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Abstract

A checklist of fishes of the order Acropomatiformes


 !"
#!#$%&'(
)'*(*$+!
+,-."'/"!0$,
0'0"'1!1""!
1"2!'345
6732)3$3
4"ACROPOMATIDAE)$8!
+96
:9      6  62  3  0"    
0"";"
"  " " 3&" 95
5"  9 3
/<3%;"
"26$6"9;6!
268  "9   "9 , 3 # ; .  "
3                    
 ;3 -5 = !,3 
!/.>9?-!9@
  3  (3  7    )    3  $          
""  =              
-."1"3
)&@A1Acropoma .1
 .    1  $'!  !3  @3  "   
)B"C""3
> ,D!>6:
0;93
Acropoma arafurensis  !"
/.:#1E8FG
436!>"373HH3H32H$686(8(6$"
"I@"18JK1!!JK&D8JK1
!!JK&3
%>17>A>/@."13
Acropoma argentistigma  
/.>$436,>"3733'83
!""0.;.3
%>17>A>/@13
Acropoma boholensis # 
 ? -  ;3  !' >"3
733'83""%".@>303
%>17>A>/@@>1I+=1#
103
Acropoma hanedai$!"%&
 -8,!,;3-3 #3 3 E" A3 @3((
""E6@L3
%>17>A>/@L3
Acropoma heemstrai' !(
   /.    )  2  FG  43     ! F>"
73(,3 ""I%.13
%I-=C1>/3
Acropoma japonicum')*!+%"
 )B$,8,#343""L13
!J'2M@,$J!M!N&>!'!3
%>17>A>/@:0;1#1L3
Acropoma leobergi ,- !"
   0.;  8  8'  43  69    L3  >"3  "  "
;=3
%>17>A>/@0@:3
Acropoma lecorneti!"++
 4 8$$ ,8 ;3  #" 3 3 ! " "
@13O03@#!(3
%>17>A>/@@#OL3
Acropoma neglectum' !(
/.)2FG43!8>"
73(,3""&)C71>3
%>17>A>/@713
Acropoma profundum !.
  /. '  4   ! F1 %" 3 8 "
"1>10;3
%."""3
Acropoma splendens /01!" "2
 +"88,8 -3 > -3 33 ! ""
)/
/.)2F2G43!(2$>"
73(,3""-1>3
%>17>A>/@1>/>/0)3
)DOEDERLEINIA 1
 1 $$! !23 43 "   
1    %P  "  ""9  "      "
-"$2-3
 L1" 8 !,,3 -3 "   
*"""3
L1"82,3-3AN"N
L1"3
48'!,83A313@3-3@33'3-3
"4"
"
>6:0;963
A=9"3
Doederleinia berycoides /345!+("2
*$28281=3)3@34$28
""L3
 1%P$$!%.3.3:3:
3'233!2"".""L3
 48'!3A313@3-3@33'
,843$""/I#+#13+=>30
!J,$M',N@JM!,N&3%3,2!'3
%>17>A>/@ >6:  0;    ."  "    @  0    
  E3        >        @
#9>/3
4"BANJOSIDAEL8
-        L       "      3
%  "  "  "  9        "  
;9<
    "3 # .
;3%!3-"3
)BANJOS .
 .    1  $'  "     7  "
C9"  1P$(#9
  "      "    "  .  $2(22  
33
 .  $2(  22  "      7  "    
3
>6:0;93
Banjos aculeatus !(
--2FG436!6866
6 F>"73('3 !
Q3
%>17>A>/@&3
Banjos banjos /6*!+.72
 7 $'(!( 73 3 3 3 13 ,  "
"1L3
.$2(223@R313@3*3""1
LA73
--2FG436!6
F>"73(' 3! " "  @ >
*:3
%>17>A>/@         
 L1#1
+.>:3
Banjos peregrinus !(
   -    -  2  F'G  43    F>"
73('3!1@"3
%>17>A>/@13

 %P  1  %P $$!2 %.3 .3
:3  :  3  '$  3  33  @            
"""   7$'(    ." L
S-(3
4"BATHYCLUPEIDAE)$8(
-9>6:0;)-53%
"9;"
9;'!8"9;
9;(!"9;9;
;#3-
"553O"!T3
3
)&@A1BATHYCLUPEA.
.$8!"."
""3
'@88'2*8$(3
Bathyclupea hoskynii 6!+"!
.$83-3@3*313(3$3'!H''!
43'""1J!M'N@8J'(M'N&>3,
$$63
%>17>A>/@>/3
Bathyclupeanikparini,- !.
0.;'!243'O">.3,'3"
":0;!J$3M@$J$3M&,,6,,3
Bathyclupea schroederi 6!"7
%.8(@3(243"")
-5!J!M@28J!8M:38$2$3
%>17>A>/@3
)NEOBATHYCLUPEA 0.;
0.;',O">.3,'3,
:8!3""3
:3
Neobathyclupea argentea /'8!+"72
)$8(133A313@3-3@3803
!;3',""I@>3:>!(,3
%>17>A>/@3
Neobathyclupea elongata/-!"(%2
82,O">.33,3((F88$G43"
"13!JM1'J,(M&'(6'2,3
%>17>A>/@#013
Neobathyclupea gracilis /9!"&+2
48!$03A313@3-33$,3!!!'43$
""/I-.">3>J8MN@2J$M!N&3
%3,(2,3
%>17>A>/@>3
Neobathyclupea japanotaiwana/,- !.2
 0.;'!436O">.3,'3
@0;!!J,,3'M@!!J83M&'$!3
%>17>A>/@L3
Neobathyclupea malayana /$!"!&2
:8!41&538!03!3"
"412J!(M12J!3$M&13!'(8'941
2J83'M1(J'83,M&3!(,!$3
%>17>A>/@>9@#3
Neobathyclupea megaceps /9!"&+2
48!$03A313@3-33$,3!!!!432
"">"3-0$JMN@'JM(N&
3%3,,2',3
%>17>A>/@:0;3
Neobathyclupea melanoptera,-' !7
0.;)0.(43!
'(S5'$3,-"23'8!J@8'3',J&2(2622
3
%>17>A>/@-"I1>/3
4"3:L1"8
-9>60;3&;31;
C31;,26"3+;
26"3E6'"
"3!"
@88'3
)CHAMPSODON )B
)B$(2"!)B"
""
/"$8,!"/""
""3
3
Champsodon atridorsalis 6*:!"7.
/@.8('3-.-33>3E"
A3@3(!43(903%!""."O3
%>17>A>/@:0;O0>3
Champsodon capensis 4!" +
78$3+313+13S33!
''032;3""1##"13
%>17>A>/@>/71E"#19
1"-3/3-3
Champsodon *mbriatus '$!" %
")8,3A3134#33!F8!G
('$03$$""0#->3-.>3
*>3A31333
%>17>A>/@*>3
Champsodon guentheri4!" +
78$3+313+13S33
!''3""03
%>17>A>/@>6:0;L03
Champsodon longipinnis$!"7.
-.8('3-.-33>3
E"A3@3(43,903#!""I-"=.03
L3
%>17>A>/@>6:0;L0>@
#3
Champsodon machaeratus :*!"".
@88'#88'3!,$438""
@2J,'M2J,2M1'2J'2J'M&Q
3
%>17>A>/@3
Champsodon nudivittis /4$1!+"%2
/"$8,03+313@313:3133!
""-"@313:33
78$3+313+13S33
!',""E>3->3>13
48'!3A313@3-3@33'$!43
""003U >31-
0J,2M@'JM'N&3
%>17>A>/@>6:0;->03
Champsodon omanensis 4!" +
78$3+313+13S33
!',032;3""1/3
%>17>A>/@:>/%.)
1=)C713>
!(6!$
$,62
.3)<)71
)C3
Champsodon pantolepis :*!"".
@88'#88'3!(43""
'.%$J''M1(J,8M&:3
%>17>A>/@>6:0;:L>
03
Champsodon sagittus :*!"".
@88'#88'3!(!43""
O">3!J$M1$J8M&:6!23
%>17>A>/@:0;>@)03
Champsodon sechellensis 4!" +
78$3+313+13S33
!'03!;3""1"3
%>17>A>/@1"->3
Champsodon snyderi;!"!
4=83.3:3--360"3E33'
133$038;32'"":."E"03L3
%>17>A>/@:0;L?1#13
7#;>@#3
Champsodon vorax ')*!+7(
!)B$(203S313+3$(2""
#183
78$3+313+13S33
!'!03!;3!""1-.-3
%>17>A>/@>60;-O@0>33
4":$88V+"78!
199";
->6:0;3#"";
"3".
"3%3&
$@3(9$".3
)APODOCREEDIA
8'$'2("
"3
/3
Apodocreedia vanderhosti 85!".+
8'$37313133!,'2(
43"">>3%"-=C3
%>17>A>/@:>/%"-=C
%13/(3
)CHALIXODYTES1=
!1=8'!("!1="
3
/3
Chalixodytes tauensis 6*;!".&
!1=8'!3A313@3-3@3$(!43'"
"7>31013
!18,23-3@3*313383$
F8,(G$843""%>3!J'M1,,J'M&1"3
%>17>A>/@>60;#>3>/-
-1037"/R
+"">3#>/ 
""4.E.(!',3/
5"3
)CREEDIA/"
/"$8$8$"$/""""3
43
Creedia alleni :!"+&
@8$!03313:338(3843""
I)>34:!J,32M1,J!8M&3
%>17>A>/@3
Creedia bilineata *#*!"+(
1?8$2L3L3>"33!'3!43
6,""E">.>3?">337".">3/.
03L'J83M@'J23,M&3
%>17>A>/@L3
Creedia haswelli /1!++!2
% 7"$$03+313@313:33(!,2,"
"@*0L.@313:33
$/"$8$03+313@313:33!!88"
"-@313:33
%>17>A>/@3
Creedia partimsquamigera :!"+&
$@8$!03313:338(3!!43'
""I#1""@313:3!!J,(M1,J(M&3
%>17>A>/@3
)CRYSTALLODYTES4
48!!8"4"
3
3
Crystallodytes cookei 9!"&
48!/30303-33$32!8
""+/>3*>3A31333
%>17>A>/@@)*>0)3
Crystallodytes enderburyensis 6*;!".&
1=8'!('43,3A313@3-3
@3$""7&">05>3
%>17>A>/@:0;3
Crystallodytes pauciradiatus :!"+%
@78$,03313:338$3
'!436""I.&>3
%>17>A>/@&>3#3
)LIMNICHTHYS:
&:8'2$"&:"
3
>60;9"3
15?388894.)3
Limnichthys fasciatus !" .
&:8'733-33,3!2$3!;3'"
"+*>3
&:"8',3S33'""
)>:3
%>17>A>/@9
L7".">/>3
Limnichthys marisrubri6' !
&4.)1W=@.
@133,$$436!""&>)
C713
%>17>A>/@)C713
Limnichthys nitidus *!"%+
&18,$3-3@3*313!33''243
""0-=C3
&1=8(3A313@3-3@332$432
"".>3.->33
%>17>A>/@">/71
0;L3
#/--&@1"""?3888283
Limnichthys orientalis #*$!"""
&?E/.888>"373'(3$43
#'#,#(#2#"".>>?">
/.L3
%>17>A>/@7".">3
Limnichthys polyactis:!"(+
&@82$@3S3L3S33,3!,!43""
:>":@S'JM12'J'2M&3
%>17>A>/@@S3."
53%"3
Limnichthys rendahli!"%+
&08,$73%3-3:3!("
".>3@S93
%>17>A>/@@S3
)MYOPSARON1.
'1.'8"'1."
3
Myopsaron nelsoni*$9 !
'1.,436',#(6863@3-3
@3131313@3'""I#6 >
/>L3
%>17>A>/@/>L3
)SCHIZOCHIRUS:
(:8''"(:"""
/3
Schizochirus insolens !" .
(:8'733-33,''03(;3!943
!!6!'""-"@313:33
%>17>A>/@3
)TEWARA)I
) )I8!!2'")  )I"""3
/3
Tewara cranwellae '<!"&&
)  )I8!!3@3S3>33(!2'03,"
"1M":*@S3
%>17>A>/@@S3/."
,3
4":0:"8'$
+6;.
-931;
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,68"5"
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-5,3-"3
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08!!(;33/3#3?
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""3
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3
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3
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"$9
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,(6(3
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>3
Epigonus bispinosus ' !+
/.)$(243,(2S5'!$3
""I-1>/J,$3'M1
'8J3(M&',!6',,3
%>17>A>/@:>/3
Epigonus carbonarius !!
/.-,(43!>"3
733,$3""I@.*>$J,$M1'J'M:
-C>
%>17>A>/@-C>3
Epigonus cavaticus (
>/.1.2!43!',>"373
3,'3""O*@>
03

