Diversity of moths (Lepidoptera: Heterocera) in the surroundings of the Bednja River, Varaždin County, Northern Croatia

Abstract

The first survey of the moth fauna on the Natura 2000 site Livade uz Bednju I to IV and its near surroundings was conducted in 2016. Four locations were selected and visited three times in different seasons. A total of 381 moth species were recorded, with the most numerous families being Geometridae (102 species), Noctuidae (94 species), Erebidae (44 species) and Crambidae (35 species). In order to put the results in perspective, a preliminary checklist of moths in the Varaždin county was created based on the literature overview and this survey, containing 603 species. 189 species recorded during this survey represent new county records. The greatest increase in the number of species was for the Noctuidae (40 species) and Geometridae (19 species) families. Some species like Diachrysia zosimi were previously recorded only a few times in Croatia. The surveyed sites can now be regarded as some of the best studied areas in Croatia in respect to their moth fauna.

Full text

NAT. CROAT. VOL. 27 No 1 111-141 ZAGREB June 30, 2018
original scientific paper/izvorni znanstveni rad
DOI 10.20302/NC.2018.27.6
DIVERSITY OF MOTHS (LEPIDOPTERA:
HETEROCERA) IN THE SURROUNDINGS

COUNTY, NORTHERN CROATIA

Hyla Association, I. Lipovac 7, HR-10000 Zagreb, Croatia (e-mail: toni.koren@hhdhyla.hr)

2712018
The first survey of the moth fauna on the Natura 2000 site Livade uz Bednju I to IV and its near
surroundings was conducted in 2016. Four locations were selected and visited three times in different
seasons. A total of 381 moth species were recorded, with the most numerous families being Geometri-
dae (102 species), Noctuidae (94 species), Erebidae (44 species) and Crambidae (35 species). In order to

based on the literature overview and this survey, containing 603 species. 189 species recorded during
this survey represent new county records. The greatest increase in the number of species was for the
Noctuidae (40 species) and Geometridae (19 species) families. Some species like Diachrysia zosimi were
previously recorded only a few times in Croatia. The surveyed sites can now be regarded as some of
the best studied areas in Croatia in respect to their moth fauna.
: wet meadows, Natura 2000, checklist, fauna
          
2712018
-

-
ometridae (120 vrsta), Noctuidae (94 vrste), Erebidae (44 vrste) te Crambidae (35 vrsta). Kako bismo



je za porodice Noctuidae (40 vrsta) i Geometridae (19 vrsta). Neke vrste poput Diachrysia zosimi



INTRODUCTION
Moths are a numerous and highly diverse group of insects. They are important plant
pollinators with more than two hundred different moth-plant interactions described so
far (, 2016). Moths also represent prey for many invertebrate and vertebrate
species ( et al., 2000;  2004).
The moth fauna of most parts of Croatia has not been the subject of systematic faunis-


112  Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesur roundingsoftheBednjariver,VaraždinCounty
containing a very limited amount of, in almost all cases, more than a century-old data are
available (e.g. , 1884; , 1910; , 1918). In the work of  (1884)

according to the common practice at that time.  (1910) lists only some dozens of
species that he recorded during his short visit to the area and  (1918) mentions only
a number of Microlepidoptera species from the area. Recent papers dealing with the moth

-
tion is extremely rich in all the insect groups, including butterflies and moths. The data
, 2006), while

the Geometridae (, 2008), Noctuidae (&, 1996) and the Sesiidae
( et al., 1997; , 2010). Nevertheless, almost all the mentioned publications
were based on specimens collected at the beginning of the 20th century. Their historical
value is unquestionable but they do not represent the current state of the biodiversity of
the area. The goal of this survey was to gain a baseline insight into the moth diversity of

collected data in perspective, the secondary goal was to create a preliminary checklist of
moth species known from the county based on the available literature and this survey.
MATERIALS AND METHODS

2 and 175 951 inhabitants. The county conta-

Toplice) and a large number of villages and small settlements. The whole area is a mosa-
ic of settlements, agricultural land, forest fragments and grasslands of different kinds.
The two largest rivers in the area are the Drava, which flows along the northern border
of the county, and the Bednja which flows into the Drava within the county. The Bednja
is a tributary of the Drava, and the whole of its course is in Croatia. The river is 133 km
-
ske toplice, Ludbreg and joins the Drava at Mali Bukovac near Ludbreg.
A total of 14 Special Protection Areas (SPAs) sites have been designated in the county,
including the Natura 2000 areas Livade uz Bednju I-V (site codes: HR2001408, HR2001409,
HR2001410, HR2001411, and HR2001412) occupying a total area of 1,534.761 ha. The
Bednja River and its valley constitute an important area of the ecological network, par-
ticularly because of the meadow and pasture habitats, and the associated species (-

meadows (with Alopecurus pratensis and Sanguisorba officinalis), Molinia meadows on
calcareous, peaty or clayey-silt-laden soils (Molinion caeruleae) and hydrophilic edges
higher green with rivers and forests (Convolvulion sepia, Filipendulion, Senecion fluviatilis).
The dominant soils are acidic brown soil on clastic rocks, luviziol on clay and partly
hydroameliorated wetland rendzinas on flysch and soft limestone (, 2015).
The Bednja River area and the surrounding habitat is one of last remaining natural sites
in the county and, consequently large parts of the remaining mesophilous grasslands are
included in the Natura 2000 network. Still, except for basic knowledge about the Natura
2000 species and habitats, almost no other available data exist for the area.

Nat.Croat.Vol.27(1),2018 113

This study took place at four localities in the Natura 2000 areas, Livade uz Bednju
I - IV, or in the near surroundings (Fig. 1). The area Livade uz Bednju V was not inclu-
ded as it was not easily accessible by car. Each locality was visited three times in di-
fferent seasons (spring, summer and autumn). On average, four hours were spent at
each locality on each date, depending on the time of year and the climate conditions.
Five to seven UV light tents were used per locality to attract moths. Most of the moths
attracted were identified in the field and released. Only in cases where the identifica-
tion was questionable were the moths collected and identified in the lab using stan-
dard identification keys (e.g. , 1997;  & , 2012;  et al.,
2012; , 1998; , 2006, 2008, 2010, 2013). Some members of the families
Erebiidae, Geometridae and Noctuidae, and all the encountered Microlepidoptera
were collected and stored in the private collection of the author. The nomenclature
follows the online database Fauna Europaea ( et al., 2014). For several species,
examination of the genital structures was carried out in order to confirm the correct
species identification.
In order to validate the results and compare them with historical records, a preli-
minary checklist of moths of the county was created based on the available literature
(, 1884; , 1910; , 1918, &, 1996,  et al.,
1997; , 2008; , 2010).
List of surveyed localities along with habitats, coordinates, altitude and dates of visits.
Locality Habitat X Y Altitude
(m a.s.l.) Dates of visit
1. 
Cvetlin Village
forest edge, wet
meadows 46,268605 15,950217 252 9.5.2016, 12.7.2016,
5.10.2016
2. Ivanec, lakes
forest edge, wet
meadows, edge of
the lakes
46,230913 16,102167 207 6.4.2016, 11.7.2016,
14.11.2016
3. Novi Marof, Ivci,
riverbank of Bednja
forest edge, wet
meadows 46,153744 16,364668 187 7.6.2016, 18.7.2016,
23.10.2016
4. Donji Martijanec,
east of the village
forest edge, wet
meadows 46,230737 16,552234 168 18.5.2016, 25.8.2016,
2.11.2016
RESULTS
During this survey we gathered data of 381 moth species within the study area. The
Geometridae are the best-represented group with a total of 102 recorded species,
followed by Noctuidae with 94 species, Erebiidae with 44 and Crambidae with 35
species. All four localities had a similar number of species and abundance of specimens
across three different seasons (Fig. 2).

was created. Overall, 189 species recorded during this survey represent new county

County is actually 603 (Appendix I).

