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Detailed biostratigraphy of the Middle Callovian - Lowest Oxfordian in the Mikhaylov reference section (Ryazan region, European part of Russia) by ammonites
Abstract
Study of the sections near the Mikhaylov (Ryazan region) has made it possible to improve the infrazonal subdivision of the Middle Callovian – Lower Oxfordian of the European Russia by ammonites. The Athleta Zone is the most complete in the studied sections. A sequence of kosmoceratid biohorizons (phaeinum, proniae, rowlstonense and kuklikum) is established here, as well as infrazonal units based on a phylogenetic sequence of species of the genus Funiferites. A study of the paleobiodiversity and frequency of occurrence of ammonites at different intervals of the section showed that the ammonite assemblages of the Athleta Zone are Sub-Boreal or Sub-Tethyan, while in the Lamberti Zone they are Boreal, and in the Lower Oxfordian they are Arctic. It is also shown that the change in ammonite composition in the sections coincides with the cycles of transgressions and regressions of Boreal basins. New species and subspecies of the ammonites Funiferites allae compressum, Cadoceras (Eichwaldiceras) intermedium, Brightia (B.) lominadzei, B. (B.) progzhellensis, B. (B.) eccentrtica, B. (Glyptia) canaliculata stankevitchae, Zieteniceras rarecostatum are described.
... Стрелками обозначены направления филогенеза. Сокращения: h -голотип; lt -лектотип, pt -паратип (двойной запятой отмечены типовые экземпляры с реконструированными значениями признаков по регрессионным формулам); ОВ -очень высокие обороты (из [Kiselev, Rogov, 2018] В филогенезе латеральное уплощение раковины сопровождается коррелятивно связанным уменьшением диаметра умбилика (рис. 1.6.28б). ...
... ЯрГПУ № М7-54; разрез «Михайловцемент-I», слой 7 (нижняя часть), подзона Grossouvrei, Д=33,1 мм; C. (E.) carinatum (Eichwald) -экз. ЯрГПУ D1-70, Разрез «Дубки» (Саратовская область), слой 1 (+30 см), биогоризонт henrici, Д = 34 мм (из [Kiselev, Rogov, 2018]). ...
... В результате исследования классических и новых разрезов, преимущественно в Центральной России и Респ. Коми, предложены новые варианты стратиграфического деления для нижнего келловея (Митта, Стародубцева, 1998;Митта, 2000б;Гуляев, 1999Gulyaev et al., 2002;Киселев, Рогов, 2007а, б), среднего-верхнего келловея и нижнего оксфорда (Киселев, 1999, 2001a2006;Киселев и др., 2003;Митта, 2003;Киселев, Меледина, 2004;Kiselev et al., 2013;Kiselev, Rogov, 2018). ...
The monograph is devoted to ammonites and infrazonal stratigraphy of the Bathonian and Callovian stages of European Russia, Ukraine and adjacent regions of the North Caucasus, as well as high-latitude regions of the Northern Hemisphere (Northern Siberia, Franz Josef Land, Southern Alaska, British Columbia, Eastern Greenland). The work affects upon the fundamental problems of infrazonal biostratigraphy and substantiates the use of infrazonal subdivisions (biohorizons) as minimal biostratigraphic units. Fundamentally new data on the evolution of ammonites in the upper part of the Middle Jurassic have been obtained. The features of evolutionary transformations in several phylogenetic lineages of cardioceratids, the ammonite family, key for the division and correlation of the Bathonian, Callovian, and Lower Oxfordian, have been revealed. As a result of the research, an important scientific problem has been solved: the regularities of the stratigraphic and biogeographic distribution of ammonites in the Bathonian and Callovian deposits of the boreal and highly boreal regions of the Northern Hemisphere have been established. The author has developed infrazonal ammonite scales for the European part of Russia and central Ukraine, as well as for key areas of the Bathonian and Callovian deposits in the Arctic, such as Northern Siberia, Southern Alaska and British Columbia. On the basis of the data obtained, a scheme for the detailed intraboreal correlation of the Bathonian and Callovian infrazonal scales and a new version of the Boreal ammonite standard has been developed. The theoretical part of the work includes the presentation of the concept of infrazonal stratigraphy, the principles of constructing the Boreal standard, methods of phylogenetic reconstructions in ammonites, the concept of morphs of intraspecific variability, conceptual foundations of the study of paleobiogeographic differentiation and invasions of extinct organisms.
