Biographical Sketch and Bibliography of Jeffrey E. Lovich

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BIOGRAPHICAL SKETCH
AND
BIBLIOGRAPHY OF JEFFREY E. LOVICH
Jeffrey E. Lovich
Southwest Biological Science Center
United States Geological Survey
SMITHSONIAN HERPETOLOGICAL
INFORMATION SERVICE
NO. 153
2018

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SMITHSONIAN HERPETOLOGICAL INFORMATION S
ERVICE
The first number of the SMITHSONIAN HERPETOLOGICAL INFORMATION SERVICE series appeared in
1968. SHIS number 1 was a list of herpetological publications arising from within or through the
Smithsonian Institution and its collections entity, the United States National Museum (USNM). The
latter exists now as little more than the occasional title for the registration activities of the National
Museum of Natural History. No. 1 was prepared and printed by J. A. Peters, then Curator-in-Charge
of the Division of Amphibians & Reptiles. The availability of a NASA translation service and
assorted indices encouraged him to continue the series and distribute these items on an irregular
schedule.
The series continues under that tradition. Specifically, the SHIS series distributes translations,
bibliographies, indices, and similar items judged useful to individuals interested in the biology of
amphibians and reptiles, and unlikely to be published in the normal technical journals. We wish to
encourage individuals to share their bibliographies, translations, etc. with other herpetologists
through the SHIS series. If you have such an item, please contact George Zug [zugg @ si.edu] for
its consideration for distribution through the SHIS series.
Our increasingly digital world is changing the manner of our access to research literature and that is
now true for SHIS publications. They are distributed now as pdf documents through two
Smithsonian outlets:
BIODIVERSITY HERITAGE LIBRARY. www.biodiversitylibrary.org/bibliography/15728
All numbers from 1 to 131 [1968-2001] available in BHL.
DIVISION’S WEBSITE. vertebrates.si.edu/herps/herps_NMNH_herppubs/herps_herps.html
Numbers 84 to 153 available as pdfs in the herpetological publications section of the website.
2018 – Division of Amphibians & Reptiles
National Museum of Natural History
Smithsonian Institution
ISSN 2331-7515
Cover Image: Gopherus agassizi from
Stebbins 1954. Amphibians & Reptiles of Western North America, Plate 26A.

Biographical Sketch
I was born to Janet Grace (née Cook) and George Edward Lovich near midnight on 30 May, 1957 in
Alexandria, Virginia. I was the first of three children that would include my sister (Cynthia - 1959),
and a brother (Robert – 1970, now also a herpetologist). Until I was seven, we lived near the Potomac
River. My friends and I spent many hours in the surrounding wetlands catching frogs and turtles,
including a large snapping turtle that my mother immediately made me return to the “swamp.” Later
we moved to a house in Alexandria, farther from the river but close to Timber Branch Creek, where I
continued my happy pursuit of salamanders, baby birds, raccoons, opossums, and other animals that
my encouraging mother tolerated. However, my favorite place was in the Allegheny Mountains of
Pennsylvania on my paternal grandparent’s farm where my father would take me on nature walks
when we visited. On one of those walks, we found an adult red-spotted newt in a nearby pond, further
cementing my interest in amphibians and reptiles. In light of my fascination with these creatures my
mother bought me a copy of Zim’s Golden Nature Guide on reptiles and amphibians in 1964, my first
book on herpetology, not counting dinosaurs. I still have that book along with many others in that
series.
I was unremarkable as a young student except for my consuming passion for wild things and wild
places. I read all the books I could find on animals, dreaming of seeing various species in their natural
habitats. I had a growing interest in turtles that would eventually translate into a career I could not yet
see or even imagine. Early experiences with box turtles and wood turtles sealed my fate. It wasn’t until
middle school that I started to have science teachers that really encouraged my interest in science. My
high school years were split between George Washington High School (9th and 10th grades) and T.C.
Williams High School (11th and 12th grades) as a result of court-ordered integration of schools in
Virginia. Those were difficult times in America due to civil unrest caused by the Vietnam War as well
as racial tensions. A now famous movie, “Remember the Titans,” documented (in Hollywood fashion)
tensions at T.C. Williams High School during those tumultuous years.
My main hobbies as a teenager were backpacking and fishing, especially in pursuit of native brook
trout in the streams of Shenandoah National Park and the Allegheny Mountains of Pennsylvania. I
came perilously close to becoming an ichthyologist as a result of that hobby and maintaining
aquariums full of various tropical fish. When not fishing, I was able to see a good bit of the world since
my father was a pilot and my mother worked for various airlines. Trips to Asia, Europe, Mexico, and
the Caribbean gave me an increasing appreciation for wildlife worldwide. Despite those experiences, I
was still looking for direction. It was my senior year physics teacher, Mr. Ferris, who encouraged me
to apply to George Mason University in nearby Fairfax, Virginia, setting me on the course to
eventually becoming a professional herpetologist.
I started at George Mason University in the fall of 1975, right after graduating from high school. I
enrolled as a biology major with no clear mentor or road map for what to do next. All I knew is that I
liked animals (not cell biology or genetics!), and I wanted a job in the outdoors. After two years of
floundering through various courses, I ended up taking a year off and moving to the family farm in
Pennsylvania to regroup and spend as much of time fishing and deer hunting as possible. During that

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time, I held jobs as a brick layer assistant, a roughneck on a gas drilling rig, and miscellaneous roles at
a coal company where my grandfather and great-grandfather had worked as miners long before. I quit
my job at the coal company to go back to George Mason University in 1978 and enrolled in an
Ichthyology class with Dr. Carl H. Ernst as the professor. I thought the class would be a breeze due to
my interest in all things related to fish. Nothing was be further from the truth as Carl made it a rigorous
and challenging course. Sometime after the first major exam, I came into class and was the first student
to arrive that morning. Carl was at his lectern, preparing for his lecture, and tersely said, “Your exam is
over there”, pointing to a pile in the front of the room. I fished mine out of the stack and was shocked
to see that I had received a “C.” As I walked slowly back to my seat, he said “I expected better from
you!” That was a turning point for me and from then on I was dead serious about my college career.
As a student in need of direction, Carl took me under his wing, seeing something in me that was worth
his investment of time. I thought, this guy has a good job, a family, a house, and a new car all because
he’s an expert on turtles. That’s what I wanted so I learned as much as I could from him in the last
years of my undergraduate experience. It was the beginning of my quest to find mentors who could
show me the way to becoming a professional herpetologist. In 1979, Carl told me about a job opening
at the National Museum of Natural History (Smithsonian) for a Museum Aide working in the Division
of Amphibians and Reptiles. As a Research Associate in the Division, Carl set me up for an interview
with Dr. Roy McDiarmid. I went in for an interview with Roy late one afternoon, dressed in a sport
coat, tie, and slacks for the meeting. To say I was overdressed is an understatement. After the
interview, Roy asked me to fill out a 3x5 card with my contact information so he could get back to me.
When I handed him the card, he told me I had the job. I was surprised at the sudden decision and asked
him why he offered to hire me when I gave him the card. He told me that I had legible handwriting and
that was a requirement for the work I’d be doing! Needless to say, I was thrilled to be working in the
Biological Survey Unit that traced its roots back to C. Hart Merriam. Later in my career I would
coauthor a paper describing the history of that unit.
As I recall, Congress had appropriated funds to inventory the collections at the Smithsonian shortly
before I stared working there. The Department of Vertebrate Zoology had selected the Division of
Amphibians and Reptiles to pilot an inventory of their turtle and crocodilian collections. There were
about 3,000 specimens of the former group and 600 of the latter. My job, along with my partner on the
project Dave Ross, another herpetologist, was to fill out a data sheet for each specimen with all the
information contained on jar labels, specimen tags, etc. Information gathered included species,
collector, date of collection, locality of collection, etc. We then went to the old leather-bound Catalogs
to confirm or add to those data and made sure the massive library of 3x5 species and locality cards
matched the information we gathered. Data sheets were turned over to a computer technician who
dutifully keypunched the information onto IBM punch cards for processing on a mainframe
somewhere at the Smithsonian. In this day of VertNet and other online databases, it’s easy to forget
that a lot of the information at our fingertips today was once only available if you visited museums and
pulled it out of written records or files. It was a great job as I got to handle all manner of species of
turtles and crocodilians from around the world. In addition, I was working with real herpetologists like
Ron Crombie, Ron Heyer, Roy, and George Zug and regularly seeing other herpetological luminaries

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who would come to the museum to use the collections or the herpetology library. It was my first job as
a professional herpetologist, and it lasted until 1981 when my contract was completed.
While I continued working on my undergraduate degree at George Mason University, I became the
first biology undergrad to convince the Department Chair, Dr. Larry Rockwood, to pay for my travel to
a professional meeting to give a paper: the Society for the Study of Reptiles and Amphibians
symposium in 1979 at the University of Utah. The talk, the first of my career, was about geographic
variation in an Asian turtle now known as Mauremys reevesii. Right before my talk, I was sitting next
to Dr. George Zug from the Smithsonian. He asked me if I was nervous and I remember saying that I
wasn’t. I survived the presentation as a first of many more to come. While at the meetings I had the
opportunity to meet Dr. R. Bruce Bury. He was a big guy with a beard and looked more like a
lumberjack than a herpetologist. I remember walking up to him and gushing that I was a student of
Carl Ernst’s working on turtles and I wanted to know how I could get a job like his. In his deep voice,
he responded, “I have to die!” It was clear that he loved his job. Later in my career when I started work
with the National Biological Survey (later absorbed into the U.S. Geological Survey), I got a job like
his, and he didn’t have to die after all!
One particularly interesting opportunity came along while I was an undergrad at George Mason
University. One of my college friends and class mates was S. Blair Hedges, another herpetologist. At
the time he had a working relationship with Richard (Dick) Highton at the University of Maryland.
Blair was scheduled to help Dick collect salamanders throughout the southeastern US for a couple
weeks but had a conflict. Blair asked if I would go with Dick to cover for him on the trip. I agreed. On
contacting Dick, he explained that we would be spending long days and nights in the field in Virginia,
Kentucky, North Carolina, Tennessee, and Georgia collecting plethodontid salamanders. The deal was
simple, if I worked hard collecting salamanders, he’d feed me all I could eat while we were on the
road. We’d be camping out and bathing in streams, as needed. One night after finishing our collecting,
we parked on a lonely dirt road somewhere in the mountains of Tennessee and prepared to bed down.
It started to rain so Dick put his sleeping bag in the back of the station wagon. I came over to do the
same. He asked what I was doing. I asked him if I could move the coolers full of salamanders out of
the back of the car so I could put my sleeping bag inside since it was raining. He told me that he didn’t
want to put the coolers outside the vehicle at night lest something bad should happen to the valuable
specimens we’d collected. Luckily it didn’t rain very hard, but that night out made me a tougher field
herpetologist! I sure learned a lot about salamanders from Dick. It was great opportunity for someone
just starting out in herpetology.
I graduated with a BS in Biology in 1982 and immediately enrolled in the Master’s degree program for
Biology at George Mason to continue learning from and interacting with Carl. He taught me how to
write my first scientific papers and how to collect data from museum specimens of turtles for
taxonomic analyses that we would continue until much later. We also visited his long-term field site
for turtle research in Lancaster, Pennsylvania where I was able to see bog turtles in the wild. It was a
productive relationship that would last until very recently when Carl finally put down his pen and
retired to spend more time with his children and grandchildren. During my time as a graduate student, I
remember Carl telling me something like, “I’ve done all I could for you. If you want to be a turtle man,

