lamellae under the first toe. Cyrtodactylus eisenmani sp. nov. is chocolate brown, has a mean SVL of 81.3 ± 5.0 mm (n = 5), 14 irregular longitudinal rows of weakly keeled tubercles at midbody between ventrolateral folds, 44–45 ventral scales between ventrolateral folds at mid-body, 4–6 enlarged femoral scales beneath each thigh, and 22–25 subdigital lamellae under the first toe. This discovery increases the total number of Cyrtodactylus found in Vietnam to fifteen.
... Comparisons were made with material housed in the Zoological Collection of the University of Natural Sciences, Hochiminh City, as well as with original published descriptions, data, and illustrations provided in broader faunal and taxonomic treatments (e.g., Smith 1935;Taylor 1963;Brown & Parker 1973;Ulber & Grossmann 1991;Ulber 1993;Darevsky & Szczerbak 1997;Das 1997Das , 2006Das & Lim 2000;Ziegler et al. 2002Ziegler et al. , 2010Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Günther & Rösler 2003;David et al. 2004;Pauwels et al. 2004;Batuwita & Bahir 2005;Grismer 2005;Grismer & Leong 2005;Kraus & Allison 2006;Nguyen et al. 2006;Youmans & Grismer 2006;Grismer et al. 2007Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Grismer & Ahmad 2008;Hayden et al. 2008;Kraus 2007Kraus , 2008Rösler et al. 2007Linkem et al. 2008;Nazarov et al. 2008;Rösler & Glaw 2008;Ngo 2008;Ngo & Bauer 2008;Oliver et al. 2008Oliver et al. , 2009Geissler et al. 2009;Mahony 2009;Welton et al. 2009Welton et al. & 2010 Paratypes. UNS 0527, subadult female, was collected by Ngo Van Tri between 19h00 and 20h00 on 26 April 2010 at the entrance of Lomyen Cave. ...
... differs from the recently described species Cyrtodactylus mandalayensis Mahony, C. nuaulu Oliver et al., C. yangbangensis Ngo & Chan, C. phuquocensis Ngo et al., C. zhaoermii Lei & Hui, C. jambangan Welton et al., and C. wayakonei Nguyen et al. in possessing femoral pores in males. Bauer et al. 2003;David et al. 2004;Rösler & Glaw 2008;Sumontha et al. 2008;Ulber 1993;Ziegler et al. 2002Ziegler et al. , 2010; and our observations on the type series of C. lomyenensis sp. nov. ...
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A new cave-dwelling bent-toed gecko, Cyrtodactylus lomyenensis sp. nov. is described from a karst forest in Gnommalath District, Khammouane Province, southern Laos. It differs from all other species of Indochinese-Thai Cyrtodactylus in the following combination of characters: maximum SVL of at least 71.2 mm; head dorsum yellowish with irregular brown blotches; presence of a brown nuchal loop reaching the posterior edge of the orbit; four narrow yellowish-cream transversal bands with irregular anterior and posterior black edges on a brown background between limb insertions; no precloacal groove; 39-40 precloacal-femoral pores in males, arranged in a continuous row; females with 32 precloacal-femoral pores in a continuous row, smaller than those of males; five postanal tubercles on each side; 16-18 subdigital lamellae on first toe; 19-23 subdigital lamellae on 4th toe; no tubercles on tail dorsum; and a median row of enlarged subcaudal scales.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e. g., Smith 1920Smith , 1921Smith , 1935Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Das 1997;Manthey & Grossmann 1997;Chan-ard et al. 1999;Das & Lim 2000;Ziegler et al. 2002;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003David et al. 2004;Pauwels et al. 2004;Batuwita & Bahir 2005;Grismer 2005;Grismer & Leong 2005;Kraus & Allison 2006;Nguyen et al. 2006;Youmans & Grismer 2006;Grismer et al. 2007Heidrich et al. 2007;Hoang et al. 2007;Kraus 2007;Orlov et al. 2007;Grismer & Norhayati 2008;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;. ...
Article
We describe a new species of the genus Cyrtodactylus from southern Vietnam, based on voucher specimens collected from Dong Nai and Ba Ria – Vung Tau provinces. Cyrtodactylus cattienensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small, with a maximum SVL of 69 mm; neck band present, extending to the posterior margin of the eye; trunk, tail and limbs bearing irregularly shaped bands: 4–6 light bands on trunk and 4–12 white bands on tail; 16–22 irregular longitudinal rows of dorsal tubercles; 28–42 longitudinal rows of ventrals at midbody; lateral folds weakly developed or absent, without enlarged tubercles; tail not depressed, not enlarged at base, scales arranged in whorls; males with 6–8 precloacal pores in angular continuous series; a patch of enlarged precloacal scales present in both sexes; 3–8 slightly enlarged femoral scales on each hind limb in both sexes; femoral pores absent; precloacal groove absent; subcaudal scales small, not transversely enlarged.
