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Gardens' Bulletin Singapore 70 (2): 307–395. 2018
doi: 10.26492/gbs70(2).2018-08 307
The plant taxa of H.N. Ridley, 5. The Gentianales
I.M. Turner1, 2, Y.W. Low2, M. Rodda2, K.M. Wong2 & D.J. Middleton2
1Singapore Botanical Liaison Ofcer,
Royal Botanic Gardens Kew,
Richmond, Surrey TW9 3AE, U.K.
i.turner@kew.org
2Singapore Botanic Gardens, National Parks Board,
1 Cluny Road, 259569, Singapore
ABSTRACT. Henry Nicholas Ridley (1855–1956) described more than 450 taxa of Gentianales,
mostly from Southeast Asia, and was the author of over 70 additional combinations in this
order. A listing of these names with place of publication and the types of the Ridley basionyms
is given here. Many names are lectotypied. The new combinations Chassalia pauciora
(Ridl.) I.M.Turner and Chassalia violascens (Ridl.) I.M.Turner are proposed for Cephaelis
pauciora Ridl. and Gartnera violascens Ridl. respecitively. Three names are newly reduced
to synonymy: Cephaelis elongata Ridl. to Chassalia bracteata Ridl., Geophila matthewii Ridl.
to Mussaenda uniora Wall. ex G.Don and Psychotria setistipula Ridl. to Ixora patens Ridl.
Keywords. Apocynaceae, Gentianaceae, lectotype, Loganiaceae, Rubiaceae, typication
Introduction
This paper continues an intermittent series on the plant taxa named by Henry Nicholas
Ridley (1855–1956). Earlier parts have covered the pteridophytes (Turner & Chin,
1998a), gymnosperms (Turner & Chin, 1998b), Zingiberales (Turner, 2000) and basal
angiosperms (Austrobaileyales, Canellales, Chloranthales, Laurales, Magnoliales,
Nymphaeales and Piperales) (Turner, 2012). In this installment focus shifts to the
Gentianales.
Ridley described many plant species, particularly from Southeast Asia. He
documented the botanical diversity of the region tirelessly and seemed prepared to
tackle all and any taxonomic group of vascular plants that came his way. Many of
his publications involve descriptions, generally including many novelties, of taxa
collected by himself or others from particular localities.
The Gentianales, as currently circumscribed (Angiosperm Phylogeny Group,
2016), include ve families: Apocynaceae (now including the Asclepiadaceae),
Gelsemiaceae (including the Malesian genus Pteleocarpa Oliv.), Gentianaceae,
Loganiaceae and Rubiaceae. The Apocynaceae and Rubiaceae are highly diverse in
Southeast Asia, so it is no surprise that Ridley described many species in these families.
In the later years of his very long career, the Rubiaceae became a particular focus of
his attention (Ridley, 1927, 1932, 1934, 1939). This work included the description
of several new genera (e.g. Asemanthia Ridl., Porterandia Ridl., Zeuxanthe Ridl.)
and outline revisions of Argostemma Wall., Porterandia Ridl., Rennellia Korth. and
Urophyllum Wall.
308 Gard. Bull. Singapore 70 (2) 2018
An enumeration of Ridley’s Gentianales taxa
The taxa that Ridley described or provided with new combinations or new names as
governed by the International Code of Nomenclature (Turland et al., 2018) are listed.
Those taxa Ridley described as new are listed against Arabic numbers, and purely
nomenclatural novelties are itemised with Roman numerals. Ridley’s combination
starts each entry and is given in bold and italics if it is still an accepted name or in italics
alone if it is now treated as a synonym. Homotypic synonyms, including basionym
where relevant, are given. If necessary, a currently accepted name is included in bold
and italics, either included in the list of synonyms if homotypic to Ridley’s taxon or
below, preceeded by ‘=’, if heterotypic. In those cases where the current identity of the
taxon is uncertain, including if the sheets have been redeterminded but that data not
published, we continue to recognise in bold and italics the name for which it is a type
but we acknowledge that in several cases this will be changed by ongoing research.
Exceptions are made where either no original material is available, or for infraspecic
taxa in difcult genera such as Argostemma. The place of publication of all names is
given and reference to important revisions and monographs are included. These were
also studied for information on earlier typications of Ridley’s names.
For each of the taxa Ridley described, the types are listed. The holotype or
lectotype is listed rst and then as many syntypes as have been located. We have
seen most of the specimens listed, but some records come from published sources,
information from correspondents or on-line databases. There are doubtless duplicates
in other herbaria and there remain a few cases where no type material has yet been
found at all. Locality and/or date information, sometimes even collector information, is
included in square brackets when the data are not repeated on all presumed duplicates
of a collection or is only given in the protologue but not on the specimens at all.
Barcodes are given, in square brackets, where available but for some type material
the specimen either has no barcode or we have not seen the specimen again if it has
more recently been barcoded. Specimen data from the protologue for which we have
found no material is included but listed as ‘(not traced)’. Where specimen information
is given in a protologue only via a collection locality and we have found no material
that might match, this is listed as ‘Unknown collector (not traced)’. A high proportion
of Ridley’s taxa require lectotypication because he rarely designated types from
among the various specimens that he cited when publishing new taxa. Specimen
citation by Ridley was, by contemporary standards, poor – he often omitted collection
numbers, very rarely noted collection dates and generally failed to state in which
herbaria material was deposited. He was also inconsistent in annotating the specimens
that he saw. Typication is therefore frequently problematic. Ridley’s career can be
divided into two main periods – the Singapore years (1888–1912) and the retirement
to Kew (1913 onwards). This information is important in making decisions relating
to typication of Ridley taxa, but experience has shown that many types one might
expect, from the date of publication, to be in Singapore are found at Kew.
In selecting lectotypes, we have generally chosen the specimen that we
considered to be of the best quality among the candidate material. To avoid much
309
The Gentinales taxa of H.N. Ridley
repetition, comments on typication in the list are generally limited to cases where the
choice of type was less straightforward.
There are specimens where a specic day and month is given for the collection
but only a year range. For example for Ixora uminalis Ridl. where the date of
collection is given as ‘22 August 1865–1866’, which is not logical but is left as written
on the label. For some localities, the current name is given in curly brackets ({}) when
it differs from what is written on the label.
Apocynaceae
1. Hoyella Ridl., J. Fed. Malay States Mus. 8(4): 62 (1917). – TYPE: Hoyella rosea
Ridl.
= Dischidia R.Br., Prodr. 461 (1810).
2. Spiladocorys Ridl., Trans. Linn. Soc. London, Bot. 3(9): 322 (1893). – TYPE:
Spiladocorys angustifolia Ridl.
= Pentasachme Wall. ex Wight, Contr. Bot. India 60 (1834).
3. Alstonia micrantha Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 95 (1918). – TYPE:
Peninsular Malaysia, Selangor, Rantau Panjang, 27 July 1914, C.B. Kloss 3 (lectotype
K [K000195837], designated by Sidiyasa (1998), or possibly holotype).
= Chilocarpus denudatus Blume, Bijdr. Fl. Ned. Ind. 16: 1025 (1826); Middleton, Fl.
Males., Ser. 1, 18: 171 (2007).
4. Alyxia angustifolia Ridl., J. Fed. Malay States Mus. 6: 161 (1915); Middleton,
Blumea 49: 29 (2000); Middleton, Fl. Males., Ser. 1, 18: 74 (2007). – TYPE:
Peninsular Malaysia, Pahang, Gunong Tahan, July 1911, H.N. Ridley 16060 (lectotype
K [K000894016], designated by Middleton (2000); isolectotypes A [A00065145], BM
[BM000508431], SING [SING0055213]).
5. Alyxia oleifolia King & Gamble var. tenuifolia Ridl., Fl. Malay Penins. 2: 333 (1923).
– TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang, 5000 ft, 20 January
1911, H.C. Robinson s.n. (lectotype K [K000894007], designated by Middleton
(2000); isolectotypes BM [BM000508433], SING [SING0055208]).
= Alyxia reinwardtii Blume, Cat. Gew. Buitenzorg (Blume) 43 (1823); Middleton,
Blumea 45: 101 (2000); Middleton, Fl. Males., Ser. 1, 18: 112 (2007).
310 Gard. Bull. Singapore 70 (2) 2018
6. Chilocarpus aurantiacus Ridl., Bull. Misc. Inform. Kew 1926: 73 (1926). – TYPE:
Indonesia, Sumatra, Siberut, 18 September 1924, C.B. Kloss 14500 (lectotype K
[K000195833], designated by Middleton (2007); isolectotype SING [SING0015891]).
= Chilocarpus costatus Miq., Fl. Ned. Ind. 2: 393 (1856); Leeuwenberg, Syst. Geogr.
Pl. 73: 133 (2002); Middleton, Fl. Males., Ser. 1, 18: 168 (2007).
7. Chonemorpha penangensis Ridl., Agric. Bull. Straits & Fed. Mal. States 10: 147
(1911). – TYPE: Peninsular Malaysia, Penang, May 1893, C. Curtis 832 [Forest Guards
leg.] (lectotype SING [SING0040281], designated by Middleton (2006); isolectotypes
SING [SING0058317, SING0040282]). Additional syntypes: Peninsular Malaysia,
Penang, road to Balik Pulau, June 1898, H.N. Ridley 9441 (BM [BM000047376]);
Perak, Gopeng, King’s Collector 6000 (not traced); Perak, Larut, King’s Collector 3636
(not traced); Perak, Temengoh, July 1909, H.N. Ridley 14271 (BM [BM000047377]);
Malacca, Brisu, Derry 543 (not traced).
= Chonemorpha fragrans (Moon) Alston, Ann. Roy. Bot. Gard. (Peradeniya) 11: 203
(1929); Middleton, Fl. Males., Ser. 1, 18: 179 (2007).
8. Chonemorpha rheedei Ridl., Agric. Bull. Straits & Fed. Mal. States 10: 146 (1911).
– TYPE: Singapore, Singapore Botanic Gardens, big climber in Plant House, 25 July
1910, Anon. s.n. (lectotype K [K000895071], designated here).
= Chonemorpha fragrans (Moon) Alston, Ann. Roy. Bot. Gard. (Peradeniya) 11: 203
(1929); Middleton, Fl. Males. Ser. 1,18: 179 (2007).
9. Cynanchum seimundii Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 300 (1922). –
TYPE: Peninsular Malaysia, Pahang, Kwala Tahan, November 1920, E. Seimund s.n.
(lectotype K [K000894968], designated by Liede (1999), or possibly holotype).
This is neither Cynanchum L. nor Gongronema (Endl.) Decne.; more likely
Cosmostigma racemosum Wight.
10. Dischidia ericiora Ridl., J. Fed. Malay States Mus. 10: 146 (1920), as
‘ericaeora’, nom. illeg., non Beccari (1886). – TYPE: Peninsular Malaysia, Kedah-
Perak boundary, Gunong Bintang, June 1917, F.M.S. Mus. Native Collectors s.n.
(lectotype K [K0000911044], designated by Rintz (1980), or possibly holotype).
= Dischidia subalata Warb., Repert. Spec. Nov. Regni Veg. 3: 345 (1907); Rintz,
Blumea 26: 96 (1980) (as ‘subulata’).
11. Dischidia fruticulosa Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 96 (1918); Rintz,
Blumea 26: 101 (1980). – TYPE: Peninsular Malayia, Perak, Gunong Kerbau at 4200
311
The Gentinales taxa of H.N. Ridley
ft, s.dat., H.C. Robinson s.n. (lectotype K [K001096329], designated by Rintz (1980),
or possibly holotype).
12. Dischidia lagenifera Ridl., J. Fed. Malay States Mus. 8(4): 63 (1917). – TYPE:
Sumatra, Ridley s.n. (not traced)
= Dischidia albida Griff., Notul. Pl. Asiat. 4: 46 (1854); Rintz, Blumea 26: 100 (1980).
Rintz (1980) referred to the type as being in K but we have not managed to nd it.
13. Dischidia lancifolia Ridl., J. Fed. Malay States Mus. 10: 102 (1920), nom. illeg.,
non Merrill (1918). – Dischidia klossii Ridl. ex Kerr, Fl. Siam. 3(1): 46 (1951). –
Dischidia acutifolia Maingay ex Hook.f. subsp. klossii (Ridl. ex Kerr) Rintz,
Blumea 26: 101 (1980). – TYPE: Thailand, Mamoh, C.B. Kloss 6721 (lectotype K
[K001096331], designated by Rintz (1980)). Additional syntype: Thailand, Tapli, C.B.
Kloss 6805 (K [K001096332]).
Ridley obviously confused Kloss’s numbers referring to 674 rather than 6721.
14. Dischidia longepedunculata Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 32 (1912),
as ‘longe-pedunculata’; Rintz, Blumea 26: 97 (1980). – TYPE: Peninsular Malaysia,
Selangor, Sempang Mines, April 1911, H.N. Ridley 15927 (lectotype K [K000911041],
designated by Rintz (1980); isolectotype SING [SING0059495]).
15. Dischidia parvifolia Ridl., J. Fed. Malay States Mus. 5: 41 (1914); Rintz, Blumea
26: 95 (1980). – TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang, January
1913, H.C. Robinson s.n. (lectotype K [K000911046], designated here).
16. Dischidia polyphylla Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 78 (1923).
– TYPE: Sumatra, Berastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000911018], designated here, or possibly holotype).
17. Dischidia pubescens Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 71 (1911). –
TYPE: Peninsular Malaysia, Temengoh Woods, July 1909, H.N. Ridley s.n. (lectotype
SING [SING0015217], designated here, or possibly holotype).
18. Dischidia rosea Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 31 (1912), nom. illeg.,
non Schlechter (1906). – Dischidia rhodantha Ridl., J. Straits Branch Roy. Asiat. Soc.
312 Gard. Bull. Singapore 70 (2) 2018
79: 97 (1918); Rintz, Blumea 26: 93 (1980). – TYPE: Peninsular Malaysia, Selangor,
Sempang Mines, April 1911, H.N. Ridley 15900 (lectotype SING [SING0059494],
designated by Rintz (1980); isolectotype K [K000911049]).
19. Dischidia singaporensis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 31 (1912);
Rintz, Blumea 26: 92 (1980). – TYPE: Singapore, Changi, Changi Police Station,
1908, H.N. Ridley s.n. (lectotype SING [SING0044741], designated by Rintz (1980)).
Additional syntype: Singapore, Bukit Mandai on a high tree, 30 May 1889, H.N. Ridley
s.n. (SING [SING0003926]) [syntype = Hoya lacunosa Blume].
20. Dischidia tomentella Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 137 (1911);
Rintz, Blumea 26: 99 (1980). – TYPE: Peninsular Malaysia, Perlis, Bukit Lagi, March
1910, H.N. Ridley 15225 (lectotype K [K000911038], designated by Rintz (1980);
isolectotype SING [SING0059492]).
21. Dischidia viridiora Ridl., J. Fed. Malay States Mus. 10: 146 (1920). – TYPE:
Peninsular Malaysia, Kelantan, Chaning, 31 January 1917, H.N. Ridley s.n. (lectotype
K [K000911048], designated here).
= Dischidia punctata (Blume) Decne. in DC., Prodr. 8: 631 (1844); Rintz, Blumea 26:
93 (1980).
22. Ervatamia coriacea Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 29 (1912). –
TYPE: Peninsular Malaysia, Selangor, Sempang, April 1911, H.N. Ridley 15739
(lectotype SING [SING0059561], designated by Leeuwenberg (1991); isolectotypes
BM [BM001014002], CAL, K [K000857392]).
= Tabernaemontana polyneura (King & Gamble) D.J.Middleton, Taxon 55: 503
(2006); Middleton, Fl. Males., Ser. 1, 18: 386 (2007).
23. Ervatamia jasminiora Ridl., Fl. Malay Penins. 2: 342 (1923). – TYPE: Peninsular
Malaysia, Pahang, Kuantan, Baloh Reserve, 13 March 1920, Forest Guard Yeop For.
Dept. F.M.S. Field Number 836 (lectotype SING [SING0056555], designated by
Leeuwenberg (1991); isolectotypes K [K000857398], KEP).
= Tabernaemontana corymbosa Roxb. ex Wall., Edwards’s Bot. Reg. 15: sub t. 1273
(1829); Leeuwenberg, Revis. Tabernaemontana, Old World Sp. 139 (1991); Middleton,
Fl. Males., Ser. 1, 18: 376 (2007).
24. Ervatamia pauciora Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 299 (1922);
Ridley, Fl. Malay Penins. 2: 342 (1923). – TYPE: Peninsular Malaysia, Selangor-
313
The Gentinales taxa of H.N. Ridley
Pahang Boundary, Ginting Simpah, March 1917, H.N. Ridley, H.C. Robinson & C.B.
Kloss s.n. (lectotype SING, designated by Leeuwenberg (1988); isolectotypes BM
[BM001014004], K [K000857403, K000857404]).
= Tabernaemontana corymbosa Roxb. ex Wall., Edwards’s Bot. Reg. 15: sub t. 1273
(1829); Leeuwenberg, Revis. Tabernaemontana, Old World Sp. 138 (1991); Middleton,
Fl. Males., Ser. 1, 18: 376 (2007).
Despite Leeuwenberg’s effective lectotypication of the SING specimen (corrected
from ‘holotype’) this specimen cannot be located in SING.
25. Ervatamia pauciora var. minor Ridl., J. Straits Branch Roy. Asiat. Soc. 86:
300 (1922); Ridley, Fl. Malay Penins. 2: 343 (1923). – TYPE: Peninsular Malaysia,
Negri Sembilan, Bukit Tangga ar 2400 ft, 20 December 1920, H.N. Ridley 13033
(lectotype SING [SING0056562], designated by Leeuwenberg (1988); isolectotypes
K [K000857399, K000857400], SING [SING0056561]).
= Tabernaemontana corymbosa Roxb. ex Wall., Edwards’s Bot. Reg. 15: sub t. 1273
(1829); Leeuwenberg, Revis. Tabernaemontana, Old World Sp. 138 (1991); Middleton,
Fl. Males., Ser. 1, 18: 376 (2007).
26. Gongylosperma lanuginosum Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 135
(1911). – Finlaysonia lanuginosa (Ridl.) Venter in Venter & Verhoeven, Ann. Missouri
Bot. Gard. 88: 565 (2001). – TYPE: Peninsular Malaysia, Kedah, Langkawi Islands,
Langkawi, near Dayang Bunting, September 1890, C. Curtis 2663 (lectotype SING
[SING0059490], designated here; ?isolectotype K [K000545859]).
There is a Curtis collection in K, which lacks a collection number and date, that may
be a duplicate of the specimen in SING and is here cited as a possible isolectotype.
27. Gymnema dissitiorum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 29 (1912), as
‘dissitiora’. – Marsdenia dissitiora (Ridl.) P.I.Forst., Austral. Syst. Bot. 8(5): 693
(1995). – TYPE: Peninsular Malaysia, Perak, Tapah, Batang Padang, November 1908,
H.N. Ridley 14058 (lectotype SING [SING0059489], designated by Forster (1995);
isolectotypes BM [BM001190935], K [K001096327]).
28. Gymnema avum Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 68 (1911), as
‘ava’. – Tylophora ridleyi M.R.Hend., Gard. Bull. Straits Settlem. 7: 111 (1933). –
Lygisma avum (Ridl.) Kerr, Bull. Misc. Inform. Kew 1939: 456 (1939). – TYPE:
Peninsula Malaysia, Perak, Kota Tampan, July 1909, H.N. Ridley 14701 (lectotype
K [K000872882], designated by Liede-Schumann & Meve (2018); isolectotype BM
[BM001190936]).
314 Gard. Bull. Singapore 70 (2) 2018
In the protologue, Ridley gives the locality of the species as from Temengoh. We have
not found any specimen matching this location but there is a collection made on the
expedition to Temengoh [Temenggor] from Kota Tampan near Lenggong. The Kew
specimen annotated by Ridley is here designated lectotype.
29. Gymnema hirtum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 29 (1912), as
‘hirta’. – Marsdenia hirta (Ridl.) P.I.Forst., Austral. Syst. Bot. 8(5): 693 (1995). –
TYPE: Peninsular Malaysia, Selangor, Semangkok Pass, [13] April 1911, H.N. Ridley
15564 (lectotype SING [SING0059488], designated by Forster (1995); isolectotype K
[K000928001]).
30. Holarrhena densiora Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 131 (1911). –
TYPE: Thailand, s. loc., N. Cantley 2746 (lectotype SING [SING0037967], designated
by Middleton (2006)). Additional syntypes: Thailand, Bangtaphan, A. Keith (not
traced); Thailand, between Singgora and Nong-chik, St. V.B. Down (not traced).
= Holarrhena curtisii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2):
446 (1908); de Kruif, Meded. Landbouwhoogeschool Wageningen 81-2: 6 (1981);
Middleton, Fl. Males., Ser. 1, 18: 198 (2007).
31. Holarrhena latifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 131 (1911). –
TYPE: Thailand, Bangtapahn, 19 March 1890, A. Keith (not traced).
= Holarrhena curtisii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2):
446 (1908); de Kruif, Meded. Landbouwhoogeschool Wageningen 81-2:6 (1981);
Middleton, Fl. Males., Ser. 1, 18: 198 (2007).
32. Holarrhena pauciora Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 132 (1911).
– TYPE: Peninsular Malaysia, Perlis, [Ginting Kabok], [March 1910], H.N. Ridley
14980 (lectotype K [K000857983], designated by Middleton (2006); isolectotypes K
[K000857983], SING [SING0056468]). Additional syntypes: Thailand, Kopah, W.
Fox (not traced); ibid., Ridley 2610 (not traced); Ban Aor to Ningchik, 9 June 1906,
St. V.B. Down (not traced).
= Spirolobium cambodianum Baill., Bull. Mens. Soc. Linn. Paris 1: 773 (1889);
Middleton, Fl. Males., Ser. 1, 18: 360 (2007).
33. Holarrhena pulcherrima Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 130 (1911). –
TYPE: Peninsular Malaysia, Perlis, Kanga, Ginting Kabok, 1 March 1910, H.N. Ridley
14982 (lectotype K [K000857435], designated by de Kruif (1981); isolectotypes BM,
SING [SING0056470]). Additional syntypes: Peninsular Malaysia, Perlis, Chupeng,
H.N. Ridley 14981 (US [US00111760]); Thailand, Setul heaths, Unknown collector
(not traced).
315
The Gentinales taxa of H.N. Ridley
= Holarrhena curtisii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2):
446 (1908); de Kruif, Meded. Landbouwhoogeschool Wageningen 81-2: 6 (1981);
Middleton, Fl. Males., Ser. 1, 18: 198 (2007).
Middleton (2007) noted that he was unable to nd the K lectotype and designated
the SING duplicate a lectotype if the K specimen was truly lost. The K specimen has
reappeared and the later lectotypication is, therefore, unnecessary.
34. Hoya brooksii Ridl., Bull. Misc. Inform. Kew 1925: 85 (1925). – TYPE: Sumatra,
Lubuk Tandai, June 1922, C.J. Brooks 7615 (lectotype K [K000613013], designated
here, or possibly holotype).
35. Hoya citrina Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 300 (1922). – Hoya
parasitica Wall. ex Wight var. citrina (Ridl.) Rintz, Malayan Nat. J. 30: 514 (1978).
– Hoya acuta Haw. var. citrina (Ridl.) D.H.Kent, Hoyan 11(1:2): i (1989). – Hoya
verticillata (Vahl) G.Don var. citrina (Ridl.) Veldkamp, Blumea 41(2): 440 (1996). –
TYPE: Peninsular Malaysia, Selangor, Batu Caves, 13 December 1920, H.N. Ridley
s.n. (lectotype K [K000895117], designated by Rintz (1978)). Additional syntypes:
Peninsular Malaysia, Perak, Ulu Bubong, June 1886, King’s Collector 10316 (K
[K000895116]); Peninsular Malaysia, Perak, Batu Kuraw, December 1884, B.
Scortechini 1626 (K [K000895118]).
36. Hoya crassifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 30 (1912), nom. illeg.,
non (J.Jacq.) Haw. (1819). – Hoya caudata Hook.f. var. crassifolia Ridl., Fl. Malay
Penins. 2: 395 (1923). – TYPE: Peninsular Malaysia, Perak, Tapah Batang Padang,
November 1908, H.N. Ridley 14059 (lectotype K [K001096330], designated by Rintz
(1978); isolectotype SING [SING0059473]).
= Hoya caudata Hook.f., Fl. Brit. India 4: 60 (1883); Rintz, Malayan Nat. J. 30: 484
(1978).
37. Hoya globiora Ridl., J. Fed. Malay States Mus. 5: 164 (1915). – TYPE: Thailand,
S.W. Koh Pennan, 26 May 1913, H.C. Robinson 5756 (lectotype K [K001129444],
designated by Rodda (2017), or possibly holotype).
= Hoya verticillata (Vahl) G.Don, Gen. Hist. 4: 128 (1837); Veldkamp et al., Blumea
40: 427 (1995).
38. Hoya occlusa Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 31 (1912). – TYPE:
Peninsular Malaysia, Selangor, Gua Batu, December 1890, H.N. Ridley s.n. (lectotype
SING [SING0059478], designated by Rintz (1978), or possibly holotype).
316 Gard. Bull. Singapore 70 (2) 2018
= Hoya coriacea Blume, Bijdr. Fl. Ned. Ind. 16: 1063 (1826); Rintz, Malayan Nat. J.
30: 495 (1978).
39. Hoya perakensis Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 70 (1911). – TYPE:
Peninsular Malaysia, Perak, Temengoh, July 1909, H.N. Ridley s.n. (lectotype SING
[SING121210], designated here). Additional syntype: Peninsular Malaysia, Perak,
Kuala Kenering, August 1909, H.N. Ridley s.n. (SING [SING121209]).
= Hoya verticillata (Vahl) G.Don, Gen. Hist. 4: 128 (1837).
40. Hoya rhodostele Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 77 (1923). – TYPE:
Sumatra, Berastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K [K000894741],
designated here). Additional syntype: Sumatra, Berastagi, 13 February 1921, H.N.
Ridley s.n. (K [K000894740]).
41. Hoya rufolanata Ridl., Fl. Malay Penins. 2: 402 (1923), as ‘rufo-lanata’. – TYPE:
Peninsular Malaysia, Perak, Sungei Siput, 11 September 1920, I.H. Burkill SFN 6332
(lectotype K [K000910079], designated by Rintz (1978)); isolectotypes BO, SING
[SING0059479]).
= Genianthus rufovelutinus King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist.
74(2): 526 (1908); Rintz, Malayan Nat. J. 30: 520 (1978); Klackenberg, Bot. Jahrb.
Syst. 117: 458 (1995).
Klackenberg (1995) designated the SING duplicate of the type collection as the
lectotype, but Rintz (1978) had already effectively chosen the K sheet.
42. Hoya sarcophylla Ridl., J. Fed. Malay States Mus. 8(4): 62 (1917). – TYPE:
Sumatra, Korinchi, Siolak Daras, 3000 ft, 17 March 1914, H.C. Robinson & C.B.
Kloss s.n. (lectotype BM [BM001190937], designated here, or possibly holotype).
43. Hoya variifolia Ridl., Bull. Misc. Inform. Kew 1926: 74 (1926). – TYPE: Sumatra,
Sipora, 27 October 1924, C.B. Kloss 14793 (holotype K [K000894735]; isotype SING
[SING005940]).
44. Hoyella rosea Ridl., J. Fed. Malay States Mus. 8(4): 62 (1917). – Dischidia hoyella
Omlor, Gen. Revis. Marsdenieae 143 (1998). – TYPE: Sumatra, Korinchi Expedition,
1914, H.C. Robinson & C.B. Kloss s.n. (lectotype BM [BM000017060], designated
by Omlor (1998); isolectotype K [K000014445]).
317
The Gentinales taxa of H.N. Ridley
45. Kopsia alba Ridl. ex M.R.Hend., Gard. Bull. Straits Settlem. 5: 78 (1930). – TYPE:
Peninsular Malaysia, Pahang, Pulau Tioman, Ayer Surin, 18 May 1927, Md Nur SFN
18948 (lectotype K [K001129442], designated by Sévenet et al. (1994); isolectotypes
BO, SING [SING0056483], UC).
= Kopsia pauciora Hook.f., Fl. Brit. India 3: 639 (1882); Middleton, Harvard Pap.
Bot. 9: 120 (2004), Fl. Males., Ser. 1, 18: 250 (2007).
46. Kopsia dasyrachis Ridl., Bull. Misc. Inform. Kew 1934: 123 (1934); Middleton,
Harvard Pap. Bot. 9: 100 (2004), Fl. Males., Ser. 1, 18: 238 (2007). – TYPE: Borneo,
British North Borneo {Sabah}, Lukan, 8 March 1930, Arsat 1211 (lectotype K
[K000195022], designated here; isolectotype K [K000852942]).
47. Kopsia singapurensis Ridl., Fl. Malay Penins. 2: 336 (1923); Middleton,
Harvard Pap. Bot. 9: 127 (2004); Middleton, Fl. Males., Ser. 1, 18: 254 (2007). –
TYPE: Singapore, Chan Chu Kang, 21 March 1894, H.N. Ridley s.n. (lectotype K
[K000852952], designated by Middleton (2004)). Additional syntypes: Singapore,
Chan Chu Kang, 1892, H.N. Ridley 59 (lectotype BM, designated by Sévenet et
al. (1994) but not found); ibid., 1890, H.N. Ridley 59 (BM [BM000541180], SING
[SING0056487]); ibid., October 1889, H.N. Ridley 59 (SING [SING0056488]);
ibid., 30 September 1889, H.N. Ridley 59 (SING [SING0054249]); ibid., 1891,
H.N. Ridley s.n. (SING [SING0056489]); ibid., 1 June 1890, H.N. Ridley s.n. (SING
[SING0056490]); Peninsular Malaysia, Johore, Kwala Kahang, 27 October 1892,
H.W. Lake & H.J. Kelsall s.n. (SING [SING0054248]); Johore, Kota Tinggi, Unknown
collector (not traced); Pahang, Rompin River, Watson s.n. (not traced).