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you need to go study with Whit Gibbons.” I graduated from George Mason with a MS Biology degree
in 1984 as did my fellow graduate student and friend Steve Gotte who landed a career in the Division
of Amphibians and Reptiles working for the U.S. Geological Survey (USGS).
Convincing Whit Gibbons to take me as a doctoral student presented some challenges, and I had no
plan B. After all, I wanted to be a “Turtle man.” Whit was planning on taking a sabbatical to the
Division of Amphibians and Reptiles at the Smithsonian during the same year that I wished to
matriculate into the University of Georgia, Institute of Ecology degree program to pursue a Ph.D. He
was in the beginning stages of preparing to write his classic book, “Life History and Ecology of the
Slider Turtle” that would be published by the Smithsonian Institution Press in 1990. It was ironic that
he was coming north to where I had worked and I wanted to go down to the South to work with him.
Even more ironic was the fact that I would eventually contribute a chapter on melanism in turtles to
that very book. I remember meeting with Whit at the Smithsonian on one of his advance trips. He was
hesitant about taking on new students at the time. He told me that he would not be able to help me in
my first year if any problems came up since he would be on sabbatical. Not wanting to take no for an
answer, I responded, “I’m very self-sufficient, I won’t need any help. What could go wrong?” I sensed
I was gaining ground as a persuasive, if not over-confident, aspiring grad student. Then, Whit said
something along the lines of, “I need my students to be good writers, I can’t accept people who don’t
like to write.” I responded that I liked to write and had publications already, knowing full well that I
didn’t hold a candle to writers like Whit. Fortunately for me, Whit agreed to take me on as his student.
I am forever grateful for that decision.
My first year at the University of Georgia was dedicated to getting as much course work completed as
possible so that I could move on to the university’s research facility at the Savannah River Ecology
Laboratory (SREL) in South Carolina where Whit worked on a daily basis. When the summer of 1985
finally came, Whit told me to come down to the lab to work until school began that fall. There I met
many notable herpetologists including Ray Semlitsch (who had just left SREL and was starting a job at
Memphis State University), Trip Lamb, Justin Congdon, Joe Pechman, David Scott, and many others.
Some were faculty at the time, others were students who later became faculty elsewhere. My job was
to learn as much as I could about the research being conducted at SREL and to help with turtle
trapping efforts and running the Ellenton Bay drift fence. I worked with two technicians at the lab at
that time, Tim Owens and Tony Mills, great guys who knew how to catch animals and have fun doing
it. Other research projects took place off of the Savannah River Site (SRS), where SREL was located,
to provide comparative data to studies that were conducted on the SRS. Studies on the SRS largely
focused on the effects of nuclear fuel production on wildlife and their habitat because SRS produced
weapon grade plutonium and tritium for the nation’s nuclear arsenal. That summer, I was also
introduced to diamond-backed terrapins on a trip to Kiawah Island to continue research on that species
that Whit started in 1983. Today, 35 years later, it is the longest continuous study on that species in the
world. It was a great summer being exposed to the biodiversity of amphibians and reptiles, and to
herpetologists in the Deep South. In 1986, I married Sharon Anne Cannon, who I met while at George
Mason University. As of 2018, we’ve been married for 32 years. We have two children, Justin (1990),
who is a Physician Assistant, and Ashley (1993), who works in the sales end of the Napa Valley wine
industry.

5
Back at the University of Georgia, I had a committee member, Dr. Ron Pulliam who wanted me to take
more classes on campus before leaving for SREL. Whit was a strong advocate for getting me back to
SREL as soon as possible so he could teach me the fine art of field herpetology and I could get on with
my dissertation research. A deal was struck. I was able to go to SREL in 1986 and do directed readings
with Ron to fulfill my course work requirements. Once a week, I would make the 125 mile drive from
Aiken, South Carolina to Athens, Georgia to meet with Ron and discuss the readings. It was a win-win
situation for me. I was able to spend quality time in stimulating discussions with Ron, a distinguished
animal ecologist, and I was able to work at SREL. Ron later went on to be the first Director of the
National Biological Survey (later Service) where I would eventually work, so technically he became
my “boss” later in my career!
I was slow to select a dissertation topic in a veritable “candy store” of herpetological opportunities, but
eventually settled on causes and consequences of sexual size dimorphism in turtles. Subsequently,
Whit and I wrote a monograph on that subject as well as a paper on how exactly to measure the
phenomenon. Sexual dichromatism in turtles was another interest, and I collaborated with Dr. C. Jack
McCoy at Carnegie Museum and Dr. William (Bill) Garstka at the University of Alabama to write a
chapter on that subject in Whit’s slider book mentioned earlier. Working at SREL presented numerous
collaborative opportunities on a diversity of topics. Dr. William Cooper spent several summers doing
research at the lab as a visiting professor. He and I collaborated with Bill on a couple of papers on
hormonal regulation of courtship behavior and melanism in slider turtles. I also collaborated with Jack
McCoy to eventually describe two new species of map turtles, Graptemys ernsti and G. gibbonsi, each
named in honor of two of my mentors to date.
There was little excuse for lack of productivity at SREL. The lab was funded as a prime contractor
with the U.S. Department of Energy. Resources were available to pursue a multitude of research
projects that met the mission of both the SRS and SREL. As a result, I was able to conduct research
and publish papers on other topics, unrelated to my dissertation work, including continuing
collaborations with Carl Ernst. Through my interactions with Whit and Justin Congdon (another of my
committee members), I learned to think critically about how herpetological research fit into the context
of ecological and evolutionary theories and predictions. When I graduated late in 1990 with my Ph.D.,
I had the foundation I needed to pursue the next phase of my career.
From December 1990 until March 1991, I had a postdoctoral position at SREL continuing my research
with Whit and others. In the late summer of 1990, I became aware of a job posting for a biologist to
work for the Bureau of Land Management (BLM) in Riverside, California analyzing demographic data
based on monitoring of Agassiz’s desert tortoise populations. The tortoise had recently been listed as
threatened under the Endangered Species Act, and a team of BLM biologists were trying to learn more
about the species to better manage populations. I began negotiations with Dr. Kristin Berry, the
supervisor for the position, and started that job on 1 April 1991. Colleagues working at BLM included
Dr. Hal Avery and Dr. William I. Boarman. Instead of working on analyzing tortoise demography, I
was directed to start a project evaluating the potential of restoring degraded tortoise habitat. This

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assignment eventually lead to a couple of publications on the sensitivity of ecological processes and
patterns to human disturbance in the California deserts, both Mojave and Sonoran.
After working in Riverside for a year, I became the Lead Wildlife Biologist for the BLM office in
Palm Springs, about 50 miles away. That job was more diverse and tortoises were only one of the
species of concern. I met local herpetologists Dr. Al Muth and Dr. Cameron Barrows and helped with
surveys of the threatened Coachella Valley fringe-toed lizard to determine population trends. I also
worked on projects to control or eradicate the invasive shrub tamarisk (or salt cedar) that infested BLM
Areas of Critical Environmental Concern in the California deserts.
In 1993, the Secretary of the Interior, Bruce Babbitt, decided that it no longer made sense for the
Department to have scientists working in separate bureaus, scattered throughout the BLM, U.S. Fish
and Wildlife Service, National Park Service, Bureau of Reclamation, etc. He decided to create a
National Biological Survey (NBS) that would combine the research scientists in the Department into a
single Bureau to facilitate collaboration and provide more integrative science to resource managers. I
served on an implementation team detail in Washington, D.C. for several weeks. We were tasked with
working out the details of how the organization would be structured and other aspects of getting the
NBS running on 1 October, 1993. This would affect Smithsonian scientists in the Biological Survey
Unit of the U.S. Fish and Wildlife Service as they were transferred to the NBS as well.
Even though I was duty-stationed in Palm Springs for the first few years of the NBS, I was
administratively attached to the National Ecology Research Center, later the Midcontinent Ecological
Science Center, and more recently, the Fort Collins Science Center. That association allowed me to
interact with other herpetologists such as Dr. R. Bruce Bury (who had just left to return to his home
turf in the Pacific Northwest) and Dr. Steve Corn. Other herpetologists were brought together in the
NBS including tortoise biologists Phil Medica and Dr. Todd Esque. The NBS got off to a rocky start
and changed their name to the National Biological Service to emphasize the fact that they were not
conducting surveys per se, but were providing biological research and services to the Department. In
October of 1996, the NBS was eliminated and all employees became part of the Biological Resources
Division of the USGS where I remain.
In 1998, I shifted my career into science management and became a Research Manager at the Western
Ecological Research Center in Sacramento, California. While there I supervised scientists at various
locations, including Piedras Blancas, California; Las Vegas, Nevada; and Tucson, Arizona. Notable
herpetologists I supervised included Dr. Norm Scott, Dr. Cecil Schwalbe, Dr. Todd Esque, and others.
It was an honor to work with those scientists as a comparatively young manager. In 2002, I became the
Center Director of the USGS Western Ecological Research Center. After 13 months in that position, I
accepted a job as the Chief of the USGS Grand Canyon Monitoring and Research Center in Flagstaff,
Arizona. My role was to coordinate USGS research on the effects of Glen Canyon Dam operations on
natural and cultural resources in Grand Canyon National Park. At the time the program received about
10 million dollars per year and launched as many as 40 river trips per year to conduct research in the
Grand Canyon. After two years in that position, I became the Deputy Director of the USGS Southwest

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Biological Science Center, also in Flagstaff. During this time I continued to publish as time permitted,
but research was not my primary assignment.
In 2008, I received a Fulbright Scholarship to visit Morocco. My in-country sponsor and collaborator
was Dr. Mohammed Znari, another herpetologist. I lectured to graduate students at Cadi Ayyad
University in Marrakech and taught them how to conduct ecological research focusing on stripe-
necked terrapins (Mauremys leprosa). After my experience there, I decided that I wished to return to
full time research and collaboration with students and other scientists. Upon my return to the States, I
asked my supervisor if I could do so and she agreed. I returned to my current position as a Research
Ecologist in 2009. Since then, my greatest satisfaction has come from exposing aspiring young
technicians, interns, and postdocs to the full cycle of science, from asking questions, collecting and
analyzing data, to publishing the results in peer-reviewed journals. Several have gone on to advanced
degrees and fulfilling careers.
During my time with the USGS, I have continued to conduct research on turtles throughout the United
States and beyond, coauthoring two editions of the “Turtles of the United States and Canada” with Carl
Ernst. I maintained collaborations with researchers in the East working on various species of map
turtles, coauthoring the description of Graptemys pearlensis in 2010. I continue my research interest in
diamond-backed terrapins started way back in 1985. Many of my collaborations on recent publications
are with my former technician Mickey Agha (currently a Doctoral Candidate at the University of
California, Davis) and my former Post-doctoral associate, Dr. Josh Ennen of the Tennessee Aquarium
Conservation Institute. Both are productive turtle biologists of note who already have impressive
resumes. I’ve had several opportunities to expand my collaboration with foreign scientists. In 2005 I
was honored to be invited as the keynote speaker at a conference on dams in Ethiopia. In 2014, I was
invited to be the keynote speaker at the Second Japanese Freshwater Turtle Symposium in Kobe, Japan
where I initiated collaboration on publications with Japanese scientists. I was honored to become an
Elected Fellow of the Linnean Society of London in March, 2015. The Linnean Society is the world’s
oldest active biological society. Charles Darwin and Alfred Russel Wallace presented papers at a
meeting of the society in 1858 outlining the theory of evolution by natural selection.
Current research is a continuation of now long-term studies on desert turtles including desert tortoises,
Mojave River populations of the Western Pond Turtle, and Diamond-backed Terrapins in South
Carolina. I couldn’t have achieved my goal of becoming a professional herpetologist without the help
of my parents, wife, professors, and mentors. The journey began at the Smithsonian. With a few
exceptions, it’s been all turtles all the time, and I wouldn’t change that for anything.
Acknowledgements
I thank Kristy Cummings, Whit Gibbons, Robert E. Lovich, and Shellie Puffer for comments and
suggestions on an earlier draft of this biography. Any use of trade, product or firm names is for
descriptive purposes only and does not imply endorsement by the U.S. Government.

