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Arrhenatheretalia elatioris uncritical checklist of Europe

April 2018
Phytologia Balcanica 24(1)

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Bulgarian Academy of Sciences

This paper presents a compilation of all syntaxa whenever related to the Arrhenatheretalia elatioris order at European level. Only those described according to the Braun-Blanquet approach are considered herein. The checklist consists of 15 alliances, 16 suballiances, 246 associations, 292 subassociations, 144 variants, 40 facies, and 36 plant communities without rank. The largest number of associations is encountered within the Arrhenatherion alliance-93, followed by Cynosurion with 85 and Polygono-Trisetion with 25 associations. The most frequently mentioned associations in Europe are Pastinaco sativae-Arrhenatheretum elatioris (Arrhenatherion) cited for 22 countries and Lolio perennis-Cynosuretum cristati (Cynosurion) cited for 20 countries. The checklist was compiled with the aim of revealing some of the existing syntaxonomical problems within the Arrhenatheretalia order. The numerous associations of some alliances suggest a suspiciously high syntaxonomical diversity. The current checklist offers a simple overview, but does not suggest a new classification system, nor any syntaxonomical revision. It could be used primarily as a starting point for any future large-scale analyses.

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PHYTOLOGIA BALCANICA 24 (1): 99 – 147, Sofia, 2018

Arrhenatheretalia elatioris uncritical checklist of Europe

Nikolay Velev

Department of Plant and Fungal Diversity and Resources, Institute of Biodiversity and

Ecosystem Research, Bulgarian Academy of Sciences, 23 Acad. G. Bonchev Str. 1113

Sofia; Bulgaria; e-mail: nvelev@bio.bas.bg

Received: September 19, 2017 ▷ Accepted: January 07, 2018

Abstract. This paper presents a compilation of all syntaxa whenever related to the Arrhenatheretalia elatioris order

at European level. Only those described according to the Braun-Blanquet approach are considered herein.

The checklist consists of 15 alliances, 16 suballiances, 246 associations, 292 subassociations, 144 variants, 40

facies, and 36 plant communities without rank. The largest number of associations is encountered within the

Arrhenatherion alliance – 93, followed by Cynosurion with 85 and Polygono-Trisetion with 25 associations.

The most frequently mentioned associations in Europe are Pastinaco sativae-Arrhenatheretum elatioris

(Arrhenatherion) cited for 22 countries and Lolio perennis-Cynosuretum cristati (Cynosurion) cited for 20

countries.

The checklist was compiled with the aim of revealing some of the existing syntaxonomical problems

within the Arrhenatheretalia order. The numerous associations of some alliances suggest a suspiciously

high syntaxonomical diversity. The current checklist offers a simple overview, but does not suggest a new

classification system, nor any syntaxonomical revision. It could be used primarily as a starting point for any

future large-scale analyses.

Key words: compilation, catalogue, mesic grasslands, nomenclature, overview, syntaxonomy, vegetation

Introduction

The order Arrhenatheretalia elatioris unites mainly

secondary mesic grasslands, which have developed in

places of former forests and are managed as hay mead-

ows and pastures (Horvat 1960; Horvatić 1963; Rod-

well & al. 2002). The order is well represented in Cen-

tral and West Europe, but it could be found in the

other parts of the continent too (Knapp 1979a, b). Ar-

rhenatheretalia elatioris Tüxen 1931 is usually subor-

dinated to the class Molinio-Arrhenatheretea Tüxen

1937 (Dierschke 1997; Päzolt & Jansen 2004; Mucina

& al. 2016). The order is typified by the alliance Tri seto-

Polygonion bistortae Br.-Bl. 1949 ex 1950, mentioned

as lectotypus (Dengler & al. 2003). At European level,

many scientists have contributed to the knowledge of

Arrhenatheretalia order. To mention only a couple of

works from the last three decades, in Central and West

Europe it is studied by Ellmauer & Mucina (1993), Di-

erschke (1995, 1997), Jansen & Päzolt (2001), Rod-

ríguez-Rojo & Sánchez-Mata (2004, 2006), Hájková &

al. (2007), Hegedüšová & Ružičková (2007), Janišová

& al. (2007, 2010), Škodová (2007), Uhliarová & al.

(2007), Rozbrojová & al. (2010), Allegrezza & Biondi

(2011), Rodríguez-Rojo & al. (2014); and in the East

and Southeast Europe by Sanda & Popescu (1991),

Kojić & al. (2004, 2008), Stančić (2000, 2008a), Zelnik

(2007), Didukh & Kuzemko (2009), Kuzemko (2009b,

2016), Velev & al. (2010, 2011a, b), Aćić & al. (2013a,

b), Šilc & al. (2014) and Shushpannikova & Yamalov

(2014). Among the latest classification systems in Eu-

rope are those proposed by Mucina & al. (2016) and

Rodríguez-Rojo & al. (2017).

Even bearing in mind all above-mentioned studies

and even assuming that supra-national studies of the

palaearctic grasslands are frequently in the scientific

100 Velev, N. • Checklist of Arrhenatheretalia order

focus (Janišová & al. 2016), still there exists a big

knowledge gap: a clear and complete syntaxonomical

scheme of Arrhenatheretalia order at European level

is lacking.

The purpose of this paper is to bring in one place

all European syntaxa whenever related to the Ar-

rhenatheretalia order. It is focused only on compiling

all appropriate information and syntaxa related to this

order, but does not suggest any classification schemes.

Such work will contribute to a more profound under-

standing of the order in European context. It can be

used as a base for further and more detailed classifica-

tion schemes, particularly below alliance level.

Material and methods

The checklist was based on literature and a review of

Internet sources. More than 500 bibliographic entities

containing information about Arrhenatheretalia order

were considered. The checklist consists of all subor-

dinated to Arrhenatheretalia syntaxa, including plant

communities without rank. Only syntaxa described

according to the Braun-Blanquet approach (Braun-Bl-

anquet 1965; Westhoff & van der. Maarel 1973) were

included. With the intention to demonstrate the full

extent of Arrhenatheretalia order in its utmost com-

pleteness, all syntaxa in this work were brought in and

presented just as taken from the publications. Orig-

inal diagnoses, nomenclature types, syntaxa relation

to higher ranks, diagnostic species and synonymy are

presented here exactly as they come from the biblio-

graphic sources, simply compiled, without any recon-

sideration or revision.

Syntaxa in the checklist are listed chronological-

ly by decrease of their rank. Syntaxa published in the

same year are listed alphabetically. All associations are

listed within the alliances directly, disregarding the

rank of suballiance, since the latter were accepted on-

ly by a few authors. Because of the varying authors’

opinions, some associations are included within two

alliances, while other are considered both as validly

published and as synonyms. The associations, related

to two or more alliances, are described in the check-

list only once, and the place of their full description is

mentioned.

A nomenclature block was prepared for every syn-

taxon, depending on the amount of available infor-

mation. Nomenclature blocks contain the following

headings:

Original name: the original name of the syntaxon

is given. This is the name comprising the original di-

agnosis, the type of the syntaxon.

Synonyms: all synonyms taken from literature and

Internet sources are compiled and listed chronologi-

cally.

Corresponding names: according to ICPN (Def. X),

the corresponding names are extracted from synony-

my and presented separately after the synonyms.

Pseudonyms: following the recommendations of

ICPN (Def. X; Recomm. 46J), all found pseudonyms

are listed chronologically after the synonyms.

Homonyms: these were validly published names,

but based on different nomenclature types and spelled

identically (ICPN: Def. IX; Art. 31, 32, 33).

Remarks: any relevant information presented, or

some controversial decisions found in synonymy are

commented.

Diagnostic species: all diagnostic species taken

from literature and Internet sources are compiled and

listed alphabetically.

Distribution: alphabetical list of all countries the

syntaxon is mentioned for.

Variations: syntaxa below association level (subas-

sociation, variant, facies) are listed alphabetically first

by countries and then chronologically. Subordinated

syntaxa always start with „typicum”, if any, regardless

of the year of publication.

References: the referred bibliographic sources are

listed chronologically.

Abbreviations used in the text followed Weber & al.

(2000):

Art. = Article (ICBN; ICPN)

Def. = Definition (ICPN)

excl. = excluso

ICBN = International Code of B otanical Nomenclature

ICPN = International Code of Phytosociological

Nomenclature

incl. = incluso

nom. ambig. = nomen ambiguum (ICPN – Art. 36)

nom. conserv. propos. = nomen conservandum prop-

ositum (ICPN – Def. XIII; Principle II; Art. 22; 52)

nom. correct. = nomen correctum (ICPN – Art. 43;

44; 48d)

nom. illeg. = nomen illegitimum (ICPN – Art. 17; 34a)

nom. inval. = nomen invalidum (ICPN – Art. 1; 3; 4;

9; 10b; 11; 17; 29b)

101Phytol. Balcan. 24(1) • Sofia • 2018

nom. invers. = nomen inversum (ICPN – Art. 42; 48;

Recomm. 42A)

nom. mut. = nomen mutatum (ICPN – Art. 48e)

nom. nud. = nomen nudum (ICPN – Recomm. 46D; 46E)

nom. prov. = nomen provisorium (ICPN – Art. 3b; 6)

nom. rejic. propos. = nomen rejiciendum propositum

(ICPN – Art. 36; Recomm. 36A)

p. p. = pro parte

p. min. p. = pro minimo parte

p. max. p. = pro maximo parte

pro syn. = pro synonymo (ICPN – Recomm. 46F)

Recomm. = Recommendation (ICPN)

Syntax. syn. = Syntaxonomical synonym (ICPN –

Def. X)

Results

The obtained checklist consists of 15 alliances, 16

suballiances, 246 associations, 292 subassociations,

144 variants, 40 facies, and 36 plant communities

without rank (Annex 1). Alliance Arrhenatherion ela-

tioris totalls 93 associations, followed by Cynosurion

cristati with 85 and Polygono bistortae-Trisetion flaves-

centis with 25 associations (Table 1).

The most often mentioned associations in Europe

are Pastinaco sativae-Arrhenatheretum elatioris (Ar-

rhenatherion) cited for 22 countries and Lolio per-

ennis-Cynosuretum cristati (Cynosurion) cited for 20

countries (cf. Annex 1). The coun-

tries richest in associations are

France (64 associations cited), fol-

lowed by Ukraine (48) and Roma-

nia (36) (Fig. 1).

Table 1. A summary of all syntaxa whenever related

to Arrhenatheretalia elatioris. Associations presented

simultaneously in different alliances are counted only once. * –

alliances recently reconsidered and extracted from synonymy as

being separate syntaxa. ** – nomen provisorum. *** – recently

described alliance, presumably related to Arrhenatheretalia.

Order

Alliance

Suballiance

Association

Subassociation

Variant

Facies

Plant community

without rank

Arrhenatheretalia elatioris

Alchemillo-Ranunculion repentis*

Arrhenatherion elatioris 4 93 148 51 20 18

Brachypodio-Centaureion nemoralis*

Cynosurion cristati 7 85 84 57 17 9

Festucion pratensis 1 21 35 20

Helictotricho-Bistortion officinalis 1 2

Pančićion serbicae 7 2

Phyteumato-Trisetion*

Poion alpinae 5 1

Polygono-Trisetion flavescentis 4 25 20 11 3 3

Potentilion-Ranunculion repentis**

Salvio pratensis-D actylidion glomeratae 3 3

Rumicion thyrsiflori 1

Trifolio-Brizion elatioris 1 3

Alliance not specified 4 3

?Anthrisco sylvestris-Arrhenatherion*** 2

Total 15 16 246 292 144 40 36

010 20 30 40 50 60 70

United Kingdom

Ukraine

Switzerland

Spain

Slovenia

Slovakia

Serbia

Russia

Romania

Portugal

Poland

Netherlands

Montenegro

Macedonia

Lithuania

Latvia

Italy

Ireland

Hungary

Germany

France

Czech Republic

Croatia

Bulgaria

Bosnia and Herzegovina

Belgium

Belarus

Austria

Albania

Number of associations

Fig. 1. Number of associations related to

Arrhenatheretalia elatioris per country.

Synonyms and plant communities without

rank not counted.

102 Velev, N. • Checklist of Arrhenatheretalia order

Discussion

The current Arrhenatheretalia catalogue is elaborated

with the aim of revealing the existing syntaxonomical

problems within this order, mostly below alliance lev-

el. It is intended mainly as a starting point for pick-

ing up some ideas for future large-scale analyses. For

this purpose, such checklist needs to be rather raw and

independent, so as not to be affiliated consciously to

any recent classification scheme. The syntaxonomi-

cal syniopsis in this paper does not advocate a new

classification system, nor any revision! On the contra-

ry, it comes to reflect mostlty the existing controver-

sy about some syntaxa, synonymy and diagnostic spe-

cies within Arrhenatheretalia. According to different

authors, many syntaxa within this order are simulta-

neously considered as validly published and as syno-

nyms. Such discrepancies should be generally resolved

by applying large-scale analyses, as simple reference

to literature will not be enough. Large-scale analy-

ses could be done best with the help of the phytoso-

ciological databases included in the European Vege-

tation Archive (EVA; http://euroveg.org/eva-database;

Chytrý & al. 2016) and the Global Index of Vegeta-

tion-Plot Databases (GIVD; http://www.givd.info/;

Dengler & al. 2011).

A closer look at the checklist (cf. Annex 1)

shows excessive syntaxonomical diversity within Ar-

rhenatheretalia even at European scale: 15 alliances

with 246 associations. With their publication Vegeta-

tion of Europe, Mucina & al. (2016) have solved this

problem to some extent, since a new alliance of such

grassland vegetation has arisen recently (cf. Rod-

ríguez-Rojo & al. 2017). Mucina & al. (2016) have pro-

posed full reconsideration of the Arrhenatheretalia or-

der in Europe, which now consists more realistically

of eight alliances only. Rodríguez-Rojo & al. (2017) al-

so invested much effort in classifying the lowland hay

meadows and pastures from West and Central Europe

into 36 associations.

Great many problems below alliance level still exist

in Arrhenatheretalia. Syntaxonomically, the Arrhena-

therion alliance has reached 93 associations. Here it

seems appropriate to recall Braun-Blanquet’s state-

ment that an alliance must unify „the most closely re-

lated associations” (Braun-Blanquet 1965: 363). How

closely then 93 associations could be related, or is such

enormous syntaxonomical diversity real? Meanwhile,

this misunderstanding is supported by many discrep-

ancies in the authors ideas about the diagnoses of

some syntaxa. Many synonyms encountered in liter-

ature are quite controversial.

Since vegetation classification is dynamic and

evolving, the content of the Arrhenatheretalia order

varies continuously. Being vitally dependent on acces-

sibility of bibliographical sources, syntaxomical diver-

sity presented in this paper may lack some published

syntaxa.

Acknowledgments. The author would like to thank all col-

leagues who supported this work by providing kindly biblio-

graphic data. Special thanks are extended to Prof. Dr. Iva Apos-

tolova, who suggested some good ideas and encouraged the

author across the years during preparation of the manuscript.

The athor gratefully acknowledges the efforts of the peer review-

ers for their helpful comments on an earlier draft of the manuscript.

Annex 1. Checklist of Arrhenatheretalia elatioris ord er.

Class MOLINIO-ARRHENATHERETEA Tüxen 1937

Order ARRHENATHERETALIA ELATIORIS Tüxen 1931

Original name: Arrhenatheretalia Tüxen 1931

nom. correct.

Synonyms: Arrhenatheretalia elatioris Pawłowski

1928 nom. nud. (Art. 2b; 8); Arrhenatheretalia Horvatić

1930; 1958 p. p.; Arrhenatheretalia Br.-Bl. 1931 (Art. 8);

Arrhenatheretalia (Pawłowski 1926) Tüxen 1937; (Trifo -

lio-) Arrhenatheretalia Oberd. 1949; Plantaginetalia ma-

joris Tüxen (1947) 1950 p. p. (Art. 8; 29a); Arrhenathere-

talia elatioris (Pawłowski 1928) Br.-Bl. 1949 ex Br.-Bl.

1950; Plantaginetalia majoris Tüxen ex von Rochow 1951

p. p.; Arrhenatheretalia Zeidler 1954 p. min. p.; Lolieta-

lia perennis Doing 1963 (Art. 8); Trifolio-Cynosureta-

lia Sougnez & Limbourg 1963 nom. inval. (Art. 2d; 3b);

Trifolio-Phleetalia (Sougnez & Limbourg 1963) Pas-

sarge 1969; Trifolio repentis-Phleetalia pratensis Passarge

1969; Cynosuro-Phleetalia pratensis Passarge 1969; Fes-

tuco-Arrhenatheretalia (Pawłowski & al. 1928) Passarge;

1969 Galio-Achilletalia millefoliae Passarge 1975; Ar-

rhenatheretalia elatioris (Pawłowski 1928) Oberdorfer

1983; Cynosuretalia cristati de Foucault 1989; Plantagi-

ni-Lolietalia Mucina 1991 (Art. 8); Poo alpinae-Triseteta-

lia Ellmauer & Mucina in Mucina & al. 1993; Plantagini-

Prunelletalia Ellmauer & Mucina 1993.

103Phytol. Balcan. 24(1) • Sofia • 2018

Corresponding names: Arrhenatheretea Br.-Bl.

1947; Arrhenatherenea elatioris de Foucault 1984;

1989; Veronico serpyllifoliae-Cynosurenalia de Fou-

cault 1989; Trisetenalia flavescentis Oberdorfer 1990;

Trifolienalia Oberdorfer 1990. According to Ellmau-

er & Mucina (1993) incl.: Phleo-Leontodion (Br.-Bl. &

Berset 1957) Dietl 1972.

Homonyms: Arrhenatheretalia Krajina 1933 (Art. 31).

Diagnostic species: Achillea millefolium, A. prat-

ensis, A. submillefoilum, Agrostis capillaris, Alchemil-

la connivens, A. crinita, A. vulgaris, A. cymatophylla

(local for Krkonoše Mountains, Czech Republic), A.

subcrenata, Alopecurus pratensis, A. utriculatus, An-

thoxanthum odoratum, Anthriscus sylvestris, Arh-

henatherum elatius, Armeria elongata, Astrantia major,

Avenula pubescens, Bellis perennis, Briza media, Bro-

mus commutatus, B. hordeaceus, B. mollis, B. racem-

osus, Campanula patula, Cardamine matthiolii, Carex

caryophyllea, C. pallescens, Carum carvi, Centaurea ja-

cea, C. phrygia, Cerastium arvense (local for Krkonoše

Mountains, Czech Republic), C. holosteoides, Cichori-

um intybus, Crepis biennis, C. capillaris, Crocus albi-

florus, C. biflorus, Cynosurus cristatus, Dactylis glom-

erata, Daucus carota, Eryngium planum, Equisetum

pratense, Euphrasia stricta, Festuca pratensis, F. r u bra ,

Fragaria vesca (local for Krkonoše Mountains, Czech

Republic), Galium mollugo agg., G. boreale, Gaudinia

fragilis, Gentiana utriculosa, Geranium pratense, Glad-

iolus imbricatus, Heracleum sibiricum, H. sphondyli-

um, Hieracium pilosella, H. umbellatum, Knautia ar-

vensis, Lathyrus pratensis, Leontodon autumnalis, L.

hispidus, Leucanthemum praecox, L. vulgare, Linum

catharticum, Lotus corniculatus, Luzula campestris,

Malva moschata, Medicago lupulina, Moenchia manti-

ca, Narcissus poeticus, Ononis pseudihircina, Orchis co-

riophora, Ornithogalum umbellatum, Pastinaca sativa,

Phleum pratense, Picris hieracioides, Pimpinella major,

P. saxifraga, Plantago media, Polygala vulgaris, Prunel-

la vulgaris, Ranunculus strigulosus, Rhinanthus alec-

torolophus, R. angustifolius, R. mediterraneus, R. mi-

nor, R. rumelicus, Rhytidiadelphus squarrosus, Rorippa

pyrenaica, Rumex acetosa, R. crispus, Saxifraga granu-

lata, Scabiosa columbaria, Scutellaria hastifolia, Sene-

cio jacobaea, Silene vulgaris, Stellaria graminea, Tara x-

um officinale, Tragopogon orientalis, T. pratensis,

Traunsteinera globosa, Trifolium aureum, T. campestre,

T. dubium, T. patens, T. pratense, T. repens, T. rubens,

Trisetum flavescens, Verbascum blattaria, Veronica ar-

vensis, V. chamaedrys, V. serpyllifolia, Vicia angustifo-

lia, V. cracca, V. sepium, Viola canina.

References: Borza (1959); Cincović (1959); Horvat

(1962; 1974); Horvatić (1963); Medwečka-Kornaš &

al. (1966); Resmeriţă (1970); Dihoru (1975); Blažková

(1979); Mititelu & al. (1986, 1988); Coldea (1987, 1990);

Ellenberg (1988); de Foucault (1989); Sanda & Popescu

(1991); Ivan & al. (1993); Ellmauer & Mucina (1993);

Oberdorfer (1993); Dierschke (1997; 1999); Kojić & al.

(1997); Krahulec & al. (1997); Sanda & al. (1998, 2001);

Stepanovich (2000); Bardat & al. (2001); Jansen & Pä-

zolt (2001); Mihăilescu (2001); Rivas-Martínez & al.

(2001, 2002); Snowarski (2002-2005); Csergő & Frink

(2003); Kaligaric & al. (2003); Burkart & al. (2004); Pä-

zolt & Jansen (2004); Rodríguez-Rojo & Sánchez-Mata

(2004); Mosek & Miazga (2006); Tomescu & al. (2006);

Zelnik (2007); Solomakha (2008); Popescu & Boruz

(2008); Stančić (2008a); Mucina & al. (2016).

Alliance ARRHENATHERION ELATIORIS Luquet 1926

Original name: Arrhenatherion Luquet 1926 (Ar-

rhenatherum elatius).

Synonyms: Arrhenatherion elatioris Br.-Bl. 1925 p.

p.; Arrhenatherion elatioris (Br.-Bl. 1925) Koch 1926;

Arrhenatherion elatioris Koch 1926 nom. nud. (Art.

2b); Arrhenatherion Horvatić 1930; 1941; 1958; Ar-

rhenatherion elatioris Tüxen 1931 (Art. 31); Alopecu-

rion utriculati Zeidler 1954 p. p.; Alopecurion pratensis

Passarge 1964 p. p.; Dauco-Arrhenatherion Passarge

1964; Brachypodio pinnati-Centaureion nemoralis Br.-

Bl. 1967; Agrostio-Festucion rubrae Passarge 1969 p.

max. p. non Puşcaru & al. 1956; Polygalo-Festucion

rubrae Passarge 1969; Phyteumato-Festucion rubrae

Passarge 1969; Dauco-Arrhenatherion Passarge 1969;

Triseto-Arrhenatherion Passarge 1969; Anthrisco-Her-

acleion Passarge 1975 p. p.; Hyperico-Vicion angustifo-

liae Passarge 1975 p. max. p.; Trifolio repentis-Lolion

perennis Dietl 1983 (Art. 5; 8). According to Ellmauer

& Mucina (1993) Alopecurion Passarge 1964 is syntax-

onomical synonym of Arrhenatherion Koch 1926. Mu-

cina & al. (2016) include Festucion pratensis Sipailova

& al. 1985 in the synonymy of Arrhenatherion.

Corresponding names: Bromo erecti-Arrhenathe-

renion Rivas Goday & Rivas-Martínez 1963; Tr ifo-

lio-Arrhenatherenion Rivas Goday & Rivas-Martínez

1963; Malvo-Arrhenatherenion Rivas Goday & Rivas-

Martínez 1963.

Annex 1. Continuation.

104 Velev, N. • Checklist of Arrhenatheretalia order

Remarks: 1. Dierschke (1997) pointed Alopecurion

pratensis Passarge 1964 as synonym of Arrhenatherion

elatioris Koch 1926 (Syn.: Arrhenatherion elatioris Lu-

quet 1926). Hájková (2007) and Hájková & al. (2007)

presented Alopecurion pratensis Passarge 1964 as a

synonym of Deschampsion cespitosae Horvatić 1930;

2. Mucina & al. (2016) accepted the synonym Brach-

ypodio-Centaureion nemoralis Br.-Bl. 1967 as a sepa-

rate syntaxon.

Diagnostic species: Achillea asplenifolia, A. colli-

na, A. millefolium, A. pannonica, A. pratensis, A. se-

tacea, Agrimonia eupatoria, Agrostis gigantea, Aju-

ga reptans, Alchemilla vulgaris agg. (всички видове

с изкл. на A. glaucescens), Alopecurus pratensis, Anth-

oxanthum alpinum, A. odoratum, Anthriscus sylvestris,

Arrhenatherum elatius, Avenula pubescens, Bellis per-

ennis, Bromus arvensis, B. mollis, B. racemosus, Cam-

panula patula, C. rapuculus, Cardaminopsis arenosa,

Carex pallescens, Carum carvi, Centaurea banatica, C.

jacea, Cerastium holosteoides, Cirsium arvense, Colchi-

cum autumnale, Coeloglossum viridae, Crepis biennis,

Cruciata laevipes, Cynosurus cristatus, Dactylis glom-

erata, Daucus carota, Equisetum arvense, Euphorbia

esula, Euphrasia stricta, Festuca arundinaceae, F. dif -

fusa, F. heterophylla, F. nigrescens, F. pratensis, F. r u -

bra, Galium mollugo agg., G. verum agg., Geranium

pratense, Heracleum sphondylium, Hieracium muro-

rum, Holcus lanatus, Hordelymus europaeus, Hyperi-

cum perforatum, Knautia arvensis agg., K. kitaibelii,

Lathyrus pratensis, Leontodon hispidus, Leucanthe-

mum ircutianum, L. vulgare, Linum catharticum, Lotus

corniculatus, Luzula campestris, L. divulgata, L. multi-

flora, L. pallescens, L. sudetica, Malva moschata, Med-

icago lupulina, Melica uniflora, Odontites vulgaris, On-

onis minutussima, O. repens, O. spinosa, Ornithogalum

umbellatum, Orobanche caryophyllacea, Pastinaca sa-

tiva, Pedicularis schizocalyx, Peucedanum carvifolia,

Phleum bertolonii, P. pratense, Phyteuma orbiculare,

Pilosella caespitosa, Pimpinella major, P. saxifraga,

Plantago lanceolata, P. m e d i a , Poa angustifolia, P. prat -

ensis, Potentilla alba, P. argentea, Prunella grandiflora,

P. vulgaris, Ranunculus acris, R. bulbosus, R. steveni, R.

strigulosus, Rhinanthus minor, Rubus caesius, Rumex

acetosa, R. thyrsiflorus, Sanguisorba minor, Saxifra-

ga bulbifera, S. granulata, Scabiosa columbaria, Sene-

cio jacobea, Silene vulgaris, Stachys officinalis, Stellaria

graminea, Taraxacum Sect. Ruderalia, Thlaspi caerule-

scens, Thuidium philibertii, T. recognitum, Tragopogon

orientalis, T. pratensis, Trifolium campestre, T. dubium,

T. flexuosum, T. hybridum, T. prаtense, T. repens, Tri-

setum flavescens, Veronica chamaedrys, V. hederifolia,

V. vindobonensis, Vicia cassubuca, V. cracca, V. sepium,

V. tenuifolia.

References: Borza (1959); Blečić & Tatić (1960);

Horvat (1962); Horvatić (1963); Moravec (1965);

Borza (1966); Medwečka-Kornaš & al. (1966); Jurko

(1974); Blažková (1979); de Foucault (1984; 1989);

Boşcaiu & Täuber (1985); Ellenberg (1988); Sanda &

Popescu (1991); Ellmauer & Mucina (1993); Ober-

dorfer (1993); Zuidhoff & al. (1995); Dierschke (1997,

1999); Kojić & al. (1997); Krahulec & al. (1997); San-

da & al. (1998, 1999); Stepanovich (2000); Rivas-Mar-

tínez & al. (2001, 2002); Šeffer & al. (2002); Snowar-

ski (2002-2005); Chytrý & Tichý (2003); Dutoit & al.

(2003); Kabucis & al. (2003); Kaligaric & al. (2003);

Burkart & al. (2004); Päzolt & Jansen (2004); Rod-

ríguez-Rojo & Sánchez-Mata (2004); Sarbu & al.

(2004); Meshinev & al (2005); Rasomavicius & al.

(2006); Hájková (2007); Hájková & al. (2007); Rodwell

& al. (2007); Uhliarová & al. (2007); Veen & al. (2007);

Solomakha (2008); Jarolímek & al. (2008a); Škodová

& Janišová (2008); Velev & al. (2010); Biondi & al.

(2014); Mucina & al. (2016).

Suballiances related to Arrhenatherion:

Suball. Centaureo jaceae-Arrhenatherenion elatioris

de Foucault 1989 nom. illeg. (Art. 22)

Synonym: Trifolio montani-Arrhenatherenion elati-

oris Rivas Goday & Rivas Mart. 1963.

Suball. Colchico autumnalis-Arrhenatherenion ela-

tioris de Foucault 1989 nom. correct. (Art. 41b)

Original name: Colchico autumnale-Arrhenathe-

renion elatioris de Foucault 1989.

Suball. Rumici obtusifolii-Arrhenatherenion elatioris

de Foucault 1989

Suball. Poo angustifoliae-Arrhenatherenion elatioris

Felzines 2012

References: Bardat & al. (2001); Ferrez (2007).

Associations related to Arrhenatherion:

Ass. Alchemillo-Trisetetum flavescentis Horvat 1931

Distribution: Bosnia and Herzegovina.

Annex 1. Continuation.

105Phytol. Balcan. 24(1) • Sofia • 2018

Ass. Anthoxantho odorati-Agrostietum tenuis Sill-

inger 1933

Remarks: for details see in alliance Cynosurion.

Ass. Holcetum lanati Issler 1934

Original name: Holcus lanatus Issler 1934.

Synonyms: Holcetum lanati Gams 1927 nom. nud.

(Art. 2b).

