Cephalopods from the Middle Carboniferous of the Donets Basin (Luhansk region, Eastern Ukraine)

Abstract

Six forms of cephalopods are described from Middle Carboniferous deposits (Man-drykinka and Mospinska Formations) of the Donets Basin. The collection includes three nautiloids: a new species Millkoninckioceras udovichenkoi sp. nov., Peripetoceras cf. globatoides Shimansky, and Ephippioceras wildi Hind. The latter was previously known only from Westphalian deposits of Britain. In the body chamber of Ephippioceras wildi a fossil was found that can be a part of jaw apparatus of this species. Peripetoceras globatoides, described by V. M. Shimansky from Serpu-khovian deposits of the central part of the East European Platform, was unknown for the Donets Basin before. A new species, Millkoninckioceras udovichenkoi, differs from other species of the genus by ornamentation consisting of the longitudinal ridges on the ventral-lateral bend, and some other morphological details, among which the most significant are the different shape of sutures, high and narrow whorls, and minor shell size. In addition, three ammonoids were described as Gas-trioceras cf. listeri (Sowerby), Gastrioceras lupinum A. Popov, and Branneroceras branneri (Smith). The first species was mentioned earlier from the Donets Basin. The second one is probably endemic and it was indicated before from younger deposits. The find of Branneroceras branneri is the first reliable indication of its presence in the Carboniferous of the Donets Basin.

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Вісник Національного науково-природничого музею
Proceedings of the National Museum of Natural History
НАЦІОНАЛЬНА АКАДЕМІЯ НАУК УКРАЇНИ · НАЦІОНАЛЬНИЙ НАУКОВО-ПРИРОДНИЧИЙ МУЗЕЙ
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ТОМ 16 2018
VOL.
Зміст Contents
палеонтологія palaeontology
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77 Surіadna, N., G. Mikitinets. Distribution
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Маркова, А. О. Особливості агресивних
взаємодій трьох видів родини Paridae з
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3p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
UDC 564.53:551.735.2(477.61)
Cephalopods from the Middle Carboniferous of the
Donets Basin (Luhansk region, EasternUkraine)
V.S.Dernov
Cephalopods from the middle Carboniferous of the Donets Basin (Luhansk region, Eastern Ukraine). —
V.S.Dernov. — Six forms of cephalopods are described from middle Carboniferous deposits (Mandrykin-
ka and Mospinska Formations) of the Donets Basin. e collection includes three nautiloids: a new species
Millkoninckioceras udovichenkoi sp. nov., Peripetoceras cf. globatoides Shimansky, and Ephippioceras wildi
Hind. e latter was previously known only from Westphalian deposits of Britain. In the body chamber of
Ephippioceras wildi a fossil was found that can be a part of the jaw apparatus of this species. Peripetoceras glo-
batoides, described by V. M. Shimansky from Serpukhovian deposits of the central part of the East European
Platform, was unknown for the Donets Basin before. A new species, Millkoninckioceras udovichenkoi, diers
from other species of the genus by orna mentation consisting of longitudinal ridges on the ventral-lateral
bend, and some other morphological details, among which the most signicant are the dierent shape of
sutures, high and narrow whorls, and minor shell size. In addition, three ammonoids were described as
Gastrioceras cf. listeri (Sowerby), Gastrioceras lupinum A.Popov, and Branneroceras branneri (Smith). e
rst species was mentioned earlier from the Donets Basin. e second one is probably endemic and it was
indicated before from younger deposits. e nd of Branneroceras bran neri is the rst reli able indication of
its presence in the Carboniferous of the Donets Basin.
Key w o r ds : cephalopods, Middle Carboniferous, Donets Basin
Introduction
Cephalopod remains are widely distributed in the Carboniferous deposits of the Donets Basin
(Dernov, 2016; Librovich, 1946, 1947; Popov, 1979; Shimansky, 1967). ey were studied by many
researchers, including A.V. Gurov, N. I. Lebedev, L. S. Librovich, A.V. Popov, D. E. Aisenverg,
T. V. Astakhova, V.I. Poletaev, and L. F. Kuzina. Nevertheless, remains of cephalopods from these
deposits require further thorough study. e relevance of such studies is justied by the large value of
cephalopods for correlating of remote sections, detailed breaking up of deposits, paleobiogeographi-
cal zoning of ancient water areas, and reconstruction of habitat conditions and burial of fossil fauna
and ora (Bogoslovskaya et al., 1999).
e purpose of this work is to describe the remains of cephalopods important for the paleobiogeo-
graphic research and stratigraphy of the middle Carboniferous deposits of the Donets Basin.
