Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico IV

Abstract

A summary of all taxonomic and nomenclatural changes to the check list of the Noctuoidea of North America north of Mexico since the last update published in 2015 is provided. A total of 64 changes are listed and discussed, consisting of 26 recently published changes and additions, and an additional 38 presented herein. One stat. n., one stat. rev., six syn. n., and two comb. n. are proposed for the first time. Orthimella Schmidt & Lafontaine nom. n. is proposed here as an objective replacement name for Himella Grote, 1874 [Noctuinae: Orthosiini], a junior homonym of Himella Dallas, 1852 [Hemiptera: Coreidae].

Full text

Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera)... 241
Additions and corrections to the check list of the
Noctuoidea (Insecta, Lepidoptera) of North America
north of Mexico IV
B. Christian Schmidt1, J. Donald Lafontaine1, James T. Troubridge2
1 Canadian National Collection of Insects, Arachnids, and Nematodes, Biodiversity Program, Agriculture and
Agri-Food Canada, K.W. Neatby Bldg., 960 Carling Ave., Ottawa, Ontario, Canada K1A 0C6 2 Hagersville,
Ontario, Canada
Corresponding authors: B.C. Schmidt (Christian.Schmidt@agr.gc.ca); J.D. Lafontaine (Don.Lafontaine@agr.gc.ca)
Academic editor: J. Adams|Received 19 July 2018|Accepted 12 September 2018|Published 8 October 2018
http://zoobank.org/8A6522C4-1220-4D31-869D-615410DC594A
Citation: Schmidt BC, Lafontaine JD, Troubridge JT (2018) Additions and corrections to the check list of the
Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico IV. In: Schmidt BC, Lafontaine JD (Eds)
Contributions to the systematics of New World macro-moths VII. ZooKeys 788: 241–252. https://doi.org/10.3897/
zookeys.788.28500
Abstract
A summary of all taxonomic and nomenclatural changes to the check list of the Noctuoidea of North
America north of Mexico since the last update published in 2015 is provided. A total of 64 changes are
listed and discussed, consisting of 26 recently published changes and additions, and an additional 38
presented herein. One stat. n., one stat. rev., six syn. n., and two comb. n. are proposed for the rst time.
Orthimella Schmidt & Lafontaine nom. n. is proposed here as an objective replacement name for Himella
Grote, 1874 [Noctuinae: Orthosiini], a junior homonym of Himella Dallas, 1852 [Hemiptera: Coreidae].
Keywords
Canada, United States, Erebidae, Noctuidae, Nolidae
Introduction
Continuing work on the taxonomy and systematics of New World Noctuoidea has
resulted in 64 additional changes to the check list of North American Noctuoidea
(Lafontaine and Schmidt 2010). In terms of the North American fauna diversity, the
ZooKeys 788: 241–252 (2018)
doi: 10.3897/zookeys.788.28500
http://zookeys.pensoft.net
Copyright B. Christian Schmidt et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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B. Christian Schmidt et al. / ZooKeys 788: 241–252 (2018)
242
current work summarizes changes with 43 species-level taxa added and nine deleted for
a net gain of 34. ese are in addition to the 115 changes made in 2011 (Lafontaine
and Schmidt 2011), and 64 made in 2013 (Lafontaine and Schmidt 2013), and the
124 made in 2015 (Lafontaine and Schmidt 2015). e new total for Noctuoidea in
North America north of Mexico is 3706 species.