Diagnostic species: Carex ovalis, Cerastium luco-

rum, Cirsium canum, Holcus lanatus, Lychnis flos-cu-

culi, Plantago lanceolata, Rumex acetosella, Sanguisor-

ba officinalis, Trifolium pratense.

Distribution: Croatia, Czech Republic (also in De-

schampsion cespitosae), Poland, Romania (in Molini-

etalia caeruleae Koch 1926, Calthion palustris Tüx-

en 1937; also in Potentillo-Polygonetalia Tüxen 1947,

Potentillion anserinae Tüxen 1937), Slovakia (in Des-

champsion cespitosae Horvatić 1930), Ukraine (as Hol-

cetum lanati Issler 1936 in Molinion Koch 1926; also in

Alopecurion pratensis Passarge 1964).

Variations: Czech Republic: var. Arrhenatherum

elatius; var. Carex hirta; var. Cirsium palustre; Ukraine:

subass. caricetosum paniceae Kuzemko 2009; subass.

festucetosum rubrae Kuzemko 2009 with two vari-

ants – var. typicum, var. Trifolium rubens.

Ass. Festucetum pratensis Soό 1938 (Art. 36)

Synonyms: Agrostio stoloniferae-Deschampsietum

cespitosae Ujvárosi 1947 p. p.; Festuceum pratensis me-

zophilim Csűrös 1970 (Art. 34a); Cirsio cani-Festuce-

tum pratensis Májovski & Ružičková 1975 p. p.

Diagnostic species: Cirsium canum, Bromus mollis,

Dactylis glomerata, Festuca pratensis, F. r ubr a, Holcus

lanatus, Phleum pratense, Poa pratensis, P. trivialis.

Distribution: Belarus (also in Festucion praten-

sis Sipaylova & al 1985), Hungary (in Deschampsion

caespitosae), Latvia, Lithuania, Romania (also in ord.

Molinietalia caeruleae and Deschampsietalia caespitos-

ae Horvatić 1956), Russia (in Festucion pratensis), Ser-

bia [in Agrostion albae (stoloniferae) Soό (1933) 1971],

Slovakia, Slovenia (as Festucetum pratensis Soό (1938)

1955 and Festucetum pratensis Soό 1958), Ukraine (in

Festucion pratensis).

Variations: Belarus: subass. typicum; subass. agros-

tidetosum vinealis sensu Stepanovich 2000; subass. alo-

pecuretosum pratensis sensu Stepanovich 2000; subass.

anthoxanthetosum odorati sensu Stepanovich 2000; sub-

ass. brometosum mollis sensu Stepanovich 2000; sub-

ass. cynosuretosum cristati sensu Stepanovich 2000; sub-

ass. dactylidetosum glomeratae sensu Stepanovich 2000;

subass. deschampsietosum cespitosae sensu Stepanovich

2000; subass. festucetosum rubrae sensu Stepanovich

2000; subass. geranietosum pratensi sensu Stepanovich

2000; subass. phleetosum pratensi sensu Stepanovich

2000; subass. poetosum pratensis sensu Stepanovich

2000; subass. poetosum trivialis sensu Stepanovich 2000;

subass. trifolietosum hybridi sensu Stepanovich 2000;

subass. trifolietosum pratensi sensu Stepanovich 2000;

Latvia: var. typicum; var. Alopecurus pratensis; var. Ga-

lium boreale; var. Medicago falcata; Lithuania: fac. typi-

cum; fac. Anthriscus sylvestris; fac. Bromopsis inermis;

fac. Deschampsia caespitosa; fac. Elytrigia repens; fac. Lo-

tus corniculatus; fac. Potentilla anserina; fac. Ranuncu-

luc acris; fac. Trifolium medium; fac. Stellaria gramin-

ea; fac. Urtica dioica; Romania: subass. transsilvanicum

Soό 1938; subass halophilum Csűrös 1947; subass. bro-

metosum commutati Soό 1957; subass. poëtosum trivia-

lis Soό 1957 (Syn.: Poa trivialis-Festuca pratensis Borza

1959); subass. ranunculetosum acris Juhász Nagy apud

Soό 1957; subass. alopecuretosum Bodrogközy 1962;

subass. brometosum mollis-hordeacei (Siroki ined.) apud

Soό 1964; subass. festucetosum pratensis Soό 1964; sub-

ass. poëtosum pratensis Soό 1964; subass. caricetosum

distantis (Horv. 1930) Pázmány 1971; subass. dactyleto-

sum glomerati Grigore 1971; subass. festucetosum valesi-

acae Gh. Popescu (1974) 1975; var. Viola elatior; var. Ga-

lium mollugo Danciu 1974; fac. trifoliosum molinieri; fac.

trifoliosum pallidi; fac. holcosum lanati; Ukraine: subass.

typicum with two variants – var. typicum, var. Agrostis

stolonifera; subass. anthoxanthetosum odorati Kuzemko

2009; subass. holcetosum lanati Kuzemko 2009; subass.

koelerietosum delavignei Kuzemko 2009.

Ass. Astrantio major-Arrhenatheretum elatioris Gi-

unochet 1939

Distribution: France.

Ass. Poëto angustifoliae-Agrostidetum capillaris Soό

& Csűrös 1947

Remarks: for details see in alliance Cynosurion.

Ass. Alchemillo-Trisetetum Horvat 1951 (Alchemilla

sp. div., Trisetum flavescens)

Diagnostic species: Alchemilla sp. div., Poa pratensis,

Trisetum flavescens.

Distribution: Croatia, Montenegro.

Ass. Arrhenatherо-Brometum erecti Balázs 1951

Diagnostic species: Arrhenatherum elatius, Bromus

erectus, Coronilla varia, Dianthus cathusianorum,

Salvia pratensis, Trifolium montanum.

Distribution: Hungary.

Annex 1. Continuation.

106 Velev, N. • Checklist of Arrhenatheretalia order

Ass. Festuco rubrae-Agrostetum capillaris Horvat

1951

Remarks: for details see in alliance Cynosurion.

Ass. Polygono bistortae-Arrhenatheretum elatioris

Oberdorfer 1952

Distribution: France.

Ass. Dactylido glomeratae-Festucetum arundinace-

ae Tüxen ex W. Lohmeyer 1953

Distribution: France.

Ass. Festucetum rubrae (Domin 1923) Válek 1956

Diagnostic species: Festuca rubra.

Distribution: Belarus, Ukraine.

Variations: Belarus: subass. typicum; subass. agros-

tidetosum tenuis sensu Stepanovich 2000; subass. anth-

oxanthetosum odorati sensu Stepanovich 2000; subass.

brizetosum mediae sensu Stepanovich 2000; subass.

brometosum mollis sensu Stepanovich 2000; subass.

cynosuretosum cristati sensu Stepanovich 2000; sub-

ass. dactylidetosum glomeratae sensu Stepanovich

2000; subass. deschampsietosum cespitosae sensu

Stepanovich 2000; subass. equisetetosum arvensi sen-

su Stepanovich 2000; subass. festucetosum pratensis

sensu Stepanovich 2000; subass. poetosum angustifoli-

ae sensu Stepanovich 2000; subass. rumecetosum thyr-

siflori sensu Stepanovich 2000; Ukraine: subass. typi-

cum; subass. brometosum mollis Kuzemko 2009 with

two variants – var. typicum, var. Koeleria delavignei;

subass. caricetosum flavae Kuzemko 2009; subass. fes-

tucetosum orientalis Kuzemko 2009.

Ass. Trifolieto-Alopecuretum pratensis Cincović

1956 (Trifolium pallidum)

Synonyms: Trifolium pallidum-Alopecurus praten-

sis Cincović 1956.

Diagnostic species: Alopecurus pratensis, Clematis

integrifolia, Poa pratensis, Ranunculus polyanthemos,

Trifolium pallidum, T. patens.

Distribution: Serbia.

Variations: subass. typicum; subass. caricetosum

Cincović 1956 (Carex praecox); subass. rhinantheto-

sum rumelici Cincović 1959; subass. trifolietosum hy-

bridi Cincović 1959.

Ass. Galio veri-Trifolietum repentis Sougnez 1957

Remarks: for details see in alliance Cynosurion.

Ass. Ophioglosso vulgati-Arrhenatheretum elatioris

P. Montserrat 1957

Remarks: According to Rodríguez-Rojo & al.

(2014) incl.: Odontito serotini-Trifolietum pratensis

O. Bolós & Masalles 1983, Rhinantho mediterranei-

Trisetetum flavescentis Vigo 1984, Tragopogono praten-

sis-Lolietum multiflori P. Montserrat 1957.

Distribution: Iberian Peninsula.

Ass. Poo-Trisetetum flavescentis Knapp ex Oberdor-

fer 1957

Original name: Poa-Trisetetum Knapp 1951 (Poa

pratensis, P. t r iv iali s , Trisetum flavescens).

Synonyms: Trisetetum flavescentis (Stebler & Schrö-

ter) Brockmann-Jerosch 1907; Trisetum flavescens-Poa

pratensis Knapp 1951 (Art. 3b); Triseto flavescentis-Po-

etum pratensis Knapp 1951; Trifolio-Festucetum rubrae

Oberdorfer 1957 nom. prov. (Art. 3b); Phyteumato-Fes-

tucetum Passarge 1968; Phleo-Agrostietum Steinbuch

1980 (Art. 1); Hyperico-Trisetetum Hundt 1980; Poo-Tri-

setetum flavescentis Knapp 1951. According to Oberdor-

fer (1993) incl.: Trifolio-Festucetum rubrae Oberdorfer

1957, Trisetetum flavescentis Tüxen 1937.

Corresponding names: Plant community Poa prat-

ensis-Trisetum sensu Dierschke 1997.

Pseudonyms: Festuco-Cynosuretum cristati narde-

tosum sensu Jurko 1974 non Tüxen in Bükker 1942;

Luzulo-Cynosuretum cristati sensu Jurko 1974 non

Meisel 1961.

Remarks: According to Hájková & al. (2007) asso-

ciations Poo-Trisetetum flavescentis Knapp ex Ober-

dorfer 1957 and Trifolio-Festucetum rubrae Ober-

dorfer 1957 are synonyms. Sanda & Popescu (1991)

accept these as different syntax and relate them to

different alliances – Poo-Trisetetum flavescentis in

Polygono-Trisetion and Trifolio-Festucetum rubrae in

Cynosurion. Krahulec & al. (1997) also accept the two

associations as separate and refer them to Arrhenathe-

rion elatioris.

Diagnostic species: Agrostis capillaris, Alchemil-

la sp. div., Anthoxanthum odoratum agg., Campanula

patula, Cardaminopsis halleri, Carum carvi, Centaurea

melanocalathia, Chaerophyllum aureum, Leucanthe-

mum vulgare agg. (L. ircutianum, L. vulgare), Persi-

caria bistorta (=Polygonum bistorta), Phleum pratense,

Poa pratensis, Rhinanthus glacialis, Rumex obtusifo-

lius, Trisetum flavescens, Veronica serpyllifolia.

Distribution: Austria [in Phyteumo-Trisetion (Pas-

sarge 1969) Ellmauer & Mucina 1993], Czech Re-

public, Germany, Hungary, Romania [according to

Dihoru (1975) and Sanda & Popescu (1991) in Polygo-

no-Trisetion], Slovakia, Ukraine.

Variations: Czech Republic: subass. poetosum triv-

ialis Blažková 1991; var. Arrhenatherum elatius; var.

Annex 1. Continuation.

107Phytol. Balcan. 24(1) • Sofia • 2018

Hypericum maculatum; var. Sanguisorba officinalis;

Romania: fac. cardaminapsosum halleri; fac. chry-

santemosum leucanthemi; Slovakia: var. Betonica of-

ficinalis; var. Hypericum maculatum; var. Sanguisor-

ba officinalis var. Trisetum flavescens; Ukraine: subass.

anthoxanthetosum odorati Kuzemko 2009; subass.

galietosum mollugo Kuzemko 2009.

Ass. Tragopogono orientalis-Lolietum multiflori P.

Montserrat 1957

Distribution: Iberian Peninsula.

Ass. Trifolio-Festucetum rubrae Oberdorfer 1957

Remarks: for details see in alliance Cynosurion.

Ass. Bromo racemosi-Cynosuretum cristati Horvatić

(1930) 1958

Synonyms: Cynosuretum cristati Horvatić 1930,

Bromus racemosus-Cynosurus cristatus Horvatić 1941,

Cynosuretum cristati Zeidler 1954, Cynosutetum cris-

tati Kojić & al. 2003 s.l.

Diagnostic species: Alopecurus utriculatus (=A.

rendlei), Cynosurus cristatus, Gaudinia fragilis, Hor-

deum secalinum, Ononis arvensis, Ophioglossum vulg-

atum, Poa trivialis, Ranunculus acris, Rhinanthus mi-

nor, Trifolium patens.

Distribution: Bulgaria (in Cynosurion), Croatia,

Montenegro, Serbia, Slovenia.

Variations: Croatia: subass. typicum; subass. hol-

cetosum lanati Horvatić 1930; subass. brometum rac-

emosi Horvatić 1963; subass. hypochoeridetosum

radicatae Hruška-Dell'Uomo 1976; subass. danthoni-

etosum Šegulja 1977; Montenegro: subass. holcetosum

lanati Horvatić 1930; Serbia: subass. typicum; subass.

brometum racemosi sensu Cincović 1956.

Ass. Colchico autumnalis-Festucetum pratensis Du-

vigneaud 1958

Distribution: Belgium, France.

Variations: Belgium and France: subass. typicum;

subass. heracleetosum sphondylii Trivaudey 1995; sub-

ass. stachyetosum officinalis Trivaudey 1995.

Ass. Lino angustifolii-Cynosuretum cristati Allorge

ex Oberdorfer & Tüxen in Tüxen & Oberdorfer 1958

Remarks: for details see in alliance Cynosurion.

Ass. Rumiceto-Agrostidetum caninae Cincović 1959

(Rumex acetosella)

Diagnostic species: Agrostis canina, Aira capillaris,

Rumex acetosella, Trifolium filiforme, Vulpia myuros.

Distribution: Serbia.

Ass. Rhinantho-Cynosuretum cristati Blečić & Tatić

1960

Original name: Alectorolopho-Cynosuretum crista-

ti Blečić & Tatić 1960 (Alectorolophus rumelicus = Rhi-

nanthus rumelicus).

Diagnostic species: Cynosurus cristatus, Ophioglos-

sum vulgatum, Poa trivialis, Rhinanthus rumelicus.

Distribution: Serbia.

Variations: subass. lupulinetosum Blečić & Tatić

1960 (Medicago lupulina); subass. auricometosum

Blečić & Tatić 1960 (Ranunculus auricomus).

Ass. Alchemillo xanthochlorae-Arrhenatheretum

elatioris (Oberdorfer 1957) Sougnez & Limbourg

1963

Diagnostic species: Alchemilla xanthochlora (A.

vulgaris agg.), Cardamine pratensis, Colchicum autum-

nale, Crepis biennis, Knautia arvensis, Luzula campes-

tris, Pimpinella major.

Distribution: Belgium, France, Slovakia.

Variations: Belgium: subass. typicum; subass. crep-

idetosum biennis (Trivaudey 1995) Trivaudey in Fer-

rez 2007; subass. knautietosum dipsacifolia (Trivaudey

1995) Trivaudey in Ferrez 2007; France: subass. typi-

cum; subass. agrostietosum capillaris (Trivaudey 1995)

Trivaudey in Ferrez 2007; subass. crepidetosum bien-

nis (Trivaudey 1995) Trivaudey in Ferrez 2007; subass.

knautietosum dipsacifolia (Trivaudey 1995) Trivaudey

in Ferrez 2007.

Ass. Pastinaco sativae-Arrhenatheretum elatioris

Passarge 1964

Original name: Pastinaco-Arrhenatheretum

(Knapp 1954) Passarge 1964 (Pastinaca sativa; Ar-

rhenatherum elatius).

Synonyms: Arrhenatheretum elatioris Br.(-Bl.) 1915

nom. ambig. (Art. 36); Arrhenatheretum elatioris Br.-Bl.

1919 sensu lato; Arrhenatheretum elatioris Br.-Bl. 1919

ap. Br.-Bl. 1948; 1949; Alopecuretum pratensis Regel

1925 nom. ambig. (Art. 36); Arrhenatheretum elatioris

Br.-Bl. ex Scherrer 1925 (Art. 31); Arrhenatheretum ela-

tioris (Br.-Bl. 1925) Koch 1926; Gaudinio-Arrhenathere-

tum Br.-Bl. 1931 (Art. 29); Аrrhentheretum еlatioris

Тüxen 1937; Arrhenatheretum medioeuropaeum (Br.-

Bl. 1919) Oberdorfer 1952 nom. illeg. (Art. 34a); Ar-

rhenatheretum medioeuropaeum Oberdorfer 1952 ap.

Oberdorfer 1957 nom. illeg. (Art. 34a); Arrhenathere-

tum subatlanticum Tüxen 1955 nom. illeg. (Art. 34a);

Centaureo nigrae-Arrhenatheretum Oberdorfer (1952)

1957; Melandrio-Arrhenatheretum Oberdorfer 1957;

Alchemillo vulgaris-Arrhenatheretum (Oberdorfer

1957) Sougnez & Limbourg 1963; Agrostio-Festucetum

Annex 1. Continuation.

108 Velev, N. • Checklist of Arrhenatheretalia order

rubrae Stockhammer 1964 (Art. 7); Polygalo-Festuce-

tum rubrae Passarge 1964; Heracleo-Arrhenatheretum

Passarge 1964; Pastinaco-Arrhenatheretum Passarge

1964; Alchemillo-Arrhenatheretum (Oberdorfer 1957)

Görs 1966 (Art. 29a); Pastinaco-Arrhenatheretum Pas-

sarge 1964; Dauco carotae-Arrhenatheretum Görs 1966;

1974 (Art. 29c); Campanulo-Lathyretum pratensis Pas-

sarge 1975; Vicio-Galietum molluginis Passarge 1975;

Dauco-Vicietum hirsutae Passarge 1975 p. max. p.;

Chrysanthemo-Rumicietum thyrsiflori Walther 1977 p.

max. p.; Lolio-Arrhenatheretum Dietl 1982 (Art. 7); Fili-

pendulo vulgaris-Arrhenatheretum Hundt & Hübl 1983

p. p.; Galio albi-Geranietum pratensis Passarge 1984 p.

p.; Tanaceto-Arrhenatheretum A. Fischer 1985 p. max.

p. (Art. 5); Tanaceto-Arrhenatheretum A. Fischer ex Ell-

mauer in Ellmauer & Mucina 1993; Centaureo scabios-

ae-Arrhenatheretum (Fartmann 1997) Passarge 1999 p.

max. p.; Filipendulo vulgaris-Ranunculetum polyanthe-

mi Hundt (1954) 1958 sensu Passarge 1999; Galio al-

bi-Alopecuretum pratensis (Steffen 1931) Hundt 1958

sensu Passarge 1999 (Art. 29a); Ranunculo repentis-Al-

opecuretum pratensis Krisch 1974 sensu Passarge 1999

p. p.; Plantagini lanceolatae-Festucetum rubrae Scamoni

sensu Passarge 1999 p. max. p.; Achilleo millefolii-Lath-

yretum nissoliae Passarge 2002.

According to Päzolt & Jansen (2004) incl.: Plant

community Artemisia vulgaris-Arrhenatherum ela-

tius sensu Dierschke 1997 p. p., Plant community Leu-

canthemum-Rumex thyrsiflorus sensu Dierschke 1997,

Plant community Phalaris-Alopecurus pratensis sensu

Passarge 1999, Plant community Festuca rubra-Agros-

tis tenuis sensu Dierschke 1997 p. p., Plant community

Galium album-Alopecurus pratensis sensu Rennwald

2002; excl.: Pastinaco-Arrhenatheretum peucedane-

tosum oreoselini Dengler 1994 p. max. p., Ranunculo

repentis-Alopecuretum pratensis phalaridetosum Pas-

sarge 1999. According to Ellmauer & Mucina (1993)

incl.: Arrhenatheretum elatioris festucetosum rubrae

Sturm 1978 (Art. 1); non: Arrhenatheretum elatixici-

fooris Braun 1915.

Corresponding names: Arrhenatheretum elatioris

Br.-Bl. 1919 subass. medioeuropaeum Horvatić 1941

ap. Horvatić-Tomažič 1941.

Remarks: 1) According to Päzolt & Jansen (2004)

Ranunculo bulbosi-Arrhenatheretum elatioris Ellmau-

er in Mucina & al. 1993 and Arrhenatheretum elati-

oris Br.(-Bl.) 1915 [Syntax. syn.: Pastinaco sativae-Ar-

rhenatheretum elatioris Passarge 1964] are synonyms,

but Vicherek & al. 2000, Hájková & al. 2007, Uhliarová

& al. 2007 and Stančić 2008a) consider these two asso-

ciationa as different syntaxa; 2) Hájková (2007), Hájk-

ová & al. 2007 and Jarolímek & al. (2008b) pointed Al-

opecuretum pratensis Regel 1925 as synonym of Poo

trivialis-Alopecuretum pratensis Regel 1925 (Deschamp-

sion). According to Päzolt & Jansen (2004) Alopecure-

tum pratensis Regel 1925 is syntaxonomical synonym

of Arrhenatheretum elatioris Br.(-Bl.) 1915 (Syn.: Past-

inaco sativae-Arrhenatheretum elatioris Passarge 1964);

3) Kovács (2004) pointed Alopecuretum pratensis Regel

1925 s. l. as synonym of Carici vulpinae-Alopecuretum

pratensis (Máthé & Kovács 1967) Soό 1971 corr. Borhi-

di 1996 [Syn.: Alopecuretum pratensis Nowinski 1928

(Art. 36); Carici-Alopecuretum pratensis Soό 1971 (Art.

3); Ranunculo repentis-Alopecuretum pratensis Ellmau-

er 1993 p. p.] and refer all these syntaxa to Deschamp-

sion caespitosae Horvatić 1930.

Diagnostic species: Achillea millefolium agg., Agros-

tis capillaris, Anthoxanthum odoratum, Arrhenatherum

elatius, Avenula pubescens, Campanula patula, Carex

sylvatica, Cerastium holosteoides, Crepis biennis, Dac-

tylis glomerata, Daucus carota, Elymus repens, Festuca

pratensis, F. rubra agg., Galium mollugo agg., Geranium

pratense, Holcus lanatus, Knautia arvensis, Leontodon

hispidus, Leucanthemum vulgare agg., Lotus cornicu-

latus, Pastinaca sativa, Peucedanum carvifolia, Planta-

go altissima, P. lanceolata, Poa pratensis agg., Ranuncu-

lus acris, Rumex acetosa, R. thyrsiflorus, Taraxum Sect.

Ruderalia, Tragopogon dubius, T. orientalis, T. pratensis,

Trifolium pratense, T. repens, Trisetum flavescens.

Distribution: Austria, Belgium, Bosnia and Her-

zegovina, Bulgaria, Croatia, Czech Republic, France,

Germany, Hungary, Lithuania (in Alopecurion praten-

sis Passarge 1964), Montenegro, Netherlands, Poland,

Portugal, Romania, Serbia, Slovakia, Slovenia, Spain,

Switzerland, Ukraine, United Kingdom.

Variations: Austria: subass. typicum Knapp 1954;

subass. alchemilletosum (Sougnez & Limbourg 1963)

Ellmauer 1995; Bulgaria: var. Central Balkan Geograph-

ic Variant; Croatia: subass. typicum sensu Ellenberg

1996; subass. hircinetosum Horvatić (1956) 1958 (Syn.:

Аrrhentheretum еlatioris var. orientalis Horvatić 1941);

subass. litorale Horvat 1961; Czech Republic: subass.

typicum sensu Oberdorfer 1952; subass. cirsietosum

cani (Vicherek 1962) Vicherek in Vicherek & al. 2000

(Orig. name: Arrhenatheretum elatioris subass. Cirsium

canum Vicherek 1962); France: subass. typicum; subass.

Annex 1. Continuation.

109Phytol. Balcan. 24(1) • Sofia • 2018

brometosum erecti Oberdorfer 1936; Poland: subass.

typicum sensu Oberdorfer 1952; subass. alopecureto-

sum pratensis Tüxen 1937; var. typicum; var. Alopecurus

pratensis; var. Armeria maritima; var. Dactylis glomera-

ta; var. Deschampsia caespitosa; var. Poa trivialis; Roma-

nia: subass. typicum sensu Oberdorfer 1952; subass. tri-

setosum flavescentis Horvatič 1930; subass. brometosum

erecti Oberdorfer 1936; subass. alopecuretosum prat-

ensis Tüxen 1937; subass. festucetosum pratensis (Soό

1940) Slavnić 1948; subass. festucetosum rubrae Tüxen

1951 (Corresp. name: Arrhenathero-Festucetum rubrae

Resmeriţă 1958; 1963); subass. festucetosum rupicolae

Eggler 1958 (festucetosum sulcatae Eggler 1958; Festu-

ca rupicola = F. sulcata) [Corresp. name: Arrhenathero-

Festucetum rupicolae Soό apud Borza 1963; Resmeriţă

1958 (Arrhenathero-Festucetum sulcatae Soό apud Bor-

za 1963; Resmeriţă 1958)]; subass. moenchietosum

manticae Borza & Boşcaiu 1960; 1963; subass. holceto-

sum lanati Csűrös 1970; fac. festucosum pratensis; fac.

dactyliosum glomeratae; fac. poosum pratensis; fac. tri-

foliosum pratensis; Serbia: subass. festucetosum arun-

dinaceae Ilijanić & Šegulja 1978; subass. salvietosum

nemorosae Ilijanić & Šegulja 1978; Slovenia: subass.

typicum sensu Passarge 1964; subass. medicagetosum

lupulinae Passarge 1964; subass. lolietosum Petras Sackl

& al. 2012; Spain: subass. geranietosum dissecti O. Bolòs

1959; subass. arrhenatheretosum O. Bolòs 1983; subass.

brometosum erecti O. Bolòs 1983; subass. holcetosum la-

nati O. Bоlòs 1983; Ukraine: subass. typicum with var.

Mentha arvensis; subass. galietosum mollugo Kuzemko

2009 with two variants – var. typicum, var. Festuca ovi-

na; subass. trifolietosum dubii Kuzemko 2009.

There are two homonyms within the varia-

tions – Arrhenatheretum elatioris Br.-Bl. ex Scherrer

1925 subass. typicum sensu Oberdorfer 1952 and Ar-

rhenatheretum elatioris Br.(-Bl.) 1915 subass. typicum

sensu Ellenberg 1996.

Ass. Trifolio-Cynosuretum cristati Veljović 1967

(Trifolium patens)

Diagnostic species: Cynosurus cristatus, Festuca

pratensis, Potentilla reptans, Rhinanthus minor, Tar ax-

acum officinale, Trifolium patens.

Distribution: Serbia.

Ass. Filipendulo-Arrhenatheretum Shimwell 1968

Distribution: United Kingdom.

Ass. Cichorietum intybi Tüxen ex Sissingh 1969

Synonyms: Cichorietum intybi Tüxen 1942 (Art.

1). According to Dengler & Wollert (2004) incl.: plant

community Artemisia vulgaris sensu Wollert 1991 p.

p., plant community Cichorium intybus sensu Pott

1995, Plantagini-Lolietum perennis cichorietosum

sensu Sýkora & al. 1996 p. max. p., plant communi-

ty Cichorium intybus sensu Weeda & Scaminée 1998.

Diagnostic species: Cichorium intybus (usually as

dominant).

Distribution: Austria, Germany (in Dauco carotae-

Melilotion Görs ex Rostański & Gutte 1971), Roma-

nia, Slovenia.

Ass. Rhinantho-Filipenduletum Ilijanić 1969

Diagnostic species: Filipendula vulgaris, Fragaria

viridis, Rhinanthus rumelicus.

Distribution: Croatia.

Ass. Agrostio castellanae-Arrhenatheretum bulbosi

Teles 1970

Distribution: Iberian Peninsula.

Ass. Trifolio-Armerietum canescentis K. Tomić 1970

Distribution: Montenegro.

Ass. Alopecuro-Arrhenatheretum (Máthe & Kovács

1960) Soό 1971

Synonyms: Arrhenatheretum elatioris alopecureto-

sum Tüxen 1937.

Diagnostic species: Alchemilla vulgaris agg., Ar-

rhenatherum elatius, Alopecurus pratensis, Geranium

pratense, Sanguisorba officinalis.

Distribution: Hungary, Romania.

Ass. Anthyllidi-Trifolietum montani Pawłowski 1971

Distribution: Ukraine.

Ass. Anthyllido vulnerariae-Festucetum rubrae

(Máthe & Kovács 1960) Soό 1971

Remarks: for details see in alliance Cynosurion.

Ass. Armerio-Trisetetum flavescentis (Feri 1974)

Rexhepi 1974

Distribution: Serbia.

Ass. Lolietum multiflorae Dietl & Lehmann 1975

Diagnostic species: Achillea millefolium, Lolium

multiflorum (обикновено като доминант), Planta-

go lanceolata, Poa trivialis, Ranunculus acris, R. re-

pens, Rumex acetosa, Taraxacum officinale, Trifolium

repens.

Distribution: Austria, Croatia, Slovenia, Switzerland.

Ass. Phyteumato orbicularis‐Arrhenatheretum elati-

oris Royer 1975

Diagnostic species: Colchicum autumnale, Narcis-

sus poeticus, Dichloropetalum carvifolia, Phyteuma or-

biculare.

Distribution: France.

Annex 1. Continuation.

110 Velev, N. • Checklist of Arrhenatheretalia order

Ass. Pimpinello majoris-Arrhenatheretum elatioris

Richard 1975

Distribution: France.

Ass. Trifolio-Poëtum pratensis (Răvărut & al. 1956)

Resmeriţă 1975 (Trifolium pratense)

Original name: Poëtum pratensis Răvărut & al. 1956.

Diagnostic species: Poa pratenisis, Trifolium prat-

ense.

Distribution: Romania (also in Agrostion stolonif-

erae Soό (1933) 1971, Molinietalia caeruleae Koch

1926).

