A checklist of Chinese crickets (Orthoptera: Gryllidea)

Abstract

A checklist of Chinese crickets, including Taiwan, is offered. Presently 331 species or subspecies have been reported including true crickets, scale crickets, ant crickets and mole crickets belonging to 6 families, 16 subfamilies and 83 genera. Modicogryllus (Modicogryllus) maculatus (Shiraki, 1930) is moved to Comidoblemmus as C. maculatus (Shiraki, 1930) comb. nov. Velarifictorus (Velarifictorus) koshunensis (Shiraki, 1911) is moved to Turanogryllus as T. koshunensis (Shiraki, 1911) comb. nov. Qingryllus Chen & Zheng, 1995 syn. is the junior synonym of Goniogryllus Chopard, 1936. Loxoblemmus angulatus Bey-Bienko, 1956 syn. is the junior synonym of Loxoblemmus appendicularis Shiraki, 1930. Cophogryllus kuhlgatzi Karny, 1908 syn. is the junior synonym of Teleogryllus (Brachyteleogryllus) occipitalis occipitalis (Serville, 1838). Velarifictorus (Velarifictorus) aspersus borealis Gorochov, 1985 syn. is the junior synonym of Velarifictorus (Velarifictorus) aspersus aspersus (Walker, 1869). Modicogryllus (Modicogryllus) latefasciatus (Chopard, 1933) syn. is the junior synonym of Velarifictorus (Velarifictorus) micado (Saussure, 1877). Velarifictorus (Velarifictorus) ornatus caudatus (Shiraki, 1930) syn. is the junior synonym of Velarifictorus (Velarifictorus) ornatus ornatus (Shiraki, 1911). Dianemobius nigrofasciatus (Matsumura, 1904) syn. is the junior synonym of Dianemobius fascipes (Walker, 1869). Polionemobius mikado (Shiraki, 1911) syn. is the junior synonym of Polionemobius taprobanensis (Walker, 1869). Vietacheta picea Gorochov, 1992, Oecanthus euryelytra Ichikawa, 2001, Oecanthus similator Ichikawa, 2001, Xabea levissima Gorochov, 1992, Pteronemobius (Pteronemobius) yezoensis (Shiraki, 1911), Metioche (Metioche) japonica (Ichikawa, 2001), Natula matsuurai Sugimoto, 2001 are the first records from China.

Full text

Accepted by D. Rentz: 21 Nov. 2017; published: 8 Jan. 2018
ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN
1175-5334
(online edition)
Copyright © 2018 Magnolia Press
Zootaxa 4369 (4): 515
–
535
http://www.mapress.com/j/zt/
Article
515
https://doi.org/10.11646/zootaxa.4369.4.4
http://zoobank.org/urn:lsid:zoobank.org:pub:48EC8B65-EB2F-4E9F-AE50-6F46E7349F31
A checklist of Chinese crickets (Orthoptera: Gryllidea)
ZHU-QING HE
School of Life Sciences, East China Normal University, Shanghai 200241, China. E-mail: zqhe@bio.ecnu.edu.cn
Abstract
A checklist of Chinese crickets, including Taiwan, is offered. Presently 331 species or subspecies have been reported in-
cluding true crickets, scale crickets, ant crickets and mole crickets belonging to 6 families, 16 subfamilies and 83 genera.
Modicogryllus (Modicogryllus) maculatus (Shiraki, 1930) is moved to Comidoblemmus as C. maculatus (Shiraki, 1930)
comb. nov. Velarifictorus (Velarifictorus) koshunensis (Shiraki, 1911) is moved to Turanogryllus as T. koshunensis (Shi-
raki, 1911) comb. nov. Qingryllus Chen & Zheng, 1995 syn. is the junior synonym of Goniogryllus Chopard, 1936.
Loxoblemmus angulatus Bey-Bienko, 1956 syn. is the junior synonym of Loxoblemmus appendicularis Shiraki, 1930.
Cophogryllus kuhlgatzi Karny, 1908 syn. is the junior synonym of Teleogryllus (Brachyteleogryllus) occipitalis occipital-
is (Serville, 1838). Velarifictorus (Velarifictorus) aspersus borealis Gorochov, 1985 syn. is the junior synonym of Ve la r -
ifictorus (Velarifictorus) aspersus aspersus (Walker, 1869). Modicogryllus (Modicogryllus) latefasciatus (Chopard, 1933)
syn. is the junior synonym of Velarifictorus (Velarifictorus) micado (Saussure, 1877). Velarifictorus (Velarifictorus) or-
natus caudatus (Shiraki, 1930) syn. is the junior synonym of Velarifictorus (Velarifictorus) ornatus ornatus (Shiraki,
1911). Dianemobius nigrofasciatus (Matsumura, 1904) syn. is the junior synonym of Dianemobius fascipes (Walker,
1869). Polionemobius mikado (Shiraki, 1911) syn. is the junior synonym of Polionemobius taprobanensis (Walker, 1869).
Vietacheta picea Gorochov, 1992, Oecanthus euryelytra Ichikawa, 2001, Oecanthus similator Ichikawa, 2001, Xabea le-
vissima Gorochov, 1992, Pteronemobius (Pteronemobius) yezoensis (Shiraki, 1911), Metioche (Metioche) japonica
(Ichikawa, 2001), Natula matsuurai Sugimoto, 2001 are the first records from China.
Key words: China, Taiwan, cricket, taxonomy, checklist
Introduction
Chinese people have been interested in crickets since Tang Dynasty (A.D. 618-907). The women in palace reared
crickets for their songs (Yin & Liu 1995). After that, this custom spread to common people. Crickets have been
reared not only for songs, but also for fighting. Jia Si-Dao (A.D. 1213-1275) and Liu Tong (A.D. 1593-1636) wrote
monographs on cricket selection, rearing and fighting (Yin & Liu 1995).
Taxonomic studies of Chinese cricket species have been reported since the 19
th
century. Walker (1869) first
described new species: Oecanthus sinensis, Plebeiogryllus spurcatus and Velarifictorus (Velarifictorus) aspersus
from Hong Kong and Macau. Subsequently, Shiraki (1911; 1930), Chopard (1936a, b; 1939), Bey-Bienko (1956)
and Gorochov (1985) also described species from China.
Xu Yin-Qi (=Hsu Y.-C.) is the first Chinese modern researcher to focus on crickets. He described 16 genera
and 34 species in 1928, and 18 genera 47 species in 1931 (Hsu 1928;1931). Hu Jing-Fu (=Wu C.-F.) reported 22
genera, 56 species from China (Wu 1935). Then taxonomic research paused until 1980s. Wu Fu-Zhen (=Woo F.-
C.), Wang Yin, Zheng Yan-Fen from Institute of Plant Protection, Chinese Academy of Agricultural Sciences,
Beijing (Wang & Woo 1992; Wang et al. 1999; Wu & Wang 1992; Wu & Zheng 1992; Zheng & Woo 1992), Xia
Kai-Ling (=Hsia K.-L.), Liu Xian-Wei, Yin Hai-Sheng from Shanghai Entomological Museum, Chinese Academy
of Sciences, Shanghai (Liu et al. 1994; Liu et al. 1993a; Liu et al. 1993b; Xia et al. 1991; Yin 1998a, b; Yin & Liu
1996; Yin & Zhang 2001), Chen Jun, Xie Ling-De, Li Kai from Shaanxi Normal University, Xi'an (Chen 1994;
Chen & Zheng 1995a, b;1996; Li et al. 2010a, b; Li & Zheng 1998; Xie 2003, 2004, 2005, Xie & Ou 2005; Xie et
al. 2006; Xie & Zheng 2003a, b, c; Xie et al. 2004; Xie et al. 2003), Yang Jeng-Tze from National Chung Hsing

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Zootaxa 4369 (4) © 2018 Magnolia Press
University, Taichung (Yang 1995, 1998; Yang & Chang 1996; Yang & Yang 2012; Yang & Yang 1994, 1995; Yang
& Yen 2001a, b) reported many cricket new species. Especially, Yin Hai-Sheng and Liu Xian-Wei published
“Synopsis on the Classification of Grylloidea and Gryllotalpoidea from China” in 1995 as one of the notable
landmark. At present, Liu Hao-Yu, Ma Li-Bin and He Zhu-Qing are still working on taxonomy of crickets, and
they reported all new species since 2006 (Gorochov & He 2017; He 2012; He & Gorochov 2015; He et al. 2010;
He et al. 2009, He et al. 2017a, He et al. 2017b, He et al. 2016, Li et al. 2010a, b; Li et al. 2007; Liu et al. 2015a;
Liu et al. 2016a; Liu et al. 2014; Liu & Shi 2007, 2011a, b, c; 2012a, b, c; 2013; 2014a, b, c; 2015a, b, c; Liu et al.
