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The paper presents new records for 19 vascular plant species from 14 Eurasian countries. Two taxa (Siphonostegia chinensis and Utricularia macrorhiza) are reported from Russia, two (Achnatherum botschantzevii and Stipa zalesskii) from Kyrgyzstan, one (Allium petraeum) from Uzbekistan, three (Crambe orientalis, Eleocharis mamillata and Geranium pratense L. fater pratense. sergievskajae) from Kazakhstan, two (Atriplex crassifolia and Petrosimonia brachyphylla) from China, one (Crambe orientalis) from Tajikistan, one (Stipa krylovii) from India, one (Agrostis lazica) from Iraq, two (Orobanche coerulescens and Orobanche zajaciorum) from Armenia, one (Phelipanche lavandulacea) from Montenegro, one (Panicum riparium) from Bosnia and Herzegovina, Romania and Sweden, one (Sporobolus vaginiflorus) from Bosnia and Herzegovina and two (Ranunculus penicillatus subsp. pseudofluitans and Scutellaria minor) from Poland. Three of the taxa presented (Crambe orientalis, Panicum riparium and Sporobolus vaginiflorus) are regarded as alien to the studied areas, whereas the remaining 16 are native elements to the flora of the countries. For each species, synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition and differentiation of the species from the most similar taxa occurring in a given country, as well as a list of recorded localities (often far from the previously known areas) are presented.

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... Stipa richteriana, on the other hand, is a native and permanent element of the Chinese flora collected from few localities in western China (see chapter New records, below). Some other taxa, treated in the Flora of China (wu & Phillips 2006) at the level of variety, such as S. klemenzii roshevitz (1924: 12) and S. krylovii roshevitz (1929: 379) are currently regarded as distinct species (Tzvelev 1976, lomonosova 1990, Nobis 2014, Nobis et al. 2016b. In a result of previous field investigation in poorly explored regions of China and revision of herbarium material collected from the country, species unlisted so far in the Flora of China were recorded and new taxa were described. ...
... Wilenskii (lectotype selected by Tzvelev (1976: 587), le!) Stipa zalesskii is a widely distributed taxon extending from central europe to the central asia (kyrgyzstan, kazakhstan, russia) (Tzvelev 1976, 2012, Martinovský 1980, Freitag 1985, Conert 1998, Nobis et al. 2016b. Stipa zalesskii was reported from China by Tzvelev (1968) and Cui (1996), but it was omitted by wu & Phillips (2006). ...
... This is the easternmost locality of this taxon. Stipa zalesskii belongs to the section Stipa and is easily distinguishable from other Chinese feather grasses of the section Stipa by having anthecia 17-23 mm long, ventral line of hairs on lemma reaching the base of the awn, vegetative shoots with sheaths shortly pubescent and leaves with a mixture of short and long hairs on the adaxial surface and the abaxial surface scabrous due to hard prickles with a more or less dense admixtue of spinules and/or short hairs (Nobis et al. 2016b). ...
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Stipa dickorei sp. nov. from the Western Tibetan Plateau (China) is described. The new species is morphologically similar to S. regeliana, but they differ from each other in the length of ligules of vegetative shoots. Stipa dickorei is also similar to S. aliena, however they differ in the shape of panicle, which is contracted with straight branches in S. dickorei, and lax with flexuous branches in S. aliena. Images of macromorphological and micromorphological structures of the new taxon are provided. Additionally, new records of S. borysthenica, S. richteriana, and S. zalesskii, species not listed in the recent Flora of China, as well as a checklist of Chinese feather grasses are also presented.