Repository abbreviations
Taxonomic changes are based on examination of material, especially type specimens,
in the following collections:
AMNH e American Museum of Natural History, New York, NY, USA
ANSP e Academy of Natural Sciences, Philadelphia, Pennsylvania, USA
NHMUK e Natural History Museum, London, UK
CNC Canadian National Collection of Insects, Arachnids, and Nematodes,
Ottawa, Ontario, Canada
CUIC Cornell University Insect Collection, Ithaca, New York, USA
KBC Knudson/Bordelon Collection at MCLB – McGuire Center for Lepidoptera
and Biodiversity, University of Florida, Gainesville, FL, USA
MNHN Muséum National d’ Histoire Naturelle, Paris, France
MSU Michigan State University, East Lansing, Michigan, USA
TLSRC Texas Lepidoptera Survey Research Collection, Houston, Texas, USA
JTTC James T. Troubridge Collection, Hagersville, Ontario, Canada
USNM National Museum of Natural History [formerly, United States National
Museum], Washington, District of Columbia, USA
Results
Corrections, additions, and changes (highlighted in bold)
930207.1 Hypoprepia lampyroides Palting & Ferguson, 2018
930215.1 Clemensia umbrata Packard, 1872
930216 Clemensia ochracea Schmidt & Sullivan, 2018
(= Clemensia patella of authors)
930233 – 930238 Chelis Rambur, 1866
(= Holoarctia, Neoarctia, Hyperborea)
930239 – 930277 Apantesis
(= Holarctia, Grammia, Notarctia)
930283 – 930289 Arctia
(= Parasemia, Acerbia, Pararctia, Platarctia, Platyprepia)
930299.1 Virbia marginata (Druce, 1885)

Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera)... 243
930409.1 Euchaetes nancyae Nagle & Schmidt, 2018
930468.1 Nyridela xanthocera (Walker, 1856)
930541.1 ursania lycimnia (Druce, 1891)
930589.1 Hypena opulenta (Christoph, 1877)
930714.1 Glympis holothermes Hampson, 1926
930723.1 Nychioptera basipallida Barnes & McDunnough, 1916
930730 Hyperstrotia villicans (Barnes & McDunnough, 1918)
930822.1 Catocala ventura Borth & Kons, 2016
930829.1 Catocala slotteni Kons & Borth, 2016
930834.1 Catocala bastropi Kons & Borth, 2017
930887.1 Bulia mexicana (Behr, 1870)
930913 Drasteria howlandii (Grote, 1865)
syn. D. tejonica (Behr, 1870)
930968.11 Mimophisma forbesi Schaus, 1940
930969.1 Tyrissa recurva Walker, 1866
931000.1 Toxonprucha scitior (Walker, 1865)
931019 Zale strigimacula (Guenée, 1852)
931020 Delete Zale obsita (Guenée, 1852)
931071.1 Eulepidotis merricki (Holland, 1902)
931143 Nycteola columbiana (H. Edwards, [1874])
931273 Delete Cobubatha ochrocraspis Hampson, 1910
931278 Delete Cobubatha hippotes (Druce, 1889)
931290 Protodeltote Ueda, 1984
931540.1 Dolocucullia poolei Crabo & Hammond, 2018
931659.1 Plagiomimicus yakama Crabo & Wikle, 2018
ssp. P. y. yakama Crabo & Wikle, 2018
ssp. P. y. mojave Wikle & Crabo, 2018
931719 Neogrotella mcdunnoughi Barnes & Benjamin, 1922
syn. N. macdunnoughi, misspelling
931815.1 Sympistis eleaner Adams, 2018
931821.1 Sympistis tenuistriga (McDunnough, 1940)
931927.1 Sympistis ferrirena Crabo, 2018
931970.1 Neotuerta collectiora Todd, 1966
932029.1 Diastema chuza (Druce, 1898)
932045.1 Helicoverpa armigera (Hübner, [1808])
932130.1 Schinia amblys (Dyar, 1913)
932139 Schinia volupia (Fitch, 1868)
syn. S. masoni (Smith, 1896)
932225.1 Elaphria hypophaea (Hampson, 1920)
932587 Eupsilia vinulenta (Grote, 1864)
syn. E. walkeri (Grote, 1864)
932588.1 Eupsilia colorado (Smith, 1903)
932588.2 Eupsilia schweitzeri Lavitt & Wagner, 2016

B. Christian Schmidt et al. / ZooKeys 788: 241–252 (2018)
244
932606.1 Chaetaglaea rhonda Stead & Troubridge, 2016
932615.1 Omphaloscelis lunosa (Haworth, [1809])
932643 Aseptis susquesa (Smith, 1908)
syn. A. monica (Barnes & McDunnough 1918)
