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ACTA ZOOLOGICA BULGARICA
Acta zool. bulg., 68 (3), 2016: 305-310
Systematics and Phylogenetics
Research Article
Introduction
The Dorcadion mniszechi species-group, as de-
ned here, includes only two species: Dorcadion
mniszechi Kraatz, 1873 and D. semibrunneum Pic,
1903. Dorcadion mniszechi (the type species of
the subgenus Cribridorcadion Pic, 1901) was de-
scribed by Kr a a t z (1873) from the Transcaucasian
Region between Iğdır Province in Turkey and the
Sevan Lake in Armenia (La z a r e v 2014). It is dis-
tributed in Armenia, Georgia and Turkey (North-
eastern Anatolia) and belongs the Southwestern
Asiatic (Anatolo-Caucasian) chorotype. Recently,
La z a r e v (2014) described two new subspecies of D.
mniszechi: D. mniszechi georgianum from Georgia
and D. mniszechi cavernosum from Armenia and
Turkey (northeastern Anatolia). Therefore, currently
D. mniszechi is divided into three subspecies.
Dorcadion semibrunneum was described by
Pi c (1903) from “Bos-Dagh” as a variety of D.
mniszechi (“D. mniszechi var. semibrunneum”). It
is known from Anatolia only. Br e u n i n g (1962) and
later Öz d i K m e n (2007) have suggested that D. semi-
brunneum is a subspecies of D. mniszechi, while
da n i L e v s K y (2010) treated D. semibrunneum as a
distinct species. Dorcadion anamasum Pic, 1934 was
Survey of the Dorcadion mniszechi Species-Group
(Coleoptera: Cerambycidae) from Turkey, with Descriptions
of Two New Subspecies of Dorcadion semibrunneum Pic, 1903
Hüseyin Özdikmen
Department of Biology, Science Faculty, Gazi University, 06500 Ankara, Turkey. E-mail: ozdikmen@gazi.edu.tr
Abstract: The species of the Dorcadion mniszechi species-group (subgenus Cribridorcadion Pic, 1901) from Tur-
key are surveyed. An identication key using characters of males is presented. Two new subspecies of
Dorcadion semibrunneum Pic, 1903 are described and gured. Dorcadion semibrunneum sivasense ssp.
n. from Karayün Village (Sivas Province) is similar to D. semibrunneum mediocreimpressum Pic, 1909
and D. semibrunneum notatum Pesarini & Sabbadini, 2012. Dorcadion semibrunneum sivrihisarense ssp.
n. from Sivrihisar, Paşakadın Village (SE of Eskişehir Province) is similar to D. semibrunneum notatum
Pesarini & Sabbadini, 2012.
Key words: Coleoptera, Cerambycidae, Dorcadion, Cribridorcadion, new subspecies, Turkey
described from “Pisidia-Taurus: Anamas” (Isparta
Province, Southwestern Anatolia) as a subspecies of
D. mniszechi, i.e. D. mniszechi ssp. anamasum (see
Pi c 1934). da n i L e v s K y (2010) assigned this species
as a subspecies of D. semibrunneum (D. semibrun-
neum anamasum Pic, 1934). However, Pe s a r i n i &
sa B B a d i n i (2012), without studying the type material,
have recognised D. anamasum as a distinct species and
provided a useful identication key to D. mniszechi
and the related taxa, including D. anamasum. In this
paper, we follow La z a r e v (2014) in recognising D.
anamasum as a junior synonym of D. semibrunneum
mediocreimpressum Pic, 1909. According to the same
author, “D. semibrunneum var. medioimpressum Pic,
1910” is a wrong orthography and an unavailable
name published as a synonym by Br e u n i n g (1962)
and da n i L e v s K y (2010).
Pe s a r i n i & sa B B a d i n i (2012) described a sub-
species of D. semibrunneum, i.e. D. semibrunneum
notatum, from Bala (Ankara Province) based on one
male and one female.
