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Revision of the tribe Cerambycini: redefinition of the genera Trirachys Hope, 1843, Aeolesthes Gahan, 1890 and Pseudaeolesthes Plavilstshikov, 1931 (Coleoptera, Cerambycidae)

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The systematics of the Australasian genera Trirachys Hope, 1843, Aeolesthes Gahan, 1890 and Pseudaeolesthes Plavilstshikov, 1931 is revised through analysis of the types-species and analytic methods of statistical similarity. Pseudaeolesthes is restored as b. genus and includes the following taxa: Pseudaeolesthes aureopilosa (Gressitt & Rondon, 1970) n. comb., Pseudaeolesthes chrysothrix (Bates, 1873) rest. comb., Pseudaeolesthes chrysothrix taiwanensis (Hayashi, 1974) n. comb., Pseudaeolesthes chrysothrix nakamurai (Kusama & Takakuwa, 1984) n. comb., Pseudaeolesthes chrysothrix kurosawai Gressitt, 1965 rest. comb., Pseudaeolesthes chrysothrix yonaguniensis Ohbayashi & Ohbayashi, 1965 rest. comb., Pseudaeolesthes chrysothrix tibetana Gressitt, 1942 rest. comb., Pseudaeolesthes malayana (Hayashi, 1979) n. comb., Pseudaeolesthes multistriata (Hayashi, 1979) n. comb., Pseudaeolesthes mutabiliaurea Chiang, 1951 rest. comb., Pseudaeolesthes psednothrix (Gressitt & Rondon, 1970) n. comb. and Pseudaeolesthes rufimembris (Pic, 1923) n. comb. Carinolesthes n. gen. for Carinolesthes pericalles (Gressitt & Rondon, 1970) n. comb., Carinolesthes aureosignata (Pic, 1915) n. comb. and Carinolesthes ningshanensis (Chiang, 1981) n. comb. is proposed. Massirachys n. gen. for Massirachys mariae (Thomson, 1878) n. comb. is proposed. Parolesthes n. gen. for Parolesthes laosensis (Gressitt & Rondon, 1970) n. comb. and Parolesthes curticornis (Hüdepohl, 1988) n. comb. is proposed. Aeolesthes includes the following species: Aeolesthes aurifaber (White, 1853), Aeolesthes gloriosa (Aurivillius, 1924) n. comb., Aeolesthes bilobulartus (Gressitt & Rondon, 1970) n. comb. and Aeolesthes vietnamensis n. sp. from Vietnam. Trirachys Hope, 1843 includes the following species: Trirachys acanthophorus Vitali, 1999, Trirachys achilles (Thomson, 1865) n. comb., Trirachys ampliatus (Gahan, 1890) n. comb., Trirachys atkinsoni Gardner, 1939, Trirachys basicornis (Gahan, 1893) n. comb., Trirachys externus (Pascoe, 1869) n. comb., Trirachys holosericeus (Fabricius, 1787) n. comb., Trirachys inexpectatus Holzshuh, 1982, Trirachys indicolus (Bates, 1891) n. comb., Trirachys indutus (Newman, 1842) n. comb., Trirachys inhirsutus (Matsushita, 1932) n. comb., Trirachys orientalis Hope, 1843, Trirachys perplexus (Gahan, 1890) n. comb., Trirachys sartus (Solsky, 1871) n. comb., Trirachys sinensis (Gahan, 1890) n. comb., Trirachys sphaericothorax Gressitt & Rondon, 1970, and Trirachys textor (Pascoe, 1869) n. comb. Aeolesthes (Pseudaeolesthes) chrysophanes Gressitt & Rondon, 1970 is transferred to the genus Elydnus Pascoe, 1869, as follows: Elydnus chrysophanes (Gressitt & Rondon, 1970) n. comb. Aeolesthes sticheri Hüdepohl, 1989 and Aeolesthes fulgens Schwarzer, 1926 are transferred to the genus Dymasius (s. str.) Thomson, 1864, as follows: Dymasius sticheri (Hüdepohl, 1989) n. comb. and Dymasius fulgens (Schwarzer, 1926) n. comb. Derolus Gahan, 1891 = Mimoderolus Pic, 1933 n. syn. and Derolus uniformis (Pic, 1933) n. comb. = Pachydissus xyliae Fisher, 1940 n. syn. are proposed.
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Revision of the tribe Cerambycini: redefinition of the genera Trirachys
Hope, 1843, Aeolesthes Gahan, 1890 and Pseudaeolesthes Plavilstshikov,
1931
(Coleoptera, Cerambycidae)
Francesco Vitali (1), Xavier Gouverneur (2), Gérard Chemin (3)
(1) Nationalmusée fir Naturgeschicht,
25, rue Münster,
L-2160 Luxembourg
(2) 3, rue de la Santé,
F-35000 Rennes
(3) 450, rue Marcel Paul,
F-94500 Champigny sur Marne
___________________
Abstract
The systematics of the Australasian genera Trirachys Hope, 1843, Aeolesthes Gahan, 1890 and
Pseudaeolesthes Plavilstshikov, 1931 is revised through analysis of the types-species and analy-
tic methods of statistical similarity. Pseudaeolesthes is restored as b. genus and includes the fol-
lowing taxa: Pseudaeolesthes aureopilosa (Gressitt & Rondon, 1970) n. comb., Pseudaeolesthes
chrysothrix (Bates, 1873) rest. comb., Pseudaeolesthes chrysothrix taiwanensis (Hayashi, 1974)
n. comb., Pseudaeolesthes chrysothrix nakamurai (Kusama & Takakuwa, 1984) n. comb.,
Pseudaeolesthes chrysothrix kurosawai Gressitt, 1965 rest. comb., Pseudaeolesthes chrysothrix
yonaguniensis Ohbayashi & Ohbayashi, 1965 rest. comb., Pseudaeolesthes chrysothrix tibetana
Gressitt, 1942 rest. comb., Pseudaeolesthes malayana (Hayashi, 1979) n. comb., Pseudaeolesthes
multistriata (Hayashi, 1979) n. comb., Pseudaeolesthes mutabiliaurea Chiang, 1951 rest. comb.,
Pseudaeolesthes psednothrix (Gressitt & Rondon, 1970) n. comb. and Pseudaeolesthes rufimembris
(Pic, 1923) n. comb. Carinolesthes n. gen. for Carinolesthes pericalles (Gressitt & Rondon, 1970)
n. comb., Carinolesthes aureosignata (Pic, 1915) n. comb. and Carinolesthes ningshanensis
(Chiang, 1981) n. comb. is proposed. Massirachys n. gen. for Massirachys mariae (Thomson,
1878) n. comb. is proposed. Parolesthes n. gen. for Parolesthes laosensis (Gressitt & Rondon,
1970) n. comb. and Parolesthes curticornis (Hüdepohl, 1988) n. comb. is proposed. Aeolesthes
includes the following species: Aeolesthes aurifaber (White, 1853), Aeolesthes gloriosa (Aurivil-
lius, 1924) n. comb., Aeolesthes bilobulartus (Gressitt & Rondon, 1970) n. comb. and Aeolesthes
vietnamensis n. sp. from Vietnam. Trirachys Hope, 1843 includes the following species: Trirachys
acanthophorus Vitali, 1999, Trirachys achilles (Thomson, 1865) n. comb., Trirachys ampliatus
(Gahan, 1890) n. comb., Trirachys atkinsoni Gardner, 1939, Trirachys basicornis (Gahan, 1893)
n. comb., Trirachys externus (Pascoe, 1869) n. comb., Trirachys holosericeus (Fabricius, 1787) n.
comb., Trirachys inexpectatus Holzshuh, 1982, Trirachys indicolus (Bates, 1891) n. comb., Trirachys
Les Cahiers Magellanes, No26, juin 2017 — 40
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indutus (Newman, 1842) n. comb., Trirachys inhirsutus (Matsushita, 1932) n. comb., Trirachys
orientalis Hope, 1843, Trirachys perplexus (Gahan, 1890) n. comb., Trirachys sartus (Solsky,
1871) n. comb., Trirachys sinensis (Gahan, 1890) n. comb., Trirachys sphaericothorax Gressitt &
Rondon, 1970, and Trirachys textor (Pascoe, 1869) n. comb. Aeolesthes (Pseudaeolesthes) chrysophanes
Gressitt & Rondon, 1970 is transferred to the genus Elydnus Pascoe, 1869, as follows: Elydnus
chrysophanes (Gressitt & Rondon, 1970) n. comb. Aeolesthes sticheri Hüdepohl, 1989 and
Aeolesthes fulgens Schwarzer, 1926 are transferred to the genus Dymasius (s. str.) Thomson,
1864, as follows: Dymasius sticheri (Hüdepohl, 1989) n. comb. and Dymasius fulgens (Schwar-
zer, 1926) n. comb. Derolus Gahan, 1891 = Mimoderolus Pic, 1933 n. syn. and Derolus uniformis
(Pic, 1933) n. comb. = Pachydissus xyliae Fisher, 1940 n. syn. are proposed.
Key-words
Coleoptera, Cerambycidae, Cerambycidae, Cerambycini, systematics.
___________________
Introduction
During the description of Trirachys acanthophorus (Vitali, 1999), the generic attribution
was doubtful because the general habitus exactly fitted Aeolesthes induta (Newman, 1842) while
the antennal structure corresponded to the genus Trirachys Hope, 1843.
According to the original description, Trirachys included only one species (T. orientalis
Hope, 1843) characterised by prothorax spined at the sides antennae and endoapically spined
from the 3rd article. On the contrary, Aeolesthes Gahan, 1890 (type-species: Hammaticherus
aurifaber White, 1853) was characterised by mutic prothorax and antennae endoapically spined
from the 5th article. Finally, Pseudaeolesthes Plavilstshikov, 1931 included only one species
(Neocerambyx chrysothrix Bates, 1873) characterised by spined prothorax and mutic antennae.
All also shared interantennal ridge and rounded procoxal cavities.
The status of these genera became more and more confusing in the subsequent years. Auri-
villius (1924) described Trirachys gloriosus since it showed a spined prothorax. But it also sho-
wed antennae spined from 5th article and a habitus extremely similar to the type-species of
Aeolesthes. Gressitt & Rondon (1970) provided erroneous and misleading differential charac-
ters, which drove to key Aeolesthes twice, to downgrade Pseudaeolesthes to subgenus level, and
to confuse the taxonomy of many species. They even provided differential diagnoses confron-
ting species of different genera and described bilobulartus as Trirachys, though it showed typi-
cal Aeolesthes-characters. Holzschuh (1982) described a Trirachys-species with antennae spi-
ned from the 3rd article but with a mutic prothorax as well. Actually, Trirachys acanthophorus
shared analogous characters. Finally, Nakamura et al. (1992) transferred Hemadius oenochrous
Fairmaire, 1889 to Aeolesthes, though it showed none of the characters of this genus.
In conclusion, several species currently classified as Trirachys or Aeolesthes often show only
a superficial likeness with their type-species. They were inserted in the respective genera only
according to one or another particular character, sometimes the antennal or the prothoracic
spines, sometimes the general habitus.