Ass. Cynosuro-Agrostidetum albae Mišić & al. 1978

nom. prov. et illeg. (Art. 3b)

Diagnostic species: Agrostis capillaris, Cynosurus

cristatus.

Distribution: Serbia.

Ass. Potentillo albae-Festucetum rubrae Blažková

1979

Diagnostic species: Avenula pubescens, Campanu-

la patula, Campanula rotundifolia agg. (C. gentilis, C.

moravica, C. rotundifolia), Erophila verna, Knautia ar-

vensis agg. (K. arvensis, K. kitaibelii), Leontodon his-

pidus, Leucanthemum vulgare agg. (L. ircutianum, L.

vulgare), Peucedanum oreoselinum, Potentilla alba, Ra-

nunculus bulbosus, Rumex thyrsiflorus, Saxifraga gran-

ulata, Thlaspi caerulescens, Trifolium dubium, T. prat -

ense, Trisetum flavescens.

Distribution: Czech Republic.

Variations: var. Festuca pratensis (Corresp. name:

Potentillo albae-Festucetum rubrae subass. festuceto-

sum pratensis Blažková 1979); var. Thymus pulegioides

(Corresp. name: Potentillo albae-Festucetum rubrae

subass. thymetosum Blažková 1979).

Ass. Anthoxantho-Brometum erecti Poldini 1980

Corresponding names: Arrhenatheretum elatioris

subass. brometosum erecti Poldini 1989.

Pseudonyms: Arrhenatheretum elatioris Poldini

1985 non Scherrer 1925.

Distribution: Italy, Croatia, Slovenia.

Variations: Croatia: var. Chrysopogoni gryllus.

Ass. Luzulo campestris-Brometum mollis de Fou-

cault 1980

Distribution: France.

Ass. Festuco rubrae-Cynosuretum cristati Petković

1981

Diagnostic species: Festuca rubra ssp. fallax (=F.

nigrescens), Cynosurus cristatus, Carum carvi, Bromus

racemosus, Lathyrus pannonicus, Poa trivialis.

Distribution: Serbia.

Variations: subass. typicum; subass. knautietosum

dinarici Petković & Tatić 1987.

Ass. Phelipaeo coeruleae-Arrhenatheretum elatioris

J.-M. & J. Gèhu 1982

Distribution: France.

Ass. Sileno montani-Arrhenatheretum elatioris J.-M.

& J. Gèhu 1982

Distribution: France.

Ass. Vicio sepium‐Arrhenatheretum O’Sullivan 1982

Diagnostic species: Centaurea nigra, Heracleum

sphondylium, Trisetum flavescens, Vicia sepium.

Distribution: Ireland.

Ass. Odontito serotini-Trifolietum pratensis O. Bolòs

& Masalles 1983

Distribution: Iberian Peninsula.

Variations: Spain: subass. typicum (Syn.: Odontito

serotini-Trifolietum pratensis O. Bolòs & Masalles 1983

subass. trifolietosum pratensis); subass. arrhenathere-

tosum elatioris O. Bolòs & Masalles 1983.

Ass. Ononido-Arrhenatheretum elatioris (Tomažič

& Horvatić 1941) Ilijanić & Šegulja (1978) 1983

Distribution: Croatia, Serbia, Slovenia.

Variations: Serbia: subass. cirsietosum cani Butorac

1989 nom. inval. (Art. 1); subass. eryngietosum camp-

estre Butorac 1989 nom. inval. (Art. 1).

Ass. Rhinantho mediterranei-Trisetetum flavescen-

tis Vigo 1984

Synonyms: Gentiano luteae-Trisetetum flavescentis

Vigo 1984.

Remarks: According to Ninot & al. (2000) these as-

sociations are separate syntaxa, not synonyms.

Diagnostic species: Crepis vesicaria subsp. taraxaci-

folia, Knautia nevadensis, Rhinanthus mediterraneus.

Distribution: Iberian Peninsula.

Variations: Spain: subass. typicum; subass. salvieto-

sum pratensis Vigo ex Vigo in Ninot & al. 2000 (Bas-

ionym: Rhinantho mediterranei-Trisetetum flavescen-

tis Vigo 1984 subass. salvietosum pratensis Vigo 1984

nom. inval.); subass. trifolietosum campestris Vigo ex

Vigo in Ninot & al. 2000 (Basionym: Rhinantho med-

iterranei-Trisetetum flavescentis Vigo 1984 subass. tri-

folietosum campestris Vigo 1984 nom. inval.).

Ass. Calamagrostietum epigeii Sapegin 1986 (Art. 31)

Homonyms: Calamagrostietum epigeji Juraszek

1928 (pointed for Poland in Epilobietea angustifolii

Tüxen & Preising 1950).

Distribution: Belarus.

Annex 1. Continuation.

111Phytol. Balcan. 24(1) • Sofia • 2018

Ass. Centaureo nemoralis‐Festucetum arundinaceae

Gruber 1986

Diagnostic species: Centaurea nemoralis (C. de-

beauxii agg.), Gaudinia fragilis, Oenanthe pimpinel-

loides, Schedonorus arundinaceus (= Festuca arundi-

nacea).

Distribution: France.

Ass. Gaudinio-Festucetum pratensis Seliškar 1986

nom. nud.

Distribution: Slovenia.

Ass. Heracleo lecoquii-Arrhenatheretum elatioris de

Foucault 1986

Distribution: France.

Ass. Lino biennis-Brometum mollis de Foucault 1986

Distribution: France.

Ass. Oenantho pimpinelloidis-Linetum biennis de

Foucault 1986

Diagnostic species: Centaurea debauxii agg., Gaud-

inia fragilis, Linum bienne, Oenanthe pimpinelloides,

Silene flos‐cuculi, Trifolium patens.

Distribution: France.

Ass. Sieglingio-Festucetum rubrae Jovanović-Dunjć

& al. 1986

Diagnostic species: Festuca rubra, Sieglingia de-

cumbens.

Distribution: Serbia.

Ass. Narcisso-Arrhenatheretum elatioris Kriczfa-

lushij & Gendey 1987

Distribution: Ukraine.

Ass. Heracleo sphondylii-Brometum mollis de Fou-

cault 1989

Distribution: Belgium, France.

Ass. Hordeo secalini- Arrhenatheretum elatioris Fri-

leux & al. 1989

Distribution: France.

Ass. Onоnido-Alopecuretum pratensis Butorac 1989

nom. inval. (Art. 1)

Distribution: Serbia.

Variations: subass. ranunculetosum aceri Butorac

1989 nom. inval. (Art. 1); subass. ranunculetosum pol-

yanthemi Butorac 1989 nom. inval. (Art. 1).

Ass. Orchido morionis-Saxifragetum granulatae

Gaume ex de Foucault 1989

Distribution: France (in Brachypodio-Centaureion

nemoralis).

Ass. Alopecuro pratensis-Arrhenatheretum elatioris

(Tüxen 1937) Julve 1993 nom. inval. (Art. 3b)

Distribution: France.

Ass. Ranunculo bulbosi-Arrhenatheretum elatioris

Ellmauer in Mucina & al. 1993

Original name: Ranunculo bulbosi-Arrhenathere-

tum Ellmauer in Mucina & al. 1993 (Arrhenatherum

elatius).

Synonyms: Arrhenatheretum elatioris Br.(-Bl.)

1915 p. p.; Arrhenatheretum elatioris Scherer 1925 non

Braun (-Bl.) 1915 (Art. 31); Arrhenatheretum elatioris

subass. ranunculetosum bulbosi Knapp 1954; Agrosti-

do-Festucetum Stockhammer 1964 (Art. 31). Accord-

ing to Uhliarová & al. (2007) incl.: Arrhenatheretum

brometosum erecti Oberdorfer 1936, Arrhenathere-

tum ranunculetosum bulbosi Knapp & Knapp 1954,

Arrhenatheretum salvietosum pratensis Hundt 1958.

According to Ellmauer & Mucina (1993) incl.: Ar-

rhenatheretum brometosum erecti Oberdorfer 1936,

Arrhenatheretum ranunculetosum bulbosi Knapp &

Knapp 1954, Arrhenatheretum salvietosum pratensis

Hundt 1958, Arrhenatheretum mesodanubiale salvie-

tosum Stockhammer 1964 (Art. 3e; 34), Arrhenathere-

tum mesodanubiale brometosum Stockhammer 1964

(Art. 3e; 34).

Remarks: According to Päzolt & Jansen (2004)

Ranunculo bulbosi-Arrhenatheretum elatioris Ell-

mauer in Mucina & al. 1993 and Arrhenatheretum

elatioris Br.(-Bl.) 1915 [Syntax. syn.: Pastinaco sativae-

Arrhenatheretum elatioris Passarge 1964] are syntax-

onomical synonyms. Vicherek & al. 2000, Hájková &

al. 2007 and Uhliarová & al. 2007 accept these associ-

ations as different syntaxa.

Diagnostic species: Anthoxanthum odoratum, Ar-

rhenatherum elatius (usually as subdominant), Avenu -

la pubescens, Briza media, Bromus erectus, Campanula

patula, Carex caryophyllea, C. montana, Clinopodium

vulgare, Coronilla varia, Daucus carota, Dianthus car-

thusianorum agg., Festuca rubra agg., Filipendula vul-

garis, Leontodon hispidus, Leucanthemum vulgare agg.

(L. ircutianum, L. vulgare), Lotus corniculatus, Lychnis

viscaria, Pimpinella saxifraga, Plantago media, Ranun-

culus bulbosus, Salvia pratensis, Sanguisorba minor, Si-

lene vulgaris, Tragopogon orientalis, Trifolium monta-

num, Trisetum flavescens.

Distribution: Austria, Bulgaria, Croatia, Czech Re-

public, Germany, Hungary, Serbia, Slovakia, Slovenia.

Variations: Germany: var. typicum; var. Linum

catharticum; var. Campanula rotundifolia; Slovakia:

var. Daucus carota; var. Fragaria vesca; var. Steris vis-

caria.

Annex 1. Continuation.

112 Velev, N. • Checklist of Arrhenatheretalia order

Ass. Ranunculo repentis-Alopecuretum pratensis

(Eggler 1933) Ellmauer in Mucina & al. 1993

Synonyms: Alopecuretum Eggler 1958 (Art. 31); Lo-

lio perennis-Alopecuretum Dietl 1983 (Art. 5). Accord-

ing to Ellmauer & Mucina (1993) incl.: Arrhenathere-

tum alopecuretosum R. Tüxen 1937, Alopecuretum

holcetosum lanati Eggler 1959, Alopecuretum poetosum

angustifoliae Eggler 1959, Ranunculetum caricetosum

brizoidis Stockhammer 1964 (Art. 3e), Arrhenathere-

tum mesodanubiale alopecuretosum Stockhammer

1964 (Art. 3e; 34); non Alopecuretum pratensis Regel

1925, Alopecuretum pratensis Steffen 1931 (Art. 31),

Agropyro-Alopecuretum Moravec 1965.

Diagnostic species: Alopecurus pratensis, Car-

damine matthiolii, C. pratensis, Carex hirta, Cerastium

holosteoides, Festuca pratensis, Glechoma hederacea,

Holcus lanatus, Lolium perenne, Lychnis flos-cuculi,

Lysimachia nummularia, Myosotis palustris, Ranuncu-

lus repens, Rumex obtusifolius, Sanguisorba officinalis.

Remarks: Dierschke (1997) refer this association to the

plant community Ranunculus repens-Alopecurus pratensis.

Distribution: Austria, Bulgaria, Czech Republic,

Germany, Hungary, Romania, Slovakia (in Deschamp-

sion), Slovenia, Ukraine (in Alopecurion pratensis).

Variations: Austria: subass. typicum Ellmauer

1995; subass. anthryscetosum Ellmauer 1995; subass.

lychnietosum Ellmauer 1995; Bulgaria: var. typicum,

var. Eryngium campestre.

Ass. Tanaceto vulgaris-Arrhenatheretum elatioris

Fischer ex Ellmauer in Mucina & al. 1993

Synonyms: Tanaceto-Arrhenatheretum Fischer

1985 (Art. 5). According to Ellmauer & Mucina (1993)

incl.: Arrhenatheretum tanacetetosum Knapp 1946

(Art. 1), Arrhenatheretum tanacetetosum Knapp 1954,

plant community Arrhenatherum elatius-Artemisia

vulgaris sensu Bornkamm 1974 (Art. 3c).

Remarks: Based on this association as holotypus,

Rodríguez-Rojo & al. (2017) propose a new alliance

within Molinio-Arrhenatheretea class – Anthrisco syl-

vestris-Arrhenatherion elatioris Rodríguez-Rojo & al.

2017, which is associated with the semi-ruderal vege-

tation of the classes Artemisietea vulgaris and Epilobi-

etea angustifolii to some extent.

Diagnostic species: Artemisia vulgaris, Cichorium

intybus, Cirsium arvense, Convolvulus arvensis, Ely-

mus repens, Linaria vulgaris, Melilotus albus, M. offic-

inalis, Salvia nemorosa (Pannonicum), Silene latifolia

ssp. alba, Tanacetum vulgare.

Distribution: Austria, Bulgaria, France, Hungary.

Ass. Centaureo carniolicae-Arrhenatheretum elatio-

ris (Oberdorfer 1964) Poldini & Oriolo 1994

Original name: Centaurea dubiae-Arrhenathere-

tum elatioris Oberdorfer 1964 (Art. 43)

Pseudonyms: Arrhenatheretum elatioris Pedrot-

ti 1964 non Scherrer 1925; Arrhenatheretum elatioris

Buffa & al. 1989 non Scherrer 1925.

Distribution: Italy, Slovenia.

Ass. Malvo moschatae-Arrhenatheretum bulbosi Tüx-

en & Oberdorfer 1958 corr. T.E. Díaz & F. Prieto 1994

Original name: Malvo moschatae-Arrhenatheretum

elatioris Tüxen & Oberdorfer 1958 (Art. 43).

Synonyms: Gaudinio fragilis-Festucetum pratensis

Br.-Bl. 1967.

Diagnostic species: Crepis capillaris, Malva mos-

chata, Rhinanthus angustifolius.

Distribution: Iberian peninsula, France.

Variations: Spain: subass. typicum sensu Tx. &

Oberd. 1958, Puente & Tascon 1981; subass. Amthyllis

dillenii Tx. & Oberd. 1958; subass. Avena sulcata Tx. &

Oberd. 1958; subass. Centaurea seussana Tx. & Oberd.

1958; subass. Polygonum bistorta Tx. & Oberd. 1958.

Ass. Poo sylvicolae-Lolietum multiflori Poldini &

Oriolo 1994

Distribution: Italy.

Ass. Malvo moschatae-Brometum mollis (Baranger

1978) Julve 1994 nom. ined. (Art. 1)

Distribution: France (in Brachypodio-Centaureion

nemoralis).

Ass. Filipendulo vulgaris-Arrhenatheretum Hundt

& Hübl ex Ellmauer 1995

Synonyms: Filipendulo vulgaris-Arrhenatheretum

Hundt & Hübl 1983.

Diagnostic species: Brachypodium pinnatum,

Bromopsis erecta, Buphthalmum salicifolium, Cam-

panula patula, Centaurea jacea s.l., Filipendula vulgar-

is, Galium boreale, Primula veris.

Distribution: Austria, Croatia, Hungary, Slovenia.

Ass. Silao silae-Colchicetum autumnalis de Foucault

1996 nom. inval. (Art. 3b)

Distribution: France.

Ass. Brizetum mediae Stepanovič 1999

Diagnostic species: Briza media.

Distribution: Belarus.

Variations: subass. typicum; subass. agrostidetosum

tenuis sensu Stepanovich 2000; subass. anthoxanthe-

tosum odorati sensu Stepanovich 2000; subass. cyno-

Annex 1. Continuation.

113Phytol. Balcan. 24(1) • Sofia • 2018

suretosum cristati sensu Stepanovich 2000; subass. fes-

tucetosum pratensis sensu Stepanovich 2000; subass.

festucetosum rubrae sensu Stepanovich 2000.

Ass. Brometum mollis Stepanovič 1999

Diagnostic species: Bromus mollis, Cardaminopsis

arenosa.

Distribution: Belarus.

Variations: subass. agrostidetosum tenuis sensu

Stepanovich 2000; subass. cynosuretosum cristati sen-

su Stepanovich 2000; subass. dactylidetosum glomera-

tae sensu Stepanovich 2000; subass. festucetosum prat-

ensis sensu Stepanovich 2000; subass. festucetosum

rubrae sensu Stepanovich 2000; subass. poetosum prat-

ensis sensu Stepanovich 2000.

Ass. Campanulo serratae-Agrostietum capillaris

Denisiuk & Korzeniak 1999

Distribution: Ukraine.

Ass. Helictotrichonetum pubescentis Stepanovič

1999

(Incl. Avenastreto-Nardetum Ružička 1955)

Diagnostic species: Helictotrichon pubescens.

Distribution: Belarus.

Variations: subass. anthoxanthetosum odorati sen-

su Stepanovich 2000; subass. brizetosum mediae sen-

su Stepanovich 2000; subass. festucetosum rubrae sen-

su Stepanovich 2000.

Ass. Centaureo fritschii-Arrhenatheretum Trinajstić

2000

Distribution: Croatia.

Ass. Cerastio holosteoides-Trisetetum flavescentis

Sanda & al. 2001

Pseudonyms: Poo-Trisetetum flavescentis auct. ro-

man.

Distribution: Romania.

Ass. Galio veri-Arrhenatheretum bulbosi (Rivas Go-

day & Borja 1961) Rivas-Martínez & al. 2001

Synonyms: Galio veri-Arrhenatheretum gudaricum

Rivas Goday & Borja 1961 nom. illeg. (Art. 34a; 39).

Distribution: Iberian Peninsula.

Variations: Spain: subass. anthoxanthetosum odo-

rati (O. Bolòs 1967) Bolòs in Ninot & al. 2000 [Syn.:

Galio veri-Arrhenatheretum gudaricum Rivas Go-

day & Borja 1961 subass. phleo-anthoxanthetosum O.

Bolòs 1967 nom. illeg. (Art. 34c)].

Ass. Lilio bulbiferi-Arrhenatheretum elatioris Ružič-

kova 2002

Diagnostic species: Allium oleraceum, Anthyl-

lis vulneraria, Aquilegia vulgaris, Arrhenatherum ela-

tius, Bromus monocladus, Carlina acaulis, Campanula

glomerata agg., Campanula serrata, Colchicum autum-

nale, Crepis alpestris, Dianthus carthusianorum agg.,

Gentiana cruciata, Jacea phrygia agg., Lilium bulbifer-

um, Plantago media, Potentilla thuringiaca, Primula

vulgaris, Rhinanrhus serotinus, Sanduisorba minor, Si-

lene nemoralis, S. vulgaris, Tragopogon orientalis, Tr i-

folium montanum, Trisetum flavescens.

Distribution: Slovakia.

Ass. Anthyllidi-Trifolietum montani Matuszkiewicz

2002

Distribution: Ukraine.

Ass. Anthyllidi-Trifolietum montani sensu Snowar-

ski 2002-2005

Diagnostic species: Anthyllis vulneraria; Trifolium

montanum.

Distribution: Poland.

Ass. Arrhenatheretum elatioris Kojić & al. 2003 sen-

su lato (non Br.-Bl. 1925)

Distribution: Serbia.

Variations: subass. arrhenatheretosum medioeuro-

paeum Kojić & al. 2003 (=Ass. Arrhenatheretum medi-

oeuropaeum Oberdorfer 1952); subass. clematetosum

integrifoliae Kojić & al. 2003 (=Ass. Arrhenatheretum

medioeuropaeum Oberdorfer 1952 subass. clemateto-

sum integrifoliae M. Vučk. 1991); subass. cisietosum

cani Kojić & al. 2003 [=Ass. Ononido arvensis- Ar-

rhenatheretum elatioris (Horvatić 1956) Ilijan-Šegulja

(1978) 1980 subass. cisietosum cani Butorac 1992];

subass. eryngietosum campestre Kojić & al. 2003 [=Ass.

Ononido arvensis- Arrhenatheretum elatioris (Horvatić

1956) Ilijan-Šegulja (1978) 1980 subass. eryngietosum

campestre Butorac 1992].

Ass. Cynosuretum cristati Kojić & al. 2003 sensu lato

Synonyms: Bromo-Cynosuretum cristati Horvatić

1930.

Distribution: Serbia.

Variations: subass. brometosum racemosi Kojić &

al. 2003 (=Ass. Bromo racemosi-Cynosuretum cristati

Horvatić 1930 subass. brometosum racemosi Horvatić

1930); subass. rhinathetosum angustifoliae Kojić & al.

2003 (=Ass. Rhinantho angustifoli-Cynosuretum cris-

tati Blečić & Tatić 1960); subass. trifolietosum paten-

si Kojić & al. 2003 (=Ass. Trifolio patensi-Cynosuretum

cristati Veljović 1967); subass. festucetosum rubrae

Kojić & al. 2003 (=Ass. Festuco rubrae-Cynosure-

tum cristati Petk. 1981); subass. knautietosum dinar-

ici Kojić & al. 2003 (=Ass. Festuco rubrae-Cynosure-

Annex 1. Continuation.

114 Velev, N. • Checklist of Arrhenatheretalia order

tum cristati Petk. 1981 subass. knautietosum dinarici

Petk. 1981); subass. pancicietosum serbicae Kojić & al.

2003 (=Ass. Festuco rubrae-Cynosuretum cristati Petk.

1981 subass. pancicietosum serbicae Gajić 1989); sub-

ass. poetosum trivialis Kojić & al. 2003 (=Ass. Poo triv-

ialis-Cynosuretum cristati R. Jov. 1955); subass. poten-

tilletosum erectae Kojić & al. 2003 (=Ass. Potentillo

erectae-Cynosuretum cristati Matović 1986); subass.

agrostietosum albae Kojić & al. 2003 (=Ass. Cynosuro

cristati-Agrostietum albae Danon-Blaž. 1978); knauti-

etosum arvensis Kojić & al. 2003; subass. potentilleto-

sum tormentillae Kojić & al. 2003; subass. genistetosum

sagittalis Kojić & al. 2003; subass. agrostietosum vul-

garis Kojić & al. 2003.

Ass. Armerio segoviensis-Arrhenatheretum bulbosi

Rodríguez-Rojo & Sánchez-Mata 2006

Distribution: Iberian Peninsula.

Ass. Lathyro tuberosi-Arrhenatheretum elatioris

Royer in Royer & al. 2006

Distribution: France.

Ass. Primulo veris-Festucetum rubrae Misset, Royer

& Didier in Royer & al. 2006

Diagnostic species: Anacamptis morio, Colchicum

autumnale, Primula veris, Vicia hirsuta.

Distribution: France.

Ass. Stellario gramineae-Festucetum rubrae Royer &

Didier in Royer & al. 2006

Distribution: France.

Ass. Triseto-Centaureetum macroptili Zelnik 2007

Diagnostic species: Centaurea macroptilon, Daucus

carota, Trisetum flavescens.

Distribution: Slovenia.

Variations: subass. avenuletosum pubescentis Zel-

nik 2007; subass. potentilletosum repentis Zelnik 2007.

Ass. Anthoxantho odorati-Festucetum pratensis

Borhidi 2012

Distribution: Hungary.

Ass. Primulo veris-Agrostietum capillaris Uhliarová

& Janišová 2014

Diagnostic species: Anemone sylvestris, Ajuga gen-

evensis, Anacamptis morio, Anemone ranunculoides,

Arabis hirsuta agg., Brachypodium pinnatum, Bri-

za media, Campanula cervicaria, C. rapunculoides,

Carex montana, Centaurea scabiosa, Convallaria ma-

jalis, Cruciata glabra, Dactylorhiza sambucina, Dian-

thus carthusianorum agg., Euphrasia rostkoviana agg.,

Hieracium sabaudum, Iris graminea, Knautia arven-

sis, Laserpitium latifolium, Lathyrus niger, Lilium mar-

tagon, Mercurialis perennis, Orchis mascula, Phleum

phleoides, Plantago media, Primula veris, Sedum sex-

angulare, Serratula tinctoria, Silene nutans, S. viscaria,

S. vulgaris, Succisa pratensis, Thalictrum lucidum, Tra-

gopogon pratensis subsp. orientalis, Trifolium rubens,

Valeriana officinalis, Vicia cassubica, Viola odorata.

Distribution: Ukraine.

Ass. Rhinantho freynii-Trisetetum flavescentis Dak-

skobler & Seliškar 2015

Diagnostic species: Helictotrichon pubescens, Med-

icago lupulina, Plantago media, Ranunculus bulbosus,

Rhinanthus freynii, Trisetum flavescens.

Distribution: Slovenia.

Variations: var. Rhinanthus minor; var. Polygala co-

mosa.

Ass. Betonico officinalis-Trifolietum pannonici Der-

zhypilsky & al. ex Zajac & al. 2016

Diagnostic species: Campanula glomerata agg.,

Gladiolus imbricatus, Helianthemum canum, Ononis

spinosa subsp. hircina, Peucedanum oreoselinum, Stel-

laria holostea, Trifolium alpestre.

Distribution: Ukraine.

Ass. Diantho deltoidis-Arrhenatheretum (Máthé &

Kovács 1960) Lengyel & al. 2016

Synonyms: Festuco rubrae-Cynosuretum ar-

rhenatheretosum Máthé & Kovács 1960; Anthyllido-

Festucetum rubrae arrhenatheretosum (Máthé & Ko-

vács 1960) Borhidi 2012.

Distribution: Hungary.

Plant communities without rank related to alliance

Arrhenatherion:

Plant community Poa bulbosa Markgraf 1932

Distribution: Albania.

Plant community Poa pratensis-Festuca rubra sen-

su Fijałk. 1962

Synonyms: Poo-Festucetum rubrae Fijałk. 1962.

Diagnostic species: Alopecurus pratensis; Festuca

rubra agg.; Holcus lanatus; Poa pratensis.

Distribution: Poland, Ukraine.

Plant community Bromus hordeaceus sensu Ell-

mauer & Mucina 1993; Ellmauer 1995

Distribution: Austria.

Plant community Artemisia vulgaris-Arrhenather-

um sensu Dierschke 1997

Synonyms: Arrhenatheretum tanacetetosum Knapp

1954; plant community Arrhenatherum elatius-Arte-

Annex 1. Continuation.

115Phytol. Balcan. 24(1) • Sofia • 2018

misia vulgaris sensu Bornkamm 1974; Tanaceto-Ar-

temisietum Fischer 1985; Micina & al. 1993; plant

community Arrhenatherum elatius; plant community

Urtica dioica-Arrhenatherum elatius sensu DG (Stot-

tele & Schmidt 1988).

Remarks: According to Päzolt & Jansen (2004)

plan community Artemisia vulgaris-Arrhenatherum

sensu Dierschke 1997 is synonym pro parte of Ar-

rhenatheretum elatioris Br.(-Bl.) 1915 [Syntax. syn.:

Pastinaco sativae-Arrhenatheretum elatioris Passarge

1964]. At the same time, Päzolt & Jansen (2004) do

not consider the synonymy, mentioned by Dierschke

(1997) – Arrhenatheretum tanacetetosum Knapp 1954;

plant community Arrhenatherum elatius-Artemisia

vulgaris sensu Bornkamm 1974; plant community Ur-

tica dioica-Arrhenatherum elatius sensu DG (Stottele

& Schmidt 1988).

Distribution: Germany.

Plant community Ranunculus repens-Alopecurus

pratensis sensu Dierschke 1997

Synonyms: Alopecurus pratensis Steffen 1931; Al-

opecuretum pratensis (Regel 1925) Steffen 1931; Plant

community Alopecurus pratensis-Galium mollugo sen-

su Hundt 1958; Alopecurion pratensis Passarge 1964;

Ranunculo repentis-Alopecuretum pratensis Krisch

1974; Ranunculo repentis-Alopecuretum pratensis

Micina & al. 1993; Trifolio repentis-Alopecuretum prat-

ensis Dietl 1995.

Remarks: According to Päzolt & Jansen (2004)

plant community Ranunculus repens-Alopecurus prat-

ensis sensu Dierschke 1997 is synonym pro parte of

Arrhenatheretum elatioris Br.(-Bl.) 1915 [Syntax. syn.:

Pastinaco sativae-Arrhenatheretum elatioris Passarge

1964].

Distribution: Germany, Poland (in Alopecuri-

on pratensis Passarge 1964), Ukraine (in Alopecurion

pratensis).

Plant community Holcus lanatus sensu Čarni 2001

Distribution: Slovenia.

Plant community Poa pratensis-Dactilis glomerata

sensu Pavlov & al. 2006

Distribution: Bulgaria.

Plant community Dactylis glomerata sensu Milkina

& Ljach 2008

Distribution: Ukraine.

Plant community Festuca pratensis sensu Milkina &

Ljach 2008

Distribution: Ukraine.

Plant community Gentiana lutea sensu Milkina &

Ljach 2008

Distribution: Ukraine.

Plant community Holcus mollis sensu Milkina &

Ljach 2008

Synonyms: Holcetum mollis Mich. 1998 nom. prov.

Distribution: Ukraine.

Plant community Pteridium aquilinum sensu Milki-

na & Ljach 2008

Distribution: Ukraine.

Plant community “Arrhenatheretum elatioris s. lat.”

sensu Šilc & Čarni 2012

Distribution: Slovenia.

Plant community “Arrhenatherion” sensu Šilc &

Čarni 2012

Distribution: Slovenia.

Plant community Dactylis-Festuca pratensis sensu

Šilc & Čarni 2012

Distribution: Slovenia.

Plant community Festuca rubra sensu Šilc & Čarni

2012

Remarks: incl. Festucetum rubrae Seliškar 1986

nom. ined.

Distribution: Slovenia.

Plant community Holcus lanatus sensu Šilc & Čarni

2012

Distribution: Slovenia.

Plant community Poa pratensis-Festuca rubra sensu

Klarzyńska & Kryszak 2015

Distribution: Poland.

Variations: var. typicum; var. Alopecurus pratensis;

var. Armeria maritima; var. Arrhenatheretum elatior-

is; var. Dactylis glomerata; var. Deschampsia caespito-

sa; var. Phalaris arundinacea.

