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Notes on some taxa of the Alvania lineata-complex with the descriptions of three new species from the Mediterranean Sea (Gastropoda: Rissoidae) Notas sobre algunos taxones del complejo de Alvania lineata, con la descripción de tres nuevas especies del Mediterráneo (Gastropoda: Rissoidae)

February 2019

Authors:

Bruno Amati

Massimo Appolloni

Comune di Roma

Quaggiotto Ermanno

The Natural History and Archaeological Museum, Vicenza, Italia

Carlo Smriglio

University RomaTre, Rome, Italy

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Some taxonomically troublesome nominal taxa of the rissoid genus Alvania Risso, 1826 are examined: Alvania disparilis Monterosato, 1890, A. peloritana (Aradas & Benoit, 1874), A. perversa F. Nordsieck, 1972 and A. dorbignyi (Audouin, 1826). The four species are here considered as valid, and for the sake of nomenclatural stability lectotypes of Alvania disparilis, Rissoa peloritana and Alvania perversa are designated. The holotype of Alvania dorbignyi is refigured and redescribed, and all Mediterranean records of this species are reassigned to A. perversa. Alvania schwartziana Brusina, 1866 is reported for the first time in Italian waters. Three new species are herein described: Alvania bartolinorum n. sp., Alvania unica n. sp. and Alvania zaraensis n. sp.

Alvania peloritana Aradas & Benoit, 1874. 12, 13: lectotype, Messina (Sicily), H 3.35 mm (MCZR-M-22150/L); 14, 15: original labels; 16, 17: paralectotype, Messina, H 3 mm (MCZR-M-22150/P); 18: original drawings (reproduced from ARADAS & BENOIT, 1874: pl. IV, fig. 16). Figuras 12-18. Alvania peloritana Aradas y Benoit, 1874. 12, 13: lectotipo, Messina (Sicilia), H 3,35 mm (MCZR-M-22150/L); 14, 15: etiquetas originales; 16, 17: paralectotipo, Messina, H 3 mm (MCZR-M-22150/P); 18: dibujos originales (reproducidos de ARADAS & BENOIT, 1874: pl. IV, fig. 16).

…

Alvania bartolinorum n. sp. 30-32: holotype, Krk I. (Croatia), 54 m depth, H 3.58 mm (MNHN IM-2000-27706); 33: paratype, Krk I. (Croatia), 54 m depth, H 3.3 mm (SB-MS); 34: paratype, Giapni Potok (Croatia), 54 m depth H 3.85 mm (BA); 35: paratype, Krk I., (Croatia), 54 m depth, H 3.45 mm (SB-MS); 36: paratype, Prvic Krk (Croatia), 54 m depth, H 3.45 mm (BA); 37: paratype, Giapni Potok (Croatia), 54 m depth,H 3.05 mm (BA); 38: paratype, Krk I. (Croatia), 54 m depth, H 3.05 mm (SB-MS); 39, 40: paratype, Krk I., (Croatia), SEM micrograph of the protoconch, and detail (CS-PM). Figuras 30-40. Alvania bartolinorum n. sp. 30-32: holotipo, Isla Krk (Croacia), profundidad 54 m, H 3,58 mm (MNHN IM-2000-27706); 33: paratipo, Isla Krk (Croacia), profundidad 54 m, H 3,3 mm (SB-MS); 34: paratype, Giapni Potok (Croacia), profundidad 54 m, H 3,85 mm (BA); 35: paratipo, Isla Krk, profundidad 54 m, H 3,45 mm (SB-MS); 36: paratipo, Prvic Krk (Croacia), profundidad 54 m, H 3,45 mm (BA); 37: paratipo, Giapni Potok, profundidad 54 m, H 3,05 mm (BA); 38: paratipo, Isla Krk, profundidad 54 m, H 3,05 mm (SB-MS); 39, 40: paratype, Isla Krk, micrografía electrónica de barrido de la protoconch, y detalle (CS-PM).

…

Alvania unica n. sp. 41, 42: holotype, Sciacca (Sicily), H 3.45 mm (MCZR-M-22155/H); 43: SEM micrograph of the holotype; 44-47: details of the protoconch; 48: original label. Figuras 41-48. Alvania unica n. sp. 41, 42: holotipo, Sciacca (Sicilia), H 3,45 mm (MCZR-M-22155/H); 43: micrografía electrónica de barrido del holotipo; 44-47: detalles de la protoconcha; 48: etiqueta original.

…

Alvania zaraensis n. sp. 49, 50: holotype, Zara (now Zadar, Croatia), H 2.6 mm (MCZR-M-22152/H); 51: original label; 52: SEM micrograph of the holotype; 53, 54: details of the protoconch and first teleoconch whorl. Figuras 49-54. Alvania zaraensis n. sp. 49, 50: holotipo, Zara (ahora Zadar, Croacia), H 2,6 mm (MCZR-M-22152/H); 51: etiqueta original; 52: micrografía electrónica de barrido del holotipo; 53, 54: detalles de la protoconcha y primera vuelta de teleoconcha.

…

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Notes on some taxa of the Alvania lineata-complex with the

descriptions of three new species from the Mediterranean

Sea (Gastropoda: Rissoidae)

Notas sobre algunos taxones del complejo de Alvania lineata, con la

descripción de tres nuevas especies del Mediterráneo (Gastropoda:

Rissoidae)

Bruno AMATI*1, Massimo APPOLLONI**, Ermanno QUAGGIOTTO***,

Carlo SMRIGLIO**** & Marco OLIVERIO*****

Recibido el 9-X-2018. Aceptado el 28-XII-2018

ABSTRACT

Some taxonomically troublesome nominal taxa of the rissoid genus Alvania Risso, 1826

are examined: Alvania disparilis Monterosato, 1890, A. peloritana (Aradas & Benoit,

1874), A. perversa F. Nordsieck, 1972 and A. dorbignyi (Audouin, 1826). The four

species are here considered as valid, and for the sake of nomenclatural stability lectotypes

of Alvania disparilis, Rissoa peloritana and Alvania perversa are designated. The holo-

type of Alvania dorbignyi is refigured and redescribed, and all Mediterranean records of

this species are reassigned to A. perversa. Alvania schwartziana Brusina, 1866 is

reported for the first time in Italian waters. Three new species are herein described: Alva-

nia bartolinorum n. sp., Alvania unica n. sp. and Alvania zaraensis n. sp.

RESUMEN

Se examinan algunos taxones nominales problemáticos de risóidos del género Alvania

Risso, 1826: Alvania disparilis Monterosato, 1890, A. peloritana (Aradas y Benoit,

1874), A. perversa F. Nordsieck, 1972 y A. dorbignyi (Audouin, 1826). Las cuatro espe-

cies se consideran aquí válidas, y con fines de mantener la estabilidad nomenclatural se

designan lectotipos de Alvania disparilis, Rissoa peloritana y Alvania perversa. El holotipo

de Alvania dorbignyi está figurado de nuevo y redescrito, y todas las citas de esta espe-

cie en el Mediterráneo se reconducen a A. perversa. Alvania schwartziana Brusina, 1866

se cita por primera vez en aguas italianas. Se describen tres nuevas especies: Alvania

bartolinorum n. sp., Alvania unica n. sp. y Alvania zaraensis n. sp.

* Largo Giuseppe Veratti, 37/D, 00146 Rome, Italy; e-mail: bruno_amati@yahoo.it

** Museo Civico di Zoologia, Via Ulisse Aldrovandi, 18, 00197 Rome, Italy; e-mail:

massimo.appolloni@comune.roma.it.