The aim of this paper is to re-establish the natural systematic relations among Trirachys and
closely related genera through the compared analysis of morphological and genital characters.
The use of statistical methods supported this task.
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Materials and methods
Species belonging to the following collections have been examined. The types belonging to
the Bishop Museum have been examined through hi-definition pictures.
AMNH: American Museum of Natural History, New York (USA).
BMNH: British Museum of Natural History, London (Great Britain).
BPBM: Bernice Pauahi Bishop Museum, Honolulu (USA).
CFV: Francesco Vitali private collection (Luxembourg).
CGC: Gérard Chemin private collection, Champigny-sur-Marne (France).
CPH: Pierre Haller private collection (Switzerland).
CXG: Xavier Gouverneur private collection, Rennes (France).
EIHU: Collection of Systematic Entomology, Hokkaido University (Japan).
MCSNG: Museo Civico di Storia Naturale, Genoa (Italy).
MNHNL: Musée national d'histoire naturelle, Luxembourg (Luxembourg).
MNHNP: Muséum national d'histoire naturelle, Paris (France).
MW: U. S. National Museum of Natural History, Washington (USA).
SMF: Natur-Museum und Forschungs-Institut Senckenberg, Frankfurt am Main (Germany).
ZSM: Zoologische Staatssammlung München (Germany).
For each available species, the presence/absence (1/0) of the following 26 characters was exa-
mined: interantennal ridge, bifurcated inteantennal ridge, furrowed interantennal ridge, femo-
ral teeth, neck, straight/bowed intergenal furrow, pronotal lateral spines, pronotal discal tuber-
cles, strong pronotal wrinkles, regular pronotal wrinkles, pronotal longitudinal furrows,
pronotal smooth field, rugosity of the scape in male, rugosity of the scape in female, antennal
spines in male, antennal spines in female, antennomeres III-IV, and/or V, and/or VI spined, rap-
port elytra length/width, truncated elytral apex, toothed elytral apex, spined elytral apex, ely-
tral ridges, elytral pubescence.
The characters of all known species were summarised in a table (Tab. 1) and subject to sta-
tistical analyses of similarity. Some species of the genus Neocerambyx Thomson, 1860 were
added for a comparison. Hemadius oenochrous Fairmaire, 1889 was also tested. Cladistic parsi-
mony analyses with Euristic algorithms NN, SPN, TBR as well as multivariable cluster analyses
with Jaccard and Dice similarity coefficients were calculated with the paired group algorithm
using Past (Hammer et al., 2006) (Fig. 24–25).
Results
According to the keyed characters (Tab. 1), the following statements can be inferred:
Les Cahiers Magellanes, No26, juin 2017 — 42
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1. Aeolesthes Gahan, 1890
Original description. – Head with a central plaque in front, with a median, more or less dis-
tinct carina occupying interantennary solcus in front, and extending behind almost to a level
with the posterior border of the upper lobes of the eyes. At the termination of this carina the
vertex bear a shallow foveolate impression. Antennae in the male much longer than the body,
with the third to fifth joints thickened at the apex, with the joints from about the fifth to the
eighth usually furnished with a minute spine at their outer apical termination. The same joints
in the female more distinctly spined externally, and each also spinosely or denticulately produ-
ced at its inner apical termination. Prothorax strongly rugose above, rounded or subangulate
and unarmed at the sides in the middle. Elytra clothed with a rich silky pubescence giving
moiré reflexions; apices truncate, with the angles spinose or dentate. Anterior cotyloid cavities
very feebly or not at all angulate on the outside. Prosternal process usually subtruncate behind.
Type-species. – Hammaticherus aurifaber White, 1853 (Gahan’s des., 1906).
Diagnosis. – Body flattened, fairly stout. Head with an interantennal ridge completely sepa-
rated from the eyes by a furrow. Scape slightly convex externally, smooth dorsally; antennae
ectoapically toothed from the 5th or 6th article and endoapically spined from the 5th article; pro-
thorax as long as wide, with or without lateral spines, dorsally with more or less regular trans-
versal wrinkles. Pronotum regularly transversely wrinkled and with a pair of furrows on the disc
delimiting a rather smooth field; procoxal cavities rounded. Elytra almost parallel-sided in both
sexes, toothed at apex; elytral pubescence giving changing pattern. Femoral apex toothed.
Species. Aeolesthes aurifaber (White, 1853) (Fig. 1); Aeolesthes gloriosa (Aurivillius, 1924)
n. comb. (Fig. 2); Aeolesthes bilobulartus (Gressitt & Rondon, 1970) n. comb. (Fig. 3); Aeolesthes
vietnamensis n. sp. (Fig. 4).
Remarks. Aeolesthes aurifaber is the only species of the genus showing toothed apices of
meso- and metafemora. Just for such character, Gahan (1890) inserted it at the first step of the
key and as first species of the list. Later, he selected aurifaber unfortunately, the most pecu-
liar species of the genus as type-species of Aeolesthes (Gahan, 1906).
This unusual character is shared only by Trirachys gloriosus, which also shares antennae spi-
ned from the 5th article. Both species are very similar between them under other examined cha-
racters, except for the pronotal spines.
The toothed femora are a peculiar autoapomorphic character, while the smooth scape is a
primitive character. The contemporaneous presence of primitive and specialised characters
implies to consider Aeolesthes as a genus paraphyletic with respect to Trirachys, as all similarity
analyses show (Fig. 24–25).
The holotype of Trirachys bilobulartus (BPBM 8291) was originally mentioned as a female
coming from Pakkading, Borikhane Prov., 18.IV.1864. Actually, it is a male collected in 1964.
According to the description, the allotype is another female, but the figured specimen is ano-
ther male different from the holotype. Finally, Pakkading and Borikhane are both districts of
the Bolikhamsai Province. Makihara et al. (2008) recorded this species from Vietnam and
quoted Rhizophora apiculata Blume as its host, but the provided pictures prove that these data
must be referred to Trirachys sinensis (Gahan, 1890).
Les Cahiers Magellanes, No26, juin 2017 — 43
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Distribution. This genus has a Philippine-Malayan-Indochinese distribution. Aeolesthes
aurifaber was recorded from Laos by Brongniart (1891) and Gressitt & Rondon (1970), but
its real occurrence remains questionable. The specimen figured by Gressitt & Rondon (1970:
fig. 12i) does not evidently belong to this species, lacking of antennal and femoral spines.
Moreover, the specimen showed in fig. 13e and identified as Trirachys orientalis seems to be the
male of Aeolesthes vietnamensis n. sp.
Examined materials. – Aeolesthes aurifaber: 1
, Malaysia, Borneo, Sabah, Crocker Range, 19-
IV-1995, in CFV; 1
, 1
, Mt. Trus Madi, 1200 m, 7/11-IV-1994, N. Kanie lgt., in CFV; 1
, ditto,
2-V-1995, in CFV; 1
, ditto, 20-IV-2006, in CXG; 1
, 3
♀♀
, ditto, III/IV-2014, ex coll. Y. Mori-
mura, in CFV; 1
, Tawau, 8-V-1996, in CFV; Sarawak, 1865-66, coll. G. Doria, in MCSNG;
2
♂♂
, 1
, Indonesia, Borneo, Kalimantan Timur, Kalimantan, V-2001, in CGC; 1
, ditto, V-
2001, ex coll. W. Heinkel, in CFV, 1
, ditto, V-2004, coll. W. Heinkel, in CFV; 2
♀♀
, ditto, Balik-
papan, XI-2002, ex coll. W. Heinkel, in CFV; Sumatra, Siboga [= Sibolga], IV-1886, E. Modi-
gliani leg., in MCSNG.
Aeolesthes gloriosa: 1
, Philippines, Luzon, Aurora, Sierra Madre, V-2013, in CGC; 1
, 1
,
Leyte, Mt. Lobi, V-2010, coll. T. Richter, in CFV; 1
, Leyte, Mt. Balacaue, VIII-2011, I. Luma-
wig leg., in CFV; 1
, Mindanao, Surigao del Sur, VI-2012, I. Lumawig leg., in CFV.
Aeolesthes bilobulartus: Holotype
, Laos, Pakkading, Borikhane, 156 m, 18-IV-1964, Tri-
rachys bilobulartus Gressitt & Rondon, Holotype, in BPBM (8291).
Aeolesthes vietnamensis n. sp. (Fig. 4)
Examined materials. Holotype
, Vietnam, Nam Trung Bo, Quang Ngai, Mt. Bato, 950
m, IV-2014, in CFV.
Diagnosis. – Female, 47 mm. Body flattened, fairly stout. Head with an interantennal ridge
completely separated from the eyes by a furrow. Scape slightly convex externally, smooth dor-
sally; antennae ectoapically toothed from the 6th article and endoapically spined from the 5th
article; prothorax as long as wide, with lateral spines. Pronotum regularly transversely wrinkled
and with a pairs of furrows on the disc delimiting a nearly smooth field; procoxal cavities roun-
ded. Elytra almost parallel-sided, toothed at apex; elytral pubescence giving changing pattern.
Femoral apex toothed.
Differential diagnosis. The new species differs from the Philippine A. gloriosa (Fig. 2), in
the antennomere V lacking the external tooth, the large and posteriorly truncate interantennal
carina (elongated and posteriorly acute in A. gloriosa), the pronotal spines (obtusely tubercu-
late in A. vietnamensis, acute in A. gloriosa), the coarser pronotal sculpture, and the elytral
pubescence, forming more uniform larger pattern.
Moreover, it differs from the Bornean A. aurifaber (Fig. 1) in the larger size (less than 40
mm in A. aurifaber), the pronotal spines, the large and posteriorly truncate interantennal carina
(elongated and posteriorly acute in A. aurifaber), and the broader prothorax.
Finally, it differs from the Laotian A. bilobulartus (Fig. 3) in the pronotal spines, the large and
posteriorly truncate interantennal carina (elongated and posteriorly acute in A. bilobulartus) and
the nearly smooth pronotal field.
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Les Cahiers Magellanes, No26, juin 2017 — 45
1. Aeolesthes aurifaber (White, 1853): 1a,
; 1b,
(CFV). 2. Aeolesthes gloriosa (Aurivillius, 1924): 2a,
; 2b,
(CFV).
3. Aeolesthes bilobulartus (Gressitt & Rondon, 1970) Holotype
(BPBM, Photo J. Yamasako). 4. Aeolesthes vietnamensis
n. sp. Holotype
(CFV).
4
2a
3
1b
2b
1a
CM 26 V2_Mise en page 1 03/05/17 17:15 Page49
2. Trirachys Hope, 1843
Original description. Genus novum Hamatichero affine. Caput porrectum, fronte rugosa.
Antennae 11-articulatae, articulo 1mo crasso, valde rugoso; 2do minimo; 3tio, 4to, et 5to spinis
armatis; quinque sequentibus gradatim longioribus et inermibus, extimo longissimo ternis praece-
dentibus haud aequali. Thorax utrinque armatus rugisque transversis impressus. Elytra apicibus 2-
spinosis, marginibus elevatis. Pedes femoribus tibiisque compressis. Tarsi articulis cordiformibus,
duobus primis simplicibus, ultimo subbilobato auricomato.
