Taxonomic notes on the Harmochirina Simon, 1903 from South and South-East Asia (Aranei: Salticidae)

Abstract

A new species – Neaetha tomkovichi sp.n. (♂♀) from India (Andhra Pradesh) – is diagnosed and described. Three new synonyms of species names are proposed: Stichius albomaculatus Thorell, 1890 and Bianor leucostictus Thorell, 1890 with Bianor angulosus (Karsch, 1879), and Bianor incitatus Thorell, 1890 with Bianor balius Thorell, 1890. The generic name Stichius Thorell, 1890 is synonymised with Bianor Peckham et Peckham, 1886. The lectotype (♀) is designated for Modunda ghigii Caporiacco, 1949, and a new combination is proposed: Ureta ghigii (Caporiacco, 1949), comb.n. (ex Modunda). New faunistic records for seven Harmochirina species of the genera Bianor Peckham et Peckham, 1886, Harmochirus Simon, 1885 and Modunda Simon, 1901 are also provided.

Full text

© ARTHROPODA SELECTA, 2019
Arthropoda Selecta 28(1): 99–112
унистические находки для семи видов Harmochirina
из родов Bianor Peckham et Peckham, 1886, Harmo-
chirus Simon, 1885 и Modunda Simon, 1901.
Introduction
According to Maddison [2015], the salticid sub-
tribe Harmochirina Simon 1903 of the tribe Plexippini
Simon, 1901 consists of 15 valid genera. Although the
majority of these genera were either recently revised in
the scope of world fauna (e.g., Logunov [2001, 2009])
or described (e.g., Prószyński [1992]; Logunov [2000;
2001]), a number of Harmochirina species remain
known from their original descriptions only. Some of
these species (e.g., Bianor balius Thorell, 1890, B.
leucostictus Thorell, 1890) were transferred to the ge-
nus Stertinius Simon, 1890 (see Metzner [2018]; WSC
[2018]), belonging to the subtribe Simaethina in the
tribe Viciriini [Maddison, 2015]. One species — Stich-
ius albo-maculatus Thorell, 1890, which is the genero-
type of the monotypic genus Stichius Thorell, 1890 —
was mentioned as a possible junior synonym of Bianor
balius (see Logunov [2001: 281]) but is still consid-
ered a valid species [Metzner, 2018; Prószyński, 2017;
WSC, 2018].
The following four Harmochirina species are of an
obscure taxonomic status: Bianor balius described from
the holotype # from Sumatra [Thorell, 1890a] and
transferred to the genus Stertinius by Roewer [1955];
Bianor leucostictus described from the holotype #
from Sumatra [Thorell, 1890b] and transferred to the
genus Stertinius by Simon [1901]; Modunda ghigii
Caporiacco, 1949 described from several syntypes
(adult $ and three subadult ##) from Kenya by Capo-
riacco [1949] and is now listed in the genus Bianor
[WSC, 2018], although we failed to find out who made
that transfer; and Stichius albo-maculatus described
from the immature # from Sumatra [Thorell, 1890a].
ABSTRACT. A new species — Neaetha tomkovi-
chi sp.n. (#$) from India (Andhra Pradesh) — is diag-
nosed and described. Three new synonyms are pro-
posed: Stichius albomaculatus Thorell, 1890 and Bi-
anor leucostictus Thorell, 1890 with Bianor angulosus
(Karsch, 1879), and Bianor incitatus Thorell, 1890
with Bianor balius Thorell, 1890. The latter species is
removed from the genus Stertinius Simon, 1890, and
its original combination is re-instated: Bianor balius
comb.rev. The generic name Stichius Thorell, 1890 is
synonymised with Bianor Peckham et Peckham, 1886.
The lectotype ($) is designated for Modunda ghigii
Caporiacco, 1949, and a new combination is proposed:
Ureta ghigii (Caporiacco, 1949), comb.n. (ex Modun-
da). New faunistic records for seven Harmochirina
species of the genera Bianor Peckham et Peckham,
1886, Harmochirus Simon, 1885 and Modunda Si-
mon, 1901 are also provided.
How to cite this article: Logunov D.V. 2019. Taxo-
nomic notes on the Harmochirina Simon, 1903 from
South and South-East Asia (Aranei: Salticidae) // Ar-
thropoda Selecta. Vol.28. No.1. P.99–112. doi:
10.15298/arthsel. 28.1.08
РЕЗЮМЕ. Диагностирован и описан новый вид
Neaetha tomkovichi sp.n. (#$) из Индии (Андхра
Прадеш). Предложены три новых синонима: Stichius
albomaculatus Thorell, 1890 и Bianor leucostictus
Thorell, 1890 с Bianor angulosus (Karsch, 1879), и
Bianor incitatus Thorell, 1890 с Bianor balius Thorell,
1890. Последний вид перенесен из рода Stertinius
Simon, 1890, и ему возвращена оригинальная ком-
бинация Bianor balius comb.rev. Родовое название
Stichius Thorell, 1890 синонимизировано с Bianor
Peckham et Peckham, 1886. Обозначен лектотип ($)
для Modunda ghigii Caporiacco, 1949, и предложена
новая комбинация: Ureta ghigii (Caporiacco, 1949),
comb.n. (ex Modunda). Также приведены новые фа-
Taxonomic notes on the Harmochirina Simon, 1903 from South and
South-East Asia (Aranei: Salticidae)
Òàêñîíîìè÷åñêèå çàìåòêè î Harmochirina Simon, 1903 èç þæíîé
è þãî-âîñòî÷íîé Àçèè (Aranei: Salticidae)
Dmitri V. Logunov
Äìèòðèé Â. Ëîãóíîâ
The Manchester Museum, University of Manchester, Oxford Road, Manchester M13 9PL, UK; email: dmitri.v.logunov@manchesre.ac.uk
KEY WORDS. Araneae, description, distribution, jumping spiders, Harmochireae, new species, Oriental
Region, Plexippini, synonymy, taxonomy.
КЛЮЧЕВЫЕ СЛОВА. Araneae, новый вид, описание, Ориентальный регион, пауки-скакунчики,
Plexippini, распространение, синонимия, таксономия, Harmochireae.