2006; Liu et al. 2016b; 2017; Liu et al. 2015b; Ma & Gorochov 2015; Ma et al. 2015a; Ma et al. 2015b; Ma et al.
2015c; Ma et al. 2016; Ma et al. 2008; Ma et al. 2015d; Ma & Zhang 2010; 2011a, b; 2012; 2013; 2015a, b; Shi &
Liu 2007; Wang et al. 2017; Wu & Liu 2017; Zhang et al. 2017a, b; Zong et al. 2017). However, reviewed books or
articles on Chinese crickets have not been reported for over 20 years except for the Nemobiinae (Liu et al. 2016a).
Here, I summarize all these researches and list the species with their distributions from China. Distribution in Hong
Kong and Macao is substituted for Guangdong province. Distribution of Beijing and Tianjin is substituted for
Hebei province. Distribution of Shanghai is substituted for Zhejiang province. Reasons for this are that these areas
are too small to be considered as geographically separate. Literature is provided for descriptions, distributions in
China, and/or generic changes and keys to species. Species representing new records are deposited in East China
Normal University (ECNU).
Results
Taxa Distribution Literature
infraorder Gryllidea
superfamily Grylloidea
family Gryllidae
subfamily Eneopterinae
genus Cardiodactylus Saussure, 1878
C. guttulus (Matsumura, 1913) Hainan, Taiwan (Robillard et al. 2013)
genus Lebinthus Stål, 1877
L. lanyuensis Oshiro, 1996 Taiwan (Oshiro 1996a)
L. yaeyamensis Oshiro, 1996 Taiwan (Ichikawa et al. 2006; Oshiro 1996b)
genus Xenogryllus Bolívar, 1890
X. marmoratus marmoratus (Haan, 1842) Anhui, Fujian, Guangxi, Jiangsu,
Zhejiang
(Yin & Liu 1995)
X. marmoratus unipartitus (Karny, 1915) Taiwan (Ichikawa et al. 2006; Karny 1915)
X. transversus (Walker, 1869) Yunnan (Yin & Liu 1995)
subfamily Euscyrtinae
genus Beybienkoana Gorochov, 1988
B. ditrapeza Liu & Shi, 2012 Hainan (Liu & Shi 2012a)
B. formosana (Shiraki, 1930) Anhui, Fujian, Guangdong, Guangxi,
Hainan, Hunan, Taiwan, Yunnan,
Zhejiang
(Shiraki 1930; Yang & Yang 2012;
Yin & Liu 1995)
B. gregaria Yang & Yang, 2012 Taiwan (Yang & Yang 2012)
B. karnyi (Shiraki, 1930) Taiwan (Shiraki 1930; Yang & Yang 2012)
B. longipennis (Liu & Yin, 1993) Yunnan (Liu & Shi 2012a; Liu et al. 1993b)
B. luteola Yang & Yang, 2012 Taiwan (Yang & Yang 2012)
B. majora Liu & Shi, 2012 Yunnan (Liu & Shi 2012a)
B. parvula Shi & Liu, 2007 Guangxi (Shi & Liu 2007)
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A CHECKLIST OF CHINESE CRICKETS (ORTHOPTERA: GRYLL IDEA)
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Taxa Distribution Literature
B. splendida Yang & Yang, 2012 Taiwan (Yang & Yang 2012)
B. trapeza Liu & Shi, 2012 Hainan (Liu & Shi 2012a)
genus Euscyrtodes Gorochov, 1987
E. ogatai (Shiraki, 1930) Taiwan (Shiraki 1930)
genus Euscyrtus Guérin-Méneville, 1844
E. (Osus) concinnus (Haan, 1844) Guangdong, Hainan, Yunnan (Yin & Liu 1995)
E. (Osus) hemelytrus (Haan, 1844) Fujian,Guangxi, Guizhou, Hainan,
Hunan, Jiangsu, Jiangxi, Shandong,
Sichuan, Taiwan, Yunnan, Zhejiang
(Yang & Yang 2012; Yin & Liu
1995)
E. (Osus) japonicus Shiraki, 1930 Taiwan (Yang & Yang 2012)
genus Patiscus Stål, 1877
P. brevipennis Chopard, 1969 Guangxi, Yunnan (Yin & Liu 1995)
P. cephalotes (Saussure, 1877) Anhui, Hainan, Taiwan, Tibet, Yunnan (Yang & Yang 2012; Yin & Liu
1995)
P. malayanus Chopard, 1969 Guangxi, Hainan, Hunan,Taiwan (Yang & Yang 2012; Yin & Liu
1995)
subfamily Gryllinae
genus Acanthoplistus Saussure, 1877
A. birmanus Saussure, 1877 Guangdong, Hainan, Jiangsu, Taiwan,
Yunnan, Zhejiang
(Zheng & Woo 1992)
A. nigritibia Zheng & Woo, 1992 Zhejiang (Zheng & Woo 1992)
A. testaceus Zheng & Woo, 1992 Yunnan (Zheng & Woo 1992)
genus Agryllus Gorochov, 1994
A. magnigenitalis He & Gorochov, 2017 Yunnan (Gorochov & He 2017)
genus Callogryllus Sjöstedt, 1910
C. yunnanus Wu & Zheng, 1992 Yunnan (Wu & Zheng 1992)
genus Capillogryllus Xie, Zheng & Liang,
2003
C. dolabripalpis Xie & Zheng, 2003 Guangxi (Xie et al. 2003)
C. exilipalpis Xie & Zheng, 2003 Guangdong (Xie et al. 2003)
genus Comidoblemmus Storozhenko &
Paik, 2009
C. excavatus Liu & Shi, 2015 Guizhou (Liu & Shi 2015c)
C. maculatus (Shiraki, 1930)
1
Taiwan (Shiraki 1930)
C. nipponensis (Shiraki, 1911) Taiwan (Liu & Shi 2015c; Shiraki 1911)
C. sororius Liu & Shi, 2015 Zhejiang (Liu & Shi 2015c)
C. yigongensis Wu & Liu, 2017 Tibet (Wu & Liu 2017)
genus Conoblemmus Adelung, 1910
C. acutifrons Chopard, 1936 Xinjiang (Chopard 1936b)
C. saussurei Adelung, 1910 Xinjiang (Adelung 1910)
genus Eumodicogryllus Gorochov, 1986
E. bordigalensis bordigalensis (Latreille,
1804)
Inner Mongolia, Xinjiang (Yin & Liu 1995)
genus Goniogryllus Chopard, 1936
2
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(Continued)
Taxa Distribution Literature
G. asperopunctatus Wu & Wang, 1992 Guangxi, Hunan, Yunnan, Zhejiang (Wu & Wang 1992)
G. atripalpulus Chen & Zheng, 1996 Shaanxi (Chen & Zheng 1996)
G. bistriatus Wu & Wang, 1992 Yunnan (Wu & Wang 1992)
G. bomicus Wu & Wang, 1992 Tibet (Wu & Wang 1992)
G. cheni Xie & Zheng, 2003 Gansu (Xie & Zheng 2003c)