Sévenet et al. (1994) designated a lectotype for Kopsia singapurensis Ridl. as ‘Ridley
59, BM!, 1892’. There is a specimen of Ridley 59 in BM, but it is dated 1890, rather than
1892. As the extant specimens of Ridley 59 were collected on various different dates it
cannot be assumed that 1892 was a misprint of 1890, especially as the 1890 specimen
in BM has no mature owers and would have been an odd choice of lectotype given
the many syntypes. Middleton (2004) therefore selected another lectotype (Ridley s.n.,
21 March 1894, K [K000852952]). If a BM specimen of Ridley 59 collected in 1892 is
found, it will have priority over the choice of Middleton (2004).
48. Lyonsia sumatrana Ridl., Bull. Misc. Inform. Kew 1925: 84 (1925). – TYPE:
Sumatra, Bencoolen, Lubok Tandai, May 1922, C.J. Brooks 7100 (lectotype K
[K000857532], designated by Middleton (1997)). Additional syntype: Sumatra,
Bencoolen, Lubok Tandai, May 1922, C.J. Brooks 7104 (K [K000857534]).
= Parsonsia alboavescens (Dennst.) Mabb., Taxon 26: 532 (1977); Middleton,
Blumea 42: 198 (1997); Middleton, Fl. Males., Ser. 1, 18: 314 (2007).
318 Gard. Bull. Singapore 70 (2) 2018
49. Marsdenia stellaris Ridl., J. Fed. Malay States Mus. 5: 40 (1914). – Dregea
stellaris (Ridl.) Ridl., Fl. Malay Penins. 2: 387 (1923). – Dittoceras stellaris (Ridl.)
Bullock, Kew Bull. 11(3): 513 (1957). – TYPE: Peninsular Malaysia, Selangor,
Gunong Mengkuang, 5000 ft, 3 February 1913, N.C. Robinson s.n. (holotype K
[K000894719]).
Although the label on the type specimen gives Robinson as the collector, Ridley
(1914) wrote that it was actually Dyak Collectors who collected the material on which
the species was based. The protologue also notes that there was only one specimen,
hence it is a holotype.
50. Melodinus cymosus Ridl., J. Fed. Malay States Mus. 10: 146 (1920). – TYPE:
Peninsular Malaysia, Malacca, Selandai, 7 April 1915, I.H. Burkill 789 (lectotype K
[K000832838), designated by Middleton (1993); isolectotype SING [SING0056496]).
= Willughbeia avescens Dyer ex Hook.f., Fl. Brit. India 3: 625 (1882); Middleton,
Blumea 38: 12 (1993); Middleton, Fl. Males., Ser. 1, 18: 429 (2007).
51. Melodinus lancifolius Ridl., Bull. Misc. Inform. Kew 1934: 123 (1934). – TYPE:
Borneo, British North Borneo {Sabah}, Kudat, December 1884, M. Fraser 70
(lectotype K [K000832945], designated here, or possibly holotype).
= Melodinus orientalis Blume, Bijdr. Fl. Ned. Ind. 16: 1026 (1826); Leeuwenberg,
Syst. & Geogr. Pl. 73: 42 (2003); Middleton, Fl. Males., Ser. 1, 18: 280 (2007).
52. Micrechites brachypetalus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 96 (1918),
as ‘brachypetala’. – TYPE: Peninsular Malaysia, Penang, Penara Bukit, May 1886, C.
Curtis 850 (lectotype K [K000857758], designated by Middleton (1994); isolectotypes
BM [BM000884880], SING [SING0056499, SING0056500]).
= Micrechites serpyllifolius (Blume) Kosterm., Reinwardtia 5: 246 (1960); Middleton,
Fl. Males., Ser. 1, 18: 287 (2007).
53. Micrechites furcatus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 95 (1918), as
‘furcata’. – TYPE: Peninsular Malaysia, Perak, overhanging Bernam River, April
1886, King’s Collector 8859 (lectotype K [K000857755], designated by Middleton
(1994); isolectotypes BM [BM000884879], K [K000857756]). Additional syntype:
Peninsular Malaysia, Perak, Relau Tujor, ?July 1888, L. Wray 2604 (K [K000857757]).
= Micrechites serpyllifolius (Blume) Kosterm., Reinwardtia 5: 246 (1960); Middleton,
Fl. Males., Ser. 1, 18: 287 (2007).
319
The Gentinales taxa of H.N. Ridley
54. Micrechites tenuifolius Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 96 (1918),
as ‘tenuifolia’. – TYPE: Peninsular Malayia, Malacca, 11 November 1867/8, A.C.
Maingay 3394 [Kew Distrib. No. 1081] (lectotype K [K000857759], designated by
Middleton (1994)). Additional syntypes: Peninsular Malaysia, Selangor, Ginting Bidai,
H.N. Ridley 7142 (not traced); Kuala Lumpur, 1905 H.N. Ridley 1857 (not traced).
= Micrechites serpyllifolius (Blume) Kosterm., Reinwardtia 5: 246 (1960); Middleton,
Fl. Males., Ser. 1, 18: 287 (2007).
55. Micrechites tubulosus Ridl., Fl. Malay. Penin. 5: 321 (1925). – Anodendron
tubulosum (Ridl.) D.J.Middleton, Blumea 39: 89 (1994); Middleton, Fl. Males., Ser.
1, 18: 142 (2007). – TYPE: Peninsular Malaysia, Perak, Maxwell’s Hill, 9 March
1924, I.H. Burkill & Md. Haniff SFN 12785 (lectotype K [K000894512], designated
by Middleton (1996); isolectotype SING [SING0056541]).
56. Neowollastonia tabernaemontanoides Wernham ex Ridl., Hooker’s Icon. Pl. 31(3):
t. 3060 (1916); Wernham, Trans. Linn. Soc. London, Bot. 9: 110 (1916). – TYPE: New
Guinea, Mt Carstensz, February 1913, C.B. Kloss s.n. (holotype BM [BM000603826]).
= Melodinus forbesii Fawc. in Forbes, Nat. Wand. East. Archip. 510 (1885);
Leeuwenberg, Syst. & Geogr. Pl. 73: 31 (2003); Middleton, Fl. Males., Ser. 1, 18: 277
(2007).
57. Spiladocorys angustifolia Ridl., Trans. Linn. Soc. London, Bot. 3(9): 322
(1893). – TYPE: Peninsular Malayia, Perak, Thaiping, Waterfall, 1892, H.N. Ridley
2885 (lectotype BM [BM001014182], designated by Rahman & Wilcock (1991);
isolectotypes K [K000872855], SING [SING0122103]). Additional syntype:
Peninsular Malayia, Pahang, Tahan River, 1891, H.N. Ridley 2589 (K [K000872856]).
= Pentasachme caudatum Wall. ex Wight, Contr. Bot. India 60 (1834); Rahman &
Wilcock, Blumea 36: 115 (1991), as ‘Pentasacme’.
58. Stephanotis parviora Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 69 (1911). –
Marsdenia ridleyi P.I.Forst., Austral. Syst. Bot. 8(5): 700 (1995). – TYPE: Peninsular
Malaysia, Perak, Tapah, Tenok Road, 1908, H.N. Ridley s.n. (lectotype SING
[SING0072733], designated by Forster (1995)). Additional syntype: Peninsular
Malaysia, Perak, Ulu Temango, July 1909, H.N. Ridley s.n. (SING [SING0072734]).
59. Trachelospermum laurifolium Ridl., J. Fed. Malay States Mus. 5: 163 (1915).
– Kibatalia laurifolia (Ridl.) Woodson, Sunyatsenia 3: 102 (1936); Rudjiman,
Agric. Univ. Wageningen Pap. 86-5: 60 (1986); Middleton, Fl. Males., Ser. 1, 18:
217 (2007). – Paravallaris laurifolia (Ridl.) Kerr in Craib, Fl. Siam. 2: 455 (1939).
320 Gard. Bull. Singapore 70 (2) 2018
– TYPE: Thailand, Koh Pennan, 28 May 1913, H.C. Robinson 5764 (lectotype K
[K000857965], designated by Rudjiman (1986); isolectotypes K [K000857964], P
[P00495719, P00495720]).
60. Trachelospermum obtusifolium Ridl., J. Bot. 62: 298 (1924). – TYPE: Peninsular
Malaysia, Pahang, on the Selangor border, Fraser {Fraser’s} Hill, 16–30 September
1922, I.H. Burkill & R.E. Holttum SFN 8898 (lectotype K [K000894517], designated
by Middleton (1996); isolectotype SING [SING0055203]).
= Anodendron wrayi King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2):
489 (1908); Middleton, Blumea 41: 63 (1996); Middleton, Fl. Males., Ser. 1, 18: 145
(2007).
61. Tylophora avescens Ridl., J. Fed. Malay States Mus. 5: 163 (1915). – TYPE:
Thailand, Koh Pennan, 26 May 1913, H.C. Robinson 5751 (lectotype K [K001096328],
designated by Liede-Schumann & Meve (2018), or possible holotype).
= Vincetoxicum indicum (Burm.f.) Mabb., Mabberley’s Pl. Book 1102 (2017).
62. Tylophora grandiora Ridl., J. Bot. 62: 299 (1924), nom. illeg., non R.Br. (1810). –
Tylophora splendida P.I.Forst., Taxon 41(4): 750 (1992). – Vincetoxicum splendidum
(P.I.Forst.) Meve & Liede, Phytotaxa 369: 163 (2018). TYPE: Peninsular Malaysia,
Pahang, between Fraser {Fraser’s} Hill and Tras Valley, 3700 ft, 18 September 1923,
R.E. Holttum SFN 11482 (lectotype SING [SING0059539], designated by Forster
(1992); isolectotypes, BM [BM001190934], K [K000894971]).
63. Tylophora squarrosa Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 30 (1912).
– TYPE: Singapore, Woodlands, June 1903, H.N. Ridley s.n. (lectotype SING
[SING0059540], designated by Liede-Schumann & Meve (2018)). Additional syntype:
Peninsular Malaysia, Johore, Tana Merah Road, 8 January 1890, H.N. Ridley 2737
(BM [BM001190933], SING [SING0059499]).
= Vincetoxicum globiferum (Hook.f.) Kuntze, Revis. Gen. Pl. 1: 424 (1891).
64. Vallaris macrantha Ridl., J. Fed. Malay States Mus. 10: 101 (1920). – Beaumontia
macrantha (Ridl.) Rudjiman, Meded. Landbouwhoogeschool 82(11): 15 (1982);
Rudjiman, Agric. Univ. Wageningen Pap. 86-5: 24 (1986). – TYPE: Thailand, Tapli,
5 November 1919, C.B. Kloss 6946 (lectotype K [K000894555], designated by
Rudjiman (1982), or possibly holotype).
321
The Gentinales taxa of H.N. Ridley
65. Voacanga havilandii Ridl., Bull. Misc. Inform. Kew 1926: 473 (1926);
Leeuwenberg, Agric. Univ. Wageningen Pap. 85-3: 63 (1985); Middleton, Fl. Males.,
Ser. 1, 18: 417 (2007). – TYPE: Borneo, Sarawak, Kuching rock road, G.D Haviland
593 [c.m.p.d.] (lectotype K [K000857348), designated by Leeuwenberg (1985);
isolectotype SAR).
66. Willughbeia dulcis Ridl., Trans. Linn. Soc. London, Bot. 3(9): 319 (1893). – TYPE:
Peninsular Malaysia, Pahang, Pekan, Pramau, 1890, H.N. Ridley 1022 (lectotype SING
[SING0056572], designated here, or possibly holotype).
= Willughbeia edulis Roxb., Pl. Coromandel 3: 77, t. 280 (1819); Middleton, Blumea
38: 11 (1993); Middleton, Fl. Males., Ser. 1, 18: 427 (2007).
i. Alstonia latifolia (King & Gamble) Ridl., Fl. Malay Penins. 2: 347 (1923). – Alstonia
angustifolia var. latifolia King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2):
441 (1908).
= Alstonia angustifolia Wall. ex A.DC., Prodr. 8: 409 (1844); Sidiyasa, Blumea Suppl.
11: 101 (1998).
ii. Dischidia klossii Ridl. ex Kerr, Fl. Siam. 3(1): 46 (1951). – Dischidia lancifolia
Ridl., J. Fed. Malay States Mus. 10: 102 (1920), nom. illeg., non Merrill (1918).
= Dischidia acutifolia Maingay ex Hook.f. subsp. klossii (Ridl. ex Kerr) Rintz,
Blumea 26: 101 (1980).
iii. Dischidia rhodantha Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 97 (1918). –
Dischidia rosea Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 31 (1912), nom. illeg.,
non Schlechter (1906).
iv. Dregea stellaris (Ridl.) Ridl., Fl. Malay Penins. 2: 387 (1923). – Marsdenia stellaris
Ridl., J. Fed. Malay States Mus. 5: 40 (1914). – Dittoceras stellaris (Ridl.) Bullock,
Kew Bull. 11(3): 513 (1957).
v. Hoya caudata Hook.f. var. crassifolia Ridl., Fl. Malay Penins. 2: 395 (1923). –
Hoya crassifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 30 (1912), nom. illeg.,
non (J.Jacq.) Haw. (1819).
= Hoya caudata Hook.f., Fl. Brit. India 4: 160 (1883); Rintz, Malayan Nat. J. 30: 484
(1978).
322 Gard. Bull. Singapore 70 (2) 2018
vi. Physostelma wrayi (King & Gamble) Ridl., Fl. Malay Penins. 2: 411 (1923). –
Hoya wrayi King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74(2): 579 (1908).
vii. Willughbeia avescens Dyer var. rufescens (Dyer ex Hook.f.) Ridl., Fl. Malay
Penins. 2: 325 (1923). – Willughbeia rufescens Dyer ex Hook.f., Fl. Brit. India 3(9):
626 (1882).
= Willughbeia angustifolia (Miq.) Markgr., Blumea 20: 414 (1972).
Gentianaceae
67. Canscora trinervia Ridl., J. Linn. Soc., Bot. 38: 316 (1908). – TYPE: Peninsular
Malaysia, Pahang, Gunong Tahan, 5000–5500 ft, 10 July 1905, L. Wray & H.C. Robinson
5507 (lectotype BM [BM001014659], designated by Thiv (2003); isolectotype SING
[SING 0045419]).
= Canscora andrographioides Griff. ex C.B.Clarke, J. Linn. Soc., Bot. 14: 431 (1875);
Thiv, Blumea 48: 9 (2003).
68. Cyrtophyllum speciosum Ridl. var. montanum Ridl., Fl. Malay Penins. 5: 322
(1925). – TYPE: Peninsular Malaysia, Pahang, Gunung Terbakar, 9 June 1923,
M.R. Henderson SFN 10992 (lectotype K, designated here; isolectotypes SING
[SING0058130, SING0058131]).
= Picrophloeus javanensis Blume, Bijdr. Fl. Ned. Ind. 16: 1020 (1826); Wong &
Sugumaran, Gard. Bull. Singapore 64: 515 (2012).
69. Fagraea caudata Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 97 (1918); Wong &
Sugau, Sandakania 8: 8 (1996). – Cyrtophyllum caudatum (Ridl.) K.M.Wong, Gard.
Bull. Singapore 64(2): 499 (2012). – TYPE: Borneo, 1853, T. Lobb s.n. (lectotype K
[K000883549], designated here, or possibly holotype).
70. Fagraea avidula Ridl., Fl. Malay Penins. 5: 322 (1925). – TYPE: Peninsular
Malaysia, Pahang, Cameron’s {Cameron} Highlands, No. 4 Camp, 16 January 1924,
M.R. Henderson 11673 (lectotype K [K000883557], designated here; isolectotypes K
[K000883556], L [L0004964], SING [SING0058129]).
= Fagraea carnosa Jack, Malayan Misc. 2(7): 81 (1822); Leenhouts, Fl. Males., Ser.
1, 6: 331 (1962); Wong & Sugau, Sandakania 8: 57 (1996); Wong & Sugumaran,
Sandakania 21: 84 (2016).
323
The Gentinales taxa of H.N. Ridley
71. Fagraea gardenioides Ridl., J. Fed. Malay States Mus. 5: 42 (1914); Leenhouts,
Fl. Males., Ser. 1, 6: 331 (1962); Wong & Sugumaran, Sandakania 21: 96 (2016).
– TYPE: Peninsular Malaysia, Selangor, Gunung Mengkuang, 22 January 1913,
H.C. Robinson s.n. (lectotype K [K000438448], designated here; isolectotype SING
[SING0052377]).
72. Fagraea gigantea Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 98 (1918). –
Cyrtophyllum giganteum (Ridl.) Ridl., Fl. Malay Penins. 2: 421 (1923). – TYPE:
Singapore, Gardens’ Jungle, 1897, H.N. Ridley 8921 (lectotype SING [SING0054758],
designated by Wong & Sugau (1996); isolectotype K [K001129448]). Additional
syntypes: Singapore, Gardens’ Jungle, H.N. Ridley 5818 (BM); Peninsular Malaysia,
Malacca, Bukit Sebukor, August 1889, R. Derry 272 (BM, SING [SING0060430]).
73. Fagraea intermedia Ridl., J. Fed. Malay States Mus. 8(4): 63 (1917). – TYPE:
Sumatra, Korinchi, Sungei Kumbang at 4500 ft, 1914, H.C. Robinson & C.B. Kloss
s.n. (lectotype BM [BM001014296], designated here). Additional syntypes: Sumatra,
Korinchi, Sungei Kumbang at 4500 ft, 1914, H.C. Robinson & C.B. Kloss 72 (BM
[BM001014297]); Sumatra, Barong Bharu, Barisan Range at 4000 ft., H.C. Robinson
& C.B. Kloss (not traced).
= Fagraea blumei G.Don, Gen. Hist. 4: 69 (1837), as ‘blumii’; Leenhouts, Fl. Males.,
Ser. 1, 6: 322 (1962).
74. Fagraea rotundifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 50: 117 (1908).
– TYPE: Peninsular Malaysia, Terengganu, Bundi, February 1904, Rostado s.n.
(lectotype K, designated by Leenhouts (1962), not traced).
= Fagraea carnosa Jack, Malayan Misc. 2(7): 81 (1822); Leenhouts, Fl. Males., Ser.
1, 6: 331 (1962); Wong & Sugumaran, Sandakania 21: 85 (2016).
Leenhouts (1962) mentioned the Rostado specimen as being at Kew but we have been
unable to trace it.
75. Gentiana loerzingii Ridl., Bull. Misc. Inform. Kew 1939: 30 (1939). – TYPE:
Sumatra, Sibolangat, Raja, Karo Plateau c. 1300 m, 24 January 1920, J.A. Lörzing
7075 (holotype BO; isotype K (fragment) [K000872093]).
76. Gentiana malayana Ridl., J. Linn. Soc., Bot. 38: 316 (1908). – Gentiana borneensis
Hook.f. subsp. malayana (Ridl.) Halda, Acta Mus. Richnov., Sect. Nat. 3(1): 45
(1995). – TYPE: Peninsular Malaysia, Pahang, Gunong Tahan, 5–6000 ft, 6 July 1905,
L. Wray & H.C. Robinson 5479 (lectotype BM [BM000959955], designated by Ho
324 Gard. Bull. Singapore 70 (2) 2018
& Liu (2001); isolectotype SING [SING0209186]). Additional syntype: Peninsular
Malaysia, Pahang, Gunong Tahan, 5–6000 ft, 5 July 1905, L. Wray & H.C. Robinson
5473 (BM [BM000959956]).
= Gentiana borneensis Hook.f., Hooker’s Icon. Pl. 15: t. 1440B (1883); Ho & Liu,
Worldw. Monogr. Gentiana 421 (2001).
77. Gentiana sumatrana Ridl., J. Fed. Malay States Mus. 8(4): 58 (1917); Ho & Liu,
Worldwide Monogr. Gentiana 331 (2001). – TYPE: Sumatra, Korinchi Peak, 10000
ft, 27 April 1914, H.C. Robinson & C.B. Kloss s.n. (lectotype K [K000872103],
designated here). Additional syntypes: Sumatra, Korinchi Peak, 10000–11000 ft, 10
May 1914, H.C. Robinson & C.B. Kloss s.n. (BM, SING [SING0240298], US); ibid.,
10500 ft, 9 May 1914, H.C. Robinson & C.B. Kloss s.n. (K [K000872104]).
viii. Cyrtophyllum giganteum (Ridl.) Ridl., Fl. Malay Penins. 2: 421 (1923). – Fagraea
gigantea Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 98 (1918).
ix. Fagraea pauciora (King & Gamble) Ridl., Fl. Malay Penins. 2: 419 (1923). –
Fagraea racemosa var. pauciora King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat.
Hist. 74(2): 609 (1908).
= Utania volubilis (Wall.) M.Sugumaran in Wong & Sugumaran, Gard. Bull. Sing. 64:
493 (2012); Sugumaran & Wong, Pl. Ecol. Evol. 147: 221 (2014).
Loganiaceae
x. Norrisia malaccensis Gardn. var. major (Soler.) Ridl., Fl. Malay Penins. 2: 414
(1923). – Norrisia major Soler. in Engler & Prantl, Nat. Panzenfam. 4(2): 37 (1892),
as ‘maior’.
Rubiaceae
78. Aleisanthia Ridl., J. Fed. Malay States Mus. 10: 138 (1920). – TYPE: Aleisanthia
rupestris (Ridl.) Ridl., designated by Tange (1996).
79. Argostemmella Ridl., J. Bot. 65: 41 (1927). – TYPE: non designatus.
= Argostemma Wall. in Roxburgh, Fl. Ind. 2: 324 (1824).
80. Becheria Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 20 (1912). – TYPE: Becheria
parviora Ridl.
325
The Gentinales taxa of H.N. Ridley
81. Gardenia section Gardeniella Ridl., J. Fed. Malay States Mus. 4: 30 (1909). –
TYPE Gardenia pulchella Ridl., nom. illeg., non G.Don (1824), designated here.
= Acranthera Arn. ex Meisn., Pl. Vasc. Gen. 162 (1840); Wong, Gard. Bull. Singapore
35: 29 (1982).
82. Klossia Ridl., J. Fed. Malay States Mus. 4: 27 (1909). – TYPE: Klossia montana
Ridl.
83. Perakanthus Robyns ex Ridl., Fl. Malay Penins. 5: 317 (1925). – TYPE:
Perakanthus velutinus (Ridl.) Robyns ex Ridl., lectotype designated by Wong (1988a).
84. Polycycliska Ridl., Bull. Misc. Inform. Kew 1926: 67 (1926). – TYPE: Polycycliska
cylindrica Ridl.
= Lerchea L., Mant. Pl. 2: 155 (1771), nom. cons.
85. Pomazota Ridl., Trans. Linn. Soc. London, Bot. 3(9): 308 (1893). – TYPE:
Pomazota sylvestris Ridl.
= Coptophyllum Korth., Ned. Kruidk. Arch. 2: 161 (1851), nom. cons.
86. Urophyllum section Pleiocarpidia Ridl., J. Bot. 70: 192 (1932). – TYPE:
Pleiocarpidia enneandra (Wight) K.Schum., lectotype designated by Bremekamp
(1940a).
87. Urophyllum section Grandiorae Ridl., J. Bot. 70: 192 (1932). – TYPE: non
designatus = Rothmannia in part.
88. Urophyllum section Axanthes Ridl., J. Bot. 70: 192 (1932). – TYPE: non
designatus.
89. Urophyllum section Tubiorae Ridl., J. Bot. 70: 192 (1932). – TYPE: non
designatus.
90. Zeuxanthe Ridl., Bull. Misc. Inform. Kew 1939: 192 (1940). – TYPE: Zeuxanthe
beccariana (Baill.) Ridl.
= Prismatomeris Thwaites, Hooker’s J. Bot. Kew Gard. Misc. 8: 268 (1856).
326 Gard. Bull. Singapore 70 (2) 2018
91. Adenosacme ava Ridl., J. Fed. Malay States Mus. 4: 29 (1909). – Mycetia ava
(Ridl.) Ridl., Fl. Malay Penins. 2: 64 (1923); Wong, Tree Fl. Malaya 4: 379 (1989).
– TYPE: Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley s.n.
(lectotype SING [SING0209192], designated here, or possibly holotype).
92. Adenosacme lanceolata Ridl., J. Fed. Malay States Mus. 4: 29 (1909), nom.
illeg., non Miq. (1857). – Mycetia lanceolata Ridl., Fl. Malay Penins. 2: 64 (1923);
Wong, Tree Fl. Malaya 4: 379 (1989). – TYPE: Peninsular Malaysia, Pahang, Telom,
November 1908, H.N. Ridley 13608 (lectotype SING [SING0209191], designated
here; isolectotype K [K000760582]).
= Mycetia fasciculata (Blume) Blume ex Korth., Ned. Kruidk. Arch. 2(2): 118 (1851).
93. Aleisanthia sylvatica Ridl., J. Fed. Malay States Mus. 10: 138 (1920); Tange,
Nordic J. Bot. 16: 565 (1996). – TYPE: Peninsular Malaysia, Kelantan, Kuala Lebir,
February 1917, H.N. Ridley s.n. (lectotype K [K000174148], designated by Tange
(1996)).
94. Amaracarpus caudatus Ridl., J. Fed. Malay States Mus. 10: 143 (1920). – TYPE:
Peninsular Malaysia, Perak, Taiping Hills, Birch’s Hill, s.dat., L. Wray 670 (lectotype
K [K000763888], designated here, or possibly holotype).
= Paralasianthus lowianus (King & Gamble) H.Zhu, Phytotaxa 202: 276 (2015).
95. Argostemma acuminatum King var. pubescens Ridl., J. Straits Branch Roy. Asiat.
Soc. 57: 52 (1911). – TYPE: Peninsular Malaysia, Perak, Temengoh, H.N. Ridley (not
traced).
= Argostemma inaequilaterum Benn., Pl. Jav. Rar. 1: 95 (1838).
96. Argostemma acuminatissimum Ridl., Fl. Malay Penins. 5: 315 (1925). – TYPE:
Peninsular Malaysia, Perak, Jor, 13 September 1924, Md Haniff SFN 14213 (lectotype
K [K000172902], designated here; isolectotype SING [SING0057771]).
97. Argostemma aequifolium Ridl., J. Straits Branch Roy. Asiat. Soc. 35: 15 (1901),
as ‘aequifolia’. – TYPE: Peninsular Malaysia, Malacca, Gunong Ledang, Padang
Batu, June 1890, H.N. Ridley s.n. (lectotype SING [SING0057773], designated here,
or possibly holotype).
327
The Gentinales taxa of H.N. Ridley
98. Argostemma albociliatum Ridl., J. Linn. Soc., Bot. 38: 311 (1908). – Argostemma
aequifolium Ridl. var. albociliatum (Ridl.) Bakh.f., Blumea 7: 333 (1953). – TYPE:
Peninsular Malaysia, Pahang, Gunong Tahan, 3300 ft, 29 May 1905, L. Wray & H.C.
Robinson 5229 (lectotype BM [BM000028717], designated here; isolectotypes K
[K000760229], SING [SING0057774]).
99. Argostemma annamiticum Ridl., J. Bot. 65: 40 (1927). – TYPE: Vietnam, Annam,
Langbian Province, Dalat, 5000 ft, April–May 1918, C.B. Kloss s.n. (lectotype BM
[BM000028592], designated here).
The specimen designated as lectotype above has no annotations by Ridley but it is the
only specimen located that ts the protologue including the eld description ‘white,
stamens yellow’. Argostemma dispar Craib should probably be reduced to a synonym
of A. annamiticum.
100. Argostemma bifolium Ridl., Fl. Malay Penins. 5: 314 (1925). – TYPE: Peninsular
Malaysia, Perak, Maxwell’s Hill, December 1902, H.N. Ridley s.n. (lectotype K
[K000760231], designated by Ridley (1927), or possibly holotype).
101. Argostemma borragineum Blume var. padangense Ridl., J. Bot. 65: 32 (1927). –
TYPE: Sumatra, Padang, Mt Singalan, June–July 1878, O. Beccari P.S. 36 (lectotype K
[K000760138], designated here; isolectotype FI-B [FI008827]). Additional syntypes:
Sumatra, Gunong Merapi at 1100 m, 23 September 1918, H.A.B. Bünnemeijer 4869 (K
[K000760137], U [U0005933]).
= Argostemma montanum Blume ex DC., Prodr. 4: 418 (1830).
102. Argostemma borragineum Blume var. breviorum Ridl., J. Bot. 65: 33
(1927). – TYPE: Sumatra, Berastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000760139], designated here, or possibly holotype).
103. Argostemma borragineum Blume var. heterophyllum Ridl., J. Bot. 65: 33 (1927).
Type: Sumatra, Korinchi, Barong Baru no. 3, Tapan, 9 June 1914, H.C. Robinson
& C.B. Kloss s.n. (lectotype BM [BM000945104], designated here; isolectotype K
[K000760140]).