%>17>A>/@03
Epigonus chilensis !
/.'!43!',F>"7
3,8""/I#'J,2M12,J2M:!,6!'3
%>17>A>/@I#3
Epigonusconstanciae/'4!++ 2
*)$$4>""-
1>"3
 08,'8;327313&53/3#33
3133'3!""$F'$G3-
1(J$M1J!M&$683
%>17>A>/@-13
Epigonus crassicaudus 8!"%"
.,/%8,88(3-3@3*3@3
1323!""IO#3
%>17>A>/@#3
#/--&@108$($8
@.@.38$(3
Epigonus ctenolepis6*;**!"+&
-=1.8$!88;3L3L3>"33
!3!""I/0;L3
%>17>A>/@E1L3
Epigonus denticulatus ;!"%
%=8,$836330313C30
#333""-16,3
%>17>A>/@>60;3
Epigonus devaneyi'!"+%
)8$,,;3!03133!83""I
@.>*>A3133!6!'23
%>17>A>/@*>3
Epigonus draco !%
/.,436',1%"3"
"1>2J'!M!'N1,$J8M'N&!86(!3
4.7322436'4533!""
":@00@)113
%>17>A>/@:0;00@)1>
O3
Epigonus elegans $-!"+7
08$(22;3303031>3
/33""@=7J,M1$J!2M:30;,3
%>17>A>/@0;3
Epigonus elongatus $-!"+7
08$(,,;3"3-.3/3>"303
/33383!""@3>/JM1,J'M&!3
%>17>A>/@@>/3
Epigonus exodon !
!/.-$,436'FS5@3!',!"
"7R>>/',6'$
%>17>A>/@>/3
Epigonusfragilis/==!"2
LL8',3;3-3#-33
3""*./>*>3
%>17>A>/@*>I->L3
Epigonus glossodontus'!"+%
)8$,43603133!83""
-"I0*/>*
>A3133!((3
%>17>A>/@*>3
Epigonus heracleus $-!"+7
08$($!;3,303031>3
/33""130;'!J!,M1(J,M:(,6(23
%>17>A>/@&I@S3
Epigonus idai ' !+
/.)$S5'!$3274, Figs. 6B, 7B, 11 (type
locality, Off Mozambique, southwestern Indian Ocean).
%>17>A>/@Southwestern Indian Ocean.
Epigonuslenimen/*1!"&%2
:"8!,!733-33833'"")
3&:!,6',3
%>17>A>/@0;@S
3
Epigonus lifouensis  !&
/.-!>"3733('43
""south of Lifou Island, Loyalty Islands, New Caledonia).
%>17>A>/@Macclesfield Bank (South China Sea), Loyalty Islands (New Caledonia) and Indonesia.
Epigonusmachaera !
/.'$43',2$8>"
73,89#7@S'!J$M12,JM:
2262'(3
%>17>A>/@@S3
Epigonusmacrops/8!" 72
+!8($$;32643,"":
1!JMN1JM,N&>3
-#.+882(;36!>"3733''3
""I#*>4J$M18J!2M:)
><3
%>17>A>/@3
7."""/.33
Epigonus marimonticolus $-!"+7

