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Accepted by B. Mantovani: 5 Dec. 2006; published: 25 Jan. 2007 45
ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN 1175-5334 (online edition)
Copyright © 2007 · Magnolia Press
Zootaxa 1398: 45–56 (20 07)
www.mapress.com/zootaxa/
The Types of Phasmida in the Zoological Institute, Russian Academy of Sciences,
St. Petersburg (ZMAS)
PAUL D. BROCK
Department of Entomology, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.
E-mail: pauldbrock@btinternet.com
Abstract
Type specimens of 67 taxa of Phasmida (including probable type specimens of 24 taxa) have been located in the Zoolog-
ical Institute, Russian Academy of Sciences, St. Petersburg. The species are listed alphabetically, with the number of
specimens, sex and locality data.
Key words: Phasmida, type species, St. Petersburg, Bey-Bienko, Brunner, Mishchenko, Redtenbacher
Introduction
In their 1906–08 monograph on Phasmida, Brunner and Redtenbacher indicated the presence of type material
of 40 species in the Zoological Museum, Academy of Science, St. Petersburg collection. These authors also
mentioned non-type specimens of species already described by other authors. In order to clarify the position
with certain material, I visited St. Petersburg in August, 2006, to examine this neglected collection and pro-
duce a list of type material.
The collection contains about 40 drawers of older identified material, mainly collected pre-1901 from
many countries, but with an emphasis on South Asia. There are also approximately 80 drawers of recently col-
lected, undetermined material, mainly from Central and South Asia (much collected by Gorochov). After a
detailed examination of older specimens and data labels, also discussions with the present curator, Andrej
Gorochov, it has been established that many specimens were obtained from and / or identified by Brunner and
Redtenbacher, whose determination labels are attached to specimens. These are often specimens described or
mentioned in their monograph (1906–08). It is believed that many specimens were purchased from the Brun-
ner collection, after his death (Gorochov, verbal information from a previous curator). Whilst much of Brun-
ner’s collection is in the Naturhistorisches Museum, Vienna (NHMW), specimens have been widely
distributed, for example by exchange (Brock, 1998), hence part of the type series is housed in depositories not
specified in the monograph. I traced a number of such specimens in St. Petersburg (see discussion for possible
origin).
Many older specimens in the St. Petersburg collection (including those described by Brunner and Redten-
bacher) originate from German sources, including natural history specimen dealers such as Hans Fruhstorfer
(1866–1922) and Ernst Heyne (1833–1905) (Asian specimens marked ‘Heyne, 901’), or expeditions by Rus-
sian entomologists. Madagascan material was almost certainly purchased from Franz Sikora (1862–63 to
1902).
Most of the definite type material i.e. mentioned in the monograph, and agreeing in measurements given,
have separate labels with numbers on (Sikora’s Madagascan material does not). In some other European col-
lections (Brock, 1998) the use of numbers linked directly with handwritten listings of species names supplied
BROCK
46 · Zootaxa 1398 © 2007 Magnolia Press
by Brunner. Unfortunately there are no documents available to inform us. The possible type material i.e. spec-
imens not listed in the monograph as being present in St. Petersburg, mostly have numbered labels, as well as
Brunner and Redtenbacher determination labels.
Very few entomologists have examined the orthopteroid collections in St. Petersburg. The Phasmida collec-
tion is mainly as curated by Nícolaj Adelung before he retired in c. 1916. Some type material has small round
labels beneath the pins. The taxonomy is in accordance with Brunner and Redtenbacher’s monograph (1906–
08), with labelled spaces left in the collection for representatives of other genera; as a result, some drawers
contain very few specimens. The next Technical Assistant / curator, E. F. Miram, identified a few phasmids,
adding his determination labels. Subsequent curators (Mishchenko, Bey-Bienko) also identified and described
a few species of interest from Asia. Gorochov and Brock (in progress) are working on describing Heteroptery-
gidae collected by Gorochov. Type material is now denoted in the collection by red labels pinned alongside
specimens.
In order to assist phasmid researchers, this is considered an appropriate time to publish a list of type mate-
rial. Photographs of all type material will be made available on the Phasmida Species File Online (author:
Paul D. Brock) http://phasmida.orthoptera.org/, which has been checked for details of type records in other
collections.
Treatment of the species is as follows:
Primary types (holotypes, lectotypes and syntypes) and secondary type material (paralectotypes and
paratypes) are listed alphabetically by their original specific or subspecific name. The information for each
type is given in the following standard sequence:
Specific name, author, date and page reference of original publication, original genus in parentheses.
Status and sex, locality, any bionomic information, date of collection, collector (in parentheses and itali-
cized, except where a specimen is from a particular person’s collection but may not have been collected by
that person the name of the person, followed by “coll.”), reference to lectotype designation, mention of any
significant damage, brief notes on whereabouts of other type material (in all cases, if applicable).