References: Cincović & Kojić (1955); Cincović

(1956, 1957, 1959); Borza (1959); Blečić & Tatić (1960);

Horvat (1962); Horvatić (1963); Moravec (1965);

Medwečka-Kornaš & al. (1966); Kornaś & Medwecka-

Kornaś (1967), Csűrös & al. (1969); Pilát (1969); Bor-

za & Raţiu (1970); Csűrös (1970); Csűrös-Káptalan

(1970); Trzcińska-Tacik & al. (1973); Horvat & al.

(1974); Blečić & Lakušić (1976); Mišić & al. (1978);

Blažková (1979); Speidel (1979); Izco & Guitian Ri-

vera (1982); Jovanović-Dunjć; R. (1983); Balátová-

Tuláčková (1985); Jovanović-Dunjć & al. (1986);

Petković & Tatić (1987); Pop & al. (1988); Sýkora &

al. (1988); Reif & al. (1989); Sanda & Popescu (1991);

Annex 1. Continuation.

116 Velev, N. • Checklist of Arrhenatheretalia order

Rodwell (1992); Ellmauer & Mucina (1993); Oberdor-

fer (1993); Poldini & Oriolo (1994); Ellmauer (1995);

Kovács (1995, 2004); Dierschke (1997; 1999); Kra-

hulec & al. (1997); Sanda & al. (1998); Kojić & al. (1998,

2004, 2005); Stepanovich (1998); Gèhu (1999); Sapegin

& Dainenko (1999); Ninot & al. (2000); Rodwell & al.

(2000; 2007); Stančić (2000); Stepanovich (2000); Stud-

er-Ehrensberger (2000); Vicherek & al. (2000); Bardat

& al. (2001); Demeter & Veen (2001); Izco & al. (2001);

Jansen & Päzolt (2001); Jermacāne & Laiviņš (2001);

Mihăilescu (2001); Rivas-Martínez & al. (2001); So-

riano (2001); Dring & al. (2002); Konrád (2002); Pop

& al. (2002); Ružičková (2002); Sendžikaitė (2002);

Snowarski (2002-2005); Čarni (2003); Chytrý & Vi-

cherek (2003); Gliga (2003); Gselman & al. (2003);

Kaligaric & al. (2003); Bardat & al. (2004); Dengler &

Wollert (2004); Drăgulescu (2004); Kojić & al. (2004);

Kulcsár (2004); Päzolt & Jansen (2004); Rodríguez-Ro-

jo & Sánchez-Mata (2004, 2006); Sarbu & al. (2004);

Tsukanova (2005); Lakušić & al. (2005); Makra (2005);

Pavlov & al. (2006); Grzelak & Bocian (2006); Mosek

& Miazga (2006; 2008); Rūsiņa (2006a; b); Šilc & Košir

(2006); Trąba & al. (2006); Apostolova & al. (2007);

Groza & al. (2007); Hájková (2007); Hájková & al.

(2007); Janišová & al. (2010); Redžić (2007); Uhliarová

& al. (2007); Zelnik (2007); Solomakha (2008); Gafta

& Mountford (2008); Milkina & Ljach (2008); Pawlac-

zyk (2008); Sendžikaitė & al. (2008); Jarolímek & al.

(2008b); Škodová & Janišová (2008); Stančić (2008a; b);

Mounford & Akeroyd (2009); Kuzemko (2009b); Tzo-

nev & al. (2009); Wolański & Trąba (2009); Becker &

Becker (2010); Chocarro & al. (2010); Collaud (2011);

Velev & al. (2011b); Lengyel & al. (2012); Petras Sackl

& al. (2012); Šilc & Čarni (2012); Aćić & al. (2013a, b);

Corriol (2013); Redžić & al. (2013); Tasenkevich & Ka-

linovych (2013); Willner & al. (2013a, b); Kosić & Brit-

vec (2014); Loidi & al. (2014); Šilc & al. (2014); Vassi-

lev & al. (2014); Dakskobler & Seliškar (2015); Fernez &

Causse (2015); Klarzyńska & Kryszak (2015); Škodová

& al. (2015); Ferrez & al. (2016); Lengyel & al. (2016);

Zajac & al. (2016); Klimantová (2017); Rodríguez-Ro-

jo & al. (2017).

Alliance CYNOSURION CRISTATI Tüxen 1947 nom.

conserv. propos.

Original name: Cynosurion cristati Tüxen 1947 –

nom. conserv. propos. versus Lolion perennis Felföldy

1942 nom. ambig. et rejic. propos.

Synonyms: Polygonion avicularis Aichinger 1933

nom. ambig. et rejic. propos. (Art. 36); Lolion peren-

nis Felföldy 1942; Potentillion anserinae Tüxen 1947 p.

p.; Lolio-Potentillion anserinae Tüxen 1947 p. p. (Art.

29c); Polygonion avicularis Br.-Bl. ex von Rochow 1951

p. p. (Art. 31); Agrostideto-Festucion rubrae Puşcaru &

al. 1956 non Passarge 1969; Achilleo-Cynosurion (Tüx-

en 1947) Passarge 1969 nom. inval. (Art. 2d; 3m; 24a);

Alchemillo-Cynosurion Passarge 1969; Thymo-Cynosu-

rion Passarge 1969; Eu-Ranunculo-Cynosurion Passarge

1969; Eu-Thymo-Cynosurion Passarge 1969; Ranuncu-

lo repentis-Cynosurion Passarge 1969; Cardamino prat-

ensis-Cynosurion Passarge 1969; Bromo mollis-Cyno-

surion Passarge 1969; Sanguisorbo minoris-Cynosurion

Passarge 1969 Lolio perennis-Plantaginion majoris Siss-

ingh 1969; Phleo-Leontodontion (Br.-Bl. & Berset 1957)

Dietl 1972 p. p. (Art. 29); Lolion perennis Resmeriţă &

Pop 1972 (Art. 31); Lolio-Cynosurion Jurko 1974; Al-

chemillo-Trifolion repentis Passarge 1976; Plantagini-

Prunellion Eliáš 1980; Trifolio repentis-Lolion perennis

Dietl 1983; Lino biennis-Gaudinion fragilis de Foucault

1989; Alchemillo xantochlorae-Cynosurion cristati (Pas-

sarge 1969) de Foucault 1989; Rumici crispi-Cynosuri-

on cristati de Foucault 1989; Ranunculo sardoi-Plan-

taginion intermediae Julve 1993. According to Päzolt &

Jansen (2004) excl.: Cichorietum intybi Tüxen ex Sissin-

gh 1969. According to Ellmauer & Mucina (1993) incl.:

Lolio-Plantagenion (Sissingh 1969) Passarge 1979; non

Plantaginion majoris Br.-Bl. 1967 (Art. 8).

Corresponding names: Gaudinio fragilis-Cyno-

surenion Rivas Goday & Rivas-Martínez 1963; Gal-

io veri-Cynosurenion Rivas Goday & Rivas-Martín-

ez 1963; Trifolio-Cynosuretalia Sougnez & Limbourg

1963; Cynosuro-Phleetalia Passarge 1969; Trifolio-

Phleetalia (Sougnez 1963) Passarge 1969; Lolio-Cy-

nosurenion Jurko 1974; Polygalo-Cynosurenion Jurko

1974; Trifolienalia Oberdorfer 1990 p. p.

Remarks: 1) Rodwell & al. (2000) accept alliances

Cynosurion cristati Tüxen 1947 and Polygonion avic-

ularis Aichinger 1933 as different syntax; 2) Ferrez

(2007) subordinate Cynosurion to Trifolio repentis-

Phleetalia pratensis passarge 1969.

Diagnostic species: Achillea millefolium, Agrostis

capillaris, A. stolonifera, Ajuga reptans, Anthoxanthum

odoratum, Atrichum undulatum, Bellis perennis, Bri-

za media, Bromus commutatus, Capsella bursa-pasto-

ris, Carex ovalis, Cichorium intybus, Cirsium arvense,

Chamomilla suaveolens, Crepis capillaris, Cynosurus

Annex 1. Continuation.

117Phytol. Balcan. 24(1) • Sofia • 2018

cristatus, Euphrasia kerneri, E. rostkoviana, Festuca ni-

grescens ssp. nigrescens, F. pratensis, F. r ubra, Gentiana

cruciata, G. lutea, Geranium molle, Gladiolus imbrica-

tus, Holcus lanatus, Hordeum secalinum, Hypochoeris

radicata, Leucanthemum vulgare, Leontodon autum-

nalis, L. saxatilis, Linum catharticum, Lolium perenne,

Luzula campestris, Odontites rubra (=O. serotina), O.

verna (=O. vulgaris), Omalotheca sylvatica, Phleum

bertolonii, P. pratense, Plantago lanceolata, P. major,

P. m e d i a , Poa annua, P. pratensis, Potentilla reptans,

Primula veris, Prunella vulgaris, Ranunculus repens,

R. sardous, Rhinanthus minor, R. rumelicus, Rumex

crispus ssp. crispus, Stellaria graminea, Thymus pan-

nonicus, T. pullegioides, Trifolium hybridum, T. prat-

ense, T. repens, Veronica filiformis, V. serpyllifolia.

References: Puşcaru & al. (1956); Medwečka-

Kornaš & al. (1966); Gergely (1969); Csűrös & al.

(1969); Jurko (1969a, b, 1974); Borza & Raţiu (1970);

Resmeriţă (1970); de Foucault (1984); Mititelu & al.

(1986, 1988); Ellenberg (1988); Coldea (1990); San-

da & Popescu (1991); Ivan & al. (1993); Oberdor-

fer (1993); Zuidhoff & al. (1995); Dierschke (1997,

1999); Kojić & al. (1997); Krahulec & al. (1997); San-

da & al. (1998); Rodwell & Dring (2000); Stepanovich

(2000); Bardat & al. (2001); Mihăilescu (2001); Ri-

vas-Martínez & al. (2001, 2002); Rodwell & al. (2002);

Snowarski (2002-2005); Chytrý & Tichý (2003); Ka-

bucis & al. (2003); Kaligaric & al. (2003); Burkart &

al. (2004); Päzolt & Jansen (2004); Rodríguez-Rojo &

Sánchez-Mata (2004); Sarbu & al. (2004); Meshinev &

al (2005); Rasomavicius & al. (2006); Ferrez (2007);

Hájková & al. (2007); Janišová & al. (2007); Veen & al.

(2007); Dajić Stevanović (2008); Mikhailovna (2008);

Solomakha (2008); Jarolímek & al. (2008a); Škodová

& Janišová (2008); Velev & al. (2010); Biondi & Blasi

(2013); Biondi & al. (2014); Mucina & al (2016).

Suballiances related to Cynosurion:

Suball. Alchemillo xantochlorae-Cynosurenion cristati

Passarge 1969

Original name: Alchemillo-Cynosurenion Passarge

1969 nom. correct. (Alchemillo-Cynosurion pro suball.

Art. 41b).

Synonyms: Eu-Ranunculo-Cynosurenion Passarge

1969 nom. correct. (Eu-Ranunculo-Cynosurion pro

suball. Art. 41b) et nom. illeg. (Art. 34b); Eu-Thymo-

Cynosurenion Passarge 1969 nom. correct. (Eu-Thy-

mo-Cynosurion pro suball. Art. 41b) et nom. illeg.

(Art. 34b).

Corresponding names: Alchemillo xantochlorae-Cy-

nosurion cristati (Passarge 1969) de Foucault 1989; Thy-

mo-Cynosurion Passarge 1969 p. p.; Ranunculo-Cynosu-

rion Passarge 1969 p. p.; Achilleo-Cynosurion (Tüxen

1947) Passarge 1969 nom. inval. (Art. 2d; 3m; 24) p. p.

Suball. Bromo mollis-Cynosurenion cristati Pas-

sarge 1969

Original name: Bromo-Cynosurenion Passarge

1969 nom. correct. (Bromo-Cynosurion pro suball.

Art. 41b).

Synonyms: Lolio-Cynosurenion Jurko 1974 (Lolio-

Cynosurion pro suball. Art. 22; 41b).

Corresponding names: Rumici crispi-Cynosurion

cristati de Foucault 1969 nom. inval. (Art. 2d; 3m; 24).

Suball. Cardamino pratensis-Cynosurenion cristati

Passarge 1969

Original name: Cardamino-Cynosurenion Pas-

sarge 1969 nom. correct. (Cardamino-Cynosurion pro

suball. Art. 41b).

Corresponding names: Ranunculo-Cynosurion Pas-

sarge 1969 p. p.

Suball. Galio veri-Cynosurenion cristati Rivas Go-

day & Rivas Mart. 1963

Synonyms: Sanguisorbo minoris-Cynosurenion

cristati Passarge 1969 nom. illeg. (Art. 22).

Corresponding names: Thymo-Cynosurion Pas-

sarge 1969 p. p.

Suball. Polygalo vulgaris-Cynosurenion cristati

Jurko 1974

Original name: Polygalo-Cynosurenion Jurko 1964

nom. correct. (Polygalo-Cynosurion pro suball. Art. 41b).

Corresponding names: Achilleo-Cynosurion (Tüx-

en 1947) Passarge 1969 nom. inval. (Art. 2d; 3m; 24)

p. p.

Suball. Agrostienion tenuis Mirkin & Naumova 1986

Suball. Cerastio arvensis-Cynosurenion cristati Bla-

si & al. 2012

Diagnostic species: Armeria canescens, Cerastium

arvense, Rhinanthus minor, Salvia pratensis, Trag op o-

gon pratensis.

Suball. Trifolio resupinati-Cynosurenion cristati

Blasi & al. 2012

Diagnostic species: Cerastium glomeratum, Convol-

vulus arvensis, Crepis neglecta, Salvia verbenaca, Trifoli-

um micranthum, T. resupinatum, Verbena officinalis.

Annex 1. Continuation.

118 Velev, N. • Checklist of Arrhenatheretalia order

Suball. Danthonio decumbentis-Caricenion insula-

ris Farris & al. 2013

Diagnostic species: Bellium bellidioides, Carex car-

yophyllea subsp. insularis, Danthonia decumbens, Lu-

zula spicata subsp. italica, Mentha insularis.

Remarks: Farris & al. (2013) described this suballi-

ance and relate it provisionally to Cynosurion

cristati alliance and Arrhenatheretalia Tüxen 1931

order. In Biondi & Blasi (2013) Cynosurion is subordi-

nated to Trifolio repentis-Phleetalia pratensis Passarge

1969 order.

References: Bardat & al. (2001); Farris & al. (2013);

Ferrez (2007); Mikhailovna (2008); Blasi & al. (2012);

Biondi & Blasi (2013).

Associations related to Cynosurion:

Ass. Festucetum rubrae Rübel 1912

Distribution: Ukraine.

Variations: var. typicum; var. Elytrigia repens.

Ass. Agrostietum vulgaris Domin 1926

Synonyms: According to Kuzemko (2009b) incl.:

Galio veri-Agrostetum tenuis Goncharenko 2003.

Diagnostic species: Agrostis tenuis.

Distribution: Ukraine.

Variations: subass. typicum with two variants – var.

typicum, var. Dianthus deltoids; subass. festucetosum

pratensis with two variants – var. typicum, var. Koeler-

ia delavignei.

Ass. Plantagini majoris-Lolietum perennis Beger

1932 nom. inver. propos.

Original name: Lolio perennis-Plantaginetum ma-

joris Beger 1932

Synonyms: Lolietum perennis Gams 1927 (Art.

7); Lolio-Plantaginetum majoris (Linkola 1921) Be-

ger 1930; Lolio perennis-Plantaginetum major-

is Beger 1930; Lolio perennis-Plantaginetum major-

is Beger 1930; Lolio perennis-Polygonetum avicularis

Br.-Bl. 1930 p. p.; Junco compressi-Plantaginetum Br.-

Bl. & Moor 1935 (Art. 2b); Lolio perennis-Matrica-

rietum suaveolentis Tüxen 1937 p. p.; Lolietum per-

ennis Felföldy 1942; Lolium perenne Sauberer 1953

(Art. 2b); Festuco rupicolae-Lolietum perennis Hadač

& Rambousková 1980; According to Päzolt & Jansen

(2004) incl.: Lolietum perennis sensu Passarge 1964;

plant community Plantago major-Trifolium repens

sensu Dierschke 1997.

Diagnostic species: Achillea millefolium, Bellis per-

ennis, Capsella bursa-pastoris, Cichorium intybus,

Dactylis glomerata, Elymus repens, Holcus lanatus,

Hordeum murinum, Lolium perenne, Plantago lanceo-

lata, P. major, Poa annua, P. pratensis agg., P. trivialis

ssp. trivialis, Polygonum aviculare agg., Ranunculus re-

pens, Stellaria media, Taraxacum Sect. Ruderalia, Tri -

folium repens.

Distribution: Austria; Belarus, Bulgaria (in Poten-

tillion anserinae Tüxen 1947), Czech Republic, France

(in Lolio perennis-Plantaginion majoris G.Sissingh

1969), Germany, Hungary (in Potentillion anserinae),

Montenegro (in Polygonion avicularis), Romania (in

Polygonion avicularis Br.-Bl. 1930), Slovakia, Slovenia,

Spain (as Lolietum perennis Gams 1927 in Lolio peren-

nis-Plantaginion maioris Sissingh 1969); Ukraine.

Variations: Belarus: subass. agrostidetosum tenuis

sensu Stepanovich 2000; subass. bromopsidetosum in-

ermis sensu Stepanovich 2000; subass. dactylidetosum

glomeratae sensu Stepanovich 2000; subass. festuceto-

sum pratensis sensu Stepanovich 2000; subass. trifoli-

etosum pratensi sensu Stepanovich 2000; subass. tri-

folietosum repentis sensu Stepanovich 2000; Bulgaria:

subass. cynodontosum Tüxen 1950; Czech Republic:

var. Medicago lupulina; var. Poa annua.

Ass. Alchemillo hybridae-Poëtum supinae Aichinger

1933 nom. invers. et nom. mut.

Original name: Poa annua ssp. varia-Alchemilla

hybrida Aichinger 1933 (Poa annua ssp. varia = Poa

supina).

Diagnostic species: Carex ovalis, Plantago major,

Poa supina, Sagina procumbens, Veronica serpyllifolia.

Distribution: Czech Republic.

Ass. Anthoxantho odorati-Agrostietum tenuis Sill-

inger 1933

Original name: Anthoxantheto-Agrostidetum ten-

uis Sillinger 1933 (Anthoxanthum odoratum; Agrostis

tenuis = A. capillaris).

Synonyms: Agrostetum vulgaris Szafer & al. 1923;

Agrostio vulgaris-Gladioletum imbricati Br.-Bl. 1930

nom. nud. (Art. 2b); Gladiolo-Agrostietum Br.-Bl.

1930; Anthoxanthetum odorati Kmoniček 1934; Festu-

co-Cynosuretum Tüxen in Büker 1942 p. p.; Gladiolo

imbricati-Agrostietum tenuis (Br.-Bl. 1930) Pawłowski

& Walas 1949; Agrostetum-tenuis hungaricum Soó

1957; Gladioleto-Agrostidetum Borza 1963 nom. nud.

According to Kuzemko (2009b) incl.: Galio-Anthox-

anthetum Tüxen & Böttscher 1969; Centaurio-Nar-

Annex 1. Continuation.

119Phytol. Balcan. 24(1) • Sofia • 2018

cissetum angustifolii Kriczfalushij & Gendey 1987;

Polytricho-Narcissetum angustifolii Kriczfalushij &

Gendey 1987.

Diagnostic species: Agrostis capillaris, Alchemilla

gracilis, A. monticola, A. walasii, Anthoxanthum od-

oratum, Anthyllis vulneraria, Briza media, Campan-

ula glomerata, Carex caryophyllea, Carlina acaulis, C.

vulgaris s. lat., Cerastium fontanum, Centaurea jacea,

C. oxylepis, Cruciata glabra, Cynosurus cristatus, Di-

anthus armeria, Euphrasia rostkoviana, Gentiana cru-

ciata, Gladiolus imbricatus, Hieracium baihinii, Leon-

todon autumnalis, L. hispidus, Leucanthemum vulgare

agg. (L. ircutianum, L. vulgare), Linum catharticum,

Lotus corniculatus, Ononis spinosa, Pimpinella saxifra-

ga, Plantago media, Polygala major, P. vulgaris, Prunel-

la laciniata, P. vulgaris, Ranunculus bulbosus, R. pol-

yanthemos, Senecio jacobea, Sieglingia decumbens,

Thymus pulegioides, Trifolium montanum, T. ochroleu-

con, T. pratense, T. repens, Viola saxatilis ssp. decorata.

Distribution: Austria (in Violion caninae Schwick-

erath 1944, class Calluno-Ulicetea Br.-Bl. & Tx. 1943),

Belarus (in Arrhenatherion), Bulgaria, Czech Repub-

lic, Hungary (also in Nardo-Agrostion Sillinger 1933),

Latvia, Lithuania, Poland (also in Arrhenatherion),

Romania (also in Polygono bistortae-Trisetion), Rus-

sia (Komi Republic), Slovakia (in Arrhenatherion),

Ukraine (also in Arrhenatherion).

Variations: Belarus: subass. typicum; subass. agros-

tidetosum tenuis sensu Stepanovich 2000; subass.

brizetosum mediae sensu Stepanovich 2000; subass. cy-

nosuretosum cristati sensu Stepanovich 2000; subass.

festucetosum pratensis sensu Stepanovich 2000; sub-

ass. festucetosum rubrae sensu Stepanovich 2000; sub-

ass. nardetosum strictae sensu Stepanovich 2000; sub-

ass. plantaginetosum lanceolatae sensu Stepanovich

2000; Czech Republic: var. Brachypodium pinnatum;

var. Crepis biennis; Latvia: subass. typicum sensu Jurko

1969 with six variants – var. typicum, var. Calama-

grostis epigeios, var. Carex panicea, var. Melampy-

rum polonicum, var. Primula veris, var. Thymus ova-

tus; subass. holcetosum lanati Rūsiņa 2006 with three

variants – var. typicum, var. Helictotrichon pubescens,

var. Festuca pratensis; subass. nardetosum strictae

Jurko 1970; subass. prov. trifolietosum arvensis sensu

Jermacāne & Laiviņš 2001; Slovakia: var. Cardaminop-

sis halleri; var. Festuca rupicola; var. Primula elatior;

var. Primula veris; var. Trifolium rubens; Ukraine: sub-

ass. typicum; subass. hieracietosum pilosellae Kuzem-

ko 2009; subass. calamagrostietosum epigei Kuzemko

2009; subass. nardetosum strictae Kuzemko 2009 with

two variants – var. typicum, var. Astrantia major.

Ass. Brizeto-Anthoxanthetum Kmoniček 1936

Synonyms: Brizetum mediae Stepanovič 1999.

Diagnostic species: Anthoxanthum odoratum, Bri-

za media.

Distribution: Ukraine.

Variations: subass. alchemilletosum gracilis Kuzem-

ko 2009; subass. luzuletosum campestris Kuzemko

2009 with two variants – var. Festuca ovina, var. Polyg-

onum bistorta; subass. trifolietosum rubentis Kuzem-

ko 2009.

Ass. Lolio perennis-Cynosuretum cristati Tüxen 1937

Original name: Lolieto-Cynosuretum (Br.-Bl. & de

Leeuw 1937) Tüxen 1937 (Lolium perenne; Cynosurus

cristatus).

Synonyms: Cynosuretum cristati Beger 1922 p. p.

(Art. 7); pastures Lolium-Cynosurus sensu Br.-Bl. & De

Leeuw 1936 (Art. 3c); Lolio-Cynosuretum Br.-Bl. & De

Leeuw 1936; Cynosuro-Lolietum Br.-Bl. & De Leeuw

1936 nom. invers.; Festuco commutatae-Cynosuretum

Tüxen 1940 (Art. 7); Festuco-Cynosuretum Tüxen in

Büker 1941, 1942; Festuco-Cynosuretum cristati Tüxen

in Büker 1942 p. p.; Lolietum perennis Safta 1943; Gal-

io veri-Trifolietum repentis Sougnez 1957 p. max. p.;

Lolio-Poetum pratensis Kutschera 1961; Luzulo-Cyno-

suretum cristati Meisel 1961 p. p.; Crepido-Cynosure-

tum Knapp 1962; Lolio-Cynosuretum mesodanubiale

Stockhammer 1964 (Art. 34); Luzulo-Cynosuretum

cristati Meisel 1966; Festuco commutatae-Cynosure-

tum Tüxen ex Büker 1942; Alchemillo-Cynosuretum

Th. Müller in Oberdorfer & al. 1967 (Art. 7); Caro-

Poëtum pratensis Blažková 1967; Alchemillo-Cyno-

suretum Th. Müller ex Passarge 1969; Alopecuro-Cy-

nosuretum Passarge 1969; Ranunculo-Cynosuretum

Passarge 1969 (Art. 7); Carici-Cynosuretum (Sougn-

ez 1963) Passarge 1969 (Art. 7); Medicagini-Cynosure-

tum Passarge 1969; Poo trivialis-Rumicetum obtusifo-

lii Hülbusch 1969 p. p.; Phleo-Leontodontetum Br.-Bl.

& Bers. in Bers. 1969 p. p.; Chrysanthemo-Rumicetum

thyrsiflori Walter 1977 p. p.; Galio-Trifolietum Schami-

née & al. 1996; Agrostio tenuis-Trifolietum repentis Pas-

sarge 1999. According to the internet site of FloraWeb

(accessed 26.03.2009) incl.: Hordeetum secalini Krisch

1974; Lolio-Cynosuretum subass. Hordeum secalinum;

plant community Elymus repens-Hordeum secalinum;

plant community Plantago coronopus.

Annex 1. Continuation.

120 Velev, N. • Checklist of Arrhenatheretalia order

Remarks: 1) Krahulec & al. (1997) accept Lolio per-

ennis-Cynosuretum cristati Tüxen 1937 and Caro-Poë-

tum pratensis Blažková 1973 as separate syntaxa; 2)

Oberdorfer (1993) and Kuzemko (2009b) accept Lo-

lio perennis-Cynosuretum cristati Tüxen 1937 and Fes-

tuco-Cynosuretum Tüxen in Büker 1942 as separate

syntaxa; 3) Ellmauer & Mucina (1993) accept Cyno-

suro-Lolietum Br.-Bl. & De Leeuw 1936 nom. invers.

and Festuco commutatae-Cynosuretum Tüxen ex Bük-

er 1942 as separate syntaxa.

Homonyms: Lolio-Cynosuretum Büker 1941.

Diagnostic species: Achillea millefolium agg., Agros-

tis capillaris, Astrantia major, Bellis perennis, Campan-

ula scheuchzeri, Carlina acaulis, Carex flacca, C. hirta,

Carum carvi, Cerastium holosteoides, Cirsium arvense,

Cynosurus cristatus, Dactylis glomerata, Elymus repens,

Euphrasia rostkoviana, Festuca pratensis, F. rubra agg.,

Genista germanica, Holcus lanatus, Leontodon autum-

nalis, L. hispidus, Linum catharticum, Lolium perenne,

Luzula campestris, Plantago lanceolata, P. major, P. m e -

dia, Pimpinella saxifraga, Poa pratensis agg., Polygala

amara, Potentilla anserina, P. erecta, Ranunculus acris,

R. sardous, Taraxacum Sect. Ruderalia, Thymus pule-

gioides, Trifolium pratense, T. repens, Veronica filiformis.

Distribution: Albania, Austria, Belarus, Belgium,

Croatia, Czech Republic, France, Germany, Hungary,

Ireland, Italy, Lithuania, Netherlands, Poland, Romania,

Serbia, Slovakia, Slovenia, Ukraine, United Kingdom.

Variatrions: Belarus: subass. agrostidetosum ten-

uis sensu Stepanovich 2000; subass anthoxanthetosum

odorati sensu Stepanovich 2000; subass. brizetosum

mediae sensu Stepanovich 2000; subass. deschampsi-

etosum cespitosae sensu Stepanovich 2000; subass. fes-

tucetosum pratensis sensu Stepanovich 2000; subass.

festucetosum rubrae sensu Stepanovich 2000; subass.

trifolietosum repentis sensu Stepanovich 2000; Czech

Republic: var. Poa trivialis; var. Plantago lanceola-

ta; France: subass. typicum; subass. ranunculetosum

bulbosi Sougnez 1951; subass. achilletosum millefolii

Trivaudei in Ferrez 2007; subass. alopecuretosum prat-

ensis Trivaudei in Ferrez 2007; Germany: subass. lu-

zuletosum campestris with three variants – var. Lotus

corniculatus, var. Ranunculus polyanthemos, var. Ve-

ronica arvensis; Italy: subass. alopecuretosum utricu-

lati Venanzoni 1992; fac. Carex acutiformis; Poland:

subass. typicum with two variants – var. typicum, var.

Agrostis gigantea; subass. ranunculetosum Grodzins-

ka, Zarzycki 1967 (Syn.: Lolio-Cynosuretum ranuncu-

letosum Tüxen & Preising 1951 p. p.); subass. anthox-

anthetosum odoratae sensu Trąba & al. 2008; subass.

dactylietosum glomeratae sensu Trąba & al. 2008; sub-

ass. festucetosum pratense sensu Trąba & al. 2008; sub-

ass. trifolietosum repentis sensu Trąba & al. 2008; sub-

ass. dactylidetosum glomeratae sensu Maćkowiak &

al. 2016 with two variants – var. typicum, var. Bro-

mus horeaceus; subass. plantaginetosum majoris sen-

su Maćkowiak & al. 2016 var. typicum; subass. tri-

folietosum repenti sensu Maćkowiak & al. 2016 with

three variants – var. typicum, var. Armeria mariti-

ma ssp. elongata; var. Poa pratensis. Romania: sub-

ass. centaureetosum induratae Borza 1941; Slovakia:

var. Plantago major; var. Thymus pulegioides; Ukraine:

subass. typicim; subass. poetosum pratensis Kuzem-

ko 2009; var. Agrostis stolonifera; var. Astrantia major;

var. Carex caespitosa; var. Trifolium dubium; var. typi-

cum (pointed as variants of Festuco-Cynosuretum cris-

tati Tüxen 1942).