... The genus Cyrtodactylus ranges throughout most of tropical Asia, from India to the Philippines and south through the Indo-Australian Archipelago, northern Australia, New Guinea, and the Solomon Islands (Youmans & Grismer 2006). With over 170 recognized species, Cyrtodactylus is the most speciose genus of gekkonid lizards Rösler & Glaw 2008;Uetz & Hallerman 2010). Since 2008, at least four new species have been described from the Indonesian Archipelago Linkem et al. 2008;Oliver et al. 2009;Iskandar et al. 2011). ...
Article
Cyrtodactylus hikidai sp. nov. is a new gekkonid species described on the basis of 13 specimens collected from MountRanai, Bunguran Island, Indonesia. This large Cyrtodactylus (female SVL up to 102.2 mm, male SVL up to 100.6 mm)differs from all other Sunda Shelf species of Cyrtodactylus by the combination of a reticulated color pattern on the dorsumof the head, body banded, enlarged precloacal scales, six pore-bearing precloacal scales in a deep groove in both malesand females, tubercles on forelimbs, hind limbs and occiput, absence of enlarged femoral scales, and absence of femoral pores.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e. g., Smith 1920Smith , 1921Smith , 1935Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Das 1997;Manthey & Grossmann 1997;Chan-ard et al. 1999;Das & Lim 2000;Ziegler et al. 2002;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003David et al. 2004;Pauwels et al. 2004;Batuwita & Bahir 2005;Grismer 2005;Grismer & Leong 2005;Kraus & Allison 2006;Nguyen et al. 2006;Youmans & Grismer 2006;Grismer et al. 2007Heidrich et al. 2007;Hoang et al. 2007;Kraus 2007;Orlov et al. 2007;Grismer & Norhayati 2008;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;. ...
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Geissler, Peter, Nazarov, Roman, Orlov, Nikolai L., Böhme, Wolfgang, Phung, Trung My, Nguyen, Truong Quang, Ziegler, Thomas (2009): A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 2161: 20-32, DOI: 10.5281/zenodo.188976
... Comparisons were made with museum material in the collections of the Institut Royal des Sciences Naturelles de Belgique, Brussels (IRSNB), Muséum National d'Histoire Naturelle, Paris (MNHN), Chulalongkorn University Museum of Zoology (CUMZ), Thailand Natural History Museum, Pathum Thani, Thailand (THNHM]), Royal Forest Department, Bangkok, Thailand (RFD), Zoological Survey of India, Kolkata (ZSI), and California Academy of Sciences, San Francisco (CAS) (see Ulber 1993;Pauwels et al. 2000b;Pauwels & Chan-ard 2006;Kunya et al. 2014Kunya et al. , 2015Panitvong et al. 2014;Sumontha et al. 2014Sumontha et al. , 2015 for specimens examined and sources of data). Original published descriptions and descriptions provided in broader faunal and taxonomic treatments (e.g., Rösler & Glaw 2008;Ziegler et al. 2010;Ngo & Grismer 2012;Nguyen et al. 2014;Nazarov et al. 2014) were also consulted. Additional specimens. ...
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A new Bent-toed Gecko, Cyrtodactylus phetchaburiensis sp. nov. is described from the Tha Yang District of Phetchaburi Province, western Thailand. It is a medium-sized Cyrtodactylus (SVL to at least 63.2 mm), with small, mostly keeled tubercles in 20 regular longitudinal rows on dorsum; 33 scales across mid-venter between lowest rows of flank tubercles; enlarged row of femoral scales present; five precloacal pores in male, femoral pores and precloacal groove absent; 5–6 broad basal lamellae and 11 narrow distal lamellae beneath digit IV of pes; and a single median row of transversely enlarged subcaudal scales present. It has a dorsal colour pattern of large, dark, diffusely-edged markings on a fawn background and a pair of dark scapular patches. The species is a member of the Central Indochinese (Thai-Myanmar) clade of Cyrtodactylus and is most closely related to C. oldhami (Theobald), from which it differs in colour pattern.
... Data used for comparison originate from specimens that we examined ourselves (see Material examined) and from literature, as follows: Smith (1935), Taylor (1963), Ulber (1993), Das (1996), Bauer (2002), Bauer et al. (2002), Bauer (2003), Bauer et al. (2003), Ziegler et al. (2002), David et al. (2004), Pauwels et al. (2004), Biswas (2007), Hoang et al. (2007), Nguyen et al. (2007), Orlov et al. (2007), Nazarov et al. (2008), Ngo (2008), Ngo and Bauer (2008), , , Bauer et al. (2009), Geissler et al. (2009), Mahony (2009), Oliver et al. (2009), Welton et al. (2009, Bauer et al. (2010), Chan and Ahmad (2010), Grismer et al. (2010), Ngo and Chan (2010), , , Nguyen et al. (2010), Shi and Zhao (2010), Sumontha et al. (2010), and Ngo and Pauwels (2010). We especially used the key to species of Vietnam provided in Ziegler et al. (2010) and the overview paper provided by Rösler and Glaw (2008). Because the holotype is an adult female, we have used characters in the comparisons, that are applicable to females or, at least, to both sexes. ...