Any further annotations. The current name followed by the current higher classification (following Otte &
Brock, 2005, except where updated online due to current thinking on two tribes) is given in square brackets
after all the other text about a taxon.
In listing the data for each specimen the name of the country, in its correct form, is given first, in capitals;
the remaining items of locality data are then listed in order of magnitude, irrespective of the order in which
they may appear on the label(s). When the country name given on the label differs from the one in current
use, the name given on the label is quoted in inverted commas, after the current name of the country of place.
The data listed for each specimen is taken from the label(s) pinned underneath the specimen. However, all
the specimens have been checked against the original descriptions to ensure that type material has been cor-
rectly interpreted. Any additional information added and marked as ’[published information]’.
Abbreviations for Depositories given in the listings are as follows:
ANSP Academy of Natural Sciences, Philadelphia, U.S.A.
BMNH The Natural History Museum, London, U.K.
ETHZ Entomologisches Institut, Eidgenössische Technische Hochschule-Zentrum, Zürich, Switzer-
land.
HNHM Hungarian Natural History Museum, Budapest, Hungary [types destroyed in fire]
ISNB Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium
MCSN Museo Civico di Storia Naturale “Giacomo Doria”, Genova, Italy
MHNG Muséum d’histoire naturelle, Geneva, Switzerland
MNHN Muséum national d’Histoire naturelle, Paris, France
MNMS Museo Nacional de Ciencias Naturales, Madrid, Spain
NHMB Naturhistorisches Museum, Basel, Switzerland
Zootaxa 1398 © 2007 Magnolia Press · 47
TYPES OF PHASMIDA IN ZMAS
NHMW Naturhistorisches Museum, Vienna, Austria
RMTO Museo Regionale di Scienze Naturali, Torino, Italy
SMNS Staatliches Museum für Naturkunde, Stuttgart, Germany
SMTD Staatliches Museum für Tierkunde, Dresden, Germany
UUZM Uppsala University Zoological Museum, Uppsala, Sweden
ZMAN Zoologisch Museum, Amsterdam, Netherlands
ZMHB Museum für Naturkunde der Humboldt-Universitat, Berlin, Germany
ZMUH Zoologischen Museum der Universität Hamburg, Germany
The following alphabetical list refers to type material confirmed as being in St. Petersburg by the original
authors and traced during the present project. Other species marked (*) refer to probable type specimens,
although ‘St. Petersburg’ was not mentioned by the original authors. Likely explanations for this are given
above.
*aculeata Redtenbacher, 1906: 25 (Antongilia)
2 probable ST: 2 MAD[AGASCAR], Antongil. There are 10 additional syntypes (4 , 5 & 1 juv.), in
MNHN (1 ), NHMW (3 & 3 ) and ZMUH (1 , 1 & 1 juv.).
= Pseudonogastris aculeata (Redtenbacher, 1906) (Bacillidae: Antongiliinae: Leprodini)
adelungi Brunner, 1907: 221 (Gratidia)
HT: [TURKMENISTAN], Transcaspien, Tsczardjui, 25.iv.[19]04 (Fischer).
= Clonaria adelungi (Brunner, 1907) (Diapheromeridae: Pachymorphinae: Gratidiini)
adspirans Redtenbacher, 1908: 499 (Asceles)
3 ST: 3 [INDONESIA], Jawa ‘Java occident.’, Bogor [Buitenzorg], 1901 (Pedaschenko) [one specimen
omits ‘Buitenzorg’]. There is an additional syntype in NHMW (1 ) [ex. St. Petersburg coll.]
= Asceles adspirans Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
albofasciata Redtenbacher, 1908: 560 (Necroscia) (Figs. 1 & 2)
HT: [INDONESIA], Bacan ‘Batjan’, ex. Heyne coll.
= Necroscia albofasciata Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
FIGURE 1. Necroscia albofasciata Redtenbacher – holotype female, dorsal view.
BROCK
48 · Zootaxa 1398 © 2007 Magnolia Press
FIGURE 2. Necroscia albofasciata Redtenbacher – holotype female, lateral view showing data labels.
aliena Brunner, 1907: 222 (Gratidia)
1 ST: 1 (not as stated) [KAZAKHSTAN], Dzhungaria, river Ili, West Kuldzha, v.1878 (A. Regel) [speci-
men is 52mm, as recorded. The 45mm mentioned has not been traced, although it was referred to by Mish-
chenko, 1937: 119].
= Junior subjective synonym of Clonaria bituberculata (Redtenbacher, 1889) (Diapheromeridae: Pachymor-
phinae: Gratidiini)
*annamensis Brunner, 1907: 205 (Cuniculina)
3 probable ST: 3 [VIETNAM], Phuc-Son, xi.–xii., ex. H. Fruhstorfer coll.