G. chongqingensis Chen & Zheng, 1995 Chongqing (Chen & Zheng 1995b)
G. chuannanensis Chen & Zheng, 1995 Sichuan (Chen & Zheng 1995a)
G. cirilinearis Xie, 2005 Gansu (Xie 2005)
G. emeicus Wu & Wang, 1992 Sichuan (Wu & Wang 1992)
G. gansuensis Xie, Yu & Tang, 2006 Gansu (Xie et al. 2006)
G. glaber Wu & Wang, 1992 Yunnan (Wu & Wang 1992)
G. jiguanshanensis (Liu, Zhang & Shi, 2017) Sichuan (Liu et al. 2017)
G. lushanensis Chen & Zheng, 1995 Jiangxi (Chen & Zheng 1995b)
G. octospinatus Chen & Zheng, 1995 Sichuan (Chen & Zheng 1995b)
G. ovalatus Chen & Zheng, 1996 Shaanxi (Chen & Zheng 1996)
G. potamini Bey-Bienko, 1956 Hubei, Sichuan (Bey-Bienko 1956; Yin & Liu 1995)
G. pubescens Wu & Wang, 1992 Sichuan (Wu & Wang 1992)
G. punctatus Chopard, 1936 Fujian, Hubei, Hunan, Guangxi,
Guizhou, Sichuan, Yunnan, Zhejiang
(Chopard 1936a; Yin & Liu 1995)
G. sexflorus Xie & Zheng, 2003 Shaanxi (Xie & Zheng 2003c)
G. striofemorus (Chen & Zheng, 1995) Shaanxi (Chen & Zheng 1995a)
G. yunnanensis Xie & Ou, 2005 Yunnan (Xie & Ou 2005)
genus Gryllodes Saussure, 1874
G. sigillatus (Walker, 1869) Anhui, Fujian, Guangdong, Guangxi,
Guizhou, Hebei, Heilongjiang, Hainan,
Hunan, Jiangsu, Jiangxi, Liaoning,
Shandong, Yunnan, Zhejiang
(Yin & Liu 1995)
genus Gryllus Linnaeus, 1758
G. (Gryllus) bimaculatus De Geer, 1773 Fujian, Guangdong, Guangxi, Hainan,
Inner Mongolia, Jiangxi, Sichuan,
Taiwan, Tibet, Yunnan, Zhejiang
(Yin & Liu 1995)
genus Gymnogryllus Saussure, 1877
G. contractus Liu, Yin & Liu, 1995 Yunnan (Liu et al. 1995)
G. dolichodens Ma & Zhang, 2011 Yunnan (Ma & Zhang 2011a)
G. extrarius Ma & Zhang, 2011 Yunnan (Ma & Zhang 2011a)
G. longus Ma & Zhang, 2011 Guangxi (Ma & Zhang 2011a)
G. odonopetalus Xie & Zheng, 2003 Yunnan (Xie & Zheng 2003a)
G. striatus Ma & Zhang, 2011 Yunnan (Ma & Zhang 2011a)
G. tumidulus Ma & Zhang, 2011 Guangdong (Ma & Zhang 2011a)
G. yunnanensis Ma & Zhang, 2011 Yunnan (Ma & Zhang 2011a)
genus Loxoblemmus Saussure, 1877
3
L. aomoriensis Shiraki, 1930 Anhui, Fujian, Guangxi, Hainan, Henan,
Hubei, Hunan, Shaanxi, Sichuan, Yunnan,
Zhejiang
(Shiraki 1930; Yin & Liu 1995)
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A CHECKLIST OF CHINESE CRICKETS (ORTHOPTERA: GRYLL IDEA)
(Continued)
Taxa Distribution Literature
L. appendicularis Shiraki, 1930
4
Guangxi, Hainan, Hunan, Jiangxi,
Sichuan, Taiwan, Yunnan
(Shiraki 1930; Yin & Liu 1995)
L. applanatus Wang, Zheng & Woo, 1999 Fujian (Wang et al. 1999)
L. arietulus Saussure, 1877 Guangxi, Hebei, Taiwan (Yin & Liu 1995)
L. detectus (Serville, 1838) Anhui, Fujian, Guangxi, Guizhou, Hebei,
Jiangsu, Jiangxi, Shaanxi, Sichuan,
Taiwan, Zhejiang
(Yin & Liu 1995)
L. doenitzi Stein, 1881 Anhui, Guangxi,Guizhou, Hebei, Henan,
Hunan, Jiangsu, Jiangxi, Liaoning,
Shaanxi, Shanxi, Shandong, Sichuan,
Zhejiang
(Yin & Liu 1995)
L. equestris Saussure, 1877 Anhui, Fujian, Guangxi, Hainan, Hebei,
Hubei, Hunan, Jiangsu, Jiangxi,
Liaoning, Sichuan, Taiwan, Tibet,
Yunnan, Zhejiang
(Shiraki 1930; Yin & Liu 1995)
L. formosanus Shiraki, 1930 Guangxi, Hainan, Taiwan, Yunnan,
Zhejiang
(Shiraki 1930; Yin & Liu 1995)
L. haani Saussure, 1877 Guangxi, Jiangsu, Taiwan, Yunnan,
Zhejiang,
(Yin & Liu 1995)
L. intermedius Chopard, 1929 Hainan, Hunan, Jiangxi,Yunnan (Yin & Liu 1995)
L. jacobsoni Chopard, 1927 Guangxi, Hainan, Hunan, Jiangxi,
Sichuan, Yunnan
(Yin & Liu 1995)
L. macrocephalus Chopard, 1967 Tibet (Yin & Liu 1995)
L. montanus Liu, Zhang & Shi, 2016 Sichuan (Liu et al. 2016b)
L. reticularus Liu, Yin & Liu, 1995 Yunnan (Liu et al. 1995)
L. taicoun Saussure, 1877 Guangxi, Hebei, Taiwan (Yin & Liu 1995)
genus Melanogryllus Chopard, 1961
M. bilineatus Yang & Yang, 1994 Taiwan (Yang & Yang 1994)
M. desertus (Pallas, 1771) Xinjiang (Yin & Liu 1995)
genus Mitius Gorochov, 1985
M. flavipes (Chopard, 1928) Yunnan (Yin & Liu 1995)
M. minor (Shiraki, 1911) Jiangsu, Zhejiang (Shiraki 1911; Yin & Liu 1995)
M. minutulus Yang & Yang, 1995 Taiwan (Yang & Yang 1995)
M. splendens (Shiraki, 1930) Taiwan (Shiraki 1930; Yin & Liu 1995)
genus Modicogryllus Chopard, 1961
M. (Modicogryllus) consobrinus (Saussure,
1877)
5
Guangxi, Taiwan, Yunnan, Zhejiang (Yin & Liu 1995)
genus Nigrogryllus Gorochov, 1983
N. sibiricus (Chopard, 1925) Heilongjiang, Jilin (Yin & Liu 1995)
genus Phonarellus Gorochov, 1983
P. (Phonarellus) flavipes Xia, Liu & Yin,
1991
Guangxi, Hainan, Yunnan (Xia et al. 1991)
P. (Phonarellus) minor (Chopard, 1959) Guangxi, Yunnan (Yin & Liu 1995)
P. (Phonarellus) ritsemae (Saussure, 1877) Guangxi, Zhejiang (Yin & Liu 1995)
genus Plebeiogryllus Randell, 1964
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(Continued)
Taxa Distribution Literature
P. guttiventris guttiventris (Walker, 1871) Fujian, Guangdong, Guangxi (Yin & Liu 1995)
P. guttiventris obscurus (Chopard, 1969) Fujian, Guizhou (Yin & Liu 1995)
P. plebejus (Saussure, 1877) Hainan, Taiwan (Yin & Liu 1995)
P. spurcatus (Walker, 1869) Guangdong (Walker 1869)
genus Svercacheta Gorochov, 1993
S. siamensis (Chopard, 1961)
6
Fujian, Guangdong, Guangxi, Guizhou,