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A new species of the gekkonid genus Cyrtodactylus Gray, 1827, Cyrtodactylus teyniei spec. nov., is described based on a single adult female from Borikhamxay Province, central Laos. It is distinguished from any other Cyrtodactylus species by the combination of the following characters: (1) SVL 89.9 mm, (2) longitudinal rows of dorsal tubercles 19, (3) ventral scales in 38 rows between ventrolateral skin folds, (4) ventrolateral skin folds without tubercles, (5) enlarged femoral and precloacal scales 54 (precloacal scales 8, plus 23 femoral scales on each side), (6) precloacal depression lacking, (7) precloacal pores 14, (8) subdigital lamellae under the firth toe 19-20, (9) enlarged median subcaudals, and (10) nuchal band absent, dorsum with blotched pattern. An extended comparison with other species of the genus Cyrtodactylus, in particular species recently described from Laos and Vietnam, is provided. Cyrtodactylus teyniei spec. nov. is the seventh species of Cyrtodactylus known to occur in Laos.
... The bent-toed geckos of the genus Cyrtodactylus are widely distributed in southeastern Asia and adjacent areas, reaching northern Australia and western Melanesia (Kraus, 2007). With more than 100 described species Cyrtodactylus is among the world's most speciose gecko genera and many new species have been described recently Heidrich, et al., 2007;Linkem, et al., 2008;Ngo & Bauer, 2008;Roesler & Glaw, 2008;Xuan, et al., 2007). Although many species have been reassigned to other genera such as Tenuidactylus, Cyrtopodion, Mediodactylus, Nactus, and Geckoella (Kluge, 1983;Kluge, 2001;Szczerbak, 1984), the number of species currently in this genus continues to grow. ...
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A new species of Cyrtodactylus described from Nyemo County, Xizang Autonomous Region, China is here referred to a new species, Cyrtodactylus zhaoermii sp. nov. It is distinguished from all other Cyrtodactylus by the following characters: dorsal surface of the body with fine granules intermixed with larger sub-conical tubercles arranged into 20 more or less regular rows; proximal subdigital lamellae transversely expanded; 19-20 subdigital lamellae on toe IV; 30-32 mid-body ventral scales; most scales in femoral region small, granular, series of 12-14 enlarged femoral scales lacking pores; a series of precloacal pores (4) present in male; tail with distinct segments, large, posteriorly directed tubercles in whorls, numbering nine to ten per caudal annulus, three on each side and three to four on the dorsum, situated at the posterior edge of each annulus; subcaudals not transversely expanded. The new species is the fourth Cyrtodactylus known from Xizang Autonomous Region.
... InterorbSc 24 18 18 bearing scales by a diastema, and we thus hereafter provide a comparison with all other species known from Cambodia, China, Laos, Myanmar, Peninsular Malaysia, Thailand and Vietnam sharing this condition (see intraspecific comparisons and latest descriptions for this set of species in , Nazarov et al. 2008, Ngo & Bauer 2008, Rösler & Glaw 2008, David et al. 2011, Grismer 2008, 2011, Luu et al. 2011, Ziegler et al. 2013, Pauwels et al. 2014a-b, Phung et al. 2014). Ngo & Bauer, 2008, C. kingsadai Ziegler, Phung, Le & Nguyen, 2013and C. ziegleri Nazarov, Orlov, Nguyen & Ho, 2008. ...
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We describe a new forest-dwelling Cyrtodactylus from Doi Inthanon, Chiang Mai Province, northern Thailand. Cyrtodactylus inthanon sp. nov. is characterized by a maximum known SVL of 87.3 mm; 18 to 20 longitudinal rows of dorsal tubercles; a continuous series of 34 to 37 enlarged femoro-precloacal scales, including four to six pitted (female) or porebearing (male) scales on each femur separated by a diastema from five pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and three to five irregular beige dorsal bands between limb insertions. The discovery of a new reptile endemic to Doi Inthanon reinforces the high importance of this mountain in terms of biodiversity conservation.
... Some of the information on character states and their distribution in other species was obtained from , Das & Lim (2000), De Rooij (1915), Dring (1979), Grismer (2008), Grismer et al. (2008); Hikida (1990), Inger & King (1961), Manthey & Grossman (1997), Rösler & Glaw (2008), Smith (1930), and Taylor (1963). Additional specimens examined are listed in the appendix. ...