There are 20 additional syntypes (13 & 7 juv.), in ISNB (5 ) and NHMW (8 & 7 juv).
= Junior subjective synonym of Medauroidea extradentata (Brunner, 1907) (Phasmatidae: Phasmatinae: Cli-
tumnini)
*angustata Redtenbacher, 1908: 562 (Necroscia)
2 probable ST: 2 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll.
There are 11 additional syntypes (9 & 2 ) in NHMW. Also, another probable syntype (1 ) in MNHN.
= Nescicroa angustata Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
anterior Brunner, 1907: 201 (Cuniculina)
HT [INDIA], Assam, ex. Heyne coll.
= Ramulus anterior (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
arrogans Brunner, 1907: 207, pl. 11: 1a–c (Prosentoria)
2 ST: 2 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. There are 11 additional syntypes (5 & 6 ) in
NHMW (4 & 3 ), SMTD (1 & 1 ) and ZMUH (2 ).
= Prosentoria arrogans Brunner, 1907 (Phasmatidae: Phasmatinae: Clitumnini)
asperrimus Redtenbacher, 1906: 42 (Obrimus)
6 ST: 4 & 2 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. There are 6 additional syntypes (3 &
3 ) in NHMW.
= Aretaon asperrimus (Redtenbacher, 1906) (Heteropterygidae: Obriminae: Obrimini)
Zootaxa 1398 © 2007 Magnolia Press · 49
TYPES OF PHASMIDA IN ZMAS
beybienkoi Bekuzin, 1960: 661 (Gratidia)
HT & up to 46 PT (number in ZMAS not confirmed). HT, 21 & 18 PT Southern KAZAKHSTAN,
Keles River basim, Karataskii District, c. 600m, near Leninsk, 29-30.vii.1957 (Bekuzin). Other PT from
KAZAKHSTAN: 1 & 1 heights of Kazgurt, 1500m, 26.vii.1957 (Bekuzin); 2 Korzhantau Mts., Kzyltal
along Ugam River Valley, 1400m, 10.vii.1957 (Bekuzin); 1 & 1 south slopes of Korzhantau, Aigyrdzhin-
ilgan, 1750m, 12.vii.1957 (Bekuzin); 1 UZBEKISTAN, Tashkent Region, Bostandyk District, Ugam River
valley, Khumsan, 20.viii.1958. There are 3 additional syntypes (1 & 2 nymphs) in the Museum of Nature,
AN Uzb.
= Clonaria beybienkoi (Bekuzin, 1960) (Diapheromeridae: Pachymorphinae: Gratidiini)
biceps Redtenbacher, 1908: 451 (Pharnacia)
1 ST: 1 [INDONESIA], Jawa ‘JAVA occident.’, Mons Gede, 4000 ft., 1898.ex. H. Fruhstorfer coll. There
are 24 additional syntypes (8 , 13 & 3 juv.) in MHNG (2 & 2 ), MNHN (1 & 2 ), NHMW (4 ,
6 & 2 juv.), SMNS (1 ), ZMHB (2 & 1 juv.) and ZMUH (1 ). HNHM material was destroyed in
fire.
= Pharnacia biceps Redtenbacher, 1908 (Phasmatidae: Phasmatinae: Pharnaciini)
buruensis Brunner, 1907: 252 (Acacus)
1 ST: 1 [INDONESIA], Bacan ‘Batjan’, ex. Heyne coll. Not traced in NHMW (Brock, 1998).
Junior subjective synonym of Orxines xiphias (Westwood, 1859) (Diapheromeridae: Necrosciinae)
*caryophyllea Redtenbacher, 1908: 510 (Marmessoidea)
2 probable ST: 2 [MALAYSIA] ‘N.Borneo’, ex. Heyne coll. There are at least 17 additional syntypes (12 ,
1 juv., 4 & alcohol material) in MNHN (3 & 1 ), NHMW (7 , 1 juv., 2 & alcohol material),
ZMHB (1 ) and ZMUH (1 & 1 ).
= Junior subjective synonym of Marmessoidea vinosa (Serville, 1838) (Diapheromeridae: Necrosciinae)
chinensis Brunner von W., 1907: 202 (Cuniculina)
1 ST: 1 CHINA, Sztschwan, 2.viii.[18]93. There are 4 additional syntypes (1 & 3 ) in MNHN (1 & 1
) and NHMW (2 ) [see Brock & Delfosse (2005: 51) for notes on confusion regarding this species].
= Ramulus chinensis (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
cornucervi Redtenbacher, 1908: 496 (Asceles)
HT: [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll.
= Asceles cornucervi Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
*cristatus Brunner, 1907: 270 (Carausius)
1 probable ST: 1 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll.
There are 3 additional syntypes (3 ) in NHMW.