114  Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesur roundingsoftheBednjariver,VaraždinCounty
Surveyed locations in the Natura 2000 areas Livade uz Bednju I-IV. Locality numbers follow Tab. 1.
Number of species (a) and specimens (b) per each locality and season. Locality numbers follow
Tab. 1.
List of recorded moth species during this survey. The locality numbers correspond to the ones
given in Tab. 1. Species for which the identification is based on the examination of the genital structu-
res are marked with an * symbol.
Species Location number Total No. of
individuals
1 2 3 4
Chimabachidae
1. Diurnea fagella 6 6
2. Diurnea lipsiella 1 1
Cossidae
3. Cossus cossus (Linnaeus, 1758) 1 5 6
4. Zeuzera pyrina (Linnaeus, 1761) 3 3
Crambidae
5. Agrotera nemoralis (Scopoli, 1763) 3 1 1 5

Nat.Croat.Vol.27(1),2018 115
Species Location number Total No. of
individuals
1 2 3 4
6. Anania coronata (Hufnagel, 1767) 3 3
7. Anania fuscalis 5 5
8. Anania lancealis 2 2
9. Anania stachydalis (Germar, 1821) 2 1 1 1 5
10. Anania verbascalis 5 1 2 8
11. Calamotropha paludella 5 5
12. Cataclysta lemnata (Linnaeus, 1758) 5 5 1 11
13. Catoptria osthelderi 5 5
14. Chrysoteuchia culmella (Linnaeus, 1758) 3 1 4
15. Crambus lathoniellus (Zincken, 1817) 6 6
16. Cynaeda dentalis 1 1
17. Diasemia reticularis (Linnaeus, 1761) 1 1
18. Ecpyrrhorrhoe rubiginalis 5 5
19. Evergestis forcalis (Linnaeus, 1758) 5 1 6
20. Evergestis pallidata (Hufnagel, 1767) 3 3
21. Metasia ophialis (Treitschke, 1829) 2 2
22. Myelois circumvoluta (Fourcroy, 1785) 3 3
23. Nomophila noctuella 1 1 2
24. Nymphula nitidulata (Hufnagel, 1767) 1 1
25. Ostrinia nubilalis 2 3 5 1 11
26. Palpita vitrealis (Rossi, 1794) 2 2
27. Paracorsia repandalis 3 3
28. Parapoynx stratiotata (Linnaeus, 1758) 3 3
29. Paratalanta pandalis 5 5
30. Pediasia contaminella 5 1 6
31. Pleuroptya ruralis (Scopoli, 1763) 5 4 5 2 16
32. Pyrausta aurata (Scopoli, 1763) 5 1 6
33. Pyrausta despicata (Scopoli, 1763) 2 2
34. Pyrausta purpuralis (Linnaeus, 1758) 1 1
35. Sclerocona acutella (Eversmann, 1842) 1 2 3
36. Scoparia pyralella 5 5
37. Sitochroa verticalis (Linnaeus, 1758) 1 1 2
38. Udea accolalis (Zeller, 1867) 1 1
39. Udea ferrugalis 4 3 5 2 14
Drepanidae
40. Drepana curvatula (Borkhausen, 1790) 2 2 4
41. Drepana falcataria (Linnaeus, 1758) 8 1 3 12
42. Falcaria lacertinaria (Linnaeus, 1758) 3 1 1 5
43. Habrosyne pyritoides (Hufnagel, 1766) 9 5 2 16
44. Polyploca ridens (Fabricius, 1787) 1 1
45. Sabra harpagula (Esper, 1786) 3 3
46. Tethea or 2 1 3
47. Thyatira batis (Linnaeus, 1758) 7 5 1 13
48. Watsonalla binaria (Hufnagel, 1767) 4 4
49. Watsonalla cultraria (Fabricius, 1775) 1 1
Elachistidae
50. Agonopterix ocellana (Fabricius, 1775) 20 20

116  Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesur roundingsoftheBednjariver,VaraždinCounty
Species Location number Total No. of
individuals
1 2 3 4
51. Agonopterix yeatiana (Fabricius, 1781) 4 4
Erebidae
52. Amata phegea (Linnaeus, 1758) 3 3
53. Arctia villica (Linnaeus, 1758) 3 3
54. Arctornis l-nigrum (Muller, 1764) 1 1
55. Callimorpha dominula (Linnaeus, 1758) 4 4
56. Calliteara pudibunda (Linnaeus, 1758) 5 5 10
57. Catocala fraxini (Linnaeus, 1758) 4 4
58. Colobochyla salicalis 6 6
59. Dysauxes ancilla (Linnaeus, 1767) 1 1
60. Eilema caniola 1 1
61. Eilema complana (Linnaeus, 1758) 6 6
62. Eilema depressa (Esper, 1787) 4 6 1 11
63. Eilema lurideola (Zincken, 1817) 2 1 3
64. Eilema sororcula (Hufnagel, 1766) 512 6 1 24
65. Euclidia (Euclidia) glyphica (Linnaeus, 1758) 1 1
66. Euplagia quadripunctaria (Poda, 1761) 1 1
67. Euproctis (Sphrageidus) similis (Fuessly, 1775) 3 1 4
68. Herminia grisealis 210 4 2 18
69. Herminia tarsicrinalis (Knoch, 1782) 5 7 3 15
70. Herminia tenuialis (Rebel, 1899) 1 1
71. Hypena proboscidalis (Linnaeus, 1758) 2 2 2 6
72. Hypena rostralis (Linnaeus, 1758) 5 1 6
73. Laspeyria exula 4 1 5
74. Lithosia quadra (Linnaeus, 1758) 4 5 1 10
75. Lygephila craccae 5 5
76. Lygephila pastinum (Treitschke, 1826) 1 1
77. Lymantria dispar (Linnaeus, 1758) 434 11
78. Lymantria monacha (Linnaeus, 1758) 1 4 5
79. Miltochrista miniata (Forster, 1771) 1 1 2
80. Minucia lunaris 1 1
81. Orgyia (Orgyia) antiqua (Linnaeus, 1758) 1 1 3 5
82. Pechipogo strigilata (Linnaeus, 1758) 3 3 6
83. Pelosia muscerda (Hufnagel, 1766) 3 3
84. Penthophera morio (Linnaeus, 1767) 5 5
85. Phragmatobia fuliginosa (Linnaeus, 1758) 2 3 4 1 10
86. Phytometra viridaria (Clerck, 1759) 4 4
87. Polyphaenis sericata (Esper, 1787) 2 5 7
88. Polypogon gryphalis 2 1 1 4
89. Polypogon tentacularia (Linnaeus, 1758) 3 5 8
90. Rivula sericealis (Scopoli, 1763) 2 5 5 5 17
91. Scoliopteryx libatrix (Linnaeus, 1758) 5 5
92. Spilosoma lubricipeda (Linnaeus, 1758) 7 1 7 6 21
93. Spilosoma lutea (Hufnagel, 1766) 4 4 7 15
94. Thumatha senex 2 2
95. Trisateles emortualis 2 2

Nat.Croat.Vol.27(1),2018 117
Species Location number Total No. of
individuals
1 2 3 4
Geometridae
96. Abraxas (Calospilos) sylvata (Scopoli, 1763) 3 3
97. Acasis viretata 2 2 4
98. Aethalura punctulata 321 24
99. Alcis repandata (Linnaeus, 1758) 5 3 8
100. Angerona prunaria (Linnaeus, 1758) 4 2 1 7
101. Ascotis selenaria 1 2 1 4
102. Asthena albulata (Hufnagel, 1767) 1 9 10
103. Asthena anseraria 2 2
104. Biston betularia (Linnaeus, 1758) 4 1 5
105. Cabera exanthemata (Scopoli, 1763) 5 2 2 1 10
106. Cabera pusaria (Linnaeus, 1758) 5 1 2 8
107. Campaea margaritaria (Linnaeus, 1761) 4 3 2 9
108. Camptogramma bilineata (Linnaeus, 1758) 1 1
109. Catarhoe cuculata (Hufnagel, 1767) 1 4 2 1 8
110. Chiasmia clathrata (Linnaeus, 1758) 3 1 4 3 11
111. Chlorissa cloraria 2 4 6
112. Chloroclysta siterata (Hufnagel, 1767) 1 1
113. Cleora cinctaria 1 1
114. Colostygia pectinataria (Knoch, 1781) 5 3 8
115. Colotois pennaria (Linnaeus, 1761) 4 4
116. Cosmorhoe ocellata (Linnaeus, 1758) 1 1
117. Cyclophora (Codonia) linearia 5 3 1 1 10
118. Cyclophora (Codonia) porata (Linnaeus, 1767) 1 1
119. Cyclophora (Codonia) punctaria (Linnaeus, 1758) 3 5 1 9
120. Cyclophora (Cyclophora) annularia (Fabricius, 1775) 8 2 4 6 20
121. Cyclophora (Cyclophora) pendularia (Clerck, 1759) 3 2 5
122. Cyclophora (Cyclophora) ruciliaria 2 1 1 4
123. Ecliptopera capitata 1 1
124. Ecliptopera silaceata 5 5
125. Ectropis crepuscularia 5 1 2 2 10
126. Ematurga atomaria (Linnaeus, 1758) 4 4 5 1 14
127. Ennomos fuscantaria (Haworth, 1809) 4 4 8
128. Ennomos quercinaria (Hufnagel, 1767) 4 4
129. Epione repandaria (Hufnagel, 1767) 3 3
130. Epirrhoe alternata (Muller, 1764) 5 6 2 13
131. Erannis defoliaria (Clerck, 1759) 2 2
132. Euchoeca nebulata (Scopoli, 1763) 3 4 2 9
133. Euphyia biangulata (Haworth, 1809) 1 1
134. Euphyia unangulata (Haworth, 1809) 4 4
135. Gandaritis pyraliata 2 2
136. Geometra papilionaria (Linnaeus, 1758) 5 5
137. Gymnoscelis rufasciata (Haworth, 1809) 1 1 2
138. Heliomata glarearia 5 5
139. Hemistola chrysoprasaria (Esper, 1795) 4 4
140. Hemithea aestivaria 4 4 8
141. Horisme radicaria (de La Harpe, 1855) 5 5