Diagnosis. Body flattened, fairly stout (2.1–2.45 as long as wide). Head with an interan-
tennal ridge completely separated from the eyes by a furrow. Scape slightly convex externally,
mostly wrinkled dorsally; antennae ectoapically toothed and endoapically spined from the 3rd,
5th or even 6th article (almost one antennomere). Prothorax as long as wide, with or without
lateral spines, dorsally with irregular transversal wrinkles; procoxal cavities rounded. Elytra
almost parallel-sided in both sexes, angulated or toothed at apex; elytral pubescence giving
changing pattern. Femoral apex mutic.
Type-species. – Trirachys orientalis Hope, 1843 (monotypic).
Species. Trirachys acanthophorus Vitali, 1999 (Fig. 6); Trirachys achilles (Thomson, 1865)
n. comb. (Fig. 7); Trirachys ampliatus (Gahan, 1890) n. comb.; Trirachys atkinsoni Gardner,
1939; Trirachys basicornis (Gahan, 1893) n. comb.; Trirachys externus (Pascoe, 1869) n. comb.;
Trirachys holosericeus (Fabricius, 1787) n. comb.; Trirachys inexpectatus Holzschuh, 1982; Trirachys
indicolus (Bates, 1891) n. comb.; Trirachys indutus (Newman, 1842) n. comb. (Fig. 8); Trirachys
inhirsutus (Matsushita, 1932) n. comb. (Fig. 9); Trirachys orientalis Hope, 1843 (Fig. 5); Trirachys
perplexus (Gahan, 1890) n. comb.;
Trirachys sartus (Solsky, 1871) n. comb. (Fig. 10); Trirachys
sinensis (Gahan, 1890) n. comb.; Trirachys sphaericothorax Gressitt & Rondon, 1970; Trirachys
textor (Pascoe, 1869) n. comb.
Remarks. – Trirachys and Aeolesthes-species with spined antennae show no other character
that can differentiate them. Antennae are spined from the 3rd or 5th (or even 6th) article without
any relation with other characters. Actually, antennal spines are a fairly unstable character in
some species since large males (e.g. T. sartus) usually tend to lose them. The pronotal spines are
another instable character that can not be considered as discriminatory being variable inside
the same species, or even tending to disappear, such as in Trirachys formosana Schwarzer, 1925
(= T. orientalis). Consequently, all Aeolesthes-species with spined antennae and without spined
femurs are merged into the genus Trirachys.
The specific validity of T. externus and T. textor, as well the systematic position of T. inhirsutus,
was already treated in Vitali (2007).
Trirachys acanthophorus was originally mentioned as coming from "Panay, Mt. Tinagung,
Dagat Lake", as written in the label. Actually, the correct spelling of this locality is Tinagong
Dagat, a mountain lake located in Negros. The species is, however, present in Panay as well,
where the first male has been collected.
Examined materials. – Trirachys acanthophorus: Holotype
, Philippines, Negros, Tinagong
Dagat, V-1994, M. Mohagan leg., ex CFV, in ZSM; 1
, Negros oriental, IV-2013, I. Lumawig
leg., in CFV; 1
, Panay, Iloilo, Leon, V-2016, I. Lumawig leg., in CFV.
Les Cahiers Magellanes, No26, juin 2017 — 46
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Trirachys achilles: Holotype
, Borneo / ex Musaeo James Thomson / Achilles Thomson
Type S. C. ap. Born. / Th. Type / C. J. Gahan vidit 1890 / Museum Paris coll. J. Thomson Paris
1952 / Holotype, in MNHNP.
Trirachys externus: Holotype
, Irian Jaya, Dorey, in BMNH; Moluccas, Halmahera I., in
MNHNP, 1
, ditto, coll. G. Goussey, in CFV; 1
, ditto, XI-2010, in CGC; Little Kay [= Kai Kecil
I.], Kühn leg., in MNHNP; Key [= Kai Is.], 1873, O. Beccari leg., in MCSNG; Aru Is., C. Ober-
thür Coll., in MNHNP; ditto, Wokan [= Tanahbesar I.], 1873, O. Beccari leg., in MCSNG;
Moluccas, 1875, O. Beccari leg., in MCSNG; West Irian Jaya, Manokwari, Mansinam I., 1875,
Coll. Bruyn, in MCSNG; 1
, Papua, Jayapura, Mamberano River, Dabra, 30-III-2008, coll. C.
A. Casadio, in CFV; Papua New Guinea, Moroka [Mts.], 3500 ft., X-1895, Antony leg., in
MNHNP; Fly River, 1876/77, L. M. D’Albertis leg., in MCSNG; Kapakapa, VI/VII-1891, L. Loria
leg., in MCSNG; Upuli, X-1890, L. Loria leg., in MCSNG; Rigo, VII-1889, L. Loria leg., in
MCSNG; Ighibirei, VII/VIII.1890, L. Loria leg., in MCSNG; Yule I., VI-1875, L. M. D’Albertis
leg., in MCSNG; 1
, [Papua, Solomon] Bougainville, P. Hastert don. 1912, in MNHNL.
Trirachys holosericeus: 1
, India, Kerala, V-1994, A. Maier leg., in CFV; 1
, N[orth] Thai-
land, Chiang Rai, Doi Chang env., 640–750 m, 11/15-V-2013, O. Mosalov leg., in CFV; 2
♀♀
,
Vietnam, Da Nang, Nuichua, Ba Na, V-2013, in CXG; Malaysia: Sarawak, 1865-66, G. Doria
coll., in MCSNG; 1
, Indonesia, Moluccas, Buru, X-1875, O. Beccari leg., in MCSNG, Ceram,
Oberthür coll., in MNHNP, ditto, Wallace leg., in MNHNP.
Trirachys indutus: 1
, Sri Lanka, Southern, Tissamaharama, 17-XI-2010, in CXG; 1
, Viet-
nam, Quang Nam, IV-2014, N. Y. Son leg., in CFV; Taiwan, Takamuka leg., in EIHU; Rebun,
Uraru [= Heng-Chun, Wu-lai], 6-IV-1938, M. Matsushita leg., in EIHU; Koshun, IV-1908, H.
Sauter leg., in MCSNG; Ohtsubo [= Chia-i-hsien, Tai-ping], in EIHU; 1
, Malaysia, Pahang,
Cameron Highlands, V-1991, C. C. Hua leg., in CFV; Sabah, Banguey island, Standinger leg., in
MNHNP; Indonesia, Kariman Is., 1913, G. Gaggino leg., in MCSNG; I. Nias, 1897-98, U. Raap
lgt., in MCSNG; Mentawei, Si Oban, IV/VIII-1894, E. Modigliani leg., in MCSNG; Enggano,
Whitehead leg., in MNHNP; ditto, Bua-Bua, V/VI-1891, E. Modigliani leg., in MCSNG; ditto,
Kifa Jac., V-1891, E. Modigliani leg., in MCSNG; 1
, Java, coll. G. Goussey, in CFV; 1
, ditto,
1996, in CGC; ditto, Sinaron, in MNHNP; 1
, Sumatra, Lampung, Mt. Pesagi, VIII-2004, ex
coll. W. Heinkel, in CFV; Philippines, Luzon, 1894, Whitehead leg., in MNHNP; ditto, Montal-
ban, 1914, G. Boettcher leg., in MNHNL; ditto, Manila, coll. Pic, in MNHNP; 1
, Laguna, Los
Baños, 2015, I. Lumawig leg., in CFV; 1
, ditto, Cagayan, S.ta Ana, IX-2014, I. Lumawig lgt., in
CFV; Samar, VI/VII-1896, Whitehead leg., in MNHNP; South Palawan, coll. Oberthür, in
MNHNP; Palawan, Standinger leg., in MNHNP; ditto, Puerto Princesa, Paragua, coll. Oberthür,
in MNHNP; 1
, 1
, Nord Palawan, 1997, in CXG; Mindanao, coll. Oberthür, in MNHNP;
ditto, Dumalon, Zimboanga, 1876, L. Laglaize leg., in MNHNP; 1
, Mindanao, Bukidnon, V-
2014, I. Lumawig leg., in CGC; 1
, ditto, Kabanglasan, IX-2014, I. Lumawig leg., in CFV; 1
,
Sibuyan, IV-1982, in CFV.
Trirachys inhirsutus: Holotype
, Caroline Is, Palau I., Korol, 24-VI-1924, S. Uchiyama leg.,
ex coll. M. Matsushita, in EIHU; Caroline Is., Palau I., Korol, 3-XII-1922, S. Uchiyama leg. in
EIHU; 1
, Caroline Is., Palau I., Korol, 15-I-1994, C. A. Casadio leg., in CFV; 1
, Caroline Is.,
Palau I., Peleliu, Hoplocerambyx brevispinis Holotype Gressitt, 1951, in MW.
Trirachys orientalis: 1
, China, Shanghai, Mt. Tianma, 2-VII-2011, in CGC; 1
, Zhejiang,
Mt. Tianmushan, 2-VII-2012, in CXG; 1
, ditto, 3-VII-2012, in CXG; 1
, 1
, ditto, Henan, Mt.
Baotianman, VI-2016, L. X. Chong, in CFV; 1
, 1
, Taiwan, Nantou, 29-VI-2005, coll. D. Duda,
in CFV; 1
, ditto, 1-VII-2006, coll D. Duda, in CFV.
Les Cahiers Magellanes, No26, juin 2017 — 47
CM 26 V2_Mise en page 1 03/05/17 17:15 Page51
Trirachys sartus: 1
, Iran, Ljutfabad, in CFV; 1
, 1
, Afghanistan, Kaboul, Darulaman, 17-
V-2011, in CGC; 2
♂♂
, 1
, ditto, VI-2011, in CXG; 1
, 1
, Turkmenistan, Askhabad, 9-IV-
1996, Matlenski leg., in CFV; 6
♂♂
, Kara Kum, Anau, 25-IV-1988, M. Kafka leg., in CFV; Uzbe-
kistan, Buchara, Repetek, 1923, in MNHNL; Buchara, 3/5-V-1977, J. Lorenc leg., in MCSNG;
1
, 1
, ditto, 30-IV-1997, in CFV.
Trirachys sinensis: 1
, China, Shaanxi, Lueang, 18/24-VI.1997, E. Kucera leg., in CFV; 1
,
Yunnan, Xinping, Mt. Ailaoshan, 1900 m, VI-2015, L. X. Chong leg., in CFV; Birma, Carin,
Asciuii Chebà (Kareen Hills), 1200-1300 m, XII-1887, L. Fea leg., in MCSNG; ditto, 900-1100 m,
1889, L. Fea leg., in MCSNG; M. Cariani [= Kareen Hills], 1898, D. Tornatore leg., in MCSNG;
Catcin Cauri, VIII/IX-1886, L. Fea leg., in MCSNG; 1
, N[orth] Thailand, Chiang Rai, Doi Chang
env., 640-750 m, 11/15-V-2013, O. Mosalov leg., in CFV; 1
, Mt. Doi Inthanon, 2/3-V-1989, H.