Ass. Loto uliginosi-Cynosuretum cristati Tüxen 1937

Diagnostic species: Cardamine pratensis, Carex

leporina, Juncus effusus, Lotus pedunculatus, Silene

flos-cuculi.

Distribution: France.

Ass. Agrostetum tenuis montanum Szafer & al. 1923;

Issler 1933; 1941 nom. illeg. (Art. 34a)

Distribution: Romania.

Variations: subass. festucetosum rubrae Anghel

& al. 1965; subass. festucetosum rupicolae Ciorcĭrlan

1968; subass. trifolietosum pratensis Păun & al. 1972;

subass. festucetum valesiacae (Ardelean 1983) Sanda

& Popescu 1991 (Corresp. name: Agrostio-Festucetum

valesiacae Ardelean 1983); fac. poosum trivialis; fac.

rhinanthosum alectorolophi; fac. moliniosum coerule-

ae; fac. trifoliosum patens.

Ass. Lolio-Cynosuretum Büker 1941

Homonyms: Lolio-Cynosuretum Br.-Bl. & De Leeuw

1936.

Diagnostic species: Bellis perennis, Lolium perenne,

Prunella vulgaris.

Distribution: Ukraine.

Variations: var. typicum; var. Festuca rubra; var.

Poa pratensis.

Ass. Deschampsietum flexuosae Issler 1942

Synonyms: Asplenio-Deschampsietum flexuosae Balász

1941; Festuceto rubrae-Deschampsietum flexuosae Raţiu

1964.

Distribution: Romania.

Annex 1. Continuation.

121Phytol. Balcan. 24(1) • Sofia • 2018

Ass. Pediculari campestris-Caricetum montanae Soό

1945

Synonyms: Agrostis tenuis-Carex montana-Festu-

ca rubra Pop & al. 1962 nom. illeg. (Art. 34c); Carex

montana-Festuca rubra-Agrostis tenuis Gergely 1964;

Csűrös-Káptalan 1964 nom. illeg. (Art. 34c).

Distribution: Romania.

Ass. Agrostio-Danthonietum provincialis Soό 1947

(Agrostis canina)

Synonyms: Agrostideto-Danthonietum Soό 1947;

Agrostis tenuis-Danthonia privincialis Bujorean & al.

1959.

Diagnostic species: Agrostis canina, A. stolonifera,

Danthonia provincialis (=D. alpina).

Distribution: Romania.

Ass. Poëto angustifoliae-Agrostidetum capillaris Soό

& Csűrös 1947

Synonyms: Poo angustifolie-Agrostietum tenuis Soό

& Csűrös 1947.

Diagnostic species: Poa pratensis var. angustifolia,

Agrostis capillaris.

Distribution: Romania (also in Arrhenatherrion).

Ass. Festuco rubrae-Agrostetum capillaris Horvat

1951

Synonyms: Festuceto rubrae-Agrostidetum ten-

uis montanum Csűrös & Resmeriţă 1960 nom. illeg.

(Art. 34a); Festuca rubra-Agrostis tenuis sensu Csűrös

& Resmeriţă 1960; Echio-Rumicetum acetosellae Soran

1962; Agrostio-Callunetum vulgaris Resmeriţă &

Csűrös 1963; Agrostis tenuis-Festuca sulcata-Festuca

pseudovina Evd. Puşcaru & al. 1963 nom. illeg. (Art.

34c); Festuco rubrae-Agrostietum capillaris Csűrös-

Káptalan 1964; Agrostideto capillaris-Festucetum rupi-

colae Csűrös-Káptalan (1962) 1964 nom. invers.; Fes-

tuceto rubrae-Agrostidetum tenuis subxerophyllum

Gergely 1964 nom. illeg. (Art. 34a); Agrostetum tenuis

biharicum Resmeriţă 1965; 1970 nom. illeg. (Art. 34a);

Agrostetum tenuis montanum apud Resmeriţă 1970

nom. illeg. (Art. 34a); Hypochoerio radicatae-Agrosri-

etum tenuis Pop & al. 1988 pro syn., nom. nud. (Art.

2a; 3a; Recomm. 46 f); Rumici acetosellae-Agrostietum

tenuis Sanda & Popescu 1981.

Corresponding names: Hypochoerio radicatae-

Agrostietum tenuis Pop & al. 1988 subass. calluneto-

sum Pop & al. 1988 [Syn.: Agrostetum tenuis biharicum

Resmeriţă 1965 nom. illeg. (Art. 34a)].

Pseudonyms: Festuca rubra-Agrostis tenuis auct.

plur. non Csűrös & Resmeriţă 1960; Festuco rubrae-

Cynosuretum auct. roman.; Festucetum rubrae auct.

roman.; Agrostetum tenuis auct. roman.

Remarks: Sanda & al. (1999) accept Festuco rubrae-

Agrostetum capillaris Horvat 1951 and Rumici aceto-

sellae-Agrostietum tenuis Sanda & Popescu 1981 (Syn.:

Echio-Rumicetum acetosellae Soran 1962) as separate

syntaxa. Festuco-Agrostietum Horvat 1962 nom. nud.

is mentioned for Slovenia.

Diagnostic species: Achillea distans, A. millefolioum

ssp. tanacetifolia, Alchemilla vulgaris, Agrostis capilla-

ris, Anthyllis vulneraria, Antennaria dioica, Anthoxan-

thum odoratum, Arnica montana, Asperula cynanchi-

ca, Campanula patula, Carlina acaulis, Centaurea

melanocalanthia, Coronilla varia, Dianthus deltoides,

Dorycnium herbaceum, Festuca rubra, F. rupicola, Fil-

ipendula vulgaris, Gentiana praecox, Helianthemum

nummularium, Hieracium aurantiacum, Hypericum

maculatum, H. perforatum, Koeleria gracilis, Luzu-

la nemorosa, Medicago falcata, Moenchia mantica,

Nardus stricta, Omalotheca sylvatica, Polygala como-

sa, Rhinanthus minor, Rumex acetosa, Salvia praten-

sis, Sanguisorba minor, Sieglingia decumbens, Stachys

germanica, Stellaria graminea, Trifolium campestre, T.

montanum, Veronica officinalis, Viola declinata.

Distribution: Bulgaria; Croatia [according to Hor-

vat (1962), Horvatić (1963) and Stančić (2000) in Ar-

rhenatherion], Montenegro (in Arrhenatherion); Ro-

mania; Serbia (also in Arrhenatherion), Slovenia? (as

Festuco-Agrostietum Horvat 1962 nom. nud.), Ukraine.

Variations: Romania: subass. typicum sensu Cold-

ea 1991; subass. coronilletosum variae Jurko 1969;

subass. festucetosum rupicolae Pázmány 1971; subass.

pteridietosum Szabό 1971; subass. agrostetosum tenuis

Coldea 1972; subass. danthonietosum (Pázmány 1971)

Coste 1975 (Corresp. names: Festuco rubrae-Agroste-

tum capillaris Horvat 1951 fac. danthoniosum provin-

cialis); subass. deschampsietosum flexuosae Raţiu &

Gergely 1976; subass. arnicetosum montanae Raţiu &

Gergely 1976; subass. nardetosum strictae Pop 1976

[Syn.: Nardetum strictae alpinum Borza 1934 nom. il-

leg. (Art. 34a) p. p.; Nardetum strictae alpinum Ghişa

1940 nom. illeg. (Art. 34a) p. p.; Nardetum strictae al-

pinum Csűrös 1955 nom. illeg. (Art. 34a) p. p.; Nar-

detum strictae alpinum Puşcaru & al. 1956 nom. illeg.

(Art. 34a); Nardetum alpigenum austro–carpaticum

Borza 1959 nom. illeg. (Art. 34a); Nardetum stric-

tae subalpinum Buia & al. 1962 nom. illeg. (Art. 34a);

Nardetum strictae alpinum Buia & al. 1962 nom. illeg.

Annex 1. Continuation.

122 Velev, N. • Checklist of Arrhenatheretalia order

(Art. 34a); Nardetum strictae alpinum Buia 1963 nom.

illeg. (Art. 34a) p. p.; Nardetum strictae montanum

Resmeriţă & Csűrös 1963 nom. illeg. (Art. 34a); Violo

declinatae – Nardetum strictae Simon 1966; Agrosteto-

Festucetum rubrae montanum nardosum Resmeriţă &

Csűrös 1966 nom. illeg. (Art. 34a); Nardetum strictae

alpinum Beldie 1967 nom. illeg. (Art. 34a) p. p.; Nar-

do–Vaccinietum Resmeriţă 1970; Nardo-Festucetum

tenuifoliae Buiculescu 1971; Festuco rubrae-Agroste-

tum capillaris Horvat 1951 var. Nardus stricta Danciu

1974; Hieracio polosellae–Nardetum strictae Pop & al.

1988 pro syn.; nom. nud. (Art. 2b; 3a; Recomm. 46f)];

subass. genistetosum sagittalis Coldea 1991 (Cor-

resp. names: Festuceto-Genistetum sagittalis Csűrös

& Resmeriţă 1960 p. p.; Festuca rubra-Genistella sag-

ittalis Gergely 1969 non Issler 1942); subass. cariceto-

sum montanae Pop & al. 2000; subass. avenuletosum

praeustae Bărbos 2006; var. typicum; var. Poa praten-

sis; var. Ranunculus acris; fac. alchemillosum mollis;

fac. genistosum sagittalis; fac. onobrychosum viciifoliae;

fac. rhinanthosum majori Ciucă 1963; fac. pteridiosum

aquilini sensu Raclaru & Zanoschi 1968 (Syn.: Festuco

rubrae-Agrostetum capillaris Horvat 1951 pteridiosum

sensu Pop 1976); fac. agrostiosum tenuis.

Remarks: Within Agrostideto capillaris-Festucetum

rupicolae Csűrös-Káptalan (1962) 1964, mentioned

as synonym, three subassociations are pointed: sub-

ass. coronilletosum variae Jurko 1967; subass. pteride-

tosum Szabό 1971; subass. caricetosum montanae Pop

& al. 2002 (Syn.: Carex montana-Festuca rubra sensu

Pop & al. 2002).

Ass. Centaureo nigrae-Cynosuretum cristati Br.-Bl.

& Tüxen 1952

Synonyms: Plant community Cynosurus cristatus-

centaurea nigra sensu Rodwell & al. 2000.

Diagnostic species: Agrostis capillaris, Centaurea

nigra agg., Hypochaeris radicata, Lotus corniculatus.

Distribution: France, Ireland, United Kingdom.

Ass. Agrostio capillaris-Poetum subcaeruleae Tüxen

1955

Distribution: France.

Ass. Brizeto-Anthoxanthetum Kmoníček 1936

Distribution: Ukraine.

Ass. Campanulo abietinae-Festucetum nigricantis

Anghel & al. 1956

Distribution: Romania.

Ass. Phleo-Leontodontetum Br.-Bl. & Berset 1956

Distribution: Switzerland.

Ass. Plantagini lanceolatae-Festucetum rubrae Sca-

moni 1956

Distribution: Ukraine.

Variations: var. typicum; var. Elytrigia repens.

Ass. Agrostio castellanae-Cynosuretum cristati Tel-

es 1957

Synonyms: Bromo commutati-Cynosuretum cristati

Teles (1963) 1966 (Art. 22; 29); Bromo commutati-Cy-

nosuretum cristati Teles 1970 (Art. 43); Bromo racemo-

si-Cynosuretum cristati Teles 1970.

Remarks: According to Rodríguez-Rojo & al.

(2014) incl.: Festuco amplae-Cynosuretum cristati Ri-

vas-Martínez ex Fuente 1986.

Distribution: Iberian Peninsula.

Ass. Alopecureto pratensis-Festucetum pseudovinae

Juhász-Nagy 1957

Original name: Alopecureto-Festucetum pseudovi-

nae Juhász-Nagy 1957.

Synonyms: Festucetum pseudovino-rupicolae

Řehořek 1971 (Art. 2b; 3b).

Diagnostic species: Alopecurus pratensis, Capsella

bursa-pastoris, Dianthus armeria, Erophila verna agg.,

Eryngium planum, Festuca pseudovina, Myosotis stric-

ta, Poa bulbosa, Potentilla argentea agg., Veronica ar-

vensis.

Distribution: Hungary, Slovakia.

Ass. Galio veri-Trifolietum repentis Sougnez 1957

Remarks: According to Zuidhoff & al. (1995) Gal-

io-Trifolietum Sougnez 1957 and Lolio-Cynozuretum

Tüxen 1937 are separate syntaxa, meanwhile Päzolt &

Jansen (2004) accept these as “Syntax. Syn. p. max p.”.

Diagnostic species: Galium verum, Medicago lu-

pulina, Pimpinella saxifraga, Plantago media, Primu-

la veris.

Distribution: Belgium (also in Arrhenatherion),

France (also in Arrhenatherion), Germany, Nether-

lands, Switzerland, United Kingdom.

Variations: Belgium: subass. typicum; France: sub-

ass. typicum.

Ass. Junco acutiflori-Cynosuretum cristati Sougnez

1957

Distribution: Belgium, France.

Ass. Trifolio-Festucetum rubrae Oberdorfer 1957

Synonyms: Festucetum rubrae semenicense Borza

1946 nom. illeg. (Art. 34a); Festucetum rubrae fallax

Puşcaru & al. 1956; Festuceto-Alchemilletum vulgaris

Csűrös & Resmeriţă 1960; Festucetum rubrae monta-

num Csűrös & Resmeriţă 1960 nom. illeg. (Art. 34a);

Annex 1. Continuation.

123Phytol. Balcan. 24(1) • Sofia • 2018

Festucetum rubrae subalpinum Csűrös & Resmeriţă

1960 nom. illeg. (Art. 34a); Festucetum rubrae mon-

tanum calcophilum Csűrös & Resmeriţă 1965 nom.

illeg. (Art. 34a); Festuceto-Alchemilletum vulgaris

Csűrös & Resmeriţă 1965; Festuca rubra-Origanum

vulgare Kovács & Moldovan 1965; Festuca rubra-Iris

graminea Kovács & Moldovan 1965; Festuca rubra-

Thymus marschallianus Kovács & Moldovan 1965;

Festuca rubra-Teucrium chamaedrys Ghişa & al. 1965;

Festuca rubra-Picea excelsa Resmeriţă 1966; Festuce-

tum rubrae xeromezophilum Resmeriţă (1965) 1970

nom. illeg. (Art. 34a); Alchemillo-Festucetum rubrae

Resmeriţă 1972 nom. illeg. (Art. 31); Festucetum ru-

brae Resmeriţă 1972; Galio veri-Festucetum rubrae

Resmeriţă 1980; Potentillo (aureae-ternatae)-Festuce-

tum rubrae Resmeriţă 1985.

Pseudonyms: Festucetum rubrae submontanum

nom. illeg. (Art. 34a) auct. rom. non (Scam. 1955)

Oberdorfer 1957.

Diagnostic species: see Poo-Trisetetum flavescentis

Knapp ex Oberdorfer 1957.

Remarks: 1) According to Oberdorfer (1993) and

Hájková & al. (2007) Trifolio-Festucetum rubrae Ober-

dorfer 1957 and Poo-Trisetetum flavescentis Knapp

ex Oberdorfer 1957 are synonyms and subordinated

to Arrhenatherion. Sanda & Popescu (1991) consider

these as different syntaxa and refer them to different

alliances – Trifolio-Festucetum rubrae to Cynosurion,

but Poo-Trisetetum flavescentis to Polygono-Trise-

tion. 2) Coldea (1987) describe Scorzonero roseae–Fes-

tucetum nigricantis (Puşcaru & al. 1956) Coldea 1987

and pointed the following synonyms: Festucetum ru-

brae fallax Puşcaru & al. 1956; Nardo–Festucetum ru-

brae fallax Puşcaru & al. 1959; Festuca supina–Festuca

rubra Puşcaru & al. 1959; Festucetum rubrae monta-

num Csűrös & Resmeriţă 1960 nom. illeg. (Art. 34a);

Festucetum rubrae subalpinum silicicolum Csűrös &

Resmeriţă 1960 nom. illeg. (Art. 34a); Festuco–Nar-

detum strictae montanum Csűrös & Resmeriţă 1960

nom. illeg. (Art. 34a); Festuco–Alchemilletum vul-

garis Csűrös & Resmeriţă 1960. Three of the syno-

nyms (Festucetum rubrae fallax Puşcaru & al. 1956;

Festucetum rubrae montanum Csűrös & Resmeriţă

1960; Festuco–Nardetum strictae montanum Csűrös

& Resmeriţă 1960) are mentioned also as synonyms

(Sanda & Popescu 1991) of Trifolio-Festucetum rubrae

Oberdorfer 1957. Thus Scorzonero roseae–Festucetum

nigricantis (Puşcaru & al. 1956) Coldea 1987 and Tr i-

folio-Festucetum rubrae Oberdorfer 1957 should be

considered as synonyms. Such consideration is not

followed by many authors (Coldea 1987, 1990; Pop &

al. 1988, 2002; Ivan & al. 1993; Mihăilescu 2001; Ni-

culescu 2004). Meanwhile, it is widespread the accept-

ance that Scorzonero roseae–Festucetum nigricantis

(Puşcaru & al. 1956) Coldea 1987 is a separate syn-

taxon, subordinated to Potentillo ternatae–Nardion

strictae Simon 1958, which is related to classes Nar-

do–Callunetea Preising 1949 (Syn.: Calluno–Ulicetea

Br.-Bl. & Tüxen ex Klika & Hadač 1944) and Juncet-

ea trifidi Hadač in Klika & Hadač 1944 (Syn.: Caricet-

ea curvulae Br.-Bl. 1948); 3) Krahulec & al. (1997) ac-

cept Trifolio-Festucetum rubrae Oberdorfer 1957 and

Poo-Trisetetum flavescentis Knapp ex Oberdorfer 1957

as separate syntaxa but relate them to Arrhenatheri-

on; 4) The synonym Festucetum rubrae subalpinum si-

licicolum Csűrös & Resmeriţă 1960, pointed as such

by Sanda & Popescu (1991), is considered as syno-

nym, by Coldea (1987), of Scorzonero roseae–Festuce-

tum nigricantis (Puşcaru & al. 1956) Coldea 1987. All

this means that Trifolio-Festucetum rubrae Oberdorfer

1957 subass. agrostidetosum Puşcaru & al. 1956 and

Scorzonero roseae–Festucetum nigricantis (Puşcaru

& al. 1956) Coldea 1987 shloud be treated as corre-

sponding names.

Distribution: Czech Republic (in Arrhenatherion);

Germany, Romania, Ukraine (also in Arrhenatherion

and Festucion pratensis).

Variations: Czech Republic: subass. alopecureto-

sum Neuhäusl 1972 (Alopecurus pratensis); subass.

carlinetosum Neuhäusl 1972 (Carlina acaulis); Roma-

nia: subass. agrostidetosum Puşcaru & al. 1956 [Syn.:

Trifolio-Festucetum rubrae Oberdorfer 1957 subass. si-

licicolum Csűrös & Resmeriţă 1960; Corresp. names:

Trifolio-Festucetum rubrae Oberdorfer 1957 fac. agros-

tosum tenius Raclaru 1959; Festucetum rubrae subalpi-

num silicicolum Csűrös & Resmeriţă 1960 nom. illeg.

(Art. 34a); Festucetum-Agrostetosum tenuis subalpi-

num Resmeriţă (1969) 1970 nom. illeg. (Art. 34a)];

subass. alchemilletosum Puşcaru & al. 1956; subass. al-

opecuretosum Puşcaru & al. 1956; subass. deschampsi-

etosum Puşcaru & al. 1956; subass. retezaticum Csűrös

& al. 1956; subass. calcipolum Csűrös & Resmeriţă

1960; subass. nardetosum Buia & al. 1962 (Corresp.

names: Trifolio-Festucetum rubrae Oberdorfer 1957

fac. nardosum strictae sensu Maloch 1932; Pop & Tre-

tiu 1958; Trifolio-Festucetum rubrae Oberdorfer 1957

Annex 1. Continuation.

124 Velev, N. • Checklist of Arrhenatheretalia order

fac. nardosum strictae sensu Resmeriţă 1963); sub-

ass. supinosum Buia & al. 1962; subass. brizetosum

mediae Raclaru 1967; Roşca & al. 1969; subass. fes-

tucetosum (rubrae)-cytisetosum nigricantis Resmeriţă

1970 nom. illeg. (Art. 34c); fac. anthoxanthosum od-

oratae; fac. caricosum montanae; fac. cynosurosum

cristati; fac. deschampsiosum flexuosae; fac. festuco-

sum pictae; fac. festucosum pratensis; fac. genistosum;

fac. narcisetosum; fac. polytrichiosum; fac. rhinantho-

sum glaberi; fac. poosum nemoralis sensu (Paucă & al.

1960) Sanda & Popescu 1991 (Corresp. names: Fes-

tuco rubrae-Poëtum nemoralis Paucă & al. 1960; Poa

nemoralis+Festuca rubra Paucă & al. 1960).

Ass. Bromo racemosi-Cynosuretum cristati Horvatić

(1930) 1958

Remarks: for details see in Arrhenatherion.

Ass. Caro verticillati-Cynosuretum cristati Bellot &

Casaseca ex Tüxen in Tüxen & Oberdorfer 1958

Synonyms: Lolio-Cynosuretum sensu Bellot &

Casaseca 1956 non Br.-Bl. & De Leeuw 1936

Corresponding names: Lolio-Cynosuretum lolieto-

sum multiflori Bellot & Casaseca 1959.

Distribution: Iberian Peninsula.

Ass. Lino angustifolii-Cynosuretum cristati Allorge

ex Oberdorfer & Tüxen in Tüxen & Oberdorfer 1958

Synonyms: Anthoxantum odoratum-Cynosurus

cristatus Allorge 1941 (Art. 2b); Lino biennis-Cyno-

suretum cristati Allorge ex Oberdorfer & Tüxen in

Tüxen & Oberdorfer 1958 nom. mut. (Art. 45).

Diagnostic species: Centaurea nigra agg., Crepis capil-

laris, Daucus carota, Gaudinia fragilis, Linum bienne.

Distribution: France (also in Arrhenatherion), Ibe-

rian Peninsula.

Ass. Merendero pyrenaicae-Cynosuretum cristati

Oberdorfer & Tüxen in Tüxen & Oberdorfer 1958

Diagnostic species: Agrostis capillaris, Colchicum

montanum, Helictochloa marginata, Thymus praecox.

Distribution: Iberian Peninsula.

Variations: Spain: subass. cynosuretosum cristati;

subass. bistortetosum officinalis Loidi & al. 2014.

Ass. Anthoxantho odorati-Festucetum pseudovinae

Juhász-Nagy 1959

Distribution: Hungary.

Ass. Agrostideto-Nardetum strictae Resmeriţă &

Texter 1956; Pauča & al. 1960 (Agrostis capillaris)

Synonyms: Agrostis tenuis-Nardus stricta Bujorean

& al. 1959.

Distribution: Romania.

Ass. Festuco rubrae-Poëtum nemoralis Pauča & al.

1960

Diagnostic species: Festuca rubra, Poa nemoralis.

Distribution: Romania.

Ass. Agrostideto tenuis-Festucetum rupicolae

Puşcaru & al. 1962

Diagnostic species: Agrostis capillaris, Briza media,

Centaurea jacea, Cynosurus cristatus, Euphrasia rostk-

oviana, Festuca rupicola, Linum catharticum, Potentil-

la argentea, Trifolium campestre.

Distribution: Romania.

Ass. Agrosti-Cynosuretum Resmeriţă 1963 (Agrostis

capillaris, Cynosurus cristatus)

Synonyms: Agrostis tenuis+Cynosurus cristatus

Evd. Puşcaru & al. 1959; Agrostideto-Cynosuretum

cristati Resmeriţă 1963.

Distribution: Romania.

Ass. Prunello vulgaris-Ranunculetum repentis Win-

terhoff 1963

Original name: Prunella vulgaris-Ranunculus re-

pens Winterhoff 1963.

Synonyms: Plantagini-Juncetum macri Oberdor-

fer 1957 nom. prov. (Art. 3b); Prunello-Plantaginetum

majoris Faliński 1963 nom. prov. (Art. 3b); Alchemil-

lo-Prunelletum vulgaris Faliński 1963 nom. prov. (Art.

3b); Alchemillo-Prunelletum vulgaris Passarge 1979

(Art. 3f).

Pseudonyms: Juncetum tenuis sensu auct. non Sch-

wickerath 1944 nom. prov.

Diagnostic species: Bellis perennis, Geranium pusil-

lum, Juncus tenuis, Leontodon autumnalis, Lolium per-

enne, Plantago major, Poa annua, Potentilla anserina,

Prunella vulgaris, Ranunculus repens.

Distribution: Austria (in Plantagini-Prunellion

Eliáš 1980), Czech Republic, Poland, Slovakia (in

Plantagini-Prunellion Eliáš 1980).

Ass. Agrostis tenuis-Festuca pseudovina Pop & al.

1964; Csűrös-Káptalan 1964

Distribution: Romania.

Ass. Agrostis tenuis-Lolium perenne Csűrös-

Káptalan 1964

Distribution: Romania.

Ass. Trifolio repenti-Lolietum perennis Krippelová

1967; Resmeriţă & Pop 1967

Synonyms: Lolio-Trifolietum repentis Resmeriţă &

al. 1967.

Corresponding names: Lolieto-Cynosuretum trifo-

liosum repentis Resmeriţă & Csűrös 1966.

Annex 1. Continuation.

125Phytol. Balcan. 24(1) • Sofia • 2018

Diagnostic species: Achillea setacea, Artemisia

austriaca, Carduus acanthoides, Erodium cicutarium,

Euphorbia cyparissias, Festuca pseudovina, Inula bri-

tannica, Lolium perenne, Lotus corniculatus, Poa bul-

bosa, Potentilla argentea, Thymus auctus, Trifolium

repens.

Remarks: According to Jarolímek & al. (2008b) this

association is synonym of Lolietum perennis Gams

1927 (Syn.: Plantagini majoris-Lolietum perennis Be-

ger 1932).

Distribution: Hungary, Romania (also in Agropy-

ro-Rumicion crispi Nordh. 1940), Serbia, Slovakia (by

the synonym Lolietum perennis Gams 1927), Ukraine.

Ass. Alchemillo monticolae-Cynosuretum cristati

Müller & Görs 1968

Distribution: Belgium, France.

Ass. Lolio-Trifolietum subterranei Bruno & Covarel-

li 1968

Distribution: Italy.

Ass. Campanulo subromboidalis-Cynosuretum cris-

tati Nègre 1969

Remarks: According to Rodríguez-Rojo & al.

(2014) incl.: Carici ornithopodae-Agrostietum capilla-

ris Villegas 1997.

Diagnostic species: Centaurea jacea, Pimpinella

saxifraga, Prunella grandiflora, Rhinanthus mediterra-

neus.

Distribution: Iberian Peninsula.

Ass. Medicagini lupulinae-Cynosuretum cristati Pas-

sarge 1969

Distribution: France.

Ass. Agrosti-Genistelletum Boşcaiu 1970 (Genistella

sagittalis)

Synonyms: Festuca rubra-Genista sagittalis An-

ghel & al. 1956; Festuca rubra+Genista sagittalis

Şerbănescu 1963.

Distribution: Romania.

Ass. Anthemido nobilis-Cynosuretum cristati Teles

1970

Distribution: Iberian Peninsula.

Ass. Anthyllido vulnerariae-Festucetum rubrae

(Máthe & Kovács 1960) Soό 1971

Original name: Festuco rubrae-Cynosuretum Tüx -

en 1940 subass. festucetosum rubrae Máthe & Kovács

1960.

Synonyms: Festuco rubrae-Cynosuretum auct.

hung. non Tüxen 1940; Soό 1957, 1962; Festuco ru-

brae-Cynosuretum nardosum Resmeriţă 1968.

Diagnostic species: Agrostis capillaris, Anthoxan-

thum odoratum, Anthyllis vulneraria agg., Campanula

patula, Carum carvi, Cynosurus cristatus, Festuca ru-

bra, Luzula campestris, Trifolium pratense.

Distribution: France, Hungary (also in Arrhenathe-

rion), Romania.

Variations: subass. festucetosum pratensis Soό 1947;

subass. festucetosum pseudovinae Soό 1964, Szabό &

Gălan 1966 (Syn.: Agrosteto-Festucetum sulcatae-vale-

siacae Szabό 1971); subass. agrostetosum tenuis (Máthe

& Kovács 1960) Soό 1964 (Syn.: Festuco-Agrostetum

tenuis Dihoru & al. 1970).

Ass. Cirsio cani-Festucetum pratensis Májovski &

Růžičková 1971

Synonyms: Deschampsietum caespitosae Hayek ex

Horvatič 1930 p. p.; Festucetum pratensis Soό 1938

(Art. 36) p. p.; Agrostio stoloniferae-Deschampsietum

caespitosae Ujvárosi 1947.

Diagnostic species: Alopecurus pratensis, Bromus

commutatus, Carex muricata agg., C. praecox, C. hir-

ta, Cirsium canum, Dactylis glomerata, Festuca prat-

ensis, Holcus lanatus, Jacea pratensis, Leucanthemum

vulgare, Lychnis flos-cuculi, Poa pratensis, P. t r i v i a l i s ,

Potentilla reptans, Rumex acetosa.

Distribution: Bulgaria, Hungary, Romania, Slova-

kia, Ukraine.

Ass. Cynosuro cristati-Trifolietum repentis Cortini

Pedrotti & al. 1973

Distribution: Italy.

Ass. Carici-Potentilletum reptantis Gamisans 1976

Distribution: Italy.

Ass. Ophioglosso vulgati-Nardetum strictae Gam-

isans 1976

Distribution: Italy.

Ass. Agrostio tenuis-Trifolietum repentis Walther

1977

Distribution: Ukraine.