*** Via Secula 13, 36023 Longare, Vicenza, Italy; e-mail: ermanno.quaggiotto@libero.it

**** Department of Science, “Roma Tre” University of Rome, Viale Marconi, 446, 00146 Rome, Italy; e-mail:

csmriglio@alice.it

***** Department of Biology and Biotechnologies “Charles Darwin”, Sapienza University of Rome, Viale

dell’Università 32, I-00185 Rome, Italy; e-mail: marco.oliverio@uniroma1.it

1Corresponding author

INTRODUCTION

Rissoidae Gray, 1847 is the most

diverse rissooidean family, with at least

38 recognised genera (Á

VILA

, 2005;

RISCIONE

& P

ONDER

, 2013; M

OLLUS

ASE

, 2018). The genus Alvania Risso,

1826 is currently represented by 314

recognised species of which 263 belong-

ing to the Recent fauna (M

OLLUSCA

ASE

2018). In the Mediterranean Sea the

genus Alvania is largely represented by

72 Recent species (M

OLLUSCA

ASE

2018) and includes several species-

complex, e.g. Alvania lineata Risso, 1826,

Alvania dictyophora (Philippi, 1844),

Alvania scabra (Philippi, 1844) and

Alvania subcrenulata (Bucquoy, Dautzen-

berg & Dollfus, 1884). Among them the

Alvania lineata-complex includes some

20 recognised Recent species (Table I),

all with non-planktotrophic develop-

ment, except for Alvania discors (Allan,

1818) which has a planktotrophic larval

stage.

Traditionally, the taxonomy of

Mediterranean rissoids has been based

almost exclusively on shell characters,

whose puzzling variation has often

caused uncertainty about the status of

some of these taxa. Although an inte-

grated approach (teleoconch, proto-

conch, soft parts morphology and

colour pattern, radula, genetics, etc...)

will certainly help in assessing their tax-

onomy, however, in most cases, the

availability of large samples and the

comparative study of type material

provide important data to improve

current taxonomic knowledge. In this

note, we focused on four nominal taxa

of this complex, of uncertain status:

Alvania dorbignyi (Audouin, 1826),

Alvania perversa F. Nordsieck, 1972,

Alvania peloritana (Aradas & Benoit,

1874) and Alvania disparilis Monterosato,

1890, and describe three additional new

species.

Description of the A. lineata group

All the species in the complex have

rather similar shells: thick, of medium-

large size for the genus, with ovate-

conical profile, wide base, and rather

strong teleoconch sculpture, the spiral

usually weaker than the axial; strong

apertural varix in mature specimens,

outer lip usually with denticles on the

inner side (except for Alvania datchaensis

Amati & Oliverio, 1987 and A. claudioi

Buzzurro & Landini, 2007); protoconch

paucispiral indicating a non-plank-

totrophic larval development, except for

Alvania discors (Allan, 1818), rarely

devoid of an apparent sculpture (e.g.

Alvania algeriana (Monterosato, 1877,

Alvania claudioi, Alvania bozcaadensis Tis-

selli & Giunchi, 2013 and Alvania zaraen-

sis n. sp.), usually sculptured by numer-

ous microtubercles randomly arranged

or fused to form parallel minute

threads, or more rarely by flattened

cordlets arranged into an orange skin

pattern (e.g. Alvania lanciae (Calcara,

1845)) or zigzag patterns (e.g. Alvania

datchaensis Amati & Oliverio, 1987).

Shell with usually uniformly coloured

background with or without more or

less fine or interrupted spiral bands. The

external soft parts have been described

for a few species and are conforming to

the general scheme of the genus Alvania

(e.g. B

RUSINA

, 1866: 25 for Alvania

schwartziana Brusina, 1866; P

ONDER

1985: 38 for Alvania lineata Risso, 1826;

MATI

& N

OFRONI

, 1985: 19; A

LBANO

ABELLI

, 2009: 56 for Alvania settepassii

Amati & Nofroni, 1985).

MATERIALS AND METHODS

The samples here studied are stored

in public and private collections, as

detailed under each species (see abbre-

viations), and in the Appendix. Pho-

tographs have been taken with a Sony

Cyber-Shot digital camera mounted on

a Kyowa KBS stereomicroscope, edited

with the Combine-Z software (H

ADLEY

2006). Most SEM photographs were

taken with a Philips XL30 at the Interde-

partmental Laboratory of Electron

Microscopy (LIME: University “Roma

Tre”, Rome); the SEM photos of the

holotype of Alvania garrafensis are cour-

tesy of Emilio Rolán (Vigo); that of the

type of Alvania dorbignyi is courtesy of

Iberus, 37 (1), 2019

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

Table I. List of the Mediterranean species of the Alvania lineata Risso, 1826 complex.

Tabla I. Lista de las especies mediterráneas del complejo Alvania lineata Risso, 1826.

Species Figures

Alvania aeoliae Palazzi, 1988 Figure 67

Alvania algeriana (Monterosato, 1877) Figure 72

Alvania aspera (Philippi, 1844) Figure 77

Alvania bartolinorum Amati & Smriglio n. sp. Figures 30-40

Alvania bozcaadensis Tisselli & Giunchi, 2013 Figures 63, 64

Alvania campanii Tisselli & Giunchi, 2013 Figures 65, 66

Alvania claudioi Buzzurro & Landini, 2007 Figure 58

Alvania colossophilus Oberling, 1970 Figure 76

Alvania datchaensis Amati & Oliverio, 1987 Figures 68, 69

Alvania discors (Allan, 1818) Figure 55

Alvania disparilis Monterosato, 1890 Figures 1-11

Alvania dorbignyi (Audouin, 1826) Figures 19-22

Alvania elisae Margelli, 2001 Figures 78, 79

Alvania fractospira Oberling, 1970 Figures 70, 71

Alvania garrafensis Peñas & Rolán, 2008 Figures 73-75

Alvania lanciae (Calcara, 1845) Figures 59-61

Alvania lineata Risso, 1826 Figure 62

Alvania peloritana (Aradas & Benoit, 1874) Figures 12-18

Alvania perversa Nordsieck, 1972 = Alvania dorbignyi AA non (Audouin, 1826) Figures 23-29

Alvania schwartziana Brusina, 1866 Figure 57

Alvania settepassii Amati & Nofroni, 1985 Figure 56

Alvania unica Amati & Quaggiotto n. sp. Figures 41-48

Alvania zaraensis Amati & Appolloni n. sp. Figures 49-54

the MNHN (Paris); those of the types of

Alvania perversa are courtesy of S. Hof

(SRI, Frankfurt am Main). Measure-

ments for the new species were taken

from 25 specimens of Alvania bartolino-

rum n. sp. and 12 specimens of Alvania

zaraensis n. sp. (Tables II-III).

Abbreviations and acronyms

AN: Andrea Nappo collection (Quartu

Sant’Elena, Italy);

AP: Attilio Pagli collection (Vinci, Flo-

rence, Italy);

AR: Alessandro Raveggi collection (Flo-

rence, Italy)

BA: Bruno Amati collection (Rome,

Italy)

CB: Cesare Bogi collection (Livorno,

Italy)

CS-PM: Carlo Smriglio–Paolo Mariottini

collection (Rome, Italy)

EQ: Ermanno Quaggiotto collection

(Longare, Vicenza, Italy)

IN: Italo Nofroni collection (Rome,

Italy)

MBAC: Museum of the Department of

Animal Biology of the University

(Catania, Italy)

CMSN: Museum of Natural History

(Milan, Italy)

MGPUF: Geological Museum and Palae-

ontology of the University (Florence,

Italy)

MCZR: Zoological Museum (Rome,

Italy)

MNHN: National Museum of Natural

History (Paris, France)

MO: Marco Oliverio collection (Rome,

Italy)

RP: Raffaele Petrone (Rome, Italy)

SG: Serge Gofas collection (University of

Malaga)

Iberus, 37 (1), 2019

SYSTEMATICS

Class G

ASTROPODA

Cuvier, 1795

Subclass C

AENOGASTROPODA

Cox, 1960

Superfamily R

ISSOOIDEA

Gray, 1847

Family R

ISSOIDAE

Gray, 1847

Genus Alvania Risso, 1826: 140

Type-species: Alvania europea Risso, 1826: 142, pl. IX, fig. 116 = Alvania cimex (Linnaeus, 1758)

(Turbo), by subsequent designation N

EVILL

, 1885: 105

Alvania disparilis Monterosato, 1890 (Figs. 1-11)

Alvania disparilis Monterosato, 1890: 146

Type material: Lectotype (MCZR–M–22155/L, Monterosato coll.) (Figs 1-4), here designated (H 2.95

mm, W 1.75 mm) and 25 paralectotypes (MCZR–M–22155/P, Monterosato coll.) (mostly fragments),

from the type locality, 1 paralectotype (MCZR–M–22155/P, Monterosato coll.) from Sciacca, Sicily.