118  Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesur roundingsoftheBednjariver,VaraždinCounty
Species Location number Total No. of
individuals
1 2 3 4
142. Horisme tersata 1 3 4
143. Horisme vitalbata 2 2
144. Hydrelia ammeolaria (Hufnagel, 1767) 3 1 4
145. Hydriomena impluviata 5 5
146. Hypomecis punctinalis (Scopoli, 1763) 5 1 1 7
147. Hypomecis roboraria 5 3 7 3 18
148. Idaea aversata (Linnaeus, 1758) 5 2 1 8
149. Idaea degeneraria 4 4
150. Idaea dimidiata (Hufnagel, 1767) 3 1 4
151. Idaea humiliata (Hufnagel, 1767) 2 2
152. Idaea muricata (Hufnagel, 1767) 1 1 2
153. Idaea ochrata (Scopoli, 1763) 2 2
154. Jodis lactearia (Linnaeus, 1758) 7 2 9
155. Lampropteryx suumata 1 1
156. Ligdia adustata 5 3 8 2 18
157. Lomaspilis marginata (Linnaeus, 1758) 6 5 2 13
158. Lomographa bimaculata (Fabricius, 1775) 2 4 3 1 10
159. Lomographa temerata 5 2 2 9
160. Lycia hirtaria (Clerck, 1759) 4 4
161. Macaria alternata 6 2 8 6 22
162. Macaria liturata (Clerck, 1759) 5 1 5 3 14
163. Macaria notata (Linnaeus, 1758) 1 4 1 6
164. Macaria signaria 3 3
165. Melanthia procellata 2 1 5 2 10
166. Mesoleuca albicillata (Linnaeus, 1758) 10 1 4 15
167. Odontopera bidentata (Clerck, 1759) 3 3
168. Operophtera brumata (Linnaeus, 1758)* 6 6
169. Ourapteryx sambucaria (Linnaeus, 1758) 3 3
170. Parectropis similaria (Hufnagel, 1767) 4 1 5
171. Pareulype berberata 5 5
172. Peribatodes rhomboidaria 1 3 4
173. Perizoma alchemillata (Linnaeus, 1758) 5 4 1 10
174. Perizoma avofasciata (Thunberg, 1792) 2 2
175. Perizoma lugdunaria 4 4
176. Petrophora chlorosata (Scopoli, 1763) 3 3
177. Phaiogramma etruscaria (Zeller, 1849) 3 3
178. Plagodis dolabraria (Linnaeus, 1767) 1 1 2
179. Plagodis pulveraria (Linnaeus, 1758) 7 1 8
180. Pungeleria capreolaria 1 1
181. Scopula (Calothysanis) immutata (Linnaeus, 1758) 5 1 6
182. Scopula (Calothysanis) incanata (Linnaeus, 1758) 10 10
183. Scopula (Calothysanis) subpunctaria 4 4
184. Scopula (Scopula) immorata (Linnaeus, 1758) 2 2
185. Scopula (Scopula) nigropunctata (Hufnagel, 1767) 6 1 7
186. Scopula (Scopula) ornata (Scopoli, 1763) 1 1
187. Scopula (Scopula) tessellaria (Boisduval, 1840) 2 2
188. Scotopteryx luridata (Hufnagel, 1767) 4 4

Nat.Croat.Vol.27(1),2018 119
Species Location number Total No. of
individuals
1 2 3 4
189. Selenia dentaria (Fabricius, 1775) 1 1 4 6
190. Selenia lunularia 2 1 1 4
191. Selenia tetralunaria (Hufnagel, 1767) 5 3 1 9
192. Thera variata 4 1 5
193. Timandra comae Schmidt, 1931 1 4 3 8
194. Xanthorhoe designata (Hufnagel, 1767) 4 4
195. Xanthorhoe ferrugata (Clerck, 1759) 6 2 11 19
196. Xanthorhoe uctuata (Linnaeus, 1758) 5 5
197. Xanthorhoe spadicearia 3 3
Gracillariidae
198. Parectopa robiniella Clemens, 1863 1 1
Hepialidae
199. Triodia sylvina (Linnaeus, 1761) 1 1
Incurvariidae
200. Incurvaria oehlmanniella 6 1 7
Lasiocampidae
201. Euthrix potatoria (Linnaeus, 1758) 3 3
202. Gastropacha (Gastropacha) quercifolia (Linnaeus, 1758) 1 1 2
203. Lasiocampa (Lasiocampa) quercus (Linnaeus, 1758) 3 3
204. Macrothylacia rubi (Linnaeus, 1758) 5 5
205. Malacosoma (Clisiocampa) neustria (Linnaeus, 1758) 1 1
206. Odonestis pruni (Linnaeus, 1758) 5 1 6
207. Phyllodesma (Phyllodesma) tremulifolia 5 5
Limacodidae
208. Apoda limacodes (Hufnagel, 1766) 3 1 1 5
209. Heterogenea asella 1 1
Noctuidae
210. Acontia (Emmelia) trabealis (Scopoli, 1763) 2 4 6
211. Acronicta (Acronicta) aceris (Linnaeus, 1758) 1 1
212. Acronicta (Acronicta) leporina (Linnaeus, 1758) 1 1
213. Acronicta (Triaena) cuspis 1 1
214. Acronicta (Triaena) psi (Linnaeus, 1758) 11 11
215. Acronicta (Viminia) euphorbiae 1 1
216. Acronicta (Viminia) rumicis (Linnaeus, 1758) 5 1 6
217. Aedia leucomelas (Linnaeus, 1758) 1 1 2
218. Agrochola (Leptologia) lota (Clerck, 1759) 2 2
219. Agrochola (Sunira) circellaris (Hufnagel, 1766) 2 4 6
220. Agrotis exclamationis (Linnaeus, 1758) 1 1
221. Agrotis segetum 5 5
222. Allophyes oxyacanthae (Linnaeus, 1758) 4 1 5
223. Amphipyra (Amphipyra) pyramidea (Linnaeus, 1758) 3 1 4
224. Anarta (Calocestra) trifolii (Hufnagel, 1766) 1 1
225. Anorthoa munda 2 2
226. Asteroscopus sphinx (Hufnagel, 1766) 6 1 7
227. Autographa gamma (Linnaeus, 1758) 4 1 1 6
228. Axylia putris (Linnaeus, 1761) 4 2 12 18
229. Callopistria juventina (Stoll, 1782) 2 2 1 5

120Koren,T.:Diversit yofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheBednjariver,VaraždinCounty
Species Location number Total No. of
individuals
1 2 3 4
230. Caradrina (Paradrina) clavipalpis Scopoli, 1763 1 1
231. Cerastis leucographa 1 1
232. Cerastis rubricosa 3 3
233. Charanyca (Charanyca) trigrammica (Hufnagel, 1766) 2 2
234. Charanyca (Rusina) ferruginea (Esper, 1785) 2 2
235. Colocasia coryli (Linnaeus, 1758) 2 4 4 5 15
236. Conistra (Conistra) ligula (Esper, 1791) 1 1
237. Conistra (Conistra) vaccinii (Linnaeus, 1761) 1 5 6
238.
Conistra (Dasycampa) erythrocephala
2 1 3
239. Conistra (Dasycampa) rubiginea 1 1
240. Cosmia (Nemus) pyralina 2 2
241. Craniophora ligustri 5 3 1 9
242. Cryphia (Euthales) algae (Fabricius, 1775) 2 3 5
243. Deltote (Deltote) bankiana (Fabricius, 1775) 4 1 10 217
244. Deltote (Deltote) uncula (Clerck, 1759) 8 8
245. Deltote (Protodeltote) pygarga (Hufnagel, 1766) 2 4 2 6 14
246. Diachrysia chrysitis (Linnaeus, 1758) 1 1
247. Diachrysia stenochrysis (Warren, 1913) 1 1
248. Diachrysia zosimi 1 1 2
249. Diloba caeruleocephala (Linnaeus, 1758) 1 1
250. Dypterygia scabriuscula (Linnaeus, 1758) 2 2
251. Elaphria venustula 5 2 11 18
252. Eucarta amethystina 3 3 2 8
253. Eucarta virgo (Treitschke, 1835) 554 14
254. Euplexia lucipara (Linnaeus, 1758) 4 4 8
255. Eupsilia transversa (Hufnagel, 1766) 2 4 1 7
256. Hydraecia micacea (Esper, 1789) 2 2
257. Ipimorpha retusa (Linnaeus, 1761) 1 1 2
258. Lacanobia (Dianobia) thalassina (Hufnagel, 1766) 1 1
259. Lacanobia (Diataraxia) oleracea (Linnaeus, 1758) 5 5
260. Lacanobia (Diataraxia) splendens 1 1
261. Lateroligia ophiogramma (Esper, 1794) 4 1 2 7
262. Macdunnoughia confusa (Stephens, 1850) 1 1
263. Meganephria bimaculosa (Linnaeus, 1767) 1 1
264. Melanchra persicariae (Linnaeus, 1761) 5 1 6
265. Mesapamea secalis (Linnaeus, 1758)* 1 1
266. Mesoligia furuncula 3 3
267. Moma alpium (Osbeck, 1778) 1 2 3
268. Mormo maura (Linnaeus, 1758) 1 1
269. Mythimna (Hyphilare) albipuncta 1 4 5
270. Mythimna (Hyphilare) congrua 1 1
271. Mythimna (Hyphilare) l-album (Linnaeus, 1767) 3 3
272. Mythimna (Mythimna) impura 1 1 1 3
273. Mythimna (Mythimna) pallens (Linnaeus, 1758) 2 1 3
274. Mythimna (Mythimna) pudorina 2 2
275. Mythimna (Mythimna) turca (Linnaeus, 1761) 5 2 7
276. Noctua comes 1 1