8
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13:25.
Lovich, J.E., S.W. Gotte, and C.H. Ernst. 1983. Clutch and egg size in the New Guinea chelid turtle
Emydura subglobosa. Herpetofauna 14:95.
Lovich, J.E. 1984. Capital trout. Virginia Wildlife 45:20 23.
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comments on taxonomic utility. Copeia 1989:304-318.

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Lovich, J.E. and J.W. Gibbons. 1990. Age at maturity influences adult sex ratio in the turtle
Malaclemys terrapin. Oikos 59:126-134.
Lovich, J.E. 1990. Spring movement patterns of two radio-tagged male spotted turtles. Brimleyana
16:67-71.
Ernst, C.H. and J.E. Lovich. 1990. A new species of Cuora (Reptilia: Testudines: Emydidae) from
the Ryukyu Islands. Proceedings of the Biological Society of Washington 103: 26-34.
Lovich, J.E., C.J. McCoy and W.R. Garstka. 1990. The development and significance of melanism
in the slider turtle. Pp. 233-254. In J.W. Gibbons, ed., Life History and Ecology of the Slider
Turtle. Smithsonian Institution Press, Washington, DC.
Lovich, J.E., A.F. Laemmerzahl, C.H. Ernst, and J.F. McBreen. 1991. Relationships among turtles
of the genus Clemmys (Reptilia:Testudines:Emydidae) as suggested by plastron scute
morphology. Zoologica Scripta 20:425–429.
Lovich, J.E., A.D. Tucker, D.E. Kling, J.W. Gibbons, and T.D. Zimmerman. 1991. Behavior of
hatchling diamondback terrapins released in a South Carolina salt marsh. Herpetological
Review 22:81–83.
Garstka, W.R., W.E. Cooper, Jr., K.W. Wasmund, and J.E. Lovich. 1991. Males sex steroids and
hormonal control of male courtship behavior in the yellow-bellied slider turtle, Trachemys
scripta. Comparative Biochemistry and Physiology 98A: 271-280.
Grant, B.W., A.D. Tucker, J.E. Lovich, A.M. Mills, P.M. Dixon, and J.W. Gibbons. 1992. The use
of coverboards in estimating patterns of reptile and amphibian biodiversity. Pp. 379–403. In
D.L. McCullough and R.H. Barrett eds., Wildlife 2001: Populations. Elsevier Applied Science,
London.
Lovich, J.E. and J.W. Gibbons. 1992. A review of techniques for quantifying sexual size
dimorphism. Growth, Development and Aging 56:269-281.

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Lovich, J.E., D.W. Herman, and K.M. Fahey. 1992. Seasonal activity and movements of bog turtles
(Clemmys muhlenbergii) in North Carolina. Copeia 1992:1107–1111.
Ross, D.A. and J.E. Lovich. 1992. Does the color pattern of two species of turtles imitate
duckweed? Journal of the Pennsylvania Academy of Science 66:39–42.
Lovich, J.E. and C.J. McCoy. 1992. Review of the Graptemys pulchra group (Reptilia, Testudines,
Emydidae), with descriptions of two new species. Annals of Carnegie Museum 61:293–315.
Lovich, J.E. 1992. Aspects of the ecology of an isolated brook trout (Salvelinus fontinalis)
population in Fairfax County, Virginia. Journal of the Pennsylvania Academy of Science
65:107–111.
Lovich, J.E. 1993. Macroclemys temminckii. Catalogue of American Amphibians and Reptiles
562.2–562.4.
Lovich, J.E. 1993. Macroclemys. Catalogue of American Amphibians and Reptiles 562.1–562.2.
Lovich, J.E. and C.J. McCoy. 1994. Graptemys gibbonsi. Catalogue of American Amphibians and
Reptiles 586.1–586.2.
Lovich, J.E. 1994. How many species of reptiles and amphibians are found in South Carolina?
Carolina Herpetology 2:1–2.
Lovich, J.E. 1994. A bad time to be a turtle. Riverside County Land Conservancy Field Report
Summer issue. 1 page.
Lovich, J.E., T.B. Egan, and R.C. de Gouvenain. 1994. Tamarisk control on public lands in the
desert of southern California: Two case studies. 46th Annual California Weed Conference,
California Weed Science Society. pp. 166–177.
Lovich, J.E. and C. J. McCoy. 1994. Graptemys ernsti. Catalogue of American Amphibians and
Reptiles 585.1–585.2.
Lovich, J.E. 1994. Biodiversity and zoogeography of non-marine turtles in Southeast Asia. Pp. 380–
391 In S.K. Majumdar, F.J. Brenner, J.E. Lovich, E.W. Miller, and J.F. Schalles eds. Biological
Diversity: Problems and Challenges. Pennsylvania Academy of Science.
Barrows, C., A. Muth, M. Fisher, and J.E. Lovich. 1995. Coachella Valley fringe-toed lizard. Pp.
137–138. In E.T. Laroe, C.E. Puckett, P.D. Doran, and M.J. Mack eds. Our Living Resources:
A report to the Nation on the Distribution, Abundance and Health of U.S. Plants, Animals and
Ecosystems. National Biological Service, Washington, D.C.
Lovich, J.E. 1995. Turtles. Pp. 118–121, In E.T. Laroe, C.E. Puckett, P.D. Doran, and M.J. Mack
eds. Our Living Resources: A report to the Nation on the Distribution, Abundance and Health
of U.S. Plants, Animals and Ecosystems. National Biological Service, Washington, D.C.
Lovich, J.E. and T. Lamb. 1995. Morphometric similarity between the turtles Kinosternon
subrubrum hippocrepis and K. baurii. Journal of Herpetology 29:621–624.
Lovich, J.E. 1995. Book review of "Liste des reptiles actuels de monde. I. Chelonii." by Patrick
David. 1994. Dumerilia (Publication de l'association des Amis du Laboratoire des Reptiles et
Amphibians du Muséum national d'Histoire naturelle de Paris.). Herpetological Review.
26:111–112.
Lovich, J.E. 1995. Turtle, tortoise and terrapin: myriad monikers. People, Land and Water (U.S.
Department of the Interior) 2:26.

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Lovich, J.E. 1995. Wildlife and weeds: Life in an alien landscape. CalEPPC News (California
Exotic Pest Plant Council newsletter) 3:4–5.
Lovich, J.E. 1995. Turtles, tortoises and terrapins: a multitude of monikers. The Tortuga Gazette
(Newsletter of the California Turtle and Tortoise Club) 31:6.
Lovich, J.E. 1995. Ecology of North American tortoises. Pp.50–53. In G. Aguirre, E.D. McCoy, H.
Mushinsky, M. Villagrán Santa Cruz, R. García, and G. C. Andreu eds., Proceedings of North
American Tortoise Conference, Mapimi Biosphere Reserve, Durango, Mexico, October 8–11,
1994. Publications de la Sociedad Herpetologica Mexicana No. 2.
Sloan, K. and J.E. Lovich. 1995. Exploitation of the alligator snapping turtle, Macroclemys
temminckii, in Louisiana: A case study. Chelonian Conservation and Biology 1:221–222.
Ernst, C.H. and J.E. Lovich. 1996. Hydromedusa maximiliani (Maximilian's snake-necked turtle).
Epizoic commensal. Herpetological Review 27:76–77.
Lovich, J.E. 1996. A brief overview of the impact of tamarisk infestation on native plants and
animals. Pp. 13–15. In J. DiTomaso and C.E. Bell eds., Proceedings of the Saltcedar
Management Workshop, Rancho Mirage, California, June 12, 1996.
Lovich, J.E. and R.E. Lovich. 1996. Decline of native brook trout (Salvelinus fontinalis)
populations along the upper West Branch of the Susquehanna River: Canaries outside the coal
mine. Journal of the Pennsylvania Academy of Science 70:55–60.
Lovich, J.E., S.W. Gotte, C.H. Ernst, J. Harshbarger, A.F. Laemmerzahl, and J.W. Gibbons. 1996.
Prevalence and histopathology of shell disease in turtles from Lake Blackshear, Georgia.
Journal of Wildlife Diseases 32:259–265.
Lovich, J.E. 1996. Possible demographic and ecologic consequences of sex ratio manipulation in
turtles. Chelonian Conservation and Biology 2:114–117.
Sloan, K.N., K.A. Buhlmann, and J.E. Lovich. 1996. Stomach contents of commercially harvested
adult alligator snapping turtles, Macroclemys temminckii. Chelonian Conservation and Biology
2:96–99.
Ernst, C. H., J.E. Lovich, A. F. Laemmerzahl, and S. Sekscienski. 1997. A comparison of plastron
scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian
Conservation and Biology 2:603–607.
Lovich, J.E. 1997. Sex, turtles and climate change. U.S. Department of the Interior People, Land
and Water 4 (November):12.
Lovich, J.E. 1997. Additional records of amphibians and reptiles from Indiana County,
Pennsylvania. Journal of the Pennsylvania Academy of Science 71:35–38.
Hinton, T.G., P. Fledderman, J. Lovich, J. Congdon, and J.W. Gibbons. 1997. Radiographic
determination of fecundity: Is the technique safe for developing turtle embryos? Chelonian
Conservation and Biology 2:409–414.
Gibbons, J.W., V.J. Burke, J.E. Lovich, R.D. Semlitsch, T.D. Tuberville, J.R. Bodie, J.L. Greene, P.
H. Niewiarowski, H.H. Whiteman, D.E. Scott, J.H.K. Pechmann, C.R. Harrison, S.H. Bennett,
J.D. Krenz, M.S. Mills, K.A. Buhlmann, J.R. Lee, R.A. Seigel, A.D. Tucker, T.M. Mills, T.
Lamb, M.E. Dorcas, J.D. Congdon, M.H. Smith, D.H. Nelson, M.B. Deitsch, H.H. Hanlin, J.A.
Ott, and D.J. Karapatakis. 1997. Perceptions of species abundance, distribution, and diversity:

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lessons from four decades of sampling on a government-managed reserve. Environmental
Management 21:259–268.
Lovich, J.E. 1997. Wildlife as weeds. Pp. 46–51. In J. Lovich, J. Randall and M. Kelly eds.,
Proceedings California Exotic Pest Plant Council Symposium 96.
Lovich, J.E. and J.W. Gibbons. 1997. Conservation of covert species: protecting species we don't
even know. Pp. 426–429. In, J. Van Abbema et al. eds., Proceedings: Conservation,
Restoration and Management of Turtles and Tortoises — an International Conference, State
University of New York, Purchase.
Lovich, J.E. and R.G. de Gouvenain. 1998. Saltcedar invasion in desert wetlands of the
southwestern United States: Ecological and political implications. Pp. 447–467. In S.K.
Majumdar, E.W. Miller, and F.J. Brenner eds., Ecology of Wetlands and Associated Systems.
Pennsylvania Academy of Science. (Reprinted in, Kelly, M., E. Wagner and P. Warner, eds.,
1999. Proceedings: California Exotic Pest Plant Council Symposium. Vol. 4:45–55).
Lovich, J.E. 1998. More facts about invasive plants by state and territory. California: Brassica
tournefortii. P. 77. In R.G. Westbrooks ed., Invasive plants: changing the landscape of
America. Federal Interagency Committee for the Management of Noxious and Exotic Weeds.
Washington, D.C.
Lovich, J.E., C.H. Ernst, R.T. Zappalorti, and D.W. Herman. 1998. Geographic variation in growth
and sexual size dimorphism of bog turtles (Clemmys muhlenbergii). American Midland
Naturalist 139:69–78.
Lovich, J.E., P. Medica, H. Avery, K. Meyer, G. Bowser, and A. Brown. 1999. Studies of
reproductive output of the desert tortoise at Joshua Tree National Park, the Mojave National
Preserve, and comparative sites. Park Science 19:22–24. (Reprinted in Tortuga Gazette 1999.
35(7):1–2, 4).
Lovich, J.E. and D. Bainbridge. 1999. Anthropogenic degradation of the southern California desert
ecosystem and prospects for natural recovery and restoration. Environmental Management
24:309–326.
Lovich, J.E. 1999. Human-induced changes in the Mojave and Colorado Desert ecosystems:
Recovery and restoration potential. Pp. 529–531, 541–542. In M.J. Mac, P.A. Opler, C.E.
Puckett Haecker and P.D. Doran eds., Status and Trends of the Nation's Biological Resources.
Department of the Interior, U.S. Geological Survey, Reston, Virginia.
Burke, V.J., J.E. Lovich, and J.W. Gibbons. 2000. Conservation of freshwater turtles. Pp. 156–179
In, M. Klemens ed., Turtle Conservation. Smithsonian Institution Press, Washington, D.C.
Lovich, J.E., R.A. Mittermeier, P.C.H. Pritchard, A.G.J. Rhodin, and J.W. Gibbons. 2000.
Powdermill Conference: Trouble for the worlds turtles. Turtle and Tortoise Newsletter 1:16–
17.
Lovich, J.E. 2000. Tamarix ramosissima Lebed, Tamarix chinensis, Tamarix gallica, Tamarix
parviflora. Pp. 312–317. In C.C. Bossard, J.M. Randall, and M.C. Hoshovsky eds., Invasive
Plants of California’s Wildlands. University of California Press, Berkeley.

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Lovich, J.E. 2000. Pennisetum setaceum Forsskal. Pp. 258–262. In C.C. Bossard, J.M. Randall, and
M.C. Hoshovsky eds. Invasive Plants of California’s Wildlands. University of California Press,
Berkeley.
DeLoach, C.J., R.I. Carruthers, J.E. Lovich, T.L. Dudley, and S.D. Smith. 2000. Ecological
interactions in the biological control of saltcedar (Tamarix spp.) in the United States: toward a
new understanding. Pp. 819–873. In N.R. Spencer ed., Proceedings of the X International
Symposium on Biological Control of Weeds. 4–14 July, 1999, Montana State University,
Bozeman, Montana.
Dudley, T.L., C.J. DeLoach, J.E. Lovich, and R.I. Carruthers. 2000. Saltcedar invasion of western
riparian areas: impacts and new prospects for control. Pp. 345–381. In E. McCabe and S.E.
Loos eds., Transactions of the 65th North American Wildlife and Natural Resources
Conference. Wildlife Management Institute, Washington, D.C.
Lovich, J.E., and R. Daniels. 2000. Environmental characteristics of desert tortoise (Gopherus
agassizii) burrow locations in an altered industrial landscape. Chelonian Conservation and
Biology 3:714–721.
Nicolai, N.C., and J.E. Lovich. 2000. Preliminary observations of the behavior of male, flat-tailed
horned lizards before and after an off-highway vehicle race in California. California Fish and
Game 86:208–212.
Gibbons, J.W., J.E. Lovich, A.D. Tucker, N.N. FitzSimmons, and J.L. Greene. 2001. Demographic
and ecological factors affecting conservation and management of the diamondback terrapin
(Malaclemys terrapin) in South Carolina. Chelonian Conservation and Biology 4:66–74.
Lovich, J.E. 2002. Overview and parting shots. Pp. 331–335. In B.Tellman ed., Invasive species in
the Sonoran Region. University of Arizona Press and the Arizona-Sonoran Desert Museum,
Tucson.
Lovich, J., and K. Meyer. 2002. The western pond turtle (Clemmys marmorata) in the Mojave
River, California, USA: highly adapted survivor or tenuous relict? Journal of Zoology, London
256:537–545.
Mittermeier, C.G., W.R. Konstant, R.E. Lovich, and J.E. Lovich. 2002. The Mojave Desert. Pp.351
356. In R. Mittermeier, C.G. Mittermeier, P. Robles Gil, G. Fonseca, T. Brooks, J. Pilgrim, and
W.R. Konstant eds., Wilderness: Earth's Last Wild Places. CEMEX, Mexico.
Lovich, J.E. 2003. Diamondback Terrapin Still Vulnerable to Human Activities. Sound Waves
(monthly newsletter produced by the U.S. Geological Survey, online from January 1999 to
present. http://soundwaves.usgs.gov/). December 2002/January 2003 issue. pp. 7–9.
Lovich, J.E. 2003. Book Review. North American box turtles: A natural history. Herpetological
Review. 33(4):332 333.
Lovich, J.E. 2005. Profiles of selected fish species found in the Grand Canyon ecosystem. Pp. 50–
53. In S.P. Gloss, J.E. Lovich, and T.S. Melis eds. The state of the Colorado River ecosystem in
the Grand Canyon. U.S. Geological Survey Circular 1282. 220 pages.
http://pubs.usgs.gov/circ/1282/

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Lovich, J.E., and T.S. Melis. 2005. Lessons from 10 years of adaptive management in Grand
Canyon. Pp. 207–220. In S.P. Gloss, J.E. Lovich, and T.S. Melis eds., The state of the
Colorado River ecosystem in the Grand Canyon. U.S. Geological Survey Circular 1282. 220
pages.
Lovich, J.E. 2005. Book Review: Introduced reptiles and amphibians of the world: unwanted exotic
species. Diversity and Distributions 11:591–593.
Lovich, J.E., and T.S. Melis. 2007. The state of the Colorado River ecosystem in Grand Canyon:
lessons from 10 years of adaptive ecosystem management. International Journal of River Basin
Management 5:207–221.
Lovich, J.E., and K.R. Beaman. 2007. A history of Gila Monster (Heloderma suspectum cinctum)
records from California with comments on factors affecting their distribution. Bulletin of the
Southern California Academy of Sciences 106:39–58.
Lovich, J.E., and K.R. Beaman. 2008. Distribution of native turtles in the arid southwestern United
States with comments on Kinosternon sonoriense: a species presumed to be lost from
California’s herpetofauna. Pp. 127–134. In R.E.Reynolds ed., The 2008 Desert Symposium and
Field Guide and Proceedings. California State University, Desert Studies Consortium and LSA
Associates, Inc..
Ernst, C.H., A.E. Laemmerzahl, and J.E. Lovich. 2008. A morphological review of the Cuora
flavomarginata complex (Testudines: Geoemydidae). Proceedings of the Biological Society of
Washington 121:391–397.
Lovich, J.E. 2009. Comments on the proposed conservation of usage of Testudo gigantea.
Schweigger,1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case
3463; see BZN 66: 34–50, 80–87, 169–186). Bulletin of Zoological Nomenclature 66(3):276.
Lovich, J.E., W. Selman, and C.J. McCoy. 2009. Graptemys gibbonsi Lovich and McCoy 1992 –
Pascagoula map turtle, Pearl River map turtle, Gibbon’s map turtle. Pp. 029.1–029.8. In A.G.J.
Rhodin, et al eds., Conservation biology of freshwater turtles and tortoises: a compilation
project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research
Monographs No. 5.
Glenn, E.P, P.L. Nagler, and J.E. Lovich. 2009. The surprising value of saltcedar. Southwest
Hydrology (May/June issue): 10–11.
Lovich, J.E., C. Drost, A.J. Monatesti, D. Casper, D. Wood, and M. Girard. 2010. New reptilian
prey items for the Sonora Mud Turtle (Kinosternon sonoriense) with a brief review of
saurophagy and ophiophagy in North American turtles. Southwestern Naturalist 55:135–138.
Ennen, J.R., J.E. Lovich, B.R. Kreiser, W. Selman, and C.P. Qualls. 2010. Genetic and
morphological variation between populations of the Pascagoula map turtle (Graptemys
gibbonsi) in the Pearl and Pascagoula Rivers with description of a new species. Chelonian
Conservation and Biology 9:98–113.