08$(,!;X"3-.3/3>"3
03/33383!""@3>/J$M1,(J!M&'6
'3
%>17>A>/@@>/3
Epigonusmarisrubri>?;*5 "
ES =E8'436!C>
L>""3,3'""@)C7
1L3
%>17>A>/@71)13
Epigonusmayeri !!
/.436!'6,>"3733,$3"
"/I8J,,M1,J,M&!8(3
%>17>A>/@I
Epigonus megalops /*6<!"!2
%17I8'',303!$;3!3A313@3-3
@3'$($""+=>J,M!N@
!J(M,N&03,!$$(3
%>17>A>/@03
7&-7E1"""#"/.(3
Epigonus notacanthus $-!"+7
08$(2';3303031>3
/33""1")=J(M1$J2M:30;
!3
%>17>A>/@0;3
Epigonus occidentalis '8!+"7
)$8(!!3((;3!(133A313@3
-3@3""I:>!23
%>17>A>/@:3
Epigonus oligolepis 1!"(.
-"82'28;3,3-3#3S33'(3!"
"41E":#3'!('J!M@
$J'M:!3
%>17>A>/@3
Epigonuspandionis/'8!++!2
0)$$3
%>17>A>/@9>/3
Epigonus parini $-!"++
8$$;3O">.3323(""
130;,!J!,M1'J'M:6'3
%>17>A>/@10;3
Epigonus pectinifer 1!"(.
-"82'$(;383-3#3S33'(3!"
"I)JM@(J,!M!N:/3,'!6,3
%>17>A>/@0;>/3
Epigonusrobustus/8!"(2
)8''3;3!7343-33
13A13373!32""13#10.
!''!'2,(29""
+!8(3
*8223-3@3*313833,
""I#010432!('(3
%>17>A>/@1#3
Epigonustelescopus/!+! 2
*7$!38;3!>"3@""@
43
7;C$('>>313""1"
>"
*#$8'!)313+3313(322"
"-1">"3
%>17>A>/@>/0;3
Epigonus thai,-89 !
0.;00.;'
43L3>"33,3!"">/1
(J''M2N@8$J(M8,N&!!6!!3
%>17>A>/@1>/3
Epigonus tuberculatus !&
/.-!,43',>"373
3(3'""'J',M18,J'N&'J',M18,JM&#6
E>>/'2!6,23
%>17>A>/@1#6E>3
Epigonus waltersensis $-!"+7
 08$($;3'303031>3
/33""-!!J(M1''J'M&
3>/2'62(3
%>17>A>/@&4S3
)FLORENCIELLA-%4
1-%48(,($"1-
%4"""3
/3"3
Florenciella lugubris !"7%
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S33!'32""&C>/J,$M1(J(M&
J(M1(JM&,3
%>17>A>/@>/3
)MICROICHTHYS7B
'7B$,"'7B"""3
303
Microichthys coccoi )!+%
'7B$,41.""-
1">"3

%>17>A>/@."1-1"-3
Microichthys sanzoi @!"%
'(1Z8,;36$3003>3@3(3,
3""1-1">"3
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-8283
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-%48(,(!"-
%4"3
9"93
Rosenblattia robusta !"7%
-%48(,(!43!3-3#3
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3
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""3
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4"""3
:0;93-"3
Sphyraenops bairdianus '!+7
)$(!,03$;3-3*3@3#3"
"#3
4822;3,#3/373133/3-31R3
/R33$3""130;(J!M1((J,$M&F
G3
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99 953
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;.3"
I""
.3
3;"."
3"".3/'
@88'0538$8.
4 #1"&>6
0;41""8$3"'3"
""-E"8823
)GLAUCOSOMA.1
0.1$'!("0
7"C3
#$2,("#"""3
/"8,!""/""
""3
48!'!,2"04"
3
43
Glaucosoma buergeri 6*!+.%
07$',>"""*3-313&
32""1L3
021$$3133021R3,'"
"1#
048!'033@313033$,!,243$"
"&->0 3
%>17>A>/@1."##
1L3
Glaucosoma hebraicum 6*!+.%
07$',>"""*3-313&
3232""*>:
93
#$2,/I7(""17
3
%>17>A>/@1."#+3
Glaucosomamagni*cum/4$1!"!%2
"/"8,-3Q3-33!!03!"
"">13
%>17>A>/@&5):1"Q3
Glaucosoma scapulare 1!++!
07"-+"$$03+313@313:33,!
!!'03!""">13
%>17>A>/@Q0L.@1:3
4"3E$2!
>60;30""0*
(@3(3
)ACANTHAPHRITIS)B
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"""3
/.E8(!'!"
/.E"""3
-E"82'"( -E""
3
43)"1=.@.88(3
Acanthaphritis barbata/*!"7&2
/.E8(!3-.-33>3E"
A3@3''!4363#I?&03L3
%>17>A>/@L3
Acanthaphritis grandisquamis ')*!++
$)B$$3-.-33>3E"A3
@3''!4363#I?&03L3
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Acanthaphritis ozawai /61!"(!2
( -E"82L37313:333,''433>
/I32J2MN18J,2MN&3
%>17>A>/@3
Acanthaphritis unoorum ;:$!""%
1=.@.88,',43,>"7
3'!3'/I*.*"0L3
%>17>A>/@L3
33738$83
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)DACTLYLOPSARON0
0888"0
""3
/3
Dactylopsaron dimorphicum 81!""
0"088O">.33!
38F22G43!3130;,J'M182J2M:3
%>17>A>/@@=710;'6(3
)ENIGMAPERCIS:"
:"8!(8":""
""
3
Enigmapercis acutirostris!""
088O">.33!32F2'G433&3
0;,JM182J!,M3
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Enigmapercis reducta *1!"&7
:"8!(0373S313@313:338!,6!(83/I
+7#"@313:33
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13A"6
"!3
)HEMEROCOETES O
%O$!2!"
11"""3
#/--&@14$#
43
Hemerocoetes artus :!"("
%@828@3S3L3S33(,8'433@S3
%>17>A>/@@S3
Hemerocoetes macrophthalmus 4!"!.
%78'3-3@3*313$3!3
2!,3#@@S3
% 78'43@&5333$31
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%>17>A>/@@S3
Hemerocoetes monopterygius /6*!+ !2
11$1">"3
'3@@S3
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313@S3
%>17>A>/@@S3
Hemerocoetes morelandi :!"("
%@828@3S3L3S33(,8433@S3
%>17>A>/@@S3
Hemerocoetes pauciradiatus 4!"!.
%78'3-3@3*313$3!32!
,3#@@S23
%>17>A>/@@S3
)MATSUBARAEA .
!48'!$2"!4"
90"!-8!(>3
'.8,!"'."
""3
1=8(22!48'!3
/3
Matsubaraea fusiformis /9!".&2
!48'!3A313@3-3@33'$243!3
#"03+=>303

'.8,!L313*A313%3S33'
436(03;36!3.;.E03>1
L3
%>17>A>/@0L3
)OSOPSARONL1.
+L1.8'("*L
1""3
3
Osopsaron formosensis *!"+%
+E18$,L3-33!$32,4363
33
%>17>A>/@3
Osopsaron karlik !"+%
+08$,L3L3>"33!3'!,$433@=7
10;,J'$M1$(J!M:3
%>17>A>/@@=710;3
Osopsaron verecundum/=1!" 2
*L1"803A313@3-33'3
(!'243!31"L3
%>17>A>/@L3
)PTEROPSARONL1"
*L1"8'2"*L
1""3
@3
Pteropsaron dabfar9 !.
*>',436!+=-
!38,$!!J@3!,'!!J&0$6$(3
%>17>A>/@03
Pteropsaron evolans =1!" 
*L1"803A313@3-33'3(!
'243311L3
%>17>A>/@L3
Pteropsaron heemstrai :!"+
*@8$L3+331>3>"31303@3,
4363/I0.7"3@1!JM1!J''M&3
%>17>A>/@I@13
Pteropsaron incisum'$!" %
*)8,3A3134#33!F8!G('2
03$23/I+">3+>3*>3A3133
%>17>A>/@*>3
Pteropsaron levitoni9 !.
*>',,43'6(+=-
!38,$!!J@3!,'!!J&0$6$(
%00@)3
Pteropsaron longipinnis !
*>',$436,F7;&>
/0-J,'3(,M1!'J,$32''M&
#":0>3
%#":0>3
Pteropsaron natalensis/:!"+2
+@8$L3+331>3>"31303@3,'
43!6'3/IE""@13
%>17>A>/@@13
Pteropsaron neocaledonicus -!"("
*47828#3>60333'
'84331>030;3
%>17>A>/@@#3
Pteropsaron springeri*=* (
*1L2!(,436,#23
&8J!(M,$N@!JM,N&@>03
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)SQUAMICREEDIA 7
$78"$7"
""3
/3
Squamicreedia obtusa *!"!
$78E313O..3*33(38
43,6(3',336#L323
%>17>A>/@3
4"HOWELLIDAE /"$88
7!29
959;;2$;
$8"9;C6;9
;!($"93->6:
0;3-5=33
)BATHYSPHYRAENOPS0
 08!!$"!0"
3
0E8$2(2 ""
% /"3
Bathysphyraenops declivifrons 1!"(7
4".82(">3/.3.3@.3
11173'('43,""0;,J!$M1!J,M&O=32(,3
%>17>A>/@:0;3