Nat.Croat.Vol.27(1),2018 121
Species Location number Total No. of
individuals
1 2 3 4
277. Noctua mbriata (Schreber, 1759) 4 1 5
278. Noctua interposita 1 1
279. Noctua orbona (Hufnagel, 1766) 4 4
280. Noctua pronuba (Linnaeus, 1758) 3 2 5
281. Ochropleura plecta (Linnaeus, 1761) 8 4 8 2 22
282. Oligia latruncula 1 1
283. Oligia strigilis (Linnaeus, 1758) 5 5
284. Orthosia (Monima) cerasi (Fabricius, 1775) 5 5
285. Orthosia (Monima) cruda 2 2
286. Orthosia (Monima) miniosa 2 2
287. Orthosia (Orthosia) incerta (Hufnagel, 1766) 1 1
288. Orthosia (Semiophora) gothica (Linnaeus, 1758) 1 1
289. Panolis ammea 1 1
290. Pseudeustrotia candidula 1 4 1 21 27
291. Senta ammea (Curtis, 1828) 5 2 7
292. Sideridis (Aneda) rivularis (Fabricius, 1775) 3 3
293. Simyra albovenosa (Goeze, 1781) 1 1
294. Spodoptera exigua 1 1
295. Tiliacea aurago 5 5
296. Trachea atriplicis (Linnaeus, 1758) 3 1 1 5
297. Tyta luctuosa 4 2 2 8
298. Xanthia (Cirrhia) icteritia (Hufnagel, 1766) 4 4
299. Xanthia (Xanthia) togata (Esper, 1788) 3 4 1 8
300. Xestia (Megasema) c-nigrum (Linnaeus, 1758) 6 6 1 13
301. Xestia (Megasema) triangulum (Hufnagel, 1766) 2 2
302. Xestia (Xestia) castanea (Esper, 1798) 1 1
303. Xestia (Xestia) stigmatica 2 2
Nolidae
304. Bena bicolorana (Fuessly, 1775) 2 2
305. Earias clorana (Linnaeus, 1761) 4 8 12
306. Meganola albula 4 7 1 12
307. Meganola strigula 1 1
308. Nola chlamitulalis 4 4
309. Nola cristatula 1 1
310. Nycteola asiatica (Krulikovsky, 1904) 1 1
311. Nycteola revayana (Scopoli, 1772) 1 1
312. Pseudoips prasinana (Linnaeus, 1758) 3 2 1 6
Notodontidae
313. Clostera anachoreta 5 7 12
314. Clostera anastomosis (Linnaeus, 1758) 1 1
315. Clostera curtula (Linnaeus, 1758) 1 5 6
316. Clostera pigra (Hufnagel, 1766) 3 1 4
317. Drymonia dodonaea 1 1
318. Drymonia melagona (Borkhausen, 1790) 1 5 6
319. Drymonia rucornis (Hufnagel, 1766) 2 2
320. Drymonia velitaris (Hufnagel, 1766) 2 2
321. Furcula bicuspis (Borkhausen, 1790) 1 1

122Koren,T.:Diversit yofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheBednjariver,VaraždinCounty
Species Location number Total No. of
individuals
1 2 3 4
322. Furcula furcula (Clerck, 1759) 8 8
323. Gluphisia crenata (Esper, 1785) 3 3
324. Notodonta dromedarius (Linnaeus, 1767) 1 1
325. Notodonta tritophus 5 5
326. Notodonta ziczac (Linnaeus, 1758) 5 5
327. Pterostoma palpina (Clerck, 1759) 5 4 9
328. Ptilodon capucina (Linnaeus, 1758) 10 111
329. Ptilodon cucullina 1 1 1 3
330. Ptilophora plumigera 1 1
331. Spatalia argentina 4 2 1 7
332. Stauropus fagi (Linnaeus, 1758) 1 7 8
Oecophoridae
333. Harpella forcella (Scopoli, 1763) 2 2
Peleopodidae
334. Carcina quercana (Fabricius, 1775) 3 5 8
Plutellidae
335. Plutella (Plutella) xylostella (Linnaeus, 1758) 5 2 7
Psychidae
336. Bijugis bombycella 1 1
Pterophoridae
337. Emmelina monodactyla (Linnaeus, 1758) 1 1
338. Pterophorus pentadactyla (Linnaeus, 1758) 4 5 9
Pyralidae
339. Aphomia sociella (Linnaeus, 1758) 2 2
340. Dioryctria abietella 4 4
341. Ematheudes punctella (Treitschke, 1833) 4 4
342. Endotricha ammealis 3 3 1 7
343. Homoeosoma sinuella (Fabricius, 1794) 4 4
344. Hypsopygia costalis (Fabricius, 1775) 4 4
345. Nephopterix angustella 4 4
346. Nyctegretis lineana (Scopoli, 1786) 1 1
347. Oncocera semirubella (Scopoli, 1763) 4 2 4 1 11
348. Phycitodes binaevella 4 1 5
349. Pyralis regalis 4 4
350. Sciota adelphella (Fischer v. Röslerstamm, 1836) 3 4 1 8
Saturniidae
351. Aglia tau (Linnaeus, 1758) 2 2
352. Antheraea (Antheraea) yamamai (Guérin-Méneville, 1861) 1 1
353. Saturnia (Eudia) pavoniella (Scopoli, 1763) 1 1
Sphingidae
354. Agrius convolvuli (Linnaeus, 1758) 1 1
355. Deilephila elpenor (Linnaeus, 1758) 4 5 9
356. Deilephila porcellus (Linnaeus, 1758) 3 3
357. Laothoe populi (Linnaeus, 1758) 1 1
358. Macroglossum stellatarum (Linnaeus, 1758) 1 1
359. Mimas tiliae (Linnaeus, 1758) 8 1 9
360. Smerinthus ocellata (Linnaeus, 1758) 7 7

Nat.Croat.Vol.27(1),2018 123
Species Location number Total No. of
individuals
1 2 3 4
361. Sphinx ligustri Linnaeus, 1758 5 5
362. Sphinx pinastri Linnaeus, 1758 5 1 6
Tineidae
363. Euplocamus anthracinalis (Scopoli, 1763) 1 1
Tortricidae
364. Acleris hastiana (Linnaeus, 1758)* 1 1
365. Agapeta hamana (Linnaeus, 1758) 120 21
366. Ancylis mierbacheriana 2 2
367. Ancylis paludana 4 4
368. Archips oporana (Linnaeus, 1758) 5 5
369. Archips xylosteana (Linnaeus, 1758) 1 1
370. Celypha lacunana 3 3
371. Cnephasia (Cnephasia) communana 4 4
372. Epiblema foenella (Linnaeus, 1758) 520 227
373. Eulia ministrana (Linnaeus, 1758) 1 1
374. Hedya salicella (Linnaeus, 1758) 20 424
375. Notocelia cynosbatella (Linnaeus, 1758) 3 1 4
376. Notocelia uddmanniana (Linnaeus, 1758) 5 5
377. Tortrix viridana Linnaeus, 1758 220 123
Yponomeutidae
378. Yponomeuta plumbella 1 1
Zygaenidae
379. Zygaena (Agrumenia) carniolica (Scopoli, 1763) 5 5
380. Zygaena (Mesembrynus) purpuralis 1 1
381. Zygaena (Zygaena) lipendulae (Linnaeus, 1758) 4 4
Number of species per locality: 193 149 151 183 381
Number of specimens per locality: 693 482 552 353 2079
DISCUSSION
With the 381 moth species recorded during this survey, the area of Bednja River can
now be regarded as one of the better surveyed Natura 2000 areas of Croatia in respect to
the moth fauna ( et al., 2011; , 2013; , 1985). Still, the
recorded number probably represents about half of the moth fauna present in the area
in comparison with the nearby best surveyed region, Podravina (, 1985).
In respect to the moth fauna of the county, which now counts 603 species, 189 spe-
cies were recorded for the first time in the area. The increase is partially because even

unpublished part includes all moth families except Geometridae & Noctuidae (and
partially Erebidae as these species were previously treated within the Noctuidae fami-
ly) (, 1996;  2008). Also within Noctuidae and Geometridae
a significant increase in the number of species present in the county was evident with
this survey. An additional 40 species of Noctuidae and 19 species of Geometridae were
recorded as new county records (, 1996;  2008). It is worth
noting that although it is rather short, this is the first moth checklist of any county in
Croatia, although hopefully not the last.