Article
Cyrtodactylus majulah is a new species of bent-toed gecko from Singapore and Pulau Bintan in the Riau Archipelago of Indonesia that had been masquerading under the nomen C. quadrivirgatus. It is differentiated from C. quadrivirgatus and all its congeners in Sundaland by having the following suite of unique morphological and colour pattern characteristics: a maximum SVL of 68.0 mm; eight or nine supralabials; seven or eight infralabials; strongly tuberculated body and limbs; 39–46 paravertebral tubercles; 43–53 ventral scales; no abrupt contact of large and small postfemoral scales; no enlarged femoral scales; 20–23 subdigital lamellae on the fourth toe; no femoral pores; no deep precloacal groove; enlarged, precloacal scales; 7–11 precloacal pores; no enlarged, median subcaudals; tubercles on the anterior portion of the tail; no reticulated pattern on the head; a blotched dorsal pattern; and lacking paired, dark, semi-lunar shaped blotches on the upper nape. This nocturnal, arboreal, forest and swamp dwelling species frequents tree trunks, leaves and low bushes and is the basal member of a monophyletic group that contains C. pantiensis and C. payacola from Peninsular Malaysia.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008, Rösler & Glaw 2008, Bauer et al. 2009, Ngo & Pauwels 2010, Ziegler et al. 2010, David et al. 2011, Iskandar et al. 2011, Ngo 2011. ...
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We describe a new species of the genus Cyrtodactylus based on two adult specimens from Huong Son limestone forest, Hanoi, Vietnam. Cyrtodactylus huongsonensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: medium-sized, with a maximum SVL of 89.8 mm; dorsal pattern consisting of dark nuchal loop, neck band and five in part irregular transverse body bands between limbs; two enlarged lateral chin-shields in contact with first postmental pair; dorsal tubercles present on occiput, body, forearms, hind limbs and tail base; 14–16 irregularly running dorsal tubercle rows; ventrals in 41–48 longitudinal rows at midbody; lateral skin folds present, without interspersed tubercles; six precloacal pores plus in total 17 femoral pores in males, which are separated by 8–12 poreless scales; enlarged femoral scales and precloacal scales present; three postcloacal spurs in males; subcaudal scales transversely enlarged. This is the 24 th species of Cyrtodactylus known from Vietnam.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Smith 1920Smith , 1921Smith , 1935Taylor 1963;Ulber & Grossmann 1991;Ulber 1993;Darevsky & Szczerbak 1997;Manthey & Grossmann 1997;Ziegler et al. 2002;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003David et al. 2004;Pauwels et al. 2004;Grismer 2005;Grismer & Leong 2005;Ngo & Grismer 2006;Nguyen et al. 2006;Grismer et al. 2007;Heidrich et al. 2007;Hoang et al. 2007;Orlov et al. 2007;Grismer & Norhayati 2008;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Bauer et al. 2009;Geissler et al. 2009). In comparison with Vietnamese congeners, Cyrtodactylus roesleri sp. ...
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We describe a third cryptic species of the genus Cyrtodactylus from Phong Nha – Ke Bang National Park, Quang Binh Province, Vietnam. Cyrtodactylus roesleri sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small, with a maximum SVL of 75.3 mm; dorsal pattern consisting of dark nuchal band and 4–5 sometimes irregularly shaped dark transversal bands between limbs; ventrals in 34–40 longitudinal rows at midbody; continuous series of 20–28 precloacal and femoral pores in males; subcaudal scales transversally enlarged. It is the 15 th new reptilian taxon described as new from the karst forests of Phong Nha – Ke Bang National Park in the past decade. In addition, it is the 19 th species of Cyrtodactylus known from Vietnam and the third Cyrtodactylus occurring in sympatry in the karst forests of Phong Nha – Ke Bang National Park.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e. g., Smith 1920Smith , 1921Smith , 1935Taylor 1963;Ulber 1993;Darevsky & Szczerbak 1997;Das 1997;Manthey & Grossmann 1997;Chan-ard et al. 1999;Das & Lim 2000;Ziegler et al. 2002;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003David et al. 2004;Pauwels et al. 2004;Batuwita & Bahir 2005;Grismer 2005;Grismer & Leong 2005;Kraus & Allison 2006;Nguyen et al. 2006;Youmans & Grismer 2006;Grismer et al. 2007Heidrich et al. 2007;Hoang et al. 2007;Kraus 2007;Orlov et al. 2007;Grismer & Norhayati 2008;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;. ...
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We describe a new species of the genus Cyrtodactylus from southern Vietnam, based on voucher specimens collected from Dong Nai and Ba Ria – Vung Tau provinces. Cyrtodactylus cattienensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small, with a maximum SVL of 69 mm; neck band present, extending to the posterior margin of the eye; trunk, tail and limbs bearing irregularly shaped bands: 4–6 light bands on trunk and 4–12 white bands on tail; 16–22 irregular longitudinal rows of dorsal tubercles; 28–42 longitudinal rows of ventrals at midbody; lateral folds weakly developed or absent, without enlarged tubercles; tail not depressed, not enlarged at base, scales arranged in whorls; males with 6–8 precloacal pores in angular continuous series; a patch of enlarged precloacal scales present in both sexes; 3–8 slightly enlarged femoral scales on each hind limb in both sexes; femoral pores absent; precloacal groove absent; subcaudal scales small, not transversely enlarged.