Jiangxi, Yunnan
(Yin & Liu 1995)
genus Tarbinskiellus Gorochov, 1983
T. orientalis (Burmeister, 1838) Guangxi (Yin & Liu 1995)
T. portentosus (Lichtenstein, 1796) Fujian, Guangdong, Guangxi, Hainan,
Hebei, Jiangxi, Qinghai, Taiwan, Tibet,
Yunnan
(Yin & Liu 1995)
genus Teleogryllus Chopard, 1961
T. (Brachyteleogryllus) commodus (Walker,
1869)
Zhejiang (Ma et al. 2015b)
T. (Brachyteleogryllus) emma (Ohmachi &
Matsuura, 1951)
Gansu, Guizhou, Hebei, Henan, Hunan,
Jiangsu, Shaanxi, Shandong, Shanxi,
Sichuan, Zhejiang
(He et al. 2017c)
T. (Brachyteleogryllus) infernalis
(Saussure, 1877)
Gansu, Hebei, Heilongjiang, Inner
Mongolia, Jilin, Liaoning, Ningxia,
Qinghai, Shanxi
(Ma et al. 2015b)
T. (Brachyteleogryllus) occipitalis
occipitalis (Serville, 1838)
7
Fujian, Guangdong, Guangxi, Guizhou,
Hainan, Jiangxi, Sichuan, Tibet, Yunnan,
Zhejiang
(He et al. 2017c)
T. (Macroteleogryllus) derelictus
Gorochov, 1985
Guangdong, Hainan, Yunnan (Ma et al. 2015b)
T. (Macroteleogryllus) mitratus
(Burmeister, 1838)
Fujiang, Guangdong, Guangxi, Hainan,
Taiwan, Yunnan
T. (Teleogryllus) fallaciosus (Shiraki, 1930) Taiwan (Shiraki 1930; Yin & Liu 1995)
genus Turanogryllus Tarbinsky, 1940
T. eous Bey-Bienko, 1956 Shandong (Bey-Bienko 1956)
T. koshunensis (Shiraki, 1911)
8
Taiwan (Shiraki 1911)
T. lateralis (Fieber, 1854) Xinjiang (Yin & Liu 1995)
T. melasinotus Li & Zheng, 1998 Shaanxi (Li & Zheng 1998)
T. rufoniger (Chopard, 1925) Guangdong, Guangxi, Hainan, Yunnan (Yin & Liu 1995)
genus Velarifictorus Randell, 1964
V. (Velarifictorus) arisanicus (Shiraki,
1930)
Taiwan (Shiraki 1930; Yin & Liu 1995)
V. (Velarifictorus) aspersus (Walker, 1869)
9
Anhui, Fujian, Guangdong, Guangxi,
Guizhou, Hainan, Hebei, Jiangsu,
Jiangxi, Shaanxi, Shandong, Sichuan,
Yunnan, Zhejiang
(Gorochov 1985)
V. (Velarifictorus) beybienkoi Gorochov,
1985
Hainan, Shandong, Zhejiang (Gorochov 1985; Yin & Liu 1995)
V. (Velarifictorus) curvinervis Xie, 2004 Guangxi (Xie 2004)
V. (Velarifictorus) flavifrons Chopard, 1966 Guangdong, Yunnan (Yin & Liu 1995)
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Taxa Distribution Literature
V. (Velarifictorus) micado (Saussure,
1877)
10
Fujian, Guangdong, Guangxi, Guizhou,
Hebei, Hunan, Jiangsu, Jiangxi, Shaanxi,
Shandong, Shanxi, Sichuan, Tibet,
Yunnan, Zhejiang
(Yin & Liu 1995)
Taiwan (Shiraki 1930)
V. (Velarifictorus) ornatus (Shiraki, 1911)
11
Fujian, Guizhou, Henan, Hubei, Hunan,
Jiangsu, Jiangxi, Shandong, Sichuan,
Taiwan, Yunnan, Zhejiang
(Shiraki 1911; Yin & Liu 1995)
genus Vietacheta Gorochov, 1992
V. harpophylla Ma, Liu & Xu, 2015 Yunnan (Ma et al. 2015c)
V. picea Gorochov, 1992 Yunnan This study
12
subfamily Itarinae
genus Itara Walker, 1869
I. (Gryllitara) denudata Ma & Zhang, 2015 Yunnan (Ma & Zhang 2015b)
I. (Itara) aperta Gorochov, 1996 Yunnan (Ma & Zhang 2015b)
I. (Itara) basidentata Ma & Zhang, 2015 Yunnan (Ma & Zhang 2015b)
I. (Itara) communis Gorochov, 1997 Yunnan (Ma & Zhang 2015b)
I. (Itara) dicrana Ma & Zhang, 2015 Hainan (Ma & Zhang 2015b)
I. (Itara) minor Chopard, 1925 Yunnan (Ma & Zhang 2015b)
I. (Itara) vietnamensis Gorochov, 1985 Guangdong, Yunnan (Ma & Zhang 2015b)
I. (Noctitara) sonabilis Gorochov, 1996 Hainan (Ma & Zhang 2015b)
genus Parapentacentrus Shiraki, 1930
P. formosanus Shiraki, 1930 Fujian, Guangxi, Taiwan (Shiraki 1930; Yin & Liu 1995)
P. fuscus Gorochov, 1988 Guangdong (Gorochov 1988b)
subfamily Landrevinae
genus Duolandrevus Kirby, 1906
D. (Duolandrevus) bannanus Zhang, Liu &
Shi, 2017
Yunnan (Zhang et al. 2017a)
D. (Eulandrevus) axinus Zhang, Liu & Shi,
2017
Hainan (Zhang et al. 2017b)
D. (Eulandrevus) coriaceus (Shiraki, 1930) Taiwan (Shiraki 1930)
D. (Eulandrevus) dendrophilus (Gorochov,
1988)
Fujian, Guangdong, Guangxi,Guizhou (Zhang et al. 2017b)
D. (Eulandrevus) gorochovi Zhang, Liu &
Shi, 2017
Guangdong (Zhang et al. 2017b)
D. (Eulandrevus) guntheri (Gorochov,
1988)
Taiwan (Gorochov 1988a)
D. (Eulandrevus) hainanensis Liu, He &
Ma, 2015
Hainan (Liu et al. 2015b)
D. (Eulandrevus) infuscatus Liu & Bi, 2010 Zhejiang (Liu & Bi 2010)
D. (Eulandrevus) unguiculatus Ma,
Gorochov & Zhang, 2015
Yunnan (M a et al. 2015a)
D. (Eulandrevus) ziyunensis Zhang, Liu &
Shi, 2017
Fujian (Zhang et al. 2017b)
D. (Jorama) kotoshoensis Oshiro, 1989 Taiwan (Gorochov 1996; Hsiung 1993)
subfamily Oecanthinae
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genus Oecanthus Serville, 1831
O. antennalis Liu, Yin & Xia, 1994 Anhui (Liu et al. 1994)
O. euryelytra Ichikawa, 2001 Fujian This study
13
O. latipennis [temper] Liu, Yin & Xia,
1994
Hebei (Liu et al. 1994)
O. longicauda Matsumura, 1904 Fujian, Guangxi, Guizhou, Heilongjiang,
Hunan, Jiangxi, Jilin, Shaanxi, Shanxi,
Sichuan, Yunnan, Zhejiang
(Liu et al. 1994)
O. similator Ichikawa, 2001 Zhejiang This study
14
O. sinensis Walker, 1869 Guangdong? (Walker 1869)
O. turanicus Uvarov, 1912 Xinjiang (Liu et al. 1994)
O. zhengi Xie, 2003 Yunnan (Xie 2003)
genus Xabea Walker, 1869
X. levissima Gorochov, 1992 Yunnan This study
15