... During field exploration across the vast area of 11 European and Asian countries as well as during taxonomic revisions based on herbarium materials of different groups of vascular plants, the authors found some species that are new to the floras of particular countries or their significant regions (provinces or republics). The paper is the continuation of the previous works (Nobis et al. 2014a(Nobis et al. , 2014c(Nobis et al. , 2015a(Nobis et al. , 2015b(Nobis et al. , 2016(Nobis et al. , 2018, which similarly to the works of Sukhorukov et al. (2016) and Sukhorukov et al. (2017) are dedicated to new national and regional vascular plant records, to broaden our knowledge on their distribution and taxonomy. The aim of this paper is to report new it occurs in a wide altitudinal rangefrom 1000 m up to 4100 m (Li and Hedge 1994). ...
... The holoparasitic Orobanchaceae in Armenia are represented by one species of Cistanche, two Diphelypaea, 15 Phelipanche and 18 Orobanche (Piwowarczyk et al. 2019). Several species have been also included in regional contributions (Aghababyan 2013;Nobis et al. 2016Nobis et al. , 2018Piwowarczyk et al. 2017a) and new species have been described as well (Piwowarczyk, Sánchez Pedraja, and Moreno Moral 2017b;Piwowarczyk et al. 2017cPiwowarczyk et al. , 2018aPiwowarczyk et al. , 2018bPiwowarczyk et al. , 2018c. ...
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The paper presents new records of 24 vascular plant species from 11 Eurasian countries. One taxon (Orobanche laxissima) is reported from Armenia; one (Epipactis condensata) from Azerbaijan; two (Phragmites americanus, Polygala multicaulis) from Belarus; one (Stipa cauca-sica) from Egypt; one (Puccinellia hauptiana) from Kyrgyzstan; three (Aquilegia xinjiangensis, Geranium saxatile, Ranunculus songaricus) from Mongolia; one (Stipa roborowskyi) from Pakistan; three (Echinochloa muricata, Erigeron acris subsp. podolicus, Hypericum majus) from Poland; six from Russia, whereof one (Zanthoxylum armatum) from the European part of Russia and five (Chaerophyllum aureum, Elsholtzia densa, Poa compressa, Ranunculus subrigi-dus, Viola sororia) from the Asian part of Russia; two (Ludwigia repens, Sagittaria latifolia) from Slovakia; and three (Rubus ambrosius, Rubus camptostachys, Rubus perrobustus) from Ukraine. For each species, synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition and differentiation of the species from the most similar taxa occurring in a given country, as well as a list of recorded localities (often far from the previously known areas), are presented. ARTICLE HISTORY
... According to Valdés et al. (2009) P. barbipulvinatum is widespread naturalized alien in C and E Europe. For the Balkan countries and neighboring Hungary species is reported for Hungary (Király et al. 2009), Slovenia (Jogan & Martinčić 2015), Croatia (Király & Alegro 2015), Romania and Bosnia and Herzegovina (Nobis et al. 2016), under the name P. riparium respectively. ...
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The paper presents three new nomenclature combinations related to representatives of the orchid family. In addition, 14 taxa (species, subspecies and hybrids) of vascular flora which are newly registered for Serbia (of which 8 are autochthonous and 6 allochtonous plants) are presented. A confirmation of old or unreliable literature records for four autochthonous and one allochthonous plant in the flora of Serbia has been done. Four plant taxa are a novelty for proper Serbia, Vojvodina, or Kosovo and Metohija, and for 7 plants, presence in certain administrative units in Serbia has been confirmed. In the last part of the paper, there is a review of 9 disputed taxa for our country. Key words: vascular flora, checklist, nomenclature, taxonomy, Serbia
... Over the last four decades, both taxa also appeared in Austria (Melzer 1978(Melzer , 1986Walter 2002;Fischer et al. 2008), and they spread further in Croatia (Horvatić & Gospodarić 1960;Marković 1973;Melzer 1985;Melzer & Bregant 1990), Italy (Pignatti 1982;Melzer 1983;Poldini 2002;Celesti Grapow 2010), and W Hungary (Kiraly & Hohla 2015). Later on, sporadic sampling in the countries neighbouring Slovenia revealed the presence of both taxa in more regions of Croatia (e.g., Slavonia, Istria, Gorski Kotar), while S. vaginiflorus alone was also found in Montenegro (Podgorica, Stešević & Jogan 2006) and Bosnia (Nobis et al. 2016). In addition, the first records of S. vaginiflorus for Serbia (Srem) and Herzegovina are herein reported. ...