)HOWELLA /"
% /"$882!'"% /""""3
0 *18!8!"0 *
1"""3
@8!!'$"@"
3
48'!("Schistoperca macrobrachium Fowler, by original
designation):
530L36#3Q880.;23153
Howella brodiei 4$1!+""
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"+*>3
% 0Q88Cybium v. 15 (no. 2): 118 (type locality, Atlantic,
44°12'N, 20°04'W, depth 1100 meters).
%>17>A>/@%#+*
>@S%#0Q88$
>=3>31"!,6
$83
Howella macrobrachium /9!".&2
48'!3A313@3-3@33'
(438""U01>[
#1>03
%>17>A>/@03
Howella pammelas /34!" &2
0 *18!03:33133,8!03'
"":>3)U>3
%>17>A>/@)U>3
Howella parini1!"(7
% 4".82(">3/.3.3@.311173'22
43""@0;!J@2(J&3
%>17>A>/@L.
Howella sherborni /:!"& 2
@8!%"733!'$43!8"
"!!J,M1!'J!M1(J'M&,J'8M&I1$,68,3
%>17>A>/@:">0;3
>9!!J6!,J@'J13L(
!""3
Howella simplex /!"&&2
!08!!3/3#3?A33!
3(843!""#0J''M@2J'!M:
233
%>17>A>/@L@)93
Howella zina1!"(7
% (4".82(">3/.3.3@.311173'2(43
""@30;8JM@'J'M&O=32'3
%>17>A>/@L3
)PSEUDOHOWELLA 4".
* 4".82(28"* 
4"."3
/3
Pseudohowella intermedia 1!"(7
* 4".82(">3/.3.3@.311173
'$43!"":30;'J$M@'J(M&O=3('83
%>17>A>/@0;3
4"008)%1$
0482'8"*!F"
3233G
+1788,8''"&!F"
8(!33G3
+/#-.)(8"&!
FG3
+1L''2"&!F
G3
+1#2'"&!F
G3
+)%1))%1
$8&!.$,,3
4."3L"
305
31."
30=."
;3
)LATEOLABRAX .
*!.1$'!"&!#$$3
""""3
&!.$,,,!"&!#$$3""
""3
:0;3301"E"8('
-"3
Lateolabrax japonicus/-!++2
&!##O$$*3@3033$,
""1L3
*!%P1%P$$!$3@
"""*!9."
L3
%>17>A>/@LE@1:3
7&-7E15E"8(%8$(2$3
Lateolabrax latus1!"%(
&!.$,!!;3

&!E"8,2L3L3>"33(3'6(,'43,,"
"1=.?03L3
%>17>A>/@L."3
Lateolabraxlyiuy/891!+%%2
%-#$''!8,33;3""#>
@#3
&!3."$,,@3-313>3@3-38"
"/1#3
*!%"$22$3!(;3""
?=7#3
*!%"$22$3!(;3!""
?=7#3
&!3?.1.88,3
%>17>A>/@1E#3
4"0)$,8
1;9""=
-@S3&"""3
-"C3+3+6<.3"3
>3+;3""
"""3/"
3,3
)CRAPATALUS)B
)B$($("()B"
""3
43
Crapatalus angusticeps /3!+(.2
&*$2'303@3S3>33(32038
;3!("")"+@S3
%>17>A>/@@S3
Crapatalus munroi 04!"+(
+&8$2-3@3-3O3'$32!43
3""%M&5#O13
%>17>A>/@O3
Crapatalus novaezelandiae ')*!+7!
()B$(3-3@3*313!323!$$2
FG033""@S3
& *$2,303@3S3>3323!'$"
"##@S3
&$$,303@3S3>332$$'3(
(,03';3'""/I0.@S3
%>17>A>/@@S3
3E88!3
%>17>A>/@3
)LEPTOSCOPUS)
&)$,8!"47"
""3
/3
Leptoscopus macropygus /6*!+.72
47$'(>"3&3,,03
!!;3'6(""0L.@313:33
&*FL343L3G$2!303@3S3>33,38
2,03(""7@S3
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"7@S3
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Lesueurina platycephala 9!" +
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  1822'43F030373133
""."3
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4"MALAKICHTHYIDAEL78
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"""3;
"3<9
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-.",23
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'%P$$!'"'%P"
""3
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9"$($-"
-"$2'+3
:0;9"?-'
=$3
Malakichthys barbatus## !
'??,2>"3733'$3!3
1"*L3
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Malakichthyselegans$#4!".&
'-?8'!$';3L31..
E.""33""1"L3
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>/313
Malakichthysgriseus B!++&
'%P1%P$$!'%.3
.3:3:3'233""."L3
108('2;3,03A313@3-33!3
'28""EL3
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'?-'43!%)'3
-3133(83!""@%:
3
%>17>A>/@3
Malakichthys mochizuki# 
'(?-',43!&3*'(3
3-3133(83!""&*.>
Q3
%>17>A>/@IQ3
Malakichthyssimilis# .
'?-','4348L3433(,
3""-03->1
03
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Malakichthyswakiyae=3$$!"%
' L*8,!!3;3-3#-3
33""E"L3
%>17>A>/@L3
#/--&@1 "E"8(3
)VERILUS 0"
50"$('"50""""3
/"$8(!"/""""3
)8(2")"
""3
)"?(=53
Verilus anomalus/4$1!+"72
/"$8('03+313@313:33"
"-"@313:33
%>17>A>/@Q)
13

(3
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88','$3
Verilus atlanticus /6*;!"+.2
-=.18$'!!,;3L3L3>"33!
3'""#1J2M@$J''M:/3('''83
%>17>A>/@#13
Verilus cynodon /4!"!2
78',3-3@3*3138323'"
",63I73A@16!3
)8($3;37343-3313
A133733!""I@1(6(3
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Verilus paci*cus/6*;!"("2
"-=.828'2;3L3L3>"33(3!"
">3L!J,M@!8J'2M&!6,3
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9#;>3
Verilus pseudomicrolepis /6*;!". 2
1=8''03A313@3-33$$3!$,
""/IO>3:>$J!$M',N@('J,M',N:$J'M,N@
('J,M,N:3
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Verilus sordidus 1!+7
50"$(,3;3(-3*3@3#3""
#3
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O=3>."3
Verilus starnesi # !7
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3
3
Ostracoberyx dorygenys 9!"&.
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'6(,3
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Ostracoberyx fowleri$!"&"
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Ostracoberyx paxtoni CD;5E;!""!
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Q3
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""$J(X1,!J,$X&I3
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4"3.$,8
"1
-.9>0;
3%""
"9"9;
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>>26',9(6$"9"'6$9
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".31**
;30"831
3O,T,3@
"3"".
=.3%6!3
!@3(3-"
"$,3
) PARAPRIACANTHUS1
*1$2(!"*
1"""3
*E=$2'2"*
E="""3
*8!"*
"""3
>6:0;3@')388'
-,@88'3"
"7."(3&3
Parapriacanthus argentea /-8!"&2
*8!7343-3313A133
73!303;33""@10.3$$
!3
%>17>A>/@-=C13
Parapriacanthusdarros841 !7
*7."($43(62FL/
143"">%M>
1"3
%>17>A>/@1"3
Parapriacanthus dispar/3!"&%2
**8!,'2F4-@*"0
SP13$3""?>1
>3
%>17>A>/@:0;>00@)1
>@#+"">3
Parapriacanthuselongatus/66*!"!!2
*-#837N&NF3G'203'
;33""/I4M>31I:M0"
O3
%>17>A>/@%"@313:30:33
3
Parapriacanthus guentheri /;4!+(!2
*E=$2O3E36E3S363)3:3'23
""713
%>17>A>/@713
Parapriacanthus kwazulu841 !7
* (7."(L/1
43!43""ES6@0.7"1
3
%>17>A>/@ES6@13
Parapriacanthusmarei!"(!
*4823#3/373133/3-31R3/R3383'
'8$432F(G3""-R>3+"R>33,3
%>17>A>/@@#3
Parapriacanthuspunctulatus 841 !7
*7."(L/
143'43!6(""0-
-=C
%ES6@1-=C1"#
-1M13
Parapriacanthusrahah 841 !7
*7."(L/1
432432""1/7"
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*.$223@313@3*3,F!'G3"
">->>3
%>17>A>/@>6:0;.13
Pempheris a3inis 66*!"!!
*--#837N&NF3G',032;3
3""0L.@1:3
%>17>A>/@*""Q->@1:3
Pempheris analis !"!
*:83@3S3>33'!2,03!(3"":
%"E>3
%>17>A>/@)7+*@.E
>39@S;*#3
Pempheris andilanaF6!"!%
*7O,243L/1
43$""I@"-3
%&-3
Pempheris argyreaF6 !%
*7O,$43!6'L/1
43$""+%@0."1"
%1"3
Pempheris bexillon !.
*!-7'!8436S5!2$3"
")#3,.#3$(2J1
'!3(2#>3
%-=C#-0#>3
Pempheris bineeshiF6 !%
*7O,43,6(L/1
43$""@>3
%>3
Pempheris bruggemanniF6 !%
*7O,43268L/
143$"":116+7R3
%7R-3
Pempheriscompressa/*!(" 2
:28O"@313:3(203,!;33""@
1:3
61$('03'F-3&31"
>"""3
*#$87Y338,3"">
13
%>17>A>/@"I@1:"
"&9@S*#37
;3
Pempheris connelliF6 !%
*7O,'436FL/
143$""ES6@11