Material and methods
e remains of middle Carboniferous cephalopods of the central part of the Donets Basin (Fig. 1)
have been collected during 2004–2016. Because of these works, and due to the help of M.I. Udovi-
chenko (Luhansk), who kindly provided a number of valuable samples and introduced some inter-
esting sites, a representative collection of cephalopod remains from middle Carboniferous deposits
was created. In this paper, we describe three nautiloids, including a new species, Millkoninckioceras
udovichenkoi Dernov, sp. nov., and three ammonoids from the Mandrykinka and Mospinska Forma-
tions (Blagodathian and Zujevian Horizons) of the Donets Basin (Table 1).
e nautiloids of the genus Millkoninckioceras Kummel, 1963 have evolute, discoidal conchs with
a subcentral siphuncle, straight suture, and whorls slowly increasing in height and width (Shimansky,
1967). e genus is known from Carboniferous and Permian deposits of the Urals (Shimansky, 1967),
the Moscow Syneclise (Makhlina et al., 2001), Western Europe (Kummel, 1967), and North America
(Miller, Kemp, 1947; Newell, 1936).
Corresponding author address: Independent researcher (Makedonivka, Ukraine); e-mail: fvitalydernov@gmail.com
2018 • vol. 16, pp 3-14
DOI: 00.0000/000-000
GEO&BIO

4GEO&BIO • 2018 • vol. 16 p-ISSN 0000-0000 e-ISSN 0000-0000
e studied material comes from the outcrops described in detail below.
A. A quarry from 1.5 km west from Volnukhine village (Lutugine district, Luhansk region). Yellow-
ish-grey siltstones above the F1 limestone (Mandrykinka Formation), which is exposed at the up-
permost ledge of the quarry. A thin layer of yellow carbonate nodules with remains of pelecypods,
gastropods, cephalopods (Millkoninckioceras udovichenkoi Dernov, sp.nov.), crinoids, and echi-
noids (Fig. 2, 1) is observed 0.3 m above the base of siltstones.
B. Exploratory ditches on the le slope of the Shchetov ravine, 1 km north from Zelenodilske vil-
lage (Antratsit district, Luhansk region). Grey mudstones and siltstones, lying 1m above the G1
limestone. In this layer, the nautiloid Gzheloceras (?) sp. and ammonoids Gastrioceras cf. listeri
(Sowerby), Gastrioceras sp., and Pseudo-
bisatoceras sp. were found (Dernov, 2016).
In addition, some remains of pelecypods,
gastropods, plant detritus, and trace fos-
sils were observed.
C. e le slope of the Rebrova ravine, 1.5
km north-west from Makedonivka village
(Lutugine district, Luhansk region). ere
is greyish-brown, ne-grained sandstone
40 m below the G2 limestone. From this
layer some remains of bryozoans, bra-
chiopods, pelecypods, gastropods, cepha-
lopods (Ephippioceras sp., Branneroceras
branneri (Smith)), arthropods, and shes
were collected.
D. e ravines near Makedonivka village (Fig.
2, 3). e exposed dark grey mudstones
contain siderite nodules and remains of a
diverse marine fauna: corals, pelecypods,
gastropods, cephalopods (Bran neroceras
branneri (Smith), Gastrio ceras cf. listeri
(Sowerby)), crinoids, arthropods, and
shes. ese mudstones lie 25 m below the
G2 limestone. Light-grey siltstones with
siderite nodules and remains of pelecy-
pods, gastropods, cephalopods (Peripeto-
ceras cf. globatoides Shimansky and Bran-
neroceras branneri (Smith)) were found
here. ese siltstones lie one metre below
the G2 limestone.
E. A small quarry and excavation of the high-
way T1320, 0.3 km north from Make-
donivka village (Fig. 2, 5). e nautiloid
Ephippioceras wildi Hind was found at
the top of the G2 limestone. e dark
grey, almost black, thick-plated limestone
contains remains of corals, bryozoans,
brachiopods, pelecypods, gastropods,
crinoids, shes, and oen stromatolites
(Dernov, 2017).
Fig. 1. The research area and the location of studied
outcrops.
Рис. 1. Район досліджень та розташування вивчених
відслонень.

5p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
F. e le slope of the Suha ravine in its
middle reaches (3 km east from Make-
donivka village). Grey and yellowish-
grey siltstone, which lies on the place of
G4
1 limestone. In this layer, the remains
of pelecypods, gastropods, cephalopods
(Gastrioceras lupinum A.Popov) and oth-
er fossils (Fig. 2, 2, 4) were found.
e technique and terminology proposed
by V.E. Ruzhenzev (Ruzhencev, Bogoslovs-
kaya, 1971) were followed during the study
and description of ammonoids, while the
examination of nautiloids follows Shiman-
sky (1967). e studied collection (No. S-4)
is stored in the Geological Museum of Taras
Shevchenko Luhansk National University.
Abbreviations used in this work: Dc —
conch’s diameter, Wh — whorl’s height,
Ww — whorl’s width, Du — umbilical di-
ameter, Wh/Dc, Du/Dc, Ww/Wh — ratios
of parameters.