... The aptychus described in this article was found in 2017 in the "Mikhailovcement" quarry in the Ryazan region, Central Russia (54.212885°N, 38.959170°E; see map in Kiselev and Rogov, 2018, fig. 1). ...
... 1). Middle to upper Callovian and lower Oxfordian beds, exposed in this quarry, were recently described in detail (Kiselev and Rogov, 2018). The aptychus was found by accident in a fragment of the body chamber (Fig. 2) of a partially flattened Peltoceras cf. ...
... The aptychus was found by accident in a fragment of the body chamber (Fig. 2) of a partially flattened Peltoceras cf. retrospinatum macroconch (Fig. 2) in a concretion of yellowish-grey, oolitic marlstone in the upper Callovian Lamberti Zone (bed 10, according to Kiselev and Rogov, 2018). Peltoceras retrospinatum (Gérard and Contaut) was originally described from the terminal horizon of the Athleta Zone, but at the Mikhailovcement locality it also was reported from the basal part of the next ammonite zone, the Lamberti Zone (see Kiselev and Rogov, 2018, p. 96). ...
... Autoren: Callomon, Dietl & Page (1988); Page (1988,1989) Index-Fossil: Kepplerites (Gowericeras) gowerianus (J. de C. Sowerby, 1827) Definierte Basis: wie Koenigi-Zone (= metorchus-Horizont) Callomon et al. (1988) (Callomon et al. 1989;Dietl 2010 (Kiselev & Rogov 2018) und in Uzhovka (Oblast Nischni Nowgorod) (Mitta 1999). ...
Kurzfassung
Diese Arbeit beschreibt Ammoniten der Gattung Kepplerites aus der Herveyi-und der Koenigi-Zone (Unter-Callovium, Mittel-Jura) von Nord-und Süddeutschland. Die meisten Stücke stammen aus dem Macrocephalenoolith der Grube Porta (Wiehengebirge, Westfalen) und aus dem Raum Hannover-Hildesheim. Es sind 11 Arten der Untergattung Gowericeras mit entsprechenden Mikrokonchen der Untergattungen Toricellites und Gulielmina. Vier Arten werden neu beschrieben: Kepplerites (Gowericeras) halleyi n. sp., K. (G.) corpernici n. sp., K. (Toricellites) besseli n. sp. und Sigaloceras (Gulielmina) fabri-cius n. sp. Alle Keppleriten sind hervorragende Leitfossilien, mit deren Hilfe die Biost-ratigraphie des Unter-Calloviums (Mittel-Jura) wesentlich verbessert wird. So wird die Abfolge der Ammoniten-Faunenhorizonte ergänzt und berichtigt und die Korrelation mit anderen Gebieten wie Frankreich, England oder der Russischen Plattform überarbeitet. Es entsteht ein detailliertes Bild der Verbreitung der Gattung Kepplerites in Raum und Zeit. Diese Daten bilden die Grundlage zur Rekonstruktion der Biogeographie und der Evolution der Keppleritinae.
Summary
This study describes ammonites of the genus Kepplerites of the Herveyi and Koenigi zones (Lower Callovian, Middle Jurassic) from Northern and Southern Germany. Most of the specimens come from the Macrocephalenoolith of the Porta mine (Westphalia), from the Hanover-Hildesheim area. There are 10 species of the subgenus Gowericeras with the corresponding microconches of the subgenus Toricellites. Four are newly described: Kepplerites halleyi n. sp., K. corpernici n. sp., K. (Toricellites) besseli n. sp. and Sigaloceras (Gulielmina) fabricus n. sp. All keppleritids are excellent guide fossils, with which the biostratigraphy of the Lower Callovian (Middle Jurassic) is significantly improved. The sequence of the ammonite faunal horizons is completed and corrected, and their correlation with other areas such as France, England or the Russian platform is revised. This creates a detailed picture of the distribution of the genus Kepplerites in space and time. The data form the basis for the reconstruction of the biogeography and the evolution of the Keppleritinae.