%13
Pempheris convexaF6 !.
*!7O'('436FL/1
43""/1>2J$X$N@
,(J!(X,N&13
%-13
Pempheris cupreaF6 !.
*7O'($43!FL/1
43""-=C3

%-=C3
PempherisdarvelliF6 !.
*7O'243'6(L/1
43""L>!J!!M!3$N@,$J!8M$3('N&)
/
%/3
Pempheris eatoniF6 !.
*7O'2(436'FL/1
43ES6@13
%-=C13
Pempheris ellipse F6 !%
*7O,L/143
$(436!""17"/3
%>17>A>/@/3
Pempheris erythraea G$!+((
*E7W$22O3@36-3O3*33
!8$3""713
*7717
."1--'L/
14!38843268""17
713
%>17>A>/@713
Pempheris familia !(
*E-2FG436!F>"73
(,39""I/ >/>L3
%>17>A>/@/>L3
Pempheris 9avicycla !.
*77717
."1--'L/
14!38'43!6(""-;>#
>#"=3
%>17>A>/@>/)E"=1"#
3
7.""*#E3'!!3
Pempheris gasparinii*86* !7
*07(S."@3,($436
"">&1=3
%."""3
Pempheris hadraF6 !%
*7O,L/143
$243'""E."
7-3
%>17>A>/@.""3
Pempheris heemstraorumF6 !%
*7O,L/1
43$843,""7->3
%>17>A>/@.""3
Pempheris hollemaniF6 !%
*7O,L/14
3$!43("""74%+
-3
%."-3
Pempheris ibo F6 !%
*7O,L/143$
!4326$"">>-=C3
%-=C13
Pempheris itoi 9!"&!
*48!3A313@3-3@33(4323"
"..>341>3
%>17>A>/@>3
PempherisjaponicaB!++&
*%P1%P$$!=3.3:3
:33,,3""."L3
1"803A313@3-33'3$!(,$3""
-.L3
%>17>A>/@L03
7&-7E111%P$$!83
Pempherisklunzingeri66*!"!!
*E=$$1=3.3:3:3$33!6
'!$F,(G03(3""E)1:9
"*0"$(3
*(-#837N&NF3G'23
""E)1:9
*E=$$
%>17>A>/@)1O)3
Pempheris kruppi F6 !%
*7O,L
/143$!!4386""7)
?3
%>17>A>/@?13
Pempheris kuriamuria F6 !%
*7O,L/1
43$!,436""17*/3
%>17>A>/@/3
Pempheris leiolepis F6 !%
*7O,L/143
$!243!""#>S=3
%>17>A>/@."""L3+33
-3-313,8,3
Pempheris malabarica-!+&!

*##O$!*3@30332
!$3"-R->3
%>17>A>/@/0.>1+.#
13
Pempheris mangula-!+"
*#$87Y338,3""
O.FO=G>3
%>17>A>/@1:>3
Pempheris megalops F6 !%
*7O,L/14
3$!$43'""+%@01"3
%>17>A>/@1"9."L8223
Pempheris micromma F6 !%
*7O,L/14
3$!843,6(""17"/3
%>17>A>/@/3
Pempheris molucca -!+"
*#$87Y338,3"
-9L>3
]61$1">"3('3"
"C>3
%>17>A>/@71>6:0;11
-"
7&-7E1""*#*8$(3
Pempheris multiradiata ;4!+("
*E=$281=3.3:3:3$3
3!6'!$F,2G3""E)1:3
*/"$$,03+313@313:33''23""
1.70L.@1:3
*-"$$03+313@313:33,',(3"
E)1:90L.@313:3
3
%>17>A>/@@1:03
Pempheris muscat F6 !%
*7O,L/143
$'432"")/#>/3
%>17>A>/@/3E"$
88"-,822""L3&373
Pempheris nesogallica-!+&!
*##O$!*3@30332
!(3""-3
%>17>A>/@1--3
Pempheris nyctereutes=-!" 
*L&803A313@3-33,3
$8!!843'3""*.3
%>17>A>/@L3
Pempheris orbis F6 !%
*7O,L/143
$'43$"")C%713
%>17>A>/@713%
8($3
Pempheris ornata =$$!""7
*#)388'($;3,!(3
*-L88(733-33'$3436!'
,3""7"!,J!M8N1!$JM,N&1833
%>17>A>/@7>E>3
Pempheris otaitensis -!+&!
*##O$!*3@30332
!'0383""1">3
%>17>A>/@3
Pempheris oualensis-!+&!
*##O$!*3@30332883
""1>3F/>3G#>3
%>17>A>/@>60;71+-C%>9
7".">+*7>3-3
Pempheris pathirana F6 !%
*7O,L/14
3$'!438""%E"040
1+.3
%>17>A>/@1+.5>9."
9*#
8(882;3
Pempheris peza F6 !%
*(7O,L/143$
''43!""0/-=C3
%>17>A>/@."""9.3
Pempheris poeyi8!++%
*$$,03A313@3-33$3,(83""
#3
%>17>A>/@#3
Pempheris rapa !""+
*-88$0313,,,;3""*"
7>40"3
%>17>A>/@."""3
Pempheris rhomboidea G$!+((
*E7W$22O3@36-3O3*33
!8$03!;3'3""713
%>17>A>/@71-1713
Pempheris rochai F6 !%
*7O,L/143
$',43!""17*/3

%>17>A>/@/9."""3
Pempheris rubricauda F6 !%
*7O,L/1
43$'(43!""@"@"O-3
%>17>A>/@."""3
Pempheris russellii 1!+++
*%"$$$4>132$$3"">3
%>17>A>/@0.3
Pempheris sarayu 8* !.
*7'L/14
3!!43""EE!
I7 *>3
%>17>A>/@."""
Pempheris sasakii /=3$$!"%2
&L*8,-3#-333$
03;3"".L3
%>17>A>/@L3
Pempheris schomburgkii )6*!+.+
*-B1.$'$*3,3
"":>3
* !##O$!*3@303O32!$
""-53
*0"$(-3*3@3#3!3""#3
*823/3#3?A333
!43,6(3"">0#F17.G3
%>17>A>/@4=3)
-53
#/--&@1"""16O=38882!4C3,
2',3
Pempheris schreineri E$!"!%
*-678,C3-3@37L332
033""4@3=3
%>17>A>/@=3
Pempherisschwenkii8!+%%
* .$,,@.3 3@3>^3$!'3"
">>3
%>17>A>/@71@:L9
>4 O3
Pempheris sergey F6 !%
*7O,L/143
$'243!!""11/-
1713
%>17>A>/@713
Pempheris shimoniF6 !%
*7O,L/143
$'843!'""1E"3
%>17>A>/@."""3%"
3
Pempheris shirleen F6 !%
*7O,L/143
$,43!,""&"*!"
>/"4713
%>17>A>/@713
Pempheris smithorum F6 !%
*7O,L/1
43$,43!(""S=#>=3
%>17>A>/@.S=-;>"5
=3
7&-7E1""L3+33-3-3113,8,
82-;>=3
Pempheris tau F6 !%
*7O,L/143$
,43!2"")C10@
&"713
%>17>A>/@713
Pempheristernay F6 !%
*7O,L/143
$,!43!$""-R"-@0.#-
1"3
%1"3E""3
Pempheris tilman F6 !%
*7O,L/143
$,'43!8""CL)C713
%>17>A>/@)C3
Pempheris tiran F6 !%
*7O,L/143$
,,43'""11713
%>17>A>/@@73
Pempheris tominagai#*6** !.
*E?'S5!28!3!!2
432""161.!'J2M'N@2J,M8N&&"3
%>17>A>/@713
Pempheris trinco F6 !%
*7O,L/143
$,2;3'""%01+.3
%>17>A>/@1+.5"3
7&-7E1""L3&3782,9
"1+.
Pempheris ufuagari#*6** !&
*E?!S5!(83!43
""E.-%>%>L3