Ass. Campanulo glomeratae-Cynosuretum cristati

Ubaldi 1978

Distribution: Italy.

Ass. Crepido capillaris-Festucetum rubrae Hülbusch

& Kiensat in Kienast 1978 nom. invers. propos.

Original name: Festuco-Crepidetum capillaris Hül-

busch & Kiensat in Kienast 1978.

Synonyms: Festuco rubrae-Crepidetum capillarim

Hülbusch & Kiensat in Kienast 1978; Bellidetum per-

ennis Gutte 1984; Trifolio repentis-Veronicetum fili-

formis N. Müller 1988.

Annex 1. Continuation.

126 Velev, N. • Checklist of Arrhenatheretalia order

Diagnostic species: Achillea millefolium agg.,

Agrostis capillaris, Cerastium holosteoides, Crepis capil-

laris, Festuca rubra agg., Hypochaeris radicata, Leonto-

don autumnalis, Lolium perenne, Plantago lanceolata,

P. major agg., Poa pratensis agg., Ranunculus repens,

Taraxacum Sect. Ruderalia, Trifolium dubium, T. re-

pens, Veronica filiformis, V. serpyllifolia.

Distribution: France, Germany.

Ass. Festuco rubrae-Deschampsietum Raţiu & Ger-

gely 1978 (Deschampsia caespitosa)

Diagnostic species: Deschampsia caespitosa, Festu-

ca rubra, Juncus articulatus, J. effusus, Nardus stricta,

Potentilla erecta, Veratrum album.

Distribution: Romania.

Ass. Galio veri-Festucetum rubrae Resmeriţă (1965)

1980

Synonyms: Festucetum rubrae xero-mezofilum

Resmeriţă 1965 nom. illeg. (Art. 34a).

Distribution: Romania.

Variations: subass. cytisetosum nigricantis Res meriţă

1970.

Ass. Luzulo campestris-Cynosuretum cristati (Meisel

1966) de Foucault 1981

Diagnostic species: Agrostis capillaris, Centaurea

jacea agg., Hypochaeris radicata, Lotus corniculatus,

Luzula campestris, Pilosella officinarum.

Distribution: Belgium, France (also in Brachypo-

dio-Centaureion nemoralis).

Ass. Eryngio campestris-Lolietum perennis Wattez 1982

Distribution: France.

Ass. Cynosuro cristati-Trifolietum repentis O. Bolòs

(1967) 1983

Synonyms: Cynosuretum pradense O. Bolòs 1967

nom. illeg. (Art. 34; 39); Cynoserutum cristati catalau-

nicum Vigo 1968 nom; illeg. (Art. 3b; 34).

Distribution: Iberian Peninsula.

Ass. Trifolietum repentis Shvergunova & al. 1984

Synonyms: Leontodonti-Trifolietum repentis (Stein-

dorsson 1945) R. Tx. 1969; incl. Junco compressi-Tri-

folietum repentis Eggler 1933, Blysmo-Juncetum com-

pressi R. Tx. 1950.

Diagnostic species: Trifolium repens.

Distribution: Belarus.

Variations: subass. agrostidetosum tenuis sensu

Stepanovich 2000; subass. cynosuretosum cristati sen-

su Stepanovich 2000; subass. lolietosum perenni sen-

su Stepanovich 2000; subass. poetosum pratensis sensu

Stepanovich 2000.

Ass. Agrostietum vinealis-tenuis Shelyag & al. 1981

ex Shelyag & al. 1985

Diagnostic species: Agrostis capillaris, A. vinealis,

Koeleria delavignei, Potentilla argentea.

Distribution: Ukraine.

Variations: var. typicum; var. Betonica officinalis;

var. Festuca pratensis.

Ass. Carici arenarie-Luzuletum campestris de Fou-

cault 1986

Distribution: France.

Ass. Festuco amplae-Cynosuretum cristati Rivas-

Martínez ex Fuente 1986

Distribution: Iberian Peninsula.

Variations: Spain: var. Juncus acutiflorus; var. Jun-

cus squarrosus.

Ass. Centaurio-Narcissetum angustifolii Kriczfa-

lushij & Gendey 1987

Distribution: Ukraine.

Ass. Polytricho-Narcissetum angustifolii Kriczfa-

lushij & Gendey 1987

Distribution: Ukraine.

Ass. Trifolio repentis-Veronicetum filiformis N. Mül-

ler 1988

Diagnostic species: Veronica filiformis.

Distribution: Austria.

Ass. Achilleo collinae-Cynosuretum cristati Biondi &

al. 1989

Distribution: Italy.

Ass. Caro carvi-Cynosuretum cristati (Pfadenhauer

1969) de Foucault 1989

Distribution: France.

Ass. Eryngio campestris-Lolietum perennis Julve

1989 nom. ined. (Art. 1)

Distribution: France.

Ass. Leontodono-Poetum pratensis Anishczenko &

L. Ishb. 1989

Variations: var. typicum; var. Elytrigia repens; var.

Festuca rubra;.

Distribution: Ukraine.

Ass. Taraxaco-Festucetum pratensis Anishczenko &

L. Ishb. 1989

Distribution: Russia.

Ass. Plantagini lanceolate-Cynosuretum cristati Co-

quillard 1993

Distribution: France.

Ass. Colchico lusitani-Cynosuretum cristati Biondi

& Ballelli 1995

Distribution: Italy.

Annex 1. Continuation.

127Phytol. Balcan. 24(1) • Sofia • 2018

Ass. Gentiano luteae-Cynosuretum cristati (de Fou-

cault 1986) Gillet in Gallandat & al. 1995

Distribution: Belgium, France.

Ass. Carici ornithopodae-Agrostietum capillaris Vil-

legas 1997

Distribution: Iberian Peninsula.

Ass. Elytrigietum repentis Stepanovič (1987) 1999

Synonyms: Rumici crispi-Agropyretum repentis He-

jný in Hejný & al. 1979; incl. Agropyro repentis-Aego-

podietum podagrariae R. Tx. 1937, Agropyro-Alopecu-

retum Moravec 1965.

Diagnostic species: Elytrigia repens.

Distribution: Belarus.

Variations: subass. agrostidetosum tenuis sensu

Stepanovich 2000; subass. aperetosum spica-ventii

sensu Stepanovich 2000; subass. bromopsidetosum

inermis sensu Stepanovich 2000; subass. dactyli-

detosum glomeratae sensu Stepanovich 2000; sub-

ass. festucetosum pratensis sensu Stepanovich 2000;

subass. poetosum pratensis sensu Stepanovich 2000;

subass. rumecetosum crispi sensu Stepanovich 2000;

subass. trifolietosum pratensi sensu Stepanovich

2000.

Ass. Poetum pratensis Stepanovič 1999

Remarks: for details see in Festucion pratensis Si-

paylova & al. 1985.

Ass. Pastinaco hirsutae-Festucetum nigrescentis

Dimitrov 2001

Diagnostic species: Armeria rumelica, Pastinaca

hirsuta, Silene roemeri.

Distribution: Bulgaria.

Variations: subass. typicum; subass. verbascetosum

longifolii Dimitrov 2001 with two variants – var. Gali-

um verum, var. Dianthus armeria.

Ass. Euphorbio cyparissiae-Festucetum rubrae Didi-

er & Royer in Royer & al. 2006

Distribution: France.

Ass. Gentiano luteae-Cynosuretum cristati de Fou-

cault & Gillet in Ferrez 2007

Diagnostic species: Agrostis capillaris, Alchemilla

monticola, Cynosurus cristatus, Gentiana lutea, Lolium

perenne, Sanguisorba minor, Stachys officinalis.

Distribution: France.

Ass. Centaureo calcitrapae–Lolietum perennis Blasi

& al. 2009

Diagnostic species: Anthemis arvensis, Carduus

nutans, Centaurea calcitrapa, Cerastium ligusticum.

Distribution: Italy.

Ass. Trifolio molinerii–Hordeetum bulbosi Blasi &

al. 2009

Diagnostic species: Hordeum bulbosum, Mentha

spicata, Trifolium incarnatum ssp. molinieri, T. stria-

tum, Vulpia ligustica.

Distribution: Italy.

Ass. Cirsio arvensis-Alopecuretum pratensis Catteau

in de Foucault & Catteau 2012

Distribution: France (in Ranunculo-Cynosurion

cristati Passarge 1969).

Ass. Festuco commutatae-Lolietum nom. nud. sensu

Šilc & Čarni 2012

Distribution: Slovenia.

Ass. Loto pedunculati-Cynosuretum cristati (Tüxen

1937) de Foucault & Julve in de Foucault & Catteau

2012

Distribution: France (in Ranunculo-Cynosurion

cristati Passarge 1969).

Ass. Loto alpini-Festucetum morisianae Farris & al.

2013

Distribution: Italy.

Ass. Phleo nodosi-Cynosuretum cristati Rodríguez-

Rojo & Fernández-González 2014

Diagnostic species: Bellis perennis, Cynosurus cris-

tatus, Phleum nodosum, Trifolium repens.

Distribution: Iberian Peninsula.

Ass. Colchico autumnalis-Festucetum rupicolae

Lengyel, Csiky, Dénes & Király in Lengyel & al. 2016

Distribution: Hungary.

Plant communities without rank related to Cyno-

surion:

Plant community Cynosurus cristatus-Trifolium

pratense Buzo 1990

Distribution: Albania.

Plant community Lolium perenne-Trifolium resupi-

natum Buzo 1990

Distribution: Albania.

Plant community Festuca ovina-Avenula pratensis

sensu Zuidhoff & al. 1995

Distribution: Belgium, France, Germany, Nether-

lands, Switzerland, United Kingdom (in Mesobromion).

Plant community Plantago major-Trifolium repens

sensu Oberdorfer 1993

Corresponding names: Lolio-Plantaginetum majo-

ris Sissingh 1969 non Beger 1930.

Distribution: Germany.

Annex 1. Continuation.

128 Velev, N. • Checklist of Arrhenatheretalia order

Plant community Plantago major-Trifolium repens

sensu Dierschke 1997

Corresponding names: Lolio-Plantaginetum ma-

joris Sissingh 1969; Lolietum perennis Mucina & al.

1993.

Remarks: According to Päzolt & Jansen (2004)

plant community Plantago major-Trifolium repens

sensu Dierschke 1997 is related to Plantagini majoris-

Lolietum perennis Beger 1932.

Distribution: Germany.

Plant community Alopecurus pratensis-Sanguisorba

officinalis sensu Rodwell & al. 2000

Distribution: United Kingdom.

Plant community Lerchenfeldia flexuosa sensu Pav-

lov & al. 2006

Distribution: Bulgaria.

Plant community Lolium perenne sensu Meleghik &

al. 2008

Distribution: Ukraine.

Plant community Poa pratensis sensu Meleghik &

al. 2008

Distribution: Ukraine.

References: Puşcaru & al. (1956); Csűrös &

Resmeriţă (1960); Horvatić (1963); Puşcaru–Soro-

ceanu (1963); Medwečka-Kornaš & al. (1966); Pop

(1968); Csűrös & al. (1969); Jurko (1969a, b; 1974); Ger-

gely (1969); Csűrös (1970); Csűrös-Káptalan (1970);

Marković (1970); Resmeriţă (1970); Trzcińska-Tacik

& al. (1973); Dihoru (1975); Blečić & Lakušić (1976);

Speidel (1979); Balátová-Tuláčková (1985); Petković

(1985); Mititelu & al. (1986, 1988); Mititelu & Dor-

ca 1987; Coldea (1987, 1990); Pop & al. (1988); Sýko-

ra & al. (1988); Reif & al. (1989); Sanda & Popescu

(1991); Rodwell (1992); Venanzoni (1992); Ellmauer

& Mucina (1993); Ivan & al. (1993); Ellmauer (1995);

Kovács (1995, 2004); Zuidhoff & al. (1995); Diersch-

ke (1997, 1999); Sanda & al. (1998, 1999); Kojić &

al. (1998); Gèhu (1999); Jermacāne (1999); Ninot &

al. (2000); Stančić (2000); Stepanovich (2000); Gon-

charenko (2001); Dimitrov (2001, 2004, 2015); Bardat

& al. (2001); Demeter & Veen (2001); Miazga & Mo-

sek (2001); Izco & al. (2001); Jansen & Päzolt (2001);

Jermacāne & Laiviņš (2001); Mihăilescu (2001); Ri-

vas-Martínez & al. (2001); Soriano (2001); Dring &

al. (2002); Pop & al. (2002); Snowarski (2002-2005);

Chytrý & Tichý (2003); Coste & Arsene (2003); Gli-

ga (2003); Neblea & Chiriţoiu (2003); Zamfirescu &

Zamfirescu (2003); Bardat & al. (2004); Drăgulescu

(2004); Grigoriu & Alda (2004); Päzolt & Jansen

(2004); Rodríguez-Rojo & Sánchez-Mata (2004); Sar-

bu & al. (2004); Lakušić & al. (2005); Meshinev & al

(2005); Pavlov & al. (2006); Apostolova & Meshiunev

(2006); Bărbos (2006); Grzelak & Bocian (2006); Mo-

sek & Miazga (2006); Rūsiņa (2006a, b); Trąba & al.

(2006); Astakhova (2007); Hájková & al. (2007); Gro-

za & al. (2007); Janišová & al. (2007, 2010); Rodwell &

al. (2007); Zaliberová & Škodová (2007); Mikhailovna

(2008); Gafta & Mountford (2008); Milkina & Ljach

(2008); Pawlaczyk (2008); Sendžikaitė & al. (2008);

Jarolímek & al. (2008b); Meleghik & al. (2008);

Škodová & Janišová (2008); Sopotlieva (2008); To-

mescu & al. (2008); Trąba & al. (2008); Blasi & al.

(2009, 2012); FloraWeb (accessed 26.03.2009); Kala-

jnxhiu & al. (2009); Kuzemko (2009b); Mounford &

Akeroyd (2009); Tzonev & al. (2009); Wolański &

Trąba (2009); Becker & Becker (2010); Chocarro &

al. (2010); Dúbravková & al. (2010); Collaud (2011);

Velev & al. (2011a); Šilc & Čarni (2012); Aćić & al.

(2013b); Farris & al. (2013); Kozyr (2013); Tasen-

kevich & Kalinovych (2013); Willner & al. (2013a,

b); Loidi & al. (2014); Rodríguez-Rojo & Fernán-

dez-González (2014); Rodríguez-Rojo & al. (2014);

Shushpannikova & Yamalov (2014); Vassilev &

al. (2014); Fernez & Causse (2015); Škodová & al.

(2015); Maćkowiak (2016); Zajac & al. (2016); Rod-

ríguez-Rojo & al. (2017).

Alliance POLYGONO BISTORTAE-TRISETION

FLAVESCENTIS Br.-Bl. & Tüxen ex Marschall 1947

nom. invers.

Original name: Triseto-Polygonion bistortae Mar-

schall 1947 (Trisetum flavescens; Polygonum bistorta =

Bistorta major).

Synonyms: Triseto flavescentis-Polygonion bistor-

tae Br.-Bl. & Tüxen 1943 nom. nud. (Art. 2; 8); Tri-

seto flavescentis-Polygonion bistortae Br.-Bl. & Tüxen

1943 ex Marschall 1947; Triseto-Polygonion bistortae

Br.-Bl. 1949 ex 1950; Rumici-Trisetion flavescentis Pas-

sarge 1969 (Art. 29c); Phyteumato-Trisetion (Passarge

1969) Ellmauer & Mucina in Mucina & al. 1993.

Corresponding names: Campanulo-Trisetenion

Dierschke 1981 nom. inval. (Art. 2b; 5;); Phyteumato-

Trisetenion Passarge 1969; Campanulo rhomboidalis-

Trisetenion flavescentis Dierschke in Theurillat 1992;

Poo alpinae-Trisetetalia Mucina & al. 1993.

Annex 1. Continuation.

129Phytol. Balcan. 24(1) • Sofia • 2018

Remarks: Mucina & al. (2016), excluded Phyteu-

mato-Trisetion (Passarge 1969) Ellmauer & Mucina in

Mucina & al. 1993 alliance from the synonymy and

presented it as a separate syntaxon. The Plygono bis-

tortae-Trisetion alliance is related to Poo alpinae-Tri-

setetalia Ellmauer & Mucina 1993 order.

Diagnostic species: Agrostis capillaris, Alchemil-

la spec. div., Anemone nemorosa, Anthoxanthum od-

oratum, Anthriscus nitidus, Arnica montana, Astran-

tia major, Bistorta major, Campanula glomerata agg.,

Campanula gentilis, C. moravica, C. rhomboidalis, C.

rotundifolia aggr., C. serrata, C. scheuchzeri, Cardami-

nopsis halleri, Carum carvi, Centaurea oxylepis, C.

phrygia, C. phrygia ssp. carpatica, C. pseudophrygia, C.

transalpina, Chaerophyllum hirsutum, C. villarsii, Cir-

sium erisithales, C. heterophyllum, Crepis aurea, C. co-

nyzifolia, C. mollis, C. pyrenaica, C. succisifolia, Crocus

albiflorus, C. discolor, C. heuffelianus, C. vernus, Cru-

ciata glabra, Dactylis glomerata, Deschampsia caespi-

tosa, Festuca diffusa, F. nigrescens, F. rubra aggr., Gali-

um saxatile, Gentianella austriaca, Geranium phaeum,

G. pratense, G. sylvaticum, Gladiolus imbricatus, Her-

acleum sphondilium, Hypericum maculatum, Knautia

dipsacifolia, K. longifolia, K. maxima, Lilium bulbifer-

um, Lychnis flos-cuculi, Meum athamanticum, Muscari

botryoides, Narcissus angustifolius, N. exsertus, N. radi-

iflorus (=N. poeticus ssp. radiiflorus), Noccaea caerule-

scens ssp. caerulescens (=Thlaspi caerulescens), Phleum

alpinum, P. rhaeticum, Phyteuma nigrum, P. orbiculare,

P. ovatum, P. spicatum, P. tetramerum, Pimpinella ma-

jor, Poa alpina, P. chaixii, Polygonum bistorta, Potentil-

la aurea, P. erecta, Primula elatior, Ranunculus carpat-

icus, R. polyanthemos agg., Rhinanthus alectorolophus,

R. pulcher, R. rumelicus, Rumex alpestris, R. arifolius,

Sanguisorba officinalis, Senecio subalpinus, Silene dioi-

ca, Stellaria graminea, Trisetum flavescens, Traunstein-

era globosa, Trifolium pratense, Trollius europaeus, Ve-

ronica chamaedrys, Vicia sepium, Viola declinata, V.

tricolor.

References: Borza (1959); Moravec (1965); Jurko

(1974); Dihoru (1975); Blažková (1985); Boşcaiu &

Täuber (1985); Ellenberg (1988); Sanda & Popescu

(1991); Oberdorfer (1993); Dierschke (1997, 1999);

Krahulec & al. (1997); Sanda & al. (1999); Pop & al.

(1988); de Foucault (1989); Mihăilescu (2001); Ri-

vas-Martínez & al. (2001); Chytrý & Tichý (2003);

Ružičková (2002); Snowarski (2002-2005); Burkart

& al. (2004); Tomescu & al. (2006); Apostolova & al.

(2007); Hájková & al. (2007); Hegedüšová & Ružičková

(2007); Solomakha (2008); Jarolímek & al. (2008a; b);

Škodová & Janišová (2008); Biondi & Blasi (2013); Bi-

ondi & al. (2014); Mucina & al. (2016).

Suballiances related to alliance Polygono-Trisetion:

Suball. Lathyro linifolii-Trisetenion flavescentis Di-

erschke 1981 nom. inval.

Original name: Lathyro linifolii-Trisetenion Dier-

schke 1981 nom. inval. (Art. 2d; 5).

Suball. Campanulo rhomboidalis-Trisetenion flaves-

centis Dierschke in Theurillat 1992

Original name: Campanulo-Trisetenion Dierschke

1981 nom. inval. (Art. 2d; 5).

Suball. Violo sudeticae-Trisetenion flavescentis Julve

1993 nom. inval.

Original name: Violo sudeticae-Trisetenion flaves-

centis Julve 1993 nom. inval. (Art. 2d; 3b).

Suball. Alchemillo monticolae-Trisetenion flaves-

centis Ferrez 2007

Associations related to alliance Polygono-Trisetion:

Ass. Anthoxantho odorati-Agrostietum tenuis Sill-

inger 1933

Remarks: for details see in alliance Cynosurion.

Ass. Meo athamantici-Festucetum rubrae Bartsch &

Bartsch 1940

Original name: Festuca rubra-Meum athamanti-

cum Bartsch & Bartsch 1940.

Synonyms: Meo-Festucetum Tüxen 1937; Meo-Fes-

tucetum rubrae Bartsch & Bartsch 1940; Geranio-Tri-

setetum flavescenstis Knapp 1951; Meo-Festucetum

Oberdorfer 1957; Alopecuro-Poëtum chaixii Blažková

1991; Meo athamantici-Cirsietum heterophylli

Blažková 1991; plant community Festuca rubra-Meum

sensu Dierschke 1997; plant community Festuca ru-

bra-Meum athamanticum sensu FloraWeb (accessed

26.03.2009).

Remarks: The synonym Geranio-Trisetetum flaves-

censtis Knapp 1951, mentioned as such by Snowarski

(2002-2005), makes Meo athamantici-Festucetum

rubrae Bartsch & Bartsch 1940 a synonym of Geranio

sylvatici-Trisetetum flavescentis Knapp ex Oberdorfer

1957. A conformation of this is the fact, that Meo-

Festucetum Oberdorfer 1957 is considered as synonym

Annex 1. Continuation.

130 Velev, N. • Checklist of Arrhenatheretalia order

of both Geranio sylvatici-Trisetetum flavescentis

(Dier schke 1997) and Meo athamantici-Festucetum

rubrae Bartsch & Bartsch 1940 [FloraWeb (accessed

26.03.2009)].

Diagnostic species: Alchemilla sp. div., Campanula

rotundifolia agg. (C. gentilis, C. moravica, C. rotundifo-

lia), Cirsium heterophyllum, Crepis mollis, Festuca ru-

bra agg., Galium saxatile, Hypericum maculatum, Me-

um athamanticum.

Distribution: Czech Republic, France, Germany,

Poland.

Ass. Heracleo lecoquii-Trisetetum flavescentis (Sus-

plugas 1935) Br.-Bl. 1948

Distribution: France.

Ass. Astrantio-Trisetetum G. Knapp & R. Knapp

1952 ex Oberdorfer 1957

Synonyms: Astrantio-Trisetetum G. Knapp & R.

Knapp 1952 nom. prov. (Art. 3b); Rumici-Trisetetum

Passarge 1969 (Art. 29); Astrantio-Brometum Machold

1991 p. p. (Art. 1).

Remarks: Astrantio majoris-Trisetetum flavescentis

(Dutoit 1924) Knapp 1951 is mentioned for France.

Diagnostic species: Carex montana, Listera ovata.

Distribution: Austria, France? [as Astrantio major-

is-Trisetetum flavescentis (Dutoit 1924) Knapp 1951],

Germany, Romania, Slovenia, Ukraine.

Variations: Slovenia: subass. typicum, subass.

buphthalmetosum Knapp & Knapp 1952, subass. trolli-

etosum Knapp & Knapp 1952.

Ass. Geranio sylvatici-Trisetetum flavescentis Knapp

ex Oberdorfer 1957

Original name: Geranio-Trisetetum Knapp 1951

(Geranium sylvaticum; Trisetum flavescens).

Synonyms: Trisetetum flavescentis Rübel 1911 sen-

su lato; Festucetum rubrae-fallacis Rübel 1911; Agrosti-

etum tenuis Rübel 1912; Trisetetum flavescentis Be-

ger 1922 (Art. 31); Agrostidetum vulgaris Szafer & al.

1923 (Art. 36); Agrostidetum vulgaris Szafer & al. 1927

(Art. 31); Trisetetum flavescentis Tüxen 1937; Tris ete-

tum meetosum athamantici Tüxen 1937; Meo-Trise-

tetum flavescentis Tüxen 1937; Festuca rubra-Meum

athamanticum J. & M. Bartsch 1940; Meo-Festuce-

tum Bartsch 1940; Trisetetum crepidetosum mollis

Moor 1942; Trisetum flavescens-Gearnium sylvaticum

Knapp 1951 (Art. 3b; 36); Astrantio-Trisetetum Ober-

dorfer 1957 p. p.; Meo-Festucetum Oberdorfer 1957;

Geranio (sylvatici)-Trisetetum Knapp ex Oberdorfer

1957; Geranio-Trisetetum Knapp ex Oberdorfer 1957;

Cirsio heterophylli-Trisetetum Hundt 1964; Centaureo

nigrae-Trisetetum Hundt 1964; Trisetetum flavescen-

tis noricum Soó 1964; Laserpitio latifoliae-Trisetetum

Hundt 1964; Trisetetum flavescentis Hundt 1964; 1969;

Melandrio-Trisetetum Moravec 1965; Cardaminop-

sido halleri-Agrostietum (Pawłowski & Walas 1949)

Moravec 1965; Cardaminopsio halleri-Agrostietum

Moravec 1965; Meo-Trisetetum Dierschke & Vogel

1981. According to internet site FloraWeb (accessed

26.03.2009) incl.: Meo-Trisetetum Tüxen 1970; Astran-

tio-Trisetetum Oberdorfer p. p.; According to Ellmau-

er & Mucina (1993) non: Trisetetum leponticum Mar-

schall 1947 (Art. 34).

Diagnostic species: Agrostis capillaris (also as dom-

inant), Alchemilla monticola, A. vulgaris agg., Alope-

curus pratensis, Anthriscus sylvestris, Arnica montana,

Bistorta major, Campanula rotundifolia agg. (C. genti-

lis, C. moravica, C. rotundifolia), C. patula, Cardami-

nopsis halleri, Carum carvi, Centaurea pseudophrygia,

Chaerophyllum aromaticum, Crepis biennis, C. mollis,

Crocus discolor, Bellis perennis, Deschampsia caespi-

tosa, Geranium sylvaticum, Lychnis flos-cuculi, Her-

acleum sphondylium, Hypericum maculatum, Pimp-

inella major, Phleum alpinum, P. pratense, Phyteuma

nigrum, P. spicatum, Rumex arifolius, Senecio auranti-

acus, Taraxacum Sect. Ruderalia, Trisetum flavescens,

Vicia cracca, V. sepium, Viola tricolor ssp. saxatilis.

Remarks: 1) Dierschke (1997) pointed Festuca ru-

bra-Meum athamanticum J. & M. Bartsch 1940 as syn-

onym. Thus Geranio sylvatici-Trisetetum flavescen-

tis Knapp ex Oberdorfer 1957 becomes a synonym of

Meo athamantici-Festucetum rubrae Bartsch & Bar-

tsch 1940; 2) Hegedüšová & Ružičková (2007) point-

ed Geranio sylvatici-Trisetetum flavescentis Knapp ex

Oberdorfer 1957 as a syntaxonomical synonym of

Agrostidetum vulgaris Szafer & al. 1923. The latter (ac-

cording to Medwečka-Kornaš & al. 1966) is given as

synonym of Gladiolo imbricati-Agrostietum capillaris

(Br.-Bl. 1930) Pawłowski & Walas 1949. Thus all three

associations become synonyms; 3) Oberdorfer (1993)

included Trisetetum flavescentis Tüxen 1937 in a big-

ger one – Poo-Trisetetum flavescentis Knapp 1951. On

the other hand, Dierschke (1997) pointed Trisetetum

flavescentis Tüxen 1937 as synonym of Geranio sylvat-

ici-Trisetetum flavescentis Knapp ex Oberdorfer 1957;

4) Ellmauer & Mucina (1993) consider Trisetetum fla-

vescentis Hundt 1964 as a separate syntaxon, not syn-

onym.

Annex 1. Continuation.

131Phytol. Balcan. 24(1) • Sofia • 2018

Distribution: Austria, Belgium, Czech Republic,

France, Germany, Hungary, Poland, Romania, Slova-

kia, Ukraine.

Variations: Austria: subass. cirsietosum hetrophylli

Dierschke ex Ellmauer 1995; Czech Republic: subass.

bellidetosum Blažková in Krahulec & al. 1997 (Bellis

perennis); subass. lathyretosum Blažková in Krahulec

& al. 1997 (Lathyrus pratensis); subass. melandrieto-

sum Blažková in Krahulec & al. 1997 (Melandrium

rubrum); subass. nardetosum Blažková in Krahulec

& al. 1997 (Nardus stricta); subass. pimpinelletosum

Blažková in Krahulec & al. 1997 (Pimpinella saxifra-

ga); var. Deschampsia caespitosa; var. Trifolium repens;

fac. alopecurosum pratensis; fac. dactyliosum glomer-

atae; fac. holcosum mollis-hypericosum maculatea;

Hungary: subass. arrhenatheretosum; subass. festuce-

tosum rubrae.

Ass. Poo-Trisetetum flavescentis Knapp ex Oberdor-

fer 1957

Remarks: for details see in Arrhenatherion.

Ass. Triseto flavescentis-Heracleetum pyrenaici Br.-

Bl. ex O. Bolòs 1957

Synonyms: Astrantio-Avenuletum pubescentis Vi-

go 1984; Triseto-Heracleetum pyrenaici Br.-Bl. in Mar-

schall 1947 (Art. 2b); Campanulo patulae-Violetum

cornutae Nègre 1972.

Pseudonyms: Heracleo-Trisetetum auct.

Distribution: France, Iberian peninsula.

Variations: Spain: subass. typicum; subass. astranti-

etosum majoris Vigo 1996 (Corresp. name: Astrantio-

Avenuletum pubescentis Vigo 1984).

Ass. Alchemillo filicaulis-Trisetetum flavescentis

Lambert 1962

Distribution: France.

Ass. Campanulo rectae-Agrostietum capillaris

(Braun 1915) Hundt 1964

Distribution: France.

Ass. Trisetum flavescens-Festuca rubra Soό 1944;

Şerbănescu 1964

Distribution: Romania.

Ass. Leontodo-Festucetum Carbiener 1966

Distribution: Belgium, France.