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Lovich, J.E., M. Znari, M.A. Ait Baamrane, M. Naimi, A. Mostalih. 2010. Biphasic geographic
variation in sexual size dimorphism of turtle (Mauremys leprosa) populations along an
environmental gradient in Morocco. Chelonian Conservation and Biology 9:45–53.
Ennen, J.R., B.R. Kreiser, C.P. Qualls, and J.E. Lovich. 2010. A morphological and molecular
reassessment of Graptemys oculifera and G. flavimaculata. Journal of Herpetology 44:544–
554.
Lovich, J.E. 2010. A preliminary review of the effects of utility-scale renewable energy
development on terrestrial wildlife in the desert southwest with emphasis on the desert tortoise.
Pp. 286–288. In R.E. Reynolds ed., Abstracts and Proceedings of the 2010 Desert Symposium
(Overboard in the Mojave: 20 million years of lakes and wetlands). California State University
Desert Studies Consortium.
Lovich, J.E., Y. Yasukawa, and H. Ota. 2011. Mauremys reevesii (Gray 1831) – Reeves’ Turtle,
Chinese Three-keeled Pond Turtle. Pp. 050.1–050.10. In A.G.J. Rhodin, et al eds.,
Conservation biology of freshwater turtles and tortoises: a compilation project of the
IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs
No. 5.
Lovich, J.E., J.C. Godwin, and C.J. McCoy. 2011. Graptemys ernsti Lovich and McCoy 1992 –
Escambia Map Turtle. Pp. 051.1–051.6. In A.G.J. Rhodin, et al eds., Conservation biology of
freshwater turtles and tortoises: a compilation project of the IUCN/SSC Tortoise and
Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5.
Lovich, J.E., J.R. Ennen, S.V. Madrak, and B. Grover. 2011. Turtles, culverts and alternative energy
development: an unreported but potentially significant mortality threat to the desert tortoise
(Gopherus agassizii). Chelonian Conservation and Biology 10:124–129.
Ennen, J.R., R.D. Birkhead, B.R. Kreiser, D.L. Gaillard, C.P. Qualls, and J.E. Lovich. 2011. The
effects of isolation on the demography and genetic diversity of long-lived species: implications
for conservation and management of the gopher tortoise (Gopherus polyphemus).
Herpetological Conservation and Biology 6:202–214.
Lovich, J.E., J.R. Ennen, S. Madrak, K. Meyer, C. Loughran, C. Bjurlin, T.R. Arundel, W. Turner,
C. Jones, and G.M. Groenendaal. 2011. Effects of wind energy production on growth,
demography and survivorship of a desert tortoise (Gopherus agassizii) population in southern
California with comparisons to natural populations. Herpetological Conservation and Biology
6:161–174. (http://www.herpconbio.org/Volume_6/Issue_2/Lovich_etal_2011.pdf).
Ennen, J.R., C.L. Loughran, and J.E. Lovich. 2011. Gopherus agassizii (Desert Tortoise). Non-
native seed dispersal. Herpetological Review (Natural History Notes) 42:266–267.
Lovich, J.E., J.R. Ennen, S. Madrak, C. Loughran, K. Meyer, T.V. Arundel, and C. Bjurlin. 2011.
Long-term post fire effects on spatial ecology and reproductive output of female desert
tortoises at a wind energy facility near Palm Springs, California. Fire Ecology 7:75–87.
Lovich, J.E., and G.B. Haxel. 2011. A previously unreported locality record for the Gila Monster.
Bulletin of the Southern California Academy of Sciences 110:59–62.
Ernst, C.H., A.F. Laemmerzahl, and J.E. Lovich. 2011. Does the "kamaroma" — plastron pattern
morph occur in both Philippine subspecies of the turtle Cuora amboinensis? Proceedings of the
Biological Society of Washington 124:259–269.

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Loughran, C.L., J.E. Ennen, and J.E. Lovich. 2011. Gopherus agassizii (Desert tortoise). Burrow
collapse. Herpetological Review 42:593.
Lovich, J.E. 2011. Gopherus agassizii (Desert Tortoise) and Crotalus ruber (Red Diamond
Rattlesnake). Burrow co-occupancy Herpetological Review. 42:421.
Lovich, J.E., and J.R. Ennen. 2011. Wildlife conservation and solar energy development in the
Desert Southwest, United States. BioScience 61:982–992.
Lovich, J.E. 2012. What do we know about the effects of climate change, especially global
warming, on desert tortoises. Pp. 105–107. R.E. Reynolds ed., The 2012 Desert Symposium
Field Guide and Proceedings (Search for the Pliocene: the southern exposure). California State
University Desert Studies Consortium.
Naimi, M., M. Znari, J.E. Lovich, Y. Feddadi, M.A. Ait Baamrane. 2012. Clutch and egg allometry
of the turtle Mauremys leprosa (Chelonia: Geoemydidae) from a polluted peri-urban river in
west-central Morocco. Herpetological Journal. 22:43–49.
Ennen, J.R., J.E. Lovich, and R. Birkhead. 2012. Graptemys pearlensis Ennen, Lovich, Kreiser,
Selman, and Qualls. Pearl River Map Turtle. Catalogue of American Amphibians and Reptiles
889.1–889.4.
Ennen, J.R., K.P. Meyer, and J.E. Lovich. 2012. Female Agassiz’s Desert Tortoise activity at a
wind energy facility in southern California: the influence of an El Niño event. Natural Science
4:30–37. doi:10.4236/ns.2012.41006.
Ennen, J.R., J.E. Lovich, K. Meyer, C. Bjurlin, and T.R. Arundel. 2012. Nesting ecology of a desert
tortoise (Gopherus agassizii) population at a utility-scale renewable energy facility in southern
California. Copeia. 2012:222–228.
Lovich, J.E., S.V. Madrak, C. Drost, A.J. Monatesti, D. Casper, and M. Znari. 2012. Optimal egg
size in a suboptimal environment: reproductive ecology of female Sonora Mud Turtles
(Kinosternon sonoriense) in central Arizona, USA. Amphibia-Reptilia 33:161–170.
Lovich, J.E., N.J. Scott, R.B. Bury, K.C. Dodd, Jr., and R.W. McDiarmid. 2012. A history of
herpetologists and herpetology in the U.S. Department of the Interior. Herpetological
Conservation and Biology. 7(Monograph 2):1–45.
Lovich, J.E., M. Agha, M. Meulblok, K. Meyer, J. Ennen, C. Loughran, S.V. Madrak, and C.
Bjurlin. 2012. Climatic variation affects clutch phenology in Agassiz's Desert Tortoise
(Gopherus agassizii). Endangered Species Research 19:63–74.
Lovich, J.E. 2013. Book review: Turtles, Tortoises and Terrapins: A Natural History. R. Orenstein.
2012. Firefly Books. Copeia. 2013:570–571.
Lovich, J.E., and J.R. Ennen. 2013. Assessing the state of knowledge of utility-scale wind energy
development and operation on non-volant terrestrial and marine wildlife. Applied Energy
103:52–60.
Lovich, J.E., and J.R. Ennen. 2013. A quantitative analysis of the state of knowledge of turtles of
the United States and Canada. Amphibia-Reptilia 34:11–23.
Agha, M., J.E. Lovich, J.R. Ennen, and E. Wilcox. 2013. Nest-guarding by female Agassiz's Desert
Tortoise (Gopherus agassizii) at a wind-energy facility near Palm Springs, California. The
Southwestern Naturalist 58:254–257.

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Underwood, E.B., S. Bowers, J.C. Guzy, J.E. Lovich, C.A. Taylor, J.W. Gibbons, and M.E. Dorcas.
2013. Sexual dimorphism and feeding ecology of diamondback terrapins (Malaclemys
terrapin). Herpetologica 69:397–404.
Lovich, J.E. 2014. Managing invasive Red-eared Sliders in Japan. Kiraku (Freshwater Turtle Data
from Japan), published by the Kobe-Suma Aquarium (7):1–2. IN ENGLISH AND JAPANESE
Beaman, K.R., Jones, J.L. and J.E. Lovich. 2014. Heloderma suspectum cinctum (Banded Gila
Monster). USA: NEVADA: Clark Co.: Lucy Gray Mountains. Herpetological Review 45:462.
Lovich, J.E., C.B. Yackulic, J. Freilich, M. Agha, M. Meulblok, K.P. Meyer, T.R. Arundel, J.
Hansen, M.S. Vamstad, and S.A. Root. 2014. Climatic variation and tortoise survival: has a
desert species met its match? Biological Conservation 169:214–224.
Ennen, J.R., M.E. Kalis, A.L. Patterson, B.R. Kreiser, J.E. Lovich, J. Godwin, and C.P. Qualls.
2014. Clinal variation or validation of a subspecies: a case study with the Graptemys nigrinoda
complex (Testudines: Emydidae). Biological Journal of the Linnean Society 111:810–822.
Lovich, J.E., J.C. Godwin, and C.J. McCoy. 2014. Graptemys pulchra Baur 1893 – Alabama Map
Turtle. Pp. 072.1–072.6. In A.G.J. Rhodin, et al eds. Conservation biology of freshwater turtles
and tortoises: a compilation project of the IUCN/SSC Tortoise and Freshwater Turtle
Specialist Group. Chelonian Research Monographs No. 5.
Lovich, J.E., J.W. Gibbons, and M. Agha. 2014. Does the timing of attainment of maturity influence
sexual size dimorphism and adult sex ratio in turtles? Biological Journal of the Linnean Society
112:142–149.
Lovich, J.E., D. Delaney, J. Briggs, M. Agha, M. Austin, and J. Reese. 2014. Black bears (Ursus
americanus) as a novel potential predator of Agassiz’s desert tortoises (Gopherus agassizii) at a
California wind energy facility. Bulletin of the Southern California Academy of Sciences
113:34–41.
Lovich, J.E., C.T. LaRue, C.A. Drost, and T.R. Arundel. 2014. Traditional cultural use as a tool for
inferring biogeography and provenance: a case study involving painted turtles (Chrysemys
picta) and Hopi Native American Culture in Arizona, USA. Copeia. 2014:215–220.
Lovich, J.E. and J.R. Ennen. 2014. Graptemys gibbonsi Lovich and McCoy Pascagoula Map Turtle.
Catalogue of American Amphibians and Reptiles 901.1–901.8.
Lovich, J.E., C.H. Ernst, E.M. Ernst and J.L. Riley. 2014. A 21-year study of seasonal and
interspecific variation of hatchling emergence in a Nearctic freshwater turtle community: to
overwinter or not to overwinter. Herpetological Monographs 28:93–109.
Godwin J., J.E. Lovich, J.R. Ennen, B.R. Kreiser, B.P. Folt, and C. Lechowicz. 2014. Hybridization
of two megacephalic map turtles (Testudines: Emydidae: Graptemys) in the Choctawhatchee
River drainage of Alabama and Florida. Copeia 2014:725–742.
Lovich, J.E., M. Agha, C.B. Yackulic, K. Meyer, C. Bjurlin, J.R. Ennen, T.R. Arundel, and M.
Austin. 2014. Nest site characteristics, nesting movements, and lack of long-term nest site
fidelity in Agassiz’s desert tortoises (Gopherus agassizii) at a wind energy facility in southern
California. California Fish and Game 100:404–416.