subfamily Pentacentrinae
genus Lissotrachelus Brunner von
Wattenwyl, 1893
L. ferrugineonotatus Brunner von
Wattenwyl, 1893
Tibet, Yunnan (Yin & Liu 1995)
genus Pentacentrus Saussure, 1878
P. acutiparamerus Liu & Shi, 2014 Yunnan (Liu et al. 2014)
P. biflexuous Liu & Shi, 2014 Guangxi (Liu et al. 2014)
P. birmanus Chopard, 1969 Tibet (Yin & Liu 1995)
P. bituberus Liu & Shi, 2011 Guangdong, Guangxi (Liu & Shi 2011c)
P. emarginatus Liu & Shi, 2014 Guangxi, Jiangxi (Liu et al. 2014)
P. formosanus Karny, 1915 Fujian, Guangxi, Taiwan, Yunnan (Karny 1915; Liu & Shi 2015b)
P. medogensis Zong, Qiu & Liu, 2017 Tibet (Zong et al. 2017)
P. multicapillus Liu & Shi, 2011 Guangxi (Liu & Shi 2011c)
P. parvulus Liu & Shi, 2014 Hainan (Liu et al. 2014)
P. sororius Zong, Qiu & Liu, 2017 Yunnan (Zong et al. 2017)
P. transversus Liu & Shi, 2015 Guangxi (Liu & Shi 2015b)
subfamily Podoscirtinae
genus Abaxitrella Gorochov, 2002
A. uncinata Ma & Gorochov, 2015 Fujian (Ma & Gorochov 2015)
genus Aphonoides Chopard, 1940
A. punctatus (Haan, 1842) Jiangxi (Yin & Liu 1995)
A. tessellatus Chopard, 1969 Hunan (Yin & Liu 1995)
A. wuyiensis Yin & Zhang, 2001 Fujian (Yin & Zhang 2001)
genus Mnesibulus Stål, 1877
M. (Mnesibulus) bicolor (Haan, 1844) Anhui, Hainan, Hunan, Jiangxi, Yunnan (Gorochov 2003b; Yin & Liu 1995)
M. (Mnesibulus) okunii Shiraki, 1930 Taiwan (Shiraki 1930)
genus Munda Stål, 1877
M. (Munda) javana (Saussure, 1878) Taiwan (Hsiung 1993)
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genus Noctitrella Gorochov, 1990
N. denticulata Liu & Shi, 2013 Yunnan (Liu & Shi 2013)
genus Phyllotrella Gorochov, 1988
P. planidorsalis Gorochov, 1988 Hainan (Yin & Liu 1995)
genus Pseudomadasumma Shiraki, 1930
P. maculata Shiraki, 1930 Taiwan (Shiraki 1930)
genus Sonotrella Gorochov, 1988
S. (Calyptotrella) quadrivittata Liu, Shi &
Ou, 2006
Yunnan (Liu et al. 2006)
S. (Sonotrella) major Liu, Yin & Wang,
1993
Yunnan (Liu et al. 1993a)
genus Trelleora Gorochov, 1988
T. consimilis Gorochov, 1988 Yunnan (Gorochov 1988a)
T. fumosa Gorochov, 1988 Yunnan (Liu & Shi 2014a)
T. kryszhanovskiji Gorochov, 1988 Yunnan (Gorochov 1988a)
genus Truljalia Gorochov, 1985
T. bispinosa Wang & Woo, 1992 Tibet (Wang & Woo 1992)
T. citri (Bey-Bienko, 1956) Guangdong, Hainan (Bey-Bienko 1956; Ma & Zhang
2015a)
T. elongata Liu & Shi, 2012 Hainan (Liu & Shi 2012b)
T. forceps (Saussure, 1878) Guangdong, Sichuan, Zhejiang (Ma & Zhang 2015a)
T. formosa He, 2012 Taiwan (He 2012)
T. hibinonis amota Gorochov, 2002 Sichuan (Liu & Shi 2012b)
T. hibinonis hibinonis (Matsumura, 1917) Guangxi, Hunan, Jiangsu, Jiangxi,
Sichuan, Yunnan, Zhejiang
(Yin & Liu 1995)
T. hubeiensis Ma & Zhang, 2015 Hubei (Ma & Zhang 2015a)
T. meloda Gorochov, 1992 Guangxi, Hainan, Yunnan (Liu & Shi 2011b; Wang & Woo
1992)
T. multiprotubera Liu & Shi, 2011 Yunnan (Liu & Shi 2011b)
T. panda Ma & Zhang, 2015 Shaanxi (Ma & Zhang 2015a)
T. tylacantha Wang & Woo, 1992 Anhui, Fujian, Guangdong, Guangxi,
Guizhou, Hubei, Hunan, Sichuan,
Zhejiang
(Wang & Woo 1992)
T. viminea Gorochov, 2002 Yunnan (Liu & Shi 2011b)
genus Valiatrella Gorochov, 2005
V. bimaculata (Chopard, 1928) Tibet (He & Gorochov 2015)
V. laminaria Liu & Shi, 2007 Guizhou (Liu & Shi 2007)
V. multiprotubera Liu & Shi, 2007 Yunnan (Liu & Shi 2007)
V. persicifolius Ma & Zhang, 2013 Hubei (Ma & Zhang 2013)
V. pulchra (Gorochov, 1985) Guizhou, Hunan (Ma & Zhang 2013)
V. sororia sororia (Gorochov, 2002) Guizhou (Ma & Zhang 2013)
genus Xuanwua He & Gorochov, 2015
X. motuoensis He & Gorochov, 2015 Tibet (He & Gorochov 2015)
genus Zamunda Gorochov, 2007
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Z. fuscirostris (Chopard, 1969) Fujian, Zhejiang (Yin & Liu 1995)
genus Zvenella Gorochov, 1988
Z. acutangulata Xia, Liu & Yin, 1991 Hainan (Xia et al. 1991)
Z. aequalis Ma & Zhang, 2012 Yunnan (Ma & Zhang 2012)
Z. decussatus Ma & Zhang, 2012 Yunnan (Ma & Zhang 2012)
Z. geniculata (Chopard, 1931) Yunnan (Gorochov 2002)
Z. scalpratus Ma & Zhang, 2012 Hainan (Ma & Zhang 2012)
Z. yunnana (Gorochov, 1985) Guangxi, Yunnan (Gorochov 1985; Yin & Liu 1995)
subfamily Sclerogryllinae
genus Sclerogryllus Gorochov, 1985
S. coriaceus (Haan, 1842) Guangxi, Taiwan, Yunnan (Yin & Liu 1995)
S. punctatus (Brunner von Wattenwyl,
1893)
Anhui, Guangxi, Hainan, Jiangsu,
Taiwan, Yunnan, Zhejiang
(Yin & Liu 1995)
S. tympanalis Yin & Liu, 1996 Hainan (Yin & Liu 1996)
family Phalangopsidae
subfamily Cachoplistinae
genus Cacoplistes Brunner von Wattenwyl,
1873
C. (Laminogryllus) brevisparamerus Wang,
Zhang, Wei & Liu, 2017
Guangxi (Wang et al. 2017)
C. (Laminogryllus) choui Liu & Shi, 2012 Guangxi (Liu & Shi 2012c)
genus Meloimorpha Walker, 1870
M. japonica japonica (Haan, 1842) Fujian, Guangxi, Hainan, Hebei, Hunan,
Jiangsu, Shandong, Taiwan, Zhejiang
(Yin & Liu 1995)
M. japonica yunnanensis (Yin, 1998) Yunnan (Gorochov 2003a; Yin 1998a)
family Phalangopsidae
subfamily Phaloriinae
genus Gorochovius Xie, Zheng & Li, 2004
G. trinervus Xie, Zheng & Li, 2004 Guangxi (Xie et al. 2004)