... Positive δ 18 O shifts, which mostly coincide with transgressions and the periods of prevalence of (Sub)boreal cephalopod faunas, could have resulted from decreased water temperatures (cf. Sahagian et al., 1996;Kiselev and Rogov, 2018). Palaeotemperatures calculated for these intervals using a constant seawater δ 18 O value of -1‰ VSMOW characteristic of non-glacial periods are, however, very low (Fig. 1). ...
... The stratigraphic significance of ostracods was subsequently studied and reflected in articles and some conference abstracts based on the materials for the Upper and Middle Jurassic of Saratov (Kiselev et al., 2006;Tesakova and Sel'tser, 2013;Tesakova et al., 2017a;, Kostroma (Tesakova et al., 2012), Ryazan , Moscow (Tesakova, 2017a), and Kursk (Tesakova, 2013c) regions, Central Poland , Southern Germany (Franz et al., 2009), and Northern Switzerland (Tesakova, 2017c). The zonal scale based on the Middle and Upper Jurassic ostracods of the RP correlating to the ammonite zones (Tesakova, 2012) and biohorizons (Tesakova, 2014a(Tesakova, , 2015Tesakova et al., 2017aTesakova et al., , 2017b is currently the most detailed of the micropaleontological RP scales, second to the ammonite standard only (Rogov, 2017;Kiselev and Rogov, 2018). The first version of this scale published in the Unified Regional Scheme of the RP Jurassic (Tesakova, 2012) was considerably reduced at the insistence of the editor-inchief; therefore, it has lost its practical significance and is not recommended for use. ...
A new forma type of palaeopathy, Forma malaspina (growth of additional double spines instead of the regular pairing with no exogenous injury) in Peltoceras schroederi Prieser [M] recorded from a condensed section (upper Callovian-lower Oxfordian) from the Ogrodzieniec Quarry (southern Poland) is reported and attributed to genetic abnormality. Microtomographical analysis of P. schroederi did not reveal any internal structures or density changes that would otherwise be suggestive of parasitism. Previous mentions by other authors of a similar duplicated spine palaeopathy in P. schroederi [M] from the Mediterranean region is also suggestive of a genetic abnormality rather than parasitism. The other palaeopathy recorded includes a transitional example of Forma juxtasulcata Géczy ‒ juxtacarinata Hölder, where the ventral region is marked by a row of single ventro-lateral tubercles rather than the usual two rows in the upper Callovian Peltoceras athleta [M] recorded from Arçay, near Loudun (western France). This palaeopathy is attributed to parasitism and also does not show any signs of exogenous injuries.
A new ammonite assemblage from the lower beds of the Ogrodzieniec Quarry (southern Poland), the only Callovian section in the middle part of the Polish Jura Chain, is described. It includes the presence of Kosmoceras rotundum (Quenstedt), followed by the first example of co-occurring micro- and macroconchs in the genus Rollierites (R. biplicatum sp. n.) and above it, the association of Euaspidoceras sp. and Peltoceratoides (Parawedekindia) gerberi Prieser. Both R. biplicatum sp. n. (m and M; microconch and macroconch) and K. rotundum are assigned to the late Callovian Lamberti Zone. P. (P.) gerberi characterizes the early Oxfordian Cordatum Zone. This is the first record of the genus Rollierites from Poland. This study extends the upper age limit of the middle Callovian Rollierites up to the late Callovian Lamberti Zone. On the basis of morphological and stratigraphical data, it is tentatively proposed that the origin of the early–middle Oxfordian Tornquistes may be in the middle–late Callovian Rollierites, rather than the previously proposed late Callovian Pachyceras. However, this is speculation, as the present data set is insufficient
The Boreal (Arctic) ammonite standard of the Jurassic system is
necessary for the effective correlation and dating of any regional Jurassic
biostratons across the Arctic region. Its development is based on principles
of continuity, complexity, completeness, as well as palaeobiochorological,
taxonomic, correlation and phylogenetic principles. The new version of the
ammonite Boreal Standard for the Bathonian and Callovian was developed
according to these principles and has a composite (synthetic) structure. It
contains a zonal and infrazonal scale, and consists of units based mainly on
ammonites of the family Cardioceratidae. It includes 14 zones and 38
biohorizons that represent fragments of the regional standard scales of East
Greenland, Northern Siberia and European Russia. This new version of the
Boreal Standard also includes 19 levels of interregional correlation for the
Panboreal Superrealm and 28 levels for the Arctic Realm.