= Carausius cristatus Brunner, 1907 (Diapheromeridae: Lonchodinae)
cylindripes Redtenbacher, 1906: 51 (Datames)
1 PLT: 1 [INDONESIA], Jawa ‘Java occident.’, Bogor ‘Buitenzorg’, 1901 (Pedaschenko). The lectotype
is in MNHN.
= Junior subjective synonym of Pylaemenes oileus (Westwood, 1859) (Heteropterygidae: Dataminae:
Datamini)
BROCK
50 · Zootaxa 1398 © 2007 Magnolia Press
*difformis Redtenbacher, 1906: 163 (Anisacantha)
5 probable ST: 2 & 3 MAD[AGASCAR], Antongil. There are 31 additional syntypes (17 , 14 (incl.
juv.) ) in MNHN (1 ), NHMW (15 & 11 (including juv.) and ZMUH (2 , 1 & 1 juv.).
= Anisacantha difformis Redtenbacher, 1906 (Anisacanthidae: Anisacanthinae: Anisacanthiini)
*dolorosa Redtenbacher, 1908: 547 (Sipyloidea)
1 probable 1 [INDONESIA], Jawa ‘Java’. There are 14 additional syntypes (7 & 5 and 2 unknown sex)
in ISNB (2 specimens) and NHMW (7 & 5 ). HNHM material was destroyed in fire.
= Sipyloidea dolorosa Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
echinata Redtenbacher, 1906: 171, pl. 6 : 10–11 (Heteropteryx)
1 ST: 1 [MALAYSIA] ‘N[or]d’ Borneo, ex. Heyne coll. There are 6 additional syntypes (2 , 2 & 2
juv.) in NHMW (2 , 1 & 2 juv.) and ZMHB (1 ).
= Haaniella echinata (Redtenbacher, 1906) (Heteropterygidae: Heteropteryginae: Heteropterygini)
echinops Redtenbacher, 1906: 25 (Antongilia)
HT MADAGASCAR mer., Fort Dauphin, 1899 (Sikora).
= Antongilia echinops Redtenbacher, 1906 (Bacillidae: Antongiliinae: Antongiliini)
*extradentatus Brunner, 1907: 193 (Clitumnus)
2 probable ST: 2 [VIETNAM], Phuc-Son, xi.–xii., ex. H. Fruhstorfer coll.
There are 12 additional syntypes (12 ) in NHMW.
= Medauroidea extradentata (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
femoralis Brunner, 1907: 262 (Lonchodes)
HT [SRI LANKA] ‘Ceylon’, 1887 (V. Griswaldt).
= Lonchodes femoralis Brunner, 1907 (Diapheromeridae: Lonchodinae)
ferruginea Redtenbacher, 1908: 569 (Calvisia)
1 ST: 1 [MALAYSIA] ‘Nord Borneo’, ex. Heyne coll. There are 6 additional syntypes (6 ) in ISNB (2 )
and NHMW (4 ).
= Calvisia ferruginea Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
grisescens Redtenbacher, 1906: 137 (Perliodes)
1 ST: 1 [PANAMA], Chiriqui, ex. Heyne coll. There are 4 additional syntypes (3 & 1 ) in NHMW (1 )
and SMTD (2 & 1 ).
= Perliodes grisescens Redtenbacher, 1906 (Pseudophasmatidae: Xerosomatinae: Prexaspini)
horridus Brunner, 1907: 244 (Menexenus)
HT NEW GUINEA, Bongu, Kaiser Wilhelmsland.
= Menexenus horridus Brunner, 1907 (Diapheromeridae: Lonchodinae)
*immundus Brunner, 1907: 270 (Carausius)
1 probable ST: 1 [SULAWESI] S. Celebes, Bua Kraeng, 5000 ft., 11.1896, ex. Coll. Fruhstorfer. There are
11 additional syntypes (6 & 5 ), as follows: NHMB (3 ) and NHMW (6 & 2 ). Not traced in MNHN.
= Junior subjective synonym of Leprocaulinus insularis insularis (Kirby, 1896) (Diapheromeridae: Necrosci-
inae)
Zootaxa 1398 © 2007 Magnolia Press · 51
TYPES OF PHASMIDA IN ZMAS
inconspicua Brunner, 1907: 223 (Gratidia)
7 ST: 1 & 2 UZBEKISTAN, Buchara, [18]96, 2 [IRAN] ‘Persia’, Kirman, viii.[18]98, 1 [IRAN]
‘Persia’, Khorasan, 26.vii.[19]01, 1 Road from Trauscaspían (Turkmenia) to Kuchan (Persia), 15.iii.[18]98.
There are 3 additional syntypes (1 & 2 ) in NHMW [ex. St. Petersburg coll.].
= Clonaria inconspicua (Brunner, 1907) (Diapheromeridae: Pachymorphinae: Gratidiini)
*inermis Redtenbacher, 1906: 68 (Canuleius)
2 ST: 1 & 1 [BRAZIL], Theresopolis. There are 15 additional syntypes (6 , 1 juv., 5 & 3 juv.) in
NHMW (4 , 1 juv., 2 & 1 juv.) and ZMUH (2 , 3 & 2 juv). Not traced in ZMHB.