... Some of the information on character states and their distribution in other species was obtained from De Rooij (1915), Smith (1930), Inger & King (1961), Taylor (1963), Dring (1979), Hikida (1990), Manthey & Grossman (1997), Das & Lim (2000), Grismer et al. (2008a); Rösler & Glaw (2008), and . Additional specimens examined are listed in the appendix. ...
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A new species of Bent-toed Gecko Cyrtodactylus guakanthanensis sp. nov. of the C. sworderi complex is described from a limestone forest in Perak, Peninsular Malaysia whose karst formations at the type locality are within an active quarry. Cyrtodactylus guakanthanensis sp. nov. can be distinguished from all other Sundaland species by having the following suite of character states: adult SVL 77.7-82.2 mm; moderately sized, conical, weakly keeled, body tubercles; tubercles present on occiput, nape, and limbs, and extend posteriorly beyond base of tail; 37-44 ventral scales; no transversely enlarged, median, subcaudal scales; proximal subdigital lamellae transversely expanded; 19-21 subdigital lamellae on fourth toe; abrupt transition between posterior and ventral femoral scales; enlarged femoral scales; no femoral or precloacal pores; precloacal groove absent; wide, dark postorbital stripes from each eye extending posteriorly to the anterior margin of the shoulder region thence forming a transverse band across the anterior margin of the shoulder region; and body bearing five (rarely four) wide, bold, dark bands. Destruction of the karst microhabitat and surrounding limestone forest will extirpate this new species from the type locality and perhaps drive it to complete extinction given that it appears to be restricted to the particular microhabitat structure of the type locality and is not widely distributed throughout the karst formations. As with plants and invertebrates, limestone forests are proving to be significant areas of high herpetological endemism and should be afforded special conservation status rather than turned into cement.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Smith 1920Smith , 1921Smith , 1935Ulber & Grossmann 1991;Darevsky & Szczerbak 1997;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009David et al. 2004David et al. , 2011Pauwels et al. 2004Pauwels et al. , 2013Nguyen S.N. et al. 2006Nguyen S.N. et al. , 2013Hoang et al. 2007;Orlov et al. 2007;Nazarov et al. 2008;Ngo 2008;Ngo & Bauer 2008;Rösler & Glaw 2008;Geissler et al. 2009;Mahony 2009;Ngo & Chan 2010;Ngo & Pauwels 2010;Ngo et al. , 2011Nguyen T.Q. et al. 2010;Sumontha et al. 2010;Schneider et al. 2011;Grismer et al. 2012;Nazarov et al. 2012;Ziegler et al. 2010Ziegler et al. , 2013Luu et al. 2014;. ...
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We describe a new species of the genus Cyrtodactylus from southern Vietnam, based on morphological and molecular datasets. Cyrtodactylus cucdongensis sp. nov. is described on the basis of seven specimens collected from Cuc Dong Cape, Khanh Hoa Province. The new species can be distinguished from the remaining bent-toed geckos by a combination of the following characters: maximum SVL 65.9 mm; 16–19 dorsal tubercle rows; 41–44 ventral scales at midbody; 5 or 6 precloacal pores in males, 4–6 pitted precloacal scales in females; no femoral pores; 6–13 enlarged precloacal scales; 5–9 enlarged femoral scales; no transversally enlarged subcaudals; dorsal pattern consisting of irregular dark bands. This is the 33rd species of Cyrtodactylus known from Vietnam.
... Comparisons. Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Rösler & Glaw 2008;Bauer et al. 2009Bauer et al. , 2010Ngo & 36), C. pulchellus Gray 1827 (33-39), C. redimiculus King 1962 (5-8+8-9), C. seribuatensis Youmans & Grismer 2006 (40-44), C. surin Chan-ard & Makchai, 2011 (34), C. tamaiensis Mahony 2009 (40), C. trilatofasciatus Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012 (41-46), C. variegatus (Blyth 1861) (32) and C. wetariensis (Dunn 1927) (11-12+12-13). ...
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We describe the third new species of the genus Cyrtodactylus from Ba Den Mountain, Tay Ninh Province, southern Vietnam based on a type series consisting of eleven specimens. Cyrtodactylus thuongae sp. nov. is distinguished from the remaining Indochinese Bent-toed Geckos by a combination of the following morphological characters: size medium, with a maximum SVL of 77.6 mm; dorsal pattern consisting of blotches between limbs and on occiput and a sometimes medially opened nuchal loop; dorsal tubercles in 16–18 irregular rows; ventrals in 29–44 longitudinal rows at midbody; lateral skin folds present, lacking tubercles; femoral pore series separated from precloacal pore series in males, 0–3 on each side; precloacal pores in males absent or only one; enlarged femoral scales and precloacal scales present, separated by a series of 15–16 smaller scales; postcloacal spurs two or three; subcaudal scales slightly enlarged. This is the 32nd described Cyrtodactylus species from Vietnam.