100 D.V. Logunov
Figs 1–4. The male palp and right chelicera of Bianor angulosus (Karsch, 1879): 1 — the palp of the holotype male of B. leucostictus
Thorell, 1890, ventral view; 2 — ditto, retrolateral view; 3 — ditto, ventral view; 4 — original data labels of the holotype male of B.
leucostictus. Scale bars: (1, 2) 0.1 mm, (3) 0.25 mm.
Рис. 1–4. Пальпа самца и правая хелицера Bianor angulosus (Karsch, 1879): 1 — пальпа голотипа самца B. leucostictus Thorell,
1890, вид снизу; 2 — то же, вид сзади-сбоку; 3 — то же, вид снизу; 4 — оригинальные этикетки голотипа самца B. leucostictus.
Масштаб: (1, 2) 0,1 мм, (3) 0,25 мм.
The aims of the present paper are: (1) to clarify the
taxonomic validity and status of the aforementioned
four species based on re-examination of their types; (2)
to redescribe and/or map two of these species; and (3)
to diagnose and describe a new species of Neaetha
Simon, 1884 from India; and (4) to provide new fau-
nistic records for seven Harmochirina species from
South and South-East Asia.
Material and methods
This work is based on the specimens borrowed from or
deposited in the following museums: ISEA — Institute for
Systematics and Ecology of Animals, SB RAS, Novosibirsk,
Russia (curator: Galina N. Azarkina); MCSN — Museo
Civico di Storia Naturale, Genova, Italy (curator: Maria
Tavano); MCVR — Museo Civico di Storia Naturale, Vero-
na, Italy (curator: Roberta Salmaso); MMUE — Manchester
Museum of the University of Manchester, Manchester, UK
(curator: Dmitri V. Logunov); MSNF — Museo di Storia
Naturale, Firenze, Italy (curator: Luca Bartolozzi); NIBGE
— the National Institute for Biotechnology and Genetic
Engineering, Faisalabad, Pakistan; OUMNH — Oxford Uni-
versity Museum of Natural History, Oxford, UK (curator:
Zoë Simmons); PCMF — Personal spider collection of Mar-
io Freudenschuss (Wien, Austria); ZMMU — Zoological
Museum of the Moscow State University, Moscow, Russia
(curator: Kirill G. Mikhailov).
A part of the digital photographs (Figs 8–10, 14–16, 32–
36) were made at the Oxford University Museum of Natural
History (UK) with a Leica M165C stereo microscope (trin-
ocular) with automated z-stepper, Camera Leica DFC495,
Leica Application Suite (software to run above), and Heli-
con Focus 5.3 x64 as a processing software. Other digital
photographs (Figs 5–7, 24–31, 42–49) were made at the
World Museum of Liverpool (UK) by means of a Canon 6d
Mark ii Camera with a Canon MP-E 65mm Lens with Heli-
con Remote ver. 3.9.7W to control the StackShot 3X Macro
Rail and camera settings, and Helicon Focus 6.8.0 as a
processing software.

101
On the Harmochirina from S and SE Asia
Figs 5–7. General appearance of Bianor angulosus (Karsch, 1879): 5 — the holotype male of B. leucostictus Thorell, 1890, lateral
view; 6 — ditto, ventral view; 7 — ditto, dorsal view. Scale bars: 1 mm.
Рис. 5–7. Общий вид Bianor angulosus (Karsch, 1879): 5 — голотип самец B. leucostictus Thorell, 1890, вид сбоку; 6 — то же,
вид снизу; 7 — то же, вид сверху. Масштаб: 1 мм.
Abbreviations used in the text: Eyes: AME — anterior
median eye, PLE — posterior lateral eye(s). Leg segments:
Fm — femur, Mt — metatarsus, Pt — patella, Tr — tarsus,
Tb — tibia. Position of spines on legs: ap — apical, d —
dorsal, pr — prolateral, rt — retrolateral, v — ventral.
Others: a.s.l. — above sea level, D — described, S —
synonymized, T — transferred. For the leg spination the
system adopted is that used by Ono [1988]. The sequence of
leg segments in measurement data is as follows: femur +
patella + tibia + metatarsus + tarsus. All measurements are
in mm.
Taxonomy
Bianor angulosus (Karsch, 1879)
Figs 1–11, Map 1.
Bianor angulosus Karsch, 1879: 553 (D$; holotype $ in the
Natural History Museum of the Humbold University; examined,
see Logunov [2001]).
Stichius albo-maculatus Thorell, 1890a: 70 (holotype: imma-
ture # in the MCSN, examined). Syn.n.
Bianor leucostictus Thorell, 1890b: 158 (D#; holotype # in
the MCSN, examined). Syn.n.
Bianor trepidans Thorell, 1895: 334 (D$; lectotype $ in the in
the Swedish Museum of Natural History in Stockholm; examined,
see Logunov [2001]). Synonymized with B. angulosus by Logunov
[2001].
Stertinius leucostictus: Simon, 1901: 839.
Bianor hotingchiehi Schenkel, 1963: 434, figs 249a–f (D#;
holotype # in the Natural History Museum in Paris; examined, see
Logunov [2001]). Synonymized with B. angulosus by Logunov
[2001].
Bianor simoni Żabka, 1985: 204, figs 30–34 (D#; holotype #
in the Institute of Zoology in Warszawa; examined, see Logunov
[2001]). Synonymized with B. angulosus by Logunov [2001].
Stichius albomaculatus: Roewer, 1955: 1662; Prószyński, 1984:
57 (#).
Bianor incitatus (nec Thorell, 1890b; misidentified): Prószyńs-
ki, 2017: 49, figs 22T1 (#).
For a complete reference list of this species see WSC [2018].
TYPES. HOLOTYPE # of Bianor leucostictus Thorell, 1890
(MCSN; Figs 1–7) from “Sumatra, Ajer Mancior, VIII-1878-Bec-
cari” [apparently, nr. Bukittinggi in Padang Province, West Sumatra
(c. 0°19′55.4″S, 100°23′06.7″E); collected by Odoardo Beccari, an
Italian naturalist who travelled for many years in Malaysia and
Indonesia]. — HOLOTYPE immature # of Stichius albo-maculatus
Thorell, 1890 (MCSN; Figs 8–11) from “Siboga, Sumatra, 1886,
E. Modigliani” [apparently, Sibolga City (c. 1°44′N, 98°46′E) in
North Sumatra, Indonesia].
OTHER MATERIAL. THAILAND: 1# (ISEA; det. G.N. Azarki-
na, 2018), Province Ta Phraya, NP La Lu (c. 14°02′N, 102°34′E),
122 m a.s.l., 1–4.08.2012, A.A. Gurina.
COMMENTS. Despite in the original description of S.
albo-maculatus T. Thorell [1890a: 70] it is stated that he
examined a single female (“Singulum exemplum femineum
nondum adultum pulchrae hujus araneolae in Sumatra (Si-
bolga) cepit Modigliani”), the holotype of S. albo-macula-