932645.1 Aseptis harpi Crabo & Mustelin, 2018
932656 Stretchia plusiaeformis H. Edwards, 1874
syn. S. plusiiformis, misspelling
932781 Orthosia tenuimacula (Barnes & McDunnough, 1913)
syn. O. mediomacula Barnes & McDunnough, 1924
syn. O. nongenerica Barnes & McDunnough, 1924
932806 move to 2924.50
932807 move to 2924.52
932785 Orthimella delis (Grote, 1874) comb. n.
932924.50 Admetovis oxymorus Grote, 1873 moved from 932806
932924.51 Admetovis icarus Crabo & Schmidt, 2018
932924.52 Admetovis similaris Barnes, 1904 moved from 932807
932937.1 Leucania clarescens Möschler, 1890
932953.1 Leucania oregona Smith, 1902
932953.2 Leucania chejela (Schaus, 1921)
932955.1 Leucania rawlinsi Adams, 2001
932961.1 Leucania latiuscula Herrich-Schäer, 1868
933115.1 Rhabdorthodes pattersoni Crabo, 2018
933181.1 Hypotrix lactomellis Wikle & Crabo, 2018
933207 Hydroeciodes serrata (Grote, 1880)
syn. H. ochrimacula (Barnes & McDunnough, 1913)
933663.1 Abagrotis benjamini Franclemont, 1955
Notes
93027.1 Hypoprepia lampyroides – is species is described in the current volume
by Palting et al. (2018).
930215.1 Clemensia umbrata – is species is raised from synonymy by Schmidt
and Sullivan (2018).
930216 Clemensia ochracea – is species is described in the current volume by
Schmidt and Sullivan (2018).
930233 – 930238 Chelis – e genera Holoarctia, Neoarctia, and Hyperborea were
subsumed under Chelis by Rönkä et al. (2016).
930239 – 930277 Apantesis – e genera Holarctia, Grammia, and Notarctia were
subsumed under Apantesis by Rönkä et al. (2016).
930283 – 930289 Arctia – e genera Parasemia, Acerbia, Pararctia, Platarctia, and
Platyprepia were subsumed under Arctia by Rönkä et al. (2016).

Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera)... 245
930299.1 Virbia marginata – the Virbia taxon from southernmost Texas previously
thought to be V. aurantiaca is in fact more closely related to V. marginata (de-
scribed from Guatemala) based on phenotype and DNA barcode, and this species
is accordingly added to the North American fauna. e western species treated as
V. marginata by Zaspel et al. (2008) is a separate, undescribed species maintained
as Virbia near marginata, as per Lafontaine and Schmidt (2010).
930409.1 Euchaetes nancyae – is species is described in the current volume by
Nagle and Schmidt (2018).
930468.1 Nyridela xanthocera – reported and photographed in southern Texas in
2017 (Krancevic 2018).
930541.1 ursania lycimnia – Several specimens of this Mexican species were col-
lected in southern Texas in 2015 by Ed Knudson. Vouchers are in KBC.
930589.1 Hypena opulenta – this species was approved for release in eastern Can-
ada and is pending approval in the United States of America as a biological
control agent of the invasive European swallow-worts (Vincetoxicum spp.). It
has become established at several locations in eastern Ontario since 2016 (R.
Bourchier, pers. comm.).
930714.1 Glympis holothermes – is species was found at Crocodile Lake National
Wildlife Refuge in the Florida Keys by David Fine in 2009.
930723.1 Nychioptera basipallida – is species was described in the genus Oxycilla
Grote by Barnes and McDunnough (1916) and has remained there until now. e
barcode results suggested the species belonged in the genus Nychioptera Franclem-
ont in the Boletobiinae and not in Oxycilla in the Rivulinae and examination of
the male genital characters conrmed the new placement as Nychioptera basipal-
lida (Barnes & McDunnough, 1916), comb. n.