genus Phaloria Stål, 1877
P. (Phaloria) karnyello (Karny, 1915) Taiwan (Hsiung 1993; Karny 1915)
P. (Phaloria) kotoshoensis (Shiraki, 1930) Taiwan (Shiraki 1930)
P. liangi Xie & Zheng, 2003 Guangdong (Xie & Zheng 2003b)
genus Vescelia Stål, 1877
V. pieli pieli (Chopard, 1939) Guangdong, Hainan (Yin & Liu 1995)
family Trigonidiidae
subfamily Nemobiinae
genus Caconemobius Kirby, 1906
C. dibrachiatus Ma & Zhang, 2015 Guangdong (Ma et al. 2015d)
genus Dianemobius Vickery, 1973
D. chibae (Shiraki, 1911) Taiwan (Liu et al. 1998; Shiraki 1911)
D. chinensis Gorochov, 1984 Fujian, Guizhou (Gorochov 1984; Liu et al. 2016a)
D. csikii (Bolivar, 1901) Hainan, Hebei, Inner Mongolia, Sichuan,
Taiwan, Yunnan
(Liu et al. 2016a)
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Taxa Distribution Literature
D. fascipes (Walker, 1869)
16
Fujian, Guangdong, Hainan, Hebei,
Hubei, Hunan, Inner Mongolia, Jiangxi,
Jilin, Shandong, Taiwan, Yunnan,
Zhejiang
(Liu et al. 2016a)
D. furumagiensis (Ohmachi & Furukawa,
1929)
Guangxi, Hebei, Inner Mongolia,
Shandong, Sichuan, Taiwan, Zhejiang
(Liu et al. 2016a)
D. jucundus Liu & Yang, 1998 Taiwan (Liu et al. 1998)
D. kimurae (Shiraki, 1911) Taiwan (Liu et al. 1998; Shiraki 1911)
D. protransversus Liu & Yang, 1998 Taiwan (Liu et al. 1998)
D. wulaius Liu & Yang, 1998 Taiwan (Liu et al. 1998)
D. zhengi Chen, 1994 Sichuan (Chen 1994)
genus Homonemobius Chopard, 1935
H. nigrus Li, He & Liu, 2010 Yunnan (Li et al. 2010a)
genus Marinemobius Gorochov, 1985
M. asahinai (Yamasaki, 1979) Hainan (Gorochov 1985)
genus Polionemobius Gorochov, 1983
P. annulicornis Li, He & Liu, 2010 Yunnan (Li et al. 2010a)
P. flavoantennalis (Shiraki, 1911) Guizhou, Jiangsu, Jiangxi, Shandong,
Taiwan, Zhejiang
(Liu et al. 2016a; Shiraki 1911)
P. taprobanensis (Walker, 1869)
17
Fujian, Guangxi, Guizhou, Hainan,
Hebei, Heilongjiang, Henan, Hubei,
Inner Mongolia, Jiangsu, Jiangxi, Jilin,
Liaoning, Shandong, Sichuan, Yunnan,
Zhejiang
(Liu et al. 2016a)
P. yunnanus Liu & Shi, 2014 Yunnan (Liu & Shi 2014b)
genus Pteronemobius Jacobson, 1904
P. (Pteronemobius) caudatus (Shiraki,
1911)
Taiwan (Liu et al. 1998; Shiraki 1911)
P. (Pteronemobius) gifuensis (Shiraki,
1911)
Heilongjiang, Jilin, Sichuan (Shiraki 1911; Yin & Liu 1995)
P. (Pteronemobius) heydenii concolor
(Walker, 1871)
Hainan, Inner Mongolia, Jilin, Tibet,
Xinjiang, Yunnan
(Yin & Liu 1995)
P. (Pteronemobius) indicus (Walker,1869) Guangxi, Hainan, Hubei, Yunnan (Liu et al. 2016a)
P. (Pteronemobius) jianfenglingensis Liu &
Shi,2016
Hainan (Liu et al. 2016a)
P. (Pteronemobius) kangdingensis Liu &
Shi, 2014
Sichuan (Liu et al. 2016a; Liu & Shi 2014c)
P. (Pteronemobius) neimongolensis Kang
& Mao, 1990
Inner Mongolia (Kang & Mao 1990)
P. (Pteronemobius) nigriscens (Shiraki,
1911)
Taiwan (Shiraki 1911)
P. (Pteronemobius) nitidus (Bolivar, 1901) Fujian, Guangxi, Hainan, Hebei, Hunan,
Jiangsu, Jiangxi, Sichuan, Yunnan,
Zhejiang
(Yin & Liu 1995)
P. (Pteronemobius) ohmachii Shiraki, 1930 Taiwan (Liu et al. 1998; Shiraki 1930)
P. (Pteronemobius) pilicornis Chopard,
1969
Fujian (Liu et al. 2016a)
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P. (Pteronemobius) qinghaiensis Yin, 1998 Qinghai (Yin 1998b)
P. (Pteronemobius) taibaiensis Deng & Xu,
2006
Shaanxi (Deng & Xu 2006)
P. (Pteronemobius) yezoensis (Shiraki,
1911)
Jinlin This study
18
P. (Pteronemobius) yunnanicus Li, He &
Liu, 2010
Yunnan (L i et al. 2010a)
genus Speonemobius Chopard, 1924
S. bifasciatus He, Lu, Wang & Li, 2016 Yunnan (He et al. 2016)
S. fulvus He, Lu, Wang & Li, 2016 Yunnan (He et al. 2016)
Speonemobius? sinensis Li, He & Liu,
2010
Zhejiang (He et al. 2016; Li et al. 2010a)
genus Stenonemobius Gorochov, 1981
S. (Ocellonemobius) bicolor bicolor
(Saussure, 1877)
Hainan (Yin & Liu 1995)
genus Taiwanemobius Yang & Chang, 1996
T. formosanus Yang & Chang, 1996 Taiwan (Yang & Chang 1996)
subfamily Trigonidiinae
genus Amusurgus Brunner von Wattenwyl,
1893
A. (Amusurgus) bispinosus He, Li, Fang &
Liu, 2010
Yunnan (H e et al. 2010)
A. (Amusurgus) fulvus Brunner von
Wattenwyl, 1893
Hainan, Yunnan, Zhejiang (He et al. 2010)
A. (Paranaxipha) fujianensis Wang, Zheng
& Woo, 1999
Fujian, Yunnan, Zhejiang (He et al. 2010; Wang et al. 1999)
A. (Paranaxipha) hainanensis He, Li, Fang
& Liu, 2010
Hainan (He et al. 2010)
A. (Usgmona) excavatus Liu, Shi & Zhou,
2015
Fujian (Liu et al. 2015a)
A. (Usgmona) genji (Furukawa, 1970) Jiangsu, Zhejiang (He et al. 2010)
genus Anaxiphomorpha Gorochov, 1987
A. biserratus Liu & Shi, 2015 Hainan (Liu & Shi 2015a)
A. brevisparamerus Liu & Shi, 2015 Yunnan (Liu & Shi 2015a)
A. longiserratus Liu & Shi, 2015 Guizhou (Liu & Shi 2015a)
A. serratiprotuberus Liu & Shi, 2015 Yunnan (Liu & Shi 2015a)
genus Homoeoxipha Saussure, 1874
H. eurylobus Ma, Liu & Zhang, 2016 Tibet (Ma et al. 2016)
H. lycoides (Walker, 1869) Yunnan, Hainan (Ma et al. 2016)
H. nigriceps Chopard, 1961 Guizhou, Hunan, Sichuan, Zhejiang (Ma et al. 2016)
H. obliterata (Caudell, 1927) Anhui, Henan (Ma et al. 2016)