102 D.V. Logunov
Figs 8–11. General appearance of Bianor angulosus (Karsch, 1879): 8 — carapace of the holotype male of Stichius albo-maculatus
Thorell, 1890, dorsal view; 9 — abdomen of S. albo-maculatus, dorsal view; 10 — carapace of S. albo-maculatus, ventral view; 11 —
original data labels of the holotype male of S. albo-maculatus. Scale bars: 0.5 mm.
Рис. 8–11. Общий вид Bianor angulosus (Karsch, 1879): 8 — головогрудь голотипа самца Stichius albo-maculatus Thorell, 1890,
вид сверху; 9 — брюшко S. albo-maculatus, вид сверху; 10 — головогрудь S. albo-maculatus, вид снизу; 11 — оригинальные
этикетки голотипа самца S. albo-maculatus. Масштаб: 0,5 мм.
tus is actually an immature (subadult) male (Figs 8–10).
This male has the dorsum with a colour pattern that is typical
of B. angulosus: viz. yellow-brown, with two pairs of elon-
gated white spots and a transverse white stripe at its front
edge (Fig. 9; cf. Fig. 7 and figs 60, 66 in Logunov [2001]).
The type locality of S. albomaculatus lies in Sumatra, from
where B. angulosus has also been recorded [Logunov, 2001].
Moreover, the mature male from Sumatra (Kampong) hith-
erto identified by T. Thorell as S. albomaculatus (deposited
in Stockholm) tuned out to be B. angulosus (see Logunov
[2001: 231]). Thus, although, the name S. albo-maculatus
could be treated as a nomen dubium (because the holotype is
an immature specimen), it is safe and better to consider it a
junior synonym of B. angulosus.
The taxonomic status of the genus Stichius itself no
longer remains unclear [Roewer, 1955: “nicht zu deuten”;
Prószyński, 2017]; this generic name is indeed a junior
synonym of Bianor.
The examined holotype male of Bianor leucostictus (Figs
1–7) possess all the diagnostic characters of and is identical
to B. angulosus (cf. figs 57, 58 in Logunov [2001]): viz., the
papal tibia as long as the cymbium (Figs 1, 2), the round
tegulum with a well-developed large membranous area, and
the same embolic length and its position (Fig. 1). Therefore,
it is safe to conclude that the name B. leucostictus is to be
synonymised with B. angulosus.
Logunov [2001: 231] re-examined the male from Sumatra
(Kampong), which was originally identified by T. Thorell as
Stichius albomaculatus, and listed it under the ‘Material’
examined for Bianor angulosus; this specimen is deposited
in the Swedish Museum of Natural History in Stockholm
(no. 1819). Earlier, the same male was also re-examined and
illustrated by Prószyński [1984: 57]. By unknown editorial
mistake, the latter Prószyński’s work was included in the
reference list of Bianor incitatus (see Logunov [2001: 236])
rather than be that of B. angulosus. Hence, the non-type
male examined and named by Thorell as S. albomaculatus
actually belongs to B. angulosus, and both illustrated records
of this male by Prószyński [1984: 57, 2017: 49, fig. 22T1]
should be included in the reference list of B. angulosus and
removed from that of B. incitatus, where they are currently
listed [WSC, 2018].
DISTRIBUTION. It is a rather widespread Oriental spe-
cies (Map 1) recorded/described to date under several names:
from Bhutan and southern India (Orissa) in the west [Lo-
gunov, 2001], southward to Sri Lanka [Karsch, 1879: sub
Ballus a.; Logunov, 2001]; eastward to throughout Bang-
ladesh and Myanmar to southern China [Schenkel, 1963;
Peng et al., 1993; Song et al., 2012, etc.: all sub. B. hot-
ingchiehi] and northern Vietnam [Żabka, 1985: sub. B. hot-
ingchiehi and B. simony] in the east, and south-eastward
throughout Thailand and southern Vietnam, to Indonesia
(Sumatra, Java) and Malaysia [Thorell, 1890a: sub. Stichius
albo-maculatus; Thorell, 1890b: sub. B. leucostictus; Lo-
gunov, 2001; Prószyński, 2017: sub. B. incitatus].
DESCRIPTION. See Logunov [2001].
Bianor balius Thorell, 1890, comb.rev.
Figs 12–17, Map 2.
Bianor balius Thorell, 1890a: 73 (D#; holotype # in the
MCSN; examined).
Bianor balius: Thorell, 1892: 256; Simon, 1901: 638.
Bianor incitatus Thorell, 1890b: 159 (D$; lectotype $ in the
Swedish Museum of Natural History in Stockholm; examined, see
Logunov [2001]). Syn.n.