Systematic palaeontology
Class Cephalopoda
Subclass Nautiloidea
Order Nautilida
Suborder Rutoceratina
Superfamily Koninckiocerataceae
Family Koninckioceratidae Hyatt, 1900
Genus Millkoninckioceras Kummel, 1963
Millkoninckioceras udovichenkoi Dernov, sp. nov. (Fig. 3, 1)
Hol ot yp e. Geological Museum of Taras Shevchenko Luhansk National University, No. 394-2015;
Donets Basin, a quarry near Volnukhine village, Mandrykinka Formation, above the F1 limestone
(Bilinguites-Cancelloceras Genus Zone); Fig. 3, 1.
Et ym o lo g y. e species is named in honour of the paleontologist Mykola Ivanovych Udovichenko.
For m. e conch is evolute discoid with a small contact furrow. e cross section of the last whorl
is longitudinally oval (Fig. 5, 2). e phragmocone’s venter is slightly convex, while the body cham-
ber’s venter of is attened. e ventral shoulders are not pronounced. Flanks are broad, very slightly
convex, insignicantly diverge to the middle of the height of the whorl, and slightly converge near by
the umbilical shoulders. e greatest width of the whorl is observed approximately on the middle of
its height. Whorls are moderately increased in height and width. Umbilical shoulder is wide, but quite
clear. e umbilical wall is at, fairly wide, inclined at an angle of approximately 45° to the symmetry
plane of the conch. e dorsal side is narrow, slightly concave. Umbilicus is attened, relatively wide,
about one third of the conch’s diameter. e body chamber occupies not less than half of the whorl.
Gas chambers are short; there are about 17 these chambers in the whorl. Position of the siphuncle is
unknown.
Fig. 2. Some studied outcrops: 1 — quarry in Volnukhine
village (outcrop А); 2 — siltstones on the place of limestone
G4
1 (outcrop F); 3 — mudstones 25 metres below of the lime-
stone G2 (outcrop D); 4 — mudstones under the siltstones
on place of the lime stone G4
1 (outcrop F); 5 —limestone
G2 (outcrop E). The height of the limestone wall — 2.5 m.
Рис. 2. Деякі вивчені відслонення:
1 — кар’єр в с. Волнухине (відслонення «А»). 2 — але-
вроліти на місці вапняку G4
1 (відслонення «F»); 3 —
глинисті сланці в 25 метрах нижче вапняку G2 (від-
слонення «D»); 4 — глинисті сланці нижче алевролітів
на місці вапняку G4
1 (відслонення «F»); 5 — вапняк G2
(відслонення «Е»). Висота стінки вапняку— 2,5 м.

6GEO&BIO • 2018 • vol. 16 p-ISSN 0000-0000 e-ISSN 0000-0000
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc Ww/Wh
394-2015 37 13 13 13 0.35 0.35 0.35 1.0
Or n am e nt a t io n . Two well-marked longitudinal narrow ribs are observed on the ventral-lateral
bend. ey are separated by a groove, which is approximately three times wider than the ribs. ese
ornamentation elements can be easily visible on both the phragmocone and the body chamber. In
addition, there are transverse thin ribs on the sink. In total, ve ribs per one millimetre of the whorl’s
length are recognizable. On the venter, the ribs form a deep sine; they cross the lateral sides directly,
and slightly deviate back on the umbilical wall.
Sutu r e. On the venter, there is a poorly visible saddle, on the lateral sides there is a weak visible
shallow lobe. On the ventral side, the saddle is small; it seems that the suture is straight (Fig. 5, 1).
Co m p ar i so n . e characteristic of ornamentation of the longitudinal ribs on the ventral-lateral
bend and some other morphological details allow us distinguishing clearly the new species from
other representatives of the genus. e new taxon diers from M. konincki (Miller and Kemp) whorl’s
height, the presence of ventral-lateral ribs and bigger size. e species group described by Newell
(1936) from the Pennsylvanian of the North America and tentatively attributed by Shimansky (1967)
to the genus Millkoninckioceras (M. jewetti, M. wyandottense, M. eliasi) signicantly diers morpho-
logically from the new species in the greater dissection of the suture line, low and wide whorls, or-
namentation, and the considerable size of conchs. From the Permian M. bibbi (Miller et Kemp) from
Texas (Miller, Kemp, 1947), the new species diers in the shape of the whorl’s cross section (in the
American species the whorls are rather low and wide, transversely elliptical). In addition, the size of
Fig. 3. Studied nautiloids: 1 — Millkonincki-
oceras udovichenkoi Dernov, sp. nov.: 1а —
lateral view, 1b — aperture view, 1c — an
enlarged portion of the sculpture on the lat-
eral side; sample No. 394-2015, outcrop А;
2 — Peripetoceras cf. globatoides Shi mansky,
1967: 2а — lateral view, 2b— ventral view,
2c— aperture view; sample No. 367, outcrop
D; 3 — Ephippioceras wildi Hind, 1910: 3а —
ventral view, 3b — lateral view, sample No.