One proposed genus name, Paralcidia Spath, 1928 in Ammonoidea is nomenclaturally invalid, as the genus group name has already been used by a different author in Lepidoptera. In accordance with Article 60 of the International Code of Zoological Nomenclature, I propose a substitute name, Alcidia Rollier, 1913, for this genus name.
The outcrops at Votcha village is the most indicative Callovian section of the Sysola river basin, which after works I. Е. Khudyaev is not examined more than 70 years. Reexamination of the section has allowed to reveal a sequence from 17 ammonite assemblages, in an interval of standard West-European Zones of Lower Callovian Koenigi – Middle Callovian Coronatum. 4 standard Zones and 4 Subzones are established, 5 new biohorizons Tcheffkini, Milaschevici khudyaevi, Milaschevici milaschevic, Praestenolobum, Stenolobumi are allocated. Among ammonites in the majority of assemblages the species, characteristic for Callovian as Central Russia, Western Europe and, simultaneously, for the Arctic basin are found, that gives additional opportunities for correlation of a boreal and subboreal scales. The new subspecies Cadoceras (Rondiceras) milaschevici khudyaevi is described.
Results of clumped isotope, oxygen isotope and elemental (Mg/Ca, Sr/Ca) analyses of exceptionally well-preserved belemnite rostra and ammonite shells from the uppermost Callovian–Upper Kimmeridgian (Lamberti–Mutabilis zones) of the Russian Platform are presented. Despite a significant decrease in belemnite δ¹⁸O values across the Upper Oxfordian–Lower Kimmeridgian, the clumped isotope data show a constant seawater temperature (ca. 16 °C) in the studied interval. The decrease in belemnite δ¹⁸O values and lower δ¹⁸O values measured from ammonite shells are interpreted as a result of the salinity decline of the Middle Russian Sea of ca. 12‰ and salinity stratification of the water column, respectively. The postulated secular palaeoenvironmental changes are linked to the inflow of subtropical, saline waters from the Tethys Ocean during a sea-level highstand at the Middle–Late Jurassic transition, and progressive isolation and freshening of the Middle Russian Sea during the Late Oxfordian–Kimmeridgian. The obtained clumped isotope data demonstrate relative stability of the Late Jurassic climate and a paramount effect of local palaeoceanographic conditions on carbonate δ¹⁸O record of shallow epeiric seas belonging to the Subboreal Province. Variations in Mg/Ca and Sr/Ca ratios of cylindroteuthid belemnite rostra, which are regarded by some authors as temperature proxies, are, in turn, interpreted to be primarily dependent on global changes in seawater chemistry. The paleoenvironmental variations deduced from clumped and oxygen isotope records of the Russian Platform correspond well with changes in local cephalopod and microfossil faunas, which show increasing provincialism during the Late Oxfordian and the Early Kimmeridgian. Based on the review of literature data it is suggested that the observed salinity decrease and restriction of Subboreal basins during the Late Jurassic played a major role in the formation of periodic bottom water anoxia and sedimentation of organic rich facies.