124Koren,T.:Diversit yofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheBednjariver,VaraždinCounty

Crambidae
Catoptria osthelderi
This species is distributed manly in central and northern Europe, with records from
France, northern Italy, Romania, Bulgaria and Yugoslavia (, 2012). The adults fly
from June till September depending on the location. The larvae feed on various mosses.
Five specimens were collected during this survey. The correct identification of this
species was confirmed by the examination of the genital structures (according to ,
2012). The only available record that may be traced back to Croatia originates from maps
present in  (2012), however, it is not mentioned in the text (except possibly as
-
hor (e.g. , 2008;  et al., 2014). Due to its presence in the nearby countries
of Slovenia and Hungary, additional records are expected in the future across northern
Croatia.
Noctuidae
Diachrysia stenochrysis
This species is distributed from Europe, through Asia to Japan. It is generally pro-
blematic, being part of a cryptic species complex, and certain specimens are hardly
distinguishable from one another. In the area of Croatia, the most similar relative is
Diachrysia chrysitis (Linnaeus, 1758). The external morphological appearance and the
genital structure are very similar in these species. They can be reliably identified only
based on the female pheromones ( et al., 2012). One of the better distinguishing
features is the shape of the golden area on the fore wing of D. stenochrysis, which is
usually well connected (rather H-shaped) while it is regularly discontinuous in D.
chrysitis. However, the connective golden streak can also be thin or interrupted in D.
stenochrysis. As the collected specimens show the typical features of D. stenochrysis, they
were included in the analysis. Further studies of the populations from Croatia needs
to be carried out in the future to obtain confirmed data about the occurrence and dis-
tribution of this species-pair.
Diachrysia zosimi

pattern, absent or local in Asia Minor. It is widespread but uncommon in Siberia and
the Pacific Region, very sporadic in western Asia but present and locally frequent or
even common in certain parts of Central and northern Central Europe. In Central Eu-
rope it is bivoltine with two more or less continuous generations from the end of May
to the end of September. In most parts of its area, the caterpillars feed on both Sangu-
isorba officinalis and Parnassia palustris L. but preferring Sanguisorba. This species was
recorded for the first time in Croatia in the Podravina (, 1985) and in Turo-
polje regions ( 1992/93). The records from the Bednja River contri-
bute to the known distribution of this species in Croatia, and indicate it may be more
common than previously thought ( et al., 2008).

Nat.Croat.Vol.27(1),2018 125
Heterogenea asella
This Limacodidae species is usually not very rare, but can be very easily overlooked
during surveys, due to its very small size, grey wings and its inconspicuous overall
appearance, especially in the cases where only Macroheterocera are studied. This Pa-
laearctic species is distributed in most parts of Europe. Only some scattered records of
this species exist from Croatia, most of them from the Podravina region (,
1985).
CONCLUSIONS
While the general trend in moth research includes ecological studies on moths (e.g.
 et al., 2013; , 2003, 2004), Croatia is still without a base-
line knowledge of moth fauna. Faunistic surveys would provide basic data for future
ecological studies and, if properly done, a baseline for further trend assessments. Such
studies are necessary, especially due to the recent changes in land use.

-
bitants lived from agriculture ( 1991); this percentage was reduced in
1991 to 13.7% only, showing a drastic decline (, 2000).
Across the whole Bednja Valley, a clear practice of either complete abandonment or
intensification in use of the grasslands is present. Even in the last few years, the chan-
ges have been very visible, and the area of wet grasslands is shrinking year by year.
With the disappearance of the habitat, plant communities, as well as the connecting
fauna (in this case the hygrophilous moths) are also endangered.
The trends in the moth fauna are not known across Europe due to the absence of
reference data from previous decades, but for some countries like the Netherlands
about one-third of the macromoth species are decreasing (, 2011).
In order to recognise such trends in Croatia in the future, more baseline surveys are
needed.
ACKNOWLEDGMENTS
I am grateful to Ronkay László for useful corrections of and additions to the manus-
cript. This moth survey was supported by the Public Institution for the Management

Received September 22, 2017

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, F., 2010: Pryaloidea (Lepidoptera) of Central Europe. Identifi cation – Distribution – Habitat –
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, K. S. & T. O. , 2003: Determinants of lepidopteran community composition and spe-
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, K. S. & T. O. , 2004: Contrasting effects of habitat quantity and quality on moth com-
munities in fragmented landscapes. Ecography 27, 3-12.

: 224; 230; (4), 18-19; 24-25.

rit Nature Park - Results of preliminary research. Entomologia Croatica , 17-32.
  


T. K oren

-
-
-







vrsta (sovice – Noctuidae) te 19 vrsta (Geometridae - grbice) što ukazuje na nedovoljnu

Diachrysia zosimi tek su relativno

-



128Koren,T.:Diversit yofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheBednjariver,VaraždinCounty

List of species Literature data This survey
Alucitidae
Alucita grammodactyla Zeller, 1841 
Bombycidae
Bombyx mori (Linnaeus, 1758) 
Chimabachidae
Diurnea fagella 
Diurnea lipsiella 
Cossidae
Cossus cossus (Linnaeus, 1758)  
Zeuzera pyrina (Linnaeus, 1761)  
Crambidae
Agriphila geniculea (Haworth, 1811) 
Agriphila tristella 
Agrotera nemoralis (Scopoli, 1763) 
Anania coronata (Hufnagel, 1767) 
Anania fuscalis 
Anania lancealis 
Anania stachydalis (Germar, 1821) 
Anania verbascalis 
Calamotropha paludella 
Cataclysta lemnata (Linnaeus, 1758) 
Catoptria osthelderi 
Chrysoteuchia culmella (Linnaeus, 1758)  
Crambus lathoniellus (Zincken, 1817) 
Crambus perlella (Scopoli, 1763) 
Cydalima perspectalis (Walker, 1859) 
Cynaeda dentalis 
Diasemia reticularis (Linnaeus, 1761)  
Ecpyrrhorrhoe rubiginalis 
Evergestis forcalis (Linnaeus, 1758) 
Evergestis pallidata (Hufnagel, 1767) 
Metasia ophialis (Treitschke, 1829) 
Myelois circumvoluta (Fourcroy, 1785) 
Nomophila noctuella  
Nymphula nitidulata (Hufnagel, 1767) 
Ostrinia nubilalis 
Palpita vitrealis (Rossi, 1794) 
Paracorsia repandalis 
Parapoynx stratiotata (Linnaeus, 1758) 
Paratalanta pandalis 
Pediasia contaminella 
Pleuroptya ruralis (Scopoli, 1763) 
Pyrausta aurata (Scopoli, 1763) 
Pyrausta despicata (Scopoli, 1763)  
Pyrausta purpuralis (Linnaeus, 1758)  
Sclerocona acutella (Eversmann, 1842) 
Scoparia pyralella 

Nat.Croat.Vol.27(1),2018 129
List of species Literature data This survey
Sitochroa verticalis (Linnaeus, 1758) 
Udea accolalis (Zeller, 1867) 
Udea ferrugalis 
Drepanidae
Drepana curvatula (Borkhausen, 1790) 
Drepana falcataria (Linnaeus, 1758) 
Falcaria lacertinaria (Linnaeus, 1758) 
Habrosyne pyritoides (Hufnagel, 1766) 
Polyploca ridens (Fabricius, 1787) 
Sabra harpagula (Esper, 1786) 
Tethea or 
Thyatira batis (Linnaeus, 1758) 
Watsonalla binaria (Hufnagel, 1767) 
Watsonalla cultraria (Fabricius, 1775) 
Elachistidae
Agonopterix ocellana (Fabricius, 1775) 
Agonopterix yeatiana (Fabricius, 1781) 
Endromiidae
Endromis versicolora (Linnaeus, 1758) 
Erebidae
Amata phegea (Linnaeus, 1758)  
Apopestes spectrum (Esper, 1787) 
Arctia villica (Linnaeus, 1758)  
Arctornis l-nigrum (Muller, 1764) 
Callimorpha dominula (Linnaeus, 1758) 
Calliteara pudibunda (Linnaeus, 1758)  
Catephia alchymista 
Catocala electa (Vieweg, 1790) 
Catocala elocata (Esper, 1787) 
Catocala fraxini (Linnaeus, 1758)  
Catocala fulminea (Scopoli, 1763) 
Catocala nupta (Linnaeus, 1767) 
Catocala promissa 
Catocala sponsa (Linnaeus, 1767) 
Colobochyla salicalis  
Cybosia mesomella (Linnaeus, 1758) 
Diacrisia sannio (Linnaeus, 1758) 
Dicallomera fascelina (Linnaeus, 1758) 
Dysauxes ancilla (Linnaeus, 1767) 
Eilema caniola 
Eilema complana (Linnaeus, 1758) 
Eilema depressa (Esper, 1787) 
Eilema lurideola (Zincken, 1817) 
Eilema sororcula (Hufnagel, 1766) 
Euclidia (Euclidia) glyphica (Linnaeus, 1758)  
Euplagia quadripunctaria (Poda, 1761) 
Euproctis (Euproctis) chrysorrhoea (Linnaeus, 1758) 
Euproctis (Sphrageidus) similis (Fuessly, 1775)  

