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Doronicum giganteum (Griseb.) D.V. Georgiev comb. et status nova – new species in the Bulgarian flora

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The basionym Doronicum pardalianches var. giganteum Grisebach was used to describe a species new to the science, phylogenetically related to Doronicum pardalianches, but distinct from all European species by its oval and elliptic-lobate bracts of the flower basket. The range of the new species covers Albania, Bulgaria, Greece, Macedonia, Serbia and Anatolian Turkey. The new species has a significant number of floral features that clearly distinguish it from other species, but also has larger sizes plant vegetative parts, which clearly justifies the name of the giant taxa with one of the highest and greatest habitus of the rest European representatives of the genus. The name of the new species in Bulgarian is a "giant wild sunflower". The English name is "giant leopard's-banes".
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Doronicum giganteum (Griseb.) D.V. Georgiev comb. et status
novae – нов вид във флората на България.
Димитър Василев Георгиев, e-mail: d_georgieff@yahoo.co.uk
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Както е видно от оригиналното описание на таксона прицветниците на Doronicum
orphanidis са ланцетни към върха стеснено-заострени. Това приближава Doronicum
orphanidis към Doronicum austriacum, вместо към Doronicum austriacum subsp. giganteum
(Griseb.) Stoj. & Stef и новия статус на видовия таксон.
В книгата си за видовия състав на флорите на Румелия и Битиния Август Гризебах
описва следния таксон:
„Doronicum Pardalianches L. var. giganteum, sexpedale, inferne subhirsutum vel
glabriusculum, superne ample paniculatum dense glanduliferum, foliis imis —, inferioribus
late ovatis denticulatis, petiolo anguste foliaceo basi auriculato, superioribus
oblongo-lanceolatis semiamplexicaulibus, ramis paniculae erecto-patentibus snbtricephalis,
involucri squamis basi ovatolanceolalis lineari-acuminatis radio aureo duplo superatis,
receptaculo convexo villoso, ligulis basi pilosis, acheniis radii glabriusculis. — Ic. n. Hayne
Arzn. 6. t. 21.: rhizoma ovatum rotundatum glabrum stoloniferum in nostris specim. deest.
IL 2. p. 263. In fagetis Scardi alt. 3000'—4360': copiose in m. Ljubatrin (substr. calc.)! Fl.
Jul. E.”
В описанието на таксона описан от Аугуст Гризебах ясно се упоменава наличието на
долни стъблови листа с дръжки, което ясно отличава посочената изменчивост от
Doronicum austriacum. От друга страна описанието „acheniis radii glabriusculis”
доближава въпросната изменчивост до Doronicum austriacum..
())*+<<=*+BC?!D(!l!C)<X9
Description: Perrenial plant 80-170(200) cm tall with peculiar unpleasant smell. Rhizome
dark, fleshy to somewhat woody, glabrous. Stems not branching, generally scape-like.
Indumentum of uniseriate, multiseriate eglandular trichomes, short-stalked and long-stalked
glandular trichomes,
scarce at the base, more abundant near the capitula. Leaves dentate to slightly dentate. Basal
leaves generally not present at flowering time; blade denticulata 10-35 cm long et 7-22(25)
cm wide, ovate-orbicular to broadly ovate with deeply cordate base, with blunt or sub-acute
apex; petiole up to 40 cm long and up to mm wide. Lower cauline leaves 8-20 cm long and 7-
15 cm wide, ovate, cordate, middle cauline leaves 6-17 cm long et 3-8 cm wide, ovate,
panduriform, sessile or rarely petiolate, amplexicaul. Upper cauline leaves 4-9 x 1.5-6 cm,
ovate to ovate-lanceolate, sometimes bract-like. Indumentum of eglandular and glandular
trichomes. Capitula 1–20, 4–8 cm diam. including rays; involucres 3-4 times
shorter than the ligulate florets. Phyllaries herbaceous, ovate or elliptic-lanceolate, generally
with acute apex; the outer 5-10 mm long and 3-6 mm wide, the inner 7-11 mm long, 3-5 mm
wide; margins ciliate. Indumentum mainly glandular, but also with uniseriate eglandular
trichomes. Receptacle pubescent. Flowers with yellow corollas. Ray flower
corollas 3-4 cm long, 1.5–5 mm wide, oblongelliptic, apex generally with three teeth.