Other material examined: Sicily. Messina, labelled, “R. Peloritana, subfoss. del Porto di Messina”,

6 sh (of which some broken) (MCZR–M–22150, Monterosato coll.); Messina, labelled “?Duccinella!

Messina!/Alvania peloritana”, 4 sh(MCZR–M–30053, Monterosato coll.); unspecified locality, 5 sh

(MCZR–M–30054, Monterosato coll.).

Type locality: Off Palermo (Sicily, Italy), ‘Funnazzi’, unspecified depth.

SB-MS: Stefano Bartolini-Maria Scape-

rrotta collection, (Florence, Italy)

SEM: scanning electron microscope

SMF: Senckenberg Museum (Frankfurt

am Main, Germany)

Teleoconch=

H: height

W: width

Ha: height of the aperture

H/W: ratio height/width

Nw: number of whorls

Nar: number of axial ribs

Nsc-ab: number of spiral cords above

the aperture

Nsc-ba: number of spiral cords on the base

Protoconch =

h: height

d: diameter of nucleus

Do: diameter of first half whorl

DM: maximum diameter

nw: number of whorls

sh: empty shell(s)

lv: live collected specimen(s) with soft

parts.

Distribution: Central Mediterranean:

off Palermo and Messina (Sicily), at

unspecified depth, possibly ca. 90-180 m

depth (ca. 50-100 ‘braccia’) for off

Palermo. The site name ‘Funnazzi’ indi-

cates an enigmatic fishing area off

Palermo, repeatedly cited by Mon-

terosato and de Gregorio (D

REGORIO

1889a: 248; 1889b: 7, 8); traces of its loca-

tion have been lost even among local

fishermen (see A

PPOLLONI ET AL

., 2018:

table 1).

Description of the lectotype: Shell (Figs

1-4) small, not solid, ovate-conic,

globose, height 2.95 mm, width 1.75

mm, H/W 1.68, aperture 1.45 mm. Pro-

toconch (Figs 3, 4) height 0.375 mm,

globose, paucispiral, of 1.6 whorls,

diameter of nucleus 0.15 mm, diameter

of first half whorl 0.25 mm, maximum

diameter 0.425 mm; sculpture of a

dozen interrupted spiral cordlets,

becoming finer and more numerous

toward the protoconch/teleoconch

boundary. Teleoconch of 3.4 convex

whorls, suture canaliculated. Axial

sculpture on the last whorl of 14 fine

orthocline ribs, half as wide as the inter-

spaces, interrupted at the suprasutural

spiral cordlet. Spiral sculpture starting

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

Figures 1-11. Alvania disparilis Monterosato, 1890. 1: lectotype, Palermo (Sicily), H 2.95 mm

(MCZR–M–22155/L); 2, 3: SEM micrograph of the lectotype; 4: detail of the protoconch sculp-

ture of the lectotype; 5: original label; 6: shell from Messina (Sicily) (MCZR–M–22150), H 2.8

mm; 7: original label of the same shell; 8: SEM micrograph of the same shell; 9-11: details of the

protoconch.

Figuras 1-11. Alvania disparilis Monterosato, 1890. 1: lectotipo, Palermo (Sicilia), H 2,95 mm

(MCZR–M–22155/L); 2, 3: micrografía electrónica de barrido del lectotipo; 4: detalle de la escultura

de la protoconcha del lectotipo; 5: etiqueta original; 6: concha de Messina (Sicilia)

(MCZR–M–22150), H 2,8 mm; 7: etiqueta original de la misma concha; 8: micrografía electrónica de

barrido de la misma concha; 9-11: detalles de la protoconcha.

200 µm

20 µm

200 µm

50 µm

100 µm

500 µm

810 11

with 2 (II and IV) visible fine spiral

cordlets on early teleoconch; on the last

whorl 9 (4+1+4) cordlets, the four adapi-

cal ones equidistant and twice as wide

as the interspaces, the others slightly

closer at first and then spaced with

increasing distance abapically; minute

tubercles at the intersection of axial ribs

and spirals cordlets, forming a quadran-

gular reticulate pattern. Coloration light

brown, with protoconch and spiral

cordlets slightly darker. Soft parts and

operculum unknown.

Remarks: The original material of Al-

vania disparilis Monterosato, 1890

(MCZR–M–22155, Monterosato coll.)

PPOLLONI ET AL

., 2018: 43, pl. 14, figs C,

D), consists of two vials: one contains 26

syntypes, mostly fragments, from ‘Fun-

nazzi’, off Palermo, Sicily, with auto-

graph label by Monterosato reading: “Al.

disparilis, Monts. mss. 1890 Palermo

1886!” (Fig. 5). The best-preserved shell

is here selected as lectotype (Figs 1-4), to

stabilize the name usage. This same shell

was figured by C

AMPANI

, B

ARTOLINI

PANU

(2012: 89, fig. 7) with erroneous

measurement (H 3.6 mm instead of H

2.95 mm) to compare with A. garrafensis

sensu C

AMPANI

, B

ARTOLINI

& S

PANU

2012 (= A. bartolinorum n. sp. this work,

not A. garrafensis Peñas & Rolán, 2008).

The other vial contains a single shell,

from off Sciacca, Sicily, also with auto-

graph label by Monterosato reading:

“Sciacca” (Figs 41-48). This shell is not

conspecific with the specimens from off

Palermo, and represents indisputably an

undescribed species, which will be de-

scribed below (Alvania unica n. sp.). Alva-

nia disparilis Monterosato, 1890 has been

considered as a deep-water form of Alva-

nia lanciae (Calcara, 1845) (K. Nicolay

and G. Schirò in

VAN

ARTSEN

, 1982: 4).

Alvania lanciae (Calcara, 1845) (:29, pl.

IV, fig. 12) (Figs 59-61) differs from

Alvania disparilis by its conspicuously

microsculptured teleoconch (lacking in A.

disparilis) and by the protoconch sculp-

ture of irregular pits, with an orange-skin

like pattern, vs. a dozen interrupted

spiral cordlets in A. disparilis. Further-

more, A. lanciae has usually a more robust

shell, with large nodules at the intersec-

tions, slightly elongated along the spirals,

gradually stronger toward the base, vs.

weaker minute tubercles in A. disparilis,

and the spiral sculpture above the aper-

ture is irregularly spaced vs. equidistant

and finer in A. disparilis.

Alvania garrafensis Peñas & Rolán,

2008 (:21, figs 8-13) (Figs 73-75) has a

weaker spiral sculpture, particularly on

the first whorls, slightly stronger only on

the base but not overrunning the axial

one (in Alvania disparilis the spiral sculp-

ture is on the whole teleoconch and the

intersection with the axials forms a

quadrangular reticulation and minute

tubercles); the axials are hardly visible

on the first whorl, are rounded and

orthocline with an evident shoulder (in

Alvania disparilis the axials are fine and

orthocline, lacking any shoulder, and

visible from the first whorl). The outline

and the protoconch sculpture are similar.

Alvania lineata Risso, 1826 (:142, pl.

IX, fig. 120) (Fig. 62) differs from Alvania

disparilis by its more robust shell reach-

ing a larger size, with a more slender

outline, with more marked and nodu-

lose sculpture (axial and spiral), with

axials reaching the base, a robust varix

and a more slender protoconch.

See also remarks under Alvania

zaraensis n. sp. for the differences with

this species.

Iberus, 37 (1), 2019

100

Alvania zaraensis Amati & Appolloni n. sp. (Figs. 49-54, Table III)

Type material: holotype (MCZR–M–22152/H), (H 2.6 mm, W 1.6 mm) and 14 paratypes

(MCZR–M–22152/P), (type loc.).

Type locality: Zara [now Zadar], Croatia.