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Agha, M., B. Augustine, J.E. Lovich, D. Delaney, B. Sinervo, M.O. Murphy, J.R. Ennen, J.R.
Briggs, R. Cooper, and S.J. Price. 2015. Using motion-sensor camera technology to infer
seasonal activity and thermal niche of the desert tortoise (Gopherus agassizii). Journal of
Thermal Biology 49–50:119–126.
Beaman, K.R., B.J. Scurlock, J.E. Lovich, and L.A. Kobelt. 2015. Heloderma suspectum cinctum
(Banded Gila Monster). USA: California: San Bernardino Co.: Mesquite Mountains.
Herpetological Review. 46:384.
Lovich, J.E. 2015. Golden eagle mortality at a wind-energy facility near Palm Springs, California.
Western Birds 46:76–80.
Lovich, J.E., J.R. Ennen, C.B. Yackulic, K. Meyer-Wilkins, M. Agha, C. Loughran, C. Bjurlin, M.
Austin, and S. Madrak. 2015. Not putting all their eggs in one basket: bet-hedging despite
extraordinary annual reproductive output of desert tortoises. Biological Journal of the Linnean
Society 115:399–410.
Agha, M.A, M.O. Murphy, J.E. Lovich, J.R. Ennen, C.R. Oldham, K. Meyer, C. Bjurlin, M. Austin,
S. Madrak, C. Loughran, L. Tennant, and S.J. Price. 2015. The effect of research activities and
winter precipitation on defensive voiding behavior of Agassiz’s Desert Tortoises (Gopherus
agassizii). Wildlife Research 41(8): 641–649.
Agha, M., J.E. Lovich, J.R. Ennen, B. Augustine, T.R. Arundel, M.O. Murphy, K. Meyer-Wilkins,
C. Bjurlin, D. Delaney, J. Briggs, M. Austin, S. Madrak and S.J. Price. 2015. Turbines and
terrestrial vertebrates: variation in tortoise survivorship between a wind energy facility and an
adjacent undisturbed wildland area in the desert southwest (USA). Environmental Management
56:332–341.
Lovich, J. 2015. Case study: road proliferation due to rapid renewable energy development. Pp. 79–
80. In K.A. Andrews, P. Nanjappa, and S.P.D. Riley eds. Roads and ecological infrastructure:
concepts and applications for small animals. Johns Hopkins University Press, Baltimore,
Maryland.
Agha, M., D. Delaney, J.E. Lovich, J. Briggs, M. Austin and S.J. Price. 2015. Nelson’s big horn
sheep (Ovis canadensis nelsoni) trample Agassiz’s desert tortoise (Gopherus agassizii) burrow
at a California wind energy facility. Bulletin of the Southern California Academy of Sciences
114:58–62.
Ennen, J.R., P.V. Lindeman, and J.E. Lovich. 2015. Intersexual allometry differences and
ontogenetic shifts of coloration patterns in two aquatic turtles, Graptemys oculifera and
Graptemys flavimaculata. Ecology and Evolution 5(11):2296–2305.
Ernst, C.H. and J.E. Lovich. 2015. Interanal seam loss in Asian turtles of the Cuora flavomarginata
complex (Testudines, Geoemydidae). Herpetological Bulletin 132:1–4.
Rhodin, A.G.J., H. Kaiser, P.P. van Dijk, W. Wüster, M. O’Shea, M. Archer, M. Auliya, L. Boitani,
R. Bour, V. Clausnitzer, T. Contreras-MacBeath, B.I. Crother, J.M. Daza, C.A. Driscoll, O.
Flores-Villela, J. Frazier, U. Fritz, A. Gardner, C. Gascon, A. Georges, F. Glaw, F.G.
Grazziotin, C.P. Groves, G. Haszprunar, P. Havaš, J.M. Hero, M. Hoffmann, M.S. Hoogmoed,
B.D. Horne, J.B. Iverson, M. Jäch, C.L. Jenkins, R.K.B. Jenkins, A.R. Kiester, J.S. Keogh, T.E.
Lacher Jr., J.E. Lovich, L. Luiselli, D.L. Mahler, D. Mallon, R. Mast, R.W. Mcdiarmid, J.
Measey, R.A. Mittermeier, S. Molur, V. Mossbrugger, R. Murphy, D. Naish, M. Niekisch, H.
Ota, J.F. Parham, M.J. Parr, N.J. Pilcher, R.H. Pine, A.B. Rylands, J.G. Sanderson, J. Savage,

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W. Schleip, G.J. Scrocchi, H.B. Shaffer, E.N. Smith, R. Sprackland, S.N. Stuart, H. Vetter, L.J.
Vitt, T. Waller, G. Webb, E.O. Wilson, H. Zaher, and S. Thomson (Corresponding Author).
2015. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for
confirmation of the availability of the generic name and for the nomenclatural validation of the
journal in which it was published. (Case 3601; see BZN 70: 234–237; 71: 30–38, 133–135,
181–182, 252–253). Bulletin of Zoological Nomenclature 72(1):65–78.
Smith, A.L., L.A. Tennant, J.E. Lovich, and T.R. Arundel. 2015. Gopherus agassizii (Agassiz’s
Desert Tortoise). Mechanical injury. Herpetological Review 46:423–424.
Zappalorti, R.T., J.E. Lovich, R.F. Farrell, and M.E. Torocco. 2015. Nest-site characteristics of bog
turtles (Glyptemys muhlenbergii) in New Jersey and Pennsylvania. Northeastern Naturalist
22:573–584.
Kern, M.M., J.C. Guzy, J.E. Lovich, J.W. Gibbons, and M.E. Dorcas. 2016. Relationships of
maternal body size and morphology with egg and clutch size in the diamondback terrapin,
Malaclemys terrapin (Testudines: Emydidae). Biological Journal of the Linnean Society
117:295–304.
Henderson, R.A., S.R. Puffer, J.E. Lovich. 2016. Gopherus agassizii (Mohave Desert Tortoise).
Nest depredation. Herpetological Review. 47:446–447.
Ennen, J.R., J.E. Lovich, and R.L. Jones. 2016. Graptemys pearlensis Ennen, Lovich, Kreiser,
Selman, and Qualls 2010 – Pearl River Map Turtle. In A.G.J. Rhodin, et al eds., Conservation
biology of freshwater turtles and tortoises: a compilation project of the IUCN/SSC Tortoise
and Freshwater Turtle Specialist Group. Chelonian Research Monographs No. 5.5(9):094.1–8.
Ennen, J.R., M. Agha, W. Matamoros, S. Hazzard, and J.E. Lovich. 2016. Using climate, energy,
and spatial-based hypotheses to interpret macroecological patterns of North America
chelonians. Canadian Journal of Zoology 94:453–461.
Ennen, J.R., J. Godwin, J.E. Lovich, B.R. Kreiser, B. Folt, and S. Hazard. 2016. Interdrainage
morphological and genetic differentiation in the Escambia Map Turtle, Graptemys ernsti.
Herpetological Conservation and Biology 11:122–131.
Ernst, C.H., A. Laemmerzahl, and J.E. Lovich. 2016. A morphological review of subspecies of the
Asian box turtle, Cuora amboinensis (Testudines, Geomydidae). Proceedings of the Biological
Society of Washington 129:144–156.
Lovich, J.E. 2016. Desert scrublands. Pp. 50–54. In L.L.C Jones, K.J. Halama, , and R.E Lovich
eds., Habitat management guidelines for amphibians and reptiles in the southwestern United
States. Birmingham, Alabama, Partners in Amphibian and Reptile Conservation. Technical
Publication HMG-5.
Lovich, J.E. and J.R. Ennen. 2016. Energy development. In. pp. 31–34 In L.L.C Jones, K.J. Halama,
and R.E Lovich eds., Habitat management guidelines for amphibians and reptiles in the
southwestern United States. Birmingham, Alabama, Partners in Amphibian and Reptile
Conservation. Technical Publication HMG-5.
Lovich, J.E., M. Agha, C.W. Painter, L. Cole, A. Fitzgerald, K. Narum, and R. Jennings. 2016.
Aspects of the reproductive ecology of female turtles in New Mexico. Western North American
Naturalist 76.

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Smith, A.L., S.R. Puffer, J.E. Lovich, L.A. Tennant, T.R. Arundel, M.S. Vamstad, and K.D.
Brundige. 2016. A potential predator-prey interaction of an American badger and an Agassiz’s
desert tortoise with a review of badger predation on turtles. California Fish and Game.
102(3):131–144.
Lovich, J.E. and K. Yamamoto. 2016. Measuring the impact of invasive species on popular culture:
a case study based on toy turtles from Japan. Humans and Nature (Museum of Nature and
Human Activities, Hyogo, Japan) 27: 1–11.
Taniguchi, M., J.E. Lovich, K. Mine, S. Ueno, and N. Kamezaki. 2017. Unusual population
attributes of invasive Red-eared Slider Turtles (Trachemys scripta elegans) in Japan: do they
have a performance advantage? Aquatic Invasions 12:97–108.
Gibbons, J.W., J. Lovich and R.M. Bowden. 2017. Turtles: Freshwater, In Reference Module in
Life Sciences, Elsevier, ISBN: 978-0-12-809633-8,
Ennen, J.R., J.E. Lovich, R.C. Averill-Murray, C. Yackulic, M. Agha, C. Loughran, L. Tennant, and
B. Sinervo. 2017. The evolution of different maternal investment strategies in two closely
related desert vertebrates. Ecology and Evolution 1–13.
Lovich, J.E., M. Quillman, B. Zitt, A. Schroeder, D. Earl Green, C. Yackulic, P. Gibbons, and E.
Goode. 2017. The effects of drought and fire in the extirpation of an abundant semi-aquatic
turtle from a lacustrine environment in the southwestern USA. Knowledge and Management of
Aquatic Ecosystems 418.
Agha, M., A.L. Smith, J.E. Lovich, D. Delaney, J.R. Ennen, J. Briggs, L.A. Tennant, S.R. Puffer, A.
Walde, T.R. Arundel, S.J. Price, B.D. Todd. 2017. Mammalian mesocarnivore visitation at
tortoise burrows in a wind farm. Journal of Wildlife Management DOI: 10.1002/jwmg.21262.
Lovich, J.E., and J.R. Ennen. 2017. Reptiles and amphibians. Pp. 97–118. In M. Perrow ed.,
Wildlife and Windfarms: Conflicts and Solutions. Vol. 1. Onshore. Pelagic Press. Exeter, U.K.
Moore-O’Leary, K.A., R.R. Hernandez, D.S. Johnston, S.R. Abella, K.E. Tanner, A.C. Swanson, J.
Kreitler, and J.E. Lovich. 2017. Sustainability of utility-scale solar energy – critical ecological
concepts. Frontiers in Ecology and the Environment.
Ennen, J.R., W.A. Matamoros, M. Agha, J.E. Lovich, S.C. Sweat, and C.W. Hoagstrom. 2017.
Hierarchical, quantitative biogeographic provinces for all North American turtles and their
contribution to the biogeography of turtles and the continent. Herpetological Monographs
31:114–140.
Lovich, J.E., R.C. Averill-Murray, M. Agha, J.R. Ennen, and M. Austin. 2017. Variation in annual
clutch phenology of Sonoran desert tortoises (Gopherus morafkai) in central Arizona.
Herpetologica 73:313–322.
Agha, M., J.R. Ennen, A.J. Nowakowski, J.E. Lovich, S.C. Sweat, B.D. Todd. 2017.
Macroecological patterns of sexual size dimorphism in turtles of the world. Journal of
Evolutionary Biology 31:336-345.
Zappalorti, R.T., A.M. Tutterow, S.E. Pittman, and J.E. Lovich. 2017. Hatching success and
predation of bog turtle (Glyptemys muhlenbergii) eggs in New Jersey and Pennsylvania.
Chelonian Conservation and Biology 16:194–202.