930730 Hyperstrotia villicans – is species was synonymized with H. nana
(Hübner, 1818) by Lafontaine and Schmidt (2015). New barcode data has shown
there are two similar species going under the name H. nana. Specimens of the
more southern of the two species, with a range extending from Pennsylvania and
Illinois southward to Florida, closely match the holotype of H. villicans. We
therefore re-instate the name Hyperstrotia villicans, stat. rev. for this taxon. e
type specimen for H. nana, like those of most names published by Hübner, is lost
or destroyed, but the rather schematic painting in Hübner (1818: 14, gs53,
54) more closely resembles the more widespread species that occurs from south-
ern Canada to Florida, so we apply the name H. nana to this species. A neotype
should be selected for H. nana when the genus Hyperstrotia Hampson is revised.
930822.1 Catocala ventura – Described as being a new species similar to but distinct
in appearance and barcodes from Catocala californiensis Brower and C. johnsoniana
Brower (Borth and Kons 2016).
930829.1 Catocala slotteni – Described from the Florida Panhandle as a southern rel-
ict species related to Catocala whitneyi Dodge from the northern prairies. Both spe-
cies are associated with leadplant (Amorpha L., Fabaceae) (Kons and Borth 2016).

B. Christian Schmidt et al. / ZooKeys 788: 241–252 (2018)
246
930834.1 Catocala bastropi – Described from western Louisiana and eastern Texas,
this new species occurs west of the known range of Catocala louiseae Bauer, which
occurs as far west as Alabama (Kons and Borth 2017).
930887.1 Bulia mexicana – Recently discovered to occur in southern Texas. One of
the two vouchers in KBC has been barcoded.
930913 Drasteria howlandii – Typical Drasteria howlandii is a species of the Great
Plains and western mountainous areas and is replaced farther south by a paler
desert form in which females look like typical D. howlandii but males have more
white in the hind wing. Richards (1939) treated them as separate species because
their range only overlapped slightly, but he also pointed out that there were many
exceptions to the “species” characters with wrong “forms” showing up in each oth-
er’s territory and suggested the two “species” hybridize where their ranges meet.
e barcodes also do not match either distribution or color forms, and unlike
other closely-related Drasteria, barcode variation is also very low; we therefore treat
D. tejonica (Behr, 1870), syn. n. as a geographic form of D. howlandii.
930968.11 Mimophisma forbesi – is species was previously known only from
Puerto Rico, but found in the Florida Keys by Jim Troubridge in 2013. e speci-
men has been barcoded.
930969.1 Tyrissa recurva – Specimens were collected at the National Key Deer Ref-
uge in the Florida Keys by David Fine and Jim Troubridge.
931000.1 Toxonprucha scitior – is species was described from northwestern Gua-
temala but is now known to occur through Mexico and into the Hill Country of
Texas. Vouchers are in the collection of Hugo Kons Jr., Florida, and the Biodiver-
sity Institute of Ontario, University of Guelph, Ontario.
931019 Zale strigimacula – e identity of this species has been a mystery for many
years. It had been reported from Florida, but specimens from Florida identied as
Z. strigimacula and those identied as Z. viridans (Guenée, 1852) were found to
represent a single undescribed species unknown from the Neotropics or elsewhere
in the Caribbean, so both species were removed from the check list of Canada and
United States (Lafontaine and Schmidt 2010) in a recent list update (Lafontaine
and Schmidt (2015). Unfortunately, the abdomen of the male lectotype borrowed
from MNHP by J. G. Franclemont for dissection has been lost. However, Neo-
tropical specimens in USNM dissected and identied as Zale strigimacula, and
presumably compared to the dissection of the lectotype, give a clue to the identity
of this species. e species occurs from Brazil northward into southern Texas and
is therefore placed back on the North American checklist; this species is, however,
not known to occur in Florida. At least one additional species belonging to the Z.
strigimacula complex is known from Texas.