Etymology: From the type locality, Zara.

Figures 49-54. Alvania zaraensis n. sp. 49, 50: holotype, Zara (now Zadar, Croatia), H 2.6 mm

(MCZR–M–22152/H); 51: original label; 52: SEM micrograph of the holotype; 53, 54: details of

the protoconch and first teleoconch whorl.

Figuras 49-54. Alvania zaraensis n. sp. 49, 50: holotipo, Zara (ahora Zadar, Croacia), H 2,6 mm

(MCZR–M–22152/H); 51: etiqueta original; 52: micrografía electrónica de barrido del holotipo; 53,

54: detalles de la protoconcha y primera vuelta de teleoconcha.

49 50

53 54 200 µm

200 µm

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

101

Distribution: Mediterranean: Adriatic

Sea, Zara [now Zadar] (Croatia),

unspecified depth; habitat unknown, it

is presumed alive in algal facies.

Description: (between parentheses

data of the holotype). Shell (Figs 49-54)

small, solid, ovate-conic, height

2.10–2.95 mm (2.6 mm), width 1.45-1.9

mm (1.6 mm), H/W 1.45-1.66 (1.62).

Protoconch (Figs 53, 54) paucispiral,

slightly globose, shiny, without appar-

ent sculpture under SEM due to the

worn condition of the specimen, of 1.25-

1.4 (1.25) whorls, height 0.25-0.35 mm

(0.30 mm), diameter of nucleus 0.125-

0.175 mm (0.137 mm), diameter of first

half whorl 0.20-0.275 mm (0.237),

maximum diameter 0.325-0.375 mm

(0.356 mm). Teleoconch of 3.7-4.8 (4)

barely convex whorls, suture deep and

canaliculated. Axial sculpture of 9-12+v

(9+v) strong orthocline ribs, as wide as

the interspaces, interrupted at the

periphery, in correspondence with the

sutural cordlet. Spiral sculpture starting

with 2 (II, IV) weak cordlets. Last whorl

with 8-10 (9) spirals; the adapical 4

above the aperture, equidistant and as

wide as the interspaces, the basal 4-6 (5)

(including the sutural one) finer and

more spaced; fourth adapical cordlet

more pronounced. Rounded tubercles at

the intersections. Microsculpture absent.

Umbilical chink barely visible. Outer lip

thickened by a broad and robust varix,

with 7-8 (7) internal elongate denticles.

Colour monochrome light beige-orange,

rarely light brown with whitish supra-

sutural band. Varix whitish with weak

orange lines in correspondence with the

Table III. Measurements of the protoconch and teleoconch of Alvania zaraensis n. sp., in mm.

Number 1 is the holotype. Abbreviations, h: height; d: diameter of nucleus; Do: diameter of first

half whorl; DM: maximum diameter; nw: number worls; H: height, W: width; RH/W: ratio

height/width; Nw: number of teleoconch whorls; Nar+v: number axial ribs+ varix; Ns: number of

spirals on the last whorl (above the aperture); Nd: number of denticles; Sts: starting number of

spiral cordlets.

Tabla III. Medidas de la protoconcha y teleoconcha de Alvania zaraensis n. sp., en mm. El número 1 corre-

sponde al holotipo. Abreviaturas, h: altura; d: diámetro del núcleo; Do: diámetro de la primera media

vuelta; DM: máximo diámetro; nw: número de vueltas; H: altura, W: anchura; RH/W: ratio

altura/anchura; Nw: número de vueltas de la teleoconcha; Nar+v: número de costillas axiales + variz; Ns:

número de espiras de la última vuelta (sobre la apertura); Nd: número de dentículos; Sts: número inicial de

cordones espirales.

Protoconch 1 2 3 4 5 6 7 8 9 10 11 12

h 0.30 0.35 0.25 —- 0.30 —- 0.30 —- 0.30 0.31 0.30 0.31

d 0.137 0.14 0.14 —- 0.175 0.14 0.125 —- 0.15 0.145 0.14 0.125

Do 0.237 0.225 0.225 —- 0.225 0.225 0.20 —- 0.275 0.245 0.26 0.225

DM 0.356 0.35 0.34 —- 0.365 —- 0.325 —- 0.36 0.365 0.375 0.35

nw 1.25 1.35 1.3 —- 1.35 —- 1.3 —- 1.35 1.35 1.4 1.3

Teleoconch 1 2 3 4 5 6 7 8 9 10 11 12

H 2.6 2.6 2.9 2.95 2.75 2.7 2.75 2.5 2.9 2.75 2.75 2.1

W 1.6 1.65 1.8 1.9 1.77 1.8 1.7 1.57 1.75 1.77 1.75 1.45

RH/W 1.62 1.57 1.61 1.55 1.55 1.50 1.62 1.59 1.66 1.55 1.57 1.45

Nw 444.8 4.8 4.6 4.6 4.6 4 4.8 4.6 4.6 3.7

Nar+v 9+v 11+v 10+v 10+v 9+v 10+v 11+v 12+v 10+v 11+v 9+v 10+v

Ns 9(4) 8(4) 9(4) 9(4) 9(4) 9(4) 10(4) 9(4) 10(4) 10(4) 8(4) 9(4)

Nd 7 no 888no 788887

Sts 222222222222

Iberus, 37 (1), 2019

102

(Right page) Figures 55-70. Alvania spp. 55: Alvania discors (Allan, 1818), Sant’Agostino, Civi-

tavecchia (Italy), H 5 mm (BA); 56: Alvania settepassii Amati & Nofroni, 1985, paratype, Torreval-

daliga, Civitavecchia (Italy), 25-30 m depth, H 4.3 mm (BA); 57: Alvania schwartziana Brusina,

1866, Pago (now Pag, Croatia), H 3.6 mm (BA); 58: Alvania claudioi Buzzurro & Landini, 2007,

Zannone I. (Italy), H 2.5 mm, (BA); 59: Alvania lanciae (Calcara, 1845), Cala Gonone (Sardinia),

0.5 m depth, H 3.25 mm, (BA); 60: Alvania lanciae (Calcara, 1845), Bastia, Corsica (France), H

3.45 mm (MCZR–M–22161); 61: Alvania lanciae (Calcara, 1845), Elba I. (Italy), H 3.58 mm

(AN); 62: Alvania lineata Risso, 1826, Salina I. (Sicily), H 3.65 mm (BA); 63, 64: Alvania boz-

caadensis Tisselli & Giunchi, 2013, Aydincik (Turkey), H 2.15 mm (BA); 65, 66: Alvania cam-

panii Tisselli & Giunchi, 2013, Bozcaada (Turkey), 8 m depth H 2.55 mm (BA); 67: Alvania

aeoliae Palazzi, 1988, Salina I. (Sicily), H 2.3 mm (BA); 68, 69: Alvania datchaensis Amati & Oliv-

erio, 1987, Aydincik (Turkey), H 2.4 mm (BA); 70, 71: Alvania fractospira Oberling, 1970, Trogyr

(Croatia), H 2.55 mm (BA); 72: Alvania algeriana (Monterosato, 1877) Algiers (Algeria), H 2.95

mm (MCZR–M–22182).