... The genus Cyrtodactylus ranges throughout most of tropical Asia, from India to the Philippines and south through the Indo-Australian Archipelago, northern Australia, New Guinea, and the Solomon Islands (Youmans & Grismer 2006). With over 170 recognized species, Cyrtodactylus is the most speciose genus of gekkonid lizards Rösler & Glaw 2008;Uetz & Hallerman 2010). Since 2008, at least four new species have been described from the Indonesian Archipelago Linkem et al. 2008;Oliver et al. 2009;Iskandar et al. 2011). ...
Article
Cyrtodactylus hikidai sp. nov. is a new gekkonid species described on the basis of 13 specimens collected from Mount Ranai, Bunguran Island, Indonesia. This large Cyrtodactylus (female SVL up to 102.2 mm, male SVL up to 100.6 mm) differs from all other Sunda Shelf species of Cyrtodactylus by the combination of a reticulated color pattern on the dorsum of the head, body banded, enlarged precloacal scales, six pore-bearing precloacal scales in a deep groove in both males and females, tubercles on forelimbs, hind limbs and occiput, absence of enlarged femoral scales, and absence of femoral pores.
... In their recent description of Cyrtodactylus thochuensis, Ngo and Grismer (2012) The dorsal pattern of Cyrtodactylus samroiyot sp. nov. is composed of three long dark bands (with only one strictly between limb insertions) separated by light interspaces of about half their width that readily distinguishes it from C. angularis (W-shaped marks), C. aurensis (light dorsal bands very short), C. bidoupimontis (blotches sometimes fused to form irregular bands), C. brevidactylus (blotches), C. buchardi (blotches), C. bugiamapensis (blotches), C. chauquangensis (three irregular bands slightly longer than inter-bands), C. chrysopylos (six irregular dark bands between limbs insertions), C. condorensis (irregular bands), C. cryptus (3-5 dark transverse bands between limbs insertions), C. durio (blotches), C. eisenmanae (light dorsal bands very short), C. elok (blotches), C. intermedius (Smith) (at least two dark bands fully within limbs insertions), C. irregularis (blotches), C. mandalayensis (blotches), C. martini (3 or four dark bands between limbs insertions), C. nigriocularis (light bands nearly not visible), C. oldhami (four longitudinal lines of spots), C. pageli (five dark bands between limb insertions), C. pantiensis (blotches), C. papilionoides (blotches), C. paradoxus Darevsky & Szczerbak (irregular narrow bands connected to a vertebral line), C. payacola (blotches), C. peguensis (spots), C. phuquocensis Ngo,Grismer & Grismer (3 dark bands between limbs insertions), C. pseudoquadrivirgatus (stripes or irregular bands, nuchal loop medially interrupted), C. quadrivirgatus (stripes), C. sanook (five or six irregular dark bands between limbs insertions), C. stresemanni (longitudinally elongated blotches), C. sumonthai (two bands fully within limbs insertions), C. surin (indistinct irregular dark bands), C. sworderi (longitudinal lines of spots), C. teyniei (blotches), C. thochuensis (three or four irregular dark bands between limb insertions), C. tiomanensis (two dark bands fully within limbs insertions), C. wakeorum (five narrow dark bands between limbs insertions), C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler (blotches or highly irregular bands, no nuchal loop), C. yangbayensis (four or five irregular dark bands between limbs insertions) and C. ziegleri (4-6 irregular dark bands between limbs insertions) (Bauer, 2002(Bauer, , 2003Rösler and Glaw, 2008;David et al., 2011;Grismer, 2011;Ngo and Grismer, 2012). ...
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We describe Cyrtodactylus samroiyot sp. nov. from a limestone relief in Sam Roi Yot District, Prachuap Khiri Khan Province, peninsular Thailand. It is characterized by a maximal known SVL of 66.9 mm; 17-18 longitudinal rows of dorsal tubercles; 33-34 longitudinal rows of ventrals across the abdomen between the ventrolateral skin folds; a continuous series of seven precloacal pores in males (six shallow precloacal pits in females); a series of slightly enlarged, poreless and pitless femoral scales; no precloacal groove nor depression; median row of transversely enlarged subcaudal scales; a complete nuchal loop; and a dorsal pattern consisting of three long dark brown bands, one above shoulders and two above abdomen.