931020 Zale obsita – As with Zale strigimacula above, there has been much confusion
as to the correct identity of this species. Specimens identied as Zale obsita from
Florida are now reidentied as the same undescribed species discussed under Z.
strigimacula, so this particular species has been the basis for the incorrect reports
of Zale strigimacula, Z. obsita, and Z. viridans from Florida. e female genitalia

Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera)... 247
of the obsita type in the NHMUK is unique in having a single elongated lobe to
the corpus bursae, unlike the gure 8-shaped bursa of species in the Z. strigimacula
complex. Alberto Zilli pointed out that the genitalia of specimens from the Galapa-
gos Islands and treated as Z. obsita by Hayes (1973) were good matches for the type
specimen from Brazil. Dissection and barcoding of specimens from the Galapagos
in the CNC conrms this identication and show that Z. obsita is known from
Brazil, Ecuador, Venezuela, Costa Rica, and Guatemala, but not from farther north.
931071.1 Eulepidotis merricki – Specimens were collected at the National Key Deer
Refuge in the Florida Keys by Jim Troubridge and David Fine. A specimen has
been barcoded.
931143 Nycteola columbiana – is species was described in the Proceedings of the
California Academy of Sciences, volume 5, 1873, with internal dates through the
volume indicating the various fascicles within it were printed in various months of
1873. However, Poole (1989: 1039) and Nye (1975: 410 [under Pseudalypia]) give
1874 as the year of publication for this volume. e year used by Nye is based on
the postal cancellation date in the library of NHMUK – the library being one of
the few that save the postal wrappers. So the date is corrected here to 1874. ere
being no internal evidence of the Edwards paper being published in 1874, the cor-
rected year is in brackets. Contributed by Lars Crabo.
931273 Cobubatha ochrocraspis – is species was added to the North American list
(Lafontaine and Schmidt 2010) on the basis of specimens identied as Cobubatha
ochrocraspis in USNM that closely resemble Cobubatha metaspilaris Walker, 1863
from Florida and the Caribbean. Examination of the holotype in the NHMUK
shows that the specimens in USNM were incorrectly associated with this name
and are Cobubatha metaspilaris, which is now known to occur in the United States
in Florida, Texas, and Arizona. e holotype of Cobubatha ochrocraspis belongs in
the genus Tripudia Grote, as Poole (1989) correctly determined. Tripudia ochro-
craspis comb. rev. occurs from Jalapa in southern Mexico to Costa Rica.
931278 Cobubatha hippotes – is species was described from Guatemala in 1889. It
was reported as C. hippotes in the Noctuidae MONA check list (Franclemont and
Todd 1983), but the species recorded in Texas is now known to be an undescribed
species related to C. hippotes.
931290 Protodeltote – Both Protodeltote and Deltote were recognized as valid genera
in Lafontaine and Schmidt (2010) following the revision by Ueda (1984), but at
the time we were unaware that Protodeltote had recently been subsumed within
Deltote as a subgenus by Fibiger et al. (2009). Despite the apparent similarity be-
tween the two genera, phylogenetic analysis shows that the two are in fact not
closely related (BCS, unpubl. data), and we therefore re-instate Protodeltote stat.
re v. as a valid genus as proposed by Ueda (1984).
931540.1 Dolocucullia poolei – is species is described in the current volume by
Crabo et al. (2018).
931659.1 Plagiomimicus yakama – is species, with two constituent subspecies, is
described in the current volume by Crabo et al. (2018).

B. Christian Schmidt et al. / ZooKeys 788: 241–252 (2018)
248
931719 Neogrotella mcdunnoughi – e species name was misspelled as macdun-
noughi following the spelling in Franclemont and Todd (1983). Contributed by
Greg Pohl & Steve Nanz.
931815.1 Sympistis eleaner – is taxon is described in Adams and Schmidt (2018)
in the current volume.