(Página derecha) Figuras 55-70. Alvania spp. 55: Alvania discors (Allan, 1818), Sant’Agostino, Civi-

tavecchia (Italia), H 5 mm (BA); 56: Alvania settepassii Amati y Nofroni, 1985, paratipo, Torrevalda-

liga, Civitavecchia (Italia), profundidad 25-30 m, H 4,3 mm (BA); 57: Alvania schwartziana

Brusina, 1866, Pago, (now Pag, Croacia), H 3,6 mm (BA); 58: Alvania claudioi Buzzurro &

Landini, 2007, Zannone I. (Italia), H 2,5 mm, (BA); 59: Alvania lanciae (Calcara, 1845), Cala

Gonone (Cerdeña), profundidad 0,5 m , H 3,25 mm, (BA); 60: Alvania lanciae, Bastia, Córcega

(Francia), H 3,45 mm (MCZR–M–22161); 61: Alvania lanciae, Elba I. (Italia), H 3,58 mm (AN);

62: Alvania lineata Risso, 1826, Salina I. (Sicilia), H 3,65 mm (BA); 63, 64: Alvania bozcaadensis

Tisselli & Giunchi, 2013, Aydincik (Turquía), H 2,15 mm (BA); 65, 66: Alvania campanii Tisselli &

Giunchi, 2013, Bozcaada (Turquía), profundidad 8 m, H 2,55 mm (BA); 67: Alvania aeoliae

Palazzi, 1988, Salina I. (Sicilia), H 2,3 mm (BA); 68, 69: Alvania datchaensis Amati & Oliverio,

1987, Aydincik (Turquía), H 2,4 mm (BA); 70, 71: Alvania fractospira Oberling, 1970, Trogyr

(Croacia), H 2,55 mm (BA); 72: Alvania algeriana (Monterosato, 1877) Argel (Argelia), H 2,95 mm

(MCZR–M–22182).

spirals. Operculum and soft parts

unknown.

Remarks: A vial in the Monterosato

collection (MCZR–M–22152) contains 32

shells from Zara (now Zadar, Croatia),

labelled (Fig. 51) “Alvania montagui

xxxxx [unreadable] di Zara”, of which 7

Alvania lineata Risso, 1826, 10 Alvania

lanciae (Calcara, 1845) and 15 Alvania

zaraensis n. sp. Actually, the new species

resembles a dwarf form of Alvania

discors (Allan, 1818) (:463, pl X, fig. 5)

(Fig. 55), but the latter has a multispiral

protoconch of 1.7-1.8 whorls vs. 1.25-1.4

of A. zaraensis n. sp., with the diameter

of nucleus 0.85-0.112 mm vs. 0.125–0.175

in A. zaraensis n. sp.; the shell is then

larger (> 5 mm vs. < 3 mm in A. zaraensis

n. sp.) and more brightly coloured and

more polychrome.

Alvania bozcaadensis Tisselli &

Giunchi, 2013 (:164, figs 1-8, 9-11, 28-29)

(Figs 61, 62) is probably the morpholog-

ically closest species, differing by the

more slender outline, the more globose

apex, the more numerous and finer

axial ribs on last whorl (13-16 vs. 9-12 in

A. zaraensis n. sp.), as wide as half the

interspaces vs. larger than interspaces

in A. zaraensis n. sp., the more globose

apex, often with lower number of

whorls (1.25 vs. 1.25-1.40 in A. zaraensis

n. sp.), and a smaller nucleus (0.100 mm

vs. 0.125-0.175 mm in A. zaraensis n.

sp.).

Alvania lanciae (Calcara, 1845) (:29,

pl. IV, fig. 12) (Figs 59-61) is similar to

the new species in particular with some

of its Central Mediterranean morphs

MATI

, 2012: 118-119 pls 1-2, figs A-G;

A-F) with robust shell and almost flat

whorls; it differs by the evident teleo-

conch and protoconch microsculptures,

absent in A. zaraensis n. sp., and for the

more globose protoconch. (A

MATI

LIVERIO

, 1987: 51, 52, pl. II, fig. 5 and

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

103

63 64 65 66

68 69 70 71 72

Iberus, 37 (1), 2019

104

Figures 73-82. Alvania spp. 73: Alvania garrafensis Peñas & Rolán, 2008: SEM micrograph of the

holotype, Mar de Cubelles, Garraf (Spain), H 2.8 mm (MNCN); 74: Detail of the protoconch;

75: Detail of the teleoconch; 76: Alvania colossophilus Oberling, 1970, Capo Greco Protaras,

Cyprus, H 3.75 mm (BA); 77: Alvania aspera (Philippi, 1844), Portoroz (Slovenia), H 3 mm (BA);

78: Alvania elisae Margelli, 2001, paratype, Capraia I. (Italy), H 3.75 mm (CS-PM); 79: Alvania

elisae Margelli, 2001, Capri I. (Italy), H 3.6 mm (MCZR-M-30060), intermediate specimen,

tending to species A. settepassii; 80: Alvania dorbignyi (Audouin, 1827), reproduced from S

AVIGNY

1817: pl. 3, 22; 81: Alvania dorbignyi (Audouin, 1827), reproduced from N

ORDSIECK

, 1972: 230,

13; 82: Alvania perversa F. Nordsieck, 1972, reproduced from N

ORDSIECK

, 1972: 230, 14.

Figuras 73-82. Alvania spp. 73: Alvania garrafensis Peñas & Rolán, 2008: micrografía electrónica de

barrido del holotipo, Mar de Cubelles, Garraf (España), H 2,8 mm (MNCN); 74: Detalle de la proto-

concha; 75: Detalle de la teleoconcha; 76: Alvania colossophilus Oberling, 1970, Capo Greco Protaras,

Chipre, H 3,75 mm (BA); 77: Alvania aspera (Philippi, 1844), Portoroz (Eslovenia), H 3 mm (BA);

78: Alvania elisae Margelli, 2001, paratipo, isla de Capraia (Italia), H 3,75 mm (CS-PM); 79:

Alvania elisae Margelli, 2001, isla de Capri (Italia), H 3,6 mm (MCZR-M-30060), ejemplar inter-

medio, que tiende hacia la especie A. settepassii; 80: Alvania dorbignyi (Audouin, 1827), reproducido

de S

AVIGNY

, 1817: pl. 3, 22; 81: Alvania dorbignyi (Audouin, 1827), reproducido de N

ORDSIECK

1972: 230, 13; 82: Alvania perversa F. Nordsieck, 1972, reproducido de N

ORDSIECK

, 1972: 230, 14.

74 75

78 79

81 82

500 µm

100 µm

1 mm

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

105

Figures 83-88. Alvania montagui (Payraudeau, 1826). 83, 84: syntype in apertural and dorsal view,

Corsica, H 4.2 mm (MNHN-IM-2000-27921): 85: syntype in apertural view, Corsica, H 3.95

mm (MNHN-IM-2000-27921); 86, 87: SEM micrograph of the same syntype as 83, 84, in aper-

tural view (86) and detail of the first whorls (87); 88: original figure (reproduced from

AYRAUDEAU

, 1826: pl. V, fig 13; drawing tipped).

Figuras 83-88. Alvania montagui (Payraudeau, 1826). 83, 84: sintipo en vista apertural y dorsal,

Córcega, H 4,2 mm (MNHN-IM-2000-27921); 85: sintipo en vista apertural, Córcega, H 3,95 mm

(MNHN-IM-2000-27921); 86, 87: micrografía electrónica de barrido del mismo sintipo que el 83,

84, en vista apertural (86) y detalle de las primeras vueltas (87); 88: figura original (reproducida de

AYRAUDEAU

, 1826: pl. V, fig. 13; dibujo volcado).

pl. III, figs 6,7; A

MATI

, 2012: 120, fig. 3,

A, B; T

ISSELLI

& G

IUNCHI

, 2013: 169, pl.

2, figs 12-14).

Alvania campanii Tisselli & Giunchi,

2013 (:165, figs 15-21, 26-27, 31-32, 36)

(Figs 63, 64) has finer axials as wide as

half the interspaces vs. robust barely

larger than interspaces ribs in A. zaraen-

sis n. sp.; its varix is less robust; the pro-

toconch is more globose, and sculptured

by granules set in series of 12-14 slightly

wavy spiral threads vs. smooth in A.

zaraensis n. sp.

Alvania elisae Margelli, 2001 (:43, figs

1-11, 16-19, pls 1-3) (Figs 78, 79) differs

in the larger shell (H 3.5-4 mm vs. H

2.1-2.95 mm in A. zaraensis n. sp.), the

protoconch has an evident microsculp-

tures vs. absent in A. zaraensis n. sp.; a

greater number of axial ribs on the last

whorl 15-16 vs. 9-12 in A. zaraensis n.

sp.

83 84 85

87 86 88

200 µm

Alvania disparilis Monterosato, 1890

(:146) (Figs 1-11) has a less robust shell

with more convex whorls, finer and

more numerous axial ribs (14-15 vs. 9-12

in A. zaraensis n. sp.).