103
On the Harmochirina from S and SE Asia
Map 1. Collecting localities of Bianor angulosus (Karsch, 1879) (circles) and Neaetha tomkovichi sp.n. (asterisk).
Карта 1. Точки находок Bianor angulosus (Karsch, 1879) (кружки) и Neaetha tomkovichi sp.n. (звездочка).
Bianor incitatus: Thorell, 1892: 259 ($); Simon, 1901: 638
($); Logunov, 2001: 236–240, figs 87–104 (T $ from Stertinius,
S#); Yin et al., 2012: 1332, figs 720a–d (#); Prószyński, Deele-
man-Reinhold, 2013: 117, figs 21–24 (#); Suguro, 2013: 5, figs
1–7 (#$).
Stertinius balius: Roewer, 1955: 1011, 1435.
Stertinius incitatus: Roewer, 1955: 1011, 1435.
Bianor carli Reimoser, 1934: 506, fig. 27 (D$; holotype $ in
the Natural History Museum in Geneve; examined, see Logunov
[2001]). Synonymized with B. incitatus by Logunov [2001].
Bianor obak Berry, Beatty et Prószyński, 1996: 220–222, figs
18–24, map 2 (D#$; holotype $ in the Bishop Museum, Honolulu,
Hawaii; examined, see Logunov [2001]). Synonymized with B.
incitatus by Logunov [2001].
Bianor maculatus (nec Keyserling, 1883; misidentified): Peng,
1989: 158, figs 1A–C (#); Peng et al., 1993: 29–30, figs 46–49
(#); Song, Zhu, Chen, 1999: 506, figs 289K, 324N (#).
TYPE. HOLOTYPE # (MCSN; Figs 12–17) from Singalang
[apparently, Singgalang Mt. (c. 0°23′24″S, 100°19′51″E)], [West]
Sumatra, [Indonesia], 1878, coll. O[dorado] Beccari.
OTHER MATERIAL. INDONESIA: 1# (MCSN), “Gunung Sitoli
Is. Nias” [apparently, Gunungsitoli, the capital city of Nias Regen-
cy of North Sumatra (c. 1°17′N, 97°37′E)], 1886, E. Modigliani
[this # was also mentioned by Thorell [1890a] in the original
description of B. balius]. — CAMBODIA: 1 $ (PCMF), Siem Reap,
Angkor Wat (13°26.370′N, 103°51.602′E), c. 6 m a.s.l., hand
collecting from a house wall, 1.12.2014, M. Freudenschuss.
DIAGNOSIS. Of the Bianor species known from both
sexes, B. balius is most similar to B. albobimaculatus (Lu-
cas, 1846) (cf. figs 4–8, 13–18, 19–27, 36–46 in Logunov
[2001]) and B. wunderlichi Logunov, 2001 (cf. figs 9–12,
28–35 in Logunov [2001]). The males of B. balius can be
distinguished from these species by the following charac-
ters: the absence of the membranous area of the tegulum
(present in both related species), the markedly more extend-
ed antero-prolateral edge of the tegulum (arrowed in Fig.
12) and the colour pattern of dorsum (Fig. 14; a pair of white
spots in B. balius which are absent from B. albobimaculatus
and two pairs of white spots and a number of irregular white
spots in B. wunderlichi). The females of B. balius were
diagnosed by Logunov [2001: sub B. incitatus], and can
reliably be distinguished from those of all other Bianor
species by the widest first loop of the insemination ducts
[Logunov, 2001: figs 94, 101, 102].
COMMENTS. To date, B. incitatus has been well-de-
scribed, illustrated and known from both sexes and many
localities, with a number of the studied samples containing
both sexes collected together (see Logunov [2001: 236–
237]). As is evident from the present study, the holotype
male of B. balius possesses all the diagnostic characters of
B. incitatus (viz., the flat, transverse-ovoid tegulum with the
visibly extended antero-prolateral edge and no membranous
area of the tegulum) and is identical to the latter species (cf.
Fig. 12 and figs 89, 90 in Logunov [2001]; figs 23, 24 in
Prószyński & Deeleman-Reinhold [2013]; or figs 23, 24 in
Berry et al. [1996: sub. B. obak]). Thus, it is safe to con-
clude that (1) B. balius comb.rev. is indeed a member of the
genus Bianor rather than of Stertinius (contra Roewer [1955])
and its original combination is to be re-instated; and (2) B.
balius is to be considered a senior synonym of B. incitatus.

104 D.V. Logunov
Figs 12–17. Copulatory organ and general appearance of the holotype male of Bianor balius Thorell, 1890: 12 — palp, ventral view;
13 — left chelicera, ventral view; 14 — body, dorsal view; 15 — ditto, lateral view; 16 — ditto, ventral view; 17 — original data labels.
Scale bars: (12, 13) 0.1 mm, (14–16) 1 mm.
Рис. 12–17. Копулятивные орган и общий вид голотипа самца Bianor balius Thorell, 1890: 12 — пальпа, вид снизу; 13 — левая
хелицера, вид снизу; 14 — тело, вид сверху; 15 — то же, вид сбоку; 16 — то же, вид снизу; 17 — оригинальные этикетки.
Масштаб: (12, 13) 0,1 мм, (14–16) 1 мм.
Although both species names were published in the same
year and month, September 1890, the exact publication date
of B. balius is September 4th, 1890 (see Thorell [1890a: 4]),
whereas the name B. incitatus was published on “pridie Kal.
Sept. 1890” [Thorell, 1890b: 132], which means “on the last
day, September 1890”; hence the name B. balius was pub-
lished slightly earlier and has a priority over the name B.
incitatus.
DISTRIBUTION. It is a rather widespread Oriental spe-
cies (Map 2) recorded/described to date under many differ-
ent names: from India (Mysore, Madya Pradesh and Megha-
laya) and Bhutan [Reimoser, 1934: sub. B. carli; Logunov,
2001: sub. B. incitatus], southward to Sri Lanka [Logunov,
2001: sub. B. incitatus]; eastward to southern China (Yun-
nan, Guanxi and Hunan) [Peng et al., 1993: sub. B. macula-
tus; Yin et al., 2012 and Logunov, 2001: both sub. B.
incitatus] and Japan (Okinawa) [Logunov, 2001 and Sug-
uro, 2013: both sub. B. incitatus], and south-eastward to
Indonesia, Malaysia [Thorell, 1890a,b, 1892; Logunov, 2001
and Prószyński, Deeleman-Reinhold, 2013: both sub. B.
incitatus] and the Caroline Islands [Berry et al., 1996: sub.
B. obak].
DESCRIPTION. MALE (the holotype; Figs 12–16). Mea-
surements: Carapace: 2.00 long, 1.73 wide, 1.10 high. Ab-
domen: 2.30 long, 1.75 wide. Ocular area: 1.25 long, 1.26
wide anteriorly, 1.68 wide posteriorly. Cheliceral length
0.80. Clypeal height 0.15. Diameter of AME 0.40. Length of
leg segments: I 1.55 + 1.08 + 1.15 + 0.90 + 0.65 (5.33); II
1.03 + 0.63 + 0.55 + 0.58 + 0.40 (3.19); III 1.03 + 0.63 +
0.68 + 0.70 + 0.40 (3.44); IV 1.23 + 0.59 + 0.70 + 0.80 +
0.48 (3.80). Leg formula I,IV,III,II. Leg spination: I: Fm d
0-0-1-2; Pt pr 0-1-0; Tb v 0-0-1-2-2ap; Mt v 2-2ap. II: Fm d
0-0-1-2; Tb pr 0-1, v 1-1; Mt v 2-2ap. III: Fm d 3ap; Tb pr
and rt 0-1, v 1ap; Mt pr and rt 2ap. IV: Fm d 1/2ap; Tb pr
and rt 0-0-1; Mt pr and rt 1ap. Coloration (in alcohol, the
specimen is visibly faded; Figs 14–16). Carapace russet and
shagreen, sparsely covered with white recumbent scales;
dark brown around eyes. Sternum yellowish russet, covered