genus Metioche Stål, 1877
M. (Metioche) bicolor (Stål, 1861) Hainan (Yin & Liu 1995)
M. (Metioche) flavipes (Brunner von
Wattenwyl, 1878)
Zhejiang (Yin & Liu 1995)
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Taxa Distribution Literature
M. (Metioche) haani (Saussure, 1878) Guizhou, Hubei, Hunan, Jiangxi,
Sichuan, Taiwan, Zhejiang
(Yin & Liu 1995)
M. (Metioche) japonica (Ichikawa, 2001) Yunnan This study
19
M. (Metioche) kotoshoensis (Shiraki, 1930) Taiwan (Shiraki 1930)
M. (Metioche) pallidicornis Stål, 1861 Guangdong (Yin & Liu 1995)
M. (Metioche) pallipes Stål, 1861 Hainan (Yin & Liu 1995)
M. (Metioche) vittaticollis vittaticollis
(Stål, 1861)
Taiwan (Yin & Liu 1995)
genus Metiochodes Chopard, 1932
M. acutiparamerus Li, He & Liu, 2010 Hainan (Li et al. 2010b)
M. denticulatus Liu & Shi, 2011 Guizhou (Liu & Shi 2011a)
M. flavescens Chopard, 1932 Yunnan (Li et al. 2010b)
M. minor Li, He & Liu, 2010 Fujian (Li et al. 2010b)
M. tibeticus Li, He & Liu, 2010 Tibet (Li et al. 2010b)
M. truncatus Li, He & Liu, 2010 Yunnan (Li et al. 2010b)
genus Natula Gorochov, 1987
N. longipennis (Serville, 1838) Hainan, Tibet, Yunnan (Yin & Liu 1995)
N. matsuurai Sugimoto, 2001 Zhejiang This study
20
N. pallidula (Matsumura, 1910) Jiangsu, Taiwan, Zhejiang (Yin & Liu 1995)
N. pravdini (Gorochov, 1985) Fujian (Gorochov 1985)
genus Paratrigonidium Brunner von
Wattenwyl, 1893
P. chloropodum He, 2017 Hainan (He et al. 2017a)
P. majusculum (Karny, 1915) Taiwan (Hsiung 1993; Karny 1915)
P. nitidum Brunner von Wattenwyl, 1893 Guangxi, Jiangxi, Tibet, Yunnan (Yin & Liu 1995)
P. striatum (Shiraki, 1911) Taiwan (Hsiung 1993; Shiraki 1911)
P. transversum (Shiraki, 1930) Taiwan (Hsiung 1993; Shiraki 1930)
P. venustulum (Saussure, 1878) Yunnan (He et al. 2017a; Yin & Liu 1995)
genus Svistella Gorochov, 1987
S. anhuiensis He, Li & Liu, 2009 Anhui (He et al. 2009)
S. bifasciata (Shiraki, 1911) Anhui, Guangxi, Hainan, Hunan, Jiangsu,
Jiangxi, Taiwan, Zhejiang
(He et al. 2009; Shiraki 1911)
S. dubia (Liu & Yin, 1993) Yunnan (He et al. 2009; Liu et al. 1993b)
S. fallax He, Li & Liu, 2009 Sichuan (He et al. 2009)
S. rufonotata (Chopard, 1932) Guangdong, Hunan, Yunnan (He et al. 2009)
S. tympanalis He, Li & Liu, 2009 Anhui, Guangxi, Tibet, Yunnan, Zhejiang (He et al. 2009)
genus Trigonidium Rambur, 1838
T. (Trigonidium) cicindeloides Rambur,
1839
Fujian, Guangdong, Guangxi, Guizhou,
Hainan, Jiangsu, Sichuan, Taiwan,
Yunnan, Zhejiang
(Yin & Liu 1995)
T. (Trigonidium) humbertianum (Saussure,
1878)
Fujian, Hainan, Yunnan (Yin & Liu 1995)
family Mogoplistidae
subfamily Mogoplistinae
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genus Arachnocephalus Costa, 1855
A. brevissimus Shiraki, 1911 Taiwan (Shiraki 1911; Yang & Yen 2001a)
genus Cycloptiloides Sjöstedt, 1910
C. orientalis Chopard, 1925 Taiwan (Yang & Yen 2001a)
genus Ectatoderus Guérin-Méneville, 1847
E. annulipedus (Shiraki, 1911) Taiwan (Shiraki 1911; Yang & Yen 2001a)
E. leuctisonus Yang & Yen, 2001 Taiwan (Yang & Yen 2001a)
genus Micrornebius Chopard, 1969
M. hainanensis Yin, 1998 Hainan (Yin 1998b)
M. perrarus Yang & Yen, 2001 Taiwan (Yang & Yen 2001a)
genus Ornebius Guérin-Méneville, 1844
O. bimaculatus (Shiraki, 1930) Guangdong, Taiwan (He et al. 2017b; Shiraki 1930; Yang
& Yen 2001a)
O. fastus Yang & Yen, 2001 Taiwan (Yang & Yen 2001a)
O. formosanus (Shiraki, 1911) Guangxi, Hainan, Taiwan, Zhejiang (He et al. 2017b; Shiraki 1911; Yang
& Yen 2001a)
O. fuscicercis (Shiraki, 1930) Taiwan (Shiraki 1930; Yang & Yen 2001a)
O. infuscatus (Shiraki, 1930) Taiwan (Shiraki 1930; Yang & Yen 2001a)
O. kanetataki (Matsumura, 1904) Guangdong, Taiwan, Zhejiang (He et al. 2017b; Yang & Yen
2001a)
O. longipennis longipennis (Shiraki, 1930) Taiwan (Shiraki 1930; Yang & Yen 2001a)
O. polycomus He, 2017 Zhejiang (He et al. 2017b)
superfamily Gryllotalpoidea Leach, 1815
family Myrmecophilidae
subfamily Myrmecophilinae
genus Myrmecophilus Berthold, 1827
M. (Myrmecophilus) formosanus Shiraki,
1930
Taiwan, Zhejiang (Shiraki 1930; Yin & Liu 1995)
M. (Myrmecophilus) quadrispina Perkins,
1899
Guangdong (Yin & Liu 1995)
M. (Myrmecophilus) sinicus Bey-Bienko,
1956
Sichuan (Bey-Bienko 1956; Yin & Liu 1995)
family Gryllotalpidae Leach, 1815
subfamily Gryllotalpinae
genus Gryllotalpa Latreille, 1802
G. breviabdominis Ma & Zhang, 2011 Henan (Ma & Zhang 2011b)
G. choui Ma & Zhang, 2010 Yunnan (Ma & Zhang 2010)
G. cycloptera Ma & Zhang, 2011 Zhejiang (Ma & Zhang 2011b)
G. dentista Yang, 1995 Taiwan (Yang 1995)
G. formosana Shiraki, 1930 Taiwan (Shiraki 1930)
G. henana Cai & Niu, 1998 Henan (Cai & Niu 1998)
G. jinxiuensis You & Li, 1990 Guangxi (You & Li 1990)
G. mabiana Ma, Xu & Takeda, 2008 Sichuan (Ma et al. 2008)
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Discussion
1 Modicogryllus (Modicogryllus) maculatus (Shiraki, 1930) is considered Comidoblemmus Storozhenko & Paik,
2009 as C. maculatus (Shiraki, 1930) comb. nov. by study of photos of type specimens on Orthoptera Species Files
Online.