= Canuleius inermis Redtenbacher, 1906 (Heteronemiidae: Heteronemiinae: Pygirhynchini)
*inquinatus Redtenbacher, 1908: 498 (Asceles)
8 probable PLT: 8 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. [one clearly belongs to a different
species]. The lectotype is in NHMW. There are 6 additional paralectotypes (4 & 2 ) in NHMW. Also a
probable syntype (1 ) in MNHN.
= Junior subjective synonym of Asceles larunda (Westwood, 1859) (Diapheromeridae: Necrosciinae)
insignis Brunner, 1907: 229 (Adelungella)
2 ST: 2 [IRAN] ‘Persia’, Kirman, 17.vi.[18]98 & 28.iii.[19]01 (Zarndny).
= Adelungella insignis Brunner, 1907 (Diapheromeridae: Pachymorphinae: Gratidiini)
insignis Redtenbacher, 1908: 474 (Parastheneboea)
2 ST: 2 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. There are 2 additional syntypes (2 ) in
NHMW.
= Junior subjective synonym of Parastheneboea laetior laetior (Brunner, 1907) (Diapheromeridae: Necrosci-
inae)
linearis Redtenbacher, 1908: 544 (Sipyloidea)
1 ST: 1 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. [This appears to be the specimen labelled with
the manuscript name ‘Platysosibia sodalis’]. There are additional syntypes (1 ) in NHMW and MNMS
(number of not known).
= Sipyloidea linearis Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
longelaminata Brunner, 1907: 200 (Cuniculina)
HT [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. [This specimen is 94mm in length excluding the
operculum (Brunner mentioned 100mm)].
= Junior subjective synonym of Prosentoria arrogans Brunner, 1907 (Phasmatidae: Phasmatinae: Clitumnini)
*margaritatus Redtenbacher, 1908: 496 (Asceles)
18 probable ST: 4 & 14 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll.
There are 7 additional syntypes (4 & 3 ) in NHMW.
= Asceles margaritatus Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
marginale Redtenbacher, 1906: 115 (Paraphasma)
2 PLT: 1 BRAZIL ‘Brasilia’, 1 [BRAZIL?] Schrerik’s Reise. The lectotype is in MCSN. There are at
least 36 additional paralectotypes (24 & 12 ) in MCSN (1 & 2 ), MNHN (number not known), MNMS
(number not known), NHMW (15 & 9 ), SMNS (3 ), SMTD (1 ) and ZMUH (4 & 1 ).
= Paraphasma marginale Redtenbacher, 1906 (Pseudophasmatidae: Stratocleinae: Stratocleini)
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52 · Zootaxa 1398 © 2007 Magnolia Press
*marginatus Redtenbacher, 1906: 81 (Presbistus)
1 probable PLT: 1 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll.
The lectotype is in NHMW. There are at least 4 additional paralectotypes (2 & 2 ), as follows: MHHN
(1 ), MNMS (number not known), NHMW (1 & 1 ) and ZMUH (1 ).
= Kerabistus (Rhadinobistus) marginatus (Redtenbacher, 1906) (Aschiphasmatidae: Aschiphasmatinae:
Aschiphasmatini)
*marmoratum Redtenbacher, 1906: 119 (Phasma)
1 probable ST: 1 COLOMBIA ‘Columbia’. There are 12 additional syntypes (6 & 6 ) in NHMW (4 &
3 ), ZMHB (1 & 3 ) and ZMUH (1 ). Also in MNMS (number not known).
= Pseudophasma marmoratum Redtenbacher, 1906 (Pseudophasmatidae: Pseudophasmatinae: Pseudophas-
matini)
*memorabilis Redtenbacher, 1906: 21 (Pseudodatames)
2 probable PLT: 1 & 1 MADAGASCAR, Antongil. The lectotype is in NHMW. There are at least 11
additional paralectotypes (5 & 5 and 1 unknown sex), as follows: ISNB (1), MNHN (2 & 1 ), NHMW
(2 & 3 [a further became a PT of Pseudodatames bicornutus Zompro, 2004], ZMUH (1 & 1 juv.).
= Pseudodatames memorabilis Redtenbacher, 1906 (Bacillidae: Antongiliinae: Pseudodatamini)
muricata Redtenbacher, 1906: 28 (Antongilia)
1 ST: 1 MADAGASCAR mer., Ranomafana, 1899 (Sikora). There are 7 additional syntypes (6 & 1 ) in
MNHN (1 & 1 ) and NHMW (5 ).