105
On the Harmochirina from S and SE Asia
Map 2. Collecting localities of Bianor balius Thorell, 1890.
Карта 2. Точки находок Bianor balius Thorell, 1890.
with white hairs. Labium and endites yellowish russet, with
yellowish tips. Chelicerae yellowish russet. Abdomen: dor-
sum light brown, covered with a shining scutum, and with a
pair of small white spots in its rear third; sides yellowish
brown; venter brownish yellow. Book-lung covers and spin-
nerets yellow. Leg I stronger and longer than other legs,
russet, with yellowish tarsi. Legs II–IV: femora brownish,
patellae and tibiae yellowish brown, metatarsi and tarsi light
yellow. Palps yellowish brownish. Palpal structure is typical
of Bianor species (cf. Logunov, 2001), as shown in Fig. 12:
RTA singular, well-developed; tegulum flat, transverse-ovoid,
with its antero-prolateral edge visibly extended; embolus
thread-like, poorly visible, originating at about seven o’clock.
FEMALE. See Logunov [2001: sub. B. incitatus].
Neaetha tomkovichi sp.n.
Figs 18–31, Map 1.
TYPES: HOLOTYPE # and paratype $ (ZMMU) from India,
Andhra Pradesh, nr. Bapatla (15.86°N, 80.49°E), 20–28.02.2014,
K.P. Tomkovich.
ETYMOLOGY. The species is named after the collec-
tor, Mr Konstantin P. Tomkovich (Moscow, Russia).
DIAGNOSIS. The new species is most similar to two
Mediterranean congeners: Neaetha membrosa (Simon, 1868)
and N. absheronica Logunov et Guseinov, 2002. From the
former species (cf. figs 1–15 in Logunov [1996]), N. tomk-
ovichi sp.n. differs in having the straight tibial apophysis
(Fig. 19; bent apically in N. membrosa), the much smaller
epigynal pocket and the central position of the copulatory
organs (Fig. 20; shifted to the epigastric furrow in N. memb-
rosa). From N. absheronica (cf. figs 4A–S in Lecigne [2016]),
the new species differs in having the slightly shorter, straight
embolus (Fig. 18; visibly bent mediad at its tip in N. absher-
onica), the round bulbus (visibly elongated-ovoid in N. ab-
sheronica), the smaller, singular epigynal pocket (Fig. 20;
notched at its distal end in N. absheronica) and the less-
chambered spermathecae (Fig. 21).
DISTRIBUTION. The type locality only (Map 1: aster-
isk). To date, the genus Neaetha has been known from the
Afrotropical Region and the Mediterranean area (including
the Near East and the Caucasus) of the Palaearctic Region
[Logunov, 2009; WSC, 2018]. It is the first record of the
genus from South Asia and the Oriental Region.
DESCRIPTION. MALE (holotype; Figs 18, 19, 24–27).
Measurements: Carapace: 1.43 long, 1.25 wide, 0.80 high.
Abdomen: 1.28 long, 0.98 wide. Ocular area: 0.90 long,
1.03 wide anteriorly, 1.18 wide posteriorly. Cheliceral length
0.48; promargin with two teeth, retromargin with one tooth.
Clypeal height 0.19. Diameter of AME 0.35. Length of leg
segments: I 1.00 + 0.60 + 0.68 + 0.68 + 0.35 (3.31); II 0.58 +
0.33 + 0.33 + 0.25 + 0.25 (1.74); III 1.23 + 0.53 + 0.60 +
0.45 + 0.33 (3.14); IV 0.70 + 0.35 + 0.31 + 0.35 + 0.30
(2.01). Leg formula I,III,IV,II. Leg spination: I: Fm d 0-0-1-
2; Tb v 2-2-2ap; Mt v 2-2ap. II: Fm d 0-1-3; Pt pr 0-1-0; Tb
pr 1-1, v 1-1ap; Mt v 2-2ap. III: Fm d 0-0-1-3; Pt pr and rt 0-
1-0; Tb d 1-0-0, pr and rt 1-1, v 2ap; Mt pr 1-1ap, rt 2-1ap, v
2ap. IV: Pt rt 0-1-0; Tb pr and rt 0-1, v 2ap; Mt pr 2ap, rt 1-
2ap. Coloration (in alcohol; Figs 24–27). Carapace yellow-
ish brownish, covered with white recumbent scales; sides
are darker (brown) with a wide yellow longitudinal marginal
band on each side (Fig. 27); thorax with a median longitudi-
nal brown stripe; there are two longitudinal thin white lines
of scales running along the ALE-PLE line on each side of
the carapace (Fig. 24). Black around eyes. Clypeus densely
covered with white hairs (Fig. 25). Sternum yellow, with a