Cypselae 2,5-5 mm long and 1,5-2 mm wide, brown, dimorphic; achenes of tubular and
ligulate florets sparse or dense hairy. Pappus 3–5 mm, white, marginal pappus of ligulate
florets pauci-setose, pappus of tubular florets pluri-setose.
Typus: Supra pagum Yundola et sub monte Belmeken. Mons Rila (Bulgaria), 1400-2440 m,
12-06-1992 l., limes silvae pineae D.V.Georgieff (Holotypus). In silvis et arbusculas strati
montani - strati subalpini et strati alpini (2450 m.). Leg. D.V. Georgieff
Doronicum giganteum (Nomen Provisorum Doronicum ovatobracteatum) - Planta perrenis
80-170(200) cm alta, erecta. Molliter pilosum. Radix cylindrica, glabra. Folia radicalia in
planta florifera nullis, in planta non florifera ovato-orbiculata, cordata, debiliter denticulata
10-35 cm lg. et 7-22(25) cm lt. Folia caulinaria denticulata; inferiora petiolata, ovato-
orbiculata, cordata; media panduriformia, ovato-orbiculata, cordata; superiora sessilia, ovata
vel ovato-oblonga. Capitulo diversa numero 1-20, 4-8 cm in diametro. Involucrum
hemisphaericum; foliola involucri ovata vel eliptico-lanceolata; involucro ligulae triplo vel
quadruplo breviore. Ligulis flavis longioribus quam bracteis. Achaeniis brunneis. Internis cum
pilis simplicibus in costis et cum vilanis pilorum scabridum. Externis sparse vel dense pilosis.
Receptaculo piloso cum facibus pilorum simplicium uniseniatorum. Pappus setosus, 1-2-
serialis, setis 3-5 mm lg.; pappus marginalis paucisetosus, pappus internis pluri-setosus.
Speciei Doronicum cataractarum Widder, D. pardalianches L. et D. austriacum Jacq. similis, a
qua praesertim differt bracteis involucralibus ovatis, foliis basalis ad 35cm lg. x 22cm lt.,
foliis caulinis inferioribus et mediis petiolatis.
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В заключение новия вид Doronicum giganteum има сходни белези както с Doronicum
pardalianches, така и с Doronicum austriacum, но от друга страна притежава белези,
характерни за D. pardalianches и D. catharactarum като наличието на дръжки на долните
стъблови листа. Налице са и белези характерни само за Doronicum giganteum без да се
повтарят в европейските представители на рода като яйцевидни до елиптично ланцетни
прицветници и диференцирана морфология на маргиналните спрямо вътрешните
плодосемки, наред със сравнително по-големия хабитус на растението от споменатите
два близки до него вида.
В нито един европейски представители на род Doronicum вариацията на
прицветниковата форма не влючва овалните очертания, а вместо това при европейските
преставители те варират от линейни през линейно-ланцетни до ланцетни, което прави
вида Doronicum giganteum сравнително обособен видов таксон наред с останалите му
видови характеристики. Хербарни образци от новия вид Doronicum giganteum са
внесени под провизорното име Doronicum ovatobracteatum в хербариумите на Кралската
ботаническа градина Кю и Берлинския хербариум.
Използвани таксони в статията:
())*+  ! <<=*+C?!E!`!b*+!kkST!
SQrv  9  ?)j+
())*+ *=*+ *?! <<=*+ BC?!D E=)F!G E=H! `! Zx<!"
!  k  SRSr!  Squu
())*+)Z)!`!g=!uuTQ!SQTp
y#
Кузманов, Б. 2012. Doronicum L.- В: Пеев, Д.(ред.). Флора на Р. България, т. 11, С. БАН:
423-426.
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Стоянов, Н., Стефанов, Б. „Флора на България“ том 2, 2-ро издание, 1933
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