= Antongilia muricata Redtenbacher, 1906 (Bacillidae: Antongiliinae: Antongiliini) [Cliquennois (in
progress), is transferring this species to a new genus in the tribe Leprodini]
muscosus Redtenbacher, 1906: 41 (Obrimus)
3 ST: 2 juv., 1 juv. [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. There are 3 additional syntypes (1
& 2 ) in NHMW.
= Junior subjective synonym of Aretaon asperrimus (Redtenbacher, 1906) (Heteropterygidae: Obriminae:
Obrimini)
nana Mishchenko, 1941: 145 (Gratidia)
HT & 2 PT: HT, 2 PT [AZERBAIJAN] Nachichevan, Transcaucasia, 2.vii.1931 (ht), 4.vii.1931 & 7–
8.vii.1932 (pt’s).
= Clonaria nana (Mishchenko, 1941) (Diapheromeridae: Pachymorphinae: Gratidiini)
*nigrotuberculata Redtenbacher, 1906: 73 (Orobia)
1 probable ST: 1 MAD[AGASCAR]. There are 19 additional syntypes (8 & 11 ), in MNHN (4 ) and
NHMW (8 & 7 ).
= Leiophasma nigrotuberculatum (Redtenbacher, 1906) (Anisacanthidae: Anisacanthinae: Anisacanthiini)
*normalis Redtenbacher, 1908: 547 (Sipyloidea)
3 probable ST: 1 [INDONESIA], Jawa ‘Java occident.’, Mons Gede, 4000 ft., 1898, ex. Coll. Fruhstorfer, 1
[SULAWESI] ‘Nord-Celebes’, Toli-Toli, xi.-xii.1895, ex. Coll. Fruhstorfer, 1 [SULAWESI] ‘Celebes.’
There are 51 additional syntypes (28 & 23 ) in NHMW (23 & 23 ), SMNS (1 ), ZMHB (2 ) and
ZMUH (2 ). HNHM material was destroyed in fire.
= Junior subjective synonym of Nescicroa poeciloptera (Rehn, 1904) (Diapheromeridae: Necrosciinae)
Zootaxa 1398 © 2007 Magnolia Press · 53
TYPES OF PHASMIDA IN ZMAS
obscurellus Redtenbacher, 1906: 137 (Perliodes)
HT: ECUADOR, Paramba.
= Olcyphides obscurellus (Redtenbacher, 1906) (Pseudophasmatidae: Stratocleinae)
*operculata Redtenbacher, 1906: 70 (Orobia)
1 probable ST: 1 MAD[AGASCAR], Antongil. There are 8 additional syntypes (3 & 5 ), in MNHN (1
& 2 ), NHMW (2 & 2 ) and ZMUH (1 ).
= Leiophasma operculatum (Redtenbacher, 1906) (Anisacanthidae: Anisacanthinae: Anisacanthiini)
oxyacantha Redtenbacher, 1908: 457 (Acrophylla)
1 PLT: 1 Locality not known (listed as ‘?’. There are specimens of several other species in the collection,
probably from the same source, clearly from AUSTRALIA). The lectotype is in NHMW, also 5 paralectoty-
pes (5 ). There is an additional paralectotype (1 juv.) in ZMUH. No specimens have been traced in
MNHN.
= Junior subjective synonym of Ctenomorpha marginipennis Gray, 1833 (Phasmatidae: Phasmatinae: Phas-
matini)
plagiatus Redtenbacher, 1906: 135 (Isagoras)
2 ST: 1 BRAZIL ‘Brasil’, 1 BRAZIL ‘Brasil’, Espirito Santo, ex. Fruhstorfer coll. (tip of abdomen bro-
ken off and missing). There are at least 16 additional syntypes (13 , 1 & 2 juv.), BMNH (1 ), MNHN (1
), MNMS (number not known), NHMW (6 , 1 & 2 juv. (sex?)), ZMHB (1 ) and ZMUH (4 ).
= Isagoras plagiatus Redtenbacher, 1906 (Pseudophasmatidae: Xerosomatinae: Prexaspini)
predtetshenskyi Bey-Bienko, 1946: 74 (Gratidia)
HT: [IRAN] ‘Persia’, Kazvin, 7.ix.193? (Predtetshensky).
= Clonaria predtetshenskyi (Bey-Bienko, 1946) (Diapheromeridae: Pachymorphinae: Gratidiini)
recessum Redtenbacher, 1908: 490 (Centema)
1 ST: 1 [INDONESIA], Jawa ‘Java occident.’, Mons Gede, 4000 ft., 1898, ex. Coll. Fruhstorfer. There are 2
additional syntypes (2 ) in NHMW (1 ) and SMNS (1 ).
= Centrophasma recessum (Redtenbacher, 1908) (Diapheromeridae: Necrosciinae)
reductus Brunner, 1907: 254 (Myronides)
HT [INDONESIA], Jawa ‘Java occident.’, Pengalengan, 4000 ft., 1893, ex. Coll. Fruhstorfer [The descrip-
tion and measurements agree with the specimen labelled with the manuscript name ‘Gratidia javana’].