%>17>A>/@%>/L
Pempheris vanicolensis -!+&!
*##O$!*3@30332
!,3""O.>31#=>330;J!2M1((J,$M&3
%>17>A>/@71E=6@@
#107".">9
-1713
Pempheris viridis F6 !%
*7O,L/143
$,$43'!""1I+713
%>17>A>/@719."3
Pempheris wilsoni F6 !%
* 7O,L/143
$(43',6'$"")/4>/3
%>17>A>/@/3
7.""
/3
Pempheris xanthomma F6 !%
*!7O,L/1
43$(43'86,"")7- 
?3
%>17>A>/@)?9."""
"43E-31&3+-",3
Pempheris xanthoptera4!"7+
*!8($L34313A3."13>OS33
$(436!3""-=E03L3
%>17>A>/@L3
Pempherisypsilychnus =$$!""7
*-L88(733-33'$3!43'
26$3""2J,8M1JM&:3
%>17>A>/@:)7
-"3
Pempheris zajonzi F6 !%
*((7O,L/143
$('43,""11>7Q
,I.?3
%>17>A>/@13
%
* Y-=$,2313>33*3
@3+"3$$,(''$F2G3"":.>
0;3
4":.$,8

-3>60;13""
*"*9"3*5
315.3%;'6'9868
"3:6,;9"26!3+;3/
;9",3131
"3>
"33
%!3&!E88!34
3
1"*>1>/0&7>@&.$2(
)EVISTIAS L
L82!("%.
1"3
:"8!!!'L3
Evistias acutirostris /66*4!+..2
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($$""@.L3
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*@.E>34."
I3
)HISTIOPTERUS .1
%.1$''$("%
.1"""3
/3
Histiopterus typus 66*4!+..
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03',""@.L3
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-"1!6!(3
%>17>A>/@H>6:0;713>."
''3
)PARAZANCLISTIUS *"
*(*"8$!!2!"*(*""
3
/3
Parazanclistius hutchinsi *"8$!
*(*"8$!73:33-33'!2,43
""77>3:!JM1,J!M&3
%>17>A>/@H:3
)PARISTIOPTERUS .
#$2"#
"1$((4
*.$2(($#3

':8#3
0:"8',$"*:""
9*3
3
Paristiopterus gallipavo *1!"..
*:"8''0373S313@313:338'!6''$
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Paristiopterus labiosus /')*!+(2
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"13
#$203S3313O3"
":0-.9
%:$8$@1:143!!03'"
"1@313:33,,6$3
%>17>A>/@H3
)PENTACEROPSIS 1
*11%P$$!!"
%7"3
*:8!,8"%7"
3
/3
Pentaceropsis recurvirostris /6*!+.%2
%7$',>"3&3!'03
;3,6(""1"3
%>17>A>/@3
)ZANCLISTIUS L
8L82!("%7"/""
3
Zanclistius elevatus /14$1!+++2
%7"/"$$$03+313@313:3133!
!!""I0L.@1:3
%>17>A>/@3
1"0&@#&7/>@&.$,8
)PENTACEROS #
*##O$8!"*
#"""3
*.$2(2"*1"
3
0L82!("*1"
3
9L82!("*)B"
3
9$L82!("*1"
3
4:"8!'3"9$-#
"3
0-:"08!8!!"0-:"
0"3
153
Pentaceros capensis -!+"
*##O$8*3@3033!!
03'!""#)*13
%>17>A>/@H1I#0C_8823
Pentaceros decacanthus')*!+%"
*)B$,8#343!""0;
/3
9$-#8,37N&N3!!
''03(;36!""O)
(6!3
%>17>A>/@H@S9#3
Pentaceros japonicus 6*!++&
*1$$!=3.3:3:33,'
""."L3
%>17>A>/@HL@S@##3
Pentaceros quinquespinis 1!"++
*$$0E"8$$O">.33$3!!,(
F$G43""M"130;,J'M1$,J$M:2,6
3
%>17>A>/@H1")=70;/3
Pentaceros richardsoni *!+..
*1$''>3S3133'F,G03
""O"#0#)*1
3
*1$((=E3.3:3-36@3#33!
8""#*13
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(9 
3
Pentaceros wheeleri/31!"+&2
* *"8$!@3S3L3S33843""
E1&1#@0;3
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"+.+>3*>3A3133
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4"0#:.$2'
:9"
3C
3
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SCOMBROPIDAE

!"#$#%&"
'()%*(%+#)"#%
,"'-'."/$'"%,('0'
'%12344"(,%
"%
$SCOMBROPS *0+-
Scombrops *0+-52""Scombrops cheilodipteroides
60'$'7$"%
Latebrus ,2""Latebrus oculatus ,'"%
Telescopias 8+4244""Telescopias gilberti 8
+'--%
*"3%%
Scombrops boops (Houttuyn, 1782)
Labrus boops 9$$:2%9% %;%9/%""
8"%
<Scombrops cheilodipteroides 60524%6/%%=$%;%/%5
"3-08"%
Scombrops dubius 42"%-%"%% %6%+$/%>+%
#%"%%"?@+$#,05)
:#%
1A+*A6>*A?32A);,2 !'7$+$#8"B&
+%+$2&"+$#%C/)2$#$"
0D$-#$((%*#$
7$%+"(#?' %
Scombrops gilberti (Jordan and Snyder, 1901)
Telescopias gilberti 8+4244"%,%>%+%3% $%/%%
"6( 0?8"%
1A+*A6>*A?328"2$900+$$-6%
Scombrops oculatus (Poey, 1860)
Latebrus oculatus ,2"%-% %9%3%&$'/%"
&$'%
1A+*A6>*A?32;2+#>+"-(
6'$-("(%,'((
"-%&$-'-"'()%
4"#3#::E"8$'
-9>60;
35
305"
9"""3%;
">`"9;!
2"9;".9<
3!3
)CYMATOGNATHUS EL0"-
E32,!"
EL0"-"
CymatognathusaureolateralisEL0"-2
EL0"-2,,
43!',(2$8FS5'223""4
. E.@1>3
%>3
)SYMPHYSANODON .
 .$22!".$22"
""3
*)B$$!8"*)B$$"
""3
L8('("L8"
3
>60;8!3
Symphysanodon andersoni *!"(.
E82'-437%3@3
2,43!("")1.J!!38M@,J,'M&
J!$M@,J,M&-38683
%>17>A>/@>/3
Symphysanodon berryi!"(
82A313@3-343134333($3
!!$43'""/I3)>>3J',M@
2!J!!M:623
*"82(((F!G43'FO">.3(3
382@13*>3
%>17>A>/@:#
O=3
Symphysanodon disii*5 +
EE$$(436FC>L
>""3'3""CL)C713
%)C3
Symphysanodon katayamai!"(
82A313@3-343134333($
3!!!433""/IE E03L3
%>17>A>/@L3
Symphysanodon maunaloae!"(
82A313@3-343134333($
3!!,43!""/I3*>3J2M@,,J,M:
J$M@,,J$M:!3
%>17>A>/@0;IL+.>*>
10;1")a=73
Symphysanodonmona4 %
1,2436!S5@388("
"-0I073
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%:/I073
Symphysanodon octoactinus!"(
82A313@3-343134333($
3!'43,""/I3)>3(J!'M@2$J,6
(M:,6(3
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(M:3
Symphysanodon parini4 %
1,43,FS5@388("
"1")a=7,J'M182J$M:0;3
%10;16"6)a=73
Symphysanodon pitondelafournaiseiCD;F1 "
QR1=O"82'F#"3!!3
1047R
->/
%1>/I7R3
Symphysanodon rhax4 %
!1,(438S5@388("
"->3
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Symphysanodon typus 8!+((
.$2206
E. ..:9@.$226$2$
3'!3""@)3F3@R313@3
*3!$2$(03!;3G3
*)B$$73#143(!803
;33""E>3#38>18
3
L803A313@3-33,83!8('243
!6'""/I<-+.E%3*>3
*>3A31333
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+=*>3>I3
0"=.3-5323
Symphysanodon xanthopterygion8* !!
!!436S5
@38((""/IQE8J,M@2,J,M&>3
%>17>A>/@11I>3
4"#:'18(
-">60;/3%;>`T>68
;>>(6231"3#9 
3
423
)CARAIBOPS0.;1=2
0.;1=1=0.;2!"
-=.1"3
Caraibops trispinosus /6*;!"+.2
-=.18$';3L3L3>"33!3
""IO=J',M@((J!2M:!3
%>17>A>/@:#1
113
)KAPERANGUS0.;1=2
60.;1=1=0.;2,"
@"3
Kaperangus microlepis /:!"&%2
@8!,;3!%"733""I
130+33
%>17>A>/@&I))3
75:"@3
)PARASCOMBROPS.
*.$$88("*."
""3
)"1=0.;2!3
Parascombrops analis/1!"%(2
E"8,2L3L3>"33(3'6(,!3"
"/-03L3
%>17>A>/@:0;/L0#1
O
Parascombrops argyreus/'$!+"(2
')#$8203A313@3-3383'
'(03!8;3!""I*>3!'28,68$
3
%>17>A>/@-9#11O9*
>3
Parascombrops glossodon6*9;*,- !(
*1=0.;2S5'(3!!
432#$48#%!#'L6@$""0-
S@3
%>17>A>/@:0;0>9
>/-"3
Parascombrops madagascariensis6*9;*,- !(
*1=0.;2S5'(3
!(43(&2%8%!%8""/I7JM1,,J(M&
%>17>A>/@-7R3
Parascombrops mochizukii6*9;*,-3 !(
*(1=0.;*21=
0.;2S5'(3!2432&8&!&'*6>"
"/I%603
%>17>A>/@3