130 Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheB ednjariver,VaraždinCounty
List of species Literature data This survey
Herminia grisealis   
Herminia tarsicrinalis (Knoch, 1782)  
Herminia tarsipennalis (Treitschke, 1835) 
Herminia tenuialis (Rebel, 1899) 
Hypena proboscidalis (Linnaeus, 1758)  
Hypena rostralis (Linnaeus, 1758)  
Hyphoraia aulica (Linnaeus, 1758) 
Idia calvaria  
Laspeyria exula   
Leucoma salicis (Linnaeus, 1758) 
Lithosia quadra (Linnaeus, 1758)  
Lygephila craccae   
Lygephila pastinum (Treitschke, 1826)  
Lymantria dispar (Linnaeus, 1758)  
Lymantria monacha (Linnaeus, 1758)  
Miltochrista miniata (Forster, 1771) 
Minucia lunaris  
Orgyia (Orgyia) antiqua (Linnaeus, 1758) 
Paracolax tristalis (Fabricius, 1794) 
Parascotia fuliginaria (Linnaeus, 1761) 
Pechipogo strigilata (Linnaeus, 1758)  
Pelosia muscerda (Hufnagel, 1766) 
Penthophera morio (Linnaeus, 1767) 
Phragmatobia fuliginosa (Linnaeus, 1758)  
Phytometra viridaria (Clerck, 1759)  
Polyphaenis sericata (Esper, 1787) 
Polypogon gryphalis 
Polypogon tentacularia (Linnaeus, 1758)  
Rivula sericealis (Scopoli, 1763)  
Scoliopteryx libatrix (Linnaeus, 1758)  
Spilosoma lubricipeda (Linnaeus, 1758)  
Spilosoma lutea (Hufnagel, 1766) 
Spilosoma urticae (Esper, 1789) 
Thumatha senex 
Trisateles emortualis  
Tyria jacobaeae (Linnaeus, 1758) 
Zanclognatha lunalis (Scopoli, 1763) 
Geometridae
Abraxas (Abraxas) grossulariata (Linnaeus, 1758) 
Abraxas (Calospilos) sylvata (Scopoli, 1763)  
Acasis viretata 
Aethalura punctulata  
Agriopis aurantiaria 
Agriopis bajaria 
Agriopis leucophaearia 
Agriopis marginaria (Fabricius, 1776) 
Alcis repandata (Linnaeus, 1758)  
Angerona prunaria (Linnaeus, 1758)  

Nat.Croat.Vol.27(1),2018 131
List of species Literature data This survey
Anticollix sparsata (Treitschke, 1828) 
Apeira syringaria (Linnaeus, 1758) 
Aplocera plagiata (Linnaeus, 1758) 
Aplocera praeformata 
Artiora evonymaria 
Ascotis selenaria  
Asthena albulata (Hufnagel, 1767)  
Asthena anseraria 
Biston betularia (Linnaeus, 1758)  
Cabera exanthemata (Scopoli, 1763)  
Cabera pusaria (Linnaeus, 1758)  
Campaea margaritaria (Linnaeus, 1761)  
Camptogramma bilineata (Linnaeus, 1758)  
Catarhoe cuculata (Hufnagel, 1767)  
Catarhoe rubidata 
Cepphis advenaria 
Chiasmia clathrata (Linnaeus, 1758)  
Chlorissa cloraria  
Chlorissa viridata (Linnaeus, 1758) 
Chloroclysta siterata (Hufnagel, 1767) 
Chloroclystis v-ata (Haworth, 1809) 
Cleora cinctaria 
Colostygia olivata 
Colostygia pectinataria (Knoch, 1781)  
Colotois pennaria (Linnaeus, 1761) 
Cosmorhoe ocellata (Linnaeus, 1758)  
Crocallis elinguaria (Linnaeus, 1758) 
Crocallis tusciaria (Borkhausen, 1793) 
Cyclophora (Codonia) linearia  
Cyclophora (Codonia) porata (Linnaeus, 1767)  
Cyclophora (Codonia) punctaria (Linnaeus, 1758)  
Cyclophora (Cyclophora) albipunctata (Hufnagel, 1767) 
Cyclophora (Cyclophora) annularia (Fabricius, 1775)  
Cyclophora (Cyclophora) pendularia (Clerck, 1759) 
Cyclophora (Cyclophora) ruciliaria 
Ecliptopera capitata 
Ecliptopera silaceata  
Ectropis crepuscularia  
Electrophaes corylata (Thunberg, 1792) 
Ematurga atomaria (Linnaeus, 1758)  
Ennomos autumnaria (Werneburg, 1859) 
Ennomos erosaria 
Ennomos fuscantaria (Haworth, 1809)  
Ennomos quercinaria (Hufnagel, 1767)  
Epione repandaria (Hufnagel, 1767)  
Epirrhoe alternata (Muller, 1764)  
Epirrhoe galiata 
Epirrhoe hastulata 

132Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheB ednjariver,VaraždinCounty
List of species Literature data This survey
Epirrhoe rivata 
Epirrhoe tristata (Linnaeus, 1758) 
Epirrita dilutata 
Erannis defoliaria (Clerck, 1759)  
Euchoeca nebulata (Scopoli, 1763) 
Euphyia biangulata (Haworth, 1809)  
Euphyia unangulata (Haworth, 1809)  
Eupithecia absinthiata (Clerck, 1759) 
Eupithecia centaureata 
Eupithecia denotata 
Eupithecia distinctaria 
Eupithecia inturbata 
Eupithecia linariata 
Eupithecia plumbeolata (Haworth, 1809) 
Eupithecia pygmaeata 
Eupithecia subfuscata (Haworth, 1809) 
Eupithecia succenturiata (Linnaeus, 1758) 
Eupithecia vulgata (Haworth, 1809) 
Gandaritis pyraliata  
Geometra papilionaria (Linnaeus, 1758) 
Gnophos (Gnophos) furvata 
Gymnoscelis rufasciata (Haworth, 1809)  
Heliomata glarearia  
Hemistola chrysoprasaria (Esper, 1795)  
Hemithea aestivaria  
Horisme corticata (Treitschke, 1835) 
Horisme radicaria (de La Harpe, 1855) 
Horisme tersata  
Horisme vitalbata  
Hydrelia ammeolaria (Hufnagel, 1767)  
Hydrelia sylvata 
Hydria cervinalis (Scopoli, 1763) 
Hydria undulata (Linnaeus, 1758) 
Hydriomena impluviata  
Hypomecis punctinalis (Scopoli, 1763)  
Hypomecis roboraria  
Idaea aureolaria 
Idaea aversata (Linnaeus, 1758)  
Idaea biselata (Hufnagel, 1767) 
Idaea camparia 
Idaea degeneraria  
Idaea dilutaria 
Idaea dimidiata (Hufnagel, 1767)  
Idaea emarginata (Linnaeus, 1758) 
Idaea humiliata (Hufnagel, 1767)  
Idaea inquinata (Scopoli, 1763) 
Idaea laevigata (Scopoli, 1763) 
Idaea moniliata 

Nat.Croat.Vol.27(1),2018 133
List of species Literature data This survey
Idaea muricata (Hufnagel, 1767)  
Idaea ochrata (Scopoli, 1763)  
Idaea rufaria 
Idaea serpentata (Hufnagel, 1767) 
Idaea straminata (Borkhausen, 1794) 
Isturgia arenacearia 
Isturgia murinaria 
Jodis lactearia (Linnaeus, 1758)  
Lampropteryx suumata 
Ligdia adustata  
Lithostege griseata 
Lomaspilis marginata (Linnaeus, 1758)  
Lomographa bimaculata (Fabricius, 1775)  
Lomographa temerata  
Lycia hirtaria (Clerck, 1759)  
Lycia zonaria 
Lythria cruentaria (Hufnagel, 1767) 
Lythria purpuraria (Linnaeus, 1758) 
Macaria alternata  
Macaria liturata (Clerck, 1759)  
Macaria notata (Linnaeus, 1758)  
Macaria signaria 
Macaria wauaria (Linnaeus, 1758) 
Melanthia procellata  
Mesoleuca albicillata (Linnaeus, 1758)  
Minoa murinata (Scopoli, 1763) 
Nebula nebulata (Treitschke, 1828) 
Nycterosea obstipata (Fabricius, 1794) 
Odontopera bidentata (Clerck, 1759) 
Operophtera brumata (Linnaeus, 1758)  
Operophtera fagata (Scharfenberg, 1805) 
Opisthograptis luteolata (Linnaeus, 1758) 
Ourapteryx sambucaria (Linnaeus, 1758) 
Pachycnemia hippocastanaria 
Parectropis similaria (Hufnagel, 1767)  
Pareulype berberata  
Pasiphila debiliata 
Pasiphila rectangulata (Linnaeus, 1758) 
Pelurga comitata (Linnaeus, 1758) 
Peribatodes rhomboidaria  
Peribatodes secundaria 
Perizoma alchemillata (Linnaeus, 1758)  
Perizoma avofasciata (Thunberg, 1792)  
Perizoma lugdunaria 
Perizoma minorata (Treitschke, 1828) 
Petrophora chlorosata (Scopoli, 1763)  
Phaiogramma etruscaria (Zeller, 1849) 
Philereme transversata (Hufnagel, 1767) 