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Lovich, J.E., S.R. Puffer, M. Agha, J.R. Ennen, K. Meyer-Wilkins, L.A. Tennant, A.L. Smith, T.R.
Arundel, K.D. Brundige and M.S. Vamstad. 2018. Reproductive output and clutch phenology
of female Agassiz's Desert Tortoises (Gopherus agassizii) in the Sonoran Desert region of
Joshua Tree National Park. Current Herpetology 37(1):40–57.
Lovich, J.E., J.W. Gibbons, and K. Green. 2018. Life history of the diamond-backed terrapin with
emphasis on geographic variation. Pp. ?-?. In W.M.Roosenburg and V.S. Kennedy eds.,
Ecology and conservation of the diamondback terrapin Malaclemys terrapin. Johns Hopkins
University Press, Baltimore, Maryland.
Lovich, J.E., and K.M. Hart. 2018. Diamond-backed terrapin taxonomy: a history of controversy
and uncertainty. Pp. ?-?. In W.M.Roosenburg and V.S. Kennedy eds,. Ecology and
conservation of the diamondback terrapin Malaclemys terrapin. Johns Hopkins University
Press, Baltimore, Maryland.
Books
Ernst, C.H., J.E. Lovich and R.W. Barbour. 1994. Turtles of the United States and Canada.
Smithsonian Institution Press, Washington, D.C. 578 pp. (682 page paperback released in
2000).
Majumdar, S.K., F. Brenner, J.E. Lovich, and J. Schalles, eds. 1994. Biological Diversity: Problems
and Challenges. Pennsylvania Academy of Science. Easton, Pennsylvania. 461 pp.
Gloss, S.P., J.E. Lovich, and T.S. Melis eds. 2005. The state of the Colorado River ecosystem in the
Grand Canyon. U.S. Geological Survey Circular 1282. 220 pages.
Ernst, C.H. and J.E. Lovich. 2009. Turtles of the United States and Canada. Second edition. Johns
Hopkins University Press. 827 pages.
Proceedings edited
J.E. Lovich, J. Randall, and M. Kelly eds. 1996. Proceedings California Exotic Pest Plant Council
Symposium '95. California Exotic Pest Plant Council, 64 pp.
J.E. Lovich, J. Randall, and M. Kelly eds. 1997. Proceedings California Exotic Pest Plant Council
Symposium '96. California Exotic Pest Plant Council, 110 pp.
Popular articles
Lovich, J.E. 1995. Turtle, tortoise and terrapin: myriad monikers. People, Land and Water (U. S.
Department of the Interior news magazine). April issue, page 26.
Lovich, J.E. 1997. Sex, turtles and climate change. People, Land and Water (U. S. Department of
the Interior news magazine). November issue, page 12.
Lovich, J.E. and K. Meyer. 2001. Pleistocene park: ice age relict turtle survives in California desert.
People, Land and Water (U. S. Department of the Interior news magazine). November issue,
page 18.
Lovich, J.E. 2001. Looking beyond the shell: a brief history of turtles. People, Land and Water (U.
S. Department of the Interior news magazine). September/October issue, page 12.

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Lovich, J.E. 2002. Terrapins and Hurricanes. People, Land and Water (U. S. Department of the
Interior news magazine). November issue, page 17.
du Bernard, M. R. with contributions from J. Lovich. 2003. Painted turtle: Meet the colorful
inhabitants of Turtle Island. Water Gardening Magazine. July/August 48–52.
Lovich, J.E. and N. Scott. 2004. Herpetology in the USGS. People, Land and Water (U. S.
Department of the Interior news magazine). March issue, 10(4):35.
Lovich, J.E. 2008. From animals to plants: creosote bush may be oldest living organism on Earth.
Daily Sun (newspaper, Flagstaff, Arizona). Monday, February 18, page A8.
Technical reports
Lovich, J.E. 1984. The status of the spotted turtle at Cedar Bog, 32 pp. Ohio Historical Society.
Lovich, J.E. 1985. The spotted turtles of Cedar Bog: A population in decline, 49 pp. Ohio
Department of Natural Resources, Division of Natural Areas and Preserves.
Lovich, J.E. 1992. Restoration and revegetation of degraded habitat as a management tool in
recovery of the threatened desert tortoise. Contract Report prepared for California Dept. Parks
and Recreation, Off-Highway Motor Vehicle Recreation Div. U.S. Dept. Interior, Bureau of
Land Management, California Desert District. 187 pp.
Drost, C. A., J.E. Lovich, S. V. Madrak, and A. J. Monatesti. 2011. Removal of non-native slider
turtles (Trachemys scripta) and effects on native Sonora mud turtles (Kinosternon sonoriense)
at Montezuma Well, Yavapai County, Arizona: U.S. Geological Survey Open-File Report
2010–1177, vi + 48 p.
Lovich, J.E. (published 2014). 2013. Assessing the long-term survival and reproductive output of
desert tortoises at a wind energy facility near Palm Springs, California - Final Project Report.
California Energy Commission. Publication number: CEC-500-2014-005. 60 pp.
Galapagos National Park Service and Galapagos Conservancy. 2013. Giant tortoise restoration
through integrated research and management: beyond rescue to full recovery. Final Report of
the International Workshop held in Galapagos July 9–13, 2012. Workshop participants
(alphabetical order): Aguilar, K., Galapagos National Park; S. Blake, Max Planck Institute; R.
Burke, Hofstra University; A. Caccone, Yale University, K. Campbell, Island Conservation; V.
Carrión, Island Conservation; O. Carvajal, Galapagos National Park; L. Cayot, Galapagos
Conservancy; Y. Chiari, Research Center in Biodiversity and Genetic Resources; D. Edwards,
Yale University; J. Flanagan, Houston Zoo; F. Franco, Galapagos National Park; J. Gibbs, State
University of New York College of Environmental Science and Forestry; O. Hamann,
University of Copenhagen; E. Hunter, State University of New York College of Environmental
Science and Forestry; P. Jaramillo, Charles Darwin Foundation; A. Llerena, Charles Darwin

23
Foundation; F. Llerena, Galapagos National Park; J. Lovich, US Geological Survey; J. Málaga,
Galapagos National Park; C. Márquez, Galapagos, Ecuador; C. Ortega, Galapagos National
Park; M. Russello, University of British Columbia; O. Ryder, San Diego Zoo; W. Tapia,
Galapagos National Park; P.P. van Dijk, IUCN TFTSG and Conservation International; F.
Villalva, Galapagos National Park. 12 pages.
Lovich, J.E. and S.R. Puffer. 2015. Establishing a long-term desert tortoise (Gopherus agassizii)
monitoring program for the Coachella Valley Conservation Commission using a population of
tortoises in Joshua Tree National Park. Annual report to the U.S. Fish and Wildlife Service
under Permit No. TE-198910-3. 3 pp. + Excel spreadsheet of transmitter records.
Lovich J.E. and S.R. Puffer. 2015. Establishing a long-term desert tortoise (Gopherus agassizii)
monitoring program for the Coachella Valley Conservation Commission using a population of
tortoises in Joshua Tree National Park. Investigator’s annual report to the National Park
Service. 2 pp.
Lovich, J.E. and S.R. Puffer. 2016. Surveys for the species of special concern, the southwestern
pond turtle (Actinemys pallida), in the Mojave Desert of California. Annual report to the
Bureau of Land Management. 5 pp.
Lovich, J.E. and S.R. Puffer. 2016. Surveys for the species of special concern, the southwestern
pond turtle (Actinemys pallida), in the Mojave Desert of California. Annual report for
California Department of Fish and Game Owned Lands, Camp Cady Wildlife Management
Area. 5 pp.
Lovich, J.E. and S.R. Puffer. 2016. Surveys for the species of special concern, the southwestern
pond turtle (Actinemys pallida), in the Mojave Desert of California. Annual report to California
Department of Fish and Wildlife under SCP-1639. 5 pp.
Lovich, J.E. and S.R. Puffer. 2016. Surveys for southwestern pond turtles (Actinemys pallida) to
minimize the impact of prescribed fire and others fuels treatment on the species at Afton
Canyon with comparative studies on population persistence at Camp Cady Wildlife
Management Area. SCP-1639. Interim survey report to California Department of Fish and
Wildlife, July 2016. 1 page.
Lovich, J.E. and S.R. Puffer. 2016. Surveys for southwestern pond turtles (Actinemys pallida) to
minimize the impact of prescribed fire and other fuels treatment on the species: Final report to
the Bureau of Land Management under IGO L15PG00166. 4 pp.
Lovich, J.E. and S.R. Puffer. 2016. Continuing research supporting a long-term desert tortoise
(Gopherus agassizii) monitoring program for the Coachella Valley Conservation Commission
using a population of tortoises in Joshua Tree National Park. Annual report to the Bureau of
Land Management. 5 pp.
Lovich, J.E. and S.R. Puffer. 2016. Continuing research supporting a long-term desert tortoise
(Gopherus agassizii) monitoring program for the Coachella Valley Conservation Commission
using a population of tortoises in Joshua Tree National Park. Investigator’s annual report to the
National Park Service. 5 pp.
Lovich, J.E. and S.R. Puffer. 2016. Continuing research supporting a long-term desert tortoise
(Gopherus agassizii) monitoring program for the Coachella Valley Conservation Commission

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using a population of tortoises in Joshua Tree National Park. Annual report to the U.S. Fish and
Wildlife Service under Permit No. TE-198910-4. 5 pp. + Excel spreadsheet of transmitter
records.
Lovich, J.E. and S.R. Puffer. 2016. Continuing research on a population of desert tortoises
(Gopherus agassizii) in Joshua Tree National Park to support the establishment of a long-term
desert tortoise monitoring program for the Coachella Valley Conservation Commission. CESA
MOU: Desert Tortoise. Annual report to the California Department of Fish and Wildlife. 6 pp.
Puffer, S.R. and J.E. Lovich. 2017. Quarterly report to Coachella Valley Conservation Commission
on FY2018 task completions for Gopherus agassizii (October–December). 1 page.
Puffer, S.R., J.E. Lovich, K.L. Cummings. 2017. Research supporting a desert tortoise (Gopherus
agassizii) monitoring program and population genetics studies within the Coachella Valley
Multiple Species Habitat Conservation Plan area. Annual report to the Bureau of Land
Management. 8 pp.
Puffer, S.R., J.E. Lovich, K.L. Cummings. 2017. Research supporting a desert tortoise (Gopherus
agassizii) monitoring program and population genetics studies within the Coachella Valley
Multiple Species Habitat Conservation Plan area. Annual report to the California Department
of Fish and Wildlife for the CESA MOU – Desert Tortoise. 8 pp.
Puffer, S.R., J.E. Lovich, K.L. Cummings. 2017. Research supporting a desert tortoise (Gopherus
agassizii) monitoring program and population genetics studies within the Coachella Valley
Multiple Species Habitat Conservation Plan area. Annual report to the U.S. Fish and Wildlife
Service under Permit No. TE-198910-5. 8 pp + Excel spreadsheet of transmitter records.
Puffer, S.R., J.E. Lovich, K.L. Cummings, S. Greely. 2017. Surveys for the species of special
concern, the southwestern pond turtle (Actinemys pallida), within the Mojave River watershed
of San Bernardino County, California. Annual report to California Department of Fish and
Wildlife under SCP-1639. 35 pp.
Lovich, J.E., M. Thomas, K. Ironside, C. Yackulic, S.R. Puffer. 2017. Spatial distribution of
estuarine diamond-backed terrapins (Malaclemys terrapin) and risk analysis from commercial
blue crab (Callinectes sapidus) trapping at the Savannah Coastal Refuges Complex, USA:
Cooperator Report. Cooperative Agreement Number G15AC00057 between the U.S.
Geological Survey and Davidson College and Cooperative Agreement Award Number
F14AC01211 from U.S. Fish and Wildlife Service to Davidson College. 24 pp. [IP-090416,
approved 10/11/17].
Puffer, S.R., J.E. Lovich. 2017. Quarterly report to Coachella Valley Conservation Commission on
FY2018 task completions for Gopherus agassizii (July-September). 1 page.
Cummings, K.L., S.R. Puffer, J.E. Lovich. 2017. Surveys for southwestern pond turtles (Actinemys
pallida) to determine population numbers and genetic affinities at Afton Canyon with
comparative studies at Mojave Narrows Regional Park, Camp Cady Wildlife Management
Area, and Camp Ironwood. SCP-1639. Interim survey report to California Department of Fish
and Wildlife, April–May 2017. 3 pp.
Lovich, J.E., T.E. Edwards, B. Kreiser, S.R. Puffer, M. Agha. 2017. A preliminary report on genetic
affinities and relatedness of Agassiz’s desert tortoises (Gopherus agassizii) at opposite ends of