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Systematic field sampling revealed that within 50 years since the first records in Slovenia, Sporobolus neglectus and S. vaginiflorus became widespread. They are two superficially similar N American annual grass species with cleistogamous spikelets and similar ecology that are confined to dry ruderal places in their European secondary range, especially along roads. The oldest records of naturalised populations of both species in Europe date back to the 1950s, when both were found for the first time in the Vipava valley (SW Slovenia). They spread slowly in the next decades to NE Italy, N Croatia, and S Austria until recently, when an explosive expansion has been observed along almost all the main roads in lowland and montane Slovenia. In addition to that, one or both of them have recently been recorded scattered in SE Europe (Hungary, Serbia, B&H, Montenegro) and W Europe (France, Switzerland). Sporobolus vaginiflorus is herein reported for the first time for Serbia, Herzegovina (in B&H), and Slavonia (in Croatia). © 2017 Institute of Botany and Botanical Garden Jevremovac, Belgrade.
... The flora of this region includes over 40 species of holoparasitic Orobanchaceae, belonging to Orobanche, Phelipanche Pomel (1874: 102), Cistanche Hoffmannsegg & Link (1809: 319), and Diphelypaea Nicolson (1975: 654) (Novopokrovskij & Tzvelev 1958, Takhtajan 1987, Tsaturyan & Gevorgyan 2009, Domina et al. 2010, Aghababyan 2013. Many of them are endemic, parasitising endemic hosts, or have their distributional center there, and through detailed research in recent years, several have been described or reported (Piwowarczyk in Nobis 2016, 2018Piwowarczyk et al. 2017bPiwowarczyk et al. , 2017cPiwowarczyk et al. , 2017dPiwowarczyk et al. , 2018b. ...
Article
Orobanche javakhetica (Orobanchaceae) is described as a new, probably endemic, species from the Lesser Caucasus in Armenia. It grows on a subalpine meadow, where it parasitises Lomelosia caucasica (Dipsacaceae). The newly-described species is very characteristic and different from other known Orobanche, however some morphological similarity may exist with species from the Orobanche subsect. Curvatae, particularly with species of the Orobanche series Krylowianae. A detailed description, illustrations, a comparison with the most similar species with identification key, and phylogenetic analysis are provided.
... Perennial grass, densely tufted; culms 20-60(90) cm with 3-4 nodes, glabrous below the nodes. Cauline leaves: sheaths shorter then internodes, glabrous or slightly scabrous, upper sheaths up to 10 mm width, encompassing the panicle in the flowering period, and for the most part during fruiting; blades glabrous to slightly scabrous, to 0.5 mm in diameter; ligules not equal with approximate range from bottom to top I -0.5-0.6 mm; II -1-1.5 mm; III -2.7-3.5 mm; IV -5.5-7 mm (often broken in herbarium China (Gansu, Hebei, Nei Mongol, Ningxia, Qinghai, Shanxi, Xinjiang, Xizang;Lu & Wu 1996;Wu & Phillips 2006), Mongolia (Grubov 1955), Kyrgyzstan (Lazkov & Sultanova 2011), Tajikistan (west Pamir;Ikonnikov 1979), North India (Ladakh; Nobis et al. 2016c). Figure 2. Specimens examined (new records to the flora of Nepal): Nepal. ...
... The paper is the continuation of previous works (Nobis et al. , 2016(Nobis et al. , 2017, which similarly to the works of Sukhorukov et al. (2016Sukhorukov et al. ( , 2017, are dedicated to new national and regional vascular plant records, to broaden our knowledge on their distribution and taxonomy. The aim of this paper is to report new records for 29 vascular plant species: 16 from Asian and 13 from European countries. ...