134 Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheB ednjariver,VaraždinCounty
List of species Literature data This survey
Philereme vetulata 
Plagodis dolabraria (Linnaeus, 1767)  
Plagodis pulveraria (Linnaeus, 1758)  
Plemyria rubiginata 
Pseudopanthera macularia (Linnaeus, 1758) 
Pseudoterpna pruinata (Hufnagel, 1767) 
Pungeleria capreolaria  
Rheumaptera subhastata (Nolcken, 1870) 
Rhodometra sacraria (Linnaeus, 1767) 
Rhodostrophia vibicaria (Clerck, 1759) 
Scopula (Calothysanis) emutaria 
Scopula (Calothysanis) immutata (Linnaeus, 1758)  
Scopula (Calothysanis) incanata (Linnaeus, 1758) 
Scopula (Calothysanis) marginepunctata (Goeze, 1781) 
Scopula (Calothysanis) subpunctaria 
Scopula (Scopula) caricaria 
Scopula (Scopula) immorata (Linnaeus, 1758)  
Scopula (Scopula) nigropunctata (Hufnagel, 1767)  
Scopula (Scopula) ornata (Scopoli, 1763)  
Scopula (Scopula) rubiginata (Hufnagel, 1767) 
Scopula (Scopula) tessellaria (Boisduval, 1840) 
Scopula (Scopula) virgulata 
Scotopteryx bipunctaria 
Scotopteryx chenopodiata (Linnaeus, 1758) 
Scotopteryx luridata (Hufnagel, 1767)  
Scotopteryx moeniata (Scopoli, 1763) 
Selenia dentaria (Fabricius, 1775)  
Selenia lunularia  
Selenia tetralunaria (Hufnagel, 1767)  
Siona lineata (Scopoli, 1763) 
Stegania cararia 
Stegania dilectaria 
Thalera mbrialis (Scopoli, 1763) 
Thera variata  
Thetidia (Antonechloris) smaragdaria (Fabricius, 1787) 
Timandra comae Schmidt, 1931  
Trichopteryx polycommata 
Triphosa dubitata (Linnaeus, 1758) 
Xanthorhoe designata (Hufnagel, 1767)  
Xanthorhoe ferrugata (Clerck, 1759)  
Xanthorhoe uctuata (Linnaeus, 1758)  
Xanthorhoe montanata 
Xanthorhoe quadrifasiata (Clerck, 1759) 
Xanthorhoe spadicearia  
Gracillariidae
Parectopa robiniella Clemens, 1863 
Hepialidae
Triodia sylvina (Linnaeus, 1761) 

Nat.Croat.Vol.27(1),2018 135
List of species Literature data This survey
Incurvariidae
Incurvaria oehlmanniella 
Lasiocampidae
Eriogaster (Eriogaster) lanestris (Linnaeus, 1758) 
Euthrix potatoria (Linnaeus, 1758) 
Gastropacha (Gastropacha) quercifolia (Linnaeus, 1758)  
Lasiocampa (Lasiocampa) quercus (Linnaeus, 1758)  
Macrothylacia rubi (Linnaeus, 1758)  
Malacosoma (Clisiocampa) neustria (Linnaeus, 1758)  
Odonestis pruni (Linnaeus, 1758)  
Phyllodesma (Phyllodesma) tremulifolia  
Limacodidae
Apoda limacodes (Hufnagel, 1766) 
Heterogenea asella 
Noctuidae
Abrostola triplasia (Linnaeus, 1758) 
Acontia (Acontia) lucida (Hufnagel, 1766) 
Acontia (Emmelia) trabealis (Scopoli, 1763)  
Acronicta (Acronicta) aceris (Linnaeus, 1758)  
Acronicta (Acronicta) leporina (Linnaeus, 1758)  
Acronicta (Hyboma) strigosa 
Acronicta (Jocheaera) alni (Linnaeus, 1767) 
Acronicta (Triaena) cuspis 
Acronicta (Triaena) psi (Linnaeus, 1758) 
Acronicta (Viminia) euphorbiae 
Acronicta (Viminia) rumicis (Linnaeus, 1758)  
Aedia leucomelas (Linnaeus, 1758) 
Agrochola (Anchoscelis) litura (Linnaeus, 1758) 
Agrochola (Leptologia) lota (Clerck, 1759) 
Agrochola (Leptologia) macilenta 
Agrochola (Sunira) circellaris (Hufnagel, 1766)  
Agrotis bigramma (Esper, 1790) 
Agrotis exclamationis (Linnaeus, 1758)  
Agrotis ipsilon (Hufnagel, 1766) 
Agrotis segetum 
Allophyes oxyacanthae (Linnaeus, 1758) 
Amphipyra (Amphipyra) livida 
Amphipyra (Amphipyra) pyramidea (Linnaeus, 1758)  
Amphipyra (Amphipyra) tragopoginis (Clerck, 1759) 
Anarta (Anarta) myrtilli (Linnaeus, 1761) 
Anarta (Calocestra) trifolii (Hufnagel, 1766)  
Anorthoa munda 
Apamea lithoxylaea 
Apamea monoglypha (Hufnagel, 1766) 
Apamea sordens (Hufnagel, 1766) 
Asteroscopus sphinx (Hufnagel, 1766) 
Autographa gamma (Linnaeus, 1758)  
Axylia putris (Linnaeus, 1761)  

136Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheB ednjariver,VaraždinCounty
List of species Literature data This survey
Bryophila (Bryoleuca) raptricula 
Bryophila (Bryophila) domestica (Hufnagel, 1766) 
Callopistria juventina (Stoll, 1782)  
Calophasia lunula (Hufnagel, 1766) 
Calophasia platyptera (Esper, 1788) 
Caradrina (Paradrina) clavipalpis Scopoli, 1763 
Caradrina (Platyperigea) kadenii Freyer, 1836 
Cerastis leucographa 
Cerastis rubricosa  
Charanyca (Charanyca) trigrammica (Hufnagel, 1766)  
Charanyca (Rusina) ferruginea (Esper, 1785)  
Colocasia coryli (Linnaeus, 1758)  
Conistra (Conistra) ligula (Esper, 1791) 
Conistra (Conistra) vaccinii (Linnaeus, 1761)  
Conistra (Dasycampa) erythrocephala 
Conistra (Dasycampa) rubiginea  
Cosmia (Calymnia) trapezina (Linnaeus, 1758) 
Cosmia (Nemus) pyralina 
Craniophora ligustri  
Cryphia (Cryphia) fraudatricula 
Cryphia (Cryphia) receptricula 
Cryphia (Euthales) algae (Fabricius, 1775)  
Cucullia (Cucullia) umbratica (Linnaeus, 1758) 
Cucullia (Shargacucullia) prenanthis Boisduval, 1840 
Cucullia (Shargacucullia) verbasci (Linnaeus, 1758) 
Deltote (Deltote) bankiana (Fabricius, 1775)  
Deltote (Deltote) uncula (Clerck, 1759)  
Deltote (Protodeltote) pygarga (Hufnagel, 1766)  
Diachrysia chrysitis (Linnaeus, 1758)  
Diachrysia chryson (Esper, 1789) 
Diachrysia stenochrysis (Warren, 1913) 
Diachrysia zosimi 
Dichagyris (Albocosta) ammatra 
Diloba caeruleocephala (Linnaeus, 1758) 
Dypterygia scabriuscula (Linnaeus, 1758)  
Egira conspicillaris (Linnaeus, 1758) 
Elaphria venustula 
Eucarta amethystina  
Eucarta virgo (Treitschke, 1835) 
Euplexia lucipara (Linnaeus, 1758)  
Eupsilia transversa (Hufnagel, 1766)  
Euxoa (Euxoa) tritici (Linnaeus, 1761) 
Hada plebeja (Linnaeus, 1761) 
Hadena (Anepia) perplexa 
Hadena (Hadena) bicruris (Hufnagel, 1766) 
Hecatera bicolorata (Hufnagel, 1766) 
Hecatera dysodea 
Heliothis ononis 