104. Argostemma corymbosum Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 67 (1923).
Argostemma borragineum Blume var. corymbosum (Ridl.) Bakh.f., Blumea 7: 333
(1953). – TYPE: Sumatra, Berastagi Hill Woods, 8 February 1921, H.N. Ridley s.n.
(lectotype K [K000760144], designated here, or possibly holotype).
328 Gard. Bull. Singapore 70 (2) 2018
105. Argostemma debile Ridl., J. Fed. Malay States Mus. 5: 35 (1914). – TYPE:
Peninsular Malaysia, Selangor, Gunong Mengkuang Lebah, 5000 ft, 14 January 1913,
H.C. Robinson s.n. (lectotype K [K000172877], designated here; isolectotypes BM
[BM000028714], SING [SING0057784]).
106. Argostemma densifolium Ridl., J. Bot. 65: 39 (1927); Bremer, Ann. Missouri
Bot. Gard. 76: 17 (1989). – TYPE: Borneo, Sarawak, Puak, September 1904, H.N.
Ridley 12448 (lectotype K [K000760169], designated here; isolectotype SING
[SING0057782]).
107. Argostemma densifolium Ridl. var. latifolium Ridl., J. Bot. 65: 40 (1927). – TYPE:
Borneo, British North Borneo {Sabah}, Bongaya, December 1897, H.N. Ridley s.n.
(lectotype SING [SING0057782], designated by Bremer (1989), or possibly holotype).
= Argostemma densifolium Ridl., J. Bot. 65: 39 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 17 (1989).
108. Argostemma denticulatum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 12 (1912).
– TYPE: Peninsular Malaysia, Selangor, Genting Simpai, April 1911, H.N. Ridley
15656 (lectotype K [K000172873], designated here, or possibly holotype).
109. Argostemma diversifolium Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 52 (1911);
Sridith, Thai Forest Bull., Bot. 27: 114 (1999). – TYPE: Peninsular Malaysia, Perak,
Lenggong, August 1909, H.N. Ridley 14478 (lectotype K [K000172898], designated
by Sridith (1999); isolectotypes BM [BM 000028710], SING [SING0057783]).
110. Argostemma elongatum Ridl., J. Fed. Malay States Mus. 6: 151 (1915). – TYPE:
Peninsular Malaysia, Pahang, Gunung Tahan, Teku River, July 1911, H.N. Ridley 16072
(lectotype K [K000172882], designated here; isolectotype BM [BM000028718]).
111. Argostemma enerve Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 13 (1912). –
TYPE: Peninsular Malaysia, Negri Sembilan, Gunong Angsi, February 1904, H.N.
Ridley s.n. (lectotype SING [SING0057780], designated here, or possibly holotype).
112. Argostemma enerve Ridl., J. Bot. 65: 28 (1927), nom. illeg., non Ridl. (1912).
Argostemma chaii B.Bremer, Ann. Missouri Bot. Gard. 76: 34 (1989). – TYPE:
Borneo, Sarawak, s.dat., Native Collector (Bureau of Science, Manila) 1227 (lectotype
K [K000760157], designated by Bremer (1989), or possibly holotype).
329
The Gentinales taxa of H.N. Ridley
113. Argostemma grandiorum Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 77 (1918).
– TYPE: Peninsular Malaysia, Perak, Gunong Kerbau, 4500 ft., June 1913, H.C.
Robinson s.n. (lectotype K [K000172909], designated here, or possibly holotype).
114. Argostemma havilandii Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 13 (1912);
Bremer, Ann. Missouri Bot. Gard. 76: 38 (1989). – TYPE: Borneo, Sarawak, Kuching,
15 May 1893, G.D. Haviland 2958 (lectotype SING [SING0057799], designated by
Bremer (1989); isolectotypes BO, K [K000760153]).
115. Argostemma hirsutum Ridl., J. Fed. Malay States Mus. 10: 139 (1920). –
TYPE: Peninsular Malaysia, Kedah-Perak boundary, Gunong Bintang, June 1917,
F.M.S. Museum Native Collector s.n. (lectotype K [K000172867], designated here, or
possibly holotype).
116. Argostemma hirtellum Ridl., J. Bot. 65: 40 (1927). – TYPE: Sumatra, Agam,
Brani, 950 m, 26 June 1918, H.A.B. Bünnemeijer 3338 (lectotype K [K000760280],
designated here; isolectotype SING [SING0057798]).
117. Argostemma hirtum Ridl., J. Straits Branch Roy. Asiat. Soc. 35: 14 (1901), as
‘hirta’. – TYPE: Peninsular Malaysia, Perak, Maxwell’s Hill, Tea Gardens, February
1891, H.N. Ridley 2927 (lectotype SING [SING0240810], designated here). Additional
syntype: Peninsular Malaysia, Malacca, Gunong Tunduk, Unknown collector (not
traced).
118. Argostemma hullettii Ridl., J. Bot. 65: 35 (1927). – TYPE: Borneo, Sarawak,
Matang, August 1884, R.W. Hullett 329 (lectotype SING [SING0057797], designated
by Bremer (1989)).
= Argostemma hameliifolium Wernham, J. Linn. Soc., Bot. 42: 89 (1914); Bremer,
Ann. Missouri Bot. Gard. 76: 27 (1989).
119. Argostemma invalidum Ridl., J. Bot. 65: 38 (1927). – TYPE: Sumatra,
Gunung Melintang, 1200 m, 5 August 1918, H.A.B. Bünnemeijer 4310 (lectotype K
[K000760273], designated here; isolectotypes SING [SING0057796], L [L0000131],
U [U0005936]). Additional syntype: Sumatra, Sibo Langit, Bukit Pasang, “Nur, Burkill
7244 in part” (not traced).
330 Gard. Bull. Singapore 70 (2) 2018
120. Argostemma involucratum Hemsl. var. hirsutum Ridl., Fl. Malay Penins. 2: 28
(1923). – Argostemma parvifolium Benn. f. hirsutum (Ridl.) B.C.Stone, Fed. Mus.
J. 26(1): 135 (1981). – TYPE: Peninsular Malaysia, Pahang, Gunung Tahan, July
1911, H.N. Ridley 16073 (lectotype, designated here, K [K001129458]). Additional
syntypes: Peninsular Malaysia, Telom, K’luang Terbang, W.D. Barnes s.n. (SING
[SING0240807]); Selangor Semangkok Pass, 21 January 1921, H.N. Ridley s.n. (K);
Selangor, Semangkok Pass track to Bukit Fraser, August 1904, H.N. Ridley 12078
(SING [SING0039491]); Pahang, Gunung Mengkuang Lebah, 6 February 1913,
Robinson s.n. (K); Perak, Gunong Kerbau, 20 March 1913, H.N. Ridley s.n. (K); Perak,
Bujong Malacca, Dec 1895, C. Curtis 3148,, (SING [SING0240808, SING024099]);
Penang, Government Hill, November 1885, C. Curtis 342 (K, SING [SING0240809,
SING0240300]); Penang, Richmond Pool, March 1915, H.N. Ridley s.n. (K); Selangor,
Menuang Gasing, Kloss (not traced); Perak, Thaiping Hills, Unknown collector (not
traced); Malacca, Mt Ophir, Unknown collector (not traced).
= Argostemma parvifolium Benn. var. involucratum (Hemsl.) Bakh.f., Blumea 7: 333
(1953).
Bakhuizen van den Brink’s varietal name has priority over Ridley’s because it is based
on the autonym (Argostemma involucratum var. involucratum) which has priority over
any variety not including the type of the species.
121. Argostemma involucratum var. sumatranum Ridl., Bull. Misc. Inform. Kew 1925:
83 (1925). – TYPE: Sumatra, Lubok Tandai, May 1922, C.J. Brooks 7190 (lectotype K
[K000760275], designated here, or possibly holotype).
= Argostemma squalens Ridl., J. Bot. 65: 37 (1927).
122. Argostemma johorense Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 11 (1912). –
TYPE: Peninsular Malaysia, Johore, Tempayan River, Easter 1908, H.N. Ridley 13270
(lectotype K [K000172872], designated here; isolectotypes BM [BM000028709],
SING [SING0057793]). Additional syntypes: Peninsular Malaysia, Johore, Sungei
Tebrau, Ridley13207 (not traced); Johore, Mt Austin, Serom near Batu Pahat, Unknown
collector (not traced).
123. Argostemma johorense Ridl. var. kelsallii Ridl., J. Straits Branch Roy. Asiat. Soc.
61: 12 (1912), as ‘Kelsalli’. – TYPE: Peninsular Malaysia, Selangor, Hulu Langat,
Bukit Hitam, January 1891, H.J. Kelsall s.n. (lectotype SING [SING00057794],
designated here, or possibly holotype).
= Argostemma johorense Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 11 (1912).
331
The Gentinales taxa of H.N. Ridley
124. Argostemma klossii Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 13 (1912). –
TYPE: Peninsular Malaysia, Johore, Ulu Sayong, June 1905, C.B. Kloss s.n. (lectotype
SING [SING0057792], designated here). Additional syntype: Peninsular Malaysia,
Negri Sembilan, Bukit Sulu, N. Cantley (not traced).
125. Argostemma lanceolatum Ridl., J. Fed. Malay States Mus. 4: 26 (1909). – TYPE:
Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley 13652 (lectotype
SING [SING0058416], designated here; isolectotype BM
[BM000028667]).
126. Argostemma longistipulum Ridl., Fl. Malay Penins. 5: 315 (1925), as
‘longistipula’; Ridley, J. Bot. 65: 39 (1927), as ‘longistipula’. – TYPE: Peninsular
Malaysia, Negri Sembilan, Gunung Angsi, 19 November 1909, H. Winkler 1775
(lectotype K [K000760225], designated here). Additional syntypes: Peninsular
Malaysia, Johore, Gunong Belumut, Ulu Madik, 23 May 1923, R.E. Holttum SFN
10637 (K [K000760228]); ibid., R.E. Holttum SFN 10657 (not traced); Negri Sembilan,
Gunung Angsi, 21 November 1923, Md. Nur SFN 11580 (K [K000760226]).
127. Argostemma monophyllum Ridl., J. Bot. 65: 27 (1927). – TYPE: India, Khasia,
Churra, 18 August 1850, J.D. Hooker & T. Thomson 1099 (lectotype K [K000031018],
designated here). Additional syntypes: India, Khasia, W. Grifth s.n. [HEIC 2874] (K);
ibid., W. Grifth s.n. [HEIC 2873] (K); Assam, North Cachar Hills, Haong, 2500 ft,
8 August 1908, W.G. Craib 176 (K).
128. Argostemma montanum Blume ex DC. var. minus Ridl., J. Bot. 65: 31 (1927),
as ‘minor’. – TYPE: Sumatra, Korinchi, Siola Daras, 4000 ft, 16 March 1914, H.C.
Robinson & C.B. Kloss s.n. (lectotype K [K001129447], designated here). Additional
syntype: Sumatra, Gunong Merapi, 1100 m, 23 September 1918, H.A.B. Bünnemeijer
4869 (K [K000760137], U [U0005933]).
= Argostemma montanum Blume ex DC., Prodr. 4: 418 (1830).
129. Argostemma montense Ridl., J. Fed. Malay States Mus. 8(4): 45 (1917). – TYPE:
Sumatra, Korinchi Peak, 26 April 1914, H.C. Robinson & C.B. Kloss 108 (lectotype
BM [BM000945105], designated here, or possibly holotype).
332 Gard. Bull. Singapore 70 (2) 2018
130. Argostemma motleyi Ridl., J. Bot. 65: 39 (1927). – TYPE: Borneo, Kalimantan,
Bangarmassing, 1857–1858, J. Motley 1174, (lectotype K [K000760171], designated
here, or possibly holotype).
= Argostemma elatostemma Hook.f., Fl. Brit. India 3: 45 (1880); Bremer, Ann.
Missouri Bot. Gard. 76: 14 (1989).
131. Argostemma moultonii Ridl., J. Bot. 65: 36 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 29 (1989). – TYPE: Borneo, Sarawak, Upper Baram, Gunong Temabok,
3000 ft, 6 November 1920, J.C. Moulton SFN 6695 (lectotype K [K000760162],
designated by Bremer (1989); isolectotype SING (not traced)). Additional syntype:
Borneo, Sarawak, Upper Baram, Gunong Temabok, 4000 ft, 5 November 1920, J.C.
Moulton SFN 6669 (K, SING (not traced)).
132. Argostemma moultonii Ridl. var. hirta Ridl., J. Bot. 65: 36 (1927). – TYPE:
Borneo, Sarawak, Upper Baram, Gunong Semabok, 7000 ft, 2 November 1920,
J.C. Moulton SFN 6654 (lectotype K [K00760163], designated by Bremer (1989);
isolectotype SING (not traced)).
= Argostemma moultonii Ridl., J. Bot. 65: 36 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 29 (1989).
133. Argostemma multinervium Ridl., J. Bot. 65: 33 (1927). – TYPE: Sumatra, Gunong
Sibayak, Bandar Baru, 6 August 1921, Md. Nur 7312 (lectotype K [K000760145],
designated here). Additional syntype: Sumatra, Berastagi, December 1921, E.M.
Burkill 123 (SING [SING0057791]).
134. Argostemma muscicola Ridl., J. Linn. Soc., Bot. 38: 310 (1908). – TYPE:
Peninsular Malaysia, Pahang, Gunong Tahan, 5000–6000 ft, 3 July 1905, L. Wray &
H.C. Robinson 5461 (lectotype BM [BM000028716], designated here; isolectotypes
K [K000172890], SING [SING0057790]).
135. Argostemma nervosum Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 77 (1918).
– TYPE: Peninsular Malaysia, Selangor, Sempang, April 1904, H.N. Ridley 15658
(lectotype K [K000172869], designated here; isolectotypes BM [BM000028713],
SING [SING0039481]).
136. Argostemma nutans King var. borneense Ridl., J. Bot. 65: 33 (1927). – TYPE:
Borneo, Sarawak, Upper Baram, Gunong Temabok, 7000 ft, 2 November 1920,
J.C. Moulton SFN 6653 (lectotype K [K000760158], designated by Bremer (1989);
333
The Gentinales taxa of H.N. Ridley
isolectotype SING [SING0057776]).
= Argostemma borragineum Blume ex DC., Prodr. 4: 417 (1830); Bremer, Ann.
Missouri Bot. Gard. 76: 32 (1989).
137. Argostemma ovatum Ridl., J. Fed. Malay States Mus. 8(4): 45 (1917). –
Argostemma borragineum Blume ex DC. f. ovatum (Ridl.) Bakh.f., Blumea 7: 333
(1953). – TYPE: Sumatra, Korinchi, Siolak Daras, 16 March 1914, C.B. Kloss s.n.
(lectotype K [K000760146], designated here, or possibly holotype).
138. Argostemma parvulum Ridl., J. Bot. 65: 36 (1927). – TYPE: Borneo, Sarawak,
Upper Baram, Lio Matu, 500 ft, 16 October 1920, J.C. Moulton SFN 6726 (lectotype
K [K000760159], designated by Bremer (1989), or possibly holotype).
= Argostemma moultonii Ridl., J. Bot. 65: 36 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 30 (1989).
139. Argostemma propinquum Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 53 (1911);
Sridith, Thai Forest Bull., Bot. 27: 89 (1999). – TYPE: Peninsular Malaysia, Perak,
Ulu Temango, July 1904, H.N. Ridley 14476 (lectotype K [K000172870], designated
by Sridith (1999); isolectotypes BM [BM000028711], SING (not traced)). Additional
syntype: Peninsular Malaysia, Trengganu, Bundi, Rostado s.n. (SING (not traced)).
Sridith (1999) referred to the type as Ridley s.n., but the specimens are numbered
14476. As Sridith gave the locality correctly, we take the error in citing the collection
number as correctible and treat the typication as effective.
140. Argostemma psychotrioides Ridl., J. Bot. 65: 38 (1927), as ‘psychotroides’;
Bremer, Ann. Missouri Bot. Gard. 76: 43 (1989). – TYPE: Borneo, Sarawak, Kuching,
G.D. Haviland 689 [c.a.b.a.] (lectotype K [K000760150], designated by Bremer
(1989); isolectotype SAR). Additional syntypes: Borneo, Lobb (not traced); Borneo,
Lowe (not traced); Borneo, Sarawak, Kuching, G.D. Haviland 668 (not traced); ibid.,
G.D. Haviland 2960 (BO n.v., L n.v., SAR n.v.).
141. Argostemma reptans Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 11 (1912). –
TYPE: Peninsular Malaysia, Kedah, Kedah Peak, rocks in stream, June 1893, H.N.
Ridley s.n. (lectotype SING [SING0058411], designated here, or possibly holotype).
142. Argostemma riparium Ridl., J. Bot. 65: 34 (1927). – TYPE: Sumatra, Siak,
Sungei Kelantan, November 1897, H.N. Ridley 8966 (lectotype K [K000760141),
designated here; isolectotype SING [SING0058410]).
334 Gard. Bull. Singapore 70 (2) 2018
143. Argostemma rugosum Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 76 (1918).
– TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang, 3600 ft, 22 January
1913, H.C. Robinson s.n. (lectotype K [K000172880], designated here, or possibly
holotype).
144. Argostemma rupestre Ridl., J. Bot. 65: 41 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 23 (1989). – TYPE: Borneo, Sarawak, on the hill near Matang, 20 February
1892, G.D. Haviland 1031 [e.d.e.r.] (lectotype K [K000760165], designated here;
isolectotypes SAR!, SING [SING0058413]).
145. Argostemma salicifolium Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 14
(1912). – TYPE: Borneo, Sarawak, Matang, July 1902, H.N. Ridley 11751
(lectotype K [K000760168], designated by Bremer (1989); isolectotypes BM, SING
[SING0058414]). Additional syntype: Borneo, Sarawak, Matang, Hullett (not traced).
= Argostemma ophirense Maingay ex Hook.f., Fl. Brit. India 3: 45 (1880); Bremer,
Ann. Missouri Bot. Gard. 76: 18 (1989).
Bremer (1989) referred to a holotype for Argostemma salicifolium, but this is effectively
a lectotypication as Ridley denitely referred to both Ridley and Hullett collections
in the protologue.
146. Argostemma sessilifolium Ridl., J. Fed. Malay States Mus. 8(4): 45 (1917). –
TYPE: Sumatra, Korinchi, Siolak Daras, 16 March 1914, H.C. Robinson & C.B. Kloss
s.n. (lectotype BM [BM000945093], designated here, or possibly holotype).
147. Argostemma squalens Ridl., J. Bot. 65: 37 (1927). – TYPE: Sumatra, Padang,
Ayer Mancior, c. 360 m, August 1878, O. Beccari P.S. 608 (lectotype K [K000760276],
designated here; isolectotypes FI-B [FI008828], L [L0000127, L0000128]).
148. Argostemma stellatum Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 67 (1923),
nom. illeg., non Craib (1911). – Argostemma stellare Ridl., J. Bot. 65: 31 (1927). –
TYPE: Indonesia, Sumatra, Berastagi, hill forest, 8 February 1921, H.N. Ridley s.n.
(lectotype, designated here, K [K000760148]).
149. Argostemma stipulaceum Ridl., J. Fed. Malay States Mus. 10(4): 139 (1920), as
‘stipulacea’. – TYPE: Peninsular Malaysia, Kelantan, Kwala Lebir, 14 February 1917,
H.N. Ridley s.n. (lectotype K [K000172876], designated by Sridith (1999), or possibly
holotype).
335
The Gentinales taxa of H.N. Ridley
= Argostemma propinquum Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 53 (1911);
Sridith, Thai Forest Bull., Bot. 27: 89 (1999).
150. Argostemma succulentum Ridl., J. Bot. 65: 28 (1927). – TYPE: Burma,
Moulmein, limestone rocks, 1839, C.S.P. Parish 126 (lectotype K [K000031023],
designated here, or possibly holotype).
151. Argostemma tenue Ridl., J. Fed. Malay States Mus. 6: 39 (1915). – TYPE:
Peninsular Malaysia, Negeri Sembilan, Gunung Tampin, 16 September 1913, H.C.
Robinson s.n. (lectotype K [K000172891], designated here, or possibly holotype).
152. Argostemma trichanthum Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 77 (1918).
– TYPE: Peninsular Malaysia, Selangor, Ulu Langat, s.dat., C.B. Kloss s.n. (lectotype
K [K000172878], designated here, or possibly holotype).
153. Argostemma triorum Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 67 (1923).
– TYPE: Sumatra, Berastagi woods, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000760277], designated here, or possibly holotype).
154. Argostemma velutinum Ridl., J. Bot. 65: 35 (1927). – TYPE: Borneo, Sarawak,
Serapi, 2800 ft, s.dat., G.D. Haviland 684 [c.l.g.d.] (lectotype K [K000760160],
designated by Bremer (1989)). Additional syntypes: Borneo, Sarawak, Matang,
August 1908, H.N. Ridley 12304 (BM, K); Sarawak, Upper Baram, Mount Marus,
Upper Baram, J.C. Moulton 165 (not traced).
= Argostemma moultonii Ridl., J. Bot. 65: 36 (1927); Bremer, Ann. Missouri Bot.
Gard. 76: 30 (1989).
155. Argostemma viscidum Ridl., J. Fed. Malay States Mus. 4: 27 (1909). – TYPE:
Peninsular Malaysia, Pahang, Telom Cascade, November 1908, H.N. Ridley 13609
(lectotype K [K000172886], designated here, or possibly holotype).
156. Asemanthia lobbii Ridl., Bull. Misc. Inform. Kew 1939: 600 (1940). –TYPE:
Borneo, Sarawak, 1857, T. Lobb s.n. (lectotype K [K000740924], designated here;
isolectotype K [K000740925]).
This is a species of Mussaenda and a combination will soon be published by Govaerts
(pers. comm.).
336 Gard. Bull. Singapore 70 (2) 2018
157. Becheria parviora Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 20 (1912). –
TYPE: Peninsular Malaysia, Pahang, Taka, Tahan, 1893, Mat s.n. (lectotype SING
[SING0059286], designated here). Additional syntypes: Peninsular Malaysia, Pahang,
Kwala Tembeling, May 1891, H.N. Ridley 8258 (SING [SING0059287]); Perak,
Tapah, June 1890, H.N. Ridley s.n. (SING [SING0058822] = Psychotria montana
Blume). {Indonesia} South of Singapore, St. Barbe isle, Lauglasse (not traced).
= Ixora umbellata Valeton ex Koord. & Valeton, Meded. Lands Plantentuin 59: 162
(1902).
158. Borreria parviceps Ridl., J. Straits Branch Roy. Asiat. Soc. 84: 299 (1921); Ridley,
Fl. Malay Penins. 2: 176 (1923). – Spermacoce parviceps (Ridl.) I.M.Turner, Novon
6: 222 (1996). – TYPE: Peninsular Malaysia, Negeri Sembilan, Bukit Tangga Pass,
23 December 1920, H.N. Ridley s.n. (lectotype SING [SING0058046], designated by
Turner (1996); isolectotypes K [K000772203], SING [SING0058047]).
159. Borreria pilulifera Ridl., J. Straits Branch Roy. Asiat. Soc. 84: 299 (1921), Fl.
Malay. Penins. 2: 176 (1923). – Spermacoce pilulifera (Ridl.) I.M.Turner, Novon 6:
222 (1996). – TYPE: Peninsular Malaysia, Selangor, Klang Gates, 1 January 1921,
H.N. Ridley s.n. (holotype K [K000772204]).
160. Canthium aciculatum Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 57 (1911);
Wong, Tree Fl. Malaya 4: 341 (1989). – TYPE: Peninsular Malaysia, Perak, Lenggong,
August 1909, H.N. Ridley 14733 (lectotype SING [SING0058926], designated here;
possible isolectotypes BM [BM001190930], K [K000763655]). Additional syntype:
Peninsular Malaysia, Kedah, Gunong Geriang, February 1910, H.N. Ridley 15085
(SING (not traced)).
The specimens in BM and K have the Ridley number 14733 but are dated July rather
than August 1909 (SING specimen). This is probably due to some error in labelling,
but it is possible they represent a separate gathering.
161. Canthium congestiorum Ridl., J. Straits Branch Roy. Asiat. Soc. 82: 185
(1920); Wong, Tree Fl. Malaya 4: 342 (1989). – TYPE: Peninsular Malaysia, Johore,
Seduah, August 1908, H.N. Ridley 13468 (lectotype K [K000763652), designated
here; isolectotype BM [BM001190931]).
162. Canthium depressinerve Ridl., J. Fed. Malay States Mus. 10: 141 (1920); Wong,
Tree Fl. Malaya 4: 341 (1989). – TYPE: Peninsular Malaysia, Kelantan, Chaning,
February 1917, H.N. Ridley s.n. (lectotype K [K001129443], designated here, or
possibly holotype).
337
The Gentinales taxa of H.N. Ridley
163. Canthium hirtellum Ridl., J. Bot. 62: 297 (1924); Wong, Tree Fl. Malaya 4: 342
(1989). – TYPE: Peninsular Malaysia, Negri Sembilan, Gunong Angsi, 23 May 1923,
Md Nur SFN 11664 (lectotype K [K000763654], designated here; isolectotype KEP
[KEP18174]).
The type collection was cited as 11669 in the protologue but this was evidently a
printing error.
164. Canthium scabridum Ridl., J. Bot. 62: 297 (1924); Wong, Tree Fl. Malaya 4:
342 (1989). – TYPE: Peninsular Malaysia, Johore, Kluang, 18 November 1922, R.E.
Holttum SFN 9433 (lectotype K [K000763653], designated here; isolectotypes SING
[SING0059272, SING0059273]).
165. Canthium trachystyle Ridl., J. Fed. Malay States Mus. 10: 95 (1920). – TYPE:
Thailand, Tapli, s.dat., C.B. Kloss 6760 (lectotype K [K000763648], designated here,
or possibly holotype).
= Canthium angustifolium Roxb., Fl. Ind. 2: 169 (1824).
166. Canthium velutinum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 19 (1912). –
Perakanthus velutinus (Ridl.) Robyns ex Ridl., Fl. Malay Penins. 5: 317 (1925);
Wong & Puff, Sandakania 16: 42 (2005). – TYPE: Peninsular Malaysia, Perak, Gunong
Keledang, 1898, H.N. Ridley 9719 (lectotype SING [SING0058915], designated here,
or possibly holotype).
167. Cephaelis albiora Ridl., J. Fed. Malay States Mus. 6: 155 (1915). – Psychotria
tahanica I.M.Turner, Asian J. Trop. Biol. 1(2): 26 (1995). – Chassalia tahanica
(I.M.Turner) A.P.Davis in Ruhsam et al., Bot. J. Linn. Soc. 157: 119 (2008). – TYPE:
Peninsular Malaysia, Pahang, Gunong Tahan, July 1911, H.N. Ridley 16075 (lectotype
K [K000761832], designated here; isolectotype BM [BM001118901]).
168. Cephaelis angustifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 88 (1918). –
Psychotria ophirensis I.M.Turner, Asian J. Trop. Biol. 1(2): 26 (1995). – Chassalia
angustifolia (Ridl.) A.P.Davis in Ruhsam et al., Bot. J. Linn. Soc. 157: 118 (2008).
– TYPE: Peninsular Malaysia, Johore, Gunong Panti, 7 Dec 1892, H.N. Ridley s.n.
(lectotype K [K000761836], designated here). Additional syntypes: Peninsular
Malaysia, Malacca, Mount Ophir, April 1888, R.W. Hullett 790 (BM); ibid., August
1867–1868, A.C. Maingay 2595 (K), ibid., 1867, A.C. Maingay 2595 (K); ibid., W.
Grifth s.n. (K); Negri Sembilan, Gunong Tampin, 20 September 1913, H.C. Robinson
s.n. (K) [referred to in the protologue as a form with broader leaves].
338 Gard. Bull. Singapore 70 (2) 2018
169. Cephaelis cuneata Korth. var. elliptica Ridl., J. Fed. Malay States Mus. 4:
39 (1909). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908, H.N.
Ridley 13636 (lectotype K [K000761831], designated here; isolectotype SING
[SING0058017]).
= Chassalia elliptica (Ridl.) A.P.Davis, Bot. J. Linn. Soc. 157: 119 (2008).
170. Cephaelis elliptica Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 89 (1918). –
Chassalia elliptica (Ridl.) A.P.Davis, Bot. J. Linn. Soc. 157: 119 (2008). – TYPE:
Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley 13636 (lectotype
K [K000761831], designated here; isolectotype SING [SING0058017]).
171. Cephaelis elongata Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 90 (1918). –
Psychotria porcata I.M.Turner, Asian J. Trop. Biol. 1(2): 27 (1995). – Chassalia
porcata (I.M.Turner) A.P.Davis in Ruhsam, Bot. J. Linn. Soc. 157: 118 (2008). –
TYPE: Peninsular Malaysia, Selangor, Semangkok Pass, April 1911, H.N. Ridley s.n.
(lectotype K [K000761834], designated here; isolectotype BM).
= Chassalia bracteata Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 23 (1912).
The types of Cephaelis elongata Ridl. and Chassalia bracteata Ridl. were both
collected from Semangkok (Fraser’s Hill) and are very similar. Cephaelis elongata is
therefore here reduced to synonymy of the older name, Chassalia bracteata.
172. Cephaelis melanocarpa Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 298 (1922),
Fl. Malay Penins. 2: 144 (1923). – Psychotria albopedunculata I.M.Turner, Asian J.