931821.1 Sympistis tenuistriga – Sympistis badistriga var. tenuistriga (McDunnough,
1940) was rst treated as a valid species in Pohl et al. (2018) based on genital and
barcode dierences.
931927.1 Sympistis ferrirena – is species is described in the current volume by
Crabo et al. (2018).
931970.1 Neotuerta collectiora – is taxon was described as a Cuban subspecies of
Neotuerta sabulosa (Felder, 1874), a species mainly occurring in Central and South
America and the Caribbean as far north as Puerto Rico. Research by Jim Trou-
bridge indicates the Cuban taxon should be raised to species status as Neotuerta
collectiora Todd, 1966, stat. n. It was collected at Crocodile Lake National Wildlife
Refuge in the Florida Keys in 2016 by David Fine.
932029.1 Diastema chuza – Diastema chuza (Druce, 1898), comb. n. was included
in the genus Eustrotia Hübner by Hampson (1910) and Poole (1989), but the bar-
codes and genitalia associate it with the genus Diastema Guenée. It has been found
in Texas in Starr County.
932045.1 Helicoverpa armigera – e Old World Bollworm, a signicant pest spe-
cies native to the eastern hemisphere, is now also established in South America.
is species was detected in Florida (Manatee County: Bradenton) in 2015, but
appears not to have become established (USDA 2017). is species has the poten-
tial to become an agricultural pest in North America (USDA 2017).
932130.1 Schinia amblys – is mainly Mexican species has recently been found in
southeastern Arizona (D. Wikle pers. comm.).
932139 Schinia volupia – Synonymy with Schinia masoni from Pogue et al. 2013.
932225.1 Elaphria hypophaea – Specimens of Elaphria fuscimacula (Grote, 1881)
from southern Texas southward have been re-identied by JDL as the central and
northern South American species Elaphria hypophaea on the basis of barcodes and
dierences in the male genitalia. Elaphria fuscimacula is a complex of three species
that occur from Florida and North Carolina to central Texas. e type locality of
Monodes fuscimacula Grote is Tallahassee, Florida.
932587 Eupsilia vinulenta – e name Eupsilia walkeri (Grote, 1864) was trans-
ferred from the synonymy of Eupsilia sidus (Guenée, 1852) to the synonymy of
Eupsilia vinulenta (Grote, 1864) by Lavitt and Wagner (2016).
932588.1 Eupsilia colorado – is name was previously treated as a synonym of
Eupsilia sidus (Guenée, 1852), but was raised to the status of a valid species by
Lavitt and Wagner (2016). It occurs in southwestern Colorado, highly isolated
from populations of Eupsilia sidus in eastern United States.
932588.2 Eupsilia schweitzeri – is new species was initially distinguished from E.
sidus by barcode and larval dierences, but also diers in details of the male geni-
talia (Lavitt and Wagner 2016).

Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera)... 249
932606.1 Chaetaglaea rhonda – is recently described species (Stead and Trou-
bridge 2016) refers to populations from the Carolinas northward to southern On-
tario previously identied as Chaetaglaea tremula.
932615.1 Omphaloscelis lunosa – A European introduction rst reported from North
America at Potomac, Maryland, 7 October 2015, by Tomas Mustelin.
932643 Aseptis susquesa – e synonymy of Aseptis monica and A. susquesa by Muste-
lin and Crabo 2015 was in inadvertently missed in Lafontaine and Schmidt 2015.
932645.1 Aseptis harpi – is species is described in the current volume by Crabo et
al. (2018).
932656 Stretchia plusiaeformis – e species name plusiaeformis was incorrectly up-
dated to plusiiformis by Lafontaine and Schmidt (2015) following Poole (1989). e
correct original spelling is plusiaeformis. Contributed by Greg Pohl & Steve Nanz.
932781 Orthosia tenuimacula – Barcodes and dissections conrm that O. medio-
macula Barnes & McDunnough, 1924, syn. n. and O. nongenerica Barnes & Mc-
Dunnough, 1924, syn. n. are color forms of Orthosia tenuimacula.