In this article, I present a survey of the taxa of
the D. mniszechi species-group from Turkey. I also
describe two new subspecies based on specimens
306
Özdikmen H.
from my collection, D. semibrunneum sivasense ssp.
n. and D. semibrunneum sivrihisarense ssp. n. A key
to the taxa of this group is presented, which is based
on the characters of male specimens.
Material and Methods
The study was based on specimens collected from
Paşakadın Village located in Eskişehir Province and
Karayün Village located in Sivas Province of Turkey in
May 2003 and April 2006, respectively. All specimens
are deposited at Gazi University of Ankara (Turkey).
All species and subspecies of the D. mniszechi
species-group are given alphabetically in the text.
The information regarding the type material for each
species is arranged according to ta v a K i L i a n (2015).
The comparison is based on the original descriptions
and the papers by Pe s a r i n i & sa B B a d i n i (2012) and
La z a r e v (2014).
Results
Genus Dorcadion Dalman, 1817
Subgenus Cribridorcadion Pic, 1901
Diagnosis of the Dorcadion mniszechi species-
group: rst antennal segment without cicatrix; up-
per side of the body almost entirely glabrous (except
sometimes the sutural stripe pubescent) and more or
less shining; elytra black to reddish-brown, entirely
covered with very rough punture or pits, sometimes
with more or less vermiculated surface; male anten-
nae surpassing elytral middle, female antennae not
reaching or hardly surpassing elytral middle; lateral
tubercles of prothorax strongly developed.
I consider that this group includes only two spe-
cies: D. mniszechi Kraatz, 1873 and D. semibrun-
neum Pic, 1903. Dorcadion mniszechi differs from D.
semibrunneum in elytra: with angular humeral costa
well-marked beyond the basal half (elytra with weak
humeral costa that vanishes before the middle in D.
semibrunneum), and in the entirely black elytra (elytra
more or less reddish-brown in D. semibrunneum).
All species group taxa of the D. mniszechi spe-
cies-group can be presented as follows:
Dorcadion mniszechi Kraatz, 1873
The species includes three subspecies.
Dorcadion mniszechi mniszechi Kraatz, 1873
(Fig. 1A)
Type material information: Syntypes ♂ &
♀, ex coll. Musaeo Mniszech, R. Oberthür coll.,
Muséum National d’Histoire Naturelle, Paris [type
locality “Caucasus” (Transcaucasian area from Iğdır
Province in Turkey to the Sevan Lake in Armenia
(La z a r e v 2014)].
Records from Turkey: NE Turkey: Kars
Province: Kağızman, Ağrı Province: Ağrı Mt.,
Erzurum Province (PL a v i L s t s h i K o v 1958); Turkey
(da n i L e v s K y & mi r o s h n i K o v 1985; Lo d o s 1998);
Sivas Province: Karayün Village (Öz d i K m e n
& ha s B e n L i 2004); Anatolia (Öz d i K m e n 2010;
da n i L e v s K y 2010).
Range: Armenia, Turkey.
Chorotype: SW-Asiatic (Anatolo-Caucasian).
Remarks: This subspecies is distributed from
the Transcaucasian area from North-eastern Turkey
to the Sevan Lake in Armenia. Two specimens (one
male and one female) of this species were collected
from the same locality near Karayün Village (Sivas
Province, Central), together with D. semibrunneum
sivasense ssp. n.
Dorcadion mniszechi georgianum Lazarev, 2014
Type material information: Holotype ♂, ex
coll. M. L. Danilevsky, coll. Zoological Institute of
the Russian Academy of Sciences, Saint-Petersbourg
[type locality Gori: Grakali (Georgia)].
Range: Georgia.
Chorotype: Georgian.
Remarks: Presently known only from
Georgia.
Dorcadion mniszechi cavernosum Lazarev, 2014
Type material information: Holotype ♂, ex
coll. M. L. Danilevsky, coll. Zoological Institute of
the Russian Academy of Sciences, Saint-Petersbourg
[type locality Arteni (Armenia)].
Records from Turkey: NE Turkey: Iğdır
Province: Kazkoparan, Kars Province: Kağızman
(Br e u n i n g 1946; La z a r e v 2014).
Range: Armenia, Turkey.
Chorotype: SW-Asiatic (Anatolo-Caucasian).
Remarks: This subspecies is distributed only in
the Transcaucasian area, from North-eastern Turkey
to Arteni in Western Armenia (La z a r e v 2014).
Dorcadion semibrunneum Pic, 1903
The species includes ve subspecies.
Dorcadion semibrunneum semibrunneum Pic,
1903
Dorcadion mniszechi var. semibrunneum Pic,
1903: 170
Type material information: Syntype, ex coll.
M. Pic, coll. Muséum National d’Histoire Naturelle,
Paris [type locality “Bos-Dagh”]. There are sev-
eral localities with this name in Turkey; according
to Br e u n i n g (1962), “Bos-Dagh” is in the İzmir
Province. Pesarini & Sabbadini (2012), based on in-
direct evidence, considered that the type locality was
in the Eskişehir Province. I agree with the approach
of Pe s a r i n i & sa B B a d i n i (2012: 63) since Pic (1903)
stated “type received from M. von Bodemeyer” in
the original description. Bodemeyer (1906) clearly
stated: “D. mniszechi var. nov. semibrunneum Pic…
Survey of the Dorcadion mniszechi Species-Group (Coleoptera: Cerambycidae) from Turkey, with Descriptions...
307
Bos-D. bei Eski-Chehir”. Thus the type locality of the
species should be the Eskişehir Province (Bozdağ).
Records from Turkey: Eskişehir Province:
Bozdağ (Pi c 1903; Bo d e m e y e r 1906; Br e u n i n g 1948,
1962); Anatolia (au r i v i L L i u s 1921; Wi n K L e r 1924-
1932; Öz d i K m e n 2010); Eskişehir Province: Bozdağ,
1 km S of Karatepe (Pe s a r i n i & sa B B a d i n i 2012).
Range: Turkey.
Chorotype: Anatolian.
Remarks: This subspecies is distributed only in
the north-west part of Eskişehir Province (Sündiken
Mts.) in Turkey. It is endemic to Turkey.
Dorcadion semibrunneum mediocreimpressum
Pic, 1909
Dorcadion semibrunneum var. mediocreim-
pressum Pic, 1909: 123.
Type material information: Holotype ♀,
ex coll. Leo Weirather (Innsbrück), M. Pic coll.,
Muséum National d’Histoire Naturelle, Paris [type
locality “Anamas Gbg.”, Turkey: Isparta, Dedegöl
Mt. according to label data, see La z a r e v (2014)].
Dorcadion semibrunneum var. medioimpres-
sum Pic, 1910: 5 [wrong subsequent spelling, una-
vailable name]
Dorcadion mniszechi ssp. anamasum Pic, 1934:
33
Type material information: Syntypes ♀♀,
ex coll. Leo Weirather (Innsbrück), M. Pic coll.,
Muséum National d’Histoire Naturelle, Paris (type
locality “Pisidia-Taurus: Anamas”, Turkey: Isparta:
Dedegöl Mt.).
Records from Turkey: Pisidia Taurus: Isparta
Province: Aksu, Anamas Mts. (Pi c 1934; Br e u n i n g
1962); Anatolia (Öz d i K m e n 2010; da n i L e v s K y
2010; Pe s a r i n i & sa B B a d i n i 2012); Afyonkarahisar
Province: Çakmaktepe pass, Isparta Province: Barla
Mt. (La z a r e v 2014).
Range: Turkey.
Chorotype: Anatolian.
Remarks: This subspecies is distributed
only in Southwestern Anatolia, from Isparta to
Afyonkarahisar. It is endemic to Turkey.
Dorcadion semibrunneum notatum Pesarini &
Sabbadini, 2012
Dorcadion semibrunneum ssp. notatum Pesarini
& Sabbadini, 2012: 62.
Type material information: Holotype ♂, coll.
Carlo Pesarini & Andrea Sabbadini, Milano (type lo-
cality “Bala”, Turkey: Ankara).
Records from Turkey: Ankara Province: Bala
(Pe s a r i n i & sa B B a d i n i 2012).
Range: Turkey.
Chorotype: Anatolian.
Remarks: This subspecies is distributed only
in the south-east part of Ankara Province, north of
Central Anatolian region of Turkey. It is endemic to
Turkey.
Dorcadion semibrunneum sivasense ssp. n. (Fig.
1B)
Type material: Holotype ♂: Turkey: Sivas
Province: Central: Karayün Village, 19.V.2003,
1440 m, leg. H. Özdikmen. Condition of the holo-
type: without front tarsi and right antenna. Paratype
♂: The locality the same as that of the holotype. The
type specimens were deposited at Gazi University in
Ankara, Turkey, coll. Hüseyin Özdikmen.
Description: Male (holotype): Length 19 mm,
width 6 mm. Tegument of body and appendages black
with reddish hues, elytra dark reddish brown and not
glossy. Head on frons and vertex completely glabrous
and not glossy, with ne longitudinal median furrow.
Punctuation on frons sparse, very ne, but recognis-
able, on vertex (only narrow part in front of anterior
margin of pronotum) and especially on temples much
denser and larger. Antennae with golden yellow hairs
and rather dense, ne punctuation. Pronotum almost
completely glabrous and only moderately glossy,
much wider than long (1.33 times as wide as long),
slightly and evenly convex, surface smooth, with ob-
tuse lateral tubercles. Pronotal punctuation sparse,
very ne but recognisable and of similar size to that
on frons, much sparser and much ner than that of
vertex. Scutellum dark reddish brown, smooth and
not glossy. Elytra without well-developed humeri,
with humeral costae limited to basal portion only,
elongated, twice as long as broad, with more or less
vermiculated surface only on disc, and with sparsely,
more or less well-spaced points, and without punc-
tuation between the points. Punctuated parts on elytra
with small circular black spots everywhere. Elytra al-
most completely glabrous, except for a sutural band
of golden-yellow hairs. Epipleura clothed with gold-
en-yellow hairs. Elytral apex rounded. Pygidium vis-
ible, dark reddish brown. Legs (especially tibiae and
tarsi) clothed with dense golden-yellow pubescence.
Abdomen dark reddish brown, sides of the segments
clothed with rather dense, recumbent, short, golden-
yellow hairs, middle looks abraded.
Female: Unknown.
Variability: Paratype identical to the holotype.
Body length of paratype 18.8 mm.
Diagnosis: The new subspecies similar to D.
semibrunneum mediocreimpressum and D. semib-
runneum notatum in terms of presence of a sutural
band on elytra. Easily distinguished from D. semi-
brunneum mediocreimpressum by a sutural band
of golden-yellow hairs on elytra (sutural band with
whitish hairs in D. semibrunneum mediocreimpres-
sum), punctuated parts on elytra with small, circular
and black spots almost everywhere (punctuated parts
308
Özdikmen H.
on elytra without black spots in D. semibrunneum
mediocreimpressum), longer body (smaller in D.
semibrunneum mediocreimpressum). Differs from
D. semibrunneum notatum by a moderately glossy
pronotal surface (pronotal surface very glossy in D.
semibrunneum notatum), punctuated parts on elytra
with small, circular and black spots almost every-
where (punctuated parts on elytra without black
spots in D. semibrunneum notatum).
The new subspecies also differs from D. semib-
runneum semibrunneum by a sutural band of golden
yellow hairs (sutural band completely absent in D.
semibrunneum semibrunneum); less vermiculated
elytral surface on basal half, the points more or less
well-spaced (elytral surface vermiculated on basal
half, the points not distinctly isolated in D. semib-
runneum semibrunneum); very ne and hardly vis-
ible pronotal punctuation on disc (very ne but ap-
preciable pronotal punctuation on disc in D. semib-
runneum semibrunneum); punctuated parts on elytra
with small, circular and black spots almost every-
where (punctuated parts on elytra without black
spots in D. semibrunneum semibrunneum), relative-
ly longer body.
Etymology: From the type locality, the Sivas
Province (Turkey).
Range: Turkey.
Chorotype: Anatolian.
Remarks: This subspecies is only known from
the type locality situated in Sivas Province in Turkey.
It is endemic to Turkey.
Dorcadion semibrunneum sivrihisarense ssp. n.
(Fig. 1C)
Type material: Holotype ♂: Turkey, Eskişehir
Province: Sivrihisar: Paşakadın Village, 13.IV.2006,
leg. H. Özdikmen. Condition of the holotype:
without left front tarsus and left antenna. Paratype
♂: The same as holotype. The specimens were de-
posited at Gazi University in Ankara, Turkey, coll.
Hüseyin Özdikmen.
Description: Male (holotype): Length 21
mm, width 7 mm. Tegument of body and append-
ages black with dark reddish hues, elytra dark red-
dish brown and glossy. Head on frons and vertex
completely glabrous and rather glossy, with a ne
longitudinal median furrow. Punctuation of head on
frons rather dense, ne, but clear, on vertex and espe-
cially on temples much denser and larger. Antennae
with black and rather dense hairs, ne punctuation.
Pronotum almost completely glabrous and very
glossy, much wider than long (1.33 times as wide
as long), slightly and evenly convex, surface smooth
and glossy; with obtuse lateral tubercles. Pronotal
punctuation rather dense, ne but clear, sparser and
ner than that of vertex. Scutellum black, smooth
and glossy. Elytra without well-developed humeri,
with humeral costae limited to basal portion only,
elongated, twice as long as broad, with more or less
vermiculated surface, with sparse, foveiform, ro-
bust and glossy punctuation, and without sculpture
between foveiform points. Elytra with complete
sutural band of black hairs, remaining parts almost
A B C
Fig. 1. A. Dorcadion mniszechi mniszechi from Karayün Village (Turkey: Sivas), B. D. semibrunneum sivasense ssp.
n. (holotype), C. D. semibrunneum sivrihisarense ssp. n. (holotype)
Survey of the Dorcadion mniszechi Species-Group (Coleoptera: Cerambycidae) from Turkey, with Descriptions...
309
completely glabrous. Epipleura clothed with black
hairs. Elytral apex rounded. Pygidium slightly vis-
ible, dark reddish brown. Legs (especially tibiae and
tarsi) clothed with dense black pubescence. Abdomen
dark reddish brown, completely clothed with rather
dense, recumbent, short, golden-yellow hairs.
Female: Unknown.
Variability: Paratype more damaged but identi-
cal to the holotype. Body length of paratype 21.2 mm.
Diagnosis: The new subspecies is closely related
to D. semibrunneum notatum but easily distinguished
by a complete sutural band of black hairs (sutural band
of golden hairs that vanishes in the basal portion in D.
semibrunneum notatum); more or less vermiculated
elytral surface on the basal half and more or less well-
spaced points (clearly vermiculated elytral surface on
the basal half and not distinctly isolated points in D.
semibrunneum notatum): legs, especially on tibiae and
tarsi, clothed with dense black pubescence similar to D.
mniszechi (with golden-yellow pubescence in D. semi-
brunneum notatum); punctuation on vertex denser and
larger than that on frons (punctuation on head the same
as almost everywhere in D. semibrunneum notatum),
pronotum 1.33 times as wide as long (pronotum ap-
proximately twice as wide as long in D. semibrunneum
notatum); relatively larger body, 21 mm, (relatively
smaller body, 19 mm, in D. semibrunneum notatum).
The new subspecies differs from D. semibrun-
neum semibrunneum by a complete sutural band of
black hairs (sutural band absent in D. semibrunneum
semibrunneum); pronotal surface between the points
very glossy (pronotal surface between the points at
most moderately glossy in D. semibrunneum semi-
brunneum); legs, especially tibiae and tarsi, clothed
with dense black pubescence (golden-yellow pubes-
cence in D. semibrunneum semibrunneum), prono-
tum 1.33 times as wide as long (pronotum 1.22 times
as wide as long in D. semibrunneum semibrunneum),
larger body, 21 mm, (smaller body, 15-17 mm, in D.
semibrunneum semibrunneum).
The new subspecies differs from D. semibrun-
neum sivasense by a complete sutural band of black
hairs (sutural band of golden-yellow hairs in D.
semibrunneum sivasense); pronotal surface between
the points very glossy (pronotal surface between the
points moderately glossy in D. semibrunneum siva-
sense); antennae and legs, especially tibiae and tarsi,
clothed with dense black pubescence (golden-yellow
pubescence in D. semibrunneum sivasense), punctu-
ated parts on elytra without black spots (punctuated
parts on elytra with small, circular black spots al-
most everywhere in D. semibrunneum sivasense),
relatively longer body, 21 mm (relatively smaller
body, 19 mm, in D. semibrunneum sivasense). The
new subspecies is the largest among the subspecies
of D. semibrunneum (body length 21 mm in male).
In addition, the new subspecies is distributed
only in Sivrihisar Mountains in the south-east part of
Eskişehir Province. On the contrary, D. semibrun-
neum semibrunneum is distributed only in Sündiken
Mountains (Bozdağ) in the north-west part of Eskişehir
Province. Dorcadion semibrunneum notatum is distrib-
uted only in Küre Mountains environs of Bala in the
south-east part of Ankara Province (Fig. 2).
Etymology: From Sivrihisar (Turkey: Eskişehir
Province), where the type locality is situated.
Range: Turkey.
Chorotype: Anatolian.
Remarks: This subspecies is distributed only in
the south-east part of Eskişehir Province (Sivrihisar
Mts.) in Turkey. It is endemic to Turkey.
An identication key for D. mniszechi
species-group (males)
1. Elytra with angular humeral costa well-marked
beyond basal half; relatively longer body (about
19-22 mm)................................................................2
-. Elytra with weak humeral costa that van-
ishes before middle; relatively smaller body (about
15-21 mm)...............................................................4
2. Elytra with moderately rough sculpture........
..................................D. mniszechi mniszechi Kraatz
-. Elytra with extremely or very rough sculpture........3
3. Elytra with extremely rough sculpture, with
shallow big conjugated dots........................................
..............................D. mniszechi georgianum Lazarev
-. Elytra with very rough sculpture, punctures deep
and conjugated........D. mniszechi cavernosum Lazarev
4. Pronotal surface only moderately glossy; rela-
tively smaller body (about 15-19 mm).........................5
-. Pronotal surface very glossy; relatively long-
er body (about 19-21 mm)...................................7
5. Elytra without a sutural band; elytral surface
on basal half vermiculated, the points not distinctly
isolated; pronotal punctuation very ne but notice-
ble…………D. semibrunneum semibrunneum Pic
-. Elytra with a sutural band; elytral surface on
basal half more or less vermiculated, the points more
or less well-spaced; pronotal punctuation entirely
absent or very ne and hardly visible on disc…....6
6. Sutural band with whitish hairs; elytral sur-
face rugged, but not vermiculated, the points well-
spaced; punctuated parts on elytra without black
spots..........D. semibrunneum mediocreimpressum Pic
-. Sutural band with golden-yellow hairs; ely-
tral surface more or less vermiculated, the points
more or less well-spaced; punctuated parts on
elytra with small, circular, black spots almost eve-
rywhere…...........D. semibrunneum sivasense ssp. n.
7. Sutural band with golden-yellow hairs; ely-
310
Özdikmen H.
tral surface on basal half clearly vermiculated, points
not distinctly isolated; legs (especially tibiae and tarsi)
clothed with dense golden yellow pubescence; body
19 mm................................................................................
..........D. semibrunneum notatum Pesarini & Sabbadini
-. Sutural band with black hairs; elytral sur-
face on basal half more or less vermiculated, the
points more or less well-spaced; legs (especially tib-
iae and tarsi) clothed with dense black pubescence
........................D. semibrunneum sivrihisarense ssp. n.
Fig. 2. Distribution patterns of Dorcadion semibrunneum semibrunneum (Sündiken Mountains, Bozdağ), D. semibrun-
neum sivrihisarense ssp. n. (Sivrihisar Mountains, Paşakadın village marked with a dot) and D. semibrunneum notatum
(Küre Mountains, env. Bala)
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Accepted: 13.05.2016