Trop. Biol. 1(2): 27 (1995). – Chassalia melanocarpa (Ridl.) A.P.Davis in Ruhsam et
al., Bot. J. Linn. Soc. 157: 118 (2008). – TYPE: Peninsular Malaysia, Negeri Sembilan,
Bukit Tangga, 20 December 1920, H.N. Ridley s.n. (lectotype K [K000761837],
designated here, or possibly holotype).
173. Cephaelis pauciora Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 71 (1923). –
Chassalia pauciora (Ridl.) I.M.Turner, comb. nov. – TYPE: Sumatra, Berastagi,
13 February 1921, H.N. Ridley s.n. (lectotype K [K001129557], designated here, or
possibly holotype).
Ridley’s plant clearly belongs with the group of Chassalia species with involucrate
inorescences, so a new combination is proposed here.
174. Cephaelis singapurensis Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 89 (1918).
– Psychotria singapurensis (Ridl.) I.M.Turner, Asian J. Trop. Biol. 1(2): 27 (1995).
339
The Gentinales taxa of H.N. Ridley
– Chassalia singapurensis (Ridl.) A.P.Davis in Ruhsam et al., Bot. J. Linn. Soc.
157: 119 (2008). – TYPE: Singapore, Bajau, 1892, H.N. Ridley 4966 (lectotype
K [K000761838], designated here; isolectotypes BM [BM001118900], SING
[SING0064264]). Additional syntypes: Peninsular Malaysia, Johore, Gunong Pulai,
December 1904, H.N. Ridley 12205 (K); Johore, Gunong Pantai, Unknown collector
(not traced).
175. Cephaelis subcoriacea Ridl., J. Fed. Malay States Mus. 5: 36 (1914). – Psychotria
subcoriacea (Ridl.) I.M.Turner, Asian J. Trop. Biol. 1(2): 27 (1995). – Chassalia
subcoriacea (Ridl.) A.P.Davis in Ruhsam et al., Bot. J. Linn. Soc. 157: 119 (2008). –
TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang Lebah, 16 January 1913,
H.C. Robinson s.n. (lectotype K [K000761833], designated here; isolectotypes BM
[BM001118899], SING [SING0058827]).
176. Cephaelis triceps Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 89 (1918). –
Psychotria triceps (Ridl.) I.M.Turner, Asian J. Trop. Biol. 1(2): 27 (1995). – Chassalia
triceps (Ridl.) A.P.Davis in Ruhsam et al., Bot. J. Linn. Soc. 157: 118 (2008), as
‘tricepa’. – TYPE: Peninsular Malaysia, Selangor, Sempang, April 1911, H.N. Ridley
15667 (lectotype K [K000761835], designated here, or possibly holotype).
177. Chassalia bracteata Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 23 (1912). –
TYPE: Peninsular Malaysia, Selangor, Semangko, August 1904, H.N. Ridley 12074
(lectotype K [K000761830], designated here, or possibly holotype).
The protologue gives the number of the collection on which this species is based as
12073. This number is evidently a printing error as it is a specimen of Pittosporum
ferrugineum W.T.Aiton at SING.
178. Chassalia minor Ridl., J. Fed. Malay States Mus. 4: 39 (1909); Wong, Tree Fl.
Malaya 4: 344 (1989). – TYPE: Peninsular Malaysia, Pahang, Gunong Beremban,
[November] 1908, H.N. Ridley 13683 (lectotype SING [SING0180010], designated
here; isolectotypes BM [BM001190962], K [K000761829]).
179. Chassalia propinqua Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 70 (1923), as
‘Chasalia’. – TYPE: Sumatra, Brastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000761884], designated here). Additonal syntype: Sumatra, Brastagi, 11 February
1921, H.N. Ridley s.n. (K [K000761883]).
340 Gard. Bull. Singapore 70 (2) 2018
180. Chassalia pubescens Ridl., J. Straits Branch Roy. Asiat. Soc. 54: 46 (1910);
Wong, Tree Fl. Malaya 4: 344 (1989). – TYPE: Peninsular Malaysia, Johore, Sedenah,
August 1908, H.N. Ridley 13531 (lectotype SING [SING0209189], designated here;
isolectotypes BM [BM001190961], K [K000761828]).
181. Coelospermum biovulatum C.B.Clarke ex Ridl., J. Straits Branch Roy. Asiat.
Soc. 79: 87 (1918). – TYPE: Peninsular Malaysia, Malacca, 31 October 1867/8,
A.C. Maingay 3053 (lectotype K [K000763733], designated by Johannson (1988), or
possibly holotype).
= Coelospermum truncatum (Wall.) Baill. ex K.Schum. in Engler & Prantl., Nat.
Panzenfam. 4(4): 136 (1891); Johansson, Blumea 33: 277 (1988).
182. Coffea malayana Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 19 (1912). –
Prismatomeris khoonmengiana Y.W.Low, Pl. Ecol. Evol. 148: 135 (2015). – TYPE:
Peninsular Malaysia, Negri Sembilan, Bukit Tampin, May 1894, J.S. Goodenough 1864
(lectotype SING [SING0166263], designated by Low (2015)). Additional syntypes:
Peninsular Malaysia, Johore, Gunong Panti, 9 December 1892, H.N. Ridley 4177
(SING [SING0166299]); Selangor, Bukit Hitam, May 1894, H.N. Ridley 7413 (SING
[SING0166295]); Kuala Lumpur, s.dat., H.N. Ridley 10217 (SING [SING0166298]);
Negri Sembilan, Gunung Berembun, 27 June 1885, ‘Cantley’ Alvins 3033 (SING
[SING0166296]); Bukit Kandang, 25 March 1885, ‘Cantley’ Alvins 1022 (SING
[SING0166297]).
183. Coffea merguensis Ridl., J. Fed. Malay States Mus. 10: 96 (1920). – Paracoffea
merguensis (Ridl.) J.-F.Leroy, Adansonia sér. 2, 11(4): 604 (1972). – Psilanthus
merguensis (Ridl.) J.-F.Leroy, Ass. Sci. Internat. Café, Colloque 9: 482 (1980). – TYPE:
Thailand, Tapli, March 1919, C.B. Kloss 6806 (lectotype K [K000466521], designated
by Leroy (1972); isolectotype SING [SING0166294]). Additional syntypes: Burma,
Tenasserim, Helfer 3016 (K); Mergui, W. Grifth 578 (K [K000466525]).
184. Coffea viridiora Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 82 (1918). –
Nostolachma viridiora (Ridl.) J.-F.Leroy ex A.P.Davis in Ruhsam et al., Bot. J. Linn.
Soc. 157: 123 (2008). – TYPE: Peninsular Malaysia, Selangor, Batu Caves, 4 March
1915, H.N. Ridley s.n. (lectotype K [K000763090], designated here). Additional
syntype: Peninsular Malaysia, Perak, Waterloo, December 1887, C. Curtis 1304 (K
[K000763089]).
185. Coptosapelta parviora Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 76 (1918).
– TYPE: Singapore, Bukit Timah, January 1909, H.N. Ridley 14117 (lectotype K
341
The Gentinales taxa of H.N. Ridley
[K000760081), designated here; isolectotype SING [SING0058045]). Additional
syntypes: Peninsular Malaysia, Penang, s.dat., C. Curtis s.n. (K [K000760082]);
Borneo, Sarawak, O. Beccari P.B. 2518 (FI-B).
186. Diplospora lasiantha Ridl., J. Fed. Malay States Mus. 6: 39 (1915); Wong, Tree
Fl. Malaya 4: 345 (1989). – TYPE: Peninsular Malaysia, Negri Sembilan, Gunung
Tampin, 20 September 1913, H.C. Robinson s.n. (lectotype K [K000763094],
designated here, or possibly holotype).
187. Diplospora minutiora Ridl., J. Fed. Malay States Mus. 10: 140 (1920). –
Hypobathrum sp. 1 of Wong, Tree Fl. Malaya 4: 346 (1989). – TYPE: Peninsular
Malaysia, Kelantan, Chaning woods, February 1917, H.N. Ridley s.n. (lectotype K
[K001129434], designated here); isolectotype SING [SING0059280]).
Wong (1989) suggested that this species belonged in Hypobathrum but did not make
a new combination for it. The rather poor type material makes it difcult to judge its
correct position.
188. Diplospora stylosa Ridl., J. Fed. Malay States Mus. 10: 94 (1920). – TYPE:
Thailand, Tasan, March 1919, C.B. Kloss 6834 (lectotype K [K000763085], designated
here; isolectotype SING [SING0059278]).
189. Gaertnera acuminata Benth. var. montana Ridl., Fl. Malay Penins. 2: 428
(1923).– TYPE: Peninsular Malaysia, Perak, Gunong Kerbau, s.dat., H.C. Robinson
s.n. (lectotype K [K000772013], designated by van Beusekom (1967)). Additional
syntypes: Peninsular Malaysia, Malacca, Mt Ophir, April 1888, R.W. Hullett 756 (K);
Pahang, Telom Ridge, H.N. Ridley (not traced); Perak, Scortechini (not traced); Kedah,
Kedah Peak, H.C. Robinson (not traced).
= Gaertnera ramosa Ridl., J. Linn. Soc., Bot. 38: 317 (1908).
190. Gaertnera brevistylis Ridl., Bull. Misc. Inform. Kew 1934: 124 (1934). –
TYPE: Borneo, British North Borneo {Sabah}, Sandakan, April 1895, C.V. Creagh
s.n. (lectotype K [K000173235], designated by van Beusekom (1967); possible
isolectotype K [K000173234]).
= Gaertnera junghuhniana Miq., Fl. Ned. Ind. 2: 383 (1857); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 626 (2009).
The second Creagh specimen at Kew does not state Sandakan as the collecting locality
or bear the collection date, but Ridley has annotated it as a type duplicate.
342 Gard. Bull. Singapore 70 (2) 2018
191. Gaertnera caudata Ridl., J. Fed. Malay States Mus. 6: 51 (1915), as ‘caudate’.
– TYPE: Peninsular Malaysia, Perak, Gunong Kerbau, 4500 ft, 14 March 1913, H.C.
Robinson s.n. (lectotype K [K001067664], designated by van Beusekom (1967)).
= Gaertnera ramosa Ridl., J. Linn. Soc., Bot. 38: 317 (1908); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 648 (2009).
192. Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009). – Gaertnera oblancelata King
& Gamble var. diversifolia (Ridl.) Beusekom, Blumea 15: 384 (1967). – TYPE:
Peninsular Malaysia, Selangor, Hulu Langat, Bukit Hitam, 1891, H.J. Kelsall 1995,
(lectotype SING [SING0059222], designated by van Beusekom (1967)). Additional
syntype: Peninsular Malaysia, Selangor, Hulu Langat, Bukit Hitam, May 1896, H.N.
Ridley 7429 (SING [SING0240238]).
193. Gaertnera intermedia Ridl., J. Fed. Malay States Mus. 6: 163 (1915). – TYPE:
Peninsular Malaysia, Selangor, Hulu Semangko, August 1904, H.N. Ridley 12080
(lectotype SING [SING0059224], designated by van Beusekom (1967)).
= Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009).
194. Gaertnera lanceolata Ridl., J. Fed. Malay States Mus. 6: 162 (1915), nom. illeg.,
non Bouton ex DC. (1845). – TYPE: Peninsular Malaysia, Pahang, Gunong Tahan,
Wray’s Camp at 3300 ft, 1 June 1905, L. Wray & H.C. Robinson 5343 (lectotype
SING [SING0059225], designated by van Beusekom (1967); isolectotypes BM
[BM000055220], K [K000772020]). Additional syntypes: Peninsular Malaysia,
Pahang, Gunong Tahan, July 1911, H.N. Ridley 16255 (K [K000173230], SING
[SING0240236]).
= Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009).
195. Gaertnera latifolia Ridl., J. Bot. 62: 299 (1924). – TYPE: Peninsular Malaysia,
Pahang, Fraser {Fraser’s} Hill, 16–30 September 1922, I.H. Burkill & R.E. Holttum
8608 (lectotype K (as ‘8606’) [K000173231], designated here; isolectotype SING
[SING0059226]).
= Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009).
The Kew duplicate of the type gathering is labelled 8606 in error, and was cited as
such by Ridley.
343
The Gentinales taxa of H.N. Ridley
196. Gaertnera ovata Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 301 (1922). – TYPE:
Peninsular Malaysia, Selangor, Semangkok Pass, 22 January 1921, H.N. Ridley
s.n. (lectotype K [K000173244], designated by van Beusekom (1967), or possibly
holotype).
= Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009).
197. Gaertnera oxyphylla Benth. var. angustifolia Ridl., Fl. Malay Penins. 2: 428
(1923). – TYPE: Peninsular Malaysia, Kedah, Kedah Peak, Padang To Seh {Padang
Tok Sheikh}, 3000 ft, 29 November 1915, H.C. Robinson & C.B. Kloss 5963 (lectotype
K [K001067663], designated here; isolectotype SING [SING0059227]).
= Gaertnera junghuhniana Miq., Fl. Ned. Ind. 2: 383 (1857); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 626 (2009).
198. Gaertnera pedicellata Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 99 (1918). –
TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang, 6 February 1913, H.C.
Robinson s.n. (lectotype K [K000772014], designated by van Beusekom (1967);
isolectotype SING [SING0059228]).
= Gaertnera ramosa Ridl., J. Linn. Soc., Bot. 38: 317 (1908); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 648 (2009).
199. Gaertnera ramosa Ridl., J. Linn. Soc., Bot. 38: 317 (1908); Malcomber &
Taylor, Ann. Missouri Bot. Gard. 96: 648 (2009). – TYPE: Peninsular Malaysia,
Pahang, Gunong Tahan, ?5000–6000 ft, 3 July 1905, L. Wray & H.C. Robinson 5458
(lectotype L [L0537712], designated by van Beusekom (1967); isolectotype BM
[BM000055231]).
A sheet in the Singapore Herbarium was referred to as holotype (an effective
lectotypication) by van Beusekom. This specimen seems to have subsequently been
donated to the Leiden Herbarium. The other sheet in SING [SING0059229] that van
Beusekom referred to as an isotype is actually labelled with a different number (5488).
Whether this represents a separate gathering or a slip of the pen is uncertain.
200. Gaertnera rigida Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 301 (1922). – TYPE:
Peninsular Malaysia, Negri Sembilan, Bukit Tangga, December 1920, H.N. Ridley s.n.
(lectotype K [K000173232], designated by van Beusekom (1967); isolectotype SING
[SING0240237]).
= Gaertnera diversifolia Ridl., J. Fed. Malay States Mus. 6: 163 (1915); Malcomber
& Taylor, Ann. Missouri Bot. Gard. 96: 614 (2009).
344 Gard. Bull. Singapore 70 (2) 2018
201. Gaertnera sessiliora Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 99 (1918).
– TYPE: Peninsular Malaysia, Negri Sembilan, Gunong Tampin, 15 September
1913, H.C. Robinson s.n. (lectotype K [K000173242], designated by van Beusekom
(1967)). Additional syntype: Peninsular Malaysia, Negri Sembilan, Gunong Tampin,
22 September 1913, H.C. Robinson s.n. (K [K000173243]).
= Gaertnera ramosa Ridl., J. Linn. Soc., Bot. 38: 317 (1908); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 648 (2009).
202. Gaertnera violascens Ridl., J. Fed. Malay States Mus. 6: 164 (1915). – Chassalia
violascens (Ridl.) I.M.Turner, comb. nov. – TYPE: Peninsular Malaysia, Pahang,
Gunong Tahan, Wray’s Camp, July 1911, H.N. Ridley 16285 (lectotype K [K001129561],
designated here; isolectotypes BM [BM000055245], SING [SING0058924]).
This species was excluded from Gaertnera Retz. by van Beusekom (1967) and referred
to Psychotria. We believe that Chassalia Comm. ex Poir is actually the correct generic
placement. The type collection is similar to Chassalia bracteata Ridl., but the longer
inorescences and at-faced rather than adaxially concave pyrenes distinguish the
two. A new combination in Chassalia for Gaertnera violascens is, therefore, proposed
here.
203. Gardenia campanula Ridl., Trans. Linn. Soc. London, Bot. 3(9): 310 (1893).
– Kailarsenia campanula (Ridl.) Tirveng., Nordic J. Bot. 3: 464 (1983). – TYPE:
Peninsular Malaysia, Pahang, Mahang, May 1890, H.N. Ridley 1316 (lectotype SING
[SING0058426], designated here, or possibly holotype).
204. Gardenia costulata Ridl., J. Bot. 72: 274 (1934). – TYPE: Borneo, Sarawak,
1865–1868, O. Beccari P.B. 1986 (lectotype K [K000173267], designated here;
isolectotypes BM [BM001124570], FI-B!, M [M0187663], P [P008432756]).
205. Gardenia didymocarpa Ridl., J. Fed. Malay States Mus. 4: 32 (1909), as
‘didymocarpus’. – Acranthera didymocarpa (Ridl.) K.M.Wong, Gard. Bull. Singapore
35: 30 (1982). – TYPE: Peninsular Malaysia, Selangor, Ginting Bidai and Bukit Kutu,
May–June 1890, H.N. Ridley 7573 (lectotype SING [SING0062183], designated
by Wong (1982); isolectotypes K [K000740912, plus one sheet with no barcode]).
Additional syntype: Peninsular Malaysia, 15th mile Pahang Track, 1897, H.N. Ridley
s.n. (SING [SING0058425]).
206. Gardenia elata Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 81 (1918); Low
& Wong, Gard. Bull. Singapore 61: 107 (2009). – Gardenia tubifera Wall. f. elata
(Ridl.) K.M.Wong, Gard. Bull. Singapore 35: 22 (1982). – TYPE: Singapore, Bukit
345
The Gentinales taxa of H.N. Ridley
Timah, 1902, H.N. Ridley 11332 (lectotype K [K000173283], designated by Low &
Wong (2009); isolectotype (SING [SING0058423]). Additional syntypes: Peninsular
Malaysia, Perak, Selama, 1894, L. Wray 4266 (K [K000763036]); Borneo, Sarawak,
Baram, C. Hose 229 (L).
207. Gardenia longituba Ridl., J. Bot. 72: 274 (1934). – TYPE: Borneo, British North
Borneo {Sabah}, Kudat, July 1885, M. Fraser 164 (lectotype K [K000173261],
designated here, or possibly holotype).
= Gardenia elata Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 81 (1918); Low & Wong,
Gard. Bull. Singapore 61: 107 (2009).
208. Gardenia pulchella Ridl., J. Fed. Malay States Mus. 4: 31 (1909), nom. illeg.,
non G.Don (1824). – Acranthera pulchella K.M.Wong, Gard. Bull., Singapore 35: 29
(1982). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley
13606 (lectotype SING [SING0062185], designated by Wong (1982); isolectotypes K
[K000740910, K000740911]).
= Gardenia virescens Ridl. (see below)
209. Gardenia virescens Ridl., J. Fed. Malay States Mus. 4: 32 (1909). – TYPE:
Peninsular Malaysia, Perak, Taiping Hills, Cottage, 5000 ft, November 1911, H.N.
Ridley s.n. (lectotype SING [SING0058424], designated by Wong (1982)).
Wong (1982) synonymised Gardenia virescens Ridl. and Gardenia pulchella Ridl.
under the name Acranthera pulchella, which he treated as a new combination.
However, as Gardenia pulchella Ridl. is a later homonym, Gardenia virescens Ridl.
has priority. A new combination will soon be published in Acranthera by Govaerts
(pers. comm.).
210. Geophila matthewii Ridl., J. Bot. 72: 275 (1934), as ‘Matthewi’. – TYPE:
Sumatra, Asahan, Boenoet, H.S. Yates 2042 (lectotype K [K001129399], designated
here, or possibly holotype).
= Mussaenda uniora Wall. ex G.Don, Gen. Hist. 3: 491 (1834); Chantaranothai,
Thai Forest Bull., Bot. 43: 60 (2015).
Geophila matthewii Ridl. is a new synonym of Mussaenda uniora Wall. ex G.Don.
211. Geophila melanocarpa Ridl., Trans. Linn. Soc. London, Bot. 3(9): 313, t. 62
(1893). – TYPE: Peninsular Malaysia, Pahang, Kota Glanggi, 1891, H.N. Ridley 2232
(lectotype SING [SING0240239], designated here, or possibly holotype).
346 Gard. Bull. Singapore 70 (2) 2018
212. Guettarda leai Ridl., J. Linn. Soc., Bot. 37: 40 (1890). – TYPE: Brazil,
Fernando Noronha, Sapate, 20 August 1887, Ridley, Lea & Ramage 88 (lectotype BM
[BM001008940], designated here; isolectotypes K [K000003585, K000003584]).
213. Hedyotis anachoreta Ridl., Bull. Misc. Inform. Kew 1939: 612 (1940). –
TYPE: Borneo, Sarawak, Mt Matang, July 1903, H.N. Ridley 11745 (lectotype K
[K000760322], designated here; isolectotype SING [SING0058321]).
214. Hedyotis everettii Ridl., Bull. Misc. Inform. Kew 1939: 611 (1940). – TYPE:
Celebes {Sulawesi}, Bonthain Peak 2100–3000 m, October 1895, A.H. Everett 36
(lectotype K [K000760320], designated here; isolectotype SING [SING0058322]).
215. Hedyotis exuosa Ridl., J. Straits Branch Roy. Asiat. Soc. 35: 15 (1910). – TYPE:
Peninsular Malaysia, Malacca, Mt Ophir, April 1888, R.W. Hullett 766 (lectotype K
[K000760359], designated here). Additional syntypes: Peninsular Malaysia, Gunong
Ledang, June 1892, H.N. Ridley 3212 (SING [SING0209184]); Peninsular Malaysia,
Gunong Mering, 9 June 1892, H.N. Ridley 3212 (SING [SING0209185]).
216. Hedyotis globiceps Ridl., Fl. Malay Penins. 2: 47 (1923). – TYPE: Peninsular
Malaysia, Langkawi Islands, Burau, November 1916, H.C. Robinson 6248 (lectotype
K [K000760529], designated here, or possibly holotype).
217. Hedyotis hirta Ridl., Bull. Misc. Inform. Kew 1926: 472 (1926); Fukuoka, S. E.
Asian Stud. 8: 314 (1970). – TYPE: Thailand, Pungah [Phangnga], 6 December 1918,
Haniff & Nur 3871 (lectotype SING [SING0058318], designated by Fukuoka (1970),
or possibly holotype).
218. Hedyotis moultonii Ridl., Bull. Misc. Inform. Kew 1939: 612 (1940). – TYPE:
Borneo, Sarawak, summit of Mt Murud, 1 December 1914, J.C. Moulton 119 (lectotype
K [K000760306], designated here; isolectotype SING [SING0058320]).
219. Hedyotis pachycarpa Ridl., J. Fed. Malay States Mus. 10: 140 (1920); Fukuoka,
S. E. Asian Stud. 8 (1970) 314. – Exallage pachycarpa (Ridl.) Bremek., Verh. Kon.
Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2): 142 (1952). – TYPE: Peninsular
Malaysia, Kelantan, Glam Woods near Kota Bahru, February 1917, H.N. Ridley s.n.
(lectotype K [K000770015], designated here). Additional syntype: Indo-China, Chiao
Shan, March 1877, L. Pierre 2032 (K).
347
The Gentinales taxa of H.N. Ridley
220. Hedyotis patens Ridl., J. Linn. Soc., Bot. 38: 311 (1908). – TYPE: Peninsular
Malaysia, Pahang, Gunong Tahan, 5000–6000 ft, 6 July 1905, L. Wray & H.C.
Robinson 5475 (lectotype BM [BM000945137], designated here; isolectotypes K
[K000760371], SING [SING0209183]). Additional syntypes: Peninsular Malaysia,
Pahang, Gunong Tahan, 4000–5000 ft, 3 June 1905, L. Wray & H.C. Robinson 5393
(BM [BM000945136], SING [SING0209182]);
221. Hedyotis pinaster Ridl., Bull. Misc. Inform. Kew 1939: 611 (1940). – TYPE:
Borneo, Sarawak, Upper Baram, Lio Matu, 16 October 1920, J.C. Moulton SFN 6734
(lectotype K [K000770007], designated here; isolectotype SING [SING0058319]).
222. Hedyotis resupinata Ridl., Bull. Misc. Inform. Kew 1926: 67 (1926). – TYPE:
Sumatra, Siberut, 17 September 1924, C.B. Kloss 14494 (lectotype K [K000770006],
designated here; isolectotype SING [SING0214034]).
223. Hedyotis rivalis Ridl., J. Fed. Malay States Mus. 6: 153 (1915). – TYPE:
Peninsular Malaysia, Pahang, Jeram Ampai, July 1911, H.N. Ridley 16341 (lectotype
K [K000760531], designated here; isolectotype SING [SING0209181]).
224. Hedyotis stelligera Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 109 (1911). –
TYPE: Thailand, Setul [Satun], heaths, March 1910, H.N. Ridley 15100 (lectotype K
[K000760530], designated here, or possibly holotype).
225. Ixora aurorea Ridl., J. Bot. 72: 251 (1934). – TYPE: Peninsular Malaysia,
Pahang, Kuala Tekam, 8 July 1917, I.H.N. Evans s.n. (lectotype K [K000763206],
designated here, or possibly holotype).
The pieces to the left and bottom, mounted on the same sheet, represent a different
gathering, Kota Tongkat, Pahang, July 1917, Evans s.n. (K [K000763281]), the locality
not mentioned in Ridley’s protologue. This plant is included in Ixora pseudojavanica
Bremek., Bull. Jard. Bot. Buitenzorg ser. III, 14: 252 (1937), although ostensibly the
same as K000763206. Study of Bremekamp’s type and materials is needed before
considering synonymy.
226. Ixora aurorea Ridl. var. major Ridl., J. Bot. 72: 251 (1934). – TYPE: Peninsular
Malaysia, Johore, Sungei Pelepah, 5 March 1928, Md. Nur SFN 20007 (lectotype K
[K001096589], designated here; isolectotype SING [SING0059237]).
= Ixora lobbii Loudon, Encycl. Pl., new ed., Suppl. 2: 1543 (1855); Wong, Tree Fl.
Malaya 4: 361 (1989).
348 Gard. Bull. Singapore 70 (2) 2018
227. Ixora candida Ridl., J. Fed. Malay States Mus. 10: 14 (1920). – TYPE: Peninsular
Malaysia, Langkawi Islands, Dayong Bunting, November 1916, H.C. Robinson
6219 (lectotype, designated here, K [K000763274]). Additional syntype: Peninsular
Thailand, Terutau Island, Telok Wau, 15 November 1916, H.C. Robinson s.n. (SING
[SING0212337]) .
= Ixora pendula Jack, Malayan Misc. 1(5): 11 (1820); Wong, Tree Fl. Malaya 4: 359
(1989).
228. Ixora celebica Ridl. ex Bremek., Bull. Jard. Bot. Buitenzorg ser. III, 14: 331
(1937). – TYPE: Celebes {Sulawesi}, Southeast Celebes, Kendari [Lepo-Lepo] 1874,
O. Beccari s.n. (holotype K [K000763362]).
The protologue mentions Lepo-Lepo, a name which is only on one of the three Beccari
s.n. sheets from 1874 from Celebes, Kendari. The sheet annotated Lepo-Lepo at K is
therefore considered the holotype. The other two sheets at K are [K000763360] and
[K000763361].
229. Ixora clerodendron Ridl., Trans. Linn. Soc. London, Bot. 3(9): 311 (1893); Wong,
Tree Fl. Malaya 4: 358 (1989). – TYPE: Peninsular Malaysia, Kwala Tembeling,
August 1891, H.N. Ridley s.n. (lectotype SING [SING0059155], designated here).
Additional syntypes: Peninsular Malaysia, Pahang, Tanjong Antan, July 1891, H.N.
Ridley s.n. (SING [SING0212336]); Pulau Chengei, 1891, H.N. Ridley s.n. (SING
[SING0212335]).
230. Ixora coriacea R.Br. ex Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 83 (1918).
– TYPE: Singapore, October 1822, N. Wallich s.n. [EIC 6151] (lectotype K-W
[K001123175], designated here). Additional syntype: Peninsular Malaysia, Penang,
G. Porter s.n. [EIC 6153] (K-W [K001123179]).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
231. Ixora crassifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 83 (1918), nom.
illeg., non Merr. (1910). – Ixora ridleyi Merr., Gard. Bull. Straits Settlem. 8: 132
(1935). – TYPE: Peninsular Malaysia, Malacca, Ayer Pannas, heavy jungle, s.dat., W.
Grifth s.n. [HEIC 2999] (lectotype K [K000763271], designated here).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
349
The Gentinales taxa of H.N. Ridley
232. Ixora curtisii Ridl., J. Bot. 72: 254 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 288 (1937). – TYPE: Sumatra, s.dat., C. Curtis s.n. (lectotype
K [K000763298], designated here, or possibly holotype).
233. Ixora cuspidata Ridl., Bull. Misc. Inform. Kew 1926: 69 (1926); Bremekamp,
Bull. Jard. Bot. Buitenzorg ser. III, 14: 239 (1937). – TYPE: Sumatra, Sipora, 27
October 1924, C.B. Kloss SFN 14798 (holotype K [K000763331]).