1421-2015, outcrop E. Scale bar equals 10
mm in 1a, 1b, 2, 3; 2 mm in 1c.
Рис. 3. Вивчені наутиліди: 1 — Millkonin-
cki oceras udovichenkoi Dernov, sp. nov.:
1а — збоку, 1b — зі сторони апертури,
c — збільшена ділянка скульптури на бо-
ковій стороні; зразок № 394-2015, відсло-
нення «А»; 2 — Peripetoceras cf. globatoides
Shi mansky,1967: 2а — збоку, 2b — вен-
трально, 2c — вигляд зі сторони аперту-
ри; зразок № 367, відслонення «D»; 3 —
Ephippioceras wildi Hind, 1910: 3а — вен-
трально, 3b— збоку; зразок №1421-2015,
відслонення «Е». Масштабний відрізок —
10 мм для 1а, 1b, 2, 3; 2 мм для 1c.

7p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
M. bibbi signicantly exceeds that of the M. udovichenkoi (for example, the conch’s diameter in the
rst species is almost 7 times greater than in the second one).
Lo c at i on . Lutugine district, Luhansk region, a quarry 1.5 km to the west from Volnukhine vil-
lage: concretions 0.3 m above the F1 limestone, Mandrykinka Formation (sample No. 394-2015). Ma-
terial was collected by M.I. Udovichenko in 2008.
Stratigraphic and geographical distribution. Upper part of the lower Bashkirian, Do-
nets Basin.
Suborder Liroceratina
Superfamily Lirocerataceae
Family Liroceratidae Miller et Youngquist, 1949
Genus Peripetoceras Hyatt, 1894
Peripetoceras cf. globatoides Shimansky, 1967 (Fig. 3, 2)
Mate r ia l . Nine fragments of conchs.
Form. e conch is semi-involute, subspherical, with rapidly increasing whorls. e cross-section
of the whorl is sub-trapezoidal. Venter is broad, at or very slightly convex. e ventral shoulder is
roundish. e anks are convex and slowly diverge from the ventral shoulder to the umbilicus. e
umbilical shoulder is almost rectangular, while the umbilical wall is at. e greatest width of the
whorl is near the umbilicus. Dorsal side is concave, about 2.5 times narrower than the whorl’s width.
e aperture shape is not established. Because of inconsistent preservation, it is not possible to count
the number of cameras in the whorl. e siphuncle could not be studied.
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc Ww/Wh
367 ≈ 32 14 21 ≈ 0.44 ≈ 0.66 1.5
414 10 17 8 1.7
415 10 15 1.5
417 11 19 9 1.7
6492 8,5 18 2.1
Or n ame n tat ion on the fossils is not observed.
e sutu r e could not be studied.
Re m ark s . e studied material is very similar to Peripetoceras globatoides Shimansky, which is
known from the Serpukhovian of the central part of the East European platform (Shimansky, 1967).
Nevertheless, we cannot assign the described fossils to the indicated taxon. due to the smaller size of
samples in comparison with the holotype, as well as the lack of possibility to compare the shape of the
sutures and the position of the siphuncle of our material and type.
Lo c at i on . Lutugine district, Luhansk region, ravine in northern vicinities of Makedonivka vil-
lage: siltstones under the G2 limestone (samples Nos. 367, 414, 415, 416, 418, 419, 420, 6492), Mospin-
ska Formation. Material was collected by the author in 2009–2016.
Stratigraphic and geographical distribution. Bashkirian Stage of the Donets Basin.
Family Ephippioceratidae Miller et Youngquist, 1949
Genus Ephippioceras Hyatt, 1884
Ephippioceras wildi Hind, 1910 (Fig. 3, 3)
1910 Ephippioceras wildi sp. nov.: Hind, p. 104, pl. IV, Fig. 2, pl. V, Fig. 1,2, pl. VI, Fig. 1.
Holo t yp e. Museum of the University of Manchester, No. W.763; UK, Lancashire, Burnley, Hap-
ton, Spa Clough, Lower Coal Measures, roof of Bullion Coal.
Mate r ia l . One fragment of the large conch.

8GEO&BIO • 2018 • vol. 16 p-ISSN 0000-0000 e-ISSN 0000-0000
For m . e conch is involute, subspherical, with rapidly increasing whorls. e cross section of
the adult whorl is low, kidney-shaped. e ventral shoulder is not pronounced. e venter forms
one hemispherical surface together with the lateral. e anks of the phragmocone are at, while
the anks of the body chamber are slightly convex. ey slowly diverge as they approach the umbili-
cal shoulder. e greatest width of turnover is observed near the umbilicus. e umbilical shoulder
rounded, but clear. e umbilical wall is narrow, at and almost perpendicular relative to the symme-
try plane of the shell. Umbilicus is wide and equals about one third of the conch’s diameter. e dorsal
side of the conch has not been studied. e length of the body chamber is not known, since only its
posterior part was preserved. Gas chambers, the available material, are short (about 9–10chambers
for the width of the conch’s whorl). e siphuncle is located between the centre and the venter.