Article
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The paper presents new records for 29 vascular plant species from 12 Eurasian countries. Seven taxa (Agave sisalana, Austrocylindropuntia subulata, Lagurus ovatus subsp. nanus, Opuntia stricta, Orobanche serbica, Oxalis articulata, Vitis × instabilis) are reported from Albania, one (Allium carinatum) from the Europaean part of Russia, six (Dipsacus fullonum, Gagea shmakoviana, Mentha × dalmatica, Thymus indigirkensis, Thymus sergievskajae, Viola × sukaczewii) from the Asian part of Russia, two (Agrostis sozanensis, Poa mustangensis) from China, two (Carex muskingumensis, Sedum rubens) from Poland, two (Crataegus macrocarpa, Dactylorhiza lapponica) from Romania, two (Oxygraphis delavayi, Ranunculus eryuanensis) from Nepal, two (Erigeron bonariensis, Ophioglossum vulgatum) from Tajikistan, one (Ranunculus olgae) from Kazakhstan, one (Najas guadelupensis) from Hungary, one (Orobanche bartlingii) from Armenia, one (Crataegus petrodavisii) from Azerbaijan and one (Amaranthus powellii) from Georgia. For each species, synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition, and differentiation the species from the most similar taxa occurring in a given country, as well as a list of recorded localities (often far from the previously known areas) are presented.
... Unfortunately, misidentification of specimens at species level is frequent in herbarium materials and in the literature, particularly in difficult groups like grasses. The first author and her collaborators revealed several such cases in Calamagrostis and Deyeuxia (Paszko & Ma 2011;Paszko 2014Paszko , 2015Paszko & Soreng 2013;Paszko et al. , 2016a as well as in Agrostis (Nobis et al. 2016;Paszko & Liu 2016). ...
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New taxonomic and distributional data are provided for three eastern Asian species of Calamagrostis (Poaceae, Agrostidinae) in Vietnam: C. abnormis (Hook. f.) Shukla, C. extremiorientalis (Tzvel.) Prob. and C. tripilifera Hook. f. The new national record of C. extremiorientalis is reported here for Vietnam, where it was previously misidentified as C. elatior (Griseb.) A. Camus or Deyeuxia petelotii Hitchc. (= C. abnormis). The presence of C. tripilifera and C. abnormis in Vietnam is confirmed. Additional new provincial records of C. extremiorientalis from Japan’s Bonin Islands (also known as the Ogasawara Islands) and C. tripilifera from Daba Mountains in the northern Chongqing Municipality (Central China) are reported here. A key for identification of Calamagrostis species in Vietnam, nomenclatural data and information about geographical, altitudinal and habitat distributions are provided. Maps showing their complete distribution in eastern Asia are presented.
... Siphonostegia chinensis is a new native species for Siberia. The species has been found in Trans-Baikal Territory (3 km and 17 km from the border with China, Inner Mongolia Autonomous Region) on mid-mountain lots of the southeastern spurs of Gazimur Range, in the lower course of the river Budyumkan (left tributary of the river Argun) (Nobis, et al., 2016). As per the zonation of Asian Russia (Baikov, et al., 2012), this territory belongs to the Baikal hemiboreal province. ...
Article
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Pollen morphology of Siphonostegia chinensis (family Orobanchaceae), collected in the Trans- Baikal Territory (Baikal hemiboreal province) was studied and illustrated with light microscopy (LM) and scanning electron microscopy (SEM). On the Trans-Baikal Territory species is found on the border area. The distinguishing feature of the pollen grains under study is the presence of 4-colpate and 6-colpate ones, which other authors have not indicated for this species.