2 Genus Goniogryllus was established on the species punctatus from Fujian, China (Chopard, 1936a). Absence
of wings is one of the important characters for this genus. Qingryllus was established on species striofemorus from
Shaanxi, China (Chen & Zheng, 1995a). Well developed wings is the main character to distinguish it from
Goniogryllus. In 1996, a Goniogryllus sp. with short wings was published as ovalatus (Chen & Zheng, 1996).
From that time, identification characters for these two genera has been ambiguous. I compared the COI genes
between G. ovalatus and Q. striofemorus, and found they were totally same. I suspect G. spp. show varied
phenotypes to be published as different species. Here, I do not make synonym of species, because more data is
needed. But Qingryllus Chen & Zheng, 1995 syn. is considered to be the junior synonym of Goniogryllus Chopard,
1936.
3 There are many species recorded from China based on the morphology of the frons in this genus. But these
features are variable due to environmental factors (He & Takeda 2014). Thus, there may be some
misidentifications. DNA markers and/or songs should be used in future research. Loxoblemmus abotus Wang ,
1992, Loxoblemmus brevipalpus Wang, 1992, Loxoblemmus neoarietulus Wang, 1992, Loxoblemmus subangulatus
Yang, 1992 have not been officially published (Hsiung 1993).
4 Loxoblemmus angulatus Bey-Bienko, 1956 is widely distributed in south China. Most research papers cite
this species. But this species has no taxonomic or morphological differences with Loxoblemmus appendicularis
Shiraki, 1930 with the latter one widely distributed in Taiwan. Thus, L. angulatus Bey-Bienko, 1956 syn. is
considered to be the junior synonym of L. appendicularis.
5 Modicogryllus (Modicogryllus) consobrinus (Saussure, 1877) is usually identified as Modicogryllus
(Modicogryllus) confirmatus (Walker, 1859) by Yin & Liu (1995).
6 Svercacheta siamensis (Chopard, 1961) is identified as Tartarogryllus minusculus Walker, 1869 by Yin &
Liu (1995).
7 Holotype of Cophogryllus kuhlgatzi Karny, 1908 is a nymph of Teleogryllus (Brachyteleogryllus) occipitalis
occipitalis (Serville, 1838). Thus, C. kuhlgatzi syn. is considered to be the junior synonym of T. (B.) occipitalis
occipitalis.
8 Velarifictorus (Velarifictorus) koshunensis (Shiraki, 1911) is considered under genus Turanogryllus
Tarbinsky, 1940 as T. koshunensis (Shiraki, 1911) comb. nov. as indicated by photos of type specimens on
Orthoptera Species Files Online.
9 Some species are widely distributed from south to north. Their life cycles can change from nymph diapause
to egg diapause, from biovoltine to univoltine, from short ovipositor to long ovipositor, which are all adaptations to
local climate. These situations also occur in many other insect groups, but they are always to be considered as one
species. Thus, here I consider Velarifictorus (Velarifictorus) aspersus borealis Gorochov, 1985 syn. as the junior
synonym of V. (V.) aspersus aspersus (Walker, 1869). Now there is no subspecies under this species.
(Continued)
Taxa Distribution Literature
G. orientalis Burmeister, 1838
21
Fujian, Guangdong, Guangxi, Guizhou,
Hainan, Hebei, Heilongjiang, Hubei,
Hunan, Inner Mongolia, Jiangsu, Jiangxi,
Jilin, Liaoning, Qinghai, Shandong,
Sichuan, Tibet, Yunnan, Zhejiang
(Yin & Liu 1995)
G. unispina Saussure, 1874 Anhui, Gansu, Hebei, Hubei, Inner
Mongolia, Jiangsu, Jiangxi, Jilin,
Liaoning, Ningxia, Shanxi, Tibet,
Xinjiang
(Yin & Liu 1995)
G. wudangensis Li, Ma & Xu, 2007 Hubei (Li et al. 2007)

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10 Modicogryllus (Modicogryllus) latefasciatus (Chopard, 1933) syn. is a phenotype of Velarifictorus
(Velarifictorus) micado (Saussure, 1877). There is a thin white stripe between stemmata in most V. (V.) micado
individuals. But sometimes there is a big white or yellow area between stemmata instead of the thin stripe. Thus, I
consider M. (M.) latefasciatus syn. as the junior synonym of V. (V.) micado. Japanese researcher consider V. (V.)
micado individuals in south Japan as V. (V.) grylloides (Chopard, 1969) or V. (V.) ryukyuensis Oshiro, 1990. But I
consider all individuals in China and Japan as one species V. (V.) micado. See discussion 9.
11 Velarifictorus (Velarifictorus) ornatus caudatus (Shiraki, 1930) syn. is the macropterous individual of V. (V.)
ornatus ornatus (Shiraki, 1911). Therefore, there are no subspecies under this species.
12 male, CHINA, Yunnan, Xishuangbanna, Menglun, 4-iv-2017, coll. He Zhu-Qing.
13 male, CHINA, Fujian, Xiamen, 13-x-2017, coll. He Zhu-Qing.
14 male, CHINA, Zhejiang, Hangzhou, 8-ix-2017, coll. He Zhu-Qing.
15 male, CHINA, Yunnan, Jinggu, 8-viii-2017, coll. He Zhu-Qing.
16 Dianemobius nigrofasciatus (Matsumura, 1904) syn. is the north group of Dianemobius fascipes (Walker,
1869). See discussion 9.
17 Polionemobius mikado (Shiraki, 1911) syn. is the north group of Polionemobius taprobanensis (Walker,
1869). See discussion 9.
18 male, CHINA, Jilin, Changbaishan, 10-vii-2016, coll. He Zhu-Qing.
19 male, CHINA, Yunnan, Xishuangbanna, Bubang, 5-viii-2017, coll. He Zhu-Qing.
20 male, CHINA, Zhejiang, Hangzhou, 1-x-2017, coll. He Zhu-Qing.
21 Holotype of Gryllotalpa oryctes Scudder, 1869 syn. is from Shanghai, China. Thus, it should be considered
as the junior synonym of Gryllotalpa orientalis Burmeister, 1838, not Gryllotalpa africana africana Palisot de
Beauvois, 1805.
Acknowledgments
This study is mainly based on Orthoptera Species Files Online and is sponsored by Shanghai Sailing Program
(grant No. 16YF1403100). I am also appreciated to Dr. Wang Han-Qiang and Chen Po-Wei for providing me with
literature.
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