932785 Orthimella Schmidt & Lafontaine, nom. n. is proposed here as an objective
replacement name for Himella Grote, 1874 [Noctuinae: Orthosiini, type species
Himella delis Grote, 1874], a junior homonym of Himella Dallas 1852 [Hemip-
tera: Coreidae], a valid genus of neotropical coreids. is action results in the fol-
lowing new combination: Orthimella delis (Grote, 1874) comb. n.
932806 – see entry for 2924.51
932807 – see entry for 2924.51
932924.51 Admetovis icarus – is species is described in the current volume by
Crabo and Schmidt (2018). e genus Admetovis is re-classied as a member of
the tribe Hadenini from its previous placement in the Orthosiini, resulting in the
re-assignment of checklist numbers from 932806 – 2807.
932937.1 Leucania clarescens – is species was described from Puerto Rico. Jim Trou-
bridge collected specimens at Bahia Honda State Park and Crocodile Lake National
Wildlife Refuge in the Florida Keys and identication was made by Cli Ferris from
the male genitalia of one these specimens. e other specimen has been barcoded.
932953.1 Leucania oregona – is species was segregated from Leucania farcta (Grote,
1881) by Lafontaine and Schmidt (2010) and recognized as a valid species because
of signicant dierences in the genitalia. Barcode results indicate it is closely related
to Leucania imperfecta Smith, 1894, and the genitalia conrm this association, so
we give it a new sequence number to reect its proper position within Leucania.
932953.2 Leucania chejela – is Caribbean and Central American species was dis-
covered at Bahia Honda State Park in the Florida Keys by Jim Troubridge in 2013.
Specimens have been dissected and barcoded.
932955.1 Leucania rawlinsi – is species was described from Jamaica, but extends as
far north as Cuba, the Bahamas, and recently was collected at the National Key Deer
Refuge in the Florida Keys by Jim Troubridge. e specimen has been barcoded.
932961.1 Leucania latiuscula – is species was described from Cuba and was col-
lected at the National Key Deer Refuge in the Florida Keys by Jim Troubridge. A
specimen has been barcoded.

B. Christian Schmidt et al. / ZooKeys 788: 241–252 (2018)
250
933115.1 Rhabdorthodes pattersoni – e genus Rhabdorthodes and the three con-
stituent species are newly described in the current volume by Crabo (2018).
933181.1 Hypotrix lactomellis – is species is described in the current volume by
Crabo et al. (2018).
933207 Hydroeciodes serrata – H. ochrimacula, syn. rev. does not dier from H. ser-
rata in structural characters or barcodes, so we consider it to be a form of H. serrata
and treat it as a synonym.
933663.1 Abagrotis benjamini – is taxon was described as a “race” of Abagrotis
crumbi. It was raised to a valid species by Goldstein and Nelson (2017).
Acknowledgements
Many friends and colleagues sent us suggestions for additions and corrections to this
list and gave us access to material in their care. We appreciate the help of James Ad-
ams, Gary Anweiler, Jerome Barbut, Charles Bordelon, Lars Crabo, Cli Ferris, Martin
Honey, Ed Knudson, Leroy Koehn, Eric Metzler, Steve Nanz, Paul Opler, Mike Pogue,
Greg Pohl, Bob Poole, Bo Sullivan, Jim Vargo, Dave Wagner, Bruce Walsh, Dave Wikle
and Alberto Zilli. Eric Maw brought to our attention the homonymy of Himella. James
Adams served as Academic Editor and he, along with Bo Sullivan and Lars Crabo who
served as reviewers, made suggestions that improved the manuscript. We especially
thank Kristie Killam (National Key Deer Refuge), Jeremy Dixon (Crocodile Lake Na-
tional Wildlife Refuge), and Eric Kiefer, Meredith Kruse, and Janice Duquesnel (Bahia
Honda State Park) for facilitating and supporting research in the reserves in their care.
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