... Perennial grass, densely tufted; culms 20-60(90) cm with 3-4 nodes, glabrous below the nodes. Cauline leaves: sheaths shorter then internodes, glabrous or slightly scabrous, upper sheaths up to 10 mm width, encompassing the panicle in the flowering period, and for the most part during fruiting; blades glabrous to slightly scabrous, to 0.5 mm in diameter; ligules not equal with approximate range from bottom to top I -0.5-0.6 mm; II -1-1.5 mm; III -2.7-3.5 mm; IV -5.5-7 mm (often broken in herbarium China (Gansu, Hebei, Nei Mongol, Ningxia, Qinghai, Shanxi, Xinjiang, Xizang;Lu & Wu 1996;Wu & Phillips 2006), Mongolia (Grubov 1955), Kyrgyzstan (Lazkov & Sultanova 2011), Tajikistan (west Pamir;Ikonnikov 1979), North India (Ladakh; Nobis et al. 2016c). Figure 2. Specimens examined (new records to the flora of Nepal): Nepal. ...
Article
Stipa krylovii is newly reported for the flora of Nepal, and this is the most southerly location yet found for this species. A full description of S. krylovii is included, along with illustrations, notes on its taxonomy and a distribution map.
... There were also published many relevant scientific papers and some new taxa, not known in our flora, were identified, or discovered (e.g. Bąk et al. 2014;Kruk et al. 2014Kruk et al. , 2015Krajewski, Płachno 2015;Pliszko 2015;Szczęśniak et al. 2015;Nobis et al. 2016). All this data has been taken into account when compiling the present list. ...
... The flora of Armenia includes over 40 species of holoparasitic Orobanchaceae belonging to Orobanche, Phelipanche, Cistanche Hoffmannsegg & Link (1809: 319), and Diphelypaea Nicolson (1975: 654) (Novopokrovskij & Tzvelev 1958, Takhtajan 1987, Domina et al. 2010, Aghababyan 2013, Piwowarczyk in Nobis et al. 2016, Piwowarczyk et al. 2017. Data concerning the distribution and host plants of these holoparasites in Armenia and beyond are still limited, and further study, including taxonomic revision and field research, is required. ...
Article
Phelipanche sevanensis (Orobanchaceae) is described as a new species from the Lesser Caucasus in Armenia. It grows on steep rocky slopes around Lake Sevan in semi-arid mountain steppe communities, where it parasitises Heracleum trachyloma, a Caucasian endemic. A detailed description, illustrations, and a comparison with the most similar species, especially P. heldreichii, are provided. We also present nomenclatural notes about P. coelestis, P. hohenackeri, P. persica and P. simplex. Three lectotypes are designated, and two new combinations are proposed.
... There were also published many relevant scientific papers and some new taxa, not known in our flora, were identified, or discovered (e.g. Bąk et al. 2014;Kruk et al. 2014Kruk et al. , 2015Krajewski, Płachno 2015;Pliszko 2015;Szczęśniak et al. 2015;Nobis et al. 2016). All this data has been taken into account when compiling the present list. ...
... The lead author and collaborators in a series of papers have clarified numerous lower-level taxonomic issues among several Eurasian taxa of Agrostis L. (Paszko 2012(Paszko , 2014bPaszko & Pendry 2013;Paszko & Soreng 2013;Paszko et al. 2015), described new species (Paszko 2014a), and published new records (Nobis et al. 2014(Nobis et al. , 2016. Recently the first author located eight duplicates (KUN, PE) representing four separate collection events of Agrostis, incorrectly identified as A. clavata Trin. ...
Article
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The first confirmed records of Agrostis scabra Willd. are reported from a northeastern Chinese province, Jilin. This extends the native range of the species into China. The recent Flora of China does not include A. scabra. The detailed morphological characters, diagnostic comparisons and habitat are presented, along with a distribution map of A. scabra in East and Northeast Asia.
... It occupies desert and semi-desert habitats, frequently on rocky or stony grounds. In Africa it has been reported from Libya, Chad, Egypt, Sudan, Eritrea, Ethiopia and Somalia (Abdallah and de Wit 1978; Miller 1984; Thulin 1993; de Wit 2000; Mart?n-Bravo 2011; Brundu and Camarda 2013; Dobignard and Chatelain 2013; Darbyshire et al. 2014; APD 2016; Hassler 2