= Myronides reductus Brunner, 1907 (Diapheromeridae: Lonchodinae)
rivalis Brunner, 1907: 194 (Clitumnus)
1 PLT: 1 [SRI LANKA] ‘Ceylon’ (Dr Bunge). The lectotype is in NHMW, also 3 additional paralectoty-
pes (3 ).
= Ramulus rivalis (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
ruficornis Redtenbacher, 1906: 86, pl. 4:9 (Dina)
6 PLT: 2 & 4 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. The lectotype is in NHMW. There
are 4 additional paralectotypes (2 & 2 ) in MNHN (1 ) and NHMW (2 & 1 ).
= Dinophasma ruficorne (Redtenbacher, 1906) (Aschiphasmatidae: Aschiphasmatinae: Aschiphasmatini)
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54 · Zootaxa 1398 © 2007 Magnolia Press
scabra Redtenbacher, 1906: 171 (Heteropteryx)
1 ST: 1 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. [The specimen has been misidentified initially,
with a data label ‘Heteropteryx gayi’ [sic]]. There are 3 additional syntypes (1 & 2 ) in ETHZ (1 ) and
NHMW (1 & 1 ).
= Haaniella scabra (Redtenbacher, 1906) (Heteropterygidae: Heteropteryginae: Heteropterygini)
*serripes Redtenbacher, 1908: 364 (Dimorphodes)
1 probable ST: 1 [INDONESIA], Bacan ‘Batjan’, ex. Heyne coll. There are 15 additional syntypes (6 , 7
& 2 juv.), as follows: MCSN (1 ), NHMW (2 & 2 ), SMTD (1 , 2 & 2 juv.), ZMAN (1 ) and
ZMHB (2 & 2 ).
= Dimorphodes prostasis serripes Redtenbacher, 1906 (Phasmatidae: Xeroderinae)
*serrulatus Brunner, 1907: 190 (Clitumnus)
3 probable ST: 3 [INDONESIA], Jawa ‘Java’. There are 41 additional syntypes (19 , 20 & 2 juv.), in
ANSP (1 ), ISNB (1 ), NHMW (6 , 4 ), RMTO (2 ), SMNS (4 ), ZMAN (1 ), ZMHB (4 , 12 &
2 juv.) and ZMUH (2 & 2 ). HNHM material from Borneo was destroyed in fire.
= Ramulus serrulatus (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
signata Redtenbacher, 1908: 485 (Diardia)
1 ST: 1 [MALAYSIA] ‘Borneo’, Kina-Balu, ex. Heyne coll. [Appears to be the specimen with a manuscript
name ‘Diesbachia incontaminata Br[unner]’]. There are 6 additional syntypes (6 ) in MCSN (1 ) and
NHMW (5 ).
= Junior subjective synonym of Diardia diardi (de Haan, 1842) (Diapheromeridae: Necrosciinae)
*simplex Redtenbacher, 1906: 26 (Antongilia)
4 probable PLT: 2 , 1 & 1 juv. MAD[AGASCAR], Antongil. The lectotype is in NHMW. There are 14
additional paralectotypes (7 , 5 , 1 juv., 1 unknown sex), in ISNB (1 sex?) MNHN (1 & 1 ), NHMW
(3 & 4 ) and ZMUH (3 & 1 juv.).
= Virgasia simplex (Redtenbacher, 1906) (Bacillidae: Antongiliinae: Leprodini)
spinulosa Brunner, 1907: 228 (Gratidia)
HT [IRAN] ‘Persia’, Bampur, 25.iv.[19]01 (Zarndny).
= Clonaria spinulosa (Brunner, 1907) (Diapheromeridae: Pachymorphinae: Gratidiini)
*squamigera Redtenbacher, 1906: 27 (Antongilia)
1 probable PLT: 1 MADAGASCAR, Antongil. The lectotype is in NHMW. There are 5 additional para-
lectotypes (3 & 2 juv.), in MNHN (1 ) and NHMW (2 & 2 juv.).
= Paronogastris squamigera (Redtenbacher, 1906) (Bacillidae: Antongiliinae: Antongiliini)
sykorai Redtenbacher, 1906: 158 (Damasippoides)
HT MADAGASCAR mer., Fort Dauphin, 1899 (Sikora).
= Damasippoides sykorai Redtenbacher, 1906 (Damasippoididae: Damasippoidinae: Damasippoidini)
trispinosa Brunner, 1907: 223 (Gratidia)
2 ST: 2 MADAGASCAR mer., Ranomafana, 1899 (Sikora) [One is 52mm, the second has the abdomen
broken off and missing, probably c. 59mm, the length given by Brunner].