Alvania peloritana (Aradas & Benoit,

1874) (:205, pl. IV, fig. 16) (Figs 12-18)

has more convex whorls vs. almost flat

in A. zaraensis n. sp.; less incise suture

vs. deeply canaliculate in A. zaraensis n.

sp.; axial ribs extending beyond the

periphery vs. interrupted at the

periphery in A. zaraensis n. sp.; larger

size (3.0-3.35 vs. 2.1-2.95 in A. zaraensis

n. sp.).

Alvania dorbignyi (Audouin, 1827)

(:171) (Figs 19-22) has a higher H/W

ratio (1.81 vs. 1.45-1.66 in A. zaraensis n.

sp.); opisthocline flexuose axials vs.

orthocline in A. zaraensis n. sp.

Alvania perversa F. Nordsieck, 1972

(:230, fig. 14) (Figs 23-29) differs in its

weaker sculpture (both axial and spiral),

with axials less elevate.

Alvania bartolinorum n. sp. (Figs 30-

40) differs in its shell less robust and

thick, with weaker sculpture (both axial

and spiral), especially the spiral one,

and in the maximum diameter of the

protoconch (0.44-0.50 mm vs. 0.325-0.375

mm A. zaraensis n. sp.).

Alvania unica n. sp. (Figs 41-48) dif-

fers in the larger size (height 3.45 mm vs.

2.10–2.95 mm in A. zaraensis n. sp.), the

finer and more numerous axials (15 vs. 9-

12 in A. zaraensis n. sp.), the suture less

canaliculate, the more numerous inner

denticles on the outer lip (10 vs. 7-8 in A.

zaraensis n. sp.), the higher protoconch,

with more whorls and wider diameter

(nw 1.5, h 0.375 mm, DM 0.475 mm vs.

nw 1.25-1.4, h 0.25-0.35 mm, DM 0.325-

0.375 mm in A. zaraensis n. sp.), the pro-

toconch sculpture of c. 10 spiral cordlets,

apparently lacking in A. zaraensis n. sp.

Specimens from some localities of

the eastern Mediterranean – Greece:

Crete, beached bioclastic sand, 2 sh (RP);

Saronikos, 1 sh (EQ); Turkey: Taşucu, 5

sh (EQ); Iskenderum Gulf, 2 sh (EQ);

Bozcaada I. 3 sh (EQ) – are very similar

to typical A. zaraensis n. sp. The shells

are often larger at the same number of

whorls, the height/width ratio is higher,

the tubercles at the intersections are

more rounded; the colour is yellowish

with two white suprasutural cords, and

the basal cords are dashed with darker

colour and the columella is darker. We

consider this material worthy of a

deeper insight for these potentially

diagnostic characters.

Iberus, 37 (1), 2019

106

ACKNOWLEDGEMENTS

Virginie Héros and Philippe Maes-

trati (MNHN, Paris) and Monica

Leonardi (CMSN, Milan) are thanked

for their very kind collaboration with

type material searches; Manuel Caballer

(project e-Recolnat; MNHN, Paris) for

the pictures of the holotype of Alvania

dorbignyi and of the syntypes of Alvania

montagui; Ronald Janssen and S. Hof

(SRI, Frankfurt am Main) for the pic-

tures of the type material of Alvania per-

versa; Paul Mazza (MGPU, Florence) for

his assistance during our visits to

examine Seguenza’s material; Emilio

Rolán (Vigo) for the pictures (SEM) of

the holotype of Alvania garrafensis;

Andrea Nappo (Quartu Sant’Elena) for

the photo of Fig. 61. A sincere thank to

Andrea Di Giulio (Dipartimento di

Scienze, “Roma Tre” University, Rome)

for the SEM photos, carried out at LIME

(Interdepartmental Laboratory of Elec-

tron Microscopy, “Roma Tre” University,

Rome); Danilo Scuderi (Catania), Italo

Nofroni (Rome), Cesare Bogi (Livorno),

Stefano Bartolini and Maria Scaperrotta

(Florence), Alessandro Raveggi (Flo-

rence) Raffaele Petrone (Rome) and

Attilio Pagli (Vinci, Florence) for their

friendly help with discussions and

materials. The authors are grateful to

Serge Gofas (Departamento de Biología

Animal, Universidad de Málaga, Spain)

and to an anonymous reviewer, for the

useful comments that helped to improve

this article.

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

107

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MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

109

Iberus, 37 (1), 2019

110

APPENDIX. LIST OF MATERIAL EXAMINED

Alvania bartolinorum n. sp. (See under species heading);

Alvania unica n. sp. (See under species heading);

Alvania zaraensis n. sp. (See under species heading);

Alvania disparilis Monterosato, 1890 (See under species heading);

Alvania peloritana (Aradas & Benoit, 1874) (See under species heading);

Alvania aeoliae Palazzi, 1988: Italy: Cannizzaro, Catania, Sicily 43 m depth, 16 sh

(BA), 30 m depth, 5 sh (BA); Salina I., Sicily, 35 m depth, vii.2002, >500 sh (BA);

Alvania algeriana (Monterosato, 1877): Algeria: Algiers, 6 syntypes

(MCZR–M–22182 Monterosato coll. ex Joly); Algiers, 5 sh (MCZR–M–22182 Mon-

terosato coll. ex Joly);

Alvania aspera (Philippi, 1844): Slovenia: Portoroz, beached, 1980, 8 sh (BA);

Greece: Zakynthos I. under stones 2-3 m depth viii.1990, 1 sh (BA);

Alvania schwartziana Brusina, 1866: Croatia: Zara (now Zadar), 4 syntypes

(MCZR–M–22326, Monterosato coll. ex Brusina); Pago (now Pag, near Zadar), 8 sh

(BA); Starigrad Paklenica, Zadar, 6 m depth, 29 sh (EQ); Italy: “Tegnùe” [reefs built by

coralline algae, bryozoans, sponges and cnidarians, with hardened sediments, typical

of shallow waters in the Gulf of Venice between 8 and 40 meters depth; the local fish-

ermen word, meaning ‘withhold’, refers to the fishing nets frequently remaining

entangled: see e.g. N

EWTON

& S

TEFANON

, 1975], Chioggia, Venice, 20 m depth, 2 sh

(EQ) (first record for Italy);

Alvania garrafensis Peñas & Rolán, 2008: only photographs of the holotype (MNCN

15.05/47516);

Alvania settepassii Amati & Nofroni, 1985: Italy: holotype (MCZR–M–TYPE00004/H)

and 42 paratypes Secche di Tor Paterno, 45 m depth (BA); Civitavecchia, 25-30 m depth,

69 paratypes (BA); Secche di Torre Flavia, Ladispoli 26 m depth 1986, 3 sh (BA); Giglio I.,

27-30 m depth, 16 sh (BA); S. Stefano I., 40 m depth, 1 sh (BA); Torre Astura, beached, 3

sh (BA); Nettuno, beached, 1982, 1 sh (BA); Torvaldaliga, Civitavecchia, 20-22 m depth,

250 sh (CS-PM); Sicily: Vendicari I., 28 m depth, 1 sh (BA); Croatia: Lastovo I. 38 m

depth, 21 sh (BA); Greece: Crete, 960 m depth, 3 sh (AP);

Alvania bozcaadensis Tisselli & Giunchi, 2013: Turkey: Bozcaada I., 12 m depth, 2

paratypes (EQ); Bozcaada I., 8-12 m depth, 23 sh (EQ); Aydincik, 4 sh (BA); Greece:

Leros I., amidst fishing nets residuals, 1985, 5 sh (CS-PM); Analipsi, Astypalea 5-6 m

depth, beached bioclastic sand, 2013, 3 sh (MO);

Alvania perversa F. Nordsieck, 1972: (See under species heading);

Alvania dorbignyi (Audouin, 1826): (See under species heading);

Alvania claudioi Buzzurro & Landini, 2007: Italy: Zannone I. 18 m depth, x.2004, 1

sh (BA); Zannone I., Approdo Romano, 35 m depth, xiv.2008, 11 sh (7 juv.) (EQ);