... Comparisons are based on the original descriptions or descriptions provided in broader faunal and taxonomic publications (e.g., Grismer et al. 2008;Rösler & Glaw 2008;Bauer et al. 2009;Ngo & Pauwels 2010;Sumontha et al. 2010;Ziegler et al. 2010;David et al. 2011;Iskandar et al. 2011;Luu et al. 2011;Ngo 2011;Ngo & Chan 2011;Schneider et al. 2011;Nazarov et al. 2012;Ngo & Grismer 2012). ...
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We describe a new species of the genus Cyrtodactylus based on five adult specimens from Dai Lanh Cape, Tuy Hoa District, Phu Yen Province, southern Vietnam. Cyrtodactylus kingsadai sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: maximum SVL of 94 mm; dorsal pattern consisting of a dark nuchal loop, continuous or partly interrupted neck band and four in part irregulartransverse body bands between limbs; internasal single; dorsal tubercles in 17–23 irregular transverse rows; ventrals in 39–46 longitudinal rows at midbody; lateral skin folds present, without interspersed tubercles; precloacal pores 7–9 plus in total 3–7 femoral pores in males (1-4 femoral pores on each side) with precloacal and femoral pore series separated from each other by 7–9 poreless scales; enlarged femoral scales and precloacal scales present; postcloacal spurs three;subcaudal scales transversely enlarged. This is the 29th species of Cyrtodactylus known from Vietnam.
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This study provides a review of reptile research in Malaysia from the years 2000-2020 to reveal trends and biases that can be leveraged to improve future research and conservation initiatives. Overall, the number of papers published shows an upward trend with an average of 10.1 papers published per year. The clear majority of papers published throughout 2000-2020 were in the field of Systematics (45%, 96 papers), followed by Inventories (32%, 67 papers), Ecology (20%, 43 papers), and Conservation (3%, 6 papers). Within the Systematics category, most papers published between 2000-2011 were solely based on morphology, while integrative taxonomic studies (morphology + DNA) began to dominate from 2012 onwards. When analysed by location, most studies were conducted in Peninsular Malaysia and its associated offshore islands (78%) while the remaining 22% of studies were exclusive to Malaysian Borneo. Our results showed that research is heterogeneous across categories, focal taxa, methods, and location. Some of these biases are inherent to the nature of the study, while others represent areas for improvement. One obvious area that could be improved is the field of conservation, which constitutes only 3% of the studies and has so far focused almost exclusively on turtles. This study highlights gaps and deficiencies in research foci and provides a roadmap for future reptile research planning in Malaysia.
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A new bent-toed gecko species of the genus Cyrtodactylus is described herein from West Bali National Park on the island of Bali, Indonesia. The Cyrtodactylus from Bali have been recognized as C. fumosus for nearly a century. However, recent detailed examination of the type material of C. fumosus has revealed that Balinese Cyrtodactylus represent at least one different species. A morphological examination of the new species and samples from different biogeographic regions revealed that the new species is similar to C. seribuatensis from Pulau Seribuat in West Malaysia. Morphologically, the new species can be distinguished from its congener by having 40-43 femoro-precloacal pores (vs 10-12 precloacal pores, 14-16 femoral pores), 40-48 ventral scales between ventrolateral folds (vs 28-39), 24-29 rows of dorsal tubercles (vs 27-35), and presence of tubercles on lateral skinfold (vs absent).
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Lizards are ancestrally diurnal, and most of them remain so. Nocturnality is common among lizards, but the environmental factors associated with lizard nocturnal activity are still unknown. Here, we contrasted the ambient temperature and productivity hypotheses, where we predicted that cold temperatures will pose a stonger limit to nocturnal species richness than diurnal lizards. Moreover, we contrasted the relative importance of annual, day and night mean temperatures to pinpoint the drivers of nocturnal lizard richness.
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The binominal Cyrtodactylus fumosus has frequently been used for populations of bent-toed geckos occurring on some Indonesian islands, including Java, Bali, Sulawesi, and Halmahera. Unfortunately, incorrect usage of this name for different geographic lineages has resulted in confusion about the true identity of C. fumosus. Examination of the type specimen and additional specimens from Rurukan and Mount Masarang, North Sulawesi Province, Indone-sia, revealed that this population is distinct from other forms heretofore called 'fumosus' by a combination of unique morphological characters. In order to stabilize the taxonomy of C. fumosus sensu stricto, and to prevent further confusion, we provide a comprehensive redescription of this species, whose distribution we herein restrict to North Sulawesi. Cyrtodactylus fumosus is one of the most distinctive species among the six bent-toed geckos recorded from Sulawesi, and it differs from Sulawesi congeners by the presence of (1) precloacofemoral scales, including three pore-bearing scales on each thigh, separated from 10 or 11 pore-bearing scales in the precloacal region by 9-11 interscales in males, (2) a precloacal groove in adult males, (3) flat dorsal tubercles in 4-7 irregularly arranged longitudinal rows at midbody, and (4) a distinct lateral fold lacking tubercles. We also provide a revised identification key to the bent-toed gecko species of Sulawesi.