Ass. Alchemillo-Festucetum rubrae Eggler

Homonyms: Alchemillo-Festucetum rubrae

Resmeriţă 1972 nom. illeg. (Art. 31).

Distribution: Poland.

Ass. Geranio-Alchemilletum crinitae Hadač & al.

1969

Original name: Geranio-Alchemilletum crinitae

Hadač & al. 1969 (Geranium phaeum; G. sylvaticum).

Synonyms: Alchemilleto-Deschampsietum caespi-

tosae Bareš & Hadač 1958 (Art. 2b); Alchemilleto-

Festucetum pratensis Bareš & Hadač 1958 (Art. 2b);

Alchemillo-Deschampsietum caespitosae Hadač &

Smola 1962 (Art. 2b); Alchemilleto-Festucetum prat-

ensis Hadač & Smola 1962 (Art. 2b); Rhinantho-Al-

chemilletum monticolae Hadač & Smola 1962 (Art.

2b); Geranio-Alchemilletum crinitae Hadač & Smola

1962 (Art. 2b); Rhinantho-Alchemilletum monticolae

Hadač & al. 1969 (Art. 3b); Alchemillo-Deschampsie-

tum caespitosae Hadač & al. 1969; Rhinantho-Alche-

milletum monticolae Hadač & al. 1969; Alchemillo-Fes-

tucetum pratensis Hadač & al. 1969; Geo-Dactylidetum

slovenicae Hadač 1981 (Art. 34a); Hyperico-Deschamp-

sietum caespitosae Hadač 1981; Rhinantho pulchri-Al-

chemilletum monticolae Hadač & al. ex Kliment 1994.

Pseudonyms: Alchemilletum pastoralis sensu

Šmarda & al. 1963; 1971 p. p. non Szafer & al. 1927;

Deschampsietum caespitosae sensu Unar & al. 1984

non Krajina 1933; Deschampsietum caespitosae typi-

cum sensu Unar & al. 1984 non Krajina 1933.

Diagnostic species: Acetosa arifolia, Bistorta major,

Campanula serrata, Conioselinum tataricum, Crepis

mollis, Dianthus superbus, Geranium sylvaticum, Hy-

pericum maculatum, Knautia maxima, Phyteuma spi-

catum, Pimpinella major, Potentilla aurea, Rhinanthus

pulcher, Senecio subalpinus, Trollius altissimus, Viola

biflora.

Distribution: Slovakia.

Variations: var. Pimpinella major ssp. rhodochla-

mys; var. Trisetum flavescens.

Ass. Meo athamantici-Trisetetum flavescentis La-

coste 1975

Distribution: France.

Ass. Phyteumo orbicularis-Trifolietum pratensis

Balcerk. 1978

Diagnostic species: Carex sempervirens, Phyteuma

orbiculare, Polygala amara ssp. brachyptera, Primula

elatior, Ranunculus oreophilus.

Distribution: Poland.

Ass. Geranio lividi-Trisetetum Knapp & Knapp ex

Dierschke 1981

Synonyms: Trisetum flavescens-Geranium lividum

Knapp & Knapp 1954 (Art. 3b).

Diagnostic species: Geranium phaeum ssp. livi-

dum, Trisetum flavescens.

Annex 1. Continuation.

132 Velev, N. • Checklist of Arrhenatheretalia order

Distribution: Austria.

Ass. Euphorbio brittingeri-Trisetetum flavescentis

de Foucault 1986

Distribution: Belgium, France.

Variations: Belgium: subass. typicum; subass. bro-

metosum erecti; subass. brometosum mollis; France:

subass. typicum; subass. brometosum erecti de Fou-

cault in Ferrez 2007; subass. brometosum mollis; sub-

ass. kanutietosum godeti Ferrez 2007; subass. trollieto-

sum europaei Ferrez 2007.

Ass. Violo cornutae-Trisetetum flavescentis (Nègre

1972) Gruber 1986

Distribution: France.

Ass. Violo luteaea-Trisetetum flavescentis (Luquet

1926) de Foucault 1986

Distribution: France.

Ass. Centaureo transalpinae-Trisetetum flavescentis

(Marschall 1947) Poldini & Oriolo 1994

Synonyms: Trisetetum insubricum Marsch. 1947 (Art.

34); Trisetetum leponticum Marschall 1947 (Art. 34).

Pseudonyms: Trisetetum flavescentis Pedrotti 1964

non Brockmann-Jerosch 1907; Trisetetum flavescen-

tis Gerdol & Piccoli 1980 non Beger 1923; Trisetetum

flavescentis Buffa & al. 1989 non Brockmann-Jerosch

1907.

Distribution: Italy.

Ass. Melandrio rubri-Phleetum alpini Blažková in

Krahulec & al. 1997

Original name: Melandrio-Phleetum alpini

Blažkova in Krahulec & al. 1997 (Melandrium rub-

rum = Silene dioica; Phleum alpinum = P. rhaeticum).

Diagnostic species: Alchemilla vulgaris s. lat., Bis-

torta major, Campanula bohemica, Cardaminopsis

halleri, Crepis conyzifolia, C. mollis, Geranium sylvat-

icum, Hypericum maculatum, Omalotheca sylvatica,

Phleum rhaeticum, Phyteuma spicatum, Poa chaixii,

Potentilla aurea, Silene dioica, S. vulgaris, Viola lutea

ssp. sudetica.

Distribution: Czech Republic.

Variations: subass. alopecuretosum Blažková in

Krahulec & al. 1997 (Alopecurus pratensis); subass.

nardetosum Blažková in Krahulec & al. 1997 (Nardus

stricta) with the following variants – var. typicum, var.

Luzula luzuloides, var. Trisetum flavescens, var. Vio-

la lutea, var. Viola lutea ssp. sudetica (Corresp. name:

Melandrio rubri-Phleetum alpini Blažková in Krahulec

& al. 1997 subass. violetosum luteae Blažková in Kra-

hulec & al. 1997).

Ass. Alchemillo xanthochlorae-Trollietum europaei

Vigo ex Vigo in Ninot & al. 2000

Original name: Alchemillo-Trollietum Vigo 1979

nom. illeg. (Art. 5).

Synonyms: Delphinio montani-Trollietum europaei

Br.-Bl. 1948 (Art. 3b); Violo cornutae-Euphorbietum

hybernae Romo 1986 p. p. (Art. 37).

Distribution: Iberian Peninsula.

Variations: Spain: subass. typicum; subass. rumi-

cetosum amplexicaulis Carrereras & al. in Ninot & al.

2000 (Basyonym: Alchemillo-Trollietum Vigo 1979

subass. rumicetosum amplexicaulis Carrereras & al.

1993 nom. invall.)

Ass. Campanulo glomeratae-Geranietum sylvatici

Ružičková 2002

Diagnostic species: Anthyllis vulneraria, Aquilegia vul-

garis, Arrhenatherum elatius, Campanula glomerata agg.,

C. serrata, Carlina acaulis, Cirsium erisithales, Colchicum

autumnale, Crocus discolor, Dianthus carthusianorum agg.,

Geranium sylvaticum, Jacea phrygia agg. (J. phrygia, J. pseu-

dophrygia, J. stenolepis), Knautia arvensis agg., Leontodon

hispidus, Lilium bulbiferum, Pimpinella major, Polygala

vulgaris, Potentilla heptaphylla, Primula elatior, Pyrethrum

corymbosum, Silene nemoralis, S. vulgaris, Tragopogon ori-

entalis, Trifolium montanum, Trisetum flavescens.

Distribution: Slovakia.

Variations: subass. typicum; subass. brometosum erec-

ti Ružičková 2002; var. Bromus erectus; var. Luzula luzu-

loides.

Ass. Crepido mollis-Agrostietum capillaris Ružič-

ková 2004

Original name: Crepido mollis-Agrostietum Ružič-

ková 2004 (Agrostis capillaris).

Diagnostic species: Cardaminopsis halleri, Crep-

is mollis, Jacea phrygia agg. (J. phrygia, J. pseudophrygia,

J. stenolepis), Luzula luzuloides, Physostegia virginiana,

Phyteuma spicatum, Pilosella aurantiaca, Poa chaixii,

Primula elatior, Ranunculus polyanthemos, Traunstein-

era globosa, Trifolium medium agg. Tromsdorfia uniflora.

Distribution: Slovakia, Ukraine.

Ass. Alchemillo monticolae-Brometum mollis Ferrez

2007

Diagnostic species: Alchemilla monticola, A. xan-

thochlora, Anthriscus sylvestris, Bromus hordeaceus ssp.

hordeaceus, Geranium sylvaticum, Heracleum sphon-

dylium ssp. sphondylium, Polygonum bistorta, Trisetum

flavescens ssp. flavescens.

Distribution: France.

Annex 1. Continuation.

133Phytol. Balcan. 24(1) • Sofia • 2018

Plant communities without rank related to alliance

Polygono-Trisetion:

Plant community Trisetum flavescens-Cynosurus

cristatus Buzo 1990

Distribution: Albania.

Plant community Festuca rubra-Agrostis tenuis sen-

su Dierschke 1997 non Glavač 1983

Distribution: Germany.

Plant community Anthoxanthum odoratum-Gerani-

um sylvaticum sensu Rodwell & al. 2000

Distribution: United Kingdom.

References: Moravec (1965); Borza (1966); Csűrös-

Káptalan (1970); Dihoru (1975); Reif & al. (1989);

Sanda & Popescu (1991); Oberdprfer (1993); Poldini

& Oriolo (1994); Ellmauer (1995); Kovács (1995); Di-

erschke (1997); Krahulec & al. (1997); Gèhu (1999);

Sanda & al. (1999); Ninot & al. (2000); Rodwell & al.

(2000); Rivas-Martínez & al. (2001); Soriano (2001);

Dring & al. (2002); Pop & al. (2002); Ružičková

(2002); Bardat & al. (2004); Sarbu & al. (2004);

Snowarski (2002-2005); Slivinska & Balashov (2006);

Ferrez (2007); Hájková & al. (2007); Hegedüšová &

Ružičková (2007); Solomakha (2008); Jarolímek & al.

(2008b); Škodová & Janišová (2008); FloraWeb (ac-

cessed 26.03.2009); Wolański & Trąba (2009); Chocar-

ro & al. (2010); Petras Sackl & al. (2012); Šilc & Čarni

(2012); Tasenkevich & Kalinovych (2013); Rodríguez-

Rojo & al. (2014); Budzhak & al. (2016).

Alliance POION ALPINAE Gams ex Oberdorfer 1950

Original name: Poion alpinae Gams 1936 nom.

nud. (Art. 2b; 8).

Synonyms: Poion alpinae Rübel 1933 (Art. 2b; 8);

Achilleo-Poion alpinae (Gams ex Oberd. 1950) Pas-

sarge 1969; Astero bellidiastri-Poion alpinae Passarge

1969; Carlino-Poion alpinae Passarge 1969; Eu-Achil-

leo-Poion alpinae Passarge 1969; Eu-Carlino-Poion al-

pinae Passarge 1969; Eu-Rumici-Poion alpinae Pas-

sarge 1969; Phleo alpini-Poion alpinae Passarge 1969;

Ranunculo-Poion alpinae Passarge 1969.

Corresponding names: Poo-Phleetalia alpini Pas-

sarge 1969; Trifolienalia Oberdorfer 1990 p. p.; Poo al-

pinae-Trisetetalia Mucina & al. 1993 p. p.

Remarks: Mucina & al. (2016) related alliance

Poion alpinae to Poo alpinae-Trisetetalia Ellmauer &

Mucina 1993 order.

Diagnostic species: Achillea sudetica, Aconi-

tum variegatum, Agrostis alpina, A. capillaris, Anth-

oxanthum odoratum, Avenula versicolor, Campanu-

la scheuchzeri, C. serrata, Crex pallescens, Cerastium

cerastoides, C. fontanum, Crepis aurea, Deschampsia

caespitosa, Euphrasia minima, E. picta, Festuca nigres-

cens, F. rubra, F. violacea, Galium anisophyllum, Geum

montanum (=Oreogeum m.), Homogyne alpina, Ligus-

ticum mutellina, Nardus stricta, Pedicularis verticillata,

Phleum alpinum, P. rhaeticum, Plantago alpina, P. atra-

ta, Poa alpina, P. m e d i a , P. supina, Potentilla aurea, P.

ternata, Prunella vulgaris, Ranunculus montanus, Sa-

gina saginoides, Senecio subalpinus, Thymus pulcherri-

mus, Trifolium badium, T. kotulae, T. pratense var. ni-

vale, T. thalii, Veronica alpina, V. serpyllifolia.

References: Horvat (1960); Jurko (1974); de Fou-

cault (1984; 1989); Ellenberg (1988); Sanda & Pope-

scu (1991); Ellmauer & Mucina (1993); Oberdorfer

(1993); Dierschke (1997; 1999); Zuidhoff & al. (1995);

Bardat & al. (2001); Burkart & al. (2004); Sarbu & al.

(2004); Meshinev & al. (2005); Hegedüšová (2007);

Jarolímek & al. (2008a); Škodová & Janišová (2008);

Biondi & Blasi (2013); Mucina & al. (2016).

Associations related to Poion alpinae:

Ass. Trifolio thalii-Festucetum violaceae Br.-Bl. in

Br.-Bl. & Jenny 1926

Distribution: France, Germany.

Ass. Alchemilletum pastoralis Szafer & al. 1927

Synonyms: Alchemilletum Szafer & al. 1923 (Art.

2b); Prunello vulgaris-Poetum alpinae Oberdorfer

1950; Agrostio vulgari-Poetum alpinae Bĕlohlávková

1980 nom. ined. (Art. 1).

Pseudonyms: Anthoxantho-Agrostietum sensu Un-

ar & al. 1984 non Silinger 1933.

Diagnostic species: Aconitum variegatum, Euphra-

sia minima agg., Galium pumillum agg., Ligusticum

mutellina, Oreogeum montanum, Phleum rhaeticum,

Poa alpina, Senecio subalpinus, Taraxacum Sect. alpi-

na, Tephroseris capitata, Thymus pulcherrimus.

Distribution: Slovakia.

Ass. Crepido aureae-Festucetum commutatae Lüdi

1948

Synonyms: Trifolietum repennis Rübel 1911; Poetum

alpinae Rübel 1911 (Art. 36); 1912; Crepido-Festucetum

rubrae Lüdi 1948; Prunello-Poetum alpinae Oberdorfer

Annex 1. Continuation.

134 Velev, N. • Checklist of Arrhenatheretalia order

1950; Poo-Prunelletum Oberdorfer 1950; Poo alpinae-

Prunelletum vulgaris Oberdorfer 1957; Poa alpina-Al-

chemilla palmata Beldie 1967; Crepido aureae-Cyno-

suretum Dietl 1972; Alchemillo-Poëtum alpinae (Beldie

1967) Resmeriţă 1972; Trifolio badii-Poetum Dietl 1982

(Art. 2b). Non: Poetum alpinae Braun 1913 (Art. 36);

Poetum alpinae sensu Smettan 1981 (Art. 36).

Diagnostic species: Alchemilla palmata, Crepis au-

rea, Festuca nigrescens, Taraxacum alpinum ssp. nigri-

cans, Trifolium badium.

Distribution: Austria, France, Germany, Romania,

Slovenia.

Ass. Crepido-Cynosuretum Knapp ex Dietl 1972

Synonyms: Crepido-Cynosuretum Knapp 1962

(Art. 2b).

Pseudonyms: Poo-Prunelletum sensu Smettan

1981.

Distribution: Austria.

Ass. Crepido aureae-Poetum alpinae Poldini & Ori-

olo 1994

Pseudonyms: Poo-Prunelletum Poldini non Oberd.

1950; Crepido-Festucetum rubrae Lausi & al. 1981 non

Ludi 1948.

Distribution: Italy.

Plant communities without rank related to Poion

alpinae:

Plant community Deschampsia caespitosa-Veratrum

album/lobelianum sensu Poldini & Oriolo 1994

Distribution: Italy.

References: Beldie (1967); Sanda & Popescu (1991);

Ellmauer & Mucina (1993); Ellmauer (1995); Oberdorfer

(1993); Poldini & Oriolo (1994); Dierschke (1997; 1999);

Gèhu (1999); Sarbu & al. (2004); Hegedüšová (2007);

Jarolímek & al. (2008b); Škodová & Janišová (2008); Flo-

raWeb (accessed 26.03.2009); Šilc & Čarni (2012).

Alliance PANČIĆION SERBICAE Lakušić 1966

Synonyms: Pancicion serbicae Lakušić 1964 (Art. 1).

Diagnostic species: Anemone narcissiflora, Astran-

tia elatior, Crepis aurea var. bosniaca, Leucanthemum

vulgare s.l., Pancicia serbica, Phleum alpinum, Pimpi-

nella serbica, Rhinanthus rumelicus.

Remarks: Mucina & al. (2016) related alliance Pan-

cicion serbicae to Poo alpinae-Trisetetalia Ellmauer &

Mucina 1993 order.

References: Lakušić & al. (2005); Šilc & al. (2014);

Mucina & al. (2016).

Associations related to Pančićion serbicae:

Ass. Ranunculo-Pančićietum serbicae Lakušić 1966

Distribution: Montenegro.

Ass. Trifolio-Polygaletum azureae Lakušić 1966

Distribution: Montenegro.

Ass. Knautio-Cynosuretum cristati Blečić & Tatić

1967

Diagnostic species: Carum carvi, Cynosurus crista-

tus, Knautia montenegrina, Ophioglossum vulgatum,

Poa trivialis.

Remarks: Blečić & Tatić (1967) refer this associa-

tion to Arrhenarherion.

Distribution: Montenegro, Serbia.

Variations: Montenegro: subass. genistetosum sag-

ittalis Blečić & Tatić 1967; subass. potentilletosum tor-

mentillae Blečić & Tatić 1967.

Ass. Pancicio-Lilietum bosniacae Bjelčić & Lakušić

1969

Distribution: Bosnia and Herzegovina.

Ass. Festuco-Pančićietum serbicae K. Tomić 1970

Distribution: Serbia.

Ass. Hypochoeri-Crepidetum conizifoliae Redžić

1990

Distribution: Bosnia and Herzegovina.

Ass. Pančićio-Centaureetum nervosae Lakušić nom.

prov.

Distribution: Serbia.

References: Blečić & Tatić (1967); Blečić & Lakušić

(1976); Kojić & al. (1998); Stešević & Petrović (2004);

Lakušić & al. (2005); Redžić (2007); Redžić & al.

(2013).

Alliance BRACHYPODIO RUPESTRIS-

CENTAUREION NEMORALIS Br.-Bl. 1967

Synonyms: Lino biennis-Gaudinion fragilis (Br.-Bl.

1967) de Foucault 1989 (Art. 29); Gaudinio fragilis-

Cynosurion cristati (Rivas Goday & Rivas-Mart. 1963)

Géhu 2006 (Art. 5).

Remarks: See the synonymy and remark №2 of Ar-

rhenatherion elatioris Luquet 1926.

References: Fernez & Causse (2015), Mucina & al.

(2016).

Annex 1. Continuation.

135Phytol. Balcan. 24(1) • Sofia • 2018

Alliance ALCHEMILLO-RANUNCULION REPENTIS

Passarge 1979

Synonyms: Potentillion anserinae Tüxen 1947;

Plantagini-Prunellion Eliáš 1980.

Remarks: 1. Chytrý & Blažková (2007) pointed

Plantagini-Prunellion Eliáš 1980 as a synonym of Cy-

nosurion cristati Tüxen 1947; 2. Biondi & al (2014)

pointed Alchemillo-Ranunculion repentis Passarge

1979 as syntax. syn. of Potentillion anserinae Tüxen

1947 within Potentillo anserinae-Polygonetalia avicu-

laris Tüxen 1947 order; 3. Mucina & al. (2016) sub-

ordinated Alchemillo-Ranunculion repentis Passarge

1979 to Arrhenatheretalia elatioris order.

References: Biondi & Blasi (2013); Biondi & al.

(2014); Mucina & al. (2016).

Alliance FESTUCION PRATENSIS Sipaylova & al.

1985

Diagnostic species: Agrostis gigantea, Carduus

crispus, Carum carvi, Daucus carota, Elymus repns,

Eryngium planum, Festuca pratensis, F. rub ra, Gera-

nium pratense, Heracleum sibiricum, Hypericum per-

foratum, Lathyrus pratensis, Oenothera rubricaulis,

Phleum prаtense, Poa pratensis, Rumex confertus, R.

crispus, R. pseudonatronatus, R. thyrsiflorus, Selinum

carvifolia, Silene nutans, Taraxacum officinale, Thalic-

trum lucidum, Trifoilum hybridum, T. prаtense, Vicia

cracca.

Remarks: This alliance is pointed in Kuzemko

(2016) and Mucina & al. (2016) as synonym of Ar-

rhenatherion elatioris Luquet 1926.

References: Stepanovich (2000); Goncharen-

ko (2001); Bobrovsky (2004); Veen & al. (2007);

Mikhailovna (2008); Solomakha (2008); Kuzemko

(2009b; 2016).

Suballiances related to Festucion:

Suball. Festucenion pratensis Mirkin & Naumova 1986

References: Mikhailovna (2008).

Associations related to Festucion pratensis:

Ass. Festucetum pratensis Soó 1938

Remarks: For details see in Arrhenatherion elatioris.

Ass. Poёtum pratensis Ravarut, Cazac & Turenschi

1956

Distribution: Ukraine.

Ass. Agrostietum albae Michalko & Petráňová 1967

Diagnostic species: Agrostis gigantea.

Distribution: Ukraine.

Variations: subass. typicum with three variants –

var. typicum, var. Alopecurus pratensis, var. Carex ni-

gra; subass. hieracietosum umbellati Kuzemko 2009

with two variants – var. Calamgrostis epigeios, var. Poa

palustris.

Ass. Deschampsio-Festucetum rubrae Mirkin & Sa-

pegin 1985

Diagnostic species: Deschampsia caespitosa, Festu-

ca rubra, Poa pratensis.

Distribution: Ukraine.

Variations: subass. typicum; subass. filipendule-

tosum ulmariae Kuzemko 2009 with three variants –

var. typicum, var. Anthoxanthum odoratum, var. Plan-

tago major.

Ass. Festuco pratensis-Deschampsietum caespitosae

Shelyag & al. in Shelyag & al. 1985

Diagnostic species: Deschampsia caespitosa, Festu-

ca pratensis, Trifolium repens.

Distribution: Ukraine.

Variations: subass. anthoxanthetosum odorati

Kuzemko 2009; subass. carisetosum nigrae Kuzemko

2009 with two variants – var. Agrostis canina; var. Poa

pratensis.

Ass. Bromopsi inermis–Alopecuretum pratensis Mir-

kin in Denisova & al. 1986

Diagnostic species: Alopecurus pratensis, Bromop-

sis inermis (= Bromus inermis).

Distribution: Russia (Komi Republic).

Ass. Festuco pratensis-Deschampsietum caespitosae

Turubanova 1986

Distribution: Russia (Komi Republic), Ukraine.

Ass. Vicio craccae-Agrostietum giganteae Mirkin in

Denisova & al. 1986

Distribution: Russia.

Ass. Agrostio gigantea-Festucetum pratensis Sipay-

lova & al. 1987

Diagnostic species: Agrostis gigantea, Festuca prat-

ensis, F. rubra, Poa pratensis.

Distribution: Ukraine.

Variations: subass. typicum; subass. deschampsieto-

sum caespitosae Kuzemko 2009 with three variants –

var. typicum, var. Carex pallescens, var. Carex praecox;

Annex 1. Continuation.

136 Velev, N. • Checklist of Arrhenatheretalia order

subass. festucetosum orientalis Kuzemko 2009; subass.

juncetosum compressi Kuzemko 2009.

Ass. Festuco pratensis-Dactylidetum glomeratae

Dymina 1989

Distribution: Russia.

Ass. Elytrigio repentis-Vicietum craccae Smetana &

al. 1997

Diagnostic species: Vicia cracca.

Distribution: Ukraine.

Ass. Vicietum craccae Smetana & al. 1997

Diagnostic species: Vicia cracca.

Distribution: Ukraine.

Ass. Dactylidetum glomeratae Stepanovič (1987) 1999

Diagnostic species: Dactylis glomerata.

Distribution: Belarus.

Variations: subass. brizetosum mediae sensu

Stepanovich 2000; subass. bromopsidetosum inermis

sensu Stepanovich 2000; subass. elytrigietosum re-

pentis sensu Stepanovih 2000; subass. festucetosum

pratensis sensu Stepanovich 2000; subass. phleetosum

pratensi sensu Stepanovich 2000; subass. poetosum

pratensis sensu Stepanovich 2000.

Ass. Phleetum pratensi Stepanovič 1999

Diagnostic species: Phleum pratense.

Distribution: Belarus.

Variations: subass. deschampsietosum cespitosae

sensu Stepanovih 2000; subass. festucetosum pratensis

sensu Stepanovich 2000; subass. festucetosum rubrae

sensu Stepanovich 2000; subass. poetosum pratensis

sensu Stepanovich 2000; subass. trifolietosum praten-

si sensu Stepanovich 2000.

Ass. Poetum pratensis Stepanovič 1999

Diagnostic species: Poa pratensis.

Distribution: Belarus, Ukraine (also in Cynosurion).

Variations: Belarus: subass. agrostidetosum ten-

uis sensu Stepanovich 2000; subass. deschampsietosum

cespitosae sensu Stepanovich 2000; subass. festucetosum

pratensis sensu Stepanovich 2000; subass. festucetosum

rubrae sensu Stepanovich 2000; subass. poetosum trivi-

alis sensu Stepanovich 2000; subass. phleetosum praten-

si sensu Stepanovich 2000; subass. trifolietosum repentis

sensu Stepanovich 2000; Ukraine: subass. typicum; sub-

ass. festucetosum rubrae Kuzemko 2009 with three var-

iants – var. typicum, var. Anthoxanthum odoratum, var.

Carex nigra; var. typicum; var. Elytrigia repens; var. Fes-

tuca rubra; var. Trifolium repens.

Ass. Trifolietum pratensi Stepanovič 1999

Diagnostic species: Trifolium pratense.

Distribution: Belarus.

Variations: subass. dactylidetosum glomeratae sen-

su Stepanovih 2000; subass. festucetosum pratensis

sensu Stepanovih 2000; subass. festucetosum rubrae

sensu Stepanovih 2000; subass. lotetosum cornicula-

ti sensu Stepanovih 2000; subass. phleetosum praten-

si sensu Stepanovih 2000; subass. poetosum pratensis

sensu Stepanovih 2000; subass. trifolietosum repentis

sensu Stepanovih 2000.

Ass. Carici vulpinae-Juncetum effusi Goncharenko

2002

Pseudonyms: Scirpetum sylvatici sensu auct. p. p.

Diagnostic species: Caltha palustris, Carex vulpina,

Crepis paludosa, Geum rivale, Juncus effusus, Scirpus

sylvaticus.

Distribution: Ukraine.

Ass. Galio veri-Agrostietum tenuis Goncharenko 2002

Pseudonyms: Anthoxantho-Agrostietum sensu

auct. p. p.

Diagnostic species: Agrostis capillaris, Briza media,

Galium verum, Luzula campestris, Rumex thyrsiflorus.

Distribution: Ukraine.

Ass. Medicago lupulinae-Phleetum pratensis Gon-

charenko 2002

Diagnostic species: Festuca orientalis, Medicago lupuli-

na, Phleum pratense, Rhinanthus vernalis, Vicia cracca.

Distribution: Ukraine.

Ass. Tussilago farfarae-Phleetum pratensis Vetluzh-

skikh 2006

Diagnostic species: Artemisia vulgaris, Dactylis

glomerata, Festuca pratensis, Phleum pratense, Picris

hieracioides, Salix caprea, S. viminalis, Tussilago far-

fara, Urtica dioica.

Variations: var. Arctium tomentosum; var. Crep-

is tectorum; var. Phalaroides arundinacea; var. Verbas-

cum thapsus; var. Pinus sylvestris.

Distribution: Russia.

Ass. Elytrigio repentis–Bromopsidetum inermis Yama-

lov 2011

Distribution: Russia (Komi Republic).

Ass. Festucetum pratensis-regelianae Kuzemko 2012

Diagnostic species: Achillea inundata, Carex dis-

tans, Festuca regeliana.

Distribution: Ukraine.

References: Stepanovich (2000); Goncharenko

(2001); Vetluzhskikh (2006); Chinkina (2006); Astak-

hova (2007); Mikhailovna (2008); Solomakha (2008);

Annex 1. Continuation.

137Phytol. Balcan. 24(1) • Sofia • 2018

Kuzemko (2009b, 2012); Kozyr (2013); Shushpannik-

ova & Yamalov (2014).

Alliance PHYTEUMATO-TRISETION Ellmauer &

Mucina 1993

Synonyms: Phyteumato-Trisetion flavescentis Hun-

dt ex Passarge 1969 (Art. 3d).

Remarks: See the synonymy and remaks of alliance

Polygono bistortae-Trisetion flavescentis Br.-Bl. & Tüx-

en ex Marschall 1947.

References: Mucina & al. (2016).

Alliance RUMICION THYRSIFLORI Micevski 1994

Diagnostic species: Achillea collina, Inula britanica,

Cinosurus cristatus, Euphrasia stricta, Geranium bru-

tium, Lolium perenne, Lotus corniculatus, Matricaria

trichophylla, Plantago lanceolata, Potentilla recta var.

pilosa, Ranunculus polyanthemos, Rumex thyrsiflorus,

Trifolium micranthum.

Remarks: This alliance has been described from

the Republic of Macedonia and classified within Tr i-

folio-Hordeetalia Horvatić 1963 order (Micevski 1994;

Melovski & Matevski 2008). Later Mucina & al. (2016)

include Rumicion thyrsiflori within Arrhenatheretalia

elatioris order.

References: Micevski (1994); Melovski & Matevski

(2008); Matevski (2010); Mucina & al. (2016).

Associations related to Rumicion thyrsiflori:

Ass. Triseto-Rumicetum thyrsiflori Micev. 1994 (Tri-

setum flavescens)

Diagnostic species: Matricaria trichophylla, Ra-

nunculus polyanthemos, Rhinanthus major, Tri setum

flavescens.

Distribution: Republic of Macedonia.

References: Micevski (1994); Matevski (2010).

Alliance POTENTILION-RANUNCULION REPENTIS

sensu Veen & al. 2007 nom. prov.

Diagnostic species: Juncus articulatus, Mentha

arvensis, Myosotis scorpioides, Potentilla anserina,

Prunella vulgaris, Ranunculus flammula, R. repens,

Sium latifolium, Veronica scutellata.

Remarks: The name of the alliance is not in accord-

ance with the rules of ICPN (Weber & al. 2000). The

alliance is mentioned for Belarus. According to Veen

& al. (2007) this syntaxon needs more investigations.

References: Veen & al. (2007).

Alliance TRIFOLIO (PRATENSE)-BRIZION

ELATIORIS Didukh & Kuzemko 2009

Diagnostic species: Briza elatior, Dianthus capita-

tus, Dorycnium herbaceum, Eryngium campestre, Inu-

la aspera, Luzula multiflora, Polygala major, Potentilla

depressa, Poterium polygonum, Primula acaulis, Pyre-

thrum corymbosum, Scabiosa columbaria, Trifolium

pratense, Vicia tenuifolia.

References: Didukh & Kuzemko (2009); Kuzemko

(2009b); Mucina & al. (2016).

Associations related to Trifolio-Brizion:

Ass. Trifolio (pratense)-Brizetum elatioris Didukh &

Kuzemko 2009

Diagnostic species: Briza elatior, Dianthus capita-

tus, Dorycnium herbaceum, Eryngium campestre, Inu-

la aspera, Luzula multiflora, Polygala major, Potentilla

depressa, Poterium polygonum, Primula acaulis, Pyre-

thrum corymbosum, Scabiosa columbaria, Trifolium

pratense, Vicia tenuifolia.

Distribution: Ukraine.

Variations: var. typicum; var. Fragaria vesca; var.

Salvia verticilata.

References: Didukh & Kuzemko (2009); Kuzemko

(2009b).

Alliance HELICTOTRICHO (COMPRESSI)-BISTORTION

OFFICINALIS Didukh & Kuzemko 2009

Diagnostic species: Ajuga genevensis, Alopecurus

vaginatus, Brachypodium pinnatum, Bromopsis cappa-

docica, B. riparia, Carex pallescens, Centaurea fusco-

marginata, Cerastium biebersteinii, Cruciata taurica,

Gentiana cruciata, Helictotrichon compressum, Hy-

pericum tauricum, Linum nervosum, Luzula campes-

tris, Pedicularis sibthorpii, Polygonum bistorta, Trifoli-

um alpestre, T. medium, Veronica gentianoides.

References: Didukh & Kuzemko (2009); Kuzemko

(2009b).

Annex 1. Continuation.

138 Velev, N. • Checklist of Arrhenatheretalia order

Associations related to Helictotricho-Bistortion:

Ass. Helictotricho (compressi)-Bistortetum officinalis

Didukh & Kuzemko 2009

Diagnostic species: Ajuga genevensis, Alopecurus

vaginatus, Brachypodium pinnatum, Bromopsis cappa-

docica, B. riparia, Carex pallescens, Centaurea fusco-

marginata, Cerastium biebersteinii, Cruciata taurica,

Gentiana cruciata, Helictotrichon compressum, Hy-

pericum tauricum, Linum nervosum, Luzula campes-

tris, Pedicularis sibthorpii, Polygonum bistorta, Trifoli-

um alpestre, T. medium, Veronica gentianoides.

Distribution: Ukraine.

Variations: var. Alchemilla tittantha; var. Myosotis

lithospermifolia.

References: Didukh & Kuzemko (2009); Kuzemko

(2009b).

Alliance SALVIO PRATENSIS-DACTYLIDION

GLOMERATAE Ubaldi & al. in Ubaldi 2003

Synonyms: Ranunculo neapolitani-Arrhenatherion

elatioris Allegrezza & Biondi 2011 (Art. 5).

Diagnostic species: Achillea collina, Arrhenather-

um elatius, Centaurea nigrescens ssp. neapolitana,

Dasypyrum villosum, Heracleum sphondylium ssp. ter-

natum, Pastinaca sativa ssp. urens, Ranunculus nea-

politanus, R. velutinus, Rhinanthus personatus.

References: Allegrezza & Biondi (2011); Biondi &

Blasi (2013); Biondi & al. (2014); Mucina & al. (2016).

Associations related to Salvio pratensis-Dactylidion

glomeratae:

Ass. Pastinaco urentis-Arrhenatheretum elatioris

Biondi & Allegrezza 1996

Diagnostic species: Inula viscosa, Tussilago farfara,

Pulicaria dysenterica, Hedysarum coronarium, Blacksto-

nia perfoliata, Senecio erucifolius, Festuca arundinacea.

Distribution: Italy.

Ass. Festuco circummediterraneae-Arrhenatheretum

elatioris Allegrezza 2003

Diagnostic species: Galium verum ssp. verum, On-

obrychis viciifolia, Festuca rubra agg., Galium mollugo

ssp. erectum, Bunium bulbocastanum, Centaurea sca-

biosa, Festuca circummediterranea, F. rupicola, Trifo-

lium incarnatum ssp. molinerii, Dasypyrum villosum.

Distribution: Italy.

Variations: subass. festucetosum circummediterra-

neae Allegrezza & Biondi 2011; subass. trisetetosum

flavescentis Allegrezza & Biondi 2011; subass. hyperi-

cetosum perforati Allegrezza & Biondi 2011.

Ass. Ranunculo neapolitani-Arrhenatheretum elati-

oris Allegrezza & Biondi 2011

Diagnostic species: Tragopogon pratensis, Ta rax -

acum officinale, Holcus lanatus, Salvia pratensis, Cy-

nosurus cristatus, Geranium dissectum, Trifolium

campestre, Galium mollugo ssp. mollugo, Herachleum

spondylium ssp. ternatum, Salvia verticillata, Tris et um

flavescens, Colchicum lusitanum.

Distribution: Italy.

Variations: subass. ranunculetosum neapolitani Al-

legrezza & Biondi 2011 (Syn.: Arrhenatheretum sensu

Pedrotti 1963; plant community Arrhenatherum ela-

tius sensu Venanzoni 1992); subass. ranunculetosum

velutini Allegrezza & Biondi 2011; subass. agropyreto-

sum repentis Allegrezza & Biondi 2011.

References: Venanzoni (1992); Allegrezza & Bion-

di (2011).

Alliance ANTHRISCO SYLVESTRIS-

ARRHENATHERION ELATIORIS Rodríguez-Rojo &

al. 2017

Diagnostic species: Anthriscus sylvestris, Cirsium

arvense, Convolvulus arvensis, Elytrigia repens, Gali-

um aparine, Potentilla reptans, Urtica dioica.

Remarks: Recently described West and Central Eu-

ropean alliance within Molinio-Arrhenatheretea class,

holotypus – Tanaceto-Arrhenatheretum Fischer ex Ell-

mauer in Mucina & al. 1993. Order relation not ex-

plicitly specified, but presumably belonging to Ar-

rhenatheretalia?

Ass. Tanaceto vulgaris-Arrhenatheretum elatioris

Fischer ex Ellmauer in Mucina & al. 1993

Remarks: for details see in alliance Arrhenatherion.

Distribution: West and Central Europe.

Ass. Vicio sepium-Arrhenatheretum O’Sullivan 1982

Remarks: for details see in alliance Arrhenatherion.

Distribution: West and Central Europe.

Plant community Urtica dioica-Arrhenatherum

sensu Rodríguez-Rojo & al. 2017

Synonyms: Arrhenatheretum-Urtica dioica sub-

community Rodwell & al. 1992.

Distribution: West and Central Europe.

Annex 1. Continuation.

139Phytol. Balcan. 24(1) • Sofia • 2018

Plant community Festuca arundinacea-Arrhena the-

rum sensu Rodríguez-Rojo & al. 2017

Synonyms: Arrhenatheretum festucetosum arundi-

naceae Zuidhoff & al. 1996.

Distribution: West and Central Europe.

References: Rodríguez-Rojo & al. (2017).

Associations subordinated to Arrhenatheretalia

elatioris, but information of alliance relation not

found

Remarks: The following association are pointed to

belong to alliances Cynosurion R. Tx. 1947 and Festu-

cion pratensis Sipaylova & al. 1985.

Ass. Festucetum ovinae Turubanova 1986

Distribution: Russia (Komi Republic).

Ass. Festuceto pratensis–Dactyletum glomeratae

Dymina in Korolyuk & Kiprijanova 1998

Distribution: Russia (Komi Republic).

Ass. Equiseto arvensis-Elytrigietum repentis Maracu-

lina 2009

Distribution: Russia (Komi Republic).

Ass. Amorio repens–Poetum pratensis sensu Shush-

pannikova & Yamalov 2014

Distribution: Russia (Komi Republic).

References: Shushpannikova & Yamalov (2014).

Plant communities without rank nor any alliance

relation, subordinated straight to Arrhenathereta-

lia elatioris

Plant community Potentilla erecta-Festuca rubra

sensu Becker & Becker 2010

Distribution: Germany.

Variations: var. Betonica officinalis; var. Trisetum

flavescens.

Plant community Agrostis vinealis sensu Shushpan-

nikova & Yamalov 2014

Distribution: Russia (Komi Republic).

Plant community Hypericum maculatum–Phleum

pratense sensu Shushpannikova & Yamalov 2014

Distribution: Russia (Komi Republic).

References: Becker & Becker (2010); Shushpannik-

ova & Yamalov (2014).

Annex 1. Continuation.

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Dec 2021
ECOL INDIC

Floristic diversity in agricultural landscapes results primarily from the diversity of semi-natural grasslands. Due to economical reasons, farmers are interested in agricultural intensification, while semi-natural grasslands are associated with low productivity. Therefore, 10 semi-natural grassland plant communities that are widespread across Central Europe and have different habitat requirements were studied: 4 from the Phragmitetea class, 4 from Molinio-Arrhenatheretea, and 2 from Festuco-Brometea. The study answers the following: (1) What is their floristic diversity and utilization value? (2) Is there a significant relationship between the diversity of vascular plant species and the utilization value of the examined communities? (3) If yes, what is the character of this relationship? The highest floristic diversity was shown by the Festuco-Brometea communities: Adonido-Brachypodietum pinnati and com. Brachypodium pinnatum. The average utilization value of the analyzed semi-natural plant communities was poor or very poor, but it varied within the plant communities, and the percentage of the relevés of at least a good utilization value was high for some communities. For most of the analyzed communities, the linear and statistically significant relation of H’ index with utilization value score (UVS) was observed. H’ index was positively related with UVS for Phragmitetum australis, Caricetum gracilis, Caricetum ripariae, and Deschampsietum caespitosae. H’ index was negatively related with UVS for Phalaridetum arundinaceae, Alopecuretum pratensis, Lolio-Cynosuretum, and Adonido-Brachypodietum pinnati. The study showed that the relationship between floristic diversity and utilization value depends on the type of plant community.

Caracterización geobotánica de algunos sintaxones de prados de siega mesófilos (all. Arrhenatherion elatioris) de Cataluña y de la Europa occidental

Article

Full-text available

Nov 2020

Gabriel Mercadal i Corominas

## Geobotanical characterization of some syntaxa from mesophilic hay meadows (all. Arrhenatherion) in Catalonia (NE Spain & SE France) and Western Europe. ## We present a summary of part of the phytogeographic results described in the author’s PhD thesis. In this case, we characterise and validate several syntaxa from mesophilic hay meadows in the Arrhenatherion elatioris alliance (ord. Arrhenatheretalia elatioris) in Catalonia (incl. NE Spain and SE France) and the Western Mediterranean. In total, we study 19 syntaxa in geobotanical terms, based on synthetic tables and factor analyses of correspondence published previously in the thesis: 1 order (Arrhenatheretalia), 1 alliance (Arrhenatherion), 2 suballiances (Eu-Arrhenatherenion, Avenulo-Arrhenatherenion), 6 associations and 9 subassociations (Trifolio-Arrhenatheretum nova subass. typicum, subass. dianthetosum nova; Tragopogono-Lolietum subass. typicum, subass. ophioglossetosum stat. nov.; Malvo-Arrhenatheretum; Galio-Arrhenatheretum subass. typicum, subass. anthoxanthetosum; Gentiano-Trisetetum; Rhinantho-Trisetetum subass. typicum, subass. centaureetosum nigrae nova, subass. centaureetosum scabiosae nova; Conopodio-Vicietum nova).

Flora Montiberica, vol. 78

Preprint

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Nov 2020

ÍNDICE Rubén RAMÍREZ-RODRÍGUEZ, Julián GARCÍA MUÑOZ & Francisco AMICH– Sobre la presencia del endemismo ibérico Delphinium bolosii en la provincia de Guadalajara, 3-6 | Descarga el artículo Gonzalo MATEO SANZ – La naturaleza en la toponimia española, VII, 7-16 | Descarga el artículo P. Pablo FERRER-GALLEGO, David BENAVENT, María FERRIOL, Alfonso GARMENDIA & Hugo MERLE – Centaurea ×masfitensis, nothosp. nov. (sect. Seridia (Juss.) DC., Asteraceae), 17-23 | Descarga el artículo Lluís SERRA LALIGA & Antoni BELDA – Papaver orientale L., nueva especie adventicia en España, 24-25 | Descarga el artículo P. Pablo FERRER-GALLEGO, Manuel PERERIRA & Emilio LAGUNA LUMBRERAS – Rhamnus alaternus var. chlorocarpus, var. nov. (Rhamnaceae), 26-28 | Descarga el artículo Agustí AGUT ESCRIG& Pello URRUTIA URIARTE – Observaciones nomenclaturales sobre Narcissus pallidulus Graells (Amaryllidaceae), 29-33 | Descarga el artículo Lluís SERRA LALIGA, Enric MARTÍ, Hilarión PEDAUYÉ& Rubén BOIX – Lafuenteo rotundifoliae-Sarcocapnetum enneaphyllae, nueva comunidad espeluncícola en la provincia murciano-almeriense, 34-40 | Descarga el artículo Arnau RIBAS SERRA, Marcello Dante CERRATO, Carles CARDONA AMETLLER, Pere Miquel MIR ROSSELLÓ & Lorenzo GIL VIVES – Aportaciones corológicas a la flora de Mallorca, 41-48 | Descarga el artículo Agustín LAHORA CANO & P. Pablo FERRER-GALLEGO – Revisión nomenclatural de Arctostaphylos uva-ursi (L.) Spreng. (Arbutoideae, Ericaceae), 49-60 | Descarga el artículo Gonzalo MATEO SANZ – Nuevas reflexiones sobre los nombres vernáculos de las plantas, 61-63 | Descarga el artículo José Luis MEDINA-GAVILÁN – First record of the exotic species Petunia axillaris (Solanaceae) from the Iberian Peninsula, 64-65| Descarga el artículo P. Pablo FERRER-GALLEGO, Roberto ROSELLÓ, Emilio LAGUNA, Diego RIVERA, Concepción OBÓN, José GÓMEZ & Juan B. PERIS – Sobre el tipo nomenclatural de Sideritis tragoriganum Lag. (Labiatae), 66-73 | Descarga el artículo Pedro GÓMEZ-MURILLO, Ángel SÁNCHEZ GARCÍA, Pablo CASTRO PRIGENT, José Félix ÁLVAREZ GONZÁLEZ & Irene ARELLANO-MARTÍN – Observaciones esporádicas de insectos polinizadores en el género Narcissus L. (Asparagales, Amaryllidaceae) en España, 74-76 | Descarga el artículo Jesús RIERA VICENT, Jaime GÜEMES HERAS & Josep Antoni ROSSELLÓ PICORNELL – Thymus × bolosii (Lamiaceae), a new wild hybrid from the eastern Iberian Peninsula, 77-80 | Descarga el artículo Gonzalo MATEO SANZ & Juan Ignacio PERIS FIGUEROLA – De flora valentina, XXI, 81-84 | Descarga el artículo Carlos AEDO PÉREZ – Una nueva combinación en Geranium L. (Geraniaceae), 85 | Descarga el artículo Juan ANTONIO RUIZ DE GOPEGUI Y VALERO & Laurentino GARCÍA CAYÓN – Híbridos naturales de la familia Orchidaceae en la Montaña Palentina (Castilla y León) y Picos de Europa (Cantabria), 86-89 | Descarga el artículo Alberto MARTÍNEZ SALDAÑA – Primera localidad de Epipogium aphyllum Sw. (Orchidaceae) en el parque nacional de Ordesa y Monte Perdido (Huesca), 90-91 | Descarga el artículo Romieg SOCA – Ophrys passionis et Ophrys marzuola (Orchidaceae) en Catalogne (Espagne), 92-103 | Descarga el artículo Gonzalo MATEO SANZ, Óscar GARCÍA CARDO & Juan Manuel MARTÍNEZ LABARGA – Nuevos datos sobre la flora de la provincia de Cuenca, XXXIII, 104-111 | Descarga el artículo Rogelio SÁNCHEZ-VILLEGAS, Manuel SÁNCHEZ-VILLEGAS, Begoña QUIRÓS DE LA PEÑA, Francisco Javier DE SANDE VELICIA, Luis Fernando ESTÉVEZ RODRÍGUEZ, José Luis ROBLES FERNÁNDEZ, Cecilia SÁNCHEZ BENZ, Lea SÁNCHEZ BENZ, Ramón ROMÁN HERNÁNDEZ, Antonino GONZÁLEZ-CANALEJO, Ricardo José MARTÍN LÓPEZ, Juan Carlos RICO JIMÉNEZ, Alberto ARRIBAS MARTÍN,Blanca MARTÍN-GARCÍA, Joaquín CASTELO SCREMMER, Gustavo MERCHÁN CRIADO, Belén HERNANDEZ DE LA TORRE BENZAL & Modesto LUCEÑO GARCÉS – Novedades corológicas para la flora vascular de la Sierra de Gredos (Sistema Central), II, 112-119 | Descarga el artículo Gabriel MERCADAL i COROMINAS – Caracterización geobotánica de algunos sintáxones de prados siega mesófilos (all. Arrhenatherion elatioris) de Cataluña y de la Europa occidental, 120-133 | Descarga el artículo

Актуализиране на класификацията на потенциалната растителност в дефилето на р. Искър между Плана и Лозенска планина

Conference Paper

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Jun 2020

Asocijacija Lolietum multiflorae Dietl et Lehmann 1975 u Hrvatskoj - Association Lolietum multiflorae Dietl et Lehmann 1975 in Croatia

Article

Full-text available

Jul 2008

Zvjezdana Stančić

SAŽETAK U radu je prvi put za Hrvatsku zabilježena asocijacija Lolietum multiflorae Dietl et Lehmann 1975. Utvrđena je na tri lokaliteta u sjeverozapadnom dijelu zemlje. Uglavnom se razvija na zasijanim livadama. Radi se o intenzivno antropogeno utjecanoj travnjačkoj vegetaciji koja se kosi četiri do šest puta godišnje i vrlo često gnoji. U flornom sastavu kao karakteristična i dominantna vrsta raste trava talijanski ljulj-Lolium multiflorum. Zajednica pripada svezi Arrhenatherion elatioris, redu Arrhenatheretalia i razredu Molinio-Arrhenatheretea. Vrlo je siromašna vrstama. Usporedba sastojina asocijacije Lolietum multiflorae iz Hrvatske, Švicarske i Austrije pokazuje vrlo sličan sastav vrsta. - ABSTRACT In the paper, the association Lolietum multiflorae Dietl et Lehmann 1975 is noted in Croatia for the first time. It has been established at three sites in the northwest part of the country. The community develops mostly on surfaces of sown meadows. This is intensively anthropogenically influenced grassland vegetation which is mown four to six times per year and often manured. In the flora composition, Lolium multiflorum grows as a characteristic and dominant species. Syntaxonomically, the association is assigned to the Arrhenatherion elatioris alliance, to the Arrhenatheretalia order and to the Molinio-Arrhenatheretea class. It is poor in species. Comparison of stands of Lolietum multiflorae from Croatia, Switzerland and Austria shows very similar species composition.

Phytosociological description of hay meadows with dominating Trisetum flavescens in the lower montane belt of north-western and western Slovenia

Article

Full-text available

Dec 2015

We conducted a phytosociological study into hay meadows on former fields on original sites of beech forests form the alliance Aremonio-Fagion in the lower montane belt of the northwestern and western Slovenia (southern Julian Alps, northern part of the Dinaric Alps) and compared them to similar, previously described meadows in Slovenia and northwestern Italy. Based on this comparison they are classified into the new association Rhinantho freynii-Trisetetum flavescentis and new habitat type, southeastern-Alpine-northern-Illyrian lower montane hay meadows – 38.239-S1.

Floristic diversity and sward use value of Lolio-Cynosuretum association in the San river valley

Article

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Jun 2008

In the present paper the floristic composition, species richness and sward use value of five subassociations distinguished within the Lolio-Cynosuretum association (typicum, trifolieto-sum repentis, festucetosum pratense, dactylidosum glomeratae and anthoxanthosum odoratae) are studied depending on the selected ecological factors. Phytosociological releves, samples of soil and samples of sward were taken for analysis. Floristic diversity and use value of the Lolio-Cynosuretum pasture sward depended on habitat conditions, fertilizing and use intensity.

PASTORAL VEGETAL ASSOCIATIONS IN THE LUCINA AND LUCAVA BASINS, SUCEAVA COUNTY (I)

Article

Full-text available

Dec 2006

As a result of the field studies undertaken on the pastures and hayfields of the Lucina Stud-Suceava County, whose surface partially includes the basin of the brooks Lucina and Lucava, various vegetal associations were identified, two of which are presented below; they are the object of a series analyses: chorological, cenotaxonomical, of the bio-forms, of the phytocoenological elements and of the ecological indices.

Syntaxonomy vegetation of Ukraine

Book

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Jun 2008

V.A. Solomakha

Syntaxonomical scheme of vegetation of Ukraine was presented and includes 965 associations, related to 70 classes according to Brown-Blanquet system. There were reflected groups of diagnostic species of syntaxons and their ecological features and spreading. It was presented a list of diagnostic species of syntaxons and index of their titles. Наведено третій варіант синтаксономії рослинності України за методом Браун-Бланке, яка включає 965 асоціацій, віднесених до 70 класів системи Браун-Бланке. Відображено блоки діагностичних видів синтаксонів, їхні екологічні особливості та поширення. Подано список діагностичних видів синтаксонів та індекс їх назв.

SOME PHYTOCOENOLOGICAL AND POPULATION STRUCTURE FEATURES OF FRITILLARIA MELEAGRIS L. IN THE UPPER ŞARD VALLEY (CLUJ COUNTY, ROMANIA)

Article

Full-text available

Mar 2003

The paper reveals the structure of a newly discovered Fritillaria meleagris population in a habitat from Cluj county, disturbed by drainage works started in the summer of 2003, after the acomplishment of the studies. The importance of the habitat is also highlighted by the presence of other red list species, like Achillea ptarmica, Dactylorhiza incarnata, Gentiana pneumonanthe, Gladiolus imbricatus, Iris sibirica, Narcissus poeticus ssp. radiiflorus, Trollius europaeus. Phytocoenological records were performed in three plant communities, developed along a moisture gradient, from wet to mesic: Caricetum acutiformis Eggler 1933, Cirsietum rivularis Nowinski 1926 and Junco-Molinietum Preising 1951. Population density, spatial and age structure of F. meleagris were investigated along three transects laid out in these communities. Morphometric data (stem height and perianth length) of individuals were also recorded and correlation between them was calculated.

Meadow vegetation of the north-east part of the Left-Bank Forest-Steppe of Ukraine

Article

Full-text available

Jan 2000

Igor V. Goncharenko

BIO-ECONOMIC ANALYSIS OF THE FLORA AND VEGETATION OF THE GRASSLANDS OF THE MEŞENDORF ECOLOGICAL FARM (BRAŞOV COUNTY, ROMANIA)

Article

Jan 1970

The paper presents the results of the botanical and phytosociological study on the natural grasslands of the Meşendorf mixed ecological farm (Transylvanian Natural Product). Floristic researches show a high floristic and phytosociological biodiversity, determined by pedo-climatic and microstational factors. The flora includes 329 species of cormophytes belonging to 111 genera of 37 botanical families. The vegetation of the grasslands comprises 12 plant associations, of which the phytocoenoses of the Festucetum pratense association are dominant. The quantitative studies performed evidence the dominance of good and very good fodder species.

Distribution of Grasses and Grasslands in Europe

Chapter

Jan 1979

R. Knapp

Meadow vegetation of the floodplains of the Vychegda and Pechora rivers. Order Arrhenatheretalia R. Tx. 1931

Article

Jan 2014

The present study continues a series of publications of the authors on diversity of the meadow vegetation in the floodplains of the Vychegda and Pechora rivers and their tributaries (Komi Republic). The floristic classification of wet meadows of the order Molinietalia Koch 1926 was published early (Shushpanikova, Yamalov, 2013). The results of floristic classification of meadow vegetation belonging to the order Arrhenatheretalia R. Tx. 1937 are discussed in this article. Analysis was based on 383 original relevés of the meadow vegetation of the order Arrhenatheretalia made in accordance with principles of the Braun-Blanquet approach (Braun-Blanquet, 1964; Mirkin and Naumova, 1998). The studies were performed in the floodplains of the Vychegda and Pechora rivers and their tributaries (Sysola, Kazhym, Pozheg, Viled, Usa, Colva, Synya, Sebys, Ilych, Syuzyu, Shchugor). The descriptions were conducted at the study plots of 10×10 m. Plant communities of small size were described within the natural limits. A location, dimensions of community, species composition, species saturation, composition of dominant and co-dominant species, total coverage and height of grass stand, aboveground phytomass production were identified. The abundance of species in meadow communities was recorded using the J. Braun-Blanquet method: r — species occurs sporadically, + — minor species, total coverage less than 1 %, 1 — 1–5 %, 2 — 6–25 % 3 — 26–50 %, 4 — 51–75 %, 5 — 76–100 %. The assessment of environmental parameters was performed by ecological methods of L. G. Ramensky (Ramensky et al., 1956). Syntaxonomic diversity of the meadow vegetation of the order Arrhenatheretalia of the Vychegda and Pechora rivers floodplains presented by 2 unions (Cynosurion R. Tx. 1947 и Festucion pratensis Sipaylova et al. 1985), 7 associations (Festucetum ovinae Turubanova 1986, Festuco pratensis–Deschampsietum caespitosae Turubanova 1986, Anthoxantho odorati–Agrostietum tenuis Sillinger 1933, Festuceto pratensis–Dactyletum glomeratae Dymina 1989 in Korolyuk et Kiprijanova 1998, Amorio repens–Poetum pratensis ass. nov., Equiseto arvensis–Elytrigietum repentis Maraculina 2009, Elytrigio repentis–Bromopsidetum inermis Yamalov 2011. Two new communities without rank are described (Agrostis vinealis, Hypericum maculatum–Phleum pratense). Most plant associations are heterogeneous by their floristic composition and they can be divided into 10 subassociations and 5 variants. The peculiarities of the floristic composition of the above mentioned associations, as well as their distribution and ecology are discussed. The distinguished associations are well differentiated by their elevation level on the floodplains. The meadow communities of Festucetum ovinae are located on the highest floodplains in the inland zone; there is Agrostis vinealis in the streamside zone of the floodplains. The meadow communities of the ass. Anthoxantho odorati–Agrostietum tenuis are located on the low floodplains in the streamside zone. The meadow communities of the associations Festuco pratensis–Deschampsietum caespitosae, Equiseto arvensis–Elytrigietum repentis, Elytrigio repentis–Bromopsidetum inermis, Festuceto pratensis–Dactyletum glomeratae are occupied the medium level floodplains. The meadow communities of the ass. Amorio repens–Poetum pratensis are found at different locations of the floodplain; they are formed under grazing. The described meadows grow on the soils with different ecological indicators. The distinguished associations are differentiated by water level (52.2–73.0), average figures — 55.9–68.9. The meadow communities of the ass. Festucetum ovinae occupy the driest sites of the floodplain; the ass. Anthoxantho odorati–Agrostietum tenuis is located in the wettest floodplains. The meadow communities of the ass. Elytrigio repentis–Bromopsidetum inermis occupy most rich and alluvial soils. They are located on the low ridges in the middle zone of the floodplain and they are formed by annual flooding on alluvial deposits. The distinguished associations are differentiated by pH (4.2–8.0), average figures — 4.9–7.0. The meadow communities with Agrostis vinealis are occupied the most acidic soil. The meadow communities of the ass. Anthoxantho odorati–Agrostietum tenuis are located on the neutral soils. The meadow communities of the most of associations occupy slightly acidic soils. The variability of moisture varies from alternating values to high values. The greatest differences in moisture are demonstrated by the communities of associations Equiseto arvensis–Elytrigietum repentis and Elytrigio repentis–Bromopsidetum inermis located on middle manes on low and medium levels of the floodplains. The associations are differ by the scale of pasture digression in range 3.3–5.0, average figures — 3.8–4.9. The meadow communities with Agrostis vinealis have the highest rates of pasture digression (4.5–5.1). The meadow communities of the ass. Anthoxantho odorati–Agrostietum tenuis have the smallest ones (3.5–3.9). A comparison of species composition of the syntaxa in Komi Republic with communities described in other regions of the Eastern Europe and the Asian part of Russia was carried out. It was shown that the meadow communities of the Komi Republic have low species diversity. Species richness of meadow communities is less than 39 species on trial area. The meadow communities of the ass. Festuco pratensis–Deschampsietum caespitosae are the richest. 195 vascular plant species are recorded in 213 geobotanical descriptions of the order Arrhenatheretalia. The aboveground phytomass varies from 20–40 g/m2 (Festucetum ovinae) to 520 g/m2 (Festuco pratensis–Deschampsietum caespitosae).

Arrhenatheretalia elatioris uncritical checklist of Europe

Abstract

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