234. Ixora elliptica R.Br. ex Ridl., Fl. Malay Penins. 2: 98 (1923). – TYPE: Peninsular
Malaysia, Penang, G. Porter s.n. [EIC 6153] (lectotype K-W [K001123179], designated
here; isolectotype K [K000763269]).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
235. Ixora uminalis Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 84 (1918);
Bremekamp, Bull. Jard. Bot. Buitenzorg ser. III, 14: 324 (1937). – TYPE: Peninsular
Malaysia, Johore, Kota Tinggi, 11 December 1892, H.N. Ridley 4165 (lectotype
SING [SING0058821], designated here; isolectotype K [K000763272]). Additional
syntypes: Peninsular Malaysia, Malacca, W. Grifth s.n. (K [×4]); Malacca, H. Cuming
2332 (K); Malacca, 22 August 1865–1866, A.C. Maingay 1297 [Kew Distribution
Number 843] (K); Perak, Gopeng, 1883, King’s Collector 4686 (SING [SING0212338,
SING0212340]); Kedah, Langkawi, Gunong Raya, February 1911, Mohamed Haniff
15480 (SING [SING0212339]).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
236. Ixora fulgida Ridl., Bull. Misc. Inform. Kew 1934: 119 (1934 [16 May 1934]);
Ridley, J. Bot. 72: 254 (1934 [September 1934]); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 245 (1937). – TYPE: Borneo, British North Borneo {Sabah},
East Coast, s.dat., C.V. Creagh s.n. (lectotype K [K000763324], designated here, or
possibly holotype).
The K Creagh specimen has ‘type’ written in Ridley’s hand. Another specimen from
British North Borneo, East Coast, Sandakan, April 1895, Creagh s.n. (K ([K000763323])
has shorter oral tubes and leaves and is the type of Ixora fugiens Bremek. Ridley’s
protologue records “Sandakan, Port Myburgh, East Coast, Tanjong Batu, Creagh (type
in Kew Herb.)” but neither actual specimen has the locality information given in such
detail.
350 Gard. Bull. Singapore 70 (2) 2018
237. Ixora havilandii Ridl., J. Bot. 72: 255 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 327 (1937). – TYPE: Borneo, Sarawak, Kuching, 29 September
1892, G.D. Haviland 1791 (lectotype K [K000763367], designated here; isolectotypes
BM [BM001190959], SING [SING0059151]).
238. Ixora keithii Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 117 (1911). – TYPE:
Thailand, Bangtaphan, 26 April 1890, A. Keith 339 (lectotype SING [SING0240277],
designated here, or possibly holotype).
239. Ixora lancifolia Ridl., J. Bot. 72: 256 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 328 (1937). – TYPE: Borneo, Sarawak, Rejang Belaga,
November 1892, G.D.Haviland 2157 (lectotype K [K000763366], designated here;
isolectotypes BM [BM001118898, BM001118897], L [L0000593]).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
240. Ixora lancisepala Ridl., J. Bot. 72: 253 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 236 (1937). – TYPE: Borneo, Sarawak, Limbang, G.D.
Haviland 694 [c.o.c.a.] [84] (lectotype K [K000763231], designated here, or possibly
holotype).
241. Ixora marsdenii Ridl., J. Bot. 72: 255 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 318 (1937). – TYPE: Sumatra, s.dat., Marsden s.n. (lectotype
K [ex herb. Benth.] [K000763282], designated here, or possibly holotype).
242. Ixora matangensis Ridl., J. Bot. 72: 254 (1934); Bremekamp, Bull. Jard. Bot.
Buitenzorg ser. III, 14: 248 (1937). – TYPE: Borneo, Sarawak, Matang, 1600 ft, G.D.
Haviland 84 [c.l.o.n.] (holotype K [K000763320]; isotype SING [SING0209195]).
= Ixora mjoebergii Merr., Sarawak Mus. J. 3: 536 (1928); Bremekamp, Bull. Jard.
Bot. Buitenzorg ser. III, 16(3): 282 (1940).
Both sheets have the identical handwritten Haviland labels; the K sheet additionally
has ‘675’, the number referred to by Ridley in the protologue, written in a different
hand.
243. Ixora micrantha Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 21 (1912), nom. illeg.,
non Roem. & Schult. (1818). – TYPE: Peninsular Malaysia, Selangor, Sempang Mines,
April 1911, H.N. Ridley 15830 (lectotype SING [SING0058818], rst step designated
by Corner (1941), second step designated here; isolectotype K [K000763267]).
= Ixora patens Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 83 (1918).
351
The Gentinales taxa of H.N. Ridley
Corner (1941) referred to Ridley 15830 as the type of Ixora micrantha Ridl. but did
not state in which herbarium the type was deposited. A second-step lectotypication
is therefore required.
244. Ixora montana Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 82 (1918), nom.
illeg., non Lour. (1790), nec Hook.f. (1880), Schlecht. (1906). – Ixora pendula Jack
var. montana K.M.Wong, Arbor. Rubiaceae Malaya 91 (1988). – TYPE: Peninsular
Malaysia, Perak, Gunong Kerbau, 6000 ft, May 1909, Mohamed Haniff 16310
(lectotype SING [SING0059293], designated here; isolectotype K [K000763275]).
245. Ixora paludosa Kurz var. koordersii Ridl., J. Bot. 72: 252 (1934). – Ixora
koordersii (Ridl.) Bremek., Bull. Jard. Bot. Buitenzorg ser. III, 14: 293 (1937). –
TYPE: Java, Besuki Reserve between Puger and Watangan, 27 February 1898, S.H.
Koorders 30011β (lectotype BO [sheet nos. 932.147a & b – i.e. a single specimen over
two sheets], designated by Bremekamp (1937); isolectotype L [L0000589]).
Ridley did not cite any specimens when proposing a new variety for a plant from
Java to which Koorders (Boomsoorten 8: 157 (1902)) had misapplied the name Ixora
barbata Roxb. In the Kew herbarium there is a mounted envelope with photographs
of the sheets under Koorders 30011 from Herbarium Bogoriense which has been
annotated by Ridley with his varietal name. Bremekamp’s effective lectotypication
of this specimen therefore seems acceptable.
246. Ixora patens Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 83 (1918). – TYPE:
Peninsular Malaysia, Selangor, Gunong Mengkuang, 5000 ft, 19 January 1893 (but see
note below), H.C. Robinson s.n. (lectotype SING [SING0058817], rst step designated
by Corner (1941), second step designated here). Additional syntype (or possibly
isolectotype): Peninsular Malaysia, Selangor, Gunong Mengkuang, 19 January 1913,
H.C. Robinson s.n. (K [K000763268]).
Ridley’s protologue states “Gunong Mengkuang Lebar 5,000 ft (Robinson)” without
any date. Robinson was likely not collecting at that locality in 1893 as he seems to have
rst visited the Malay Peninsula in 1901 (Van Steenis-Kruseman, 1950); the dates and
notes on both SING and K sheets are in Ridley’s hand, and the date of the SING sheet
is likely wrong. It is possible that the SING and K sheets are duplicates of the same
gathering. Corner (1941) referred to the Robinson collection as type but did not state
in which herbarium the type was deposited. Therefore a second-step lectotypication
is required.
352 Gard. Bull. Singapore 70 (2) 2018
247. Ixora plumea Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 117 (1911). – TYPE:
Peninsular Malaysia, Perlis, Telor Jambu, March 1910, H.N. Ridley 15097 (lectotype
SING [SING0209196], designated here; isolectotype K [K000763214]). Additional
syntypes: Peninsular Malaysia, Perlis, Bukit Rajah Wang, March 1910, H.N. Ridley
15098 (K); Thailand, Bangtaphan, Keith (not traced).
= Ixora nigricans R.Br. ex Wight & Arn., Prodr. Fl. Ind. Orient. 1: 428 (1834);
Bremekamp, Bull. Jard. Bot. Buitenzorg ser. III, 14: 282 (1937); Wong, Tree Fl.
Malaya 4: 359 (1989).
248. Ixora pumila Ridl., J. Fed. Malay States Mus. 10: 142 (1920). – TYPE: Peninsular
Malaysia, Pahang, Gunong Senyum, 23 June 1917, I.H.N. Evans s.n. (lectotype K
[K000763227], rst step designated by Corner (1941), second step designated here).
Additional syntype (or possibly an isolectotype): Peninsular Malaysia, Pahang,
Gunong Senyum, June 1917, I.H.N. Evans 13226 (SING [SING0059283]).
= Ixora clerodendron Ridl., Trans. Linn. Soc. London, Bot. 3(9): 311 (1893); Wong,
Tree Fl. Malaya 4: 358 (1989).
Corner (1941) referred to an Evans collection as type of Ixora pumila Ridl. but did
not state in which herbarium the type was deposited. A second-step lectotypication
is therefore required.
249. Ixora robinsonii Ridl., J. Fed. Malay States Mus. 6: 154 (1915); Wong, Tree
Fl. Malaya 4: 360 (1989). – TYPE: Peninsular Malaysia, Pahang, Gunong Tahan,
29 May 1905, L. Wray & H.C. Robinson 5304 (lectotype BM [BM000832519],
rst step designated by Corner (1941), second step designated here; isolectotypes K
[K000763216], SING [SING0059282]).
Corner (1941) referred to a Wray & Robinson collection as type of Ixora robinsonii
Ridl. but did not state in which herbarium the type was deposited. A second-step
lectotypication is therefore required.
250. Ixora stenophylla Ridl., Trans. Linn. Soc. London, Bot. 3(9): 311 (1893), nom.
illeg., non (Korth.) Kuntze (1891). – Ixora salicina Ridl., J. Bot. 72: 253 (1934) – TYPE:
Pahang, rocky banks of Tahan, H.N. Ridley 2215, 1891 (lectotype K [K000763209],
designated here; isolectotypes BM, SING [SING0058819]).
= Ixora lobbii Loudon var. stenophylla Corner, Gard. Bull. Straits Settlem. 11: 216
(1941); Wong, Tree Fl. Malaya 4: 361 (1989). – TYPE: Peninsular Malaysia, Pahang,
rocky banks of Tahan, 1891, H.N. Ridley 2215 (lectotype K [K000763209], designated
here; isolectotypes BM, SING [SING0058819]).
353
The Gentinales taxa of H.N. Ridley
251. Klossia montana Ridl., J. Fed. Malay States Mus. 4: 28 (1909). – TYPE:
Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley 13911 (lectotype
SING [SING0043411], designated here; isolectotype K [K000740711]). Additional
syntypes: Peninsular Malaysia, Selangor, Ginting Bidai, 2–3000 ft, May 1895, H.N.
Ridley 7411 (K, SING [SING0209193]); Pahang Track [‘Track to Semangko Pass
from Kuala Kubu’], 1897, H.N. Ridley s.n. (SING [SING0209194]).
252. Lasianthus barbellatus Ridl., J. Straits Branch Roy. Asiat. Soc. 77: 187 (1917).
– Lasianthus trichophlebus Hemsl. subsp. barbellatus (Ridl.) H.Zhu, Acta Phytotax.
Sin. 39(2): 144 (2001). – TYPE: Peninsular Malaysia, Pahang, Pulau Tioman, Joara
{Juara} Bay, 22 June 1915, I.H. Burkill SFN 976 (lectotype SING [SING0059247],
designated here; isolectotype K [K000763901]).
= Lasianthus trichophlebus Hemsl., J. Linn. Soc., Bot. 23: 388 (1888); Zhu, Syst. &
Geogr. Pl. 72: 92 (2002); Zhu et al., Blumea 57: 90 (2012).
253. Lasianthus barbiger Ridl., J. Fed. Malay States Mus. 8(4): 52 (1917); Zhu et al.,
Blumea 57: 16 (2012). – TYPE: Sumatra, Korinchi Expedition, 1914, H.C. Robinson
& C.B. Kloss s.n. (lectotype BM [BM000945594], designated here; isolectotype K
[K000763996]).
254. Lasianthus bractescens Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 85 (1918). –
TYPE: Peninsular Malaysia, Selangor, Batu Caves, 17 February 1896, H.N. Ridley 8231
(lectotype K [K000763941], designated here; isolectotype SING [SING0240243]).
= Lasianthus cyanocarpus Jack, Trans. Linn. Soc. London 14: 125 (1823); Zhu et al.,
Blumea 57: 33 (2012).
255. Lasianthus bractescens Ridl. var. rosulatus Ridl., J. Straits Branch Roy. Asiat.
Soc. 79: 85 (1918). – TYPE: Peninsular Malaysia, Perak, Telok Pinang, 1898,
H.N. Ridley 9728 (lectotype SING [SING0240241], designated here). Additional
syntypes: Peninsular Malaysia, Perak, Lenggong, August 1909, H.N. Ridley 14340
(SING [SING0240242]); Perak, Temengoh, July 1909, H.N. Ridley s.n. (SING
[SING0240240]).
= Lasianthus cyanocarpus Jack, Trans. Linn. Soc. London 14: 125 (1823); Zhu et al.,
Blumea 57: 33 (2012).
256. Lasianthus caudiculatus Ridl., J. Fed. Malay States Mus. 6: 48 (1915), as
‘Laudiculatus’. – TYPE: Peninsular Malaysia, Perak, Gunong Kerbau, 4500 ft, 14
March 1913, H.C. Robinson s.n. (lectotype K [K000763902], designated here, or
possibly holotype).
354 Gard. Bull. Singapore 70 (2) 2018
= Lasianthus montanus King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73:
127 (1904).
Wong (1989: 373) treated Lasianthus caudiculatus as a synonym of L. montanus King
& Gamble. However, Zhu et al. (2012: 94–95) consider both names as dubious.
257. Lasianthus chryseus Ridl., J. Fed. Malay States Mus. 10: 143 (1920); Zhu et
al., Blumea 57: 28 (2012). – TYPE: Singapore, Gardens’ Jungle, February 1889, H.N.
Ridley 140 (lectotype K [K000763913], designated by Zhu et al. (2012)).
258. Lasianthus conspicuus Ridl., J. Fed. Malay States Mus. 4: 36 (1909); Zhu et al.,
Blumea 57: 30 (2012). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908,
H.N. Ridley 13628 (lectotype SING [SING0059764], designated here; isolectotypes
BM [BM001190977], K [K000763890]).
259. Lasianthus crassifolius Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 85 (1918).
– TYPE: Peninsular Malaysia, Selangor, Sempang Mines, April 1911, H.N. Ridley
15689 (lectotype K [K000763897], designated by Zhu et al. (2012)). Additional
syntype: Peninsular Malaysia, Malacca, A.C. Maingay (not traced).
= Lasianthus pergamaceus King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73:
123 (1904); Zhu et al., Blumea 57: 73 (2012).
260. Lasianthus crassinervis Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 59 (1911);
Zhu et al., Blumea 57: 31 (2012). – TYPE: Peninsular Malaysia, Perak, Temango,
July 1909, H.N. Ridley 14487 (lectotype SING [SING0059246], designated here;
isolectoptyes BM [BM001191346], K [K000763929]).
261. Lasianthus avinervius Ridl., J. Fed. Malay States Mus. 6: 154 (1915); Zhu et
al., Blumea 57: 41 (2012). – TYPE: Peninsular Malaysia, Pahang, Gunong Tahan,
July 1911, H.N. Ridley 16121 (lectotype SING [SING0059765], designated here;
isolectotype K [K000763909]).
262. Lasianthus glaber Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 59 (1911); Zhu
et al., Blumea 57: 45 (2012). – TYPE: Peninsular Malaysia, Perak, Temango, July
1909, H.N. Ridley 14490 (lectotype BM [BM001190978], designated here, or possibly
holotype).
355
The Gentinales taxa of H.N. Ridley
263. Lasianthus glaberrimus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 86 (1918).
– TYPE: Peninsular Malaysia, Selangor, Semangkok Pass, 1897, H.N. Ridley 8574
(lectotype K [K000763889], designated here). Additional syntypes: Peninsular
Malaysia, Malacca, 26 October 1865/6, A.C. Maingay 1698 (K); Penang, West Hill,
August 1886, C. Curtis 964 (K); Penang, s.dat., C. Curtis 964 (K); Kedah, Kedah
Peak, June 1893, H.N. Ridley 5549 (K).
= Paralasianthus lowianus (King & Gamble) H.Zhu, Phytotaxa 202: 276 (2015).
264. Lasianthus glaberrimus Ridl., J. Fed. Malay States Mus. 10: 98 (1920), nom. illeg.,
non L. glaberrimus Ridl. (1918), as ‘glaberrima’. – TYPE: Thailand, Tasan, [March
1919], C.B. Kloss 7033 (lectotype K [K000777019], designated here; isolectotype
SING [SING0059256]).
= Lasianthus longipedunculatus R.Parker, Bull. Misc. Inform. Kew 1925: 430 (1925);
Zhu, Acta Phytotax. Sin. 29: 134 (2001).
265. Lasianthus hirtus Ridl., J. Fed. Malay States Mus. 4: 36 (1909); Zhu et al.,
Blumea 57: 48 (2012). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908,
H.N. Ridley 14009 (lectotype SING [SING0059261], designated here; isolectotypes
BM [BM001190976], K [K000777044]).
266. Lasianthus maingayi Hook.f. var. hirtus Ridl., Fl. Malay Penins. 2: 160 (1923), as
‘hirta’. – TYPE: Peninsular Malaysia, Negeri Sembilan, Bukit Tangga, 17 December
1920, H.N. Ridley s.n. (lectotype K [K000763939], designated here, or possibly
holotype).
= Lasianthus ridleyi King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 115
(1904); Zhu et al., Blumea 57: 79 (2012).
267. Lasianthus mollis Ridl., J. Fed. Malay States Mus. 10: 144 (1920); Zhu et al.,
Blumea 57: 62 (2012). – TYPE: Peninsular Malaysia, Malacca, Tebong woods, 19
January 1917, H.N. Ridley s.n. (lectotype SING [SING0059241], designated here);
isolectotypes BM [BM001190975], K [K000763928]).
268. Lasianthus myrtifolius Ridl., J. Fed. Malay States Mus. 4: 35 (1909); Zhu
et al., Blumea 57: 64 (2012). – TYPE: Peninsular Malaysia, Pahang, Telom ridge,
November 1908, H.N. Ridley 13630 (lectotype SING [SING0059239], designated
here; isolectotypes BM, K [K000763914]).
356 Gard. Bull. Singapore 70 (2) 2018
269. Lasianthus pendulus Ridl., J. Fed. Malay States Mus. 4: 37 (1909), as ‘pendula’;
Zhu et al., Blumea 57: 73 (2012). – Litosanthes pendulus (Ridl.) Deb & M.Gangop.,
Candollea 44(1): 222 (1989). – Lasianthus capitatus Blume subsp. pendulus
(Ridl.) H.Zhu, Acta Bot. Yunnan. 22: 398 (2000). – TYPE: Peninsular Malaysia,
Pahang, Gunong Berambun, November 1908, H.N. Ridley 14007 (lectotype SING
[SING0059231], designated here; isolectotypes BM, K [K000763885]).
270. Lasianthus politus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 85 (1918); Zhu et
al., Blumea 57: 75 (2012). – TYPE: Peninsular Malaysia, Selangor, Sempang, April
1911, H.N. Ridley 15690 (lectotype K [K000763927], designated here; isolectotype
BM [BM001190974]).
271. Lasianthus robinsonii Ridl., J. Fed. Malay States Mus. 4: 37 (1909); Zhu et al.,
Blumea 57: 81 (2012). – Litosanthes robinsonii (Ridl.) Deb & M.Gangop., Candollea
44(1): 222 (1989). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908,
H.N. Ridley 13631 (lectotype SING [SING0059762], designated here; isolectotypes
BM, K [K000763884]).
272. Lasianthus salicifolius Ridl., J. Fed. Malay States Mus. 4: 35 (1909). –
TYPE: Peninsular Malaysia, Pahang, Telom Ridge, November 1900, H.N. Ridley
14008 (lectotype SING [SING0059270], designated here; possible isolectotype K
[K000763910]).
= Lasianthus angustifolius King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73:
126 (1904); Zhu et al., Blumea 57: 13 (2012).
273. Lasianthus sepalinus Ridl., Bull. Misc. Inform. Kew 1934: 120 (1934). – TYPE:
Borneo, British North Borneo {Sabah}, Bongaya, Labuk Bay, 1887, H.N. Ridley 9073
(lectotype K [K000777119], designated here, or possibly holotype).
= Lasianthus borneensis Merr., Univ. Calif. Publ. Bot. 15: 292 (1929); Zhu et al.,
Blumea 57: 19 (2012).
274. Lasianthus sordidus Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 60 (1911). –
TYPE: Peninsular Malaysia, Perak, Temango, July 1909, H.N. Ridley 14488 (lectotype
BM [BM001190973], designated here; isolectotype K [K000763925]).
= Lasianthus ellipticus Wight, Calcutta J. Nat. Hist. 6: 507 (1846); Zhu et al., Blumea
57: 35 (2012).
357
The Gentinales taxa of H.N. Ridley
275. Lasianthus stipularis Blume var. hirtus Ridl., Fl. Malay Penins 2: 153 (1923).
– TYPE: Peninsular Malaysia, Selangor, Kanching, December 1920, H.N. Ridley s.n.
(lectotype K [K000763942], designated here, or possibly holotype).
276. Lasianthus velutinus Ridl., J. Fed. Malay States Mus. 10: 144 (1920). – TYPE:
Peninsular Malaysia, Kelantan, Chaning Woods, 4 February 1917, H.N. Ridley s.n.
(lectotype SING [SING0059265], designated here; isolectotype K [K000763926]).
= Lasianthus ellipticus Wight, Calcutta J. Nat. Hist. 6: 507 (1846); Zhu et al., Blumea
57: 35 (2012).
277. Lasianthus venulosus Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 23 (1912), nom.
illeg., non (Wight & Arn.) Wight (1846). – TYPE: Singapore, Bukit Panjang, 1906,
H.N. Ridley 12542 (lectotype K [K000763924], designated here). Additional syntype:
Singapore, Chan Chu Kang, H.N. Ridley 6146 (SING [SING0012081]).
= Lasianthus ellipticus Wight, Calcutta J. Nat. Hist. 6: 507 (1846); Zhu et al., Blumea
57: 35 (2012).
278. Lasianthus villosus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 86 (1918). –
Litosanthes villosus (Ridl.) Deb & M.Gangop., Candollea 44(1): 223 (1989). – TYPE:
Peninsular Malaysia, Perak, Tapah, November 1908, H.N. Ridley 14074 (lectotype K
[K000763886], designated here; isolectotype BM [BM001190979]).
279. Lasianthus vulcanicus Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 71 (1923);
Zhu et al., Blumea 57: 94 (2012). – TYPE: Sumatra, Berastagi, Sibayak Volcano, Karoo
Plains, 11 February 1921, H.N. Ridley s.n. (lectotype K [K000777109], designated
here, or possibly holotype).
280. Mussaenda angustisepala Ridl., Fl. Malay Penins. 2: 60 (1923); Chantaranothai,
Thai Forest Bull., Bot. 43: 53 (2015). – TYPE: Peninsular Malaysia, Pahang, Kwala
Tembeling, 1891, H.N. Ridley 2188 (lectotype K [K000740943], designated by
Chantaranothai (2015)). Additional syntypes: Peninsular Malaysia, Perak, Pluss River,
s.dat., L. Wray 44 (K); Perak, Gopeng, September 1880, King’s Collector 731 (K);
Perak, Kal, June 1886, King’s Collector 10345 (K[×2]); Kelantan, Kuala Lebir, July
1904, Dr J.D. Gimlette s.n. (K, SING [SING0240246]).
281. Mussaenda celebica Ridl., Bull. Misc. Inform. Kew 1939: 598 (1940). – TYPE:
Celebes {Sulawesi}, Malino, 280 m, 4 April 1921, H.A.B. Bünnemeijer 10772
(holotype K [K000740933]).
358 Gard. Bull. Singapore 70 (2) 2018
282. Mussaenda cordifolia Wall. ex G.Don var. pubescens Ridl., Fl. Malay Penins.
2: 62 (1923). – TYPE: Peninsular Malaysia, Selangor, Sempang Mines, April 1894,
H.N. Ridley 15660 (lectotype K [K001129453], designated here; isolectotype SING
[SING0240245]).
= Mussaenda cordifolia Wall. ex G.Don, Gen. Hist. 3: 491 (1834).
283. Mussaenda hirsuta Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 68 (1923).
– TYPE: Sumatra, Berastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000740928], designated here, or possibly holotype).
284. Mussaenda lanuginosa Ridl., Bull. Misc. Inform. Kew 1939: 597 (1940).
– TYPE: Borneo, Sarawak, near Kuching, 22 March 1893, G.D. Haviland 2870
(holotype K [K000740927]).
285. Mussaenda malaccensis Ridl., Fl. Malay Penins. 2: 59 (1923). – TYPE: Peninsular
Malaysia, Malacca, s.dat., A.C. Maingay s.n. [Kew Dist. No. 835/2] (lectotype K
[K000740944], designated here, or possibly holotype).
286. Mussaenda motleyi Ridl., Bull. Misc. Inform. Kew 1939: 599 (1940). – TYPE:
Borneo, Kalimantan, Bangarmassing [Banjermasin], 1857–1858, J. Motley 695
(holotype K [K000740923]).
287. Mussaenda mutabilis Hemsl. var. montana Ridl., Fl. Malay Penins. 2: 59 (1923).
– Asemanthia maingayi (Hook.f.) Ridl. var. montana (Ridl.) A.K.Sinha & B.Mitra, J.
Bombay Nat. Hist. Soc. 78(2): 428 (1981). – TYPE: Peninsular Malaysia, Malacca,
Mount Ophir, Padang Batu, H.N. Ridley 3215 (lectotype CAL, designated by Sinha &
Mitra (1981); isolectotype SING [SING0240244]).
= Mussaenda maingayi (Hook.f.) Hemsl. ex T.Durand & B.D.Jacks. Index Kew.
Suppl. 1: 284 (1903).
288. Mussaenda purpurascens Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 18 (1912);
Chantaranothai, Thai Forest Bull., Bot. 43: 57 (2015). – TYPE: Peninsular Malaysia,
Selangor, Sempang, April 1911, H.N. Ridley s.n. (lectotype K [K000740941],
designated here, or possibly holotype).
289. Mussaenda sanderiana Ridl., Gard. Chron. 46: 34 (1909); Chantaranothai, Thai
Forest Bull., Bot. 43 (2015) 58. – TYPE: Laos, 1909, H.N. Ridley 14129 [leg. W.
359
The Gentinales taxa of H.N. Ridley
Micholitz] (lectotype SING [SING0058851], designated here; isolectotype SING
[SING0058852]).
290. Mussaenda spectabilis Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 78 (1918).
– TYPE: Peninsular Malaysia, Pahang, Pulau Tioman, June 1916, C.B. Kloss s.n.
(lectotype K [K000740947, K001129441 – i.e. a single specimen over two sheets],
designated here, or possibly holotype).
291. Mycetia angustifolia Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 68 (1923).
– TYPE: Sumatra, Berastagi, 11 February 1921, H.N. Ridley s.n. (lectotype K
[K000760574], designated here, or possibly holotype).
292. Mycetia minor Ridl., Bull. Misc. Inform. Kew 1926: 67 (1926). – TYPE: Sumatra,
Sipora, 10 October 1924, C.B. Kloss 14686 (lectotype K [K000760573], designated
here; isolectotype SING [SING0240247]).
293. Myrioneuron capitatum Ridl., J. Fed. Malay States Mus. 10: 93 (1920), as
‘capitata’. – Keenania capitata (Ridl.) Craib, Fl. Siam. 2: 82 (1932). – TYPE:
Thailand, Tasan, s.dat., C.B. Kloss 7053 (lectotype K [K001067653], designated here,
or possibly holotype).
294. Myrioneuron microcephalum Ridl., J. Fed. Malay States Mus. 7: 43 (1916). –
Rennellia microcephala (Ridl.) K.M.Wong, Arbor. Rubiaceae Malaya 111 (1988). –
TYPE: Peninsular Malaysia, Kedah, Gurun, December 1915, H.C. Robinson & C.B.
Kloss 6180 (lectotype K [K001067645], designated here; isolectotype SING).
295. Oldenlandia rosea Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 110 (1911). –
TYPE: Thailand, Setul, heaths, 1910, H.N. Ridley 15466 (lectotype K [K000760532],
designated here; isolectotype BM [BM001190972]).
= Hedyotis kurzii Merr., Pap. Michigan Acad. Sci. 19: 195 (1934).
296. Oldenlandia stolonifera Ridl., Bull. Misc. Inform. Kew 1939: 613 (1940). –
TYPE: Borneo, Sarawak, Mt Singhi, 11 December 1892, G.D. Haviland 2151
(holotype, K [K000760303]; isotype SING [SING0058332]).
360 Gard. Bull. Singapore 70 (2) 2018
297. Ophiorrhiza arenicola Ridl., Sarawak Mus. J. 1(2): 31 (1912). – TYPE: Borneo,
Sarawak, Puak, September 1905, H.N. Ridley 12449 (lectotype K [K000740650],
designated here; isolectotype SING [SING0240248]).
298. Ophiorrhiza avenis Ridl., J. Fed. Malay States Mus. 8(4): 48 (1917). – TYPE:
Sumatra, Barong Baru, W. side of Barisan Range at 4000 ft, 4 June 1914, H.C.
Robinson & C.B. Kloss s.n. (lectotype BM [BM000945188], designated here, or
possibly holotype).
299. Ophiorrhiza axillaris Ridl., Sarawak Mus. J. 1(2): 30 (1912). – TYPE: Borneo,
Sarawak, Matang, s.dat., G.D. Haviland 84 [c.l.r.p.] [687] (lectotype K [K000740649],
designated here; isolectotype SING [SING0240250]). Additional syntype: Borneo,
Sarawak, Matang, s.dat., Hullett s.n. (SING [SING0240251]).