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc Ww/Wh
1421-2015 ≈ 90 54 95 29 0.6 ≈ 1.05 0.32 1.76
ere is no or na m ent a ti o n on the mold.
Sutu r e. e ventral saddle is high, tapers slowly as it comes near the apex. Its height is 2.5 times
less than the width of its base. e ventral saddles of adjacent lobes are quite distant from each other.
On the anks, there is a shallow wide lobe (Fig. 5, 3).
Co m p ar i so n . e described species diers from the closely related species of Ephippioceras fer-
ratum (Cox, 1857) from the Lower and Middle Carboniferous and E.clitellarium (Sowerby, 1840)
from the Lower — Upper Car boniferous, in the shape of the suture (in Ephippioceras wildi the ventral
saddle has a more rounded top and a lesser of height). In addition, the Ephippioceras wildi is diers
signicantly from other species of the genus in the shape of cross section of the whorl and wider
umbilicus.
Re m ark s . Two fragments of the nautiloid mold (sample Nos. 405-2015 and 413-2015) were ob-
tained from the sandstone in 40 m below the G2 limestone. ese specimens resemble Ephippioceras
wildi. We dene the nautiloid from sandstone as Ephippioceras sp.
In the body chamber of the described conch, fragments of brachiopod shells and fragments of
stems of crinoids, introduced there by the current during its postmorality stay on the surface of the
seabed, were found. Moreover, in the last gas chamber there is a small brachiopod shell, which got
here because of the violation of the integrity of the last septa. A deformed fossil was also found in
the body chamber in the form of a thin plate, presumably oval, on the surface of which frequent thin
concentric ribs are observed (Fig. 4). e length of this fragment is 17 mm. e mentioned fossil may
be a part of the jaw apparatus of the described nautiloid.
Lo c at i on . Lutugine district, Luhansk region, a small quarry 0.3 km north from Makedonivka
village, G2 limestone, Mospinska Formation (sample Nos. 1421-2015). Material was collected by the
author in 2014.
Str a ti g rap h ic an d g e og r aph ic a l d is t r i but i on . In addition to the Donets Basin, this spe-
cies is known from the Westphalian A of Britain.
Subclass Ammonoidea
Order Goniatitida Hyatt, 1884
Suborder Goniatitina Hyatt, 1884
Superfamily Gastriocerataceae Hyatt, 1884
Family Gastrioceratidae Hyatt, 1884
Genus Gastrioceras Hyatt, 1884
Gastrioceras cf. listeri (Sowerby, 1812) (Fig. 5, 3)
Mate r ia l . Two fragments of dierent conchs and one incomplete mold.

9p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
Form. It is dicult to estimate about
the shell’s shape. e venter is a single con-
vex surface with lateral sides; the ventral
shoulder is not expressed. e umbilical
shoulder is somewhat angular, while the
wall is wide and slightly convex. Cross sec-
tion of the whorl is low. e greatest width
of the whorl is observed near the umbilical
shoulder. e umbilicus is deep, wide.
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc
655-2015 13.5 25
827-2015 7.5 2.5 3.5 0.33 0.47
Or n am e nt a t io n . e conch’s surface is covered with gentle striae forming a shallow sine on the
venter, and a weak protrusion on the ventral-lateral part of the whorl. e nature of the striae on other
parts of the shell is unknown. Sharp tubercles are located on the umbilical shoulder. On 5 mm of the
umbili cal shoulder account for about two tubercles (whorl height is 13.5 mm). On the conch, there
are clearly no ticeable constrictions, but it is not possible to count them.
e shape of the su tur e has not been studied because of poor preservation of the samples.
Re m ar k s. Insucient preservation of the material did not allow investigating the suture and, ac-
cordingly, to attribute the studied material with those of G. listeri.
Lo c at i on . Lutugine district, Luhansk region, 2.5 km north from Zelenodilske village: siltstones
over limestone G1 (No. 827-2015), Dyakivska Series; a ravine 3 km east of Makedonivka village: argil-
lite in 25 m below the G2 limestone (Nos. 552-2015, 655-2015), Mospinska Formation. Material was
collected by the author in 2006–2014.
Stratigraphic and geographical distribution. Gastrioceras listeri (Sowerby, 1812) is
known from the goniatite zone G2 of Britain, Portugal, the Netherlands, Belgium, Germany, and Po-
land. is species was also noted earlier from the Donets Basin (Popov, 1979) in limestones G1–G2
and G4 of the Mospinska Formation and in the roof of coal h4
2 layer of the Smolyanynovka Formation.
