... Pterosaurs were the first vertebrates to fly, and many workers have reported on wing structure and aerodynamics. Relatively fewer workers have reported on trackway production and terrestrial locomotion (Stokes, 1957; Padian, 1983a Padian, , b, 1984 Padian, , 1985 Padian, , 2003 Padian and Olsen, 1984; Padian and Rayner, 1993; Bennett, 1990 Bennett, , 1997a Logue, 1994; Lockley et al., 1995 Lockley et al., , 1997 Mazin et al., 1995; Unwin, 1987 Unwin, , 1988 Unwin, , 1989 Unwin, , 1997 Hwang et al., 2002; Wellnhofer, 1975 Wellnhofer, , 1978 Wellnhofer, , 1988 Wellnhofer, , 1991 Chatterjee and Templin, 2004). Even fewer have compared pterosaur pedal skeletons (Wild, 1978; Wellnhofer, 1974 Wellnhofer, , 1978 Wellnhofer, , 1991 Unwin, 1989). ...