Hirasawa leg., in CFV; 1
, 1
, Umphang. XI-1993, in CGC; 1
, 1
, Laos, Houa Phan, Ban Saleui,
1-V-2012, X. Gouverneur leg., in CFV; 1
, 1
, Laos, Houaphan, Mt. Phu Phan, 1-VI-2011, loc.
collector, in CXG; 1
, 1
, ditto, 1-V-2012, in CXG; 1
, Luang Prabang, Nong Khiaw, 22-IV-2015,
X. Gouverneur leg., in CXG; 1
, Taiwan, Tainan, 21-III-1909, S. Matsumura leg., in EIHU.
Trirachys sphaericothorax: Holotype, Laos, Houa Khong, Houay Say, 1200 m, 1.VI.1965
Trirachys sphaericothorax Gressitt & Rondon, J. A. Rondon coll., in BPBM (8292).
Trirachys textor: Holotype, Moluccas, Ternate I., in MNHNP; ditto, C. Oberthür Coll., in
MNHNP; ditto, Ançoy Coll., in MNHNP; ditto, 1861, E. Deyrolle Coll., in MNHNP; ditto, 1875,
A. A. Bruijn Coll., in MCSNG; 1
, ditto, in CFV; ditto, 1877, in MNHNP; ditto, 1931, A. A.
Argod-Vallon Coll., in MNHNP; ditto, 1903, J. Waterstradt leg., in MNHNP;
, ditto, II-2013,
coll. G. Aji, in CFV; ditto, La Glaize, in MNHNP; Gilolo [= Halmahera, 1858], A. R. Wallace
Coll., in MNHNP; Batchian [= Bacan] 1902, J. Waterstradt leg., in MNHNP; Obi Major [= Obi,
Obira], 1902, J. Waterstradt leg., in MNHNP; 1
, Kelang, IX-2010, in CGC; Moluccas [without
further indications], 1878, A. Raffray & R. Maindron leg., in MNHNP. New Guinea, West Irian
Jaya, Ramoi [= Sorong], II-1875, O. Beccari leg., in MCSNG; 2
♂♂
, Schouten Is., Biak I., Korem,
3-XII-1993, C. A. Casadio leg., in CFV.
3. Dymasius sticheri (Hüdepohl, 1989) n. comb. (Fig. 11)
= Aeolesthes sticheri Hüdepohl, 1989.
Remarks. – Due to the elongated prothorax and the mutic antennae, this species is transfer-
red to the genus Dymasius (s. str.) Thomson, 1864.
Examined materials. holotype
, Borneo, Sabah, Mt. Rinagsian, IV-1986, in ZSM; PARA-
TYPES: ditto, IV-1986, in ZSM; ditto, VI-1986 (ZSM); ditto, VII-1986, in ZSM; ditto, Kimanis
Road, 7th mile, VI-1986, in ZSM; ditto, 18th mile, IV-1986, K. E. Hüdepohl coll., in ZSM; 1
,
Kalimantan, Balikpapan, IX-2007, ex. coll. W. Heinkel, in CFV.
4. Dymasius fulgens (Schwarzer, 1926) n. comb. (Fig. 12)
Aeolesthes fulgens Schwarzer, 1926: 7 (Philippines, Mindanao: Surigao, SMF)
Remarks. – Aeolesthes fulgens is characterised by mutic antennae, interantennal carina posteriorly
bifurcated, pronotum evidently longer than wide and strongly convergent elytra. This set of charac-
ters fits the genus Dymasius Thomson, 1864, to which the species is consequently transferred.
Les Cahiers Magellanes, No26, juin 2017 — 48
CM 26 V2_Mise en page 1 03/05/17 17:15 Page52
Actually, Dymasius fulgens (Schwarzer, 1926) n. comb. is closely related to the Bornean
D. cuneatulus Holszchuh 2005, which might be a subspecies or even a synonym of the Philip-
pine species.
Examined material. Holotype
, Philippinen, Mindanao, Surigao, Aeolesthes fulgens
Holotypus, coll. B. Schwarzer, in SMF.
5. Pseudaeolesthes Plavilstshikov, 1931 rest. gen.
Original description Generi Aeolesthes Gah. simillimus, sed pronoto lateraliter spinoso.
Antennis corpore longioribus (
) vel parum brevioribus (
); articulo ceteris longiore, articulo
quarto longiore et fere aequali; articulis 3°-5° vel apice incrassatis sed non dentatis. Pro-
noto trasverso, vel subtrasverso, antice fortiter angustato ante basim et post apicem constricto et
satis profunde transverse sulcato; lateribus rotundato-dilatato, dente spiniformi, satis longe pro-
ducto acuteque armato; disco irregulariter fortiter ruguloso. Elytris satis longis, apice truncatis
angulo suturali in spinam longam producto.
Type-species. – Neocerambyx chrysothrix Bates, 1873.
Diagnosis. – Body convex, elongated. Head with an interantennal ridge posteriorly bifurcate
and delimiting a triangular interocular space. Scape slightly concave externally, smooth dor-
sally; antennae endoapically mutic and ectoapically toothed from the 5th or 6tharticle. Protho-
rax as long as wide, often toothed at the sides, dorsally with strong longitudinal wrinkles and
6 raised tubercles forming a hexagon; prosternal intercoxal process not tuberculate; procoxal
cavities rounded. Elytra considerably elongated, apically narrowed in males, parallel-sided in
females; elytral pubescence giving changing pattern, more or less condensed along longitudinal
stripes. Femoral apex mutic.
Species and subspecies. – Pseudaeolesthes aureopilosa (Gressitt & Rondon, 1970) n. comb.
(Fig. 15); Pseudaeolesthes chrysothrix (Bates, 1873) rest. comb. (Fig. 13); Pseudaeolesthes
chrysothrix taiwanensis (Hayashi, 1974) n. comb.; Pseudaeolesthes chrysothrix nakamurai
(Kusama & Takakuwa, 1984) n. comb.; Pseudaeolesthes chrysothrix kurosawai Gressitt, 1965
rest. comb.; Pseudaeolesthes chrysothrix yonaguniensis Ohbayashi & Ohbayashi, 1965 rest. comb.;
Pseudaeolesthes chrysothrix tibetana Gressitt, 1942 rest. comb.; Pseudaeolesthes malayana
(Hayashi, 1979) n. comb.; Pseudaeolesthes multistriata (Hayashi, 1979) n. comb.; Pseudaeolesthes
mutabiliaurea Chiang, 1951 rest. comb.; Pseudaeolesthes rufimembris (Pic, 1923) rest. comb.
(Fig. 14).
Remarks. – Pseudaeolesthes appears as a split group using all statistical methods (Fig. 24–25).
In order to separate this taxon (downgraded to subgenus of Aeolesthes), Gressitt & Ron-
don (1970) keyed the shape of the prosternal process at apex, “somewhat projecting”
(Pseudaeolesthes), or “horizontal, more or less vertical” (Aeolesthes). The body shape, not com-
pletely fitting the previous schema, was mentioned as an additional character. Nonetheless, the
prosternal shape is ambiguous and completely illusory character.
Les Cahiers Magellanes, No26, juin 2017 — 49
CM 26 V2_Mise en page 1 03/05/17 17:15 Page53
Actually, Pseudaeolesthes is well characterised by mutic antennae, scape slightly concave,
externally smooth dorsally, elytra elongated (2.45–3 as long as wide) and apically narrowed in
males. Moreover, the interantennal ridge is posteriorly bifurcate, connecting the upper eyes
lobes and delimiting a triangular interocular space, while it is a simple carina in Aeolesthes and
Trirachys. All these characters are also present in Aeolesthes psednothrix Gressitt & Rondon,
1970, which must be transferred to Pseudaeolesthes.
This clade includes more primitive species showing prothoracic spines, irregularly wrin-
kled pronotum, mutic elytral apex and pubescence forming changing pattern (chrysothrix,
rufimembris), and more specialised ones showing mutic prothorax, elytral spines, and shorter
antennae (mutabiliaurea, aureopilosa). Some characters are also shared by Aeolesthes or Trirachys;
however, elytral shape and interantennal ridge remain constant, while the pronotal hexagonal
granulation is peculiar of this genus.
According to the description, Pseudaeolesthes mutabiliaurea might be the senior synonym of
Pseudaeolesthes aureopilosa. The species was described on a female (Chiang, 1951), but the
author was possibly confused by the short antennae of this species.
Examined materials. Pseudaeolesthes aureopilosa: Holotype
, Laos, Xieng Khouang,
Plaine des Jarres, 1000 m, 28-III-1964, Aeolesthes (Pseudaeolesthes) aureopilosa Gressitt & Ron-
don, J. A. Rondon coll., in BPBM (8287); 1
, ditto, Houaphan, Mt. Phu Phan, III-2016, loc. coll.,
in CGC; 1
, ditto IV. 2016, in CGC; 1
, Vietnam, Mt. Fan-si-Pan, Cha-pa, 2400 m, 8/29-V-1993,
Sinjaev & Simonov leg., ex coll. A. Schintlmeister, in CFV; 1
, ditto, Lam Dong, IV-2013, L. X.
Xun leg., in CFV; 1
, ditto, Thua Thien Hue, Mt. Bach Ma, 1400 m, III-2016, in CGC.
Pseudaeolesthes c. chrysothrix: 1
, Japan, Honshu, Saitama, Iida, Ogawa-Maki, 16-VII-1987,
Izumiyama leg., in CFV; 1
, Niigata, Toyosoka-shi, Takamori-no-oka, 19-VII-1996, in CFV; 1
,
Osaka, Baba Ibaragi, 17-VI-1982, T. Ochi leg., in CFV; 1
, ditto, 10-VI-1983, in CFV; 1
, Iba-
rachi, Mito, Mt. Mito-yama, 2-VII-1991, M. Hasegawa leg., in CFV; 1
, Kanagawa, Yokohama,
Toshuka, 16-VII-1985, O. Furuta leg., in CFV; 1
, Kyushu, Nagasaki, Higashisonagi-machi, 15-
VII-1989, M. Furukawa leg., in CFV; 1
, Kagoshima, Yamura, 14-VII-1999, M. Furukawa leg.,
in CFV; 2
♂♂
, Tsushima I., Izuhara, Mt. Taitera-san, 21/25-VII-1991, M. Hasegawa leg., in CFV;
1
, Kōchi, Kōchi, Kaganoi. 14-VI-2013, in CGC.
Pseudaeolesthes c. nakamurai: 1
, 1
, Japan, Okinawa, Ishigaki-Jima I., Mt. Omoto-dake,
13-II-2015, T. Nakata leg., in CFV
Pseudaeolesthes c. taiwanensis: 1
, 1
, Taiwan, Nan-To, 7-III-2007, in CGC.
