Content uploaded by Bernd Jaeger
Author content
All content in this area was uploaded by Bernd Jaeger on Feb 17, 2017
Content may be subject to copyright.
273
VERNATE 31/2012
Revision of the immaculate species of the Anthracus annamensis group from the East
Palaearctic and Oriental Regions (Insecta: Coleoptera: Carabidae: Harpalini)
Bernd Jaeger
S. 273
-
308
Summary
The Anthracus annamensis group is introduced and
characterized. A revision of the immaculate spe-
cies of this species group from the Oriental and East
Palaearctic Regions is presented. Acupalpus deroga-
tus Walker, 1858 and Acupalpus glabrus Louwerens,
1956 are transferred to the genus Anthracus (comb.
nov.). Anthracus derogatus (Walker, 1858), furvus
(Andrewes, 1947) and A. glabrus (Louwerens, 1956)
are redescribed. Four new species are described:
Anthracus wrasei nov. sp. from China (type locality:
China: Yunnan: 4 km E Dali old town, shore terrain of
Er Hai Lake, 2020 m, 25°42’N /100°11.5’E), A. schuhi
nov. sp. and A. indicus nov. sp. from India (type locali-
ty: India: Maharashtra: Lonvala, 80 km E Bombay) and
A. latus nov. sp. from Thailand, Laos, Vietnam, China
and Myanmar (type locality: Thailand: Chiang Mai
prov.: Ban San Pakia, 1700 m). New distributional data
are provided for Anthracus furvus (Andrewes) (rst
records for Vietnam and China, Yunnan prov.), and for
A. glabrus (Louwerens) (rst record for Bali).
Zusammenfassung
Revision der einfarbigen Arten der Anthracus
annamensis-Gruppe aus der Ostpaläarktis und
der orientalischen Region (Insecta: Coleoptera:
Carabidae: Harpalini)
Die Anthracus annamensis-Gruppe wird vorgestellt
und charakterisiert. Die durch einfarbige Flügeldecken
ausgezeichneten, orientalischen und ostpaläarktischen
Arten der Artengruppe werden revidiert. Für Anthracus
furvus (Andrewes, 1947), derogatus (Walker, 1858) und
glabrus (Louwerens, 1952) werden Redeskriptionen
vorgelegt, wobei die beiden letztgenannten Arten in
die Gattung Anthracus umgesetzt werden. Vier neue
Arten werden beschrieben: A. wrasei nov. sp. (loc. typ.:
China: Yunnan: 4 km E Dali old town, shore terrain of
Er Hai Lake, 2020 m, 25°42’N /100°11.5’E) aus China,
A. schuhi nov. sp. (loc. typ.: Indien: Maharasthra:
Lonvala, 80 km E Bombay) und A. indicus nov. sp.
(loc. typ.: Indien: Maharasthra: Lonvala, 80 km E
Bombay) aus Indien und A. latus nov. sp. (loc. typ.:
Thailand: Chiang Mai prov.: Ban San Pakia, 1700 m”)
aus Thailand, Laos, Vietnam, China und Myanmar.
Neue Verbreitungsdaten werden für Anthracus fur-
vus (Andrewes) (Erstnachweise für Vietnam und
China, Yunnan prov.) und A. glabrus (Louwerens)
(Erstnachweis für Bali) vorgelegt.
Key words: Acupalpus, redescriptions, new species,
rst records, India, China, South East Asia
Introduction
The closely related Stenolophina genera Acupalpus La-
treille, 1829 and Anthracus Motschulsky, 1850 comprise
16 species which were described or recorded from the
Oriental region, excluding Acupalpus ovatulus Bates,
1889 that actually belongs to the genus Stenolophus (Ka-
taev pers. communication) and A. assamicus Jedlička,
1964 that represents a species of the subtribe Tachyina
of the tribe Bembidiini according to an examination of
the holotype.
Six of the Oriental species, Anthracus annamensis
(Bates, 1889) and furvus (Andrewes, 1947), Acupalpus
derogatus Walker, 1858, nesophilus Andrewes, 1936,
haemorrhous Louwerens, 1952 and glabrus Louwerens,
1952, and additionally Anthracus biplagiatus (Boheman,
1858) from E China and furvinus (Darlington, 1968)
from New Guinea belong to a complex of closely related
species recently mentioned as the “Acupalpus annamen-
sis group” by Jaeger (2010: 147) and here introduced as
a peculiar species group of the genus Anthracus.
Until now most of the Oriental taxa of this species
group were known only from their original descrip-
tions and subsequent records in world and regional
catalogues, in collection lists or other faunistic papers
274
(e.g. Bates 1892, andrewes 1926, 1928, 1930, 1933,
1947, Csiki 1932, Landin 1954, Jedlička & ChûJô
1964, 1966, Lorenz 1998, 2005, Jaeger & kataev
2003, Chaudari 2005). Redescriptions or taxonomic
notes were provided only for A. derogatus (Walker) by
Bates (1886) and andrewes (1919). A key, or modern
redescriptions characterising and guring the essentials
characters of the male aedoeagi of the formerly de-
scribed species are missing to date. Moreover, the study
of older and recent collections from India and SE Asia
revealed that the A. annamensis group comprises vari-
ous undescribed species externally often very close to
formerly described taxa. Thus, a reliable identication
of the species was impossible without type compari-
son, and consequently most former faunistic records
remained doubtful.
This rst contribution on the taxa of the Anthracus
annamensis group provides a general diagnosis of the
species group, a revision of the immaculate species,
containing redescriptions of A. derogatus Walker, A.
furvus Andrewes and glabrus Louwerens, as well as
descriptions of four new species from China, India and
SE Asia, and an identication key. The species with
maculate elytra, related to A. annamensis (Bates) and
nesophilus (Andrewes) will be treated in future papers
(Jaeger in prep.).
Material and Methods
The material examined is deposited in the following
public institutions and private collections:
BMNH Natural History Museum, London, Great
Britain, Chr. Taylor, Cl. Gent and B. Garner
IZAS Institute of Zoology, Chinese Academy of
Sciences, Beijing, China, Dr. Meiying Lin
MHNG Museum d´Histoire Naturelle, Geneva, Swit-
zerland, Dr. I. Löbl and Dr. G. Cuccudoro
NHMB Naturhistorisches Museum Basel, Bale,
Switzerland, Dr. M. Brancucci
NHMW Naturhistorisches Museum Wien, Vienna,
Austria, Dr. H. Schönmann and Dr. H.
Schillhammer
NHRS Swedish Museum of Natural History, Stock-
holm, Sweden, Dr. B. Viklund and Dr. J. Ferrer
NME Naturkundemuseum Erfurt, Germany, Dipl.-
Biol. M. Hartmann
NMP Narodny Muzeum v Praze, Prague, Czech
Republic, Dr. J. Hájek
RMNH Nationaal Naturhistorische Museum „Natu-
ralis“, Leiden, The Netherlands, F. van Assen
ZISP Zoological Institute, Russian Academy of Sci-
ences, St. Petersburg, Russia, Dr. B. M. Kataev
cBUL Coll. P. Bulirsch, Prague, Czech Republic
cJAE Coll. B. Jaeger, Berlin, Germany
cKME Coll. R. Kmeco, Litovel, Czech Republic
cSCHM Coll. J. Schmidt, Admannshagen, Germany
cWEI Coll. A. Weigel, Wernburg, Germany
cWRA Coll. D.W. Wrase, Berlin, Germany
Label data of examined type material are cited in full
using a “/” to separate different lines of the label in
question. If not otherwise stated labels are printed in
black on white paper. Collection data of revised non-
type material are presented according to the following
form: “locality, month. year, collector(s) (number of
specimens
-
deposition)”.
Measurements, as dened in Jaeger (2009: 1636),
were taken at magnications of 30 times (body length,
elytra and pronotum partly) or 70 times (head and prono-
tum) using an ocular micrometer attached to a stereomi-
croscope Nikon SMZ 1500.
The following abbreviations were used in the text and/
or in the legends and in Table 1: BL
-
body length, HW
-
width of the head including eyes, HWbE
-
width
of head between inner margins of eyes, PL
-
length
of pronotum, PW
-
width of pronotum at its broadest
point, EL
-
length of elytra, EW
-
width of elytra, HT
-
Holotype, PT
-
Paratype, hw
-
handwritten.
Microsculpture was examined at magnication of 100
times.
Dissections were made, using standard techniques;
genitalia and mouthparts were preserved in Euparal on
acetate strips and pinned beneath the specimens from
which they had been removed.
Descriptions/redescriptions do not include general
characters of the genus and do not repeat features al-
ready mentioned in the diagnosis of the species group,
except when these characters show distinct modica-
tions or variation between different species.
Photographs of habitus, head and pronotum were tak-
en using a Canon EOS 60d DSLR camera tted with
a Promicron photo adapter attached to a Nikon SMZ
1500 stereomicroscope. Photographs of male and fe-
275
male genitalia, and mouthparts were taken using the
above-mentioned equipment attached to a Leica DM
LB2 transmitted light microscope. In both cases sev-
eral single images from different focal planes were
taken, which were later combined, using the Combine
ZP (©Alan Hadley) and Auto-Montage Essentials
(©Synoptics Ltd) software. The head and pronotum of
mounted specimens are often distinctly bent downward
or aligned to the left or right of centre and therefore not
at the same focal plane as the elytra. If so, separate im-
ages of each body part were produced, which were sub-
sequently combined with Adobe Photoshop software.
Thus, the nal images presented here, may often not
reect the original mounting condition.
The distribution maps were generated using the online
mapping software SimpleMappr (©David P. Shorthouse).
The Anthracus annamensis group
Diagnosis: General appearance Acupalpus-like, with
pronotal sides usually roundly or rectilinearly narrowed
posteriad, rarely very faintly sinuate, and posterior an-
gles usually widely rounded, or sometimes an indistinct
obtuse angle is suggested. Antennomere 2 sparsely pu-
bescent. Ligula with two long ventral setae inserted
subapically and two additional shorter setae inserted
dorso-laterally just posteriad of ventral ones (Figs 66
-
70). Mentum and submentum fused, at least laterally,
only at middle divided by diffuse edge. Prosternum
medially with 4
-
6 biseriately arranged medium-long
setae and just posterior to apical margin with a row of
5
-
10 medium-long setae. Prosternal process with 1
-
3
distinct setae (often broken, and then insertion points
not or only hardly visible). Abdominal sternites 4
-
6 in
most species with rather dense and long pubescence.
Protarsomeres 2
-
4 of males moderately to markedly
dilated and with distinct biseriately arranged adhesive
hairs beneath, protarsomere 4 often bilobed. Mesotar-
someres 2
-
4 of males weakly to moderately dilated,
segment 1
-
4 or 2
-
4 with biseriately arranged adhesive
hairs beneath. Tarsomere 5 of pro-, meso- and metatar-
someres without pairs of ventrally arranged setae.
Remarks: Members of the Anthracus annamensis group
share a number of distinctive characters, such as men-
tum and submentum fused, prosternum medially with
longer, erect setae, and tarsomere 5 without ventro-
laterally arranged pairs of setae, with the Asian Acupal-
pus subgenera Subacupalpus Habu, Pseudanthracus
Habu, Palcuapus Habu, Setacupalpus Habu and the
genus Anthracus. Though their general appearance re-
sembles typical species of the genus Acupalpus, mem-
bers of the A. annamensis group are obviously closer
related to the genus Anthracus than to the previously
mentioned Acupalpus subgenera, because they share
another important character, the peculiar chaetotaxy
of the ligula, which is quadrisetose in Anthracus and
the A. annamensis group instead of being bisetose in
Acupalpus and most other Stenolophina, except the ge-
nus Parabradycellus Ito. The latter genus has doubtless
no closer relations the Acupalpus/Anthracus lineage
because it differs in various important features from the
mentioned genera and occupies an outstanding position
within the subtribe.
Though the modied ligular chaetotaxy has been
evolved twice in Stenolophina it constitutes obviously
an apomorphic character state, which can be hypothe-
sized to be the autapomorphy for a monophyletic group
comprising the genus Anthracus and the A. annamensis
group. Taking this hypothesis in to consideration and
following SChauberger (1930), Csiki (1932) and
Lorenz (1998, 2005) who already placed or listed a
part of the species in the subgenus, respectively genus
Anthracus, the A. annamensis group is here regarded a
distinctive species group within the genus Anthracus. A
formal taxonomic rank is not proposed here because a
phylogenetic analysis of Acupalpus and related genera on
a worldwide base is needed to clarify the phylogenetic
relations within this genus group and from these results
the nal taxonomic rank of the involved supraspecic
taxa should be determined.