Gastrioceras lupinum A. Popov, 1979 (Fig. 5, 1)
1979 Gastrioceras lupinum sp. nov.: Popov, p. 83–84, plate Х, Figs 9–11.
Holo t yp e. Federal State Budgetary Institution “A.P. Karpinsky Russian Geological Research In-
stitute,” No. 3223; Donets Basin, Sorochya girder, Bashkirian Stage, H5 – І1 limestones.
Mate r ia l . Six fragments of moldes and imprints of conchs.
Fig. 4. Fragment of nautiloid jaw apparatus (?)
from the body chamber of Ephippioceras wildi
Hind, 1910; sample No. 1424-2015, outcrop E,
scale barequals 10 mm.
Broken or deformed sides of the fossil are
marked by dotted line; extant edges are marked
by thin solid lines and concentric sculpture is
shown by thick lines.
Рис. 4. Фрагмент щелепного апарату на-
утиліди (?) із житлової камери Ephippioceras
wildi Hind, 1910; зразок № 1424-2015, від-
слонення Е, масштабний відрізок — 10 мм.
Пунктиром позначені зламані чи деформо-
вані сторони фосилії; тонкою суцільною
лінією — краї фосилії, що збереглися; тов-
стою лінією — концентрична скульптура.

10 GEO&BIO • 2018 • vol. 16 p-ISSN 0000-0000 e-ISSN 0000-0000
For m . e shape of the conch cannot be established; Popov (1979) pointed out that it is pachy-
conical, very involute. Flanks are slightly convex; slightly divergent at coming to the umbilicus. e
umbilicus has medium size. e umbilical shoulder is sharp and angular.
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc
492-2015 ≈12 6 4 0.5 0.36
1356-2015 11 4 3.5 0.36 0.32
e orn ame nt a t io n is represented by thin ribs forming a small protrusion on the ventro-lateral
bend, and wide shallow sinus on the anks. Between the ribs, there are lines repeating the congura-
tion of the ribs. On the umbilical shoulder, the ribs turn into sharp small tubercles. At 1 mm of the
umbilical shoulder (at the whorl’s height of 6 mm), there are two or three tubercles. In addition to the
transverse ribs, weakly noticeable lyres are observed on the ventral-lateral bend. ere are constric-
tions on the studied fragments of conchs, however it is impossible to calculate exactly their number
(Popov (1979) pointed out that there are three of them on the whorl).
Sutu r e is not observed due to insucient preservation of the material.
Co m p ar i s o n . Due to the characteristic ornamentation, this species diers quite sharply from
close taxa.
Fig. 5. The shape of the suture and the cross
section of the whorl of some studied nautiloids
and ammonoids: 1 — suture line of Ephippioce-
ras wildi Hind, 1910 (Wh= 54 mm, Ww= 95
mm, sample No.1424-2015, outcrop E); 2 — su-
ture of Millkoninckio ceras udovichenkoi Dernov,
sp. nov. (Wh = 13 mm, Ww = 13 mm, sample
No. 394-2015, outcrop А). 3 —cross section of
body chamber of Millkoninckioceras udovichen-
koi Dernov, sp. nov. (sample and outcrop are the
same). 4,5 — conchs of Branneroceras branneri
(Smith), 1896 (sample No. 4036 (4), outcrop D,
scale bar equals 3 mm; sample No. 502 (5), out-
crop D, scale bar equals 5mm).
Рис. 5. Форма лопатевої лінії та попереч-
ний переріз завитку деяких вивчених на-
утилоідей та аммоноідей: 1 — лопатева лі-
нія Ephippioceras wildi Hind, 1910 (В=54 мм,
Ш=95 мм, зразок № 1424-2015, відслонення E).
2 — лопатева лінія наутиліди Millkoninckio-
ceras udovichenkoi Dernov, sp. nov. (В = 13
mm, Ш = 13 mm, зразок № 394-2015, від-
слонення А); 3 —поперечний перетин жит-
лової камери наутилоідеї Millkoninckioceras
udovichenkoi Dernov, sp. nov. (зразок та від-
слонення ті самі). 4,5 — черепашки аммоно-
ідей Branneroceras branneri (Smith), 1896 (зра-
зок № 4036 (4), відслонення D, масштабний
відрізок 3мм; зразок № 502 (5), відслонення
D, масштабний відрізок 5 мм).

11p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
Lo c at i on . Suha ravine, Lutugine district, Luhansk region, 3 km east from Makedonivka village:
siltstone on the place of the G4
1 limestone and adjacent slates (sample Nos. 515-2015, 517-2015, 611-
2015, 644-2015, 1320-2015, 1356-2015), Mospinska Formation. Material was collected by the author
in 2006–2014.