106 D.V. Logunov
Figs 18–23. Copulatory organs of Neaetha tomkovichi sp.n., the holotype male (18, 19) and the paratype female (20–23): 18 —
bulbus, ventral view; 19 — male palp, retrolateral view; 20 — epigyne, ventral view; 21 — spermathecae, dorsal view; 22 — female right
chelicerae, ventral view; 23 — diagrammatic course of the insemination ducts. Scale bars: 0.1 mm.
Рис. 18–23. Копулятивные органы Neaetha tomkovichi sp.n., голотип самец (18, 19) и паратип самка (20–23): 18 — бульбус,
вид снизу; 19 — пальпа самца, вид сбоку-сзади; 20 — эпигина, вид мнизу; 21 — сперматека, вид сверху; 22 — правая хелицера
самки, вид снизу; 23 — схематический ход осеменительных канальцев. Масштаб: 0,1 мм.
brown band along its margins, covered with white hairs.
Labium brown. Endites yellowish brownish, with white apex-
es. Chelicerae brownish yellow, anteriorly covered with
sparse white scales. Abdomen yellow, with a brown reticu-
lated pattern (Figs 24, 27). Book-lung covers brown. Spin-
nerets contrastingly grey-brown. Leg I larger than others and
visibly swollen; its Fm brownish yellowish, the remaining
segments darker, orange-brown and covered with black and
white hairs. Legs II-IV yellow, with brownish rings and
patches. Male palp: Fm and Pt yellow, Tb, cymbium and
bulbus brownish; palpal structure as in Figs 18, 19: both
palps of the holotype are visibly expanded; Tb relatively
short, with a wide and straight tibial apophysis directed
anteriad; bulbus round; embolus straight.
FEMALE (paratype; Figs 20–23, 28–31). Measurements:
Carapace: 1.93 long, 1.53 wide, 0.94 high. Abdomen: 2.00
long, 1.78 wide. Ocular area: 1.05 long, 1.23 wide anterior-
ly, 1.38 wide posteriorly. Cheliceral length 0.60; promargin
with two teeth, retromargin with one tooth (Fig. 22). Clypeal
height 0.20. Diameter of AME 0.43. Length of leg seg-
ments: I 0.93 + 0.58 + 0.63 + 0.45 + 0.33 (2.92); II 0.70 +
0.45 + 0.38 + 0.30 + 0.26 (2.09); III 1.48 + 0.73 + 0.73 +
0.55 + 0.40 (3.89); IV 0.83 + 0.40 + 0.43 + 0.43 + 0.35
(2.44). Leg formula III,I,IV,II. Leg spination: I: Fm d 1ap;
Tb v 2-2-2ap; Mt v 2-2ap. II: Fm d 1ap; Pt pr 0-1; Tb v 1-1;
Mt v 2-2ap. III: Fm d 0-0-0-1-1-3; Pt pr and rt 0-1-0; Tb pr
1-1, rt 1-1-1, v 1ap; Mt pr 1-2ap, rt 2-2ap. IV: Pt rt 0-1-0; Tb
rt 1-1; Mt pr 2ap, rt 1-2ap. Coloration (in alcohol; Figs 28–
31), as in the male, but differs as follows: body colour
lighter, predominatly yellow, abdomen ventrally and its sides
yellow, book-lung covers and palps yellow. Epigyne and
spermathecae as in Figs 20–22: epigynal plate flat, with a
small blind-ending epigynal pocket in its centre; copulatory
openings small, covered with round flaps; insemination dusts
and receptacles thick-walled and merged in a kind of a
multi-chambered mass, which is typical of all Neaetha spe-
cies.
Ureta ghigii (Caporiacco, 1949), comb.n.
Figs 32–41.
Modunda ghigii Caporiacco, 1949: 484, fig. 96 (D$; lectotype
$ in the MCVR; examined).
TYPES: LECTOTYPE $ (MCVR, 647; Figs 32–34, 37–41; desig-
nated here) from [Kenya], Elmenteita [Lake], 100 km N [apparent-
ly, Baringo County (c. 0°30′16.1″N, 36°10′55.5″E)], m 1550 [a.s.l.],
XII.1945, Toschi Menegnetti. PARALECTOTYPE subadult # (MSNF,
647; Figs 35, 36; designated here) from Kenya, Elmenteita, XII.1945,
[Toschi] Menegnetti.
DIAGNOSIS. The female of U. ghigii (Figs 37–39)
differs from that of U. quadrispinosa (Lawrense, 1928) (cf.
figs 173–175 in [Wesołowska, Haddad, 2013] and figs 37,
38 in Azarkina & Foord [2013: sub Euophrys q.]) in having
the epigynal notch in which the singular epigynal pocket is
situated (the notch is absent and the double epigynal pocket
in U. quadrispinosa), the larger copulatory openings form-

107
On the Harmochirina from S and SE Asia
Figs 24–31. General appearance of Neaetha tomkovichi sp.n., the holotype male (24–27) and the paratype female (28–31): 24, 31 —
body, dorsal view; 25, 28 — clypeus, frontal view; 26 — body, ventral view; 27 — ditto, lateral view; 29 — abdomen, dorsal view; 30 —
carapace, ventral view. Scale bars: (24, 26, 27, 29–31) 1 mm, (25, 28) 0.5 mm.
Рис. 24–31. Общий вид Neaetha tomkovichi sp.n., голотип самец (24–27) и паратип самка (28–31): 24, 31 — тело, вид сверху;
25, 28 — клипеус, вид спереди; 26 — тело, вид снизу; 27 — то же, вид сбоку; 29 — брюшко, вид сверху; 30 — головогрудь, вид
снизу. Масштаб: (24, 26, 27, 29–31) 1 мм, (25, 28) 0,5 мм.
ing almost a round central depression and the straight in-
semination ducts directed laterad (convoluted and directed
anteriad in U. quadrispinosa).
COMMENTS. With the absence of the male, it is not
easy to assign M. ghigii to a correct genus. Yet, it is obvious
that the species belongs neither to Modunda Simon, 1901,
as none of the four diagnostic characters of the latter genus
(see Logunov [2001: 270]) can be found in M. ghigii, nor to
the Harmochireae (sensu Logunov [2001, 2009]). The spe-
cies seems to belong to the subtribe Thiratoscirtina of the
tribe Aelurillini (sensu Maddison [2015]) containing 14 Afro-
tropical genera, and, in my opinion, could be provisionally

108 D.V. Logunov
Figs 32–36. General appearance of Ureta ghigii (Caporiacco, 1949), the lectotype female (32–34) and the paralectototype immature
male (35, 36): 32, 36 — body, dorsal view; 33 — ditto, lateral view; 34, 35 — ditto, ventral view. Scale bars: 1 mm.
Рис. 32–36. Общий вид Ureta ghigii (Caporiacco, 1949), лектотип-самка (32–34) паралектотип, ювенильный самец (35, 36): 32,
36 — тело, вид сверху; 33 — тоже, вид сбоку; 34, 35 — тоже, вид снизу. Масштаб: 1 мм.
assigned to the genus Ureta Wesołowska et Haddad, 2013.
M. ghigii is a medium-sized salticid (body length 4.88 mm),
with unidentate chelicerae (Fig. 40) and the conformation of
the copulatory organs being similar to that of the Ureta
generotype – U. quadrispinosa from South Africa — name-
ly, in having the well-developed epigynal pocket, the wid-
ened entrances of the copulatory openings and the rather
long insemination ducts ended up by the retort-shaped re-
ceptacles (Figs 37–39; cf. figs 173–175 in Wesołowska &
Haddad [2013] and figs 37–39 in Azarkina & Foord [2013:
sub Euophrys q.]). Therefore, it seems reasonable to pro-
pose a new combination: Ureta ghigii (Caporiacco, 1949),
comb.n.
DISTRIBUTION. The type locality only.
DESCRIPTION. MALE unknown.
FEMALE (lectotype; Figs 32–34, 37–41). Measurements:
Carapace: 2.25 long, 1.58 wide, 0.95 high. Abdomen: 2.63
long, 2.38 wide. Ocular area: 1.03 long, 1.28 wide anterior-
ly, 1.29 wide posteriorly. Cheliceral length 0.58; promargin
with two teeth, retromargin with one tooth (Fig. 40). Clypeus
not marked. Diameter of AME 0.40. Length of leg segments:
I 1.15 + 0.68 + 0.70 + 0.60 + 0.38 (3.51); II 1.00 + 0.63 +
0.58 + 0.55 + 0.35 (3.11); III 1.08 + 0.55 + 0.60 + 0.70 +
0.45 (3.38); IV 1.38 + 0.68 + 0.90 + 0.88 + 0.48 (4.32). Leg
formula IV,I,III,II. Leg spination: I: Fm d 0-1-1-1; Tb pr 0-
0-1, v 2-2ap; Mt v 2-2ap. II: Fm d 0-1-1-2; Tb v 1-0; Mt v 2-
2ap. III: Fm d 0-1-1-3; Tb pr and rt 0-1, v 1ap; Mt pr and rt
2ap, v 1-0-0. IV: Fm d 0-1-1-1; Tb v 1-0-2ap; Mt pr and rt
1ap, v 2ap. Coloration (in alcohol, the specimen is visibly
faded; Figs 32–34). Carapace yellowish brown, shagreen,
with a pale yellow line in the middle of the thorax; eye field
darker (brown); carapace sparsely covered with white re-
cumbent scales. Sternum and chelicerae yellowish brown.
Endites and labium yellowish brown, with white apexes.
Abdomen slightly flattened and faded, greyish; dorsal co-
lour pattern of reddish scales is poorly visible due to the
poor condition of the specimen. Book-lung covers and spin-
nerets yellowish. All legs and palps yellowish, with no co-