300. Ophiorrhiza bracteata Ridl., J. Fed. Malay States Mus. 8(4): 47 (1917), nom.
illeg., non Korth. (1851). – Ophiorrhiza ridleyi Merr., Pap. Michigan Acad. Sci. 19:
198 (1934). – TYPE: Indonesia, Sumatra, Sungei Kumbang at 5000 ft, 18 March 1914,
H.C. Robinson & C.B. Kloss s.n. (lectotype BM [BM000945186], designated here).
Additional syntype: Indonesia, Sumatra, Siolak Daras at 4400 ft, 16 March 1914, H.C.
Robinson & C.B. Kloss s.n. (BM [BM000945187]).
301. Ophiorrhiza communis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 16 (1912). –
TYPE: Peninsular Malaysia, Bukit Tampin, J.S. Goodenough 1860 (lectotype SING
[SING0051290], designated here). Additional syntypes: Peninsular Malaysia, Selangor,
Rawang, May 1896, H.N. Ridley 7404 (SING [SING0051287]); Selangor, Ginting
Bidai, May 1896, H.N. Ridley 7402 (K [K000740586], SING [SING0051294]); ibid.,
H.N. Ridley 7407 (SING [SING0051293]) (var. with pink s); Selangor, Sempang,
H.N. Ridley 12075 (SING [SING0043414]); Selangor, Semangkok Pass, H.N. Ridley
8580 (SING [SING0240249]); Selangor, Klang Gates, H.N. Ridley 13413 (SING
[SING0051289]); Perak, Dindings, Pangkor, Cantley 8345 (not traced); Telok Pinang,
Cantley 9765 (SING [SING0051291]); Pahang, Telom, November 1908, H.N. Ridley
13642 (K, SING [SING0051295]); Pahang, Kuala Lipis, May 1903, A.D. Machado
11577 (SING [SING0051288]).
302. Ophiorrhiza costata Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 15 (1912). –
TYPE: Peninsular Malaysia, Pahang, Telom, November 1908, H.N. Ridley 13644
(lectotype K [K000740594], designated here; isolectotypes SING [SING0240264,
SING0240263]). Additional syntypes: Peninsular Malaysia, Selangor, Bukit Hitam,
Kelsall 1908 (not traced); Selangor, Semangkok Pass, H.N. Ridley 8584 (SING
[SING0240262]).
361
The Gentinales taxa of H.N. Ridley
303. Ophiorrhiza deexa Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 65 (1923). –
TYPE: Sumatra, Berastagi Woods, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000740647], designated here; isolectotype K [K000740648]).
304. Ophiorrhiza discolor R.Br. ex G.Don var. elegans Ridl., Fl. Malay Penins. 2:
36 (1923). – TYPE: Peninsular Malaysia, Kelantan, Glam Wood near Kota Bahru,
19 February 1917, H.N. Ridley s.n. (lectotype K [K001067657], designated here;
isolectotype SING [SING0240258]).
= Ophiorrhiza discolor R.Br. ex G.Don, Gen. Hist. 3: 522 (1834).
305. Ophiorrhiza exserta Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 66 (1923).
– TYPE: Sumatra, Berastagi, 13 February 1921, H.N. Ridley s.n. (lectotype K
[K000740644], designated here; possible isolectotype K [K000740645]).
The second sheet in K does not bear the exact date of the chosen lectotype but may be
a duplicate collection.
306. Ophiorrhiza brillosa Ridl., J. Straits Branch Roy. Asiat. Soc. 33: 22 (1900). –
TYPE: Borneo, Sarawak, Penrissen, 4000 ft, Shelford (not traced).
307. Ophiorrhiza avescens Ridl., J. Fed. Malay States Mus. 8(4): 48 (1917). –
TYPE: Sumatra, Sandaran Agong at 2450 ft, 25 May 1914, H.C. Robinson & C.B.
Kloss 75 (lectotype BM [BM000945179], designated here; possible isolectotype K
[K000740640]).
The specimen in K does not bear a collection number but otherwise the details are the
same as the BM specimen.
308. Ophiorrhiza fontinalis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 55 (1912). –
TYPE: Thailand, Rawei Island [Ko Ra Wi], April 1911, H.N. Ridley 15895 (lectotype
K [K000740589], designated here; isolectotype BM [BM000832520]). Additional
syntype: Peninsular Malaysia, Kedah, Langkawi Islands, Telaya Tujoh, April 1911,
H.N. Ridley 15895b (K [K000740590]).
309. Ophiorrhiza fruticulosa Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 15 (1912),
nom. illeg., non Nadeaud (1873). – Ophiorrhiza fruticosa Ridl., Fl. Malay Penins. 2: 41
(1923). – TYPE: Peninsular Malaysia, Selangor, Gua Batu [Batu Caves], top of rocks,
December 1896, H.N. Ridley 8237 (lectotype SING [SING0043413], designated here;
362 Gard. Bull. Singapore 70 (2) 2018
isolectotype K [K000740579]). Additional syntype: Peninsular Malaysia, Selangor,
Gua Batu [Batu Caves], H.N. Ridley 8583 (SING [SING0240257]).
The lectotype collection was cited incorrectly as 8287 in the protologue.
310. Ophiorrhiza havilandii Ridl., Sarawak Mus. J. 1(2): 30 (1912). – TYPE:
Borneo, Sarawak, Mt Kowon, 21 September 1892, G.D. Haviland 1709 (lectotype
K [K000740638], designated here; isolectotype SING [SING0240253]). Additional
syntypes: Borneo, Sarawak, Matang, July 1903, H.N. Ridley 11758 (K [K000740639],
SING [SING0240254]); Borneo, Sarawak, Braang, 24 November 1888, G.D. Haviland
57 (SING [SING0240255]).
311. Ophiorrhiza ixora Ridl., J. Fed. Malay States Mus. 8(4): 46 (1917). – TYPE:
Sumatra, Sungei Kumbang at 4500 ft, 29 March 1914, H.C. Robinson & C.B. Kloss
s.n. (lectotype BM [BM000945177], designated here; isolectotype K [K000740637]).
312. Ophiorrhiza kingiana Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 17 (1912).
– TYPE: Peninsular Malaysia, Perak, Gopeng, April 1884, King’s Collector 5853
(lectotype SING [SING0043412], designated here).
313. Ophiorrhiza klossii Ridl., J. Linn. Soc., Bot. 41: 291 (1913). – TYPE: Peninsular
Malaysia, Selangor, Menuang Gasing, Ulu Langat, February 1912, C.B. Kloss s.n.
(lectotype K [K000740593], designated here; isolectotype BM [BM000832521]).
314. Ophiorrhiza lancifolia Ridl., J. Fed. Malay States Mus. 5: 162 (1915). – TYPE:
Thailand, Koh Samui, 16 May 1913, H.C. Robinson 5739 (lectotype K [K000740666],
designated here, or possibly holotype).
315. Ophiorrhiza longerepens Ridl., Bull. Misc. Inform. Kew 1929: 258 (1929), as
‘longe-repens’. – TYPE: Peninsular Malaysia, Kelantan, Gua Panjang at Gua Ninik,
21 October 1927, M.R. Henderson SFN 19524 (lectotype K [K000740576], designated
here; isolectotypes K [K000740575], SING [SING0051286]).
316. Ophiorrhiza major Ridl., Fl. Malay Penins. 2: 36 (1923). – TYPE: Peninsular
Malaysia, Kelantan, Chaning Woods, 4 February 1917, H.N. Ridley s.n. (lectotype K
[K000740597], designated here). Additional syntypes: Peninsular Malaysia, Tomoh,
s.dat., A. Machado s.n. (SING [SING0240276]); Kelantan, Chaning Woods, 8
363
The Gentinales taxa of H.N. Ridley
February 1917, H.N. Ridley s.n. (K [K000740596]).
= Ophiorrhiza remotiora Ridl., J. Fed. Malay States Mus. 10: 140 (1920).
The protologue lists a particularly large number of syntypes, most of which we have
omitted.
317. Ophiorrhiza nervosa Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 18 (1912). –
TYPE: Peninsular Malaysia, Selangor, Pahang track, May 1903, A.D. Machado s.n.
(lectotype SING [SING0051284], designated here; isolectotypes K [K000740574],
SING [SING0051285]).
318. Ophiorrhiza nutans Ridl., J. Fed. Malay States Mus. 8(4): 47 (1917), nom.
illeg., non Clarke (1880). – TYPE: Sumatra, Siolak Daras at 3000 ft, 19 March 1914,
H.C. Robinson & C.B. Kloss s.n. (lectotype BM [BM000945176], designated here;
isolectotypes K [K000740629], SING [SING0240261]).
319. Ophiorrhiza pallidula Ridl., Fl. Malay Penins. 2: 38 (1923). – TYPE: Peninsular
Malaysia, Selangor Ulu Gombak, December 1920, H.N. Ridley s.n. (lectotype K
[K000740588], designated here; isolectotypes SING [SING0240269, SING0240270]).
Additional syntypes: Peninsular Malaysia, Pahang, Tahan river woods, August 1891,
H.N. Ridley s.n. (SING [SING0240273]); Peninsular Malaysia, Selangor, top of Batu
Caves; Ginting Bidai, May 1895, H.N. Ridley 7406 (SING [SING0240272]); Selangor,
Semangkok, August 1906, H.N. Ridley 12076 (SING [SING0240271]); Perak, Ipoh,
December 1895, C. Curtis 3147 (SING [SING0240274]).
320. Ophiorrhiza pilosissima Ridl., J. Fed. Malay States Mus. 8(4): 47 (1917). – TYPE:
Sumatra, Korinchi, Siolak Daras at 3000 ft, 16 March 1914, H.C. Robinson & C. B.
Kloss s.n. (lectotype BM [BM000945173], designated here, or possibly holotype).
321. Ophiorrhiza remotiora Ridl., J. Fed. Malay States Mus. 10: 140 (1920). –
TYPE: Peninsular Malaysia, Pahang, Gunong Senyum, June 1917, I.H.N. Evans s.n.
(lectotype K [K000740595], designated here; isolectotypes BM [BM001190971],
SING [SING0240268]). Additional syntype: Peninsular Malaysia, Pahang, Jerantut,
Kota Tonkat, Unknown collector (not traced).
322. Ophiorrhiza reticulata Ridl., Sarawak Mus. J. 1(2): 31 (1912), nom. illeg., non
Korth. (1851). – Ophiorrhiza sarawakensis Merr., J. Arnold Arbor. 35: 155 (1954).
– TYPE: Borneo, Sarawak, Bau, July 1893, H.N. Ridley 11756 (lectotype SING
364 Gard. Bull. Singapore 70 (2) 2018
[SING0062089], designated here; isolectotype K [K000740627]). Additional syntypes:
Borneo, Sarawak, Jambusan, September 1905, H.N. Ridley 12364 (K [K000740626],
SING [SING0062090]); Sarawak, Braang or Bau, limestone rocks, G.D. Haviland
8495 [c.d.f.f.] (SING [SING0062088]).
323. Ophiorrhiza rosea Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 55 (1911), nom.
illeg., non Hook.f. (1880). – Ophiorrhiza rosacea Ridl., Fl. Malay Penins. 2: 38
(1923). – TYPE: Peninsular Malaysia, Perak, Temango, July 1909, H.N. Ridley 14482
(lectotype K [K000740592], designated here; isolectotype BM, SING [SING0240267].
Additional syntype: Peninsular Malaysia, Perak, Temango, July 1909, H.N. Ridley
14483 (SING [SING0240266]).
324. Ophiorrhiza scabrella Ridl., Fl. Malay Penins. 2: 39 (1923). – TYPE:
Thailand, Adang islands, Pulau Butang, April 1911, H.N. Ridley 15894 (lectotype K
[K000740587], designated here; isolectotype SING [SING0240265]).
325. Ophiorrhiza singapurensis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 17
(1912). – TYPE: Singapore, Bukit Timah, October 1883, R.W. Hullett 335 (lectotype
SING [SING0030403], designated here). Additional syntypes: Singapore, Bukit
Timah, 1891, H.N. Ridley 2866 (BM [BM000945172], K [K000740585]); ibid., H.N.
Ridley 6899 (SING [SING0172367]); Pulau Damar, 1911, H.N. Ridley s.n. (SING
[SING0030399]); Pulau Ubin, March 1885, R.W. Hullett 424 (SING [SING0030393]);
Jurong, H.N. Ridley 10763 (not traced); Peninsular Malaysia, Johore, Gunong Pulai,
December 1904, H.N. Ridley 12204 (K, SING [SING0051280]); Johore, Gunong
Panti, December 1892, H.N. Ridley 4172 (K, SING [SING0051281]); Johore, Sungei
Tebrau, s.dat., H.N. Ridley 13221 (SING [SING0051279]); Negri Sembilan, Bukit
Sulu, Cantley 3296 (SING [SING0241772]); Perak, Dindings, Lumut, March 1896,
H.N. Ridley 7195 (SING [SING0051278]); Sumatra, Lingga Island, Hullett (not
traced).
The lectotype collection was incorrectly cited as 2855 in the protologue.
326. Ophiorrhiza stylosa Ridl., J. Fed. Malay States Mus. 8(4): 48 (1917). – TYPE:
Sumatra, Barong Baru, 6 June 1914, H.C. Robinson & C.B. Kloss 97 (lectotype BM
[BM000945171], designated here, or possibly holotype).
327. Ophiorrhiza subcrenata Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 66 (1923).
– TYPE: Sumatra, Bandar Baru, Gunong Sebayok, 6 August 1921, Md. Nur SFN 7345
(lectotype K [K000740625], designated here; isolectotypes SING [SING0240259,
SING0240260]).
365
The Gentinales taxa of H.N. Ridley
328. Ophiorrhiza tenuis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 16 (1912), as
‘tenue’. – TYPE: Peninsular Malaysia, Selangor, Dusun Tua, 20 May 1895, H.N.
Ridley s.n. (lectotype SING [SING0051283], designated here, or possibly holotype).
Additional syntypes: Peninsular Malaysia, Perak, Telor Pinang, October 1898, H.N.
Ridley 9767 (SING [SING0051282]); Perak, Tambun, Unknown collector (not traced).
329. Ophiorrhiza tenuis Ridl. var. minor Ridl., Fl. Malay Penins. 2: 42 (1923). –
TYPE: Peninsular Malaysia, Selangor, Ulu Gombak, 10 March 1915, H.N. Ridley s.n.
(lectotype K [K000740577], designated here, or possibly holotype).
= Ophiorrhiza tenuis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 16 (1912).
330. Paederia tomentosa Blume var. subglabra Ridl., J. Straits Branch Roy. Asiat.
Soc. 59: 121 (1911). – TYPE: Peninsular Malaysia, Perlis, Bukit Lagi, H.N. Ridley s.n.
(lectotype SING [SING0240279], designated here, or possibly holotype).
= Paederia foetida L., Mant. Pl. 1: 52 (1767).
331. Palicourea insularis Ridl., J. Linn. Soc., Bot. 27: 41 (1890). – Erithalis insularis
(Ridl.) Zappi & T.S.Nunes, Kew Bull. 55(3): 655 (2000). – TYPE: Brazil, Fernando
Noronha, 1887, Ridley, Lea and Ramage 86 (lectotype BM [BM000949953],
designated by Zappi & Nunes (2000); isolectotype K [K000016496]).
= Erithalis fruticosa L., Syst. Nat. ed. 10, 2: 930 (1759).
332. Pavetta indica L. var. canescens Ridl., Fl. Malay Penins. 2: 100 (1923). – TYPE:
Singapore, October 1822, N. Wallich s.n. [EIC 6181] (lectotype K-W [K001123242],
designated here).
= Pavetta wallichiana Steud. ex Bremek., Repert. Spec. Nov. Regni Veg. 37: 84
(1934).
Note that Pavetta indica L. var. canescens Pit., Fl. Indo-Chine 3: 332 (1924), a variety
from Annam in Vietnam, is a later homonym which was renamed Pavetta pitardii
Bremek., Repert. Spec. Nov. Regni Veg. 37: 83 (1934).
333. Pavetta naucleiora R.Br. ex G.Don var. longifolia Ridl., Fl. Malay Penins.
2: 101 (1923). – TYPE: Peninsular Malaysia, Kelantan, Chaning, Kelantan River,
February 1917, H.N. Ridley s.n. (lectotype K [K001067638], designated here, or
possibly holotype).
= Pavetta napieri (Ridl.) Bremek., Repert. Spec. Nov. Regni Veg. 37: 96 (1934).
366 Gard. Bull. Singapore 70 (2) 2018
334. Pavetta pauciora Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 297 (1922),
nom. illeg., non Blume ex DC. (1830). – TYPE: Peninsular Malaysia, Selangor, Batu
Caves, December 1920, H.N. Ridley s.n. (lectotype K [K001067636], designated here;
isolectotypes SING [SING0058846, SING0240261 (dated 13 December 1920)]).
= Pavetta sylvatica Blume, Bijdr. Fl. Ned. Ind. 16: 953 (1826); Bremekamp, Repert.
Spec. Nov. Regni Veg. 37: 86 (1934).
335. Petunga conifera Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 81 (1918). –
Hypobathrum coniferum (Ridl.) Kiew in Kiew & Turner, Gard. Bull. Singapore 55:
182 (2003). – TYPE: Singapore, Gardens’ Jungle, 1899, H.N. Ridley 10722 (lectotype
K [K001067665], designated here; isolectotype SING [SING0045448]).
336. Petunga hirta Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 69 (1923). –
Hypobathrum longifolium (DC.) K.M.Wong var. hirtum (Ridl.) K.M.Wong, Arbor.
Rubiaceae Malaya 77 (1988), as ‘hirta’. – Hypobathrum hirtum (Ridl.) Mulyan. &
Ridsdale, Reinwardtia 12: 107 (2002). – TYPE: Sumatra, Sibolangat, Bukit Kluang, 4
August 1921, Md. Nur SFN 7405 (lectotype SING [SING0058021], designated here,
or possibly holotype).
337. Polycycliska cylindrica Ridl., Bull. Misc. Inform. Kew 1926: 67 (1926). –
TYPE: Sumatra, Siberut, 9 September 1924, C.B. Kloss 11439 (lectotype SING
[SING0011439], designated by Axelius (1987); isolectotype K [K000760598]).
= Lerchea bracteata Valeton, Icon. Bogor. 4: t. 382 (1914); Axelius, Blumea 32: 99
(1987).
The type gathering was incorrectly referred to as 11489 in the protologue.
338. Pomazota sylvestris Ridl., Trans. Linn. Soc. London, Bot. 3(9): 308 (1893).
– Coptophyllum sylvestre (Ridl.) I.M.Turner, Novon 6: 222 (1996). – TYPE:
Peninsular Malaysia, Pahang, Pulau Tawar, 1891, H.N. Ridley 2230 (lectotype SING
[SING0058441], designated by Turner (1996)).
339. Porterandia catappifolia Ridl., Bull. Misc. Inform. Kew 1939: 596 (1940). –
TYPE: Borneo, Sarawak, 1865–1868, O. Beccari P.B. 2599 (holotype K [K000740990];
isotypes FI-B!, P [P00836435]).
340. Porterandia glabrifolia Ridl., Bull. Misc. Inform. Kew 1939: 596 (1940). – TYPE:
Borneo, Sarawak, 1865–1868, O. Beccari P.B. 1686 (holotype K [K000740989];
isotypes A [A00094993], FI-B!).
367
The Gentinales taxa of H.N. Ridley
341. Porterandia grandifolia Ridl., Bull. Misc. Inform. Kew 1939: 595 (1940). –
TYPE: Borneo, Sarawak, near Kuching, 25 October 1892, G.D. Haviland 1892 [Garai
leg.] (lectotype K [K000740992], designated here; isolectotypes K [K000740993], L
[L0625767], SAR!, SING [SING0052703]).
342. Porterandia minor Ridl., Bull. Misc. Inform. Kew 1939: 594 (1940). – TYPE:
Borneo, Sarawak, Rejang Belaga, November 1892, G.D. Haviland 2161 (holotype K
[K000740994]; isotypes SAR, SING [SING0052704]).
343. Porterandia pauciora Ridl., Bull. Misc. Inform. Kew 1939: 597 (1940). –
TYPE: Borneo, Sarawak, Niah, May 1892, G.D. Haviland & C. Hose 3421 (holotype
K [K000740988]; possible isotype SAR (labelled 3421A)).
344. Porterandia sessiliora Ridl., Bull. Misc. Inform. Kew 1939: 597 (1940). –
Bungarimba sessiliora (Ridl.) K.M.Wong, Sandakania 15: 50 (2004). – TYPE:
Borneo, Sarawak, Kuching, 9 October 1894, [G.D.Haviland & C. Hose] 1961 A
(holotype K [K000786007]).
345. Prismatomeris andamanica Ridl., Bull. Misc. Inform. Kew 1939: 607 (1940).
– Prismatomeris fragrans Geddes subsp. andamanica (Ridl.) J.T.Johanss., Opera
Bot. 94: 45 (1987). – TYPE: India, Andaman Islands, Long Island, June 1915, C.E.
Parkinson 601 (holotype K [K000031590]).
346. Prismatomeris brachypus Ridl., Bull. Misc. Inform. Kew 1939: 602 (1940);
Johansson, Opera Bot. 94: 37 (1987). – TYPE: Philippines, Luzon, Province of
Cagayan, January 1912, H.M. Curran Forestry Bureau No. 17784 (lectotype K
[K000763849], designated by Johansson (1987); isolectotypes BM [BM000796390,
BM000796392], P, US [US00050617]).
347. Prismatomeris grifthii Ridl., J. Fed. Malay States Mus. 10: 96 (1920); Johansson,
Opera Bot. 94: 32 (1987). – TYPE: Thailand, Tasan, [5 November 1919], C.B. Kloss
6940 (lectotype K [K000763866], designated by Johansson (1987); isolectotype SING
[SING0058839, SING0058840]). Additional syntype: Burma, Mergui, W. Grifth (K).
348. Prismatomeris malayana Ridl., J. Fed. Malay States Mus. 10(2): 96, 97, 142
(1920). – Prismatomeris tetrandra (Roxb.) K.Schum. subsp. malayana (Ridl.)
J.T.Johanss., Opera Bot. 94: 29 (1987). – TYPE: Thailand, Delisle Island off Renong,
s.dat., C.B. Kloss 6647 (lectotype K [K001067637], designated by Johansson (1987)).
368 Gard. Bull. Singapore 70 (2) 2018
349. Prismatomeris multiora Ridl., Bull. Misc. Inform. Kew 1939: 603 (1940). –
Prismatomeris tetrandra (Roxb.) K.Schum. subsp. multiora (Ridl.) Y.Z.Ruan, Acta
Phytotax. Sin. 26(6): 447 (1988). – TYPE: Thailand, Doi Angka Noi, near summit at
1540 m, 27 June 1927, H.B.G. Garrett 390 (holotype K [K000763867, K000763869 –
i.e. a single specimen over two sheets]; isotypes ABD, C, E, K [K000763868]).
= Prismatomeris tetrandra (Roxb.) K.Schum. in Engler & Prantl, Nat. Panzenfam.
4(4): 138 (1891); Johansson, Opera Bot. 94: 26 (1987).
350. Prismatomeris ovalifolia Ridl., Bull. Misc. Inform. Kew 1939: 603 (1940). –
TYPE: Sumatra, Anambas Islands, Tanjong Suka, Siantan on rocks, 3 April 1928,
M.R. Henderson 20228 (lectotype K [K000763859], designated by Johansson (1987);
isolectotypes K [K000763858], SING [SING0058844]).
= Prismatomeris tetrandra (Roxb.) K.Schum. subsp. malayana (Ridl.) J.T.Johanss.,
Opera Bot. 94: 29 (1987).
351. Prismatomeris parkinsonii Ridl., Bull. Misc. Inform. Kew 1939: 605 (1940).
– TYPE: Burma, Tenasserim, Nyaungbinkin, 20 February 1926, C.E. Parkinson
1933 (lectotype K [K000031589], designated by Johansson (1987); isolectotype K
[K000031588]).
= Prismatomeris tetrandra (Roxb.) K.Schum. subsp. malayana (Ridl.) J.T.Johanss.,
Opera Bot. 94: 29 (1987).
352. Prismatomeris parviora Ridl., Bull. Misc. Inform. Kew 1939: 605 (1940). –
TYPE: Peninsular Malaysia, Perlis, Chupeng, March 1912?, H.N. Ridley 15000
(holotype K [K000763870]; possible isotypes BM [March 1910], SING [1911?]).
= Prismatomeris tetrandra (Roxb.) K.Schum. subsp. malayana (Ridl.) J.T.Johanss.,
Opera Bot. 94: 29 (1987).
The date on the holotype is difcult to determine. Consequently, it is hard to tell if the
specimens of Ridley 15000 are duplicates of a single collection, hence a holotype and
isotypes, or if they were collected on different dates.
353. Prismatomeria tetrandra (Roxb.) K.Schum. var. philippinensis Ridl., Bull. Misc.
Inform. Kew 1939: 603 (1940). – TYPE: Philippines, Luzon, Prov. of Tayabas, March
1911, M. Ramos Bur. Sci. 13315 (lectotype K [K000763850], designated by Johansson
(1987); isolectotypes BM, CAL, E, F, G, L, LD, MO, S).
= Prismatomeris tetrandra (Roxb.) K.Schum. in Engler & Prantl, Nat. Panzenfam.
4(4): 138 (1891); Johansson, Opera Bot. 94: 26 (1987).
369
The Gentinales taxa of H.N. Ridley
354. Psychotria arborea Ridl., J. Fed. Malay States Mus. 8(4): 51 (1917), nom. illeg.,
non Hiern (1877), nec Sessé & Moc. (1897). – TYPE: Sumatra, Korinchi Peak, 24 April
1914, H.C. Robinson & C.B. Kloss 65 (lectotype BM [BM000945356], designated
here, or possibly holotype).
355. Psychotria ascendens Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 61 (1911).
– TYPE: Peninsular Malaysia, Perak, Temengoh [Temenggor], July 1909, H.N.
Ridley 14485 (lectotype K [K000777155], designated by Turner & Kumar (2018);
isolectotypes BM, SING [SING0234546]).
= Psychotria polytricha Miq., Fl. Ned. Ind. 2: 287 (1857).
356. Psychotria atroviridis Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 297 (1922). –
TYPE: Peninsular Malaysia, Negri Sembilan, Bukit Tangga, 22 December 1920, H.N.
Ridley s.n. (lectotype K [K000777130], designated here, or possibly holotype).
357. Psychotria auriculata Ridl., J. Fed. Malay States Mus. 10: 97 (1920), nom. illeg.,
non Griseb. (1866). – Psychotria klossii Craib, Fl. Siam. 2: 191 (1934). – TYPE:
Thailand, Tasan, s.dat., C.B. Kloss 6853 (lectotype K [K001067641], designated here).
Additional syntype: Thailand, Tasan, s.dat., C.B. Kloss 6945 (not traced).
358. Psychotria brachybotrys Ridl., J. Fed. Malay States Mus. 4: 38 (1909), nom.
illeg., non P. brachybotrya Müll.Arg. (1881). – Psychotria tahanensis Ruhsam, Bot.
J. Linn. Soc. 157: 116 (2008). – TYPE: Peninsular Malaysia, Selangor, Semangko
Pass, 1904, H.N. Ridley 12072 (lectotype K [K000777150], designated by Turner &
Kumar (2018)). Additional syntype: Peninsular Malaysia, Gunung Berumban at 6000
ft elevation, Unknown collector (not traced).
359. Psychotria cantleyi Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 24 (1912). –
TYPE: Peninsular Malaysia, Johore, Mount Austin, Kwala Sembrong, 1892, H.W.
Lake & H.J. Kelsall s.n. (lectotype SING [SING0234543], designated by Turner
& Kumar (2018)). Additional syntypes: Singapore, Chua Chu Kang, Unknown
collector (not traced); Peninsular Malaysia, Negri Sembilan, Buruang, M.V. Alvins
2053 (SING [SING0241770]); Bukit Sulu, M.V. Alvins 1978 (SING [SING0241771]);
Selangor, Semangkok Pass, 15th mile Pahang Track, 1891, H.N. Ridley 8573 (SING
[SING0234541]); Malacca, Chenana Puteh, M.V. Alvins 844 (SING [SING0241769]);
Pahang, Kota Glanggi, 1891, H.N. Ridley s.n. (SING [SING0234540]); Pahang,
Sungei Tahan, 21 July 1893, H.N. Ridley 5830 (SING [SING0234539]); Perak, Tapah,
H.N. Ridley 14072 (K, SING [SING0234542]).
370 Gard. Bull. Singapore 70 (2) 2018
= Psychotria scortechinii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 74:
7 (1906).
The syntypes collected by Alvins are credited to Cantley in the protologue.
360. Psychotria cuprea Ridl., J. Fed. Malay States Mus. 8(4): 51 (1917). – TYPE:
Sumatra, Korinchi Peak 7300 ft, 2 May 1914, H.C. Robinson & C.B. Kloss 186
(lectotype BM [BM000945343], designated here; isolectotype K [K000777222]).
361. Psychotria elegans Ridl., Sarawak Mus. J. 1(2): 32 (1912). –TYPE: Borneo,
Sarawak, Matang, August 1904, H.N. Ridley 12294 (lectotype K [K000777217],
designated here). Additional syntype: Borneo, Sarawak, Matang, 800–900 ft, 19 June
1893, [Hewitt] (SING [SING0241773]).
362. Psychotria fraseri Ridl., Bull. Misc. Inform. Kew 1934: 120 (1934). – TYPE:
Borneo, British North Borneo {Sabah}, Kudat, 10 November 1884, M. Fraser 54
(holotype K [K000777204]).