25
the Coachella Valley in California. U.S. Geological Survey Cooperators Report (Final report to
the Coachella Valley Conservation Commission). 16 pp. [IP-088375, approved 7/24/17].
Puffer, S.R and J.E. Lovich. 2017. Standard field work on turtles and tortoises. IACUC Protocol
Number 16-002. IACUC Protocol Annual Report to Northern Arizona University. 4 pp.
Lovich, J.E. and S.R. Puffer. 2017. Developing an effective Agassiz’s desert tortoise monitoring
program. Final report to the Coachella Valley Conservation Commission. U.S. Geological
Survey Cooperators Report. 26 pp. [IP-088374, approved 7/21/17].
Lovich, J.E., S.R. Puffer, K.L. Cummings, S. Greely. 2018. Feasibility study for re-establishing
southwestern pond turtles and Mojave tui chubs to Afton Canyon ACEC. U.S. Geological
Survey Cooperators Report to the Bureau of Land Management under IAA No. L16PG00229.
30 pp. [IP-093437, approved 16 Jan, 2018].
Selected web publications and films
Lovich, J.E. 1996. A brief review of the impacts of tamarisk, or saltcedar, on biodiversity in the
New World.
Lovich, J.E. 1997. Turtles and global change. Also published with same title as hard copy in the
Tortuga Gazette (newsletter of the California Turtle and Tortoise Club) 1999. 35:6–7.
Lovich, J.E. 1997. Saltcedar invasion in desert wetlands of the southwestern United States:
Ecological and political implications. Woody Plant Wetland Workshop Saltcedar, Russian
Olive.
Fenn, D., P. Garvin, and J. Lovich. 2003. Executive Producers. Sonoran Desert: fragile land of
extremes. USGS Open File Report 03-305. A 28 minute video (DVD) featuring USGS
scientists and their cooperators conducting research on the flora and fauna of the Sonoran
Desert.
Renewable Energy Global Innovations, Key Scientific Articles. 2013.
Fact sheets
Assessing desert tortoise survival and reproduction at a wind energy facility near Palm Springs,
California. 2012. California Energy Commission, Public Interest Energy Research (PIER)
Program, Environmental Area Research.
Published U.S. Geological Survey data releases
Lovich, J.E. 2016. Turtle reproductive ecology data, New Mexico, 2012-2013: U.S. Geological
Survey data release, http://dx.doi.org/10.5066/F7N014N2.

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Lovich, J.E., 2017. Population attributes Red Eared Slider Japan data: U.S. Geological Survey data
release, https://dx.doi.org/10.5066/F71C1V2B.
Lovich, J.E., Ennen, J.R., Averill-Murray, R.C., and Agha, M., 2017. Desert Tortoise reproductive
ecology and precipitation, Mojave and Sonoran Deserts—Data: U.S. Geological Survey data
release, https://dx.doi.org/10.5066/F7JS9NN9.
Thomas, M., J.E. Lovich, K. Ironside, C. Yackulic, and S.R. Puffer. 2018. Spatial distribution and
risk analysis data for Diamond-backed Terrapins relative to crab trapping, Savannah Coastal
Refuge Complex, USA. U.S. Geological Survey data release,
https://doi.org/10.5066/F7PN94W3.
Lovich, J.E. and S.R. Puffer. 2018. Reproductive ecology data for female Agassiz's Desert Tortoises
(Gopherus agassizii) in Joshua Tree National Park. USA. U.S. Geological Survey data release,
https://doi.org/10.5066/F7JW8D4B.

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Curriculum vita – J.E. Lovich
Birthplace: Alexandria, Virginia; May 30, 1957.
Marital Status: Married
Education:
B.S. George Mason University 1982
M.S. George Mason University 1984
Ph.D. University of Georgia 1990
Positions Held:
National Museum of Natural History, Division of Amphibians and Reptiles
1979–1980 Museum Assistant
George Mason University
1981–1983 Teaching Assistant
University of Georgia
1984–1986 Teaching Assistant
University of Georgia, Savannah River Ecology Laboratory
1985–1990 Graduate Student
1990–1991 Post-Doctoral Research Coordinator
Bureau of Land Management,
1991–1992 Wildlife Biologist, California Desert District Office
1992–1993 Lead Wildlife Biologist, Palm Springs-South Coast Resource Area
National Biological Survey (later Service)
1993–1996 Research Wildlife Biologist
U.S. Geological Survey (USGS)
1996–1998 Station Leader, USGS Western Ecological Research Center, Canyon Crest Field
Station, University of California, Riverside
1998–2002 Research Manager, USGS Western Ecological Research Center, Sacramento,
California
2002–2003 Center Director, USGS Western Ecological Research Center, Sacramento, California
2004 (in part) Acting Chief Scientist for Biology, USGS Headquarters, Reston, Virginia
2003–2005 Chief, USGS Grand Canyon Monitoring and Research Center, Flagstaff, Arizona
2005–2009 Deputy Center Director, USGS Southwest Biological Science Center,
Flagstaff, Arizona
2009–present Research Ecologist
University of California, Riverside
1997–2000 Associate Professor (Courtesy)
Northern Arizona University
2005–present Associate Professor (Courtesy)
Arizona Invasive Species Advisory Council
2005–2006 Appointed by the Governor of Arizona
IUCN/SSC Tortoise and Freshwater Turtle Specialist Group
2009–present

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Cadi Ayyad University
2010–2012 Visiting Professor (Master’s Program in Ecological Engineering & Methodologies
of Biodiversity Analysis and Management), Marrakech, Morocco
Awards and Special Recognition:
1981 Grant in Aid of Research. Sigma Xi
1983 Conservation Scholarship. Fairfax Audubon Society
1985 Eugene P. Odum Coastal Conservation Award. Sierra Club Gulf Coast Regional
Conservation Committee
1989 Second place for best student paper award from the Herpetologists League
1989-1990 Graduate Student Education Program scholarship, Savannah River Ecology
Laboratory
1990 Student Research Award, Association of Southeastern Biologists
1993 Outstanding Performance Evaluation from the Palm Springs-South Coast Resource Area,
Bureau of Land Management
1993 Certificate of Recognition from Secretary of the Interior, for outstanding contributions
toward the establishment of the National Biological Survey
2008 Fulbright Senior Specialists Award to teach and conduct research at Cadi Ayyad
University, Marrakech, Morocco
2011 The Wildlife Society award for outstanding book in wildlife ecology and management,
“Turtles of the United States and Canada. Second Edition”
2013, 2014 Performance Awards, U.S. Geological Survey
2015 Elected Fellow of The Linnean Society of London
Fieldwork: United States (Alabama, Arizona, California, Georgia, Mississippi, New Mexico, Ohio,
Pennsylvania, South Carolina, Virginia), Japan, Morocco
Professional Organizations
American Society of Ichthyologists and Herpetologists [1986–2003)]
Editorial Board (1994–1995)
California Exotic Pest Plant Council (now California Invasive Plant Council)
Board Member (1992–1997)
Chelonian Research Foundation [1993 – present]
Editorial Review Board, Chelonian Conservation and Biology (1993–present)
Editor (2011–present)
Herpetological Society of Japan [2015–present]
Associate Editor (2015–present)
Herpetologists’ League [1982–2006]
Executive Council (2000–2003)
International Herpetological Symposium, Inc. [1993–2001]
Associate Editor, Herpetological Natural History (1995–1996)
Society for the Study of Amphibians and Reptiles [1981–2010]
Associate Editor, Herpetological Review (1997–2000)

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Endnote – Contributions to the history of the Division of Amphibians & Reptiles – USNM
As one grows older, there comes a desire to record one’s past experiences and also the history of one’s
workplace. None of us in the USNM Division of Amphibians and Reptiles has expressed a desire to
write a divisional history for the past half-century. As an alternative, I am encouraging colleagues who
have been associated with the division to create autobiographical sketches. Although such sketches
will not provide a detailed history of divisional activities, each offers a unique perspective of past
divisional activities and insights into each author’s contribution to the division and, of course, a
window into the author’s personality.
The SHIS series is an obvious outlet. SHIS has been a facet of the division’s contribution of research
information to the herpetological community since its establishment in 1968 by James A. Peters.
May 2018 GRZ
Previously published contributions to divisional history
SHIS 1. A list of the herpetological publications of the United States National Museum,
1853-1965. James A. Peters 1965 [revised 1968].
SHIS 42. A revised list of the herpetological publications of the National Museum of
Natural History (USNM) 1853-1978. Ronald I. Crombie 1979.
SHIS 51. Biography and bibliography of James A. Peters. Frances J. Irish & George R.
Zug 1982.
SHIS 101. Herpetological publications of the National Museum of Natural History
(USNM), 1853-1994. Ronald I. Crombie 1994.
SHIS 147. Biographical sketch and bibliography of W. Ronald Heyer. W. Ronald Heyer
& Miriam H. Heyer 2016.
SHIS 148. Biographical sketch and bibliography of James B. Murphy. James B. Murphy
2016.
SHIS 149. Biographical sketch and bibliography of C. Kenneth Dodd, Jr. C. Kenneth
Dodd, Jr. 2016.
SHIS 150. Biographical sketch and bibliography of Carl H. Ernst. Carl H. Ernst 2016.
SHIS 151. Biographical sketch and bibliography of Richard Highton. Richard
Highton 2017.
SHIS 152. Biographical sketch and bibliography of Robert P. Reynolds. Robert P.
Reynolds 2017.