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We describe two new species of the Cyrtodactylus irregularis complex both based on phylogenetic analysis of 654 bp of COI mtDNA gene and morphological analyses of voucher specimens from Binh Phuoc and Lam Dong provinces, southern Vietnam. Cyrtodactylus bugiamapensis sp. nov. is described from the monsoon tropical forests of Bu Gia Map National Park, Binh Phuoc Province, and is distinguished from the remaining representatives of the C. irregularis complex by a combination of the following characters: (1) size medium, with a maximum SVL of 76.8 mm; (2) original tail relatively thin, longer than body; (3) presence of enlarged femoral scales without femoral pores; (4) preclocal groove lacking; (5) 36-46 longitudinal rows of ventral scales at midbody; (6) males with 7-11 precloacal pores in an angular continuous series; (7) absence of enlarged subcaudals; (8) dorsal pattern consisting of a dark neck band which can be medially divided, and irregular dark brown spots with bright white edges. Cyrtodactylus bidoupimontis sp. nov. is described from mountainous evergreen tropical forests of Bidoup - Nui Ba National Park, Lam Dong Province, and is most similar to C. irregularis sensu stricto from which it is distinguished by a combination of the following characters: (1) absence of enlarged, strongly keeled conical tubercles on the dorsal tail-base; (2) presence of flat rounded smooth to weakly keeled dorsal tubercles; (3) pallid dorsal head surface pattern lacking distinct dark brown irregular spots with light edges; and (4) elongated limbs. Phylogenetic analyses revealed the presence of a number of cryptic allopatric species within the C. irregularis complex. Long geological history and complicated relief of the Lang Bian plateau and surrounding areas might have shaped the present diversity within the C. irregularis complex. COI DNA-barcoding appears to be a useful tool to reveal cryptic diversity within the genus Cyrtodactylus.
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We describe a new forest-dwelling Cyrtodactylus (Gekkonidae) from Doi Suthep, Chiang Mai Province, northern Thailand. Cyrtodactylus doisuthep sp. nov. is characterized by a maximal known SVL of 90.5 mm; 19 or 20 longitudinal rows of dorsal tubercles; a continuous series of 34 or 35 enlarged femoro-precloacal scales, including six or seven pitted scales on each femur (male and females) separated by a diastema from six pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and six or seven irregular thin beige dorsal bands between limb insertions.
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The paper describes a new cave-dwelling species, Cyrtodactylus sanook, from Tham Sanook, southern Thailand, characterized by a maximum SVL of 79.5 mm; 18-20 longitudinal rows of dorsal tubercles at midbody; a continuous series of enlarged femoral and precloacal scales, no femoral pores, three or four precloacal pores in males (no pores in females), no precloacal groove nor depression; 19-20 subdigital lamellae on 4th toe; transversally enlarged subcaudal plates; and 6-7 irregular pale narrow dorsal bands between limb insertions. It seems endemic to this cave and is the 7th Thai Cyrtodactylus species that is known only from a cave environment.
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Fieldwork conducted throughout Timor-Leste in September 2004 and July 2009 resulted in a collection or recording of 263 herpetological specimens (100 amphibians, 163 reptiles), comprising at least seven species of frogs and toads, 20 species of lizards, seven species of snakes, two species of turtles, and one species of crocodile. Among the amphibians, the most frequently encountered species were toads (Duttaphrynus melanostictus), rice paddy frogs (genus Fejervarya), and rhacophorid treefrogs (Polypedates cf. leucomystax). All three variants of rice paddy frogs encountered represent undescribed species similar to Fejervarya verruculosa from neighboring Wetar Island. Records of Fejervarya cancrivora and Fejervarya limnocharis for Timor Island are apparently errors based on misidentification. We obtained voucher specimens for a total of 147 lizards and voucher photographs only for four specimens of Varanus timorensis. Aside from geckos frequently associated with human habitations (e.g., Gehyra mutilata, Gekko gecko, Hemidactylus frenatus, Hemidactylus platyurus), we discovered an as yet undescribed species of bent-toed gecko, genus Cyrtodactylus, in the Same valley. Our specimens of Hemidactylus platyurus are the first record of this species from Timor-Leste. Commonly encountered skinks included four-fingered skinks (genus Carlia), wedge skinks (genus Sphenomorphus), and night skinks (genus Eremiascincus). Notable among the 15 snakes collected was the frequency of pitvipers (Cryptelytrops insularis), which amounted to over 25% of all snakes. Our specimen of the wolfsnake Lycodon subcinctus is the first record of this species for Timor-Leste. Based on these findings, it appears that the biodiversity of amphibians and reptiles in this remote corner of Wallacea is much greater than previously thought, particularly with respect to scincid lizards. The detail we provide in the species accounts is designed to allow the use of this report as a preliminary field guide to the amphibians and reptiles of Timor-Leste. However, survey work is ongoing.
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