Nat.Croat.Vol.27(1),2018 137
List of species Literature data This survey
Heliothis peltigera 
Heliothis viriplaca (Hufnagel, 1766) 
Hoplodrina ambigua 
Hoplodrina superstes (Ochsenheimer, 1816) 
Hydraecia micacea (Esper, 1789) 
Ipimorpha retusa (Linnaeus, 1761)  
Lacanobia (Dianobia) contigua 
Lacanobia (Dianobia) suasa 
Lacanobia (Dianobia) thalassina (Hufnagel, 1766)  
Lacanobia (Diataraxia) oleracea (Linnaeus, 1758)  
Lacanobia (Diataraxia) splendens 
Lateroligia ophiogramma (Esper, 1794) 
Leucania (Leucania) comma (Linnaeus, 1761) 
Leucania (Leucania) obsoleta 
Lithophane (Lithophane) ornitopus (Hufnagel, 1766) 
Luperina testacea 
Macdunnoughia confusa (Stephens, 1850)  
Mamestra brassicae (Linnaeus, 1758) 
Meganephria bimaculosa (Linnaeus, 1767) 
Melanchra persicariae (Linnaeus, 1761)  
Mesapamea secalis (Linnaeus, 1758)  
Mesoligia furuncula  
Mniotype satura 
Moma alpium (Osbeck, 1778) 
Mormo maura (Linnaeus, 1758)  
Mythimna (Hyphilare) albipuncta  
Mythimna (Hyphilare) congrua 
Mythimna (Hyphilare) l-album (Linnaeus, 1767)  
Mythimna (Mythimna) impura 
Mythimna (Mythimna) pallens (Linnaeus, 1758)  
Mythimna (Mythimna) pudorina 
Mythimna (Mythimna) turca (Linnaeus, 1761)  
Mythimna (Mythimna) vitellina 
Naenia typica (Linnaeus, 1758) 
Noctua comes 
Noctua mbriata (Schreber, 1759)  
Noctua interposita 
Noctua orbona (Hufnagel, 1766) 
Noctua pronuba (Linnaeus, 1758)  
Ochropleura plecta (Linnaeus, 1761)  
Oligia latruncula  
Oligia strigilis (Linnaeus, 1758)  
Orthosia (Monima) cerasi (Fabricius, 1775) 
Orthosia (Monima) cruda 
Orthosia (Monima) miniosa 
Orthosia (Orthosia) incerta (Hufnagel, 1766)  
Orthosia (Semiophora) gothica (Linnaeus, 1758)  
Panemeria tenebrata (Scopoli, 1763) 

138 Koren,T.:Diversityofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheB ednjariver,VaraždinCounty
List of species Literature data This survey
Panolis ammea 
Peridroma saucia 
Phlogophora meticulosa (Linnaeus, 1758) 
Plusia festucae (Linnaeus, 1758) 
Polia nebulosa (Hufnagel, 1766) 
Pseudeustrotia candidula  
Pyrrhia umbra (Hufnagel, 1766) 
Rhyacia simulans (Hufnagel, 1766) 
Senta ammea (Curtis, 1828) 
Sideridis (Aneda) rivularis (Fabricius, 1775)  
Simyra albovenosa (Goeze, 1781) 
Spaelotis ravida 
Spodoptera exigua  
Subacronicta megacephala 
Tholera cespitis 
Tholera decimalis (Poda, 1761) 
Tiliacea aurago  
Trachea atriplicis (Linnaeus, 1758)  
Tyta luctuosa  
Valeria oleagina 
Xanthia (Cirrhia) gilvago 
Xanthia (Cirrhia) icteritia (Hufnagel, 1766)  
Xanthia (Xanthia) togata (Esper, 1788) 
Xestia (Megasema) c-nigrum (Linnaeus, 1758)  
Xestia (Megasema) ditrapezium 
Xestia (Megasema) triangulum (Hufnagel, 1766) 
Xestia (Xestia) castanea (Esper, 1798) 
Xestia (Xestia) stigmatica 
Nolidae
Bena bicolorana (Fuessly, 1775) 
Earias clorana (Linnaeus, 1761)  
Earias vernana (Fabricius, 1787) 
Meganola albula  
 
Nola chlamitulalis 
Nola cristatula 
Nycteola asiatica (Krulikovsky, 1904) 
Nycteola revayana (Scopoli, 1772) 
Pseudoips prasinana (Linnaeus, 1758)  
Notodontidae
Cerura (Cerura) vinula (Linnaeus, 1758) 
Clostera anachoreta 
Clostera anastomosis (Linnaeus, 1758) 
Clostera curtula (Linnaeus, 1758) 
Clostera pigra (Hufnagel, 1766) 
Drymonia dodonaea 
Drymonia melagona (Borkhausen, 1790) 
Drymonia rucornis (Hufnagel, 1766) 

Nat.Croat.Vol.27(1),2018 139
List of species Literature data This survey
Drymonia velitaris (Hufnagel, 1766) 
Furcula bicuspis (Borkhausen, 1790) 
Furcula furcula (Clerck, 1759) 
Gluphisia crenata (Esper, 1785) 
Notodonta dromedarius (Linnaeus, 1767) 
Notodonta tritophus 
Notodonta ziczac (Linnaeus, 1758) 
Phalera bucephala (Linnaeus, 1758) 
Pterostoma palpina (Clerck, 1759) 
Ptilodon capucina (Linnaeus, 1758) 
Ptilodon cucullina 
Ptilophora plumigera 
Spatalia argentina 
Stauropus fagi (Linnaeus, 1758) 
Oecophoridae
Harpella forcella (Scopoli, 1763) 
Peleopodidae
Carcina quercana (Fabricius, 1775) 
Plutellidae
Plutella (Plutella) xylostella (Linnaeus, 1758) 
Psychidae
Bijugis bombycella 
Canephora hirsuta (Poda, 1761) 
Pterophoridae
Emmelina monodactyla (Linnaeus, 1758) 
Pterophorus pentadactyla (Linnaeus, 1758)  
Pyralidae
Aphomia sociella (Linnaeus, 1758) 
Dioryctria abietella 
Ematheudes punctella (Treitschke, 1833) 
Endotricha ammealis 
Ephestia elutella 
Homoeosoma sinuella (Fabricius, 1794) 
Hypsopygia costalis (Fabricius, 1775) 
Nephopterix angustella 
Nyctegretis lineana (Scopoli, 1786) 
Oncocera semirubella (Scopoli, 1763)  
Phycitodes binaevell 
Phycitodes inquinatella (Ragonot, 1887) 
Plodia interpunctella 
Pyralis regalis 
Sciota adelphella (Fischer v. Röslerstamm, 1836) 
Saturniidae
Aglia tau (Linnaeus, 1758) 
Antheraea (Antheraea) yamamai (Guérin-Méneville, 1861) 
Saturnia (Eudia) pavoniella (Scopoli, 1763)  
Saturnia (Saturnia) pyri 

140 Koren,T.:Diversit yofmoths(Lepidoptera:Heterocera)inthesurroundingsoftheBednjariver,VaraždinCounty
List of species Literature data This survey
Sesiidae
Bembecia albanensis (Rebel, 1918) 
Bembecia ichneumoniformis 
Bembecia megillaeformis 
Chamaesphecia euceraeformis (Ochsenheimer, 1816) 
Chamaesphecia hungarica (Tomala, 1901) 
Chamaesphecia nigrifrons (Le Cerf, 1911) 
Chamaesphecia tenthrediniformis 
Paranthrene tabaniformis 
Pyropteron triannuliformis (Freyer, 1843) 
Sesia apiformis (Clerck, 1759) 
Synanthedon cephiformis (Ochsenheimer, 1808) 
Synanthedon conopiformis (Esper, 1782) 
Synanthedon culiciformis (Linnaeus, 1758) 
Synanthedon formicaeformis (Esper, 1783) 
Synanthedon myopaeformis (Borkhausen, 1789) 
Synanthedon spheciformis 
Synanthedon stomoxiformis 
Synanthedon tipuliformis (Clerck, 1759) 
Synanthedon vespiformis (Linnaeus, 1761) 
Sphingidae
Acherontia atropos (Linnaeus, 1758) 
Agrius convolvuli (Linnaeus, 1758)  
Deilephila elpenor (Linnaeus, 1758)  
Deilephila porcellus (Linnaeus, 1758)  
Hemaris tityus (Linnaeus, 1758) 
Hyles euphorbiae (Linnaeus, 1758) 
Laothoe populi (Linnaeus, 1758)  
Macroglossum stellatarum (Linnaeus, 1758)  
Mimas tiliae (Linnaeus, 1758)  
Smerinthus ocellata (Linnaeus, 1758)  
Sphinx ligustri Linnaeus, 1758  
Sphinx pinastri Linnaeus, 1758 
Tineidae
Euplocamus anthracinalis (Scopoli, 1763) 
Tortricidae
Acleris hastiana (Linnaeus, 1758) 
Agapeta hamana (Linnaeus, 1758) 
Ancylis mierbacheriana 
Ancylis paludana 
Archips oporana (Linnaeus, 1758) 
Archips xylosteana (Linnaeus, 1758) 
Celypha lacunana 
Celypha rivulana (Scopoli, 1763) 
Clepsis rurinana (Linnaeus, 1758) 
Cnephasia (Cnephasia) communana 
Epiblema foenella (Linnaeus, 1758)  
Eulia ministrana (Linnaeus, 1758) 

Nat.Croat.Vol.27(1),2018 141
List of species Literature data This survey
Hedya salicella (Linnaeus, 1758)  
Notocelia cynosbatella (Linnaeus, 1758) 
Notocelia uddmanniana (Linnaeus, 1758) 
Pandemis dumetana (Treitschke, 1835) 
Tortrix viridana Linnaeus, 1758 
Yponomeutidae
Argyresthia (Argyresthia) bonnetella (Linnaeus, 1758) 
Yponomeuta plumbella 
Zygaenidae
Zygaena (Agrumenia) carniolica (Scopoli, 1763) 
Zygaena (Mesembrynus) purpuralis 
Zygaena (Zygaena) lipendulae (Linnaeus, 1758)  
Total: 414 381