109
On the Harmochirina from S and SE Asia
Figs 37–41. Copulatory organ of the lectotype female of Ureta ghigii (Caporiacco, 1949): 37 — epigyne, ventral view; 38 —
spermathecae, dorsal view; 39 — ditto, ventral view; 40 — right chelicerae, ventral view; 41 — original data label from two sides. Scale
bars: 0.1 mm.
Рис. 37–41. Копулятивные орган лектотипа-самки Ureta ghigii (Caporiacco, 1949): 37 — эпигина, вид снизу; 38 — сперматека,
вид сверху; 39 — тоже, вид снизу; 40 — правая хелицера, вид снизу; 41 — оригинальная этикетка с двух сторон. Масштаб: 0,1 мм.
lour pattern. Epigyne and spermathecae as shown in Figs
37–39: epigynal plate with two large and deep copulatory
openings; proximal edge of the epigynal plate overhangs the
epigastric furrow and bears a deep notch having the well-
developed, singular pocket in the middle; the insemination
ducts with wide, well-developed entrances followed by nar-
row tubes ended up with retort-shaped receptacles bearing
the fertilization ducts.
New faunistic records
The following sections present new faunistic records
for seven Harmochirina species, of which the majority,
except for Bianor albobimaculatus and Harmochirus
brachiatus, remain known to date from few localities
only.
Bianor albobimaculatus (Lucas, 1846)
MATERIAL. PAKISTAN: 1 #, 1 $ (MMUE), Lahore [c.
31°32′42.9″N, 74°19′54.9″E], rice field, 9.08.2005, A. Butt; 1 #,
2 $$ (ZMMU), NWF, Peshawar, Forestry Campus of Agricultural
University [c. 34°01′00.0″N, 71°28′44.6″E], 14–26.08.2004, S.V.
Ovtchinnikov; 1 # (MMUE), Pakistan, Khushâb Distr. [c. 32°
19′40.1"N, 72°18′30.5"E], in grass, 2014, G. Rasool; 2 ##, 1 $
(ISEA, 001.8258), Gahkuch Paeen (36.176°N, 73.762°E), 1861 m
a.s.l., rural environments, 10.09.2018, D. Jablonski; 1 #, 2 $$
(MMUE), Karachi (?) [c. 25°13′34.8″N, 67°06′11.9″E], 2014, M.
Kazim; 1 #, 1 $, 2 imm. (NIBGE; det. G.A. Blagoev, 2018),
Punjab, Sahiwal (30.65°N, 73.1°E), 173 m a.s.l., 12.08.2010, Q.
Abbas; 5 ##, 9 $$, 3 imm (NIBGE; det. G.A. Blagoev, 2018),
Punjab, Faisalabad, NIBGE (31.3833°N, 73.0167°E), 182 m a.s.l.,
12.11.2010, S. Akhtar; 1 # (NIBGE; det. G.A. Blagoev, 2018),
Punjab, Sargodha (32.0667°N, 72.6667°E), 192 m a.s.l., 14.06.2010,
S. Akhtar; 1 # (NIBGE; det. G.A. Blagoev, 2018), Punjab, Lahore,
Changamanga (31.0833°N, 73.95°E), 197 m a.s.l., 17.07.2010, S.
Akhtar; 2 imm. (NIBGE; det. G.A. Blagoev, 2018), Punjab, Nan-
kana Sahib (31.45°N, 73.7°E), 193 m a.s.l., 28.05.2010, S. Akhtar;
1 imm. (NIBGE; det. G.A. Blagoev, 2018), Punjab, Sheikhupura,
Pekhi (31.6167°N, 73.8667°E), 208 m a.s.l., 29.04.2010,S. Akhtar;
1 imm. (NIBGE; det. G.A. Blagoev, 2018), Punjab, Faisalabad,
PARS (31.033°N, 73°E), 180 m a.s.l., 29.04.2012, S. Akhtar. —
INDIA: 2 ## (ZMMU), Uttarakhand, nr. Rishikesh (29.976°N,
78.209°E), c. 300 m a.s.l., 14–16.04.2012, K.P. Tomkovich.
DISTRIBUTION. This is one of the best-known and
widespread Bianor species, recorded from South Africa, the
Mediterranean, throughout the Arabian Peninsula and Near
East to Central Asia and NW India (Punjab and Uttarak-
hand) [Logunov, 2001; present data]. Although hitherto the
species was formally reported from Pakistan [Logunov, 2001:
227], the present records provide exact locality data for the
first time.
Bianor pseudomaculatus Logunov, 2001
MATERIAL. INDIA: 1 #, 1 $ (ZMMU), Assam, Chapar, Cham-
pamati River (26.323°N, 90.461°E), c. 400 m a.s.l., 1–3.01.2014,
K.P. Tomkovich; 1 # (ZMMU), Meghalaya, Sohra area, nr. Tyrne
(25.246°N, 91.672°E), lowland jungle, 300–400 m a.s.l., 18–
21.12.2013, K.P. Tomkovich. — CAMBODIA: 2 ## (PCMF), Koh
Kong/Tatai (11°34.624′N, 103°07.684′E), c. 10 m a.s.l., 02.12–
05.12.2014, J. Nigl.
DISTRIBUTION. Bhutan, NE India (Assam, Megha-
laya), southern Vietnam (Ho Chi Minh) [Logunov, 2001;
present data]. New species record to Cambodia.
Bianor punjabicus Logunov, 2001
Figs 42–49.
MATERIAL. PAKISTAN: 2 ##, 1 $ (MMUE), Punjab Prov.,
Khushab Distr., Jahlar Lake [c. 32°30′04.1″N, 72°05′11.0″E], on