Pseudaeolesthes c. tibetana: Holotype
, Tibet Prov. / China / F. 4722 /Pseudaeolesthes
chrysothrix tibetanus Holotype, in AMNH; 1
, 1
, China, Guangxi, Dayao-Shan, Jinxiu, 100 km
SE Liuzhou, 1200 m, IV-2005, V. Siniaev’s team leg., in CFV.
Pseudaeolesthes psednothrix: Holotype
Laos, Phou Khao Khoay, 1040 m, N of Vientiane,
9-IV-1964, Aeolesthes (Pseudaeolesthes) psednothrix Gressitt & Rondon, J. A. Rondon coll., in
BPBM (8290).
Pseudaeolesthes rufimembris: Holotype
/ Tonkin, Mt. Manson IV-V, 2-3000 (ft), H.
Frühstorfer / Pseudaeolesthes Plav. [handwritten by M. Pic] / Aolesthes rufimembris Pic [handw-
ritten by M. Pic] / type [handwritten by M. Pic] / Museum Paris, coll. M. Pic, in MNHNP; 1
,
Vietnam, Vinh Phu, Tam Dao, loc. coll., in CFV; 1
, 1
, ditto, V-1995, in CGC; 1
, ditto, 15/25-
VI-1996, in CFV; 1
, ditto, 1000 m, 27-VII-2013, M. Pejcha lgt., in CFV; 2
♀♀
, Yenbai, Mucang
Chai, 1700 m, V-2016, T. Luong leg., in CFV; 1
, Laos, Houaphan, Ban Salaui, 1-V-2014, in
CGC; 1
, 2
♀♀
, Houaphan, Mt. Phu Phan, 1-V-2012, in CXG.
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Les Cahiers Magellanes, No26, juin 2017 — 51
5. Trirachys orientalis Hope, 1843: 5a,
; 5b;
(CFV). 6. Trirachys acanthophorus Vitali, 1999 Hol otype
(ZSM). 7.
Trirachys achilles (Thomson, 1865): 7a, Holotype
(MNHNP); 7b, labels of the holotype;. 8. Trirachys indutus (New-
man, 1842): 8a,
; 8b;
(CFV).
8a
7b
8b
7a
6
5b5a
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Les Cahiers Magellanes, No26, juin 2017 — 52
9. Trirachys inhirsutus (Matsushita, 1932): 9a, Holot ype
(EIHU); 9b,
(CFV); 9c, labels of the holotype. 10.
Trirachys sartus (Solsky, 1871): 10a,
; 10b,
(CFV). 11. Dymasius sticheri (Hüdepohl, 1989)
(CFV). 12. Dymasius
fulgens (Schwarzer, 1926) Holoty pe
(SMF, Photo A. Skale).
12
10b
9b
10a
9a
11
9c
CM 26 V2_Mise en page 1 03/05/17 17:15 Page56
6. Carinolesthes n. gen.
Type species. – Aeolesthes (Pseudaeolesthes) pericalles Gressitt & Rondon, 1970.
Diagnosis. – Body convex, elongated. Head with an interantennal ridge posteriorly bifurcate
but disappearing, delimiting a hardly guessable triangular interocular space. Scape slightly
concave externally, smooth dorsally; antennae endoapically mutic and ectoapically compressed
(
) or toothed (
) from the 7th article, antennomeres III to V or VI strongly inflated and
pubescent, especially in females. Prothorax as long as wide, with or without lateral spines, dor-
sally transversely wrinkled; prosternal intercoxal process not tuberculate; procoxal cavities
rounded. Elytra elongated, apically narrowed in males, parallel-sided in females, with longitu-
dinal smooth ridges on the disc; elytral pubescence giving changing pattern condensed along
longitudinal stripes. Femoral apex mutic.
Etymology. – From “carina” and Aeolesthes (gender feminine).
Species. – Carinolesthes aureosignata (Pic, 1915) n. comb. (Fig. 17); Carinolesthes ningshanensis
(Chiang, 1981) n. comb.; Carinolesthes pericalles (Gressitt & Rondon, 1970) n. comb. (Fig. 18)
Examined materials. – Carinolesthes pericalles: Holotype
and Allotype
, Laos, Xieng
Khouang, Plaine des Jarres, 1000 m, 28-III-1964, J. A. Rondon coll., Aeolesthes (Pseudaeolesthes)
pericalles Gressitt & Rondon, Holotype, Allotype, in BPBM (8286); 3
♂♂
, 3
♀♀
, Vietnam, Lào
Cai, Mt. Sapa, 1600 m, IV-2015, N. Son leg., in CFV; 1
, ditto IX-2015, in CFV; 1
, 1
, Laos,
Houaphan, Ban Saleui, 1-V-2014, loc. collector, in CGC; 1
, 1
, ditto, in CXG; 1
, ditto, in
CFV.
Carinolesthes aureosignata: Holotype
, Aeolesthes/ A. aureosignatus / Pic / type
[China] Ht Yunnan, Museum Paris / Yunnan / P. Guerry, 1924, Muséum Paris, Holotype
(in MNHNP).
Remarks. Carinolesthes n. gen. is related to Pseudaeolesthes, from which it differs in the
pronotal sculpture (with fine transverse rather than strong longitudinal wrinkles), the elytra
peculiarly sculptured by longitudinal smooth ridges and the inflated basal antennomeres.
The Vietnamese Carinolesthes aureosignata differs from C. pericalles, in primitive characters
linking it to Pseudaeolesthes: pronotum spined at sides, antennomeres III-V feebly inflated and
antennomere VI nearly normal. In contrast, the posteriorly bifurcate interantennal ridge (linear
in Pseudaeolesthes) and the ridged elytra belong patently to Carinolesthes n. gen.
According to the original descriptions (Chiang, 1981), the Chinese Aeolesthes ningsha-
nensis differs from aureosignata in the golden pubescence, the transverse pronotum and the
rounded elytral apices. The inflated antennomeres III-V and the ridged elytra belong to Cari-
nolesthes n. gen.
7. Elydnus chrysophanes (Gressitt & Rondon, 1970) n. comb. (Fig. 20)
= Aeolesthes (Pseudaeolesthes) chrysophanes Gressitt & Rondon, 1970: 64, Fig. 12g. (Cam-
bodia, Koh Rong Sanlem I., Saracens Bay, in BPBM).
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CM 26 V2_Mise en page 1 03/05/17 17:15 Page57
Examined material. Holotype
, Laos, Baie de Khompong Som, Iles des Saracems, 5-XII-
1963, at light, Aeolesthes (Pseudaeolesthes) chrysophanes Gressitt & Rondon, J. A. Rondon coll.,
in BPBM (8288).
Remarks. – This species is characterised by mutic antennae and interantennal furrow. The
pronotum shows a single median furrow, while the elytral pubescence is longitudinally striped
and obscured along the suture. All these characters belong to Elydnus Pascoe, 1869; actually, this
species is extremely similar to the type-species E. amictus Pascoe 1869.
Distribution. – The holotype is mentioned as collected in both Laos and Cambodia from the
“Iles des Saracems, Baie de Khompong Som” [sic!]. Actually, Saracens is a bay located on the
isle Koh Rong Sanlem, in front of the Kampong Som Bay, in Cambodia. Thus, the correct loca-
lity is: Cambodia, Koh Rong Sanlem I., Saracens Bay. The species does not belong to the Lao-
tian fauna, but to the Cambodian one.
8. Massirachys n. gen.
Type-species. – Pachydissus mariae Thomson, 1878 (monotypic).
Diagnosis. – Body size large. Head elongated, temples with feebly convergent sides, distinctly
longer than hind upper lobes of eyes and not distinguished from the neck; interantennal ridge
feebly furrowed. Scape slightly convex externally, smooth dorsally; antennae ectoapically too-
thed from the 6th article, endoapically mutic and pubescent at the inner side in males, endoa-
pically spined from the 5th article in females. Prothorax as long as wide, apically restricted,
mutic at sides, thinly and almost regularly transversely wrinkled on the disk; procoxal cavities
rounded. Elytra convex-sided, apex narrow, each armed with two minute spines, elytral pubes-
cence giving changing pattern. Femoral apex mutic.
Species. – Massirachys mariae (Thomson, 1878) n. comb. (Fig. 19)
Etymology. – From the genera Massicus and Trirachys (gender masculine).
Remarks. – The Bornean Aeolesthes mariae (Thomson, 1878) evolved some peculiar charac-
ters making its habitus analogous to that of the genus Massicus Pascoe, 1867: large body size,
elongated head and temples without a distinct neck, elytra more elongated and convex-sided,
elytral apex narrower than it occurs to other related genera. The interantennal ridge is less deve-
loped and feebly furrowed but always present. Males have antennomeres without endoapical
spines, bowed and inner pubescent, while females have spined antennae. Such peculiar charac-
ters justify the institution of a new genus.
Examined materials. Holotype
, Borneo / ex Musaeo James Thomson / Mariae Thom-
son Type T. C. 34 30 Borneo / Th[omson]. Type / C. J. Gahan vidit 1890 / Museum Paris coll.
J. Thomson Paris 1952 / Holotype, in MNHNP; 1
, Malaysia, Borneo, Sabah, Crocker Range,
25-I-1989, in CFV.
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Les Cahiers Magellanes, No26, juin 2017 — 55
13. Pseudaeolesthes chrysothrix (Bates, 1873): 13a,
; 13b,
(CFV). 14. Pseudaeolesthes rufimembris (Pic, 1923): 14a,
(CFV); 14b, Holoty pe
(MNHNP); 14c, labels of the holotype. 15. Pseudaeolesthes aureopilosa (Gressitt & Rondon,
1970): 15a,
(CFV); 15b Holotype
(BPBM, Photo J. Yamasako).
15b
15a
14b
14a
13b
14c
13a
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Les Cahiers Magellanes, No26, juin 2017 — 56
16. Pseudaeolesthes psednothrix (Gressitt & Rondon, 1970) Holotype
(BPBM, Photo J. Yamasako); 17. Carinolesthes aureo-
signata (Pic, 1915): 17a, Holotype
(MNHNP); 17b, labels of the holotype. 18. Carinolesthes pericalles (Gressitt & Rondon,
1970): 18a, Holotype
; 18b, Allotype
(BPBM, Photo J. Yamasako). 19. Massirachys mariae (Thomson, 1878): 19a,
Holotype
(MNHNP); 19b, labels of the holotype.
19b
19a
18b
17b
18a
17a
16
CM 26 V2_Mise en page 1 03/05/17 17:15 Page60
Les Cahiers Magellanes, No26, juin 2017 — 57
20. Elydnus chrysophanes (Gressitt & Rondon, 1970) Holotype
(BPBM, Photo J. Yamasako). 21. Parolesthes laosensis
(Gressitt & Rondon, 1970) Holotype
(BPBM, Photo J. Yamasako); 22. Pachydissus langsonius Fairmaire, 1895 incertae
sedis: 22a, claimed Holotype
(MNHNP); 22b, labels of the holotype. 23. Derolus uniformis (Pic, 1933): 23a, Holotype
(MNHNP); 23b, labels of the holotype; 23c,
(CFV).