= Clonaria trispinosa (Brunner, 1907) (Diapheromeridae: Pachymorphinae: Gratidiini) [Cliquennois (pers.
com.) is working on this species and believes that it belongs to the tribe Leprodini in Antongiliinae]
Zootaxa 1398 © 2007 Magnolia Press · 55
TYPES OF PHASMIDA IN ZMAS
*unispinosa Redtenbacher, 1908: 532 (Diacanthoidea)
1 probable ST: 1 [MALAYSIA] ‘B[ritish] N[orth] Borneo’, Labuan, ex. Swinhoe & Heyne colls. There are
6 additional syntypes (1 & 5 ) in NHMW.
= Diacanthoidea unispinosa Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
ussurianum Bey-Bienko, 1960: 690 (Baculum)
2 ST: 2 [RUSSIA], Southern Ussuriland, Middle stretch of Daubikhe River, Anuchinskii district, 28–
30.viii.1957 (E. S. Sinchilina).
= Ramulus ussurianus (Bey-Bienko, 1960) (Phasmatidae: Phasmatinae: Clitumnini)
uvaroviana Mishchenko, 1937: 119 (Gratidia)
HT & 1 PT. HT TURKMENISTAN, V. Keltetshinar, 16.viii.1928 (Maximov). PT: 1 IRAN, Khorasan,
Hassan-Khurda, 24.iv.1858 (Kaizerling & Binert).
= Clonaria uvaroviana (Mishchenko, 1937) (Diapheromeridae: Pachymorphinae: Gratidiini)
*vaginata Redtenbacher, 1906: 70 (Orobia)
1 probable ST: 1 MAD[AGASCAR], Antongil. There are 11 additional syntypes (3 , 8 ), in ANSP (1 ),
ISNB (1 ), MNHN (4 ), NHMW (1 & 2 ) and ZMUH (1 & 1 ).
= Leiophasma vaginatum (Redtenbacher, 1906) (Anisacanthidae: Anisacanthinae: Anisacanthiini)
No type specimens of the following species were traced:
amica Bey-Bienko, 1959: 1818, fig. 4 (Sipyloidea)
CHINA, Mt. Tingushan [Possibly misplaced or may have been forwarded to Beijing].
= Sipyloidea amica Bey-Bienko, 1959 (Diapheromeridae: Necrosciinae)
irregularis Brunner 1907: 195 (Clitumnus)
juv. [INDONESIA], Jawa ‘Java’, Bogor ‘Buitenzorg’ [Specimen apparently retained in NHMW].
= Ramulus irregularis (Brunner, 1907) (Phasmatidae: Phasmatinae: Clitumnini)
spinipennis Redtenbacher, 1908: 521 (Aruanoidea)
[MALAYSIA] Nord-‘Borneo’ [Possibly a misidentified specimen of a related species, but not traced with
any degree of certainty].
= Scionecra spinipennis (Redtenbacher, 1908) (Diapheromeridae: Necrosciinae)
vittata Redtenbacher, 1908: 532 (Diacanthoidea)
[INDONESIA], Jawa ‘West-Java’ [Not traced, despite checking of species with manuscript names, also
undetermined material].
= Diacanthoidea vittata Redtenbacher, 1908 (Diapheromeridae: Necrosciinae)
Thunberg (1915) did not specify that type material was in St Petersburg, although the journal Mémoires de
l’Académie impériale des sciences de Saint-Pétersbourg implies a possible link. However, none of his type
material was found. Brock (2002) stated that all of Thunberg’s type material is lost, except for three species in
UUZM.
Discussion
The unknown factor is how so much probable type material became deposited in the St. Petersburg collection.
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56 · Zootaxa 1398 © 2007 Magnolia Press
There are various possibilities:
1) Brunner and Redtenbacher omitted to mention St. Petersburg in their monograph. This is unlikely
because so many species are involved, but possible for a few specimens, such as those originating from
Heyne.
2) Material was purchased or exchanged at some stage. Specimens do not include the NHMW collection
numbers on the attached Brunner & Redtenbacher determination labels, hence are more likely to originate
from Brunner’s personal collection. This is arguably the most likely explanation, confirmed by comment from
a past curator, albeit verbal.
3) Prior to publication of the monograph, Redtenbacher is known to have worked on the orthopteroid col-
lection in the Zoological Museum, St. Petersburg and may have added labels.
4) As an alternative to option 3) above, Brunner and Redtenbacher may have examined material in
Vienna, sometime after 1901 (last collection dates), but before publication of the first part of the monograph
in 1906. This option is supported by numbered material, a method used in several other European museum
collections.
Acknowledgements
I would like to thank Andrej Gorochov (ZMAS) for his assistance with the collection, including history, trans-
lation of data labels in Russian and comments on the draft manuscript. Nicolas Cliquennois (Madagascar) also
kindly commented on the manuscript and provided information on Madagascan species.
References
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