110 D.V. Logunov
Figs 42–49. General appearance of Bianor punjabicus Logunov, 2001 from Pakistan (Jahlar Lake), male (42–45) and female (46–49):
42, 47 — body, dorsal view; 43, 48 — ditto, lateral view; 44, 49 — ditto, ventral view; 45, 46 — clypeus, frontal view. Scale bars: 1 mm.
Рис. 42–49. Общий вид Bianor punjabicus Logunov, 2001 из Пакистана (оз. Яхлар), самец (42–45) и самка (46–49): 42, 47 —
тело, вид сверху; 43, 48 — то же, вид сбоку; 44, 49 — то же, вид снизу; 45, 46 — клипеус, вид спереди. Масштаб: 1 мм.
Typha, 28.10.2012, G. Rasool; 1 $ (MMUE), Pakistan, Lahore [c.
31°32′42.9″N, 74°19′54.9″E], in grass, 16.05.2013, G. Rasool.
DISTRIBUTION. NE Afghanistan, NW India (Punjab)
[Logunov, 2001] and NE Pakistan [present data]. New spe-
cies record to Pakistan, and the second one after the original
description.
COMMENTS. Although B. punjabicus can easily be
distinguished from its closest congener B. angulosus by the
conformation of the male palp (see Logunov [2001] for
further details), the males of both species can also be sepa-
rated by their body colour pattern: viz. by the presence of
two round white spots on the eye field in B. punjabicus

111
On the Harmochirina from S and SE Asia
(absent in B. angulosus) and three pairs of large white round
spots on the dorsum in B. punjabicus (two pairs of elongated
spots/bands in B. angulosus) (cf. Figs 7, 9 and 42). The
clypeus of B. punjabicus also contains two transverse white
streaks of scales (Fig. 45), not known in other Bianor spe-
cies for which the males have been described (see Logunov
[2001, 2009]).
Harmochirus brachiatus (Thorell, 1877)
MATERIAL. THAILAND: 1 # (ISEA; det. G.N. Azarkina, 2018),
Province Nakhon Ratchasima (Korat), Nong Bun Nak Vil. (14°42′N,
102°27′E), 170 m a.s.l., 20–23.07.2012, A.V. Korshunov.
DISTRIBUTION. From Bhutan in the north-west, east-
ward to southern Vietnam (Dak Lak Province) and Taiwan,
and south-eastward to Indonesia and Malaysia (Borneo) [Lo-
gunov, 2001]. The new record lies within the outlined range
of the species.
Harmochirus insulanus (Kishida, 1914)
MATERIAL. VIETNAM: 1 # (MMUE), Tuyen Quang Province,
c. 5 km E of Na Hang (22°20′59″N, 105°25′36″E), c. 290 m a.s.l.,
beaten from understorey vegetation, 4–13.11. 2015, D.V. Logunov.
DISTRIBUTION. Japan (Honshu, Shikoku, Kyushu),
China (Gansu, Hainan, Guandong) [Logunov et al., 1997;
Logunov, 2001] and northern Vietnam (present data). New
species record to Vietnam.
Harmochirus zabkai Logunov, 2001
MATERIAL. PAKISTAN: 1 # (ZMMU), N vicinity of Islama-
bad [c. 33°47′24.4″N, 73°06′21.8″E], national park, 800 m a.s.l.,
24.08.2005, S.V. Ovtchinnikov; 2 $$ (ZMMU), same locality,
5.09.2004, S.V. Ovtchinnikov; 2 $$ (MMUE), North-West Fron-
tier Prov., c. 1.5 km N of Islamabad (33°44′30″N, 73°03′E), c. 800
m a.s.l., forest (under stones), 10.07.2003, S.V. Ovtchinnikov; 1 #
(ISEA, 001.8259), Choa Saidan Shah, Chakwal (32.761°N,
73.145°E), 741 m a.s.l., rocky habitat, 15.09.2018, D. Jablonski.
— SRI LANKA: 3 $$ (OUMNH), “Ceylon, jar 1867” (no exact
locality and date).
DISTRIBUTION. Pakistan (present data), India (Pun-
jab, Maharashtra, Tamil Nadu, Karnataka), Sri Lanka (present
data), southern Vietnam (Dong Nai Province), and Malaysia
(Malaya) [Logunov, 2001]. New species record to Sri Lanka
(although without an exact locality).
Modunda staintoni (O. Pickard-Cambridge, 1872)
MATERIAL. PAKISTAN: 1 # (MMUE), Punjab Prov., Uchali
Lake [c. 32°33′49.2″N, 72°01′02.5″E], on Typha vegetation, 29.10.
2012, G. Rasool; 1 imm (CBGO; det. G.A. Blagoev, 2018), Pun-
jab, D.G. Khan, Fort Minro (29.9167°N, 69.9667°E), 1850 m a.s.l.,
18.05.2010, S. Akhtar; 1 $ (NIBGE; det. G.A. Blagoev, 2018),
Punjab, Nankana Sahib (31.45°N, 73.7°E), 193 m a.s.l., 28.05.2010,
S. Akhtar; 1 # (NIBGE; det. G.A. Blagoev, 2018), Islamabad,
Rawal Dam, Lake View Park (33.716°N, 73.129°E), 534 m a.s.l.,
22.07.2013, S. Swar.
DISTRIBUTION. From Egypt, throughout the Near East,
eastward to NW India (Punjab) [Logunov, 2001] and NE
Pakistan [present data]. New species record to Pakistan.
Acknowledgements. I want to thank Luca Bartolozzi
(MSNF), Roberta Salmaso (MCVR) and Maria Tavano
(MCSN), for giving me an opportunity to study the type
specimens deposited in their museums. I also wish to thank
Zoë Simmons (Oxford, UK), Tony Hunter and Gary Hedges
(both Liverpool, UK) for allowing me to use the digital
facilities at the Oxford University Museum of Natural Histo-
ry (UK) and the World Museums of Liverpool (UK) corre-
spondingly. I am sincerely grateful to Galina Azarkina (No-
vosibirsk, Russia) for her kind help with preparing the maps
and for providing me with additional faunistic records for B.
albobimaculatus, B. angulosus, H. brachiatus and H. zab-
kai. Daniel Jablonski (Bratislava, Slovakia) is thanked for
allowing us to study some of his spider collections from
Pakistan. Finally, I would like to thank Muhammad Ashfaq
and Gergin A. Blagoev from the Centre for Biodiversity
Genomics, University of Guelph, Ontario, Canada for pro-
viding me with additional faunistic records for B. albobi-
maculatus and M. staintoni from Pakistan.
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