20 21 22a
22b
23a
23b
23c
CM 26 V2_Mise en page 1 03/05/17 17:15 Page61
9. Parolesthes n. gen.
Type species. – Aeolesthes laosensis Gressitt & Rondon, 1970.
Description. – Body convex, flattened, fairly stout. Head with an interantennal ridge posteriorly
bifurcate and delimiting a triangular interocular space. Scape slightly convex externally, wrinkled
dorsally; antennae ectoapically toothed and endoapically mutic in both sexes. Prothorax mutic or
with a minuscule tubercle at sides, with two longitudinal furrows on the disc delimiting a squa-
red field; prosternal intercoxal process not tuberculate; procoxae rounded. Elytra parallel-sided in
both sexes, toothed at apex; elytral pubescence giving changing pattern. Femoral apex mutic.
Etymology. – From the ancient Greek “para” (near) and Aeolesthes (gender feminine).
Species. – Parolesthes curticornis (Hüdepohl, 1988) n. comb.; Parolesthes laosensis (Gressitt &
Rondon, 1970) n. comb. (Fig. 21).
Remarks. The Aeolesthes-species with mutic antennae form a small group that can be
inserted into none of the previous taxa due to the occurrence of opposite characters.
In fact, this clade shares with Pseudaeolesthes divided interantennal ridge and mutic anten-
nae and with Trirachys globular pronotum, dorsally flat elytra and wrinkled scape in males.
Hence, it seems to be as a link between such genera, but many specialised characters (elytral
spines, pronotum with longitudinal furrows and smooth area) make doubtful to consider it as
a primitive branch of Trirachys. In other words, the contemporaneous presence of primitive and
specialised characters implies to consider this taxon as a paraphyletic genus. However, we are
conscious that the genus is possibly polyphyletic and deserves further analyses.
Though mentioned as a female in the original description, the holotype of Aeolesthes
laosensis (BPBM 8289), as well the figured specimen, is a male (Fig. 21). It was also mentioned
as coming from Muong Khong, Sithandone Prov. [sic!], but the correct spelling is Muang
Khong (Khong distr., Champasak prov.). Siphandone is a surrounding locality corresponding
to the Khone Phapheng Falls.
Examined materials. – Parolesthes curticornis: Holotype
, Malaysia, Borneo, Sabah, Kima-
nis Road 18th mile, V-1986, K. E. Hüdepohl coll., Aeolesthes curticornis Holotypus (in ZSM).
Parolesthes laosensis: Holoty pe
, Laos, Siphandone, Muang Khong, 87 m, 27-XII-1965,
Aeolesthes laosensis Gressitt & Rondon, Holotype, in BPBM (8289); 2
♂♂
, 4
♀♀
, Vietnam,
Nam Trung Bo, Quang Ngai, Mt. Bato, 950 m, IV-2014, in CFV; 5
♀♀
, Vietnam, Tam Dao,
1000 m, VII-2010, in CFV; 1
, China, Yunnan, Jinghong, Mengwavy, V-2012, coll. P. Demez,
in CFV.
10. Pachydissus langsonius Fairmaire, 1895 incertae sedis (Fig. 22)
Pachydissus langsonius Fairmaire, 1895: 176 (Vietnam: Lang Son, in MNHNP).
Aeolesthes langsonius Aurivillius, 1912: 47; Plavilstshikov, 1931: 77.
Les Cahiers Magellanes, No26, juin 2017 — 58
CM 26 V2_Mise en page 1 03/05/17 17:15 Page62
Original description. – Oblongus sat convexus, subaenescenti- fuscus, brunneo et lutoso cinereo
nitide sericans et mutans; fronte bisulcata, medio carinula nigra nitida signata, oculis supra sat
approximatis, antennis 5 corpore longioribus, articulis intermedibus inermibus, 1° crasso, irregula-
riter rugoso plicato; prothorace transverso, lateribus rotundato, basi constricto, dorso grosse ac irre-
gulariter plicato, spatio medio fere laevi et sulcis 2 obliquo arcuatis antice approximatis limitato;
elytris minus latis, postice attenuatis, apice utrinque sinuatis, angulo suturali spinoso, dorso obso-
letissime longitudinaliter, impresso; prosterno antice fortiter transversim plicato, abdomine parcius
pubescente, medio denudato. Long. 27 mm.
Remarks. – The examination of the holotype Pachydissus langsonius Fairmaire, 1895 revea-
led that it does not differ substantially from Cerambyx holosericeus Fabricius, 1787. This “holo-
type” is strangely characterised by spined antennae (Fig. 22), whereas the author highlighted
articulis intermedibus inermibus, also insisting in the differential diagnosis: “les 5e, 6eet 7earti-
cles des antennes ne sont pas épineux à l’extrémité”.
However, though some specimens with mutic antennae identified by Fairmaire as
Pachydissus langsonius” are preserved in the IRSNB, the holotype does not match the descrip-
tion. In our opinion, the presumed “holotype” is a false and, waiting for further analyses, this
species is provisionally considered as incertae sedis.
Examined material. Holotype
, Vietnam, Langson, coll. L. Fairmaire, in MNHNP.
11. Mimoderolus Pic, 1933
Original description. – Grandis, parum elongatus, nitidus, griseo–holosericeo, in elytris unifor-
miter, pubescens, nigro–piceus, pedibus rufis; antennis, in mare, elytris valde longioribus, articulis
3–8 intus longe ciliatis, 6–8 diverse arcuatis, 9 inciliatis et corpore brevioribus; thorace parum elon-
gato, lateraliter sinuato, diverse et pro parte reducte plicato, ante medium supra minute bitubercu-
lato; elytris thorace latioribus, parum elongatis, postice attenuatis, apice truncatis et dentatis,
minutissime punctatis et uniformiter sericeo pubescentibus, lateraliter distincte carinulatis; femo-
ribus infra minute carinatis. Long. 30–32 mill. Tonkin.
On peut établir pour cette espèce un sous-genre nouveau (s.-g. Mimoderolus) qui sera carac-
térisé par la structure des antennes du
et le revêtement pubescent fin et uniforme des élytres.
Type species. – Aeolesthes (Mimoderolus) uniformis Pic, 1933 (Fig. 23)
Remarks. – Pic (1933) described Mimoderolus as a monotypic subgenus of Aeolesthes. The
taxon, 3 cm long, is characterised by inferiorly pubescent, mutic antennae and ridged ventral
side of the femora. The interantennal space is finely furrowed.
This set of characters does not evidently belong to Aeolesthes or closely related genera, but to
Derolus Gahan, 1891, for which the author did not provide any difference. The type also corres-
ponds to the Burmese Pachydissus xyliae Fisher, 1940, which Gressitt & Rondon (1970) trans-
ferred to the genus Derolus. Consequently, the following taxonomic changes are introduced:
Derolus Gahan, 1891 = Mimoderolus Pic, 1933, n. syn.
Derolus uniformis (Pic, 1933) n. comb. = Pachydissus xyliae Fisher, 1940, n. syn.
Les Cahiers Magellanes, No26, juin 2017 —59
CM 26 V2_Mise en page 1 03/05/17 17:15 Page63
Distribution. – The species is widespread in the mountains from Myanmar to Laos and Vietnam.
Examined materials. Holotype
[Vietnam], Tonkin, Hoo-Binh, coll. M. Pic, Museum
Paris, Aeolesthes sg. Mimoderolus uniformis n. sp. [handwritten by M. Pic], type [handwritten
by M. Pic], in MNHNP; 1
, 1
, Vietnam, Lang Dong, VI-2014, in CFV; 3
♀♀
, Vietnam, Lao
Cai, Mt. Sapa, 1600 m, IX-2015, N. Son leg., in CFV; 1
, Vietnam, Con Dao island, IV-2012, in
CPH; 1
, Laos, Luang Prabang, Kiew Mak Nao, VI-2016, loc. coll., in CXG; 1
, Laos, Mt. Phu
Phan, in CGC.
12. Hemadius oenochrous Fairmaire, 1889
Remarks. – On the basis of a superficial likeness, Nakamura et al. (1992) transferred Hemadius
oenochrous Fairmaire, 1889 to Aeolesthes. Though Yu et al. (2002) re-transferred it to Neocerambyx,
Hua (2002) and Hua et al. (2009) keep consider this species as a member of the genus Aeolesthes.
Löbl & Smetana (2010) revalidated again this species in the genus Hemadius, though misspel-
ling it as Hemadius oenochroa (sic!).
This species has no interantennal ridge but a simple furrow, a fact that excludes any belon-
ging to Aeolesthes. Moreover, it shares with Neocerambyx perfectly mutic antennae, inflated
basal antennomeres, and large body size. Nonetheless, the typical deep interantennal fovea of
Neocerambyx is nearly lacking and the species shows a peculiar pronotal tooth at each side of
the prothorax.
For these reasons, the restoration of Hemadius Fairmaire, 1889 and of its original combina-
tion Hemadius oenochrous Fairmaire, 1889 are deemed as correct.
Examined materials. 1
Laos, Houaphan, Ban Saleui, 1-V-2014, in CGC; 2
♀♀
, 1-V-2014,
Houaphanh, Mt. Phu Phan, loc. coll., in CXG; 1
, 1
, Taiwan, Nan-To, V-2007, in CGC; 1
,
1
, ditto, Liu-she, VIII-2007, ex. coll. W. Duda, in CFV; 1
, 1
, ditto, Puli, IV-2014, D. Wang,
in CFV; 4
♂♂
, 3
♀♀
, ditto, Ren’ai Wu-she, V-2015, in CFV.
The taxonomic status of several Oriental Cerambycini needs further revisions, but a key is
hereafter proposed in order to facility the identification of the genera previously treated.
1. Body size larger (66–70 mm); temples distinctly longer than the hind upper lobes of eyes;
antennae endoapically mutic in male, spined in female .................................. Massirachys n. gen.
-. Body size smaller (less than 54 mm); temples distinctly shorter than the hind upper lobes
of eyes; antennae mutic or spined in both sexes ........................................................................... 2
2. Prothorax as long as broad or transverse .............................................................................. 3
-. Prothorax evidently longer than broad .................................................................................. 8
3. Head with a posteriorly bifurcate interantennal ridge; antennae endoapically smooth ... 4
-. Head with a simple interantennal ridge; antennae endoapically spined ............................ 7
4. Body hairless ............................................................................. Pseudopachydissus Pic, 1933
Les Cahiers Magellanes, No26, juin 2017 — 60
CM 26 V2_Mise en page 1 03/05/17 17:15 Page64
-. Body covered with a dense pubescence ................................................................................. 5
5. Elytra with smooth longitudinal ridges; antennae ectoapically toothed from the 7th arti-
cle; antennomeres III to V or VI strongly inflated and pubescent ................ Carinolesthes n. gen.
-. Elytra without smooth longitudinal ridges; antennae ectoapically toothed from the 5th or
6th article; antennomeres III-VI normal ........................................................................................ 6
6. Body dorsally flat; pronotum globular, with irregular transversal wrinkles and two longi-
tudinal furrows at base; scape convex above, wrinkled in male ....................... Parolesthes n. gen.