An accurate chronostratigraphy of the timing and magnitude of global sea-level trends and their short-term variations is an indispensable tool in high-resolution correlations, exploration, and paleoenvironmental and geodynamic models. This paper is a reappraisal of the Jurassic sealevel history in view of recent updates in time scales and a large body of new chronostratigraphic data accrued since 1998, when the last such synthesis was presented. A review of the Jurassic sea-level history has also been keenly awaited by explorationists given that the Jurassic continues to be a major exploration target for the industry. As in previous eustatic models of this period, the updated Jurassic sealevel curve remains largely Eurocentric due to the limitations imposed by biostratigraphic correlation criteria (provinciality of ammonite and microfossil zones), though it can now be extended to some parts of the Tethys toward the east. The updated long-term curve indicates that there was a general rise of sea level through the Jurassic that began close to a level similar to or below the present-day mean sea level (pdmsl) in the early Jurassic, culminating in the peak high in the late Kimmeridgian-early Tithonian interval, before stabilizing in the earliest Cretaceous at ∼110 m above pdmsl. Within this long-term trend are relative secondorder highs in the Toarcian and Aalenian, and at Bathonian-Callovian and Kimmeridgian-Oxfordian boundaries. Superimposed are 64 third- and fourthorder fluctuations of which 15 are considered major with base-level falls of more than 75 m, although precise amplitudes of drawdowns are often difficult to establish. Higher resolution fourth-order cyclicity (∼410 k.y.) is also observable in many Jurassic sections whenever sedimentation rates were high. Causes for the third-order cyclicity, in the absence of major ice sheets in the Jurassic, remains enigmatic.
Review of zonal and infrazonal subdivision of the Kimmeridgian and Volgian stage of southern part of the Moscow Syneclise, accompanied by the detailed description of key sections and all stratigraphic units (zones, subzones and biohorizons) of the region, is provided. New biohorizons are introduced: crenulatus (Lower Kimmeridgian), tenuicostatum, arkelli (Lower Volgian), gerassimovi, virgatus, rarecostatus (Middle Volgian), praeokensis, interjectum, transitionis (Upper Volgian), and characteristics of biohorizons mentioned previously by the author but never described in details (evolutus, tenuicostatus, cheremkhensis, subfulgens, involutus, catenulatum, nodiger, milkovensis horizons) is provided. Dimorphism and polymorphism patterns in Kimmeridgian and Volgian ammonites are discussed. Updated diagnoses are provided for Virgatitidae, Garniericeratinae and Craspedites (Trautscholdiceras). New species, which are used as indexes of ammonite horizons, are described: Virgatites rarecostatus sp. nov., Craspedites (C.) praeokenis sp. nov., C. (Trautscholdiceras) transitionis sp. nov., Kachpurites evolutus sp. nov., K. tenuicostatus sp. nov. and K. involutus sp. nov.. Evolution of Late Volgian craspeditids is reviewed. Two eudemic subfamilies which evolved in the Late Volgian in the Middle Russian Sea (Craspeditinae and Garniericeratinae) were characterized by significantly different evolutionary rates through time: at the beginning of Late Volgian time evolutionary rates in garniericeratins was very high, while during the Nodiger Chron craspeditins evolved much quicker than garniericeratins. Although early members of two lineages strongly resemble each other, major trends in evolution of these subfamilies were different. At the beginning of the Late Volgian garniericeratins predominated in ammonoid assemblages, and their shells were quickly evolving from platycones to oxycones; this was accompanied by gradual reduction of variability and loss of sculpture. In the both discussed lineages intraspecific variability and evolutionary rates were positively correlated with relative abundance within ammonite assemblages. However, intraspecific variability of some species was especially high (Kachpurites tenuicostatus) or very low (Garniericeras interjectum). Speciation in Craspeditinae led to the appearance of cadicones, and these changes in shape were accompanied by gradual strengthening of the sculpture (with appearance of strong nodes in C. (Trautscholdiceras)). At the beginning of the Late Volgian craspeditins were relatively uncommon (5-10%), but became predominate during the Nodiger Chron (90-98% of all ammonite records). Changes of shell size through the evolution of garniericeratins were irregular, while craspeditins showed strong trend to decrease of both median and maximal sizes through their evolution. Late Volgian ammonites, belonging to eudemic craspeditid lineages inhabiting the Middle Russian Sea, became extinct in the latest Late Volgian. During the latest Chron of the Late Volgian (Singularis Chron) ammonoid assemblages of the Middle Russian Sea were represented by the immigrant taxa only (Volgidiscus and Shulginites).