Stratigraphic and geographical distribution. Bashkirian Stage, Donets Basin. Occurs in
shales near the G4
1 limestone. Popov (1979) established this species in the upper part of the Smoly-
anynovka Formation (limestones H5 – І1 (?) and H6). In the shale above the G3 limestone in Suha ra-
vine, there are remains of ammonoids, very similar to G. lupinum. Fragmentation and a small number
of materials, as well as minor dierences in sculpture, did not allow us to assign these ammonoids to
the above-mentioned species.
Superfamily Schistocerataceae Schmidt, 1929
Family Schistoceratidae Schmidt, 1929
Genus Branneroceras Plummer et Scott, 1937
Branneroceras branneri (Smith), 1896 (plate 5, Figs 4, 5; plate 6, Figs 2, 4, 5)
1896 Gastrioceras branneri sp. nov.: Smith, p. 257, 258, pl. 23, Figs 1–6.
1903 Gastrioceras branneri Smith: Smith, р. 83, pl.11, Figs8–13.
1923 Gastrioceras branneri Smith: Schindewolf, text Fig. 12c.
1937 Branneroceras branneri (Smith): Plummer, Scott, p. 219 – 221, pl. 11, Figs 1–7.
1938 Branneroceras branneri var. branneri (Smith): Miller, Moore; pp. 348–350, pl. 44, Figs 5–12.
1938 Branneroceras branneri var. halenseMiller et Moore: Miller, Moore; pp. 350–351, pl.44, Figs 13–14.
1940 Branneroceras branneri var. branneri (Smith): Bisat, p. 332, Figs 3A, 3B.
1944 Gastrioceras branneri Smith: Miller, Owen, pp. 422–423, pl. 63, Figs 1, 2, pl. 65, Figs1, 2.
1948 Gastrioceras branneri branneri Smith: Miller, Downs, pl. 103, Figs 10, 11.
1950 Branneroceras branneri (Smith): Руженцев, Fig. 48d.
1958 Branneroceras branneri (Smith): Miller, Furnish, p. 262, pl. 34, Figs 5, 6.
1960 Branneroceras branneri (Smith): Руженцев, Fig. 92b
1964 Gastrioceras (Branneroceras) branneri (Smith): Gordon, p. 253–255, pl. 27, Figs 16–23, 27 – 30, text Fig. 76.
1968 Branneroceras branneri Smith: McCaleb, p. 60 – 65, pl. 8, Fig. 1–16, pl. 9, Fig. 1–19.
1971 Branneroceras branneri (Smith) Plummer et Scott: Wagner-Gentis, p. 348–349, pl. 5, Fig. 16, pl. 6, Fig. 17.
1975 Branneroceras branneri (Smith): Nassichuk, p. 142 – 144, pl. 16, Fig. 1, 3, 4, 7, 9, 10, 15, 16.
1977 Branneroceras branneri (Smith): Saunders, Manger, Gordon, pl. 5, Figs 14–17.
1980 Branneroceras branneri (Smith): Manger, Saunders, text-Fig. 10, D–G.
1999 Branneroceras branneri (Smith): Fujikawa, Ishibashi, Nakornsri, pl. 1, Figs 2–8.
Holo t yp e. United States National Museum, No. 26439; USA, Pilot Mountain, Valley Springs,
sandstone of Morrow age (location No. 1275A10).
Ma t er i al . 15 samples, including molds, conchs and their fragments of dierent state of preservation.
For m . e conch is subdiskoconic, evolute, with a moderately wide and broad umbilicus. e
cross-section of the whorl is low. Venter is slightly convex and wide. e ventral shoulder is round-
ish. e anks are convex, moderately divergent at coming to the umbilicus. e greatest value of the
whorl’s width is observed near the umbilical shoulder. e umbilical shoulder is angular, while the
umbilical wall is slightly convex. Umbilicus is moderately broad and wide, deep, stepped. Early whorls
had rounded coiling.
Di m ens i ons (in mm) and ratios:
No. sample Dc Wh Ww Du Wh/Dc Ww/Dc Du/Dc
609 14 5 7 0.35 0.50
656 30 8 14 0.27 0.47
544 30 8 12 14 0.27 0.40 0.47
669 26 8.5 13 13 0.33 0.50 0.50
3776 13 3.5 6 7 0.27 0.46 0.54
710 18 5 10 0.28 0.55
725 7 13.5
315 25 8 10 12 0.32 0.40 0.48
350 17 5 9 0.29 – 0.53
338 31 10 14 16 0.32 0.45 0.52
335 40 11 15 17 0.28 0.38 0.43

12 GEO&BIO • 2018 • vol. 16 p-ISSN 0000-0000 e-ISSN 0000-0000
e orn a me n t at i on is represented by thin frequent sharp transverse ribs forming a moderate
sine on the ventral side, and a weak protrusion on the ventral shoulder. On the anks, the ribs deviate
slightly. ere are also single spiral, which when crossing with the transverse ribs, forms a characteris-
tic reticulate ornament. On the umbilical shoulder, the transverse ribs are connecting and form short,
massive ribs. At 10 mm of the umbilical shoulder (with a diameter of conch is 26 mm) comes about
6–7 ribs. ere are three constrictions per whorl.