Alvania campanii Tisselli & Giunchi, 2013: Turkey: Bozcaada I., 12 m depth, 2

paratypes (EQ), Bozcaada I., 8 m depth, 2 sh (BA), 2 sh (CS-PM); Bozcaada I., 8-12 m

depth, 12 sh (EQ);

Alvania colossophilus Oberling, 1970: 1 syntype (NMBE); Croatia: Lastovo I. 38 m

depth, 31 sh (BA); Turkey: Kash, 26.v.1992, 50 m depth, 20 sh (BA); Aydincik, 10 m

depth ix.1990 41 sh (BA); Cyprus: Cape Greco, Protaras, xi.2011, 7 sh (BA); Macronis-

sos, beached xii.1988, 2 sh (BA);

Alvania datchaensis Amati & Oliverio, 1987: holotype and paratypes: MCZR, BA,

MO; Cyprus, Cape Greco, Protaras, ix.2011, 11 sh (BA); Turkey: Aydincik 9-10 m depth

1990 86 sh (BA);

Alvania discors (Allan, 1818): Italy: Trieste, 1980, 1 sh (BA); Sant’Agostino, Civi-

tavecchia, Rome, 0.50 m depth on Posidonia oceanica 1980, 26 sh (BA); Torre Astura,

Rome 1976-77, 64 sh (BA); Porto Cesareo, 55 sh (BA); Sicily: Palermo, 166 sh

(MCZR–M–22145); Ognina, 19 sh (MCZR–M– 22306); Porto Palo di Capopassero, 6 sh

(BA); Lampedusa I., Cala Calandra, 30 m depth, iv.1981, 29 sh (BA); Vendicari I., 28 m

MATI ET AL

.: Notes on the Alvania lineata-complex and three new Mediterranean species

111

depth, 7 sh (BA); Capo Asparano, 13 sh (BA); Canizzaro, 30 m depth, 1 sh (BA);

Magnisi (SR), beached bioclastic sand, legit J.J. Oberling (13.iii.1970), 30 sh (BA); Pan-

telleria I., Scauri, 13 m depth, 45 sh (BA); Sardinia: unspecified locality, viii.1981, 14 sh

(BA); Santa Caterina, 1978, 8 sh (BA); Calasetta S. Antioco, 10 sh (BA); Maddalena I., 1

sh (BA); Croatia: Trogyr viii.1989 5-6 m depth, 58 sh (BA); San Lorenzo, 6 m depth, 25

sh (BA); Umag, 1980, 200 sh (BA); France: Verghia, Ajaccio, Corsica, 3 m depth, 4 sh

(BA); Greece: unspecified locality, 15 sh (BA); Zakinthos I., 2-3 m depth, 24 sh (BA);

Turkey: Aydincik, 15 sh. (BA); Malta: undefined locality, 81 sh (BA); undefined local-

ity, 8 sh (BA); Algeria: Oran, viii.1977, 4 sh (BA); Tunisia: Gulf of Gabès, Kerkennah,

beach, 1974, 25 sh (BA); Gulf of Gabès, Kerkennah, loc. Sidi Youssef, 49 sh (BA); Gulf

of Gabès, Kerkennah, beach, 32 sh (BA); Gulf of Gabès, Kerkennah, loc. Sidi Youssef,

vii.2015, 350 sh (BA);

Alvania montagui (Payraudeau, 1826): photographs of the 2 syntypes, Corsica

(MNHN-IM-2000-27921);

Alvania lanciae (Calcara, 1845): lectotype (Amati, 2012) and numerous (>200) para-

lectotypes, Pantelleria I. (MZP–1935); lectotype (Amati, 2012) and 7 paralectotypes of

Alvania consociella Monterosato, 1884, Corsica (MCZR–M–22159), 7 paralectotypes,

Villefranche (Villefranche-sur-Mer) (MCZR–M–30126 cabinet of type material);

France: St. Raphael 49 sh (MCZR–M–22183), original labels reading “Alvania arguta

(ex Locard coll.)” and “A. locardi Monts. mss (ex Caziot coll.)”; Verghia, Ajaccio,

Corsica, 3 m depth, viii.1988, 4 sh (BA); Suasset les Pins, v.1982, 2 sh (IN); Spain:

Palamos, 1978-79, 2 sh (BA); undefined locality, 2 sh (IN); Sardinia: Cala Gonone,

Dorgali, viii.2010, 40°16’58.34”N, 9°38’15.30”E, 0.50 m depth, 42 lv on brown algae,

273 lv and sh on Posidonia oceanica (BA); undefined locality, 30 sh (Fra Piero legit, 1863,

Monterosato coll., MCZR–M–30056); undefined locality, 7 m depth, 7.viii.1981, 6 sh

(BA); Carloforte, Calasapone, 1 m depth, bioclastic sand, viii.2009, 7 sh (BA); Calasetta

S. Antioco, 1977, 22 sh (BA); ‘Punta Cavallo’ (=Capo Coda Cavallo), 30 sh (Fra Piero

legit, 1903, Monterosato coll., MCZR–M–30057); S. Pietro I., 8 sh (IN); S. Pietro I., local-

ity Punta, 1-4 m depth, ix.1986, 17 sh (BA); Gulf of Aranci, viii.1977, 42 sh (IN); Porto

Ferro, Sassari, viii.1980, 20 sh (IN); Maddalena I., Punta Marginetto, 41°15’14.70”N,

9°24’49.26”E, iii.1980, 25 sh (BA); Costa Serena, 1.5 m depth, vii.1991, 5 sh (BA);

Stintino, viii.1977, 110 sh (IN); Isola Piana, Cagliari, vii.1984, 35 sh (IN); Golfo Aranci,

1985, 154 sh (BA); Capo Pecora, Cagliari, 1-2 m depth, vii.1984, 56 sh (IN); Bocche di

Bonifacio 100-200 m depth, 1977, 1 sh (IN); Sicily: Magnisi, 130 sh (MCZR–M–22168);

Magnisi, beached bioclastic sand, legit Oberling (13.iii.1970), 10 sh (BA); Ognina, Sira-

cusa, 10.x.1982, beached bioclastic sand, 16 sh (BA); Punta Asparano, Siracusa, ix.1985,

14 sh (BA); north of Trapani, xii.1981, 12 sh (IN); S. Giuliano, Trapani, iv.1983, 12 sh

(IN); Siracusa, iv.1982, 7 sh (IN); Lampedusa I., viii.1984, 8 sh (IN); Pantelleria I., local-

ity Scauri, 12 vi.1991, 13 m depth, 3 sh (BA); Italy: Porto Maurizio, Imperia, 22 sh (Sul-

liotti legit, Monterosato coll., MCZR–M–30117); Sanremo, Imperia, 6 sh (IN); 4 km

NW of Capraia I., 140 m depth, 1978, 1 sh (MCZR–M ex Mauro Pizzini coll.); Baratti

Gulf, Livorno 10 m depth, viii.1983, 4 sh (IN); Antignano, Livorno, 1.5 m depth Posido-

nia oceanica meadow on muddy bottom, viii.1978, 1 lv (BA); Giannutri I., 18 m depth,

vi.1983, 1 sh (IN); Giannutri I., Grosseto 47 m depth, vi.1982, 1sh (IN); Sant’Agostino,

Civitavecchia, Rome, 1977, beached bioclastic sand, 250 sh (IN); Marina di San-

t’Agostino, Civitavecchia, Rome, 1978, 5 sh (BA); Santa Marinella, Rome, beached bio-

clastic sand, 1977, 13 sh (BA); Torre Astura, Nettuno, Rome, 1977, beached bioclastic

sand, 10 sh (BA); Torre Astura, Nettuno, Rome, 1977, 4 sh (IN); Marina di S. Nicola,