Parascombrops nakayamai6*9;*,- !(
*1=0.;2S5'(3!8
4324$#84!4'/672""L"-
;.E3
%>17>A>/@:0;L01IO
:43
Parascombrops ohei6*9;*,- !(
*1=0.;2S5'(3'43
#()2)8*!)'16A""L"!!J'M@
!!J!$M&33
%>17>A>/@L3
Parascombrops parvidens6*9;*,- !(
*1=0.;2S5'(3'!
432*$8)#!*'O6?!"">1
%>17>A>/@:0;0>3
Parascombrops pellucidus66!++"
*.$$88(3;3L3133,$
3!""(3-%($39"
J$M@8J,M&(,3
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3@3(""13J,M@'!J,'3,M&J(M@
'!J,M&-3,'(3
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#3
Parascombrops philippinensis/')*!++ 2
)B$$,73#143(
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:8!8(;3,41&53""
1919+1>13,!!!6'2
3
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Parascombrops serratospinosus/*!"!2
17I7I8'''3!$;3
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!J''M'N@J',M!N&03,!(,'3
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Parascombrops spinosus /6*;!". 2
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!(J!$M@2J2M@,'J2M:2'J''M:3:0;
L*#3
Parascombrops yamanouei6*9;*,-3 !(
*1=0.;*1=
0.;2S5'(354, Figs. 7M, 10F, 13M, 15S-U, 30 ""
Dong-gang, Pingtung, southwestern Taiwan).
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1$(%'1b*$23
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9'%P
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"""3
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17I"3
>6:0;9
1=0.;235E"8(3
Synagrops bellus /'8!+"72
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-3@3""@3)-5$J'MN@$(J!(MN:
3!82$3
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@#A1)-519
(83
Synagrops japonicus/B!++&2
'%P$$!'=3.3:3:33,
""."L3
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 33 883 1       & 
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343($!,6!'(3
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&7#"O36,3
4C 3:3  7343  1    %3#3  +C3  ,3  #  '(  6  03  >
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4  *3  :3  823 @    13  0      "    @
103,8'86(83
4  *3:3  8!3  #      "      0    
 3;0333
 " 333  NN  <"    0     
33A313@3-3@3365T6!$$3
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3  #          @  #3  %
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3'69(8628>6O>>3
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03#8$833'6,!3
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'3 # 3   *0 " &53
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" ;13 03 @3  1"  >"
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L%313318,3+;"%3%
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43,2!6'(,3
L%3131"L3/383;L303A313
@3-3!$868!3
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"0>3S5'223,6((3
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* 0  0            ."  
3733-3'$26!3
-"3#3,3#.;*3-
S"@3(62!3
@.>33>-3.*3*.38$(3>;I
03L-4"77#3406
!(83
@L3 133#3)  -3O3*3:3(34  :343
L:"1*.@3L365T6223
/"L3%3$$,3@0L.;3 03+3
13@313:386!,6!'',6''23
/.-33;
  #            0  
8$(0&3>"373,$6$3
/. -3 3        0 &
    1  0;              3
>"3733,8'6,'3
/.  -3  '3             "  
01>31%"3886
'3
/.  -3,3              ;
0&0;31%"3 6
23
/.  -3(3  O"      0  &
#;##1
0;31%"3226$(3
/.  -3  $3  4          ;  0
&#>>
/31%"3!'!6'$3
/. -3 83 7    ;  I  -
>@0;3L3L3>"3((3286$(3
/.-3#3"3!3765"
0.;                
0&30.33#37332'86,!3
/.-303*3-33 ""
"0&31%"3
23!6(3
/.  -3  :36L3  #    )3  13$3    0
& "" # #
#:0;3S5''2(3'6,3
/.  -3    3  4.3  3  7          ;
   -=.    1.  8$!      
0&3>"373,$!$$6!83
/.  -3    %3  )3  $3              
0>/3>"7(,6'3
42HH3H32H$626,8,63
/.-3/3)3$3; 
0 &  :> /   
3S5'!$3(683
/.  -3    *3  >3  3             
1I03>"373
3'8$6$,3
/.  -3    *3  -3  3           
  ;  0 &      -C  >
;3>"373,$,,6(3
/.-3  *3 -3 3  !   
;    0  &    7R    >
/3S5@3!',!$'6$$3
/.-3*3-3!3;
>60;   ;     0
&3>"73(3'F6G!6!3
/. -3  *3 -323      "
   0      O  1  0;
/3#"3'3'!2!6!2,3
/.  -3    @3  @."3  (3  4          
;0&    8( 
1131%"3286$!3
/.-3:L33#E3&3383%
1
0.;$;
?-1#13>"37383
HH3H32H$686(8(6$
 /3 -3 E3 &3 # L3 3 -. L3 &3 )330" 
"1090"
C3#3!($6(!
0@3O33@338$(3;
&>/3"3
-/>03/338!,!6,23
0  @3O3    3@3  3  8$(3                
7;C0&
1&0;"N #N3
3>6.@11172!68'3
05 L373 * %343  )373   *"L3&38$83 S  
3 O3 23 0 0"=  #3 # 
)303133,86,!3
0*33)3+337(3*
    ;      >      
03SE"@3,(,6,3
0"43$2,3&3313&3*3@3-'2,6(3
0.;3-323"!FG
*00"3L>""3'23
$,((6,2$3
7 L3  &3  E3 E3 3'37    ;    *
00  >      
"*##3L/1436
'3
7L3&313O3."3(30";
*       >  /        ;  
3L/1436'3
5
7 L3&   A3  13 888     >    
;  $0          1    -
>30.3-33#3733(6$3
7L3&33#3O3'34;*0
0        >  /3  L      /  1
43(6$!3
7 L3&3  3#3O3 ,3 %  "6     ;
 * 0 0   ."     
>/3L/143$6223
7*3837%3&3- P1;5
86!3``O>>>343EO.6*1.
3(386'3
1=:3:33-30.;3237)B$$2
         * .  $$8  
                  0
3S5'(362'3
1. E3 3 '          
0    #      /  >  L3
@-@1131@3
''86((3
1:3+3-33#323#"
 5         
;3#23!,6,,3
1 O3 )3 )3 %3 L3'3 1"   6 
 ;      "3   
1":@36T6(3
1O3)3-3137388,344;41
:33*.@3863
143$$!3W=E4LM>>33=3.3:3
:33,!6,3
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A taxonomic review of the genus Banjos (Perciformes: Banjosidae), previously restricted to a single species, Banjos banjos (Richardson 1846), recorded from the northwestern Pacific Ocean from the South China Sea north to Japan, as well as Lombok (Indonesia), New Caledonia and Australia, resulted in the recognition of three species, including B. banjos (northwestern Pacific Ocean, Indonesia and western Australia), Banjos aculeatus sp. nov. (eastern Australia) and Banjos peregrinus sp. nov. [northern Australia (Timor Sea)]. Records of B. banjos from New Caledonia probably also represent B. aculeatus, which is clearly distinct from other congeners in having a relatively long, strongly serrated spine at the posteroventral angle of the preopercle and an entirely dusky membrane on the spinous dorsal fin in juveniles < ca. 70 mm SL, in addition to slightly longer first and second dorsal-fin spines. Banjos peregrinus is characterized by a relatively greater head length, orbit diameter, postorbital length and pre-pelvic-fin length, as well as poorly developed serration of the exposed margin of the cleithrum. Within B. banjos, a population from the southeastern Indian Ocean, including Indonesia and western Australia, is regarded as a distinct subspecies (Banjos banjos brevispinis ssp. nov.), distinguishable from B. b. banjos from the northwestern Pacific Ocean by a relatively narrow least interorbital width, and shorter second and eighth dorsal-fin spines. Ontogenetic morphological changes within the genus and the status of the holotype of Anoplus banjos Richardson 1846 are discussed in detail.
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Bioluminescent organs have evolved many times within teleost fishes and exhibit a wide range of complexity and anatomical derivation. Although some bioluminescent organs have been studied in detail, the morphology of the bacterial light organs in glowbellies (Acropoma) is largely unknown. This study describes the anatomy of the bioluminescent organs in Haneda's Glowbelly (Acropoma hanedai) and the Glowbelly (Acropoma japonicum) and places the evolution of this light‐producing system in the context of a new phylogeny of glowbellies and their relatives. Gross and histological examination of the bioluminescent organs indicate that they are derived from perianal ectodermal tissue, likely originating from the developmental proctodeum, contrary to at least one prior suggestion that the bioluminescent organ in Acropoma is of endodermal intestinal derivation. Additionally, anterior bioluminescent organ development in both species is associated with lateral spreading of the bacteria‐containing arms of the bioluminescent organ from an initial median structure. In the context of a 16‐gene molecular phylogeny, the bioluminescent organ in Acropoma is shown to have evolved within the Acropomatidae in the ancestor of Acropoma. Further, ancestral‐states reconstruction demonstrates that the bioluminescent organs in Acropoma evolved independently from the light organs in related howellid and epigonid taxa which have esophageal or intestinally derived bioluminescent organs. Across the acropomatiforms, our reconstructions indicate that bioluminescent organs evolved independently four or five times. Based on the inferred phylogeny of the order where Acropoma and Doederleinia were separated from other traditional acropomatids, the familial taxonomy of the Acropomatidae was modified such that the previously described Malakichthyidae and Synagropidae were recognized. We also morphologically diagnose and describe the family Lateolabracidae.