Sutu r e. e ventral lobe is wide, splits by a wide siphuncle saddle into two narrow branches;
outer saddle rounded, the lateral lobe is wedge-shaped.
Co m p ar i s o n . e described taxon diers from Branneroceras hillsi Nassichuk, 1975 by lower
whorls, short and massive umbilical ribs, as well as higher and narrower siphuncle saddle of the su-
ture. It diers from B. nicholasi Nassichuk, 1975 in form of the whorl and smaller height, as well as
more robust umbilical ribs and narrow outer saddle.
Re mar ks . Librovich (1947) indicated Branneroceras branneri (Smith) without a description from
the upper part of the Mandrykinka Formation, along with lower Bashkirian ammonoids (e.g., Bilin-
guites superbilingue (Bisat)). e late Bashkirian genus Branneroceras was mistakenly dened as genus
Cancelloceras (early Bashkirian).
Lo c at i on . Lutugine district, Luhansk region, ravines in the northern vicinities of Makedonivka
village: siltstone directly under the G2 limestone (Nos. 338, 502, 544, 669, 725), Mospinska Formation.
In the same place, as well as girders 3 km east and 5 km west from the village: argillite 25 m below
limestone G2 (no. 315, 335, 350, 352, 501, 656, 3776, 4036, 4213), Mospinska Formation. Beams in the
northern vicinities of Makedonovka village: sandstone 40meters below the G2 limestone (No. 710),
Mospinska Formation. Material was collected by the author in 2006–2016.
Stratigraphic and geographi-
ca l di st ribut io n. e upper part of
the Bashkirian Stage (Branneroceras-
Gastrioceras Genus Zone) of Western
Europe, China, Japan, Uzbekistan, the
Canadian Arctic Archipelago, and the
USA.
Fig. 6. Studied ammonoids: 1— Gastrio-
ceras lupinum A. Popov, 1979: 1а, 1b —
lateral view, sample No. 611-2015 (1а),
No. 611-2015 (1b), outcrop F; 2, 4, 5 —
Branneroceras branneri (Smith), 1896: 2,
4а, 5 — lateral view, 4b — ventral view,
sample No. 725 (2), No. 3776 (4), No. 669
(5), outcrop D; 3— Gastrioceras cf. listeri
(Sowerby, 1812): 3а — lateral view, 3b —
aper ture view, sample No. 655–2015, out-
crop D. The scale bar equals 5 mm in 1
and 4, and 10 mm in 2, 3, 5.
Рис. 6. Вивчені аммоноідеї: 1— Gastrio-
ceras lupinum A. Popov, 1979: 1а, 1b —
збоку, зразок № 611-2015 (1а), № 644-
2015 (1b), відслонення F; 2, 4, 5— Bran-
neroceras branneri (Smith), 1896: 2, 4а,
5— збоку, 4b — вентрально, зразок №
725 (2), № 3776 (4), № 669 (5), відслонен-
ня D; 3— Gastrioceras cf. listeri (Sowerby,
1812): 3а — збоку, 3b— з боку перего-
родки, зразок № 665-2015, відслонення
D. Масштабний відрізок — 5 мм для 1
та 4, 10мм для 2, 3, 5.

13p-ISSN 0000-0000 e-ISSN 0000-0000 GEO&BIO • 2018 • том 16
Conclusions
Six species of cephalopods were described from middle Carboniferous deposits of the Donets Ba-
sin. e described nautiloids include one newly established species (Millkoninckioceras udovichenkoi)
and two taxa previously unknown for the Donets Basin (Peripetoceras cf. globatoides, Ephippioceras
wildi). ree species of ammonoids were recorded: Gastrioceras cf. listeri, Gastrioceras lupinum, Bran-
neroceras branneri. Gastrioceras lupinum shows certain details of ornamentation not mentioned in
the original description of this taxon. Ornamentation is represented by thin ribs, which form a small
protrusion on the ventral-lateral bend, and a wide shallow sinus on the anks. Between the ribs, there
are lines that repeat the conguration of the ribs. On the umbilical shoulder, the ribs turn into sharp
small tubercles.
e presence of the genera Branneroceras and Gastrioceras in the sediments of the Mospinska For-
mation evidence its belonging to the Branneroceras-Gastrioceras Genus Zone. e studied materials
conrm the close connection of the Middle Carboniferous water areas of the Donets Basin and the
Moscow Syneclise, as well as the Donets Basin and Western Europe.
Acknowledgements
e author is grateful to M. I.Udovichenko for assistance in the course of research and samples, to
I.V. Zagorodniuk and A. V. Bratishko for their assistance in the publication of this article, as well as
to Oleksandr Kovalchuk for a signicant contribution in improving its content.
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