Within the genus Anthracus members of the annamensis
group differ from typical western Palaearctic species
by the different general appearance with shorter anten-
nae, less elongate elytra, and pronotal sides without or
with only a weak sinuation before the posterior angles,
which are usually no more than suggested. From members
of the Anthracus angusticollis group which comprises
ve species from Ethiopian, Palaearctic and Oriental
regions, they differ additionally by the unmodied me-
dian line of the pronotum and the non-serrate lateral
margin of the elytra.
Included Taxa: The Anthracus annamensis group
comprises 11 species (including those described here)
from the East Palaearctic and the Oriental Regions, A.
276
furvinus (Darlington, 1968) from New Guinea and an
unknown number of species described from the Ethio-
pian Region that are still placed in the genus Acupalpus.
Revision of the immaculate species of the A. anna-
mensis group
Anthracus derogatus (Walker, 1858)
(Figs 1
-
3, 22
-
24, 78)
Acupalpus derogatus Walker, 1858: 204 (type locality:
“Ceylon”).
Acupalpus derogatus Walker: Bates 1886: 80 [descrip-
tive notes & faunistic record], andrewes 1919: 190 [re-
description]; andrewes 1928: 138 [catalogue Ceylon].
Acupalpus (s.str.) derogatus Walker: Csiki 1932: 1244
[world catalogue], Lorenz 1998: 337 [world cata-
logue], Lorenz 2005: 359 [world catalogue]
Type material: Holotype: 1 % (BMNH) labelled
[Fig 3] “Type / HT” (circular label with red margin);
“Ceylon” (circular label, hw); “Acupalpus / deroga-
tus Walk / A. N. Hist (Type)” (hw); “derogatus” (hw);
“Acupalpus / derogatus / stet Wlk / Teste Bates” [hw]
and my label “Anthracus /derogatus / (Walker, 1858) /
det. B. Jaeger 2012”.
Remarks: The holotype is in fair condition. It was origi-
nally pinned and later glued on a small paper strip. The
right anterior leg, the left antennomeres 3
-
11 are miss-
ing, and the right elytron was somewhat damaged by
former pinning. I dissected the aedoeagus and mounted
the specimen on a new card.
Redescription: General appearance as gured (Fig. 1).
Body length 3.1 mm; width 1.2 mm.
Head blackish brown with clypeus, labrum and man-
dibles (inner margins and apices dark brown) reddish.
Pronotum dark reddish brown with lateral margins pal-
er. Elytra blackish brown, leaving rst interval, base,
lateral margins and the apex (vaguely) paler reddish
brown. Legs and palpi, and antennae entirely yellow-
ish to pale reddish brown. Ventral surface dark reddish
brown, with pro- and metepisterna somewhat darker,
epipleura brown. Upper surface shiny, elytra slightly
iridescent.
Head (Figs 1
-
2) including eyes 0.80 times as wide as
pronotum, with eyes only weakly prominent (head 1.46
times as wide as head between eyes). Labrum almost
rectilinear at apical margin. Mandibles medium sized,
not markedly prolonged, left mandible faintly obtuse
at apex. Antennae 2.35 times as long as pronotum and
0.89 times as long as elytra. Microsculpture on labrum
and clypeus distinctly isodiametric, on anterior half
of head almost obliterated, only traces of isodiametric
meshes are visible, on posterior half faintly isodiametric
to short transverse.
Pronotum (Figs 1
-
2) 1.31 times as wide as long, 1.25
times as wide as head, widest in second quarter, lateral
seta inserted a little posterior to beginning of second
quarter. Apical margin faintly emarginated, with lateral
border indistinct and widely interrupted, not prolonged
upon lateral quarters. Anterior angles narrowly rounded
at tips, not projecting forward. Sides convex in anterior
half, markedly narrowed to posterior angles, the latter
obtuse. Basal margin almost rectilinear medially, ob-
liquely produced to posterior angles. Lateral furrows
evenly narrow in apical two thirds, markedly widened
at basal third, where they are fused with the latero-basal
impressions. Latero-basal impressions rather small,
distinctly delimited from pronotal disc and median part
of base, fused with basal and lateral margin. Basal im-
pressions and other surface of pronotum impunctate.
Median line ne on disc, disappearing before reaching
basal and apical margins. Anterior transverse impres-
sion only suggested. Microsculpture on disc with very
faintly impressed, shorter to longer transverse meshes,
which become more distinct at medial part of base. Latero-
basal impressions and lateral furrows with course iso-
diametric to short transverse meshes.
Elytra (Fig. 1) with sides moderately widened poste-
riorly, widest just at or just posterior to middle, 1.52
times as long as wide, 2.64 times as long and 1.32 times
as wide as pronotum. Elytral striae distinctly impressed
and impunctate, scutellar striole rather long. Intervals
weakly convex, narrowed and moderately convex
at apex. Basal pore at beginning of scutellar striole
present, interval 3 in third quarter with one setiferous
pore, adjoining stria 2. Microsculpture on scutellum
isodiametric, on elytral intervals with indistinct, very
lightly impressed transverse lines. Macropterous.
Metepisterna long and narrowed posteriad, at inner mar-
gin about 1.5 times longer than wide at basal margin.
Prosternum medially with 7 and posterior to apical
margin with about 5 longer seta. Setae at prosternal
277
process missing, but probably present and later broken.
Abdominal sternites 4
-
6 with distinct pubescence, in
sternites 4 and 5 laterally somewhat reduced. Last visible
abdominal sternite of the male holotype very faintly emar-
ginate at apex, at apical margin with two longer setae.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 moderately bilobed. Mesotar-
someres 2
-
4 weakly dilated and with adhesive hairs on
ventral surface, mesotarsomere 4 somewhat emarginated
but not clearly bilobed.
Median lobe of aedoeagus (Figs 22
-
24) rather small
with general appearance and structures of internal sac
as gured. The apical portion (in lateral view) is rather
distinctly bent ventrally. The apex is somewhat thick-
ened but without any distinct hook. The internal sac
(lateral aspect) has one larger and one medium sized
tooth in apical half, two medium sized teeth in upper
basal half and 3 smaller spines in lower basal half.
Comparisons: A. derogatus can be separated easily from
other immaculate species of the A. annamensis group by
the combination of following characters: small size, head
rather wide between eyes, the latter rather at, pronotum
markedly narrowed posteriorly, and specic size, shape
and inner structures of median lobe of aedoeagus.
Distribution: A. derogatus is so far known only from
Sri Lanka (waLker 1858, Bates 1886: 80). However,
because the species is macropterous it is obviously
more widely distributed and occurs probably also in
southern India.
Anthracus furvus (Andrewes, 1947)
(Figs 4
-
6, 25
-
35, 66, 71, 76)
Acupalpus furvus Andrewes, 1947: 8, 17
-
18, plate 1,
g 5 (type locality: Myanmar: Kambaiti, 7000 ft).
Acupalpus furvus Andrewes: Landin 1954: 457
[faunistics].
Anthracus furvus (Andrewes): Lorenz 1998: 339
[world catalogue], 2005: 360 [world catalogue].
Type material: Holotype: 1 % (NHRS) labelled
“N.E. BURMA / Kambaiti, 7000 ft. / 12/4 1934 / R.
MALAISE” [date hw], “Typus“ [red label], “Acupalpus
/ furvus / Type Andr. / H.E. Andrewes det.” [rst three
lines hw Andrewes], “Acupalpus / (Palcuapus) / furvius
[Sic!]/ Andrewes / Det. N. ITO 1994” [rst three lines
hw Ito], “6384 / E91”, “Naturhistoriska / Riksmuseet
/ Stockholm / Loan no 465/93”, “NRM Sthlm / Loan
1105/09” and “Acupalpus / furvus / Andrewes, 1947 /
det. B. Jaeger 2011”.
Paratypes: 1 % (NHRS) labelled “N.E. BURMA /
Kambaiti, 2000 m / 20/3 1934 Malaise”, “Paratypus“
[red label], “furvus / Andr.”, “6385 / E91”, “NRM Sthlm
/ Loan 1106/09” and “Acupalpus / furvus / Andrewes,
1947 / det. B. Jaeger 2011”. 1 % (BMNH) labelled
“Para- / type” [circular label with yellow margin],
“N.E. BURMA / Kambaiti, 7000 ft / 12
-
15/4 1934 / R.
Malaise” [date hw], “Acupalpus / furvus / cotype Andr.
/ H.E. Andrewes det.” [rst three lines hw Andrewes]. 1
& (BMNH) labelled “Para- / type” [circular label with
yellow margin], “N.E. BURMA / Kambaiti, 7000 ft /
28/3 1934 / R. Malaise” [date hw], “Acupalpus / furvus
/ cotype Andr. / H.E. Andrewes det.” [rst three lines
hw Andrewes]. 1 & (BMNH) labelled “N.E. BURMA
/ Kambaiti, 7000 ft / 22/3 1934 / R. Malaise” [date
hw], “Co- / type” [circular label with green margin],
“Acupalpus / furvus / cotype Andr. / H.E. Andrewes det.”
[rst three lines hw Andrewes], “H.E. Andrewes Coll.
/ B.M. 1945
-
97”. All paratypes from BMNH with my
label “Anthracus / furvus / (Andrewes, 1947) / det. B.
Jaeger 2012”Remarks: andrewes (1947: 17
-
18) did
not designate a type (in sense of a holotype) in the text
part of the description. However, according to the legend
of the plate 1 the image no 5 is that of the “type” which I
accept as the valid holotype designation.
The holotype has been remounted and dissected by N.
Ito and is now in poor condition compared with the
image given by Andrewes. It has been “embedded” in
glue, the mandibles were broken off, and parts of anten-
nae and tarsi are missing. The aedoeagus was embed-
ded on a separate card, with mounting medium turbid,
but so far visible the internal sac was everted.
The paratypes are in perfect or fair condition. Some of
them have missing antennomeres, tarsomeres or legs.
Redescription: Remark: measurements and indices giv-
en below are based on populations from the type locality
Kambaiti (near the border to Yunnan) and from Yunnan.
General appearance as gured (Fig. 4). Body length
3.4
-
4.1 mm; width 1.3
-
1.6 mm.
Dorsal surface mainly dark to blackish brown, leav-
ing clypeus dark reddish, and labrum, mandibles (in-
ner margins and apices blackish), pronotal and elytral
lateral margins and rst interval of elytra paler reddish
278
brown. Head posterior to eyes sometimes indistinctly
reddish brown. Legs and palpi pale or dark yellowish
brown, antennae brown to dark brown, with rst two,
rarely with only rst antennomere paler. Ventral surface
mainly dark brown, with gula, pro-, meso and metaster-
num paler, sometimes also posterior half of abdominal
sternites somewhat lightened. Upper surface shiny,
elytra with faint iridescence.
Head (Figs 4
-
6) including eyes 0.75
-
0.83 times as
wide as pronotum, with eyes moderately prominent
(head 1.45
-
1.58 times as wide as head between eyes).
Labrum almost rectilinear at apical margin. Mandibles
medium sized, not distinctly prolonged and curved, left
mandible faintly obtuse at apex. Antennae 2.19
-
2.39
times as long as pronotum and 0.77
-
0.85 times as long
as elytra. Microsculpture on labrum and clypeus dis-
tinctly isodiametric to short transverse, on anterior half
of head almost obliterated, only traces of isodiametric
meshes are visible, on posterior half faintly isodiametric.
Pronotum (Figs 4
-
6) 1.25
-
1.37 times as wide as long,
1.21
-
1.33 times as wide as head, widest in second
quarter, lateral seta inserted a little posterior to begin-
ning of second quarter. Apical margin almost rectilinear
or faintly emarginate, lateral border interrupted, reach-
ing medial third. Anterior angles narrowly rounded
at tips, not or only weakly projecting forward. Sides
convex in anterior half, rectilinearly narrowed to the
widely rounded posterior angles. Basal margin weakly
arcuate medially, obliquely produced to posterior
angles. Lateral furrows evenly narrow, sometimes
weakly widened posteriorly, where they are fused with
the latero-basal impressions. Latero-basal impressions
rather small, distinctly delimited from pronotal disc and
median part of base, fused with basal and lateral mar-
gin, or separated from the latter by a small convexity.
Basal impressions and other surface of pronotum im-
punctate. Median line ne on disc, disappearing before
reaching basal and apical margins. Anterior transverse
impression obliterated or suggested. Microsculpture
on disc with faintly impressed, short to long transverse
meshes, which become more distinct at medial part of
base. Latero-basal impressions and lateral furrows with
course isodiametric to short transverse meshes.