?= Psychotria bangueyensis Merr., Philipp. J. Sci. 29: 423 (1926).
As indicated by W. Meijer on the type specimen, Psychotria fraseri Ridl. is probably
a synonym of P. bangueyensis Merr.
363. Psychotria grifthii Hook.f. var. angustifolia Ridl., Fl. Malay Penins. 2: 137
(1923). – TYPE: Peninsular Malaysia, Kedah, Langkawi, Burau Woods, April 1911,
H.N. Ridley 15823 (lectotype K [K001067640], designated here).
= Psychotria angulata Korth., Ned. Kruidk. Arch. 2(2): 243 (1851).
364. Psychotria lanceolaria Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 297 (1922).
– TYPE: Peninsular Malaysia, Selangor, Batu Caves, 27 December 1920, H.N. Ridley
s.n. (lectotype K [K000777149], designated here, or possibly holotype).
365. Psychotria lasiocephala Ridl., J. Fed. Malay States Mus. 10: 97 (1920). – TYPE:
Thailand, Tapli, s.dat., C.B. Kloss 6728 (lectotype K [K000777247], designated here,
or possibly holotype).
366. Psychotria megacarpa Ridl., J. Fed. Malay States Mus. 6: 48 (1915). – TYPE:
Peninsular Malaysia, Perak, Gunong Kerbau 3500 ft, 20 March 1913, H.C. Robinson
s.n. (lectotype K [K000777134], designated here; isolectotype BM).
371
The Gentinales taxa of H.N. Ridley
367. Psychotria minutiora Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 88 (1918),
nom. illeg., non Miq. (1876). – Psychotria sohmeriana I.M.Turner, Asian J. Trop.
Biol. 1(2): 26 (1995). – TYPE: Peninsular Malaysia, Selangor-Pahang border, Ginting
Simpah, March 1917, H.N. Ridley, H.C. Robinson & C.B. Kloss s.n. (lectotype K
[K000777129], designated here, or possibly holotype).
368. Psychotria multinervia Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 70 (1923).
– TYPE: Sumatra, Berastagi, Karoo Plateau, 8 February 1921, H.N. Ridley s.n.
(lectotype K [K000777190], designated here). Additional syntype: Sumatra, Berastagi,
10 February 1921, H.N. Ridley s.n. (K [K000777189]).
369. Psychotria patula Ridl., J. Fed. Malay States Mus. 8(4): 51 (1917), nom. illeg.,
non Schult. (1819). – Psychotria kumbangii Ruhsam in Ruhsam et al., Bot. J. Linn.
Soc. 157: 116 (2008). – TYPE: Sumatra, Sungei Kumbang 4–5000 ft, 29 March 1914,
H.C. Robinson & C.B. Kloss s.n. (lectotype BM [BM000945505], designated here;
isolectotype K [K000777193]).
370. Psychotria penduliora Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 70 (1923).
– TYPE: Sumatra, Berastagi Woods, 5500 ft, 8 February 1921, H.N. Ridley s.n.
(lectotype K [K000777187], designated here).
371. Psychotria rudis Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 87 (1918). – TYPE:
Peninsular Malaysia, Kedah, Kedah Peak 2800–4000 ft, December 1915, H.C.
Robinson & C.B. Kloss s.n. (lectotype K [K000777140], designated here, or possibly
holotype).
372. Psychotria sarmentosa Blume var. montana Ridl., Fl. Malay Penins. 2: 131
(1923). – Psychotria euosma I.M.Turner, Phytotaxa 361: 188 (2018). – TYPE:
Peninsular Malaysia, Pahang, Gunong Tahan, July 1911, H.N. Ridley 16068 (lectotype
K [K000777156], designated by Turner & Kumar (2018)). Additional syntype:
Peninsular Malaysia, Malacca, Mt Ophir, A.C. Maingay 2747 (K).
= Psychotria lutescens Craib, Bull. Misc. Inform. Kew 1932: 481 (1932).
373. Psychotria setistipula Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 88 (1918). –
TYPE: Peninsular Malaysia, Selangor, Gunong Mengkuang, 5 February 1913, H.C.
Robinson s.n. (lectotype K [K000777138], designated here, or possibly holotype).
= Ixora patens Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 83 (1918).
372 Gard. Bull. Singapore 70 (2) 2018
Psychotria setistipula Ridl. is here listed as a new synonym of Ixora patens Ridl.
374. Psychotria siamensis Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 120 (1911).
– TYPE: Peninsular Malaysia, Kedah, Pulau Langkawi, September 1890, C.
Curtis 2631 (lectotype SING [SING0234537], designated here; isolectotype SING
[SING0234538]). Additional syntype: Thailand, Setul heaths, H.N. Ridley 15096 (not
traced).
= Psychotria adenophylla Wall. in Roxburgh, Fl. Ind. 2: 166 (1824).
The specimen [SING0234538] has been labelled 2636 in a later hand but this is almost
certainly a mistake for 2631.
375. Psychotria sumatrensis Ridl., Bull. Misc. Inform. Kew 1926: 70 (1926). – TYPE:
Sumatra, Sipora, 9 October 1924, C.B. Kloss 14666 (lectotype K [K001067646],
designated here, or possibly holotype).
376. Psychotria vulpina Ridl., J. Fed. Malay States Mus. 10: 142 (1920). – TYPE:
Peninsular Malaysia, Kelantan, Kuala Lebir, February 1914, H.N. Ridley s.n. (lectotype
K [K000777142], designated here, or possibly holotype).
377. Randia fragrantissima Ridl., J. Straits Branch Roy. Asiat. Soc. 50: 115 (1908).
– Oxyceros fragrantissimus (Ridl.) K.M.Wong, Malayan Nat. J. 38: 43 (1984).
– TYPE: Singapore, Gardens’ Jungle, 1896, H.N. Ridley 5664 (lectotype SING
[SING0001133], designated by Wong (1984)). Additional syntypes: Singapore, Bukit
Timah, February 1908, H.N. Ridley 13022 (K [K000742990, K000742991], SING
[SING0001112, SING0172368]); Singapore, Pulau Tekong, 1891, H.N. Ridley s.n.
(SING [SING0207034]); Peninsular Malaysia, Malacca, Selandor, M.V. Alvins 484
[‘Cantley’s Collector’ in protologue] (SING [SING0001115]); Malacca, Ayer Panas,
R. Derry 1056 (SING [SING0001116]); Malacca, Bukit Bruang, R. Derry 574 [274 in
protologue] (SING [SING0240284]).
= Oxyceros bispinosus (Griff.) Tirveng., Nordic J. Bot. 3: 466 (1983); Ridsdale,
Reinwardtia 12: 296 (2008).
378. Randia hirsuta Ridl., J. Fed. Malay States Mus. 10: 110 (1920). – TYPE: Peninsular
Malaysia, Kelantan, Chaning Woods, February 1917, H.N. Ridley s.n. (lectotype SING
[SING0059557], designated by Wong (1984); ?isolectotype K [K000742851]).
= Benkara armigera (K.Schum.) Ridsdale, Reinwardtia 12: 298 (2008).
373
The Gentinales taxa of H.N. Ridley
379. Randia incurva Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 79 (1918). – TYPE:
Peninsular Malaysia, Penang, West Hill 1000 ft, May 1886, C. Curtis 818 (lectotype
SING [SING0059630, SING0059631, SING001134 – i.e. one specimen over three
sheets), designated by Wong (1984); K [K000742988]).
= Oxyceros bispinosus (Griff.) Tirveng., Nordic J. Bot. 3: 466 (1983).
380. Randia klossii Ridl., J. Fed. Malay States Mus. 10: 94 (1920), as ‘klossi’. – TYPE:
Thailand, Tasan, March 1919, C.B. Kloss 6872 (lectotype SING [SING0001135],
designated here; isolectotypes K [K001067650], SING [SING0059642]).
= Oxyceros kunstleri (King & Gamble) Tirveng., Nordic J. Bot. 3: 466 (1983);
Ridsdale, Reinwardtia 12: 294 (2008).
381. Randia oocarpa Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 79 (1918). –
Catunaregam oocarpa (Ridl.) Tirveng., Nordic J. Bot. 3: 456 (1983). – TYPE:
Peninsular Malaysia, Kedah, Langkawi, Burau, April 1911, H.N. Ridley 15816 (lectotype
K [K000786005], designated here; isolectotypes BM, SING [SING0197723]).
Additional syntypes: Peninsular Malaysia, Pahang, Pekan, 4 May 1890, H.N. Ridley
1118 (SING [SING0197720]); Pekan, Relau Tujor, July 1888, L. Wray 2599A (K
[K000786004]); Perak, Thaiping, March 1884, B. Scortechini 95 (BM, K); Kedah,
Langkawi, Coah, March 1892, C. Curtis 2799 (BM, K); Perlis, Kanga, March 1910,
H.N. Ridley 15007 (K, SING [SING0058016, SING0197722]).
382. Randia parvula Ridl., J. Fed. Malay States Mus. 10: 94 (1920). – Oxyceros
parvulus (Ridl.) T.Yamaz., J. Jap. Bot. 45: 339 (1970). – TYPE: Thailand, Koh Gah,
s.dat., C.B. Kloss 6608 (lectotype K [K000742849], designated by Wong (1984), or
possibly holotype).
= Benkara armigera (K.Schum.) Ridsdale, Reinwardtia 12: 298 (2008).
383. Randia pauciora Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 112 (1911). –
Oxyceros pauciorus (Ridl.) Tirveng., Nordic J. Bot. 3(4): 466 (1983). – TYPE:
Thailand, Pungah, July 1893, C. Curtis s.n. (lectotype SING (not traced), designated
by Wong (1984), or possibly holotype).
= Oxyceros kunstleri (King & Gamble) Tirveng., Nordic J. Bot. 3: 466 (1983).
384. Randia roxburghii Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 80 (1918). –
TYPE: Singapore, September 1822, N. Wallich 54 [EIC 8284D] (lectotype K-W
[K001125142], designated here). Additional syntypes: Singapore, September 1822,
N. Wallich 55 [EIC 8284D] (K-W [K001125141]); Singapore, T. Lobb (not traced);
374 Gard. Bull. Singapore 70 (2) 2018
Bangladesh, Chittagong, N. Wallich [EIC 8284B] (K-W [K001125139]); India, Assam,
Jenkins (not traced).
= Oxyceros kunstleri (King & Gamble) Tirveng., Nordic J. Bot. 3: 466 (1983).
385. Randia uncata Ridl., J. Straits Branch Roy. Asiat. Soc. 59 (1911) 113. – TYPE:
Thailand, Bangtaphan, 22 March 1890, A. Keith 101 (lectotype SING [SING0059629],
designated here, or possibly holotype).
= Oxyceros horridus Lour., Fl. Cochinch. 151 (1790); Ridsdale, Reinwardtia 12: 293
(2008).
386. Rennellia speciosa (Wall. ex Kurz) Hook.f. var. angustifolia Ridl., Fl. Malay
Penins. 2: 120 (1923). – TYPE: Peninsular Malaysia, Kedah, Gunong Bintang, C.B.
Kloss (not traced).
= Rennellia elliptica Korth., Ned. Kruidk. Arch. 2(2): 257 (1851).
387. Saprosma pubescens Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 23 (1912). –
TYPE: Peninsular Malaysia, Pahang, Tahan woods, 23 July 1891, H.N. Ridley 2210
(lectotype SING [SING0058739], designated here, or possibly holotype).
388. Saprosma pubescens Ridl. var. hirsuta Ridl., J. Straits Branch Roy. Asiat. Soc.
61: 23 (1912). – TYPE: Peninsular Malaysia, Malacca, foot of Ophir (on SING sheet)/
Gunong Ledang, lower part (on K sheet), June 1892, H.N. Ridley 3217 (lectotype
SING [SING0209190], designated here; isolectotype K [K000761914]).
389. Saprosma saxicola Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 22 (1912);
Davis & Bridson, Blumea 49: 60 (2004). – TYPE: Peninsular Malaysia, Perak, Bukit
Kamuning, February 1904, H.N. Ridley 11884 (lectotype K [K001067654], designated
here, or possibly holotype).
The sheet Perak, Bukit Kamuning, February 1904, H.N. Ridley s.n. (SING
[SING0058738]) could be from the same gathering, and hence an isolectotype, but
lacks a number.
390. Streblosa bracteata Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 64 (1911);
Bremekamp, J. Arnold Arbor. 28: 166 (1947). – TYPE: Borneo, Sarawak, Jambusan,
September 1905, H.N. Ridley 12437 (lectotype SING [SING0057803], designated
here). Additional syntype: Borneo, Sarawak, Bau, July 1903, H.N. Ridley 11741
(SING [SING0057802]).
375
The Gentinales taxa of H.N. Ridley
Razamandimbison et al. (2014) proposed an expanded genus Psychotria to include
a number of genera previously maintained as distinct. These included Streblosa.
The taxonomy of Streblosa is poorly known, so we refrain from any nomenclatural
alterations here.
391. Streblosa hirta Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 63 (1911). – TYPE:
Peninsular Malaysia, Pahang, Taka Tahan, 1893, H.N. Ridley s.n. (lectotype SING
[SING0241774], designated here).
= Streblosa polyantha Korth., Ned. Kruidk. Arch. 2(2): 246 (1851).
We have failed to locate any specimens from Penang or Perak, the only localities given
by Ridley. A specimen from Taka Tahan, Pahang, from 1893 is labelled as Streblosa
hirta in Ridley’s hand in SING and is here selected as lectotype.
392. Streblosa microcarpa Ridl., Fl. Malay Penins. 2: 148 (1923); Bremekamp, J.
Arnold Arbor. 28: 164 (1947). – TYPE: Peninsular Malaysia, Penang, 1822, N. Wallich
s.n. [EIC 8344] (lectotype K [K000761891], rst step designated by Bremekamp
(1947), second step designated here). Additional syntypes: Peninsular Malaysia, Perak,
Temengoh, July 1909, H.N. Ridley 14318 (SING [SING0240288]); Perak Gopeng,
November 1880, King’s Collector [Kunstler] 957 (K); Penang, road to Pantai Acheh,
August 1885, C. Curtis 405 (K, SING [SING0240287, SING0240286]).
Bremekamp (1947) referred to the Wallich collection under the number 8344 as the
type of Streblosa microcarpa Ridl. but did not state in which herbarium the type was
deposited, only that GH contained an example. A second step lectotypication is
therefore made here.
393. Streblosa pubescens Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 63 (1911). – TYPE:
Peninsular Malaysia, Penang, Balik Pulau, June 1898, H.N. Ridley 9425 (lectotype
SING [SING0240292], designated here). Additional syntypes: Peninsular Malaysia,
Johore, Gunong Panti, December 1892, H.N. Ridley s.n. (SING [SING0240290]);
Perak, Temengoh Woods, H.N. Ridley s.n. (SING [SING0240291]); Perak, Dindings
at Telok Serah, March 1895, H.N. Ridley s.n. (SING [SING0240289]).
= Streblosa polyantha Korth., Ned. Kruidk. Arch. 2(2): 246 (1851).
394. Stylocoryna sylvicola Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 69 (1923),
as ‘Stylocoryne’. – Tarenna sylvicola (Ridl.) Merr., Contr. Arnold Arbor. 8: 158
(1934). – TYPE: Sumatra, Berastagi, February 1921, H.N. Ridley s.n. (lectotype K
[K000763172], designated here, or possibly holotype).
376 Gard. Bull. Singapore 70 (2) 2018
395. Tarenna arborescens Ridl., J. Bot. 72: 271 (1934). – TYPE: Borneo, Sarawak,
Kuching, 3 March 1893, G.D. Haviland 2979 (lectotype K [K001129556], designated
here).
396. Tarenna axillaris Ridl., J. Bot. 72: 273 (1934). – Tarennoidea axillaris (Ridl.)
Tirveng. & Sastre, Mauritius Inst. Bull. 8(4): 91 (1979). – TYPE: Borneo, Sarawak,
Kuching River, 1893, G.D. Haviland 2973 (lectotype K [K000173182], designated by
Tirvengadum & Sastre (1979), or possibly holotype).
397. Tarenna bartlettii Ridl., J. Bot. 72: 273 (1934). – TYPE: Borneo, Sarawak,
Kuching, 21 April 1893, E. Bartlett s.n. (lectotype K [K000173174], designated here,
or possibly holotype).
398. Tarenna calcarea Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 295 (1922). – TYPE:
Peninsular Malaysia, Perak, Ipoh, limestone rocks, 26 January 1921, H.N. Ridley s.n.
(lectotype K [K001067655], designated here, or possibly holotype).
= Tarenna adangensis (Ridl.) Ridl., Fl. Malay Penins. 2: 107 (1923); Wong, Tree Fl.
Malaya 4: 412 (1989).
399. Tarenna crassifolia Ridl., J. Bot. 72: 257 (1934). – TYPE: Borneo, Sarawak,
Kuching, G.D. Haviland 1053 [Garai leg.] [e.b.d.h] (lectotype K [K000173180],
designated here, or possibly holotype).
400. Tarenna debilis Ridl., J. Bot. 72: 272 (1934). – TYPE: Borneo, Sarawak, Baram,
July 1894, G.D. Haviland & C. Hose 3428A (lectotype K [K000173179], designated
here, or possibly holotype).
401. Tarenna evansii Ridl., Fl. Malay Penins. 2: 103 (1923). – TYPE: Peninsular
Malaysia, Pahang, Gunong Senyum, Evans (not traced).
402. Tarenna glabra Ridl., J. Fed. Malay States Mus. 10: 141 (1920), nom. illeg.,
non Merr. (1914). – Tarenna glaberrima Ridl., J. Bot. 72: 257 (1934). – TYPE:
Peninsular Malaysia, Kelantan, Chaning, 31 January 1917, H.N. Ridley s.n. (lectotype
K [K001129553], designated here; isolectotype K [K001129554]).
377
The Gentinales taxa of H.N. Ridley
403. Tarenna hosei Ridl., J. Bot. 72: 272 (1934). – TYPE: Borneo, Sarawak, Baram,
Mt Trekan 1000 ft, July 1895, C. Hose 636 (lectotype K [K000173177], designated
here, or possibly holotype).
404. Tarenna lancifolia Ridl., Fl. Malay Penins. 5: 316 (1925). – TYPE: Peninsular
Malaysia, Kedah, Kedah Peak, 4 August 1919, Md. Haniff & Md. Nur SFN 4954
(lectotype SING [SING0058436], designated here by Ng Xin Yi & Wong Khoon Meng;
isolectotypes K [K001067652], KEP [KEP181745]). Additional syntype: Peninsular
Malaysia, Pahang, Fraser {Fraser’s} Hill, [21 September 1922], I.H. Burkill & R.E.
Holttum SFN 8686 (K [K001067651), SING [SING0058811]).
= Tarenna adpressa (King) Merr., Philipp. J. Sci. 17: 472 (1921).
405. Tarenna oblanceolata Ridl., J. Bot. 72: 256 (1934). – TYPE: Borneo, Sarawak, fort
on Matang, G.D. Haviland 1032 [Garai leg.] (lectotype K [K000173173], designated
here). Additional syntype or possible isolectotypes: Borneo, Sarawak, fort on Matang,
G.D. Haviland 1032 [Garai leg.] [e.d.e.b.] (K [K000173172], SING [SING0240293]).
406. Tarenna rudis Ridl., Fl. Malay Penins. 2: 103 (1923). – TYPE: Peninsular
Malaysia, Selangor, Kuala Lumpur, Weld’s Hill Forest Reserve, 27 January 1919,
Forester Hamid, For. Dept. F.M.S. eld no. 2952 (lectotype K [K001067660],
designated here; isolectotype SING [SING0240278]). Additional syntype: Peninsular
Malaysia, Selangor, Kuala Lumpur, Weld’s Hill Forest Reserve, 20 December 1918,
Climber Ahmad, For. Dept. F.M.S. eld no. 2593 (K [K001067659]).
The 1919 collection was mistakenly given as 2592 in the protologue.
407. Timonius diffusus Ridl., J. Fed. Malay States Mus. 4: 32 (1909); Wong, Kew
Bull. 43: 503 (1988). – TYPE: Peninsular Malaysia, Pahang, Telom, November 1908,
H.N. Ridley 13611 (lectotype SING [SING0058053, in part, including female material
only], designated by Wong (1988a); isolectotype K [K000763415, in part]).
408. Timonius hirsutus Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 81 (1918), nom.
illeg., non Merrill (1917). – Timonius atropurpureus Craib, Fl. Siam. 2: 132 (1932). –
Timonius ridleyi Merr., J. Arnold Arbor. 18: 132 (1937), nom. illegit, super. – Antirhea
atropurpurea (Craib) Chaw in Wong, Kew Bull. 43: 493 (1988). – TYPE: Peninsular
Malaysia, Kedah, Langkawi Islands, C. Curtis 2544 (lectotype SING [SING0209187],
rst step designated by Wong (1988b), second step designated here; isolectotype SING
[SING0209188]). Additional syntype: Peninsular Malaysia, Kedah, Langkawi Islands,
Pulau Dayang Bunting, November 1916, H.C. Robinson 6229 (K [K000763444]).
378 Gard. Bull. Singapore 70 (2) 2018
Wong (1988b) designated Curtis 2544 in SING as the lectotype, but as there are two
specimens under this number in SING, a second-step lectotypication is made here,
Although Curtis 2544 only comprises leafy branches without owering or fruiting
material, it is sufciently diagnostic and no epitype is proposed; further, the other
syntype Robinson 6229 is white-hairy and in that aspect atypical of what Ridley and
Craib had intended.
409. Timonius montanus Ridl., J. Linn. Soc., Bot. 38: 312 (1908); Wong, Kew Bull.
43: 506 (1988). – TYPE: Peninsular Malaysia, Pahang, Gunong Tahan 5–6000 ft, 9 July
1905, L. Wray & H.C. Robinson 5499 (lectotype SING [SING 0209197], designated
by Wong (1988a); isolectotypes BM [BM001190970], K [K000763585]).
410. Timonius oreophilus Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 22 (1912);
Wong, Kew Bull. 43: 506 (1988). – TYPE: Peninsular Malaysia, Selangor, Bukit
Hitam, 21 May 1896, H.N. Ridley 7399 (lectotype, designated by Wong (1988a),
SING [SING0209198]; isolectotype K [K000763584]). Additional syntypes:
Peninsular Malaysia, Kedah, Kedah Peak (a form with very narrow leaves), Unknown
collector (not traced); Pahang, Kluang Terbang, 1900, W.D. Barnes 10886 (SING
[SING0209199]); Selangor, Bukit Hitam, May 1896, H.N. Ridley 7397 (K, SING
[SING0209200]); Malacca, Mt. Ophir, Hullett 837 (not traced).
411. Timonius vaccinioides Wernham ex Ridl., Hooker’s Icon. Pl. 31: t. 3054 (1916
[June 1916]); Wernham, Trans. Linn. Soc., Bot. 9: 74 (1916 [August 1916]). – TYPE:
New Guinea, Utakwa River to Mt. Carstensz [Camp Pundok Padang], 18 February
1913, C.B. Kloss s.n. (holotype BM [BM000945275]).
412. Timonius wallichianus (Korth.) Valeton var. penangianus Ridl., Fl. Malay Penins.
2: 116 (1923). – Timonius corneri K.M.Wong var. penangianus (Ridl.) K.M.Wong,
Kew Bull. 43: 502 (1988). – TYPE: Peninsular Malaysia, Penang, Balik Pulau, June
1898, H.N. Ridley 9398 (lectotype SING [SING0058052], designated by Wong
(1988a)). Additional syntypes: Peninsular Malaysia, Penang, Government Hill, March
1890, C. Curtis s.n. (SING); Penang, N. Wallich s.n. [EIC 6217] (K[×2]).
413. Uncaria lanosa Wall. var. parviora Ridl., J. Straits Branch Roy. Asiat. Soc.
59: 109 (1911). – Uncaria parviora (Ridl.) Ridl., J. Straits Branch Roy. Asiat. Soc.
79: 75 (1918). – TYPE: Peninsular Malaysia, Perlis, Chupeng, March 1910, H.N.
Ridley 15019 (lectotype K [K00072990], designated by Turner (2018); isolectotypes
L (fragment) [L0001518], SING [SING0240294]).
= Uncaria homomalla Miq., Fl. Ned. Ind. 2: 343 (1857).
379
The Gentinales taxa of H.N. Ridley
414. Urophyllum bracteolatum Ridl., J. Bot. 70: 221 (1932). – TYPE: Borneo,
Sarawak, Mt Singhi, 11 December 1892, G.D. Haviland 2016 [Garai leg.] (lectotype
K [K000740847], designated by Bremekamp (1940b), or possibly holotype).
415. Urophyllum bullatum Ridl., J. Bot. 70: 195 (1932). – TYPE: Borneo, Sarawak,
Matang, January 1915, H.N. Ridley s.n. (lectotype K [K000740815], designated here;
isolectotype BM [BM000945203]).
416. Urophyllum capituligerum Ridl., J. Bot. 70: 222 (1932). – Pleiocarpidia
capituligera (Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 216 (1940). – TYPE:
Borneo, Sarawak, Saribas, Paku, 17 November 1893, G.D. Haviland & C. Hose 3415
(lectotype K [K000740771], designated by Bremekamp (1940a); isolectotypes SAR,
U [U0006341]).
417. Urophyllum castaneum Ridl., J. Bot. 70: 220 (1932). – TYPE: Borneo, Sarawak,
1865–1868, O. Beccari P.B. 1424 (lectotype K [K001067644], designated here, or
possibly holotype; isolectotype FI-B!).
418. Urophyllum cephalotes Ridl., J. Bot. 70: 222 (1932). – Pleiocarpidia cephalotes
(Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 223 (1940). – TYPE: Borneo, Sarawak,
Kuching, G.D. Haviland 1043 [Garai leg.] [e.d.b.b.] (lectotype K [K000740767],
designated by Bremekamp (1940a); isolectotype SAR!).
419. Urophyllum congestiorum Ridl., J. Bot. 70: 224 (1932). – TYPE: Borneo,
Sarawak, Kuching, 16 February 1894, G.D. Haviland & C. Hose 3416 (lectotype K
[K001067642], designated here; isolectotype BM [BM001118895]).
420. Urophyllum coriaceum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 18 (1912), nom.
illegit, non Miq. (1861). – Urophyllum leucophlaeum Ridl., J. Bot. 70: 196 (1932). –
TYPE: Peninsular Malaysia, Malacca, Mt Ophir, December 1898, H.N. Ridley 10105
(lectotype SING [SING0240295], designated here, or possibly holotype).
421. Urophyllum coriaceum Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 78 (1918),
nom. illegit, non Miq. (1861), nec Ridl. (1912). – TYPE: Peninsular Malaysia,
Pahang, Gunong Tahan, Wray’s Camp, July 1911, H.N. Ridley 16247 (lectotype K
[K001067643], designated here). Additional syntype: Peninsular Malaysia, Pahang,
Gunong Tahan, July 1911, H.N. Ridley 16070 (K [K000740836]).
= Urophyllum leucophlaeum Ridl., J. Bot. 70: 196 (1932).
380 Gard. Bull. Singapore 70 (2) 2018
422. Urophyllum creaghii Ridl., J. Bot. 70: 193 (1932). – Praravinia creaghii (Ridl.)
Bremek., Recueil Trav. Bot. Néerl. 37: 253 (1940). – TYPE: Borneo, British North
Borneo {Sabah}, s.dat., C.V. Creagh s.n. (lectotype K [K000740806], designated by
Bremekamp (1940b), or possibly holotype).
423. Urophyllum grandifolium Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 68 (1923).
– Maschalocorymbus grandifolius (Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 182
(1940). – TYPE: Sumatra, Berastagi, 8 February 1921, H.N. Ridley s.n. (lectotype K
[K000740722], designated here, or possibly holotype).
424. Urophyllum havilandii Ridl., J. Bot. 70: 194 (1932). – Praravinia havilandii
(Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 263 (1940). – TYPE: Borneo, [Sabah],
Gaya Island, G.D. Haviland 1435 [e.k.d.] (lectotype K [K000740798], designated by
Bremekamp (1940b)). Additional syntype or possible isolectotype: Borneo, [Sabah],
Gaya Island, 19 April 1892, G.D. Haviland 1435 (SING [SING0058125]).
425. Urophyllum lasiocarpum Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 19 (1912).
– TYPE: Peninsular Malaysia, Perak, Bujong Malacca, September 1898, H.N. Ridley
9726 (lectotype K [K000740844], designated here).
The type sheet in K is labelled as part of the type from Singapore. We have not been
able to trace the remaining specimen in SING, so here designate the piece in K as
lectotype.
426. Urophyllum longipetalum Ridl., J. Bot. 70: 193 (1932). – TYPE: Borneo,
Sarawak, Kuching, 16 September 1892, G.D. Haviland 1662 [Garaman leg.] (lectotype
K [K001067643], designated by Bremekamp (1940b), or possibly holotype).
427. Urophyllum lowei Ridl., J. Bot. 70: 195 (1932). – Rhaphidura lowei (Ridl.)
Bremek., Recueil Trav. Bot. Néerl. 37: 187 (1940), as ‘lowii’. – TYPE: Borneo,
Sarawak, Sarawak River, G.D. Haviland 1837 [b.z.e.v.] (lectotype K [K000740779],
designated here). Additional syntypes: Borneo, Sarawak, Near Kuching, 1892,
G.D. Haviland 1837 (K [K000740780]); ibid., October 1892, G.D. Haviland 1837
(SING [SING0058121]); ibid., 12 November 1892, G.D. Haviland 1837 (BM
[BM000945237]).