Rome, 23.ii.1978, 4 sh (MCZR–M ex Mauro Pizzini coll.); Gulf of Napoli, bioclastic

sand, iii.1979, 1 sh (BA); Procida I., Napoli, 4.vi.1978, 1 sh (BA); Procida I., Napoli,

ix.1981, 5 m depth, 39 sh, (IN); Scilla, Reggio Calabria, xii.1987, 1 m depth amidst

algae on the pier, 5 lv (IN); Santa Trada, Reggio Calabria, 1987, 1 m depth, 4 sh (IN);

Scilla, Reggio Calabria, viii1987, 7 m depth, 4 sh (IN); Taranto, 2 sh (Monterosato coll.,

MCZR–M–30058); Croatia: Punte Bianche (Velirat ‘Veli Rat’, Dugi Otok) 83 sh

Iberus, 37 (1), 2019

112

(MCZR–M–22168); Trogyr, viii.1989, 6 m depth in Posidonia oceanica meadow, 7 lv

(BA); Salvore 1 sh, (BA); Salvore 40-60 m depth, 2 sh (IN); Umag, iii.1980, 82 sh (BA);

Slovenija: Portoroz, 1977, beached bioclastic sand, 5 sh, (BA); Greece: undefined local-

ity in the Jonian Sea, beached bioclastic sand, 18 sh (BA); Lagonissi, Athens, vi.1982,

25 sh (IN); Astypalea, 25 m depth, 5 sh (CS-PM); Mediterranean Sea, undefined locali-

ties, 6 sh (Monterosato coll., MCZR–M–30128).

Alvania fractospira Oberling, 1970: 1 syntype (NMBE); Croatia: Trogir, 6 m depth

Posidonia oceanica meadow ix.1989, 2 sh (BA); Greece: undefined localities of the Ionian

Sea, beached, 5 sh (BA).

Alvania lineata Risso, 1826: Spain: Fuengirola, 30 m depth, viii.1985, 45 sh (BA);

Malaga, 5-10 m depth 11.xi.2009, 30 sh (BA); Tunisia: Gulf of Gabès, Kerkennah, 1974,

24 sh (BA); Gulf of Gabès, Kerkennah, loc. Sidi Youssef, vii.2015, 200 sh (BA); Sardinia:

San Teodoro, 1 sh (BA); Cala Gonone, Dorgali, Sardinia 1 m depth, viii.2010, 25 lv

(BA); Maddalena I., Sardinia 1989 24 m depth, 1 sh (BA); Gulf Aranci, Sardinia, 1985, 3

sh (BA); Oristano, Sardinia, viii.1978, 4 sh (BA); Cala Gonone, Dorgali, Sardinia 2-4 m

depth, viii.2010, 61 sh (BA); Sicily: Salina I., 35 m depth, vii.2002, 500 sh (BA); Levanzo

I., Punta Altarella 31 m depth, 3.vi.1991, 6 sh (BA); Marettimo I., Egadi, Sicily, 19 sh

(BA); Ognina, Siracusa, Sicily, 13 sh (BA); Magnisi, Eastern Sicily, Beached, legit Ober-

ling (13.iii.1970), 35 sh, (BA); Portopalo di Capo Passero, Sicily, 1 sh (BA); Lampedusa

I., Capo Grecale, vii.2009 50 m depth 10 sh (BA); Cannizzaro, Catania, Sicily, 35/43 m

depth, 61 sh (BA); Ognina, Siracusa, Sicily 1984, washing algae, 24 lv (BA); Lampe-

dusa I., 30 m depth, 53 sh (BA); Lampedusa I., Cala Calandra 30 m depth, 30.iv.1991,

46 sh (BA); Vendicari I., 28 m depth, 23 sh (BA); Ragusa, Secca of Roventa, Sicily 27 m

depth, vi.1989, 8 sh (BA); Marettimo I., Egadi I., 10 sh (BA); Malta: Malta I., beached

bioclastic sand, 1 sh (BA); Italy: Giglio I. 27-36 m depth, 45 sh (BA); Capraia I. 200-280

m depth, 1979, 1 sh (BA); Torre Astura, Rome, 1977, 1 sh (BA); San Felice Circeo,

Latina, 30 m depth, 10 sh (BA); Pozzano Reggio Calabria, 40 m depth, viii.1986, 1 sh

(BA); Secche of Tor Paterno, Rome, 30 m depth, 1.x.1988, 4 sh (BA); Secche of Tor

Paterno, Rome, 45 m depth, 18 sh (BA); Santo Stefano I., Ventotene I. 40 m depth, 14

sh (BA);Ventotene I., 1979, 25 sh (BA); Capraia I., Livorno 140-180 m depth, 1978, 16 sh

(BA); Secca di Torre Flavia 26 m depth, 1986, 64 sh (BA); Scilla (RC), 41 m depth,

16.xii.2006 legit Marconcini, 48 sh (IN); Croatia: Umag, beached bioclastic sand, 73 sh

(BA); Umag, 1978, 7 sh (BA); San Lorenzo, 5 sh (BA); Slovenija: Portoroz, beached bio-

clastic sand, 1 sh (BA); Greece: undefined locality, 29 sh (BA); Crete, 34 sh

(MCZR–M–30127); Crete, beached bioclastic sand, 5 sh (BA); Mediterranean Sea,

undefined localities, 75 sh (MCZR–M–22149);

Alvania elisae Margelli, 2001: Italy: 1 paratype, Capraia I., 2-15 m depth (CS-PM, ex

M. A. Fontana Angioy coll., Rome); Capri I., unspecified depth, 7 sh

(MCZR–M–30060); Elba I., Capo di Fonza, 25 m depth, 80 sh (EQ), 7 sh (BA); Elba I.,

Capo di Stella, 20 m depth, 13 sh (EQ); Elba I., 3 sh (MO).

Contributo alla conoscenza della malacofauna marina delle isole adriatiche. 2. Nota sui micromolluschi conchiferi rinvenuti sulla costa meridionale ed orientale dell’isola di Krk [Veglia] (Croazia, Mar Adriatico Nord-Orientale) / Contribution to the knowledge of the marine Malacofauna of Adriatic Islands. 2. Note on the shelled micromolluscs found around the southern and eastern coasts of Krk Island (Croatia, North-Eastern Adriatic Sea)

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Nov 2020

Le conoscenze sulla malacofauna del Mare Adriatico orientale, sebbene sia stata investigata sin dalla metà XIX secolo, risultano ancora parziali. Presentiamo qui i risultati dell’esame di sedimenti prelevati nel piano circalitorale della parte orientale e meridionale dell’isola di Krk (Croazia). In totale sono state rinvenute conchiglie appartenenti a 372 specie, alcune delle quali mai o raramente censite per il settore orientale adriatico o per le acque croate. / The knowledge of the eastern Adriatic Sea malacofauna, although this has been investigated since the mid-19th century, is still partial. The outcomes of the examination of shelled micromolluscs found in sediments of the circalittoral zone from SE Krk Island (Croatia) are here given. 372 species were found, some of which have never been recorded before for the E Adriatic Sea and Croatia

Contributo alla conoscenza della malacofauna marina delle isole adriatiche. 1. Nota sui micromolluschi marini conchiferi rinvenuti sulla costa settentrionale dell’isola di Lastovo [Lagosta](Croazia, Mar Adriatico Sud-Orientale) / Contribution to the knowledge of the marine Malacofauna of Adriatic Islands. 1. Note on the shelled marine micromolluscs found around the northern coast of Lastovo Island (Croatia, South-Eastern Adriatic Sea)

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Jun 2018

The outcomes of the examination of shelled micromolluscs found in sediments of the circalittoral zone from N Lastovo Island (Croatia) are here given. 407 species was found, many of which constitute are new records for the E Adriatic Sea and Croatia, among them: Parachiton africanus (Nierstrasz, 1906), Sinezona semicostata Burnay & Rolán, 1990, Dikoleps umbilicostriata (Gaglini, 1987), V

Notes on some taxa of the Alvania lineata-complex with the descriptions of three new species from the Mediterranean Sea (Gastropoda: Rissoidae) Notas sobre algunos taxones del complejo de Alvania lineata, con la descripción de tres nuevas especies del Mediterráneo (Gastropoda: Rissoidae)

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