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Epigonus okamotoi Fricke, 2017 was originally described on the basis of a single specimen collected from New Britain, Papua New Guinea during one of the exploratory cruises (campaign: MADEEP) in 2014 organized under the Tropical Deep-Sea Benthos program. However, there are no clear differences in the meristic and morphometric characters between the holotype of the new species and specimens of E. draco Okamoto, 2015, including two additional specimens of the species found in the ichthyological collections in the NTUM. The genetic distance (p-distance) between the two “species” at the COI locus was negligible. Accordingly, the holotype of E. okamotoi is considered to be a specimen of E. draco, and the former nominal species is reduced to a junior synonym of E. draco. In addition, we rediagnose and report new distributional records for E. atherinoides (Gilbert, 1905) and E. lifouensis Okamoto & Motomura, 2013 in the West Pacific.
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The fishes of the genus Epigonus Rafinesque, 1810 in the Western Indian Ocean (including the Red Sea and the Gulf of Aden) are reviewed. Twelve species of the genus are recognized: Epigonus angustifrons Abramov & Manilo, 1987; E. denticulatus Dieuzeide, 1950; E. elongatus Parin & Abramov, 1986; E. exodon Okamoto & Motomura, 2012; E. lenimen (Whitley, 1935); E. macrops (Brauer, 1906); E. marimonticolus Parin & Abramov, 1986; E. marisrubri Krupp, Zajonz & Khalaf, 2009; E. pectinifer Mayer, 1974; E. robustus (Barnard, 1927); E. telescopus (Risso, 1810); E. waltersensis Parin & Abramov, 1986, and two new species, E. bispinosus n. sp. and E. idai n. sp. Epigonus bispinosus and E. idai belong to the E. constanciae group, defined as having a pungent opercular spine, more than 45 pored lateral-line scales, and lacking an isolated dorsal fin spine between the first- and second dorsal fins. Epigonus bispinosus differs from other members of the group in having two small spines on the symphysis of lower jaw, a pair of ribs on the last abdominal vertebra, 10 + 15 vertebrae, 31–33 gill rakers and lacking a maxillary mustache-like process. Epigonus idai differs from other members of the group in having a pair of ribs on the last abdominal vertebra, 4–5 tiny projections present on symphysis of lower jaw, 10 + 15 vertebrae, strongly ctenoid scales, and 28–29 gill rakers, and in lacking a maxillary mustache-like process. A key to the species, photographs, diagnoses, and distributions in the Western Indian Ocean are given.
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Pempheris familia sp. nov. is described on the basis of two specimens collected from the Ogasawara Islands, Japan. The new species is most similar to Pempheris japonica Döderlein in Steindachner and Döderlein 1883, endemic to Japanese and Korean waters, in having adherent scales with strong ctenii on the lateral and ventral surfaces of the body, each scale expanded basally and distally due to central narrowing, the abdomen with a U-shaped cross-sectional outline, a large ventral fenestra between the coracoid and cleithrum, 10 dorsal-fin soft rays and 35 or 36 anal-fin soft rays. However, Pempheris familia can be distinguished from P. japonica by the following combination of characters: 84–88 pored lateral-line scales; 14 or 15 scale rows above the lateral line; 50–55 predorsal scales; 26 circumpeduncular scales; and a distinct blackish blotch on the pectoral-fin base. Pempheris familia appears to be endemic to the Ogasawara Islands.
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An ongoing review of the fishes of the basal percoid family Acropomatidae has revealed that the genus SynagropsGünther, 1887 as it is currently understood is not a natural group. Species with a serrated pelvic-fin spine are here placed in the resurrected genus Parascombrops Alcock, 1889 (type-species: Parascombrops pellucidus Alcock, 1889), and the new, monospecific genus Caraibops n. gen. (type-species: Synagrops trispinosus Mochizuki & Sano, 1984). Parascombrops is unique amongst Acropomatidae in the combination of the presence of vacant 8th interneural space, a predorsal formula /0+0/0+2/ and an epaxialis attachment type 1. Caraibops n. gen. shares none of these characters and further differs from Parascombrops by an anal-fin formula of III + 9 (vs II + 7 or III + 6), and the absence of denticles on the ectopterygoid. Parascombrops is revised and now contains a total of 13 species, including 7 new: P. analis (Katayama, 1957), P. argyreus (Gilbert & Cramer, 1897), P. glossodon n. sp., P. madagascariensis n. sp., P. mochizukii n. sp., P. nakayamai n. sp., P. ohei n. sp., P. parvidens n. sp., P. pellucidus Alcock, 1889, P. philippinensis (Günther, 1880), P. serratospinosus (Smith & Radcliffe, 1912), P. spinosus (Schultz, 1940) and P. yamanouei n. sp. Synagrops adeni Kotthaus, 1970 and S. malayanus Weber, 1913 are treated as synonyms of P. pellucidus and P. philippinensis, respectively. Lectotypes are designated for P. philippinensis and S. malayanus. The main characters used to distinguish between the species of Parascombrops are: serration of other fin spines, number of gill rakers and pseudobranchial filaments, head profile, presence or absence of ridges on the preopercle, shape of 1st anal-fin pterygiophore, dentition on vomer, palatines and ectopterygoids, orbit diameter, pectoral-fin length, maximal body depth and otolith morphology. The genus Synagrops is here confined to two species, S. japonicus (Döderlein, 1883) and S. bellus (Goode & Bean, 1896), characterized by the apomorphic character of an otic capsule with a posteriorly open myodome, a basioccipital fossa and a very specialized otolith morphology. Synagrops is also characterized by the absence of pelvic-fin spine serrations. Two other species without a serrated pelvic-fin spine, originally described in Synagrops, are removed from this genus. Synagrops microlepis Norman, 1935 is separated into the monotypic Kaperangus n. gen., the only genus in the family with two supraneurals (cf. three in all other taxa). The second, Synagrops pseudomicrolepis Schultz, 1940 is re-assigned to the genus Verilus. The geographic distribution of Parascombrops as currently composed is discussed, and is shown to be primarily of West Pacific nature, with few species in the Indian Ocean and one in the tropical West-Atlantic (P. spinosus). The West Atlantic species Parascombrops spinosus is very closely related to P. mochizukii from the tropical northwestern Pacific, and the implications of this disjunct distribution are discussed. The high degree of speciation now recognized in Parascombrops species of the West-Pacific indicates that a diverse ecological adaptation within an overall pseudoceanic habitat may have played a major role in speciation, which would have remained obscured without adequate taxonomic resolution. Fossil, otolith-based records are also briefly discussed in the context. The extant Parascombrops argyreus and P. oheiare reported from the Pliocene of Japan, and Caraibops trispinosus has been recorded from the Pliocene of Venezuela.