Elytra (Fig. 4) with sides moderately widened posteri-
orly, widest just at or just posterior to middle, 1.55
-
1.64
times as long as wide, 2.73
-
2.90 times as long and
1.30
-
1.41 times as wide as pronotum. Elytral striae
distinctly impressed and impunctate, scutellar striole
long. Intervals rather at, narrowed and very weakly con-
vex at apex. Basal pore at beginning of scutellar striole
present, interval 3 in third quarter with one setiferous
pore, adjoining stria 2. Microsculpture on scutellum
isodiametric, on elytral intervals with very lightly
impressed transverse lines. Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.5 times longer than wide at basal margin.
Prosternum medially with 5
-
6 and posterior to apical
margin with a row of 8 longer setae. Prosternal process
with at least 1 distinct seta (very often broken). Abdomi-
nal sternites 4
-
6 with distinct pubescence. Last visible
abdominal sternite of males and females very faintly
emarginate at apex, with two setae at apical margin in
males and four in females.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 markedly bilobed. Mesotar-
someres 1
-
4 moderately dilated and with adhesive hairs
on ventral surface, mesotarsomere 4 moderately bilobed.
In females pro- and mesotarsomeres unmodied.
Median lobe of aedoeagus (Figs 25
-
35) with general
appearance and structures of internal sac as gured.
Apex in lateral aspect with a small indistinct hook.
Internal sac (lateral aspect) composed of one smaller
and one larger apical tooth, two larger subapical tooth,
with one of them usually orientated ventrad and the
other dorsad, a row of 5
-
8 dorsally arranged medium
sized teeth (the last tooth usually orientated dorsad) and
sometimes 1
-
3 smaller ventrally arranged teeth. More
rarely, in specimens from S Vietnam, 12 km N Dalat
(Figs 34
-
35) and single specimens from Yunnan (Fig
33), both subapical teeth are orientated ventrad.
Female genitalia as gured (Fig. 71) without signicant
specic characters.
Variation: Populations from Myanmar, Yunnan and N
Vietnam shows a normal individual variation in exter-
nal features, as well as in shape and inner structures of
the median lobe of the aedoeagus. In contrast to those, a
small series of somewhat teneral specimens from S Viet-
nam (N Dalat) differs externally, as well as in inner struc-
tures of the aedoeagus. The specimens are larger on aver-
age (3.9
-
4.6 mm), differ in some body proportions (e.g.
AL/PL, EL/PL, EL/EW) and in the pronotal shape with
sides more rectilinearly narrowed to base and the lateral
furrows somewhat widened posteriorly. In this respect the
279
%&
S Vietnam specimens are more similar to A. wrasei than
to Yunnan populations of A. furvus. The inner structures
of the aedoeagus of the two males are modied concern-
ing size and orientation of the apical and subapical teeth
(Figs 29, 34
-
35), but the differences are less distinct
than those between A. furvus and wrasei, and addition-
ally there is no discrepancy in shape of the apical hook of
the median lobe. Thus, the S Vietnam population is here
regarded as a geographical form of A. furvus. Additional
material from the same area is necessary to determine the
nal taxonomic position of this population.
Comparisons: A. furvus is in general appearance very
similar to A. wrasei, which were collected together with
the former in the Chinese province Yunnan at the west-
ern shore of the Er Hai Lake and at Xue Shan. It can be
separated easily from this species by the structures of
the internal sac without a ventrally arranged row of dis-
tinct medium sized teeth, and the different shape of the
aedoeagal apex with has only a small hook (Figs 25
-
35).
Externally both species are very similar, but at least the
A. furvus populations from Myanmar, Yunnan and N
Vietnam can be distinguished from A. wrasei by their
smaller size, somewhat shorter elytra and antennae (see
ratios EL/PL, EL/EW, AL/PL, AL/EL), the pronotum
which is at the average shorter and has the lateral fur-
rows not or less distinctly widened in front of posterior
angles, and the head being blackish brown or only in-
distinctly reddish posterior to eyes instead having dis-
tinct reddish spots. In contrast, the A. furvus populations
from S Vietnam and those of A. wrasei from Yunnan
have no signicant external differences, though they dif-
fer markedly in the shape of the apex of the median lobe
and its inner structures (Figs 29, 34, 35 respectively 36
-
41). There are only less valuable, minor external differ-
ences in the shape of pronotum with sides usually less
distinctly narrowed posteriorly. However, according to
present knowledge it remains unsure whether both spe-
cies actually occur sympatrically in S Vietnam.
From A. latus nov. sp. which has a similar distribution
and occurs sympatrically, at least in S and N Vietnam A.
furvus can be separated easily by its different general ap-
pearance (Figs 4, 19), and the shape of the aedoeagal me-
dian lobe and its internal structures (Figs 25
-
35, 59
-
65).
Other similar species occur allopatrically farther west
(A. schuhi) or south (A. glabrus). They differ from A.
furvus by various external and aedoeagal features (for
details see under these species).
Distribution: A. furvus occurs in S Vietnam and more
widely in N Vietnam, W Yunnan and NE Myanmar (Fig.
76). It has been collected at altitudes of 1200
-
2500 m.
In addition to the type material from E Myanmar the
following material has been studied.
Examined Material:
China: Yunnan: Baoshan Pref., Gaoligong Shan,
31 km SE Tengchong, 2200
-
2250 m, 24°53’11”N,
98°45’22”E, (pasture in sec. forest, under moss/shrubs)
[09],VIII. 2009, Wrase (4
-
cWRA); Baoshan Pref.,
Gaoligong Shan, E pass, 36 km SE Tengchong, 2200
m, 24°49’32”N, 98°46’06”E, (farm land, under moss,
grass, shrubs, under stones) [13], VIII. 2009, Wrase (2
-
cWRA); Baoshan Pref., Gaoligong Shan, E pass, 36 km
SE Tengchong, 2230 m, 24°49’32”N, 98°46’06”E, (pas-
ture in prim. decid. forest, under stones/wood) [08], VIII.
2009, Wrase (1
-
cWRA); Baoshan Pref., mount. range
14 km E Tengchong, 1850 m, 25°00’28”N, 98°38’07”E,
(eld edge, compost, debris sifted) [16a],VI. 2007,
Schülke (1
-
cWRA); Baoshan Pref., mount. range 22 km
S Tengchong, 1750 m, 24°49’29”N, 98°29’27”E, (loamy
banks of shponds) [18], VI. 2007, Wrase (7
-
cWRA);
Baoshan Pref., mount. range 22 km S Tengchong, 1750
m, 24°49’29”N, 98°29’27”E, (loamy banks of sh-
ponds) [19A], VIII. 2009, Wrase (1
-
cWRA); Dali Bai
Nat. Auton. Pref., 1 km W Dali old town, creek val-
ley at foothill of Diancang Shan, 2170m, 25°41.9’N,
100°08.4’E, (loamy banks of shponds) [18],VIII.
-
IX.
2003, Wrase (4
-
cWRA, ZISP); Dali Bai Nat. Auton.
Pref., 4 km E Dali old town, shore terrain of Er Hai
Lake, 2020 m, 25°42’N, 100°11.5’E, young willows
with knotgrass, under vegetation/in roots [16],VIII.
-
IX.
2003, Wrase (8
-
cWRA, ZISP); Shore of Erhai Hu lake,
1970 m, 25°42.4’N/100°12.1’E, (elds close to shore,
under stones) [CH 24], VI. 2007, Ruzicka (1 female with
doubtful identication
-
cWRA); Dali Bai Nat. Auton.
Pref., 36 km N Dali, 2158 m, 26°01’20’’N / 100°08’14’’E
(ruderal pasture with pines, shrubs, small brooks, und.
stones/litter sifted) [04], VIII. 2009, Wrase (1
-
cWRA);
Dali Bai Nat. Auton. Pref., Măo Jiao Shan, E pass,
2525 m, 58 km NE Dali, 25°56’41”N, 100°40’05”E,
(eld edge with shrubs, in soil) [26], IX. 2009, Wrase
(6
-
cWRA); Dali Bai Nat. Auton. Pref., mount. range E
Weishan, 12 km NE Weishan, 2630
-
2660 m, 25°17’02
-
15”N, 100°22’23
-
30”E, (slopes with itration springs,
under stones) [54], IX. 2009, Wrase (1
-
cWRA); Dali
280
Bai Nat. Auton. Pref., foothills of Diancang Shan W
Dali, 2050 m, 25°42’12’’N, 100°08’26’’E, (stone and
gravel pit) [34], VI. 2007, Schülke (1
-
cWRA); Dali
Bai Nat. Auton. Pref., NE bank of Er Hai, 27 km N Dali,
1980 m, 25°57’N, 100°09’E (embankment, under plants/
litter) [33], VI. 2007, Wrase (1
-
cWRA); Dali Bai Nat.
Auton. Pref., Wuliang Shan, 9 km SW Weishan, 2450
-
2500 m, 25°10’14’’N, 100°14’22’’E (W. slope, sec. oak/
pine for., pasture, und. stones) [35],VI. 2007, Wrase (1
-
cWRA); Dehong Dai Aut. Pref., mount. range 31 km E
Luxi, 2280 m, 24°29’31”N, 98°52’58”E, (grassland/pas-
ture, uinder stones/shrubs, in moss/litter) [19], VI. 2007,
Wrase (2
-
cWRA); Jizu Mts., 2300 m,VII. 1995, Bolm
(18
-
NHMB, cJAE, cWRA); Lijiang, V.
-
VI. 2010,
Kucera (1
-
cWRA); Lincang Pref., Bangma Shan, 20
km NW Lincang, 2210 m, 23°58’25”N, 99°54’36”E,
(water reservoir, in/on loamy soil of bank) [37], IX.
2009, Wrase (25
-
cWRA, cJAE, NME); Lincang Pref.,
Bangma Shan, 20 km NW Lincang, 2210 m, 23°58’25”N,
99°54’36”E, (bank of water reservoir, devast. forest, lit-
ter & ferns sifted [37], IX. 2009, Schülke (4
-
cWRA);
Lincang Pref., Xue Shan, 48 km N Lincang, 2070 m,
24°19’03”N, 100°07’13”E, (slope close to small pond,
in loamy soil/plant roots) [45], IX. 2009, Wrase (21
-
cWRA, cJAE); Ailaoshan Mt. Range, W Shuitangzhen
Town, 24°07’18’’ N / 101°27’44’’ E, 1205
-
1965 m, V.
2011, Belousov, Kabak, Korolev (1
-
ZISP); Jlanchuan
Co., Yuhua reservoir, 26.42694°N, 99.96504°E, 2437 m,
VII. 2007, Kataev & Liang (2
-
ZISP); Kunming, 1900
m, III. 1955, Kryzhanovskij (1 male
-
ZISP, examined
and communicated by B. Kataev).
Vietnam: Sapa (Lao Cai), 22°20’N, 103°50’E, V
-
VI.
1991, Jendek (2
-
NHMW); Lao Cai Province, 250 km
from Hanoi, bearing 310, SaPa vill. env., Hoang Lien
Son N. Park, ~ 1250 m, VI
-
VII. 1998, Napolov (1
-
ZISP); Lao Chai pr., Sa Pa env., ca. 1650 m, 22°19.52N
103°50.35E, V. 1999, Ahrens (2
-
cSCHM); 12 km N
Dalat, Lang Bian, 1580
-
1750 m, 12°03’N, 108°27’E,
IV. 1995, Pacholatko & Dembicky (8
-
NHMW, cJAE).
Anthracus wrasei nov. sp. (Figs 7
-
9, 36
-
41, 72, 77)
Type material: Holotype: 1 % (cWRA) labelled
“CHINA (N-Yunnan) Dali Bai / Nat. Aut. Pref., 4 km
E Dali / old town, shore terrain / of Er Hai Lake, 2020
m”, “25°42’N/100°11.5’E (young / willows with knot-
grass, / under vegetation/in roots /, 27.VIII./2.IX. 2003
Wrase [16]”, “Acupalpus / ? furvus Andr. / det. B. Kataev
2006” [rst two lines and year hw], “COLL. WRASE
/ BERLIN” and “HOLOTYPE % / Anthracus / wrasei
spec. nov. / des. B. Jaeger 2012”.
Paratypes: 1 %, 1 & (cWRA, cJAE) with same labels
as the holotype, but the male with “Acupalpus / fur-
vus Andr. / det. B. Kataev 2006” and the female with
“Acupalpus / (Setacupalpus) /furvus Andr. / det. B.