There are various Haviland 1837 sheets available, which vary in details of collection
date. An undated (but communicated in 1890) specimen in K, which is annotated as
type in Ridley’s hand, is here designated as lectotype.
381
The Gentinales taxa of H.N. Ridley
Ridley referred to the collector as Lowe not Low, so Bremekamp’s change to
lowii should not be followed.
428. Urophyllum macranthum Ridl., J. Malayan Branch Roy. Asiat. Soc. 1: 69 (1923).
– Leucolophus macranthus (Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 188 (1940).
– TYPE: Sumatra, Berastagi Woods, 11 February 1921, H.N. Ridley s.n. (lectotype K
[K000740778], designated here, or possibly holotype).
429. Urophyllum melanocarpum Ridl., J. Bot. 70: 226 (1932). – TYPE: Borneo,
Sarawak, Mt Bongoh near Tegura, 19 December 1892, G.D. Haviland 2004 (lectotype
K [K000740874], designated here, or possibly holotype).
430. Urophyllum motleyi Ridl., J. Bot. 70: 225 (1932). – TYPE: Borneo, [Kalimantan],
Bangarmassing, 1857–1858, J. Motley 1109 (lectotype K [K000740878], designated
here, or possibly holotype).
431. Urophyllum neriifolium Ridl., J. Bot. 70: 225 (1932). – TYPE: Borneo, Sarawak,
Matang, G.D. Haviland 84 [669] [c.l.p.f.] (lectotype K [K000740881], designated
here, or possibly holotype).
432. Urophyllum paniculatum Ridl., J. Bot. 70: 223 (1932). – TYPE: Borneo,
Sarawak, Kuching, 22 May 1893, G.D. Haviland 2986 (lectotype BM [BM001118896],
designated here; isolectotypes SAR, SING [SING0057807], U [U0006343]).
Bremekamp (1940b) effectively lectotypied the species to the Haviland 2986
duplicate in K, but this specimen could not be found. We therefore propose the sheet
in BM as lectotype.
433. Urophyllum pellacalyx Ridl., J. Bot. 70: 194 (1932). – Crobylanthe pellacalyx
(Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 192 (1940). – TYPE: Borneo, Sarawak,
Kuching, 8 March 1893, G.D. Haviland 2965 (lectotype K [K000740776], designated
here, or possiby holotype).
434. Urophyllum pilosum Ridl., J. Bot. 70: 219 (1932). – Pleiocarpidia pilosa (Ridl.)
Bremek., Recueil Trav. Bot. Néerl. 37: 226 (1940). – TYPE: Borneo, Sarawak, Rejang,
Kapit, August 1893, G.D. Haviland 2962 [Kalong leg.] (lectotype K [K000740850],
designated by Bremekamp (1940a); isolectotype SAR).
382 Gard. Bull. Singapore 70 (2) 2018
435. Urophyllum rigidum Ridl., J. Bot. 70: 196 (1932). – TYPE: Peninsular Malaysia,
Selangor-Pahang border, Fraser {Fraser’s} Hill, 16–30 September 1922, I.H. Burkill
& R.E. Holttum SFN 8668 (lectotype K [K000740835), designated here; isolectotypes
A [A00094717], SING [SING0057806]).
= Urophyllum leucophloeum Ridl., J. Bot. 70: 196 (1932).
436. Urophyllum sericeum Ridl., J. Bot. 70: 221 (1932). – TYPE: Java, Mt Gede,
14 February 1915, H.N. Ridley s.n. (lectotype K [K000740724], designated here;
isolectotype BM [BM000945234]).
= Urophyllum corymbosum (Blume) Korth., Ned. Kruidk. Arch. 2(2): 194 (1851).
437. Urophyllum sessiliorum Ridl., Fl. Malay Penins. 2: 69 (1923). – TYPE:
Peninsular Malaysia, Negri Sembilan, Bukit Tangga, 22 December 1920, H.N. Ridley
s.n. (lectotype K [K000740824], designated here, or possibly holotype).
438. Urophyllum shelfordii Ridl., J. Bot. 70: 197 (1932). – TYPE: Borneo, Sarawak,
1865–1868, O. Beccari P.B. 1434 (lectotype K [K000740777], designated here;
isolectotype FI-B).
= Crobylanthe pellacalyx (Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 192 (1940).
439. Urophyllum sumatranum Ridl., J. Bot. 70: 197 (1932). – Didymopogon
sumatranum (Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 181 (1940). – TYPE:
Sumatra, Mt Singalan, June–July 1878, O. Beccari P.S. 348 (lectotype K [K000740727],
designated here; isolectotypes BM [BM000945233], FI-B, L).
440. Urophyllum vulcanicum Ridl., J. Bot. 70: 224 (1932). – TYPE: Sumatra, Sibayak
Volcano woods, Berastagi, Karoo Plain, February 1921, H.N. Ridley s.n. (lectotype K
[K000740879], designated here, or possibly holotype).
441. Urophyllum yatesii Ridl., J. Bot. 70: 220 (1932). – Maschalocorymbus yatesii
(Ridl.) Bremek., Recueil Trav. Bot. Néerl. 37: 182 (1940). – TYPE: Sumatra, East
Coast, s.dat., H.S. Yates 1325 (lectotype K [K000740721], designated here; isolectotype
NY [NY00133475]).
442. Webera adangensis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 54 (1912). – Tarenna
adangensis (Ridl.) Ridl., Fl. Malay Penins. 2: 107 (1923). – TYPE: Thailand, Pulau
Adang, April 1911, H.N. Ridley 15897 (lectotype SING [SING0058812], designated
here; isolectotypes BM [BM001118893, BM001118894], K [K001067656]).
383
The Gentinales taxa of H.N. Ridley
443. Webera insularis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 55 (1912). – Tarenna
insularis (Ridl.) Ridl., Fl. Malay Penins. 2: 107 (1923). – TYPE: Thailand, Pulau
Adang, Pulau Nipis, April 1911, H.N. Ridley 15896 (lectotype SING [SING0058813],
designated here; isolectotypes BM [BM001118891], K [K001067658]).
444. Webera napieri Ridl., J. Fed. Malay States Mus. 4: 34 (1909). – Pavetta napieri
(Ridl.) Bremek., Repert. Spec. Nov. Regni Veg. 37: 96 (1934). – TYPE: Peninsular
Malaysia, Negeri Sembilan, Bukit Tanga, 10 February 1903, W.J. Napier s.n.
(lectotype SING [SING0058435], designated by Bremekamp (1934)). Additional
syntype: Peninsular Malaysia, Pahang, Telom, 1908, H.N. Ridley 13639 (K, SING
[SING0240280]).
445. Webera pulchra Ridl., J. Fed. Malay States Mus. 4: 33 (1909). – Tarenna pulchra
(Ridl.) Ridl., Fl. Malay Penins. 2: 104 (1923). – TYPE: Peninsular Malaysia, Pahang,
Telom, November 1908, H.N. Ridley 13638 (lectotype SING [SING0058440],
designated here; isolectotypes BM [BM001118892], K [K001067661], SING
[SING0058439]).
446. Webera salicina Ridl., J. Fed. Malay States Mus. 4: 31 (1909). – Tarenna salicina
(Ridl.) Ridl., Fl. Malay Penins. 2: 104 (1923). – Pavetta salicina (Ridl.) Bremek., Repert.
Spec. Nov. Regni Veg. 37: 103 (1934). – TYPE: Peninsular Malaysia, Pahang, Telom,
8 November 1908, H.N. Ridley 13610 (lectotype SING [SING0058845], designated
by Bremekamp (1934); isolectotypes BM [BM001190981], K [K000763506]).
447. Xanthophytum biceps Ridl., Fl. Malay Penins. 5: 316 (1925). – TYPE: Peninsular
Malaysia, Pahang, Ulu Tahan, 7 December 1924, I.H. Burkill & Md. Haniff SFN 17463
(lectotype K [K000761892], designated here).
The species was excluded from Xanthophytum Reinw. ex Blume by Axelius (1990). Its
correct generic position remains uncertain but is treated here as accepted, pending a
new combination being made or it being placed in synonymy.
448. Xanthophytum capitatum Ridl., J. Fed. Malay States Mus. 10: 139 (1920), nom.
illeg., non Valeton (1910). – Xanthophytum capitellatum Ridl., Fl. Malay Penins. 5:
316 (1925). – TYPE: Peninsular Malaysia, Pahang, Pehi River woods, Chaning, 3
February 1917, H.N. Ridley s.n. (lectotype K [K000760618], designated by Axelius
(1990), or possibly holotype).
384 Gard. Bull. Singapore 70 (2) 2018
449. Xanthophytum rupestre Ridl., J. Straits Branch Roy. Asiat. Soc. 54: 44 (1910). –
Aleisanthia rupestris (Ridl.) Ridl., J. Fed. Malay States Mus. 10: 138 (1920); Tange,
Nordic J. Bot. 16: 565 (1996). – TYPE: Peninsular Malaysia, Selangor, Klang Gates,
August 1908, H.N. Ridley 13414 (lectotype K [K000174147], designated by Tange
(1996); isolectotype BM [BM001190932]).
450. Zeuxanthe moultonii Ridl., Bull. Misc. Inform. Kew 1939: 610 (1940). – TYPE:
Borneo, Sarawak, Upper Baram, Lio Matu, 16 October 1920, J.C. Moulton SFN 6719
(holotype K [K000763853]; isotype SING [SING0059766]).
= Prismatomeris beccariana (Baill. ex K.Schum.) J.T.Johanss., Opera Bot. 94: 57
(1987).
451. Zuccarinia cordata Ridl., Bull. Misc. Inform. Kew 1926: 68 (1926). – TYPE:
Sumatra, Sipora, 8 October 1924, C.B. Kloss 14641 (lectotype K [K000763112],
designated here; isolectotypes K [K000763113], SING [SING0240297]).
= Zuccarinia macrophylla Blume, Bijdr. Fl. Ned. Ind. 16: 1007 (1826); Wong, Tree
Fl. Malaya 4: 425 (1989).
xi. Argostemma sect. Pomangium (Reinw.) Ridl., J. Bot. 65: 20 (1927). – Pomangium
Reinw., Syll. Pl. Nov. 2: 10 (1828).
= Argostemma Wall. in Roxb., Fl. Ind. 2: 324 (1824).
xii. Asemanthia (Stapf) Ridl., Bull. Misc. Inform. Kew 1939: 599 (1940). – Mussaenda
section Asemanthia Stapf, Trans. Linn. Soc. ser. 2, 4 (1894) 173.
= Mussaenda L., Sp. Pl. 177 (1753).
xiii. Porterandia Ridl., Bull. Misc. Inform. Kew 1939: 593 (1940). – Randia section
Anisophyllea Benth. & Hook.f., Gen. Pl. 2(1): 89 (1873).
xiv. Aleisanthia rupestris (Ridl.) Ridl., J. Fed. Malay States Mus. 10: 138 (1920), as
‘rupestre’; Tange, Nordic J. Bot. 16: 565 (1996). – Xanthophytum rupestre Ridl., J.
Straits Branch Roy. Asiat. Soc. 54: 44 (1910).
xv. Amaracarpus saxicola (Ridl.) Ridl., J. Fed. Malay States Mus. 10: 143 (1920). –
Saprosma saxicola Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 22 (1912); Davis &
Bridson, Blumea 49: 60 (2004).
385
The Gentinales taxa of H.N. Ridley
xvi. Argostemma lanceolarium Ridl., J. Bot. 65: 36 (1927). – Argostemma lanceolatum
Valeton, Icon. Bogor. 4: t. 388 (1914), nom. illeg., non Ridl. (1909).
xvii. Argostemma stellare Ridl., J. Bot. 65: 31 (1927). – Argostemma stellatum Ridl.,
J. Malayan Branch Roy. Asiat. Soc. 1: 67 (1923), nom. illeg., non Craib (1911).
xviii. Argostemmella brachyanthera (Stapf) Ridl., J. Bot. 65: 41 (1927). – Argostemma
brachyantherum Stapf, Trans. Linn. Soc. London, Bot. 4(2): 168 (1894); Bremer,
Ann. Missouri Bot. Gard. 76: 36 (1989).
xix. Argostemmella gracilis (Stapf) Ridl., J. Bot. 65: 41 (1927). – Argostemma gracile
Stapf, Trans. Linn. Soc. London, Bot. 4(2): 168 (1894); Bremer, Ann. Missouri Bot.
Gard. 76: 35 (1989).
xx. Asemanthia coccinea Ridl., Bull. Misc. Inform. Kew 1939: 600 (1940). –
Mussaenda coccinea Stapf, Trans. Linn. Soc. London, Bot. 4: 170 (1894), nom. illeg.,
non (Vahl) Poir. (1797).
A new name in Mussaenda will soon be published by Govaerts (pers. comm.).
xxi. Asemanthia maingayi (Hook.f.) Ridl., Bull. Misc. Inform. Kew 1939: 600 (1940).
– Acranthera maingayi Hook.f., Fl. Brit. India 3: 92. 1880.
= Mussaenda maingayi (Hook.f.) Hemsl. ex T.Durand & B.D.Jacks., Index Kew.
Suppl. 1: 284 (1903).
xxii. Asemanthia spectabilis (Ridl.) Ridl., Bull. Misc. Inform. Kew 1939: 600 (1940).
– Mussaenda spectabilis Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 78 (1918).
xxiii. Borreria laevicaulis (Miq.) Ridl., J. Straits Branch Roy. Asiat. Soc. 86: 298
(1922). – Bigelovia laevicaulis Miq., Fl. Ned. Ind. 2: 335 (1857).
= Spermacoce ocymoides Burm.f., Fl. Indica 34 (1768).
xxiv. Gaertnera acuminata Benth. var. oxyphylla (Benth.) Ridl., Fl. Malay Penins.
2: 428 (1923). – Gaertnera oxyphylla Benth., J. Proc. Linn. Soc., Bot. 1: 112 (1857).
= Gaertnera junghuhniana Miq., Fl. Ned. Ind. 2: 383 (1857); Malcomber & Taylor,
Ann. Missouri Bot. Gard. 96: 626 (2009).
386 Gard. Bull. Singapore 70 (2) 2018
xxv. Ixora gigantea (King & Gamble) Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 84
(1918); nom. illeg., non Rechinger (1907). – Ixora grandifolia Zoll. & Moritzi var.
gigantea King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 82 (1904).
= Ixora grandifolia Zoll. & Moritzi in A.Moritzi, Syst. Verz. 65 (1846); Wong, Tree
Fl. Malaya 4: 364 (1989).
In a very concisely written passage, Ridley proposed Ixora gigantea as the name of a
species split off from the Ixora grandifolia complex. He mentions the variety gigantea
of King and Gamble but does not make it explicit that he is raising this to species rank.
However, this does seem implicit and it is acceptable to consider it as a comb. & stat.
nov., as otherwise the species name would be validated by the description provided
by King and Gamble. The combination remains illegitimate as it is a later homonym.
xxvi. Ixora humilis King & Gamble var. scortechinii (King & Gamble) Ridl., Fl. Malay.
Penins. 2: 95 (1923). – Ixora scortechinii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2,
Nat. Hist. 73: 80 (1904).
xxvii. Ixora pendula Jack var. opaca (R.Br. ex G.Don) Ridl., Fl. Malay. Penins. 2: 96
(1923). – Ixora opaca R.Br. ex G. Don, Gen. Hist. 3: 573 (1834); Bremekamp, Bull.
Jard. Bot. Buitenzorg ser. III, 14: 291 (1937).
xxviii. Ixora salicina Ridl., J. Bot. 72: 253 (1934). – Ixora stenophylla Ridl., Trans.
Linn. Soc. London, Bot. 3(9): 311 (1893), nom. illeg., non (Korth.) Kuntze (1891).
= Ixora lobbii Loudon var. stenophylla Corner, Gard. Bull. Straits Settlem. 11: 216
(1941).
xxix. Lasianthus maingayi Hook.f. var. subglaber (King) Ridl., Fl. Malay Penns. 2:
160 (1923), as ‘subglabra’. – Lasianthus avicans King & Gamble var. subglaber
King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 116 (1904).
= Lasianthus reticulatus Blume, Bijdr. Fl. Ned. Ind. 16: 1000 (1826).
xxx. Lasianthus pilosus Wight var. retosus (Wight) Ridl., Fl. Malay Penins. 2: 157
(1923), as ‘setosus’. – Lasianthus retosus Wight, Calcutta J. Nat. Hist. 6: 506 (1846).
= Lasianthus pilosus Wight, Calcutta J. Nat. Hist. 6: 506 (1846).
xxxi. Morinda elliptica (Hook.f.) Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 86
(1918). – Morinda citrifolia L. var. elliptica Hook.f., Fl. Brit. India 3: 156 (1880).
387
The Gentinales taxa of H.N. Ridley
xxxii. Morinda ridleyi (King & Gamble) Ridl., Fl. Malay Penins. 2: 118 (1923). –
Morinda umbellata L. var. ridleyi King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat.
Hist. 73: 88 (1904). – Gynochthodes ridleyi (King & Gamble) Razam. & B.Bremer,
Adansonia, sér. 3, 33: 294 (2011).
xxxiii. Morinda scortechinii Ridl., Fl. Malay Penins. 2: 118 (1923). – Morinda
umbellata L. var. scortechinii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist.
73: 88 (1904).
= Gynochthodes umbellata (L.) Razam. & B.Bremer, Adansonia, sér. 3, 33: 296
(2011).
xxxiv. Mycetia ava (Ridl.) Ridl., Fl. Malay Penins. 2: 64 (1923). – Adenosacme ava
Ridl., J. Fed. Malay States Mus. 4: 29 (1909).
xxxv. Mycetia lanceolata Ridl., Fl. Malay Penins. 2: 64 (1923). – Adenosacme
lanceolata Ridl., J. Fed. Malay States Mus. 4: 29 (1909), nom. illeg., non Miq. (1857).
= Mycetia fasciculata (Blume) Blume ex Korth., Ned. Kruidk. Arch. 2(2): 118 (1851).
xxxvi. Mycetia scortechinii (King & Gamble) Ridl., Fl. Malay Penins. 2: 63 (1923). –
Adenosacme scortechinii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72:
197 (1904).
= Mycetia cauliora Reinw., Syll. Pl. Nov. 2: 9 (1825).
xxxvii. Ophiorrhiza fruticosa Ridl., Fl. Malay Penins. 2: 41 (1923). – Ophiorrhiza
fruticulosa Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 15 (1912), nom. illeg., non
Nadeaud (1873).
xxxviii. Ophiorrhiza rosacea Ridl., Fl. Malay Penins. 2: 38 (1923). – Ophiorrhiza
rosea Ridl., J. Straits Branch Roy. Asiat. Soc. 57: 55 (1911), nom. illeg., non Hook.f.
(1880).
xxxix. Perakanthus pauciorus Robyns ex Ridl., Fl. Malay Penins. 5: 317 (1925). –
Canthium pauciorum King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73(3):
58 (1904), nom. illeg., non Blanco (1837), nec (Schweinf. ex Hiern) Baill. (1878),
(Klotzsch ex Eckl. & Zeyh.) Kuntze (1898). – TYPE: Perak, s.dat., Scortechini 62
(lectotype SING [SING0212321], designated here; isolectotypes SING [SING0212322,
SING0212323]).
388 Gard. Bull. Singapore 70 (2) 2018
This species requires a new name when the appropriate genus designation becomes
clear.
xl. Perakanthus velutinus (Ridl.) Robyns ex Ridl., Fl. Malay Penins. 5: 317 (1925);
Wong & Puff, Sandakania 16: 42 (2005). – Canthium velutinum Ridl., J. Straits Branch
Roy. Asiat. Soc. 61: 19 (1912).
xli. Petunga oribunda (King) Ridl., Fl. Malay Penins. 2: 85 (1923). – Petunga
roxburghii var. oribunda King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72: 223 (1904).
= Hypobathrum racemosum (Roxb.) Kurz, Prelim. Rep. Forest Pegu, App. B 59
(1875).
xlii. Porterandia anisophylla (Jack ex Wall.) Ridl., Bull. Misc. Inform. Kew 1939:
594 (1940). – Gardenia anisophylla Jack ex Roxb., Fl. Ind. 2: 561 (1824).
xliii. Porterandia scortechinii (King & Gamble) Ridl., Bull. Misc. Inform. Kew 1939:
595 (1940). – Randia scortechinii King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat.
Hist. 72: 210 (1904).
xliv. Porterandia subsessilis (Valeton) Ridl., Bull. Misc. Inform. Kew 1939: 595
(1940). – Gardenia anisophylla Jack ex Roxb. var. subsessilis Valeton, Bot. Jahrb.
Syst. 44: 558 (1910).
xlv. Prismatomeris javanica (Valeton) Ridl., Bull. Misc. Inform. Kew 1939: 606
(1940); Johansson, Opera Bot. 94: 52 (1987). – Prismatomeris albidiora Thwaites
var. javanica Valeton, Bull. Inst. Bot. Buitenzorg 8: 6 (1901).
xlvi. Prismatomeris lepidophloia (Miq.) Ridl., Bull. Misc. Inform. Kew 1939: 601
(1940). – Coffea lepidophloia Miq., Fl. Ned. Ind., Eerste Bijv. 548 (1861).
= Prismatomeris glabra (Korth.) Valeton, Bot. Jahrb. Syst. 44: 569 (1910).
xlvii. Prismatomeris neurophylla (Miq.) Ridl., Bull. Misc. Inform. Kew 1939: 606
(1940). – Coffea neurophylla Miq., Fl. Ned. Ind. 2: 1079 (1859).
= Prismatomeris glabra (Korth.) Valeton, Bot. Jahrb. Syst. 44: 569 (1910).
389
The Gentinales taxa of H.N. Ridley
xlviii. Psychotria obovata Wall. ex Ridl., Fl. Malay Penins. 2: 129 (1923), nom. illeg.,
non Ruiz & Pavon (1799), nec Hemsl. (1881). – Psychotria polycarpa (Miq.) Hook.f.,
Fl. Brit. India 3: 165 (1880).
xlix. Psychotria pachyphylla Ridl., Fl. Malay Penins. 2: 129 (1923). – Psychotria
sarmentosa Blume var. pachyphylla King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat.
Hist. 74(1): 6 (1906).
l. Randia dasycarpa (Kurz) Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 113 (1911). –
Gardenia dasycarpa Kurz, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 41: 312 (1872).
= Catunaregam tomentosa (Blume ex DC.) Tirveng., Taxon 27: 515 (1979).
li. Randia stenopetala (King & Gamble) Ridl., Fl. Malay Penins. 2: 79 (1923). –
Gardenia stenopetala King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72: 218
(1904).
= Kochummenia stenopetala (King & Gamble) K.M.Wong, Malayan Nat. J. 38: 32
(1984).
lii. Rennellia elongata (King & Gamble) Ridl., Bull. Misc. Inform. Kew 1939: 608
(1940). – Rennellia speciosa (Wall. ex Kurz) Hook.f. var. elongata King & Gamble, J.
Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 90 (1904).
liii. Rennellia morindiformis (Korth.) Ridl., Bull. Misc. Inform. Kew 1939: 609
(1940). – Tribachya morindiformis Korth., Ned. Kruidk. Arch. 2(2): 255 (1851), as
‘morindaeformis’.
liv. Tarenna adangensis (Ridl.) Ridl., Fl. Malay Penins. 2: 107 (1923). – Webera
adangensis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 54 (1912).
Note: Tarenna curtisii (King) Ridl., Fl. Malay Penins. 2: 107 (1923) is an isonym of T.
curtisii (King) F.N.Williams, Bull. Herb. Boissier, sér. 2, 5: 952 (1905).
lv. Tarenna glaberrima Ridl., J. Bot. 72: 257 (1934). – Tarenna glabra Ridl., J. Fed.
Malay States Mus. 10: 141 (1920), nom. illeg., non Merr. (1914).
390 Gard. Bull. Singapore 70 (2) 2018
lvi. Tarenna grandifolia (Hook.f.) Ridl., Fl. Malay Penins. 2: 104 (1923). – Webera
grandifolia Hook.f., Fl. Brit. India 3: 105 (1880).
= Tarenna odorata (Roxb.) B.L.Rob., Proc. Amer. Acad. Arts 45: 405 (1910); Wong,
Tree Fl. Malaya 4: 410 (1989).
lvii. Tarenna insularis (Ridl.) Ridl., Fl. Malay Penins. 2: 107 (1923). – Webera
insularis Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 55 (1912).
lviii. Tarenna longifolia (G.Don) Ridl., Fl. Malay Penins. 2: 104 (1923). – Ixora
longifolia G.Don, Gen. Hist. 3: 573 (1834). – Pavetta longifolia (G.Don) Miq., Fl.
Ned. Ind. 2: 275 (1857).
lix. Tarenna napieri (Ridl.) Ridl., Fl. Malay Penins. 2: 102 (1923). – Webera napieri
Ridl., J. Fed. Malay States Mus. 4: 34 (1909).
= Pavetta napieri (Ridl.) Bremek., Repert. Spec. Nov. Regni Veg. 37: 96 (1934).
lx. Tarenna papillulosa (King) Ridl., Fl. Malay Penins. 2: 10 (1923). – Stylocoryna
adpressa var. papillulosa King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72: 200 (1904).
= Tarenna adpressa (King) Merr., Philipp. J. Sci. 17: 472 (1921).
lxi. Tarenna pulchra (Ridl.) Ridl., Fl. Malay Penins. 2: 104 (1923). – Webera pulchra
Ridl., J. Fed. Malay States Mus. 4: 33 (1909).
lxii. Tarenna salicina Ridl., Fl. Malay Penins. 2: 104 (1923). – Webera salicina Ridl.,
J. Fed. Malay States Mus. 4: 33 (1909).
= Pavetta salicina (Ridl.) Bremek., Repert. Spec. Nov. Regni Veg. 37: 103 (1934).
lxiii. Tarenna stellulata (Hook.f.) Ridl., Fl. Malay Penins. 2: 106 (1923). – Webera
stellulata Hook.f., Fl. Brit. India 3: 104 (1880).
lxiv. Tarenna sumatrana (Boerl.) Ridl., Bull. Misc. Inform. Kew 1926: 69 (1926). –
Webera sumatrana Boerl. in Veth, Midden-Sumatra 4(2): 21 (1884).
lxv. Tarenna wallichii (Hook.f.) Ridl., Fl. Malay Penins. 2: 106 (1923). – Webera
wallichii Hook.f., Fl. Brit. India 3: 105 (1880).
391
The Gentinales taxa of H.N. Ridley
lxvi. Tarenna wrayi (King) Ridl., Fl. Malay Penins. 2: 105 (1923). – Webera wrayi
King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 69 (1904).
lxvii. Tarenna yappii (King) Ridl., Fl. Malay Penins. 2: 105 (1923). – Webera yappii
King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 70 (1904).
lxviii. Timonius peduncularis (Wall. ex G.Don) Ridl., Fl. Malay Penins. 2: 113 (1923).
– Guettarda peduncularis Wall. ex G.Don, Gen. Hist. 3: 551 (1834).
= Timonius avescens (Jacq.) Baker, Fl. Mauritius 144 (1877).
lxix. Uncaria parviora (Ridl.) Ridl., J. Straits Branch Roy. Asiat. Soc. 79: 75 (1918).
– Uncaria lanosa Wall. var. parviora Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 109
(1911).
= Uncaria homomalla Miq., Fl. Ned. Ind. 2: 343 (1857).
lxx. Urophyllum enneandrum (Wight) Ridl., J. Bot. 70: 193 (1932). – Axanthes
enneandra Wight, Calcutta J. Nat. Hist. 7: 144 (1847).
lxxi. Urophyllum leucophlaeum Ridl., J. Bot. 70: 196 (1932). – Urophyllum coriaceum
Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 18 (1912), nom. illeg., non Miq. (1861).
lxxii. Vangueria grisea (King & Gamble) Ridl., J. Straits Branch Roy. Asiat. Soc. 59:
116 (1911). – Canthium griseum King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat.
Hist. 73: 62 (1904).
= Meyna grisea (King & Gamble) Robyns, Bull. Jard. Bot. État Bruxelles 11: 240
(1928).
lxxiii. Zeuxanthe beccariana (Baill.) Ridl., Bull. Misc. Inform. Kew 1939: 610 (1940).
– Morinda beccariana Baill., Bull. Mens. Soc. Linn. Paris 1(26): 205 (1879).
= Prismatomeris beccariana (Baill.) J.T.Johanss., Opera Bot. 94: 57 (1987).
lxxiv. Zeuxanthe prismatomeriformis (Merr.) Ridl., Bull. Misc. Inform. Kew 1939:
611 (1940). – Rennellia prismatomeriformis Merr., Univ. Calif. Publ. Bot. 15: 296
(1929).
= Prismatomeris beccariana (Baill. ex K.Schum.) J.T.Johanss., Opera Bot. 94: 57
(1987).
392 Gard. Bull. Singapore 70 (2) 2018
ACKNOWLEDGEMENTS. We thank the staff and volunteers at the Herbarium of the
Singapore Botanic Garden for their help with specimens. We thank Ng Xin Yi of the National
Parks Board, Singapore, for providing the lectotypication and correct identity of Tarenna
lancifolia Ridl. We also thank Seah Wei Wei for clarifying the status of Morinda elliptica
(Hook.f.) Ridl. Two anonymous reviewers provided valuable comments on the manuscript.
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