-. Body dorsally convex; pronotum elongated, without transversal wrinkles and longitudi-
nal basal furrows, with 6 raised tubercles forming a hexagon; scape slightly concave above,
smooth in male ..................................................................... Pseudaeolesthes Plavilstshikov, 1931
7. Meso- and metafemora toothed at apex; scape smooth ................. Aeolesthes Gahan, 1890
-. Meso- and metafemora mutic at apex; scape mostly wrinkled .......... Trirachys Hope, 1843
8. Apex of the antennomeres IV-V with pores; elytral pubescence more or less condensed
forming longitudinal bands ........................................................................... Elydnus Pascoe, 1869
-. Apex of the antennomeres IV-V without pores .......................... Dymasius Thomson, 1864
Le statut taxonomique de plusieurs Cerambycini orientaux nécessite d'autres révisions, mais une
clé est proposée ci-dessous afin de faciliter l'identification des genres précédemment traités.
1. Corps de grande taille (66–70 mm); tempes distinctement plus longues que les lobes
supérieurs des yeux; face endoapicale des antennes inerme chez le mâle, épineuse chez la
femelle) ............................................................................................................... Massirachys n. gen.
-. Corps plus petit (moins de 54 mm); tempes distinctement plus courtes que les lobes supé-
rieurs des yeux; antennes inermes ou épineuses chez les deux sexes ........................................... 2
2. Pronotum aussi long que large, ou transverse ...................................................................... 3
-. Pronotum visiblement plus long que large ........................................................................... 8
3. Tête dotée d'une carène postérieurement dédoublée entre les antennes; face endoapicale
des antennes inerme ......................................................................................................................... 4
-. Tête avec une carène interantennaire simple ; face endoapicale des antennes épineuse ... 7
4. Corps glabre ............................................................................. Pseudopachydissus Pic, 1933
-. Corps couvert d'une dense pubescence ................................................................................. 5
5. Élytres avec des carènes longitudinales lisses ; face ectoapicale des antennes dentée à par-
tir du 7earticle; antennomères III à V ou VI fortement globuleux et pubescents ......................
.......................................................................................................................... Carinolesthes n. gen.
Les Cahiers Magellanes, No26, juin 2017 — 61
CM 26 V2_Mise en page 1 03/05/17 17:15 Page65
-. Élytres sans carènes longitudinales lisses; face ectoapicale des antennes dentée à partir du
5eou du 6eantennomère; antennomères III-VI normaux ........................................................... 6
6. Corps dorsalement aplati; pronotum globuleux, doté de crêtes transversales irrégulières
et de deux sillons longitudinaux à la base du disque ; scape dorsalement convexe, ridé chez le
mâle ...................................................................................................................... Parolesthes n. gen.
-. Corps dorsalement convexe, pronotum allongé, sans crêtes transversales et sillons longi-
tudinaux basaux, avec six tubercules élevés disposés en hexagone ; dessus du scape légèrement
concave, lisse chez le mâle .................................................... Pseudaeolesthes Plavilstshikov, 1931
7. Méso- et métafémurs à l'apex épineux ; scape lisse ......................... Aeolesthes Gahan, 1890
-. Méso- et métafémurs à l'apex inerme; surface dorsale du scape principalement ridée .........
.......................................................................................................................... Trirachys Hope, 1843
8. Apex des antennomères IV-V avec des pores; pubescence des élytres plus ou moins
condensée, formant des bandes longitudinales ............................................ Elydnus Pascoe, 1869
-. Apex des antennomères IV-V sans pores ..................................... Dymasius Thomson, 1864
Acknowledgements
Many thanks to the Hokkaido University Systematic Entomology, Faculty of Agriculture,
Sapporo (Japan), for the pictures of the type of Aeolesthes inhirsuta Matshushita, 1932, to Jun-
suke Yamasako, Ehime University, Tarumi, Matsuyama (Japan), for sharing the pictures of the
types belonging to the Bernice Pauahi Bishop Museum, Honolulu (USA), to Andre Skale, Hof
an der Saale (Germany), for sharing on the forum www.cerambycoidea.com the picture of the
type of Dymasius fulgens Schwarzer, 1926, to Thierry Deuve and Azadeh Taghavian, Muséum
national d'histoire naturelle, Paris (France), for their warm welcome and their high availability,
to Kiyoshi Matsuda, Takarazuka (Japan) and Michiaki Hasegawa, Toyohashi Museum of Natu-
ral History (Japan), for sending of material, to Shulin Yang, Guizhou Normal University
(China) for sending Chiang’s (1951) paper, to Koji Mizota, Miyagi University of Education
(Japan), for sending many Japanese papers, to Ira and Ernestina Repetto, Genoa (Italy) for the
translation of Chiang’s (1981) description.
References
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Schenkling S. ed., Berlin: 574 pp.
Aurivillius (C.), 1924. – Neue oder wenig bekannte Coleoptera Longicornia. 19. Arkiv för
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Chinese Longicorn Beetles (1406 species) in Color. Sun Yat-sen University Press, Guangzhou: 474
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loidea. Apollo Books, Stenstrup: 924 pp.
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Les Cahiers Magellanes, No26, juin 2017 — 63
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Les Cahiers Magellanes, No26, juin 2017 — 64
acanthophorus
indutus
achilles
basicornis
pericalles
aureosignata
ningshanensis
rufimembris
chrysothryx
aureopilosa
catherinae
raddei
paris
oenochrous
curticornis
laos ens is
mariae
bilobulartus
vietnamensis
gloriosa
aurifaber
orientalis
inhirsutus
indico lus
sartus
sinensis
ampliatus
textor
externus
holosericeus
0,2
0,3
0,4
0,5
0,6
0,7
0,8
0,9
1,0
oenochr ous
paris
catherinae
raddei
mariae
aurifaber
bilobulartus
glor iosa
vietnamensis
externus
textor
ampliatus
holosericeus
indu tus
acanthophorus
basicornis
sinensis
inhirsutus
orientalis
achilles
indic olus
sartus
curticornis
laos ens is
aureopilosa
chrysothryx
rufimembris
aureosignata
nings han ens is
pericalles
25. Cladistic parsimony analyses with Euristic algorithm TBR.
24. Diagram of the distances based on Jaccard’s similarity coefficients.
CM 26 V2_Mise en page 1 03/05/17 17:15 Page68
Les Cahiers Magellanes, No26, juin 2017 — 65
Tab. 1. Keyed characters:
Body large
femurs toothed
H interantennal ridge
PT long furrows
PT smooth field
A scape
wrinkled
M
A scape
wrinkled
F
A spined in M
A spined in F
A 3
-
4 spined
A 5 spined
A 6 spined
A globose
acanthophorus 1
0
1
0 0 1 1 0 0 0 0 1 1
1
1
1
1 1
1
1
0
1 1 0 1 0
achilles 1
0
1
0 1 1 0 0 0 1 0 1 1
1
1
1
1 0
1
1
0
1 1 0 1 0
ampliatus 1
0
1
0 0 1 0 0 0 0 0 1 1
1
1
1
1 0
1
1
0
1 1 1 1 0
aureopilosa 0
0
1
1 1 1 0 0 1 0 1 1 0
0
0
0
0 0
0
0
0
1 1 1 1 0
aureosignata 0
0
1
1 0 1 0 1 0 1 0 0 0
0
0
0
0 0
0
0
1
1 0 0 1 1
aurifaber 1
1
1
0 0 1 0 0 0 1 0 1 0
0
0
1
1 0
1
1
0
1 1 0 1 0
basicornis 1
0
1
0 0 1 0 0 0 0 0 1 0
1
1
1
1 0
1
1
0
1 1 0 1 0
bilobulartus 1
1
1
0 0 1 0 0 0 1 0 1 0
0
0
1
1 0
1
1
0
1 1 0 0 0
chrysothryx 0
0
1
1 1 1 0 1 1 0 1 1 0
0
0
0
0 0
0
0
0
1 0 0 1 0
curticornis 1
0
1
0 1 1 1 0 0 0 0 1 0
0
0
0
0 0
0
0
0
1 1 0 1 0
externus 1
0
1
0 0 1 1 0 0 0 0 1 1
1
1
1
1 0
0
1
0
1 1 1 1 0
gloriosa 1
1
1
0 0 1 0 1 0 1 0 1 1
0
0
1
1 0
1
1
0
1 1 0 1 0
holosericeus 1
0
1
0 0 1 0 0 0 0 0 1 1
1
1
1
1 0
1
1
0
1 1 0 1 0
indicolus 1
0
1
0 0 1 0 0 0 0 0 1 0
1
1
1
1 0
0
1
0
1 0 0 1 0
indutus 1
0
1
0 0 1 1 0 0 0 0 1 1
1
1
1
1 0
1
1
0
1 1 0 1 0
inhirsutus 1
0
1
0 0 1 0 0 0 0 0 1 0
1
1
1
1 0
1
1
0
1 1 0 0 0
laosensis 1
0
1
1 1 1 0 0 0 0 0 1 1
1
0
0
0 0
0
0
0
1 1 0 1 0
mariae 1
0
1
0 1 0 0 0 0 1 0 0 0
0
0
0
1 0
1
1
0
1 1 0 1 0
ningshanensis 0
0
1
1 0 1 0 1 0 0 0 0 0
0
0
0
0 0
0
0
1
0 0 0 1 1
orientalis 1
0
1
0 0 1 0 1 0 0 0 1 0
1
1
1
1 1
1
1
0
1 1 0 1 0
pericalles 0
0
1
1 1 1 0 0 0 0 0 0 0
0
0
0
0 0
0
0
1
1 0 0 1 1
rufimembris 0
0
1
1 1 1 0 1 1 0 1 1 0
0
0
0
0 0
0
0
0
1 0 0 1 0
sartus 1
0
1
0 0 1 0 0 0 0 0 1 0
1
1
0
1 0
0
1
0
1 0 0 1 0
sinensis 1
0
1
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CM 26 V2_Mise en page 1 03/05/17 17:15 Page69
... We examined most of the literature citing Aurivillius's "Neue oder wenig bekannte Coleoptera Longicornia" published in the journal "Arkiv för zoologi", and gathered the results herein. a) Citation using the journal's page numbers: Aurivillius 1912Aurivillius , 1922Aurivillius , 1923Aurivillius , 1928Breuning 1939Breuning , 1940Breuning , 1944Breuning , 1950Breuning , 1956Breuning , 1958Breuning -1969Quentin 1956;Podaný 1968Podaný , 1971Gressitt and Rondon 1970;Rondon and Breuning 1970;Hüdepohl 1985Hüdepohl , 1988Hüdepohl , 1989Hüdepohl , 1990Lee 1987;Niisato 1989Niisato , 2007Nakamura et al. 1992;Napp 1993;Nýlander 1998;Makihara 1999;Heffern 2002;Makihara and Woro 2002;Morati and Huet 2004;Ohbayashi and Niisato 2007;Bousquet et al. 2009;Morati and Bentanachs 2009;Bentanachs et al. 2010;Sudre et al. 2010;Juhel 2011;Jiroux 2011;Vitali 2011;Weigel and Skale 2011;Vitali and Vitali 2011;Wallin et al. 2014;Lin 2015;Vitali et al. 2017. b) Citation using both Aurivillius's and the journal's page numbers, but considering journal numbers as more important: Löbl and Smetana 2010;Heffern 2011;Viktora 2013Viktora , 2015aViktora , 2019Viktora and Tichý 2017; Yang 2019. ...
... ;Lingafelter and Hoebeke 2002;Hüdepohl and Heffern 2004;Heffern 2005;Makihara and Woro 2002;Monné 2005a Monné , b, 2012 2019a, b, d;Monné and Napp 2005;Yokio and Niisato 2009;Vives 2011;Monné et al. 2016;Monné and Monné 2017;Bezark 2019).However, since Zoological Record cited 1924 for this work, so do the Titan database (Tavakilian and Chevillotte 2019). Less than half of authors used 1924 as the publication date(Lee 1987;Hüdepohl 1990;Heffern 2011;Lingafelter et al. 2014;Nakamura et al. 2014;Viktora and Tichý 2016;Vitali et al. 2017;Lin and Yang 2019). The first author of this paper Mei-Ying Lin was confused in this case (chose 1924 in the catalogue byLin and Yang (2019) and an earlier version of this paper, but finally decided to choose the earlier date of "Tryckt den 31 december 1923"). ...
Article
Full-text available
Aurivillius’s work entitled “Neue oder wenig bekannte ColeopteraLongicornia” was published in parts over a period of over four decades. There were two page numbers on most pages of these publications, one ordered by Aurivillius, the other by the journal. Historically, different authors have used different page numbers, and sometimes different years for these publications, which has caused chaos in the citations. Herein, accurate dates of publications for this work, and correct page numbers that should be used are provided and discussed.
... Notes on the type locality. Vitali et al. [2017] have corrected the type locality of S. chrysophanes comb. n. which was confused in its original description [Gressitt, Rondon, 1970], proposing the following: Cambodia, Koh Rong Sanloem Island, Saracen Bay. ...
... Through the courtesy of Dr. Andre Skale (Hof, Germany), I have received a good picture of the holotype male of this species, although such is also available in Vitali et al. [2017]. ...
Article
Full-text available
Taking into account the new data presented in this paper, the tribe Cerambycini in the fauna of Asia contains almost 60 genus-group taxa and about 335 species, being the largest compared to the faunas of the other parts of the world. Serious problems in the development of a reasonable supraspecific classification of the tribe are noted. Reviews of such taxonomically confused genera as Elydnus Pascoe, 1869, Imbrius Pascoe, 1866, Zatrephus Pascoe, 1857, and Zegriades Pascoe, 1869, as well as keys to their constituent species are given. The following new genera and species are described, two new statuses change and new combinations are established: Falsopachydissus gen. n., F. foveiscapus (Holzschuh, 2011), comb. n., Mimimbrius gen. n., M. dembickyi sp. n. (Southern Thailand), M. geminatus (Holzschuh, 2005), comb. n., M. micaceus (Pascoe, 1858), comb. n., M. subargenteus (Gressitt et Rondon, 1970), comb. n., Pascoetrephus gen. n., P. hefferni sp. n. (Eastern Malaysia), P. klimenkoi sp. n. (Eastern Malaysia), P. inscitus (Pascoe, 1857), comb. n., P. ranongensis (Holzschuh, 2009), stat. n. et comb. n., Spinidymasius gen. n., S. chrysophanes (Gressitt et Rondon, 1970), comb. n., S. crinicornis (Hüdepohl, 1989), comb. n., S. dembickyi (Holzschuh, 2003), comb. n., S. grossescapus (Hüdepohl, 1989), comb. n., S. huedepohli (Vives, 2005), comb. n., S. ochraceovittatus (Hüdepohl, 1989), comb. n., S. pascoei (Gahan, 1891), comb. n., S. sericatus (Pascoe, 1869), comb. n., S. tawauanus (Vives et Heffern, 2016), comb. n., Elydnus rufulus Holzschuh, 2016, stat. n. These new genera are also reviewed, with keys to their species given. The following new species are described: Dymasius makarovi sp. n. (Western Malaysia), D. murzini sp. n. (Sri Lanka), Elydnus barclayi sp. n. (Southern Thailand and Western Malaysia), E. tatianae sp. n. (Vietnam), E. vitalii sp. n. (Vietnam), Imbrius fedorenkoi sp. n. (Southern Vietnam), I. klimenkoi sp. n. (Eastern Malaysia), I. solodovnikovi sp. n. (Eastern Malaysia), Massicus ivani sp. n. (Eastern Malaysia), M. valentinae sp. n. (Western Malaysia), Sebasmia indochinensis sp. n. (Thailand and Vietnam), Zatrephus golovatchi sp. n. (Southern Vietnam), Zegriades olemehli sp. n. (Eastern Malaysia). The genus Sebasmia Pascoe, 1859 is reported from Indochina for the first time. New records of a number of species from other genera are given as well, thus one way or another expanding their distribution areas, sometimes very significantly so. Provisional considerations on the taxonomy of the genus Pachydissus Newman, 1838 and on the systematic position of Plavichydissus Pic, 1946 are presented. Based on a comparison of the holotypes of Dymasius macilentus (Pascoe, 1859) and D. strigosus J. Thomson, 1864, their synonymy traditionally used in the literature is questioned as requiring indisputable evidence. The lectotype male of Dymasius minor Gahan, 1906 is designated. Abundant pictures of the species studied, including numerous type specimens, are provided.
... Insect pests were mainly identified based on their morphology in taxonomic and other works as follows: Lepidoptera-keys [37][38][39][40]; Coleoptera-keys [41][42][43][44]; Hemiptera-keys [45,46]; Orthoptera-keys [47]. Phylogenetic analyses using the cytochrome c oxidase subunit I were undertaken to help confirm the identity of Batocera lineolata [48], Euwallacea fornicatus [31], and Tapinolachnus lacordairei [49]. ...
Article
Full-text available
The planted forest area in Vietnam increased from 3.0 to 4.4 million hectares in the period 2010–2020, but the loss of productivity from pests and diseases continues to be a problem. During this period, frequent and systematic plantation forest health surveys were conducted on 12 native and 4 exotic genera of trees as well as bamboo across eight forest geographic regions of Vietnam. Damage caused by insects and pathogens was quantified in the field and laboratory in Hanoi. The threats of greatest concern were from folivores (Antheraea frithi, Arthroschista hilaralis, Atteva fabriciella, Hieroglyphus tonkinensis, Lycaria westermanni,Krananda semihyalina, and Moduza procris), wood borers (Batocera lineolata, Euwallacea fornicatus, Tapinolachnus lacordairei, Xyleborus perforans, and Xystrocera festiva), sap-sucking insects (Aulacaspis tubercularis and Helopeltis theivora) and pathogens (Ceratocystis manginecans, Fusarium solani, and Phytophthora acaciivora). The number of new and emerging pests and pathogens increased over time from 2 in 2011 to 17 in 2020, as the damage became more widespread. To manage these pests and diseases, it is necessary to further invest in the selection and breeding of resistant genotypes, improve nursery hygiene and silvicultural operations, and adopt integrated pest management schemes. Consideration should be given to developing forest health monitoring protocols for forest reserves and other special-purpose forests.
... Adult body length was measured along the midline from the anterior of the eye to the apex of the elytra, and width was measured across the dorsal plate at the widest point. Identification to species was based on keys in Thomson (1864), Vitali et al. (2017) and Miroshnikov (2018). Thirty adult specimens of Tapinolachnus were deposited in the insect collection of the Forest Protection Research Centre -Vietnamese Academy of Forest Sciences, 46 Duc Thang Ward, Bac Tu Liem District, Hanoi, Vietnam with numbers coded from CT001 to CT030. ...
Article
Chukrasia tabularis (Meliaceae) has been planted for many decades in home gardens and plantations in Vietnam. Recently, growers have become concerned with attack by stem borers. Tapinolachnus lacordairei J. Thomson 1864 (Coleoptera: Cerambycidae) is identified as a new indigenous pest causing damage to 15 to 30 year-old stands of C. tabularis in Tuyen Quang, Hoa Binh and Thanh Hoa provinces. The leaves of infested trees prematurely senesce or wilt followed by tree death. In 2019, the infestation level was 12.5–22.5% and the damage index was 0.33–0.55. Adults emerge from February to May, mate within 1–3 days and the females lay 20–40 eggs on the bark of healthy and infested trees. The larvae feed first in the phloem and sapwood, and then in the heartwood where they make pupal chambers. Larvae take up to one year to become fully grown. To mitigate against further loss of C. tabularis, cost-effective control strategies need to be developed and steps taken to reduce the risk of dispersal of the pest beyond its native range in SE Asia.
Article
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This datasheet on Trirachys holosericeus covers Identity, Overview, Distribution, Dispersal, Hosts/Species Affected, Diagnosis, Biology & Ecology, Environmental Requirements, Natural Enemies, Impacts, Prevention/Control, Further Information.
Article
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The members of longhorn beetles are basically phytophagous and many are wood boring causing significant damage to many plants. This study discusses 28 species of such beetles belonging to 17 tribes, 24 genera and four subfamilies of agriculturally importance. Of these 13 species belong to the Lamiinae, 12 to Cerambycinae, two of Prioninae and one of Lepturinae. These had been collected from agriculture lands, fruits and plantation orchards. Brief descriptions of these along with synonyms, host range and distribution details are provided, of which 15 species are new reports from different states of India.
Article
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The Catalogue includes all 78 Cerambycidae species of Afghanistan fauna known up to 2019 with the references to the original descriptions; 22 species were not mentioned for Afghanistan in Palaearctic Cerambycidae Catalogue by Löbl & Smetana (2010). Bibliography of each species usually includes the geographical information from corresponding publications. Many new taxonomy positions published after 2010 are used here without special remarks. Agapanthia (Epoptes) dahli ustinovi Danilevsky, 2013 stat. nov. is downgraded from the species level. Two species are described as new Phytoecia (Parobereina) pashtunica sp. n. from Afghanistan and Phytoecia (Parobereina) heinzi sp.n. from Pakistan.
Article
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Hoplocerambyx inhirsutus (Matsushita, 1932) is transferred to the genus Aeolesthes Gahan, as it was originally described: Aeolesthes inhirsuta Matsushita, 1932 rest. status. Pachydissus externus Pascoe, 1869 is rehabilitated as valid species inside of the genus Aeolesthes, as follows: Aeolesthes externa (Pascoe, 1869) rest. status, n. comb. This species is also considered as an older synonym of Pachydyssus frenchi n. syn. Aeolesthes? nishikawai Hayashi, 1975 is transferred to the genus Nadezhdiella Plav., as follows: Nadezhdiella nishikawai (Hayashi, 1975) n. comb.
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