Kataev 2006” [rst three lines and year hw]. 1% (cWRA)
labelled “CHINA (Yunnan) Lincang Pref. / Xue Shan /
48 km N Lincang 2070 m / 24°19’03”N / 100°07’13”E
/ (slope close to small pond, / in loamy soil/plant roots)
/ 12.IX. 2009 D.W.Wrase [45]”. All paratypes addition-
ally with my label “PARATYPE % or & / Anthracus /
wrasei spec. nov. / des. B. Jaeger 2012”
Remarks: The holotype and the paratypes are in per-
fect condition without missing body parts, the paratype
from Xue Shan is moderately teneral, with somewhat
deformed pronotum and right elytron.
Description: General appearance as gured (Fig. 7).
Body length 4.4
-
4.5 mm (holotype 4.4 mm); width
1.6
-
1.7 mm.
Head mainly blackish, posterior to eyes with two dark
reddish spots. Clypeus dark reddish, labrum and mandi-
bles (inner margins and apices blackish) paler reddish.
Pronotum dark to blackish brown, with lateral margins
paler brownish, sometimes also middle part of base
and apical margin indistinctly brownish. Elytra almost
blackish, leaving the lateral margin and the rst interval
dark reddish brown. Sometimes posterior to shoulders
and at sides more dark brown than blackish. Legs and
palpi yellowish brown, antennae dark brown, with rst
two antennomeres paler. Ventral surface mainly black-
ish brown, with posterior part of prosternum, metaster-
num, and sometimes also abdominal sternites somewhat
paler. Upper surface shiny, elytra moderately iridescent.
Head (Figs 7
-
9) including eyes 0.78
-
0.82 times as
wide as pronotum, with eyes moderately prominent
(head 1.49
-
1.55 times as wide as head between eyes).
Labrum almost rectilinear at apical margin. Mandibles
medium sized, not distinctly prolonged and curved, left
mandible faintly obtuse at apex, not thickened or trun-
cate. Antennae 2.45
-
2.52 times as long as pronotum
and 0.83
-
0.86 times as long as elytra. Microsculpture
on labrum and clypeus distinct and almost isodiametric,
on anterior half of head almost obliterated in males, in
281
%&
females with lightly impressed isodiametric meshes, on
posterior half faintly isodiametric in both sexes.
Pronotum (Figs 7
-
9) 1.29
-
1.34 times as wide as long,
1.22
-
1.27 times as wide as head, widest in second quar-
ter, lateral seta inserted at or a little posterior to beginning
of second quarter. Apical margin almost rectilinear or
slightly emarginate, lateral border interrupted, reaching
medial third. Anterior angles narrowly rounded at tips,
not or only weakly projecting forward. Sides convex in
anterior half, rectilinearly narrowed to posterior angles,
which are obtuse but at least suggested, instead of being
widely rounded. Basal margin weakly arcuate medially,
obliquely produced to posterior angles. Lateral furrows
evenly narrow in apical two thirds, becoming distinctly
widened at posterior third, where they are fused with
the latero-basal impressions. Latero-basal impressions
rather small, distinctly delimited from pronotal disc and
median part of base, fused with basal and lateral margin.
Basal impressions and other surface of pronotum im-
punctate. Median line ne on disc, disappearing before
reaching basal and apical margins. Anterior transverse
impression obliterated or suggested. Microsculpture
with faintly impressed, short to long transverse meshes
on disc, at latero-basal impressions and lateral furrows
with course isodiametric meshes.
Elytra (Fig. 7) with sides weakly widened posteriorly,
widest just posterior to middle, 1.62
-
1.65 times as long
as wide, 2.89
-
3.02 times as long and 1.32
-
1.40 times
as wide as pronotum. Elytral striae distinctly impressed
and impunctate, scutellar striole long. Intervals rather
at, narrowed and very weakly convex at apex. Basal
pore at beginning of scutellar striole present, interval 3
in third quarter with one setiferous pore, adjoining stria
2. Microsculpture on scutellum almost isodiametric, on
elytral intervals with indistinct, very lightly impressed
transverse lines. Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.6
-
1.7 times longer than wide at basal
margin. Prosternum medially with 5
-
6, and posterior
to apical margin with a row of 8
-
10 longer setae. Pros-
ternal process with at least 1 distinct seta at middle (ad-
ditional setae probably broken). Abdominal sternites
4
-
6 with distinct pubescence. Last visible abdominal
sternite of males moderately emarginate and that of fe-
males very faintly emarginate at apex, at apical margin
with two longer setae in males and four in females.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 markedly bilobed. Mesotar-
someres 1
-
4 moderately dilated and with adhesive hairs
on ventral surface, mesotarsomere 4 moderately bilobed.
In females pro- and mesotarsomeres unmodied.
Median lobe of aedoeagus (Figs 36
-
41) with general
appearance and structures of internal sac as gured.
Apex in lateral aspect with a rather long and distinct
hook. Internal sac (lateral aspect) composed of 2
-
4 two
larger apical tooth, and a row of 8
-
10 dorsally arranged
large to medium sized teeth, and a row of 6
-
9 large to
medium sized ventrally arranged teeth.
Females genitalia as gured (Fig. 72) without signi-
cant specic characters.
Comparisons: A. wrasei is in general appearance most
similar to A. furvus and they were collected together at
the eastern shore of the Er Hai Lake and at Xue Shan in
the Chinese province of Yunnan. It can be distinguished
easily from this species by the different structures of the
internal sac and the apex of the median lobe (Figs 36
-
41)
(for details see under furvus). Externally both taxa are very
similar but A. wrasei differs at least from the Yunnan pop-
ulations of A. furvus by characters given under A. furvus.
From A. latus which probably occurs sympatrically in
southern Yunnan, A. wrasei differs by the different gen-
eral appearance and the shape of the pronotum (Figs
7
-
9, 19
-
21), and the different shape of the median lobe
and the structures of the internal sac of the latter (Figs
36
-
41, 59
-
65).
Other similar species occur allopatrically farther west
(A. schuhi) or south (A. glabrus). They differ from A.
wrasei by various external and aedoeagal features (for
details see under these species).
Etymology: The species is dedicated to my friend and
colleague David Wrase, specialist of Harpalini, who
collected this and other interesting Anthracus speci-
mens during his collecting trips to China.
Distribution: A. wrasei is so far known from two lo-
calities in W Yunnan (Fig. 77) where it occurs together
with A. furvus. Because the species has fully developed
hind wings, it is obviously more widely distributed and
has probably a similar distribution as the latter species.
Anthracus glabrus (Louwerens, 1952)
(Figs 10
-
12, 42
-
48, 67, 74, 79)
Acupalpus glabrus Louwerens, 1952: 213
-
214 (type
locality: Indonesia: E Java: Idjen Plateau, Blawan)
282
Acupalpus glabrus Louwerens: Lorenz 1998: 338
[world catalogue], 2005: 3590 [world catalogue].
Remarks: Louwerens did not provide an etymology when
naming the species „glabrus“ but referred probably to the
glabrous dorsal surface, or to the glabrous prosternal
process. If so, the correct latin name of the species
would be the adjective „glaber“ instead of „glabrus“.
The name given by Louwerens is here regarded as an
incorrect latinization, rather than a lapsus calami, and
therefore has not been emended. (ICZN Article 32.5.1).
Type material: Holotype: 1 % (RMNH) labelled “H.
LUCHT, Java / K. O. Blawan / 900/1500 Mr. / Idjen
Plateau / 2.III. / 1941” [last two lines vertical, hw],
“Museum Leiden / ex. collection / C. J. Louwerens /
rec. 1979”, “Typus / Acupalpus / glabrus Louw. / det.
C.J. Louwerens” [rst three lines hw Louwerens, rst
line in red ink] and “type” [on red paper].
Paratype: 1 & (RMNH) labelled “H. LUCHT, Java / K. O.
Blawan / 900/1500 Mr. / Idjen Plateau / 2.III. / 1941” [last
two lines vertical, hw], „Museum Leiden / ex. collection /
C. J. Louwerens / rec. 1979”, “PARATYPUS” [red paper];
“Acupalpus / glabrus Louw. / det. C. J. Louwerens” [rst
two lines hw Louwerens] and “Paratype” [red paper].
The holotype and the paratype additionally with my la-
bel: “Acupalpus / glabrus / Louwerens, 1952 / det. B.
Jaeger 2012”.
Remarks: The holotype is in good condition except left
metatibia and tarsomeres are missing. The paratype has
the antennomeres, left protarsomeres 3
-
5, and the left
metatibia and tarsomeres missing.
The remaining two paratypes mentioned in the descrip-
tion should be deposited in the Zoological Museum
Bogor (West Java) according to the introduction of
Louwerens’ contribution.
Redescription: General appearance as gured (Fig.
10). Body length 3.8
-
4.3 mm (holotype 3.9 mm); width
1.5
-
1.7 mm.
Colour: Dorsal surface dark brown to blackish brown,
leaving clypeus, labrum, mandibles (inner margins and
apices blackish), pronotal and elytral lateral margins
and rst interval of elytra reddish brown. Legs and palpi
pale or yellowish brown, antennae dark brown, with
rst, sometimes also second antennomere paler. Ventral
surface mainly brown or reddish brown, with head, pro-
and metepisterna, and sternites laterally darker. Upper
surface shiny, elytra with weak iridescence.
Head (Figs 10
-
12) including eyes 0.76
-
0.81 times as
wide as pronotum, with eyes moderately prominent
(head 1.44
-
1.55 times as wide as head between eyes).
Labrum almost rectilinear at apical margin, sometimes
faintly emarginated at middle. Mandibles medium sized,
not distinctly prolonged and curved, left mandible
moderately obtuse at apex. Antennae 2.27
-
2.44 times
as long as pronotum and 0.76
-
0.81 times as long as
elytra. Microsculpture on labrum and clypeus distinctly
isodiametric to short transverse, on anterior half of head
almost obliterated, only traces of isodiametric meshes
are visible, on posterior half rst faintly isodiametric,
later, in front of pronotal apical margin meshes become
transversely stretched.
Pronotum (Figs 10
-
12) 1.28
-
1.36 times as wide as long,
1.23
-
1.31 times as wide as head, widest in second
quarter, lateral seta inserted a little posterior to begin-
ning of second quarter. Apical margin almost rectilinear
to weakly emarginate, lateral border reaching medial
third. Anterior angles narrowly rounded at tips, not or
weakly projecting forward. Sides convex in anterior
third to half, rectilinearly narrowed to posterior angles,
which are obtuse and at least suggested. Basal margin
almost straight or very weakly arcuate medially, ob-
liquely produced to posterior angles. Lateral furrows
evenly narrow anteriorly, becoming weakly widened
and somewhat reexed posteriorly, where they are
fused with the latero-basal impressions. Latero-basal
impressions small, distinctly delimited from the prono-
tal disc and median part of base, fused with basal and
lateral margin. Basal impressions and other surface of
pronotum impunctate, medial part of base often, and
anterior part of pronotum sometimes with some vertical
wrinkles. Median line ne on disc, disappearing before
reaching basal and apical margins. Anterior transverse
impression suggested. Microsculpture on disc with
faintly impressed, long-transverse meshes, which
become more distinct and short transverse in front of
anterior impression and at medial part of base. Latero-
basal impressions and lateral furrows with distinct iso-
diametric meshes.
Elytra (Fig. 10) with sides moderately widened posteri-
orly, widest just at or just posterior to middle, 1.51
-
1.62
times as long as wide, 2.93
-
3.06 times as long and
1.40
-
1.50 times as wide as pronotum. Elytral striae
283
distinctly impressed and impunctate, scutellar striole
long. Intervals rather at, narrowed and very weakly
convex at apex. Basal pore at beginning of scutellar
striole present, interval 3 in third quarter with one setif-
erous pore, adjoining stria 2. Microsculpture on scutel-
lum isodiametric, on elytral intervals with very lightly
impressed transverse lines. Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.7 times longer than wide at basal margin.
Prosternum medially with 4
-
5 and posterior to apical
margin with a row of 3
-
4 medium long setae (often
broken and then insertion points difcult to observe).
Prosternal process with 2
-
3 distinct setae (often broken).
Abdominal sternite 4 with moderate pubescence in me-
dial two thirds, and 5
-
6 completely with distinct pubes-
cence. Last visible sternite of males and females very
weakly emarginate at apex, at apical margin with two
longer setae in males and four in females.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 markedly bilobed. Mesotar-
someres 1
-
4 moderately dilated with adhesive hairs on
ventral surface. Mesotarsomere 4 moderately bilobed.
In females pro- and mesotarsomeres unmodied.
Median lobe of aedoeagus (Figs 42
-
48) with general
appearance and structures of internal sac as gured.
Apex (lateral aspect) with a short but distinct hook. In-
ternal sac composed of 1 large and 1
-
2 smaller subapi-
cal teeth (sometimes completely reduced, gs 43, 48)
and 4
-
5 smaller teeth arranged in medial portion, and
1
-
2 somewhat larger teeth in basal portion.
Females genitalia as gured (Fig. 74) without signi-
cant specic characters.
Comparisons: A. glabrus resembles in colour and gen-
eral appearance A. furvus, wrasei and schuhi, which are
distributed farther north according to present knowledge.
From all these species it can be distinguished easily by
the different shape of the median lobe of the aedoeagus
and the structures of the internal sac (Figs 42
-
48). Ex-
ternally it differs from A. furvus (excluding S Vietnam
specimens) by the pronotum which is more distinctly
and more rectilinearly narrowed posteriad and the elytra
which are longer and wider compared with the pronotum
(see ratios EL/PL, EW/PW in table 1). From S Vietnam
populations of A. furvus it can be separated by the nar-
rower head (see ratios HW/PW), comparatively shorter
and wider elytra and related proportions (see ratio EL/
EW, EW/PW). From A. wrasei it can be distinguished by
its smaller size, longer antennae (see ratios AL/PL resp.
AL/EL), somewhat shorter elytra (see ratio EL/EW) and
the pronotum which is more slender relative to the elytra
(see ratio EW/PW). From A. schuhi it can be separated
by the pronotal shape with sides more distinctly nar-
rowed posteriad, the larger, less distinctly produced eyes,
and various body proportions as PW/PL, EW/PW, EL/
PL, EL/EW, HW/HWbE (for details see table 1).
Distribution: Until now the species was known only
from East Java but is here recorded from Bali for the
rst time (Fig. 79). Because the species has fully de-
veloped hind wings it is obviously more widely distrib-
uted. In addition to the type material mentioned above
the following material has been examined:
Examined Material:
Indonesia: Bali: Bedugul reg., Tamblingan Lakes N.P.,
alt, 1200m, IX. 2004 (18
-
cBUL, cJAE).
Anthracus schuhi nov. sp.
(Figs 13
-
15, 49
-
54, 69, 78)
Type material: Holotype: 1 % (NHMW) labelled
“INDIA: Maharashtra / Lonvala 13.9.1991 / (80 km
E Bombay) / leg. R. SCHUH” and “HOLOTYPE %
/ Anthracus / schuhi spec. nov. / des. B. Jaeger 2012”
[red paper].
Paratypes: 2 %% (NHMW, cJAE) with same locality
label as the holotype and “PARATYPE % / Anthracus
/ schuhi spec. nov. / des. B. Jaeger 2012” [red paper].
Remarks: The holotype is in good condition except for
left antennomeres 9
-
11 which are missing. The para-
types are in moderate condition and have some anten-
nomeres and/or tarsomeres missing.
Description: General appearance as gured (Fig. 13). Body
length 4.0
-
4.1 mm (holotype 4.0 mm); width 1.5 mm.
Colour: Head and pronotum dark brown with lateral
margins of the latter somewhat paler, elytra blackish
brown, with lateral margin and rst interval somewhat
lighter. Clypeus, labrum and mandibles (inner margins
and apices blackish) reddish brown. Legs and palpi yel-
lowish brown, antennae brown, with rst two antenno-
meres somewhat paler. Ventral surface mainly brown
to dark brown, with head, pro-, mes- and metepisterna
somewhat darker. Upper surface shiny, elytra with
weak iridescence.
284
Head (Figs 13
-
15) including eyes 0.77
-
0.79 times
as wide as pronotum, with eyes distinctly prominent
(head 1.61
-
1.63 times as wide as head between eyes).
Labrum almost rectilinear at apical margin. Mandibles
not markedly prolonged and curved, left mandible
weakly obtuse at apex. Antennae 2.38
-
2.39 times as
long as pronotum and 0.86
-
0.88 times as long as elytra.
Microsculpture on labrum and clypeus distinct and al-
most isodiametric, on anterior half of head almost ob-
literated, only traces of isodiametric meshes are visible,
on posterior half of head rst faintly isodiametric, later
(in front of pronotal apical margin) the meshes become
transversely stretched.
Pronotum (Figs 13
-
15) 1.22
-
1.25 times as wide as
long, 1.27
-
1.30 times as wide as head, widest in second
quarter, lateral seta inserted a little posterior to be-
ginning of second quarter. Apical margin weakly to
moderately emarginate, lateral border reaching medial
third. Anterior angles narrowly rounded at tips, weakly
projecting forward. Sides convex in anterior third, rec-
tilinearly or roundly narrowed to the widely rounded
posterior angles. Basal margin almost straight or very
weakly arcuate medially, obliquely produced to poste-
rior angles. Lateral furrows evenly narrow anteriorly,
becoming weakly to moderately widened posteriorly,
where they are fused with the latero-basal impressions.
Latero-basal impressions medium sized, distinctly de-
limited from the pronotal disc and median part of base,
fused with basal and lateral margin. Median line ne
on disc, disappearing before reaching basal and apical
margins. Anterior transverse impression only suggest-
ed. Basal impressions and other surface of pronotum
impunctate. Microsculpture on disc with faintly im-
pressed, long-transverse meshes, at basal impressions
and in lateral furrows with distinctly impressed isodia-
metric meshes.
Elytra (Fig. 13) with sides very weakly widened poste-
riorly, widest about at middle, 1.62
-
1.63 times as long
as wide, 2.63
-
2.77 times as long and 1.32
-
1.36 times
as wide as pronotum. Elytral striae distinctly impressed
and impunctate, scutellar striole long. Intervals rather
at, narrowed and very weakly convex at apex. Basal
pore at beginning of scutellar striole present, interval 3
in third quarter with one setiferous pore, adjoining stria
2. Microsculpture on scutellum isodiametric, on elytral
intervals with very lightly impressed transverse lines.
Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.4 times longer than wide at basal margin.
Prosternum medially with 5
-
6 and posterior to api-
cal margin with a row of 6 medium long setae (often
broken and then insertion points difcult to observe).
Prosternal process with 2
-
3 distinct setae (broken in
the holotype). Abdominal sternite 4
-
6 with distinct,
rather long and dense pubescence. Last visible sternites
of males very weakly emarginate at apex, at apical margin
with two longer setae in males.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 markedly bilobed. Mesotar-
someres 1
-
4 moderately dilated with adhesive hairs on
ventral surface. Mesotarsomere 4 moderately bilobed.
Median lobe of aedoeagus (Figs 49
-
54) with general
appearance and structures of internal sac as gured.
Apex in lateral aspect without any distinct hook. Apical
lamella (dorsal aspect) rather wide and long, at apical
margin widely rounded. Internal sac with 3
-
4 smaller
subapical teeth, and a band of closely arranged medium
sized teeth in basal half.
Etymology: The species is dedicated to Rudolf Schuh,
Vienna, specialist of Cucujoidea, who collected this
and another new Anthracus species in India.
Comparisons: Anthracus schuhi is in colour and general
appearance similar to A. furvus, wrasei and glabrus
which occur farther east or southeast. It differs from
these species by the shape of aedoeagal median lobe
and its peculiar internal structures (Figs 49
-
54), the
head with rather strongly produced eyes, the shape of
the pronotum with sides less distinctly narrowed poste-
riorly and hind angles widely rounded and various body
proportions e.g. HW/HwbE, PW/PL, EW/PW, EL/PL
(for details see table 1).
Distribution: So far known only from the type locality
in West India, Maharashtra province (Fig. 78). Because
the species has fully developed hind wings it is probably
more widely distributed.
Anthracus indicus nov. sp.
(Figs 16
-
18, 55
-
58, 68, 73, 78)
Type material: Holotype: 1 % (NHMW) labelled “INDIA:
Maharashtra / Lonvala 13.9.1991 / (80 km E Bombay) /
leg. R. SCHUH” and “HOLOTYPE % / Anthracus /
indicus spec. nov. / des. B. Jaeger 2012” [red paper].
285
Paratypes: 1 %, 2 && (NHMW, cJAE) with same lo-
cality label as the holotype and “PARATYPE % or &
/ Anthracus / indicus spec. nov. / des. B. Jaeger 2012”
[red paper].
Remarks: The holotype is in perfect condition, two of
the paratypes have some antennomeres and tarsomeres
missing.
Description: General appearance as gured (Fig. 16).
Body length 3.3
-
3.4 mm (holotype 3.3 mm); width
1.2
-
1.3 mm.
Colour: Head and pronotum reddish brown or brown,
the former sometimes slightly darker, elytra dark brown
(in one PT almost blackish brown), with lateral margin
and rst interval reddish brown. Clypeus, labrum and
mandibles (inner margins and apices blackish) pale red-
dish brown. Legs, palpi and antennae entirely yellowish
brown, sometimes antennomeres 3
-
11 with a suggested
infuscation. Ventral surface brown to reddish brown.
Pronotum weakly, elytra markedly iridescent.
Head (Figs 16
-
18) including eyes 0.75
-
0.79 times as
wide as pronotum, with eyes small and moderately
prominent (head 1.51
-
1.54 times as wide as head be-
tween eyes). Labrum almost rectilinear at apical mar-
gin. Mandibles not markedly prolonged and curved,
left mandible rather sharp at apex. Antennae 2.32
-
2.42
times as long as pronotum and 0.88
-
0.93 times as long
as elytra. Microsculpture on labrum and apical half of
clypeus distinct and almost isodiametric, on anterior
half of head almost obliterated, only traces of isodia-
metric meshes are visible in females, on posterior half
rst faintly isodiametric, later, in front of pronotal apical
margin, the meshes become transversely stretched.
Pronotum (Figs 16
-
18) 1.21
-
1.27 times as wide as long,
1.27
-
1.32 times as wide as head, widest in second quar-
ter, lateral seta inserted a little in front of widest point.
Apical margin almost rectilinear to faintly emarginate,
in one PT moderately emarginate, lateral border reaching
medial third. Anterior angles narrowly rounded at tips,
only weakly projecting forward. Sides convex in anteri-
or third, rectilinearly or roundly narrowed to the widely
rounded posterior angles. Basal margin almost straight
or very weakly arcuate medially, obliquely produced
to posterior angles. Lateral furrows evenly narrow
anteriorly, moderately to distinctly widened posteriorly,
where they are fused with the latero-basal impressions.
Latero-basal impressions rather small, distinctly delim-
ited from the pronotal disc and median part of base, and
fused with basal and lateral margin. Basal impressions
and other surface of pronotum impunctate. Median line
ne on disc, disappearing before reaching basal and api-
cal margins. Anterior transverse impression weak or only
suggested. Microsculpture on disc with a mixture of very
faintly impressed lines and long-transverse meshes, at
basal impressions and in lateral furrows with distinctly
impressed isodiametric meshes.
Elytra (Fig. 16) with sides weakly widened posteriorly,
widest at middle, 1.58
-
1.59 times as long as wide, 2.61
-
2.69 times as long and 1.34
-
1.38 times as wide as prono-
tum. Elytral striae distinctly impressed and impunctate,
scutellar striole long. Intervals rather at, narrowed and
very weakly convex at apex. Basal pore at beginning of
scutellar striole present, interval 3 in third quarter with
one setiferous pore, adjoining stria 2. Microsculpture
on scutellum isodiametric, on elytral intervals with very
lightly impressed transverse lines. Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.4 times longer than wide at basal mar-
gin. Prosternum medially with 4 and posterior to api-
cal margin with a row of 6 medium long setae (often
broken and then insertion points difcult to observe).
Prosternal process with at least 1 seta (others probably
broken). Abdominal sternite 4 in medial two thirds and
5
-
6 entirely with distinct, rather long and dense pubes-
cence. Last visible sternite of males very weakly emar-
ginate at apex, at apical margin with two longer setae in
males and four setae in females.
Protarsomeres 1
-
4 of males markedly dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 markedly bilobed. Mesotar-
someres 2
-
4 moderately dilated with adhesive hairs on
ventral surface. Mesotarsomere 4 moderately bilobed.
Pro- and mesotarsomeres of females unmodied.
Median lobe of aedoeagus (Figs 55
-
58) with general
appearance and structures of internal sac as gured.
Apex in lateral aspect almost straight, without any
hook. Apical lamella (dorsal aspect) rather wide and
short, at apical margin widely rounded. The internal sac
has only one medium sized subapical tooth.
Females genitalia as gured (Fig. 73) without signi-
cant specic characters.
Comparisons:Acupalpus indicus occurs sympatrically
with A. schuhi and differs from this and most other
immaculate species of the A. annamensis group by its
286
small size and its pronotal colour, which is markedly
paler than that of the elytra, and the more or less uni-
coloured antennae. From A. derogatus which is also
rather small in size it can be separated by the different
colour and shape of head and pronotum (Figs 1, 17
-
18).
In addition, the median lobe of the aedoeagus (Figs
55
-
58) has a relatively short and wide apical plateau,
which is straight in lateral view and has no capitulum
or hook, and the internal sac is rather simple with only
one small subapical tooth.
Etymology: The name “indicus” refers to the known
distribution of the species.
Distribution: Known only from the type locality in
West India, Maharashtra province (Fig. 78). Because
the species has fully developed hind wings it is probably
more widely distributed.
Anthracus latus nov. sp.
(Figs 19
-
21, 59
-
65, 70, 75, 77)
Type material: Holotype: 1 % (NMP) labelled “NW
Thailand, 25.iv.
-
/ Chiang Mai prov.
-
7.v. / BAN SAN
PAKIA 1996 / Sv. Bilý leg., 1700m” and “HOLOTYPE
% / Anthracus / latus spec. nov. / des. B. Jaeger 2012”.
Paratypes: Vietnam: 5 %%, 5 && (NHMW, cJAE)
labelled “S-VIETNAM: 17.
-
21.4. / 12km N Dalat
1995 / Lang Bian”; “12°03’n 108°27’e / 1580
-
1750
m / Pacholatko & Dembicky”. 1& (NHMW) labelled
“N-Vietnam 25.V.
-
10.VI. / SAPA (Lao Cai) / 22°20’N
103°50’E / leg. E. Jendek 1991”.
China: 1 % (NME) labelled “CHINA: S-YUNNAN
/ (Xishuangbanna) / 37 km NW Jinghong / vic. Guo
Men Shan” and “ N22°14.43 E100°36.12 / 1100m,
06.IV.2009 MF / leg. L.Meng rice fallow”. 1 % (cWEIG)
labelled “CHINA: S-YUNNAN / (Xishuangbanna) /
20 km NW Jinghong / vic. Man Dian (NNNR)” and
“ N22°07.80 E100°40.05 / 730m, 06.IV.2009 BF /
leg. L.Meng forest”. 1 & (IZAS) labelled “CHINA:
S-YUNNAN / (Xishuangbanna) / 37 km NW Jinghong
/ vic. Guo Men Shan” and “ N22°14.48 E100°36.22 /
1080m, 06.IV.2009 EKL / leg. L.Meng prim. forest”.
Laos: 1 % (NMP) labelled “LAOS, Hua Phan prov.,
2001 / 20°15’N 104°02’E, 26.iv.
-
11.v. / Ban Saluei,
Phu Phan Mt. env. / Jan Bezděk leg., 1500
-
2000 m”.
Thailand: 11 %%, 3 && (NMP, cJAE) with same label
data as the holotype. 2 %%, 2 && (NMP, cJAE) la-
belled “NW Thailand, Chieng Dao, / Ban San Pakia, 5.
-
10. v. / 2004, 1200m, Sv. Bilý leg.”. 2 %%, 1 & (cKME,
cWRA) labelled “THAI-N, 1
-
15.v.1998, / Chiang Mai
prov., / 19°19’N 98°50’E, / SAN PAKIA, 1400m, / Vit
Kubáň leg.”, 1 % additionally with “COLL. WRASE
/ BERLIN” [green label]. 2 && (cKME, cJAE) “N.
THAILAND 19°19’N / Chiang Mai prov. 98°50’E / San
Pakia, 1400m / 1.
-
15.5.1998 / Michael Bednařik leg.”
and 1 additionally with “Collectio / Rudolf KMECO”.
6 %%, 1 &, (NMP, cJAE) labelled “NW THAILAND,
1200m / Mae Hong Son pr. / BAN SI LANG, 20.-22.v. /
Sv. Bilý leg. 1996”. 2 && (NHMB) “NW THAILAND
9.
-
16.5. 1991 / MAE HONG SON / BAN HUAI PO
1600m / LEG. PACHOLÁTKO”; 2 %%, 1 & (MHNG)
labelled “THAILAND: Chiang Mai / Doi Angkhang
10km W / Fang 1450m 22.IV.87 / P. Schwendinger”.
4 %% (cKME, cWRA, cJAE) “N. THAILAND / 50
km NW of Chiang Rai / Mae Salong env. / 1.5.2003 O.
Šafránek leg.”, 3 of them additionally with “Collectio /
Rudolf KMECO / Czech Republic” and 1 with “COLL.
WRASE / BERLIN” [green label].
Myanmar: 1 % (cWRA) labelled “MYANMAR
(Chin State) / Chin Hills / Avocado Plantage /
21°23’34.7”N/093°52’29.4”E / 30.VI.
-
01.VII.2008
M. Langer”, “COLL. WRASE / BERLIN” [with black
frame], “Acupalpus / spec. / D.W. Wrase det. ‘11” [rst
two lines hw]; “COLL. WRASE / BERLIN” [green la-
bel].
All Paratypes additionally with my label “PARATYPE
% or & / Anthracus / latus spec. nov. / des. B. Jaeger
2012”.
Remark: The holotype is in perfect condition, except
right antennomere 11 is missing. Some of the paratypes
have missing antennomeres or tarsomeres.
Description: General appearance as gured (Fig. 19).
Body length 3.5
-
4.4 mm (holotype 4.1 mm); width
1.4
-
1.7 mm.
Dorsal surface mainly dark to blackish brown, with
pronotum often somewhat paler, and clypeus, labrum,
mandibles (inner margins and apices blackish), lateral
margins of pronotum and elytra, and rst interval of the
latter paler reddish brown. Legs, palpi and rst two an-
tennomeres yellowish brown, remaining antennomeres
darkened. Ventral surface brown, with mouthparts, epi-
pleura, pro-, meso- and metasternum, and abdominal
sternites medially pale brown. Upper surface shiny,
elytra slightly iridescent.
287
Head (Figs 19
-
21) including eyes 0.70
-
0.76 times as
wide as pronotum, with eyes moderately prominent
(head 1.50
-
1.59 times as wide as head between eyes).
Labrum with apical margin rectilinear or slightly emar-
ginate at middle. Mandibles not markedly prolonged
and curved, left mandible weakly obtuse at apex.
Antennae rather short only 2.02
-
2.26 times as long
as pronotum and 0.72
-
0.79 times as long as elytra.
Microsculpture on labrum and clypeus distinct and
almost isodiametric, on anterior half of head almost ob-
literated, only traces of isodiametric meshes are visible
in females, on posterior half of head rst isodiametric,
later (in front of pronotal apical margin) the meshes be-
come transversely stretched.
Pronotum (Figs 19
-
21) rather broad, 1.35
-
1.40 times as
wide as long, 1.32
-
1.43 times as wide as head, widest
in second quarter, lateral seta inserted a little posterior
to beginning of second quarter. Apical margin almost
straight or faintly emarginate, lateral border distinctly
prolonged to middle, leaving the medial third or quarter
unbordered. Anterior angles narrowly rounded at tips,
quite clearly projecting forward. Sides convex in ante-
rior third, rectilinearly or roundly narrowed to the widely
rounded posterior angles. Basal margin almost straight
or very weakly convex medially, and weakly produced
to posterior angles. Lateral furrows evenly narrow ante-
riorly, becoming moderately widened posteriorly, where
they are fused with the latero-basal impressions. Latero-
basal impressions medium sized, clearly delimited from
the pronotal disc and median part of base, fused with
basal and lateral margin. Basal impressions and other
surface of pronotum impunctate, the former sometimes
with some irregular wrinkles or rugosities. Median line
ne on disc, disappearing before reaching basal and
apical margins. Anterior transverse impression weak to
obsolete. Microsculpture on disc with faintly impressed,
long-transverse meshes, at basal impressions and in lat-
eral furrows with distinctly impressed short transverse to
irregularly isodiametric meshes.
Elytra (Fig. 19) with sides weakly widened posterior-
ly, widest at middle, 1.51
-
1.62 times as long as wide,
2.77
-
2.96 times as long and 1.29
-
1.36 times as wide
as pronotum. Elytral striae distinctly impressed and
impunctate, scutellar striole long. Intervals rather at,
narrowed and weakly convex at apex. Basal pore at be-
ginning of scutellar striole present, interval 3 in third
quarter with one setiferous pore, adjoining stria 2. Mi-
crosculpture on scutellum isodiametric, on elytral inter-
vals with lightly impressed transverse lines or indistinct
meshes. Macropterous.
Metepisterna long and narrowed posteriad, at inner
margin about 1.5
-
1.6 times longer than wide at basal
margin. Prosternum medially with 4
-
6 and posterior to
apical margin with a row of 8 medium long setae (often
broken and then insertion points difcult to observe).
Prosternal process with at least 1 seta (very often
broken). In contrast to other immaculate species treated
above abdominal sternite 4 glabrous, or only with few
single hairs in medial part, 5 with weak and scattered,
6 with moderately dense pubescence (hairs shorter and
less densely arranged than in furvus and other species).
Last visible sternite of males weakly emarginate at
apex, at apical margin with two longer setae in males
and four setae in females.
Protarsomeres 1
-
4 of males moderately dilated and with
distinct, biseriately arranged adhesive hairs on ventral
surface. Protarsomere 4 weakly bilobed. Mesotarsomeres
2
-
4 moderately dilated with adhesive hairs on ventral sur-
face. Mesotarsomere 4 not or only very weakly bilobed.
Median lobe of aedoeagus (Figs 59
-
65) with external
shape and structures of internal sac as gured. Apex in
lateral aspect with distinct capitulum. Internal sac with
one large tooth in apical half, and with an accumulation
of some medium sized and various smaller teeth medially.
Females genitalia as gured (Fig. 75) without signi-
cant specic characters.
Etymology: The Latin adjective latus, meaning broad,
refers to the broad body of the species.
Comparisons: Acupalpus latus differs at rst glance
from all other immaculate species by its robust, rather
broad body. It can be distinguished particularly by its
rather wide pronotum (PW/PL > 1.34), rather short an-
tennae (AL/PL < 1.27), the lateral border of the prono-
tal apical margin which is distinctly prolonged towards
the mid line, the reduced pubescence of abdominal ster-
nite 4 with only a few single hairs or glabrous, instead
of being moderately to distinctly pubescent as in other
species, and the completely different apex of aedoeagal
median lobe and its characteristic inner structures.
Distribution: The species occurs from S Vietnam
across N Vietnam, northern Laos, southern Yunnan,
northern Thailand to west Myanmar (Fig. 77). It has
been collected in mountainous areas at altitudes of
730
-
2000 m.
288
Key to immaculate species of A. annamensis group from the East Palaearctic and Oriental Regions
1 Smaller species, 3.1
-
3.4 mm, with general appearance, shape of head and pronotum as gured (Figs. 1
-
2, 16
-
18). Antennae with antennomeres 3
-
11 not markedly darkened. Median lobe (Figs 22
-
24, 55
-
58) 2
-
Larger species, mostly 3.5
-
4.6 mm, when smaller then general appearance and median lobe different (Figs 4
-
15,
19
-
21, 25
-
54, 59
-
65). Antennae with antennomeres 3
-
11 markedly darkened 3
2 Head and pronotum blackish brown, pronotum markedly narrowed posteriad (Fig. 1), head with eyes 1.46 times
as wide as head between eyes, elytra shorter (EL/EW < 1.53), pronotum broader (PW/PL > 1.30). Median lobe
and internal structures as gured (Figs 22
-
24). 3.1 mm. So far known only from Sri Lanka
A. derogatus (Walker, 1858)
-
Head and pronotum reddish to reddish brown, pronotum less markedly narrowed posteriad (Figs 16
-
18), head
with eyes 1.51
-
1.54 times as wide as head between eyes, elytra longer (El/EW > 1.57), pronotum narrower (PW/
PL < 1.28). Median lobe and internal structures different (Figs 55
-
58). 3.3
-
3.4 mm. So far known only from the
type locality in western India A. indicus nov. sp.
3 General appearance with pronotum and elytra rather broad (Figs 19
-
21), fourth abdominal sternite glabrous or
with few single hairs only. Median lobe and internal structures as gured (Figs 59
-
65). 3.5
-
4.4 mm. Widely
distributed (Fig. 77) from S Vietnam in the southeast to western Myanmar in the west. A. latus nov. sp.
-
General appearance with pronotum and elytra more slender (Figs 4
-
15), fourth abdominal sternite with distinct
pubescence. Median lobe and internal structures different (Figs 25
-
54). 4
4 Known from Indonesian Isles Java and Bali. General appearance and shape of pronotum as gured (Figs 10
-
12)
Elytra rather long and broad relative to pronotum (EW/PW > 1.39, EL/PL >2.93). Median lobe and internal
structures as gured (Figs 42
-
48). 3.8
-
4.2 mm. A. glabrus (Louwerens, 1952)
-
Species from India or continental SE Asia. General appearance and shape of pronotum different (Figs 4
-
9, 13
-
15). Elytra usually shorter and more slender relative to pronotum (EW/PW < 1.42, EL/PL < 2.95). Median lobe
and internal structures different (Figs 25
-
41, 49
-
54). 5
5 Species from western India. Head with eyes more distinctly prominent (HW/HWbE > 1.60) and pronotum rather
long (PW/PL < 1.26) and less distinctly narrowed to posterior angles (Figs 13
-
15). Median lobe and internal struc-
tures as gured (Figs 49
-
54). 4.0
-
4.1 mm. A. schuhi nov. sp.
-
Species with distribution farther east. Head with eyes less prominent (HW/HWbE < 1.59) and pronotum shorter
and broader (PW/PL > 1.24) and more distinctly narrowed to posterior angles (Figs 4
-
9). Median lobe and inter-
nal structures as gured (Figs 25
-
41) 6
6 Species larger, 4.3
-
4.5 mm. Head with two distinct dark reddish spots behind eyes. Pronotum (Figs 7
-
9) with
sides rectilinearly narrowed to base and lateral furrows more distinctly widened posteriorly. Antennae markedly
longer relative to pronotum (AL/PL > 2.44). Apex of aedoeagal median lobe (Figs 36
-
38, 41) with a rather long
distinct hook. Internal structures in addition to a distinct dorsal row of teeth also with a ventrally arranged row
of medium sized teeth. Known from two localities in the Chinese province Yunnan. A. wrasei nov. sp.
-
Species smaller, 3.3
-
4.1 mm. Head usually entirely blackish brown or more rarely indistinctly dark reddish
behind eyes. Pronotum (Figs 4
-
6) less distinctly narrowed to base and lateral furrows not or less distinctly
widened posteriorly. Antennae shorter relative to pronotum (AL/PL < 2.40). Apex of aedoeagal median lobe
(Figs 30
-
35) with a short indistinct hook. Internal sac in addition to the dorsal teeth row only with 1
-
3 ventrally
arranged teeth. Known from E Myanmar, Yunnan, N Vietnam and S Vietnam. Members of the latter population
are markedly larger (3.9
-
4.6 mm) and represent a distinct geographical form, which cannot be separated from
the preceding species by the aforementioned external characters, but differs clearly by the shape of median lobe
and structures of the internal sac (Figs 29, 34, 35). A. furvus (Andrewes, 1947)
289
Acknowledgements
I thank all curators and collection staff mentioned in
the material section, who lent the material, which forms
the basis for this paper. I am also very grateful to Dr.
Roger Booth (London) for the linguistic revision of the
nal draft and other valuable comments. Furthermore, I
thank Dr. Hans Fery (Berlin) for the important hints on
some nomenclatorical problems.
References
andreweS, h. e. (1919): VI. On the types of Oriental Carabidae in the
British Museum, and in the Hope Department of the Oxford Uni-
versity Museum.
-
The Transactions of the Entomological Society
London 1919: I+II: 209
-
217.
-
(1926): A Catalogue of Philippine Carabidae.
-
The Philippine
Journal of Science 31/3: 345
-
361.
-
(1928): A Catalogue of the Carabidae of Ceylon.
-
Spolia Zey-
lanica 14 (2): 135
-
195.
-
(1930): Catalogue of Indian Insects. Part 18
-
Carabidae.
-
Cal-
cutta: Government of India. Central Publication Branch: 389 pp.
-
(1933): Catalogue of the Carabidae of Sumatra.
-
Tijdschrift voor
Entomologie 76: 319
-
382.
-
(1947): Entomological Results from the Swedish Expedition 1934
to Burma and British India. Coleoptera: Carabidae. Collected by
René Malaise.
-
Arkiv för Zoologi 38 a. (n:o 20): 1
-
49.
bateS, h.w. (1886): On the Geodeaphagus Coleoptera collected my Mr.
George Lewis in Ceylon.
-
The Annals and Magazine of Natural
History (5) 17: 68
-
212.
-
(1889): Contributions à la Faune Indo-Chinoise. 3e Mémoire.
-
Annales de la Société entomologique de France (6) 9: 261
-
286.
-
(1892): Viaggio di Leonardo Fea in Birmania e regioni vicine.
XLIV. List of the Carabidae.
-
Annali del Museo Civico di Storia
Naturale di Genova 32: 267
-
428.
Chaudhary, a. (2005): A Checklist of Family Carabidae (Insecta: Co-
leoptera) from Nepal.
-
Journal of the Natural History Museum
22: 79
-
98.
Csiki, e. (1932): CaraBidae: Harpalinae VI. in: Junk w. & S. SChen-
kling (edS.): Coleopterorum CataloguS. parS 121.
-
Berlin:
w. Junk, Pp. 1023
-
1278.
Jaeger, B. (2009): Two new species of Psychristus subgenus Psychris-
tus Andrewes 1930 from Southeast Asia, India and Pakistan, with
additions to the Psychristus discretus group (Col., Carabidae).
-
Linzer biologische Beiträge 41/2: 1635
-
1657.
-
(2010): A new species of the Acupalpus subgenus Subacupal-
pus Habu, 1978 from Nepal and Yunnan, and a redescription of
Acupalpus sikkimensis Andrewes, 1930 (Coleoptera, Carabidae,
Harpalini).
-
Entomologische Blätter 106: 145
-
158.
Jaeger, b. & b.m. kataev (2003): Carabidae: Harpalini: Subtribe
Stenolophina. Pp. 397
-
406.
-
In: löbl, i. & a. Smetana (edS),
Catalogue of Palaearctic Coleoptera, Vol. 1.
-
Stenstrup: Apollo
Books, 819 pp.
Jedlička, a. & m. ChûJô (1964): 3. Family Carabidae.
-
Nature and
Live in Southeast Asia 3: 167
-
175.
Jedlička, a. & m. ChûJô (1966): Coleoptera of East Nepal. 3. Familie
Harpalidae.
-
Journal of the College of Arts and Sciences, Chiba
University 4 No. 4: 536
-
538.
Landin, B.-o. (1954): Entomological Results from the Swedish expe-
dition 1934 to Burma and British India. Coleoptera: Carabidae
collected by René Malaise.
-
Arkiv för Zoologi 8 (3): 399
-
472.
Lorenz, w. (1998): Systematic List of extant Ground Beetles of the
World (Insecta Coleoptera “Geadephaga”: Trachypachidae and
Carabidae incl. Paussinae, Cicindelinae, Rhysodinae).
-
Tutzing:
W. Lorenz, 502 pp.
-
(2005): Systematic List of extant Ground Beetles of the World
(Insecta Coleoptera “Geadephaga”: Trachypachidae and Carabi-
dae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition.
-
Tutzing: w. Lorenz, 530 pp.
louwerenS, C.J. (1952): New Carabidae from the Malay Archipelago
(Col.) (3rd Communication on Oriental Carabidae).
-
Treubia 21
(2): 211
-
229.
SChauberger, e. (1930): Zur Kenntnis der paläaerktischen Harpalinen
(Sechster Beitrag).
-
Koleopterologische Rundschau 15 (5/6):
193
-
209.
waLker, F. (1858): XXIII.
-
Characters of some apparently undescribed
Ceylon Insects. Order Coleoptera.
-
The Annals and Magazine of
Natural History (3) 2: 202
-
209.
Author‘s address:
Bernd Jaeger
Zingster Str. 40
D
-
13051 Berlin
Germany
e-mail: acupalpus.bj@gmx.de
290
Table 1. Variation of ratios among specimens of the A. annamensis group
species n BL HW/HWbE AL/PL AL/EL HW/PW
A. derogatus (%)1 3.1 1.46 2.35 0.89 0.80
A. furvus (%%)10 3.7-4.1 (3.9) 1.48-1.58 (1.54) 2.19-2.36 (2.28) 0.78-0.85 (0.82) 0.75-0.80 (0.77)
A. furvus (&&)10 3.4-4.1 (3.8) 1.45-1.55 (1.51) 2.19-2.39 (2.26) 0.77-0.83 (0.79) 0.76-0.83 (0.79)
A.furvus (Dalat) (%%)2 3.9-4.1 (4.0) 1.51-1.52 (1.51) 2.57-2.58 (2.58) 0.88-0.91 (0.89) 0.81-0.81 (0.81)
A. furvus (Dalat) (&&)6 4.1-4.6 (4.3) 1.49-1.54 (1.51) 2.38-2.62 (2.49) 0.80-0.88 (0.84) 0.81-0.83 (0.82)
A.wrasei (%%)3 4.3-4.4 (4.3) 1.49-1.55 (1.53) 2.45-2.52 (2.48) 0.84-0.86 (0.85) 0.78-0.82 (0.80)
A. wrasei (&&)1 4.5 1.51 2.51 0.83 0.80
A. glabrus (%%)10 3.8-4.3 (4.0) 1.49-1.54 (1.51) 2.27-2.44 (2.36) 0.77-0.81 (0.80) 0.76-0.81 (0.79)
A. glabrus (&&)10 3.9-4.2 (4.1) 1.44-1.55 (1.47) 2.27-2.35 (2.31) 0.76-0.78 (0.77) 0.78-0.81 (0.80)
A. schuhi (%%)3 4.0-4.1 (4.0) 1.61-1.63 (1.62) 2.38-2.39(2,38) 0.86-0.88 (0.87) 0.77-0.79 (0.78)
A. indicus (%%)2 3.3 1.52-1.53 (1.52) 2.39-2.42 (2.40) 0.91-0.93 (0.92) 0.75-0.77 (0.76)
A. indicus (&&)2 3.3-3.4 (3.4) 1.51-1.54 (1.52) 2.32 0.88 0.78-0.79 (0.78)
A. latus (%%)10 3.5-4.4 (4.0) 1.50-1.58 (1.53) 2.08-2.26 (2.18) 0.72-0.79 (0.77) 0.72-0.76 (0.73)
A. latus (&&)10 3.8-4.4 (4.2) 1.50-1.59 (1.53) 2.02-2.16 (2.07) 0.72-0.74 (0.73) 0.70-0.75 (0.73)
PW/HW PW/PL EW/PW EL/PL EL/EW
A. derogatus (%)1 1.25 1.31 1.32 2.64 1.52
A. furvus (%%)10 1.25-1.33 (1.30) 1.27-1.33 (1.30) 1.30-1.38 (1.34) 2.73-2.86 (2.80) 1.57-1.64 (1.61)
A. furvus (&&)10 1.21-1.31 (1.27) 1.25-1.37 (1.30) 1.35-1.41 (1.37) 2.78-2.90 (2.84) 1.55-1.61 (1.59)
A. furvus (Dalat) (%%)2 1.24-1.24 (1.24) 1.30-1.32 (1.31) 1.36-1.39 (1.38) 2.85-2.94 (2.90) 1.60-1.61 (1.61)
A. furvus (Dalat) (&&)6 1.20-1.23 (1.22) 1.27-1.33 (1.30) 1.35-1.41 (1.39) 2.93-3.02 (2.96) 1.62-1.65 (1.63)
A. wrasei (%%)3 1.22-1.27 (1.24) 1.29-1.34 (1.31) 1.32-1.39 (1.37) 2.89-2.94 (2.92) 1.62-1.65 (1.63)
A. wrasei (&&)1 1.26 1.32 1.40 3.02 1.64
A. glabrus (%%)10 1.23-1.31 (1.27) 1.28-1.33 (1.31) 1.40-1.47 (1.42) 2.93-3.00 (2.96) 1.54-1.62 (1.58)
A. glabrus (&&)10 1.24-1.28 (1.26) 1.26-1.36 (1.33) 1.41-1.50 (1.46) 2.95-3.06 (3.01) 1.51-1.58 (1.56)
A. schuhi (%%)3 1.27-1.30 (1.28) 1.22-1.25 (1.24) 1.32-1.36 (1.34) 2.63-2.77 (2.71) 1.62-1.63 (1.63)
A. indicus (%%)2 1.30-1.32 (1.31) 1.22-1.23 (1.23) 1.34-1.34 (1.34) 2.61-2.61 (2.61) 1.59-1.59 (1.59)
A. indicus (&&)2 1.27-1.28 (1.27) 1.21-1.27 (1.24) 1.34-1.38 (1.36) 2.64-2.69 (2.67) 1.58-1.59 (1.59)
A. latus (%%)10 1.32-1.40 (1.36) 1.36-1.40 (1.38) 1.29-1.34 (1.31) 2.77-2.96 (2.84) 1.54-1.62 (1.56)
A. latus (&&)10 1.33-1.43 (1.37) 1.35-1.40 (1.38) 1.30-1.36 (1.33) 2.77-2.92 (2.83) 1.51-1.58 (1.55)
291
%&
Figs. 1
-
3. Anthracus derogatus (Walker). Habitus, head, pronotum and orignal labels. (1
-
3) Holotype.
292
Figs. 4
-
6. Anthracus furvus (Andrewes). Habitus, head and pronotum. (4) China, Yunnan, Xue Shan; (5) Paratype; (6) Vietnam, Sa Pa
293
Abbildungslegenden:
Figs. 1
-
3. Anthracus derogatus (Walker). Habitus, head, pronotum and
orignal labels. (1
-
3) Holotype.
Figs. 4
-
6. Anthracus furvus (Andrewes). Habitus, head and pronotum.
(4) China, Yunnan, Xue Shan; (5) Paratype; (6) Vietnam, Sa Pa
Figs. 7
-
9. Anthracus wrasei nov. sp. Habitus, head and pronotum. (7-8)
Paratypes, China, Yunnan, Er Hai Lake; (9) Holotype.
Figs. 10
-
12. Anthracus glabrus (Louwerens). Habitus, head and prono-
tum. (10, 12) Bali, Tamblingan Lakes; (11) Holotype.
Figs. 13
-
15. Anthracus schuhi nov. sp. Habitus, head and pronotum.
(13) Holotype; (14
-
15) Paratypes.
Figs. 16
-
18. Anthracus indicus nov. sp. Habitus, head and pronotum.
(16) Holotype; (17
-
18) Paratypes.
Figs. 19
-
21. Anthracus latus nov. sp. Habitus, head and pronotum. (19)
Holotype; (20) Paratype, S Vietnam, 12 km N Dalat; (21) Paratype,
Thailand, Ban San Pakia.
Figs. 22
-
29. Median lobe of aedoeagus, lateral and dorsal aspect.
(22
-
24) Anthracus derogatus (Walker), Holotype; (25
-
29) A. furvus
(Andrewes) (25, 28 Yunnan, Xue Shan; 26 Yunnan, Er Hai Lake; 27
Yunnan, Jizu Mts.; 29 S Vietnam, 12 km N Dalat).
Figs. 30
-
35. Anthracus furvus (Andrewes). Median lobe of aedoea-
gus lateral and dorsal aspect. (30) N Vietnam, Sapa; (31) Paratype,
Myanmar, Kambaiti; (32) Yunnan, Bangma Shan; (33) Yunnan, W Dali;
(34
-
35) S Vietnam, 12 km N Dalat.
Figs. 36
-
41. Anthracus wrasei nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (36, 39) Holotype; (37, 40, 41) Paratype, Yunnan,
Er Hai Lake; (38) Paratype, Yunnan, Xue Shan.
Figs. 42
-
48. Anthracus glabrus (Louwerens). Median lobe of ae-
doeagus, lateral and dorsal aspect. (42
-
44, 46
-
48) Bali, Tamblingan
Lakes;(45) Holotype.
Figs. 49
-
54. Anthracus schuhi nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (50, 52) Holotype; (49, 51, 53, 54) Paratypes.
Figs. 55
-
58. Anthracus indicus nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (55
-
56) Holotype; (57
-
58) Paratype.
Figs. 59
-
65. Anthracus latus nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (59) Paratype, Yunnan, Guo Men Shan; (60)
Holotype; (61) Paratype, Thailand, Doi Angkhang; (62
-
64) Paratypes, S
Vietnam, 12 km N Dalat; (65) Paratype, Thailand, Mae Salong.
Figs. 66
-
70. Ligula, dorsal aspect. (66) Anthracus furvus (Andrewes);
(67) A. glabrus (Louwerens); (68) A. indicus nov. sp.; (69) A. schuhi nov.
sp.; (70) A. latus nov. sp.
Figs. 71
-
75. Hemisternite and gonocoxa. (71) Anthracus furvus
(Andrewes); (72) A. wrasei nov. sp.; (73) A. indicus nov. sp.; (74) A.
glabrus (Louwerens); (75) A. latus nov. sp.
Figs. 76
-
77. Distribution. (76) Anthracus furvus (Andrewes); (77) A.
wrasei nov. sp. (triangles) and A. latus nov. sp. (circles).
Figs. 78
-
79. Distribution. (78) Anthracus indicus nov. sp. (black star);
A. schuhi nov. sp. (white circle); A. derogatus (Walker) (black square);
(79) A. glabrus (Louwerens).
.
Figs. 7
-
9. Anthracus wrasei nov. sp. Habitus, head and pronotum. (7-8) Paratypes, China, Yunnan, Er Hai Lake; (9) Holotype.
294
Figs. 10
-
12. Anthracus glabrus (Louwerens). Habitus, head and pronotum. (10, 12) Bali, Tamblingan Lakes; (11) Holotype.
295
Figs. 13
-
15. Anthracus schuhi nov. sp. Habitus, head and pronotum. (13) Holotype; (14
-
15) Paratypes.
296
Figs. 16
-
18. Anthracus indicus nov. sp. Habitus, head and pronotum. (16) Holotype; (17
-
18) Paratypes.
297
Figs. 19
-
21. Anthracus latus nov. sp. Habitus, head and pronotum. (19) Holotype; (20) Paratype, S Vietnam, 12 km N Dalat; (21) Paratype, Thailand,
Ban San Pakia.
298
Figs. 22
-
29. Median lobe of aedoeagus, lateral and dorsal aspect. (22
-
24) Anthracus derogatus (Walker), Holotype; (25
-
29) A. furvus (Andrewes) (25, 28 Yunnan, Xue Shan; 26 Yunnan, Er Hai Lake; 27 Yunnan,
Jizu Mts.; 29 S Vietnam, 12 km N Dalat).
299
Figs. 30
-
35. Anthracus furvus (Andrewes). Median lobe of aedoeagus lateral and dorsal aspect. (30) N Vietnam, Sapa; (31) Paratype, Myanmar, Kambaiti; (32) Yunnan, Bangma Shan; (33) Yunnan, W Dali; (34
-
35)
S Vietnam, 12 km N Dalat.
300
Figs. 36
-
41. Anthracus wrasei nov. sp. Median lobe of aedoeagus, lateral and dorsal aspect. (36, 39) Holotype; (37, 40, 41) Paratype, Yunnan, Er Hai Lake; (38) Paratype, Yunnan, Xue Shan.
301
Figs. 42
-
48. Anthracus glabrus (Louwerens). Median lobe of aedoeagus, lateral and dorsal aspect. (42
-
44, 46
-
48) Bali, Tamblingan Lakes;(45) Holotype.
302
Figs. 49
-
54. Anthracus schuhi nov. sp. Median lobe of aedoeagus, lateral and dorsal aspect. (50, 52) Holotype; (49, 51, 53, 54) Paratypes.
303
Figs. 55
-
58. Anthracus indicus nov. sp. Median lobe of aedoeagus, lateral and dorsal aspect. (55
-
56) Holotype; (57
-
58) Paratype.
304
Figs. 59
-
65. Anthracus latus nov. sp. Median lobe of aedoeagus, lateral and dorsal aspect. (59) Paratype, Yunnan, Guo Men Shan; (60) Holotype; (61) Paratype, Thailand, Doi Angkhang; (62
-
64) Paratypes, S
Vietnam, 12 km N Dalat; (65) Paratype, Thailand, Mae Salong.
305
Figs. 66
-
70. Ligula, dorsal aspect. (66) Anthracus furvus (Andrewes); (67) A. glabrus (Louwerens); (68) A. indicus nov. sp.; (69) A. schuhi nov. sp.; (70)
A. latus nov. sp.
306
Figs. 71
-
75. Hemisternite and gonocoxa. (71) Anthracus furvus (Andrewes); (72) A. wrasei nov. sp.; (73) A. indicus nov. sp.; (74) A. glabrus (Louwerens);
(75) A. latus nov. sp.
307
Abbildungslegenden:
Figs. 1
-
3. Anthracus derogatus (Walker). Habitus, head, pronotum and
orignal labels. (1
-
3) Holotype.
Figs. 4
-
6. Anthracus furvus (Andrewes). Habitus, head and pronotum.
(4) China, Yunnan, Xue Shan; (5) Paratype; (6) Vietnam, Sa Pa
Figs. 7
-
9. Anthracus wrasei nov. sp. Habitus, head and pronotum. (7-8)
Paratypes, China, Yunnan, Er Hai Lake; (9) Holotype.
Figs. 10
-
12. Anthracus glabrus (Louwerens). Habitus, head and prono-
tum. (10, 12) Bali, Tamblingan Lakes; (11) Holotype.
Figs. 13
-
15. Anthracus schuhi nov. sp. Habitus, head and pronotum.
(13) Holotype; (14
-
15) Paratypes.
Figs. 16
-
18. Anthracus indicus nov. sp. Habitus, head and pronotum.
(16) Holotype; (17
-
18) Paratypes.
Figs. 19
-
21. Anthracus latus nov. sp. Habitus, head and pronotum. (19)
Holotype; (20) Paratype, S Vietnam, 12 km N Dalat; (21) Paratype,
Thailand, Ban San Pakia.
Figs. 22
-
29. Median lobe of aedoeagus, lateral and dorsal aspect.
(22
-
24) Anthracus derogatus (Walker), Holotype; (25
-
29) A. furvus
(Andrewes) (25, 28 Yunnan, Xue Shan; 26 Yunnan, Er Hai Lake; 27
Yunnan, Jizu Mts.; 29 S Vietnam, 12 km N Dalat).
Figs. 30
-
35. Anthracus furvus (Andrewes). Median lobe of aedoea-
gus lateral and dorsal aspect. (30) N Vietnam, Sapa; (31) Paratype,
Myanmar, Kambaiti; (32) Yunnan, Bangma Shan; (33) Yunnan, W Dali;
(34
-
35) S Vietnam, 12 km N Dalat.
Figs. 36
-
41. Anthracus wrasei nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (36, 39) Holotype; (37, 40, 41) Paratype, Yunnan,
Er Hai Lake; (38) Paratype, Yunnan, Xue Shan.
Figs. 42
-
48. Anthracus glabrus (Louwerens). Median lobe of ae-
doeagus, lateral and dorsal aspect. (42
-
44, 46
-
48) Bali, Tamblingan
Lakes;(45) Holotype.
Figs. 49
-
54. Anthracus schuhi nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (50, 52) Holotype; (49, 51, 53, 54) Paratypes.
Figs. 55
-
58. Anthracus indicus nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (55
-
56) Holotype; (57
-
58) Paratype.
Figs. 59
-
65. Anthracus latus nov. sp. Median lobe of aedoeagus, lat-
eral and dorsal aspect. (59) Paratype, Yunnan, Guo Men Shan; (60)
Holotype; (61) Paratype, Thailand, Doi Angkhang; (62
-
64) Paratypes, S
Vietnam, 12 km N Dalat; (65) Paratype, Thailand, Mae Salong.
Figs. 66
-
70. Ligula, dorsal aspect. (66) Anthracus furvus (Andrewes);
(67) A. glabrus (Louwerens); (68) A. indicus nov. sp.; (69) A. schuhi nov.
sp.; (70) A. latus nov. sp.
Figs. 71
-
75. Hemisternite and gonocoxa. (71) Anthracus furvus
(Andrewes); (72) A. wrasei nov. sp.; (73) A. indicus nov. sp.; (74) A.
glabrus (Louwerens); (75) A. latus nov. sp.
Figs. 76
-
77. Distribution. (76) Anthracus furvus (Andrewes); (77) A.
wrasei nov. sp. (triangles) and A. latus nov. sp. (circles).
Figs. 78
-
79. Distribution. (78) Anthracus indicus nov. sp. (black star);
A. schuhi nov. sp. (white circle); A. derogatus (Walker) (black square);
(79) A. glabrus (Louwerens).
.
Figs. 76
-
77. Distribution. (76) Anthracus furvus (Andrewes); (77) A. wrasei nov. sp. (triangles) and A. latus nov. sp. (circles).
308
Figs. 78
-
79. Distribution. (78) Anthracus indicus nov. sp. (black star); A. schuhi nov. sp. (white circle); A. derogatus (Walker) (black square);
(79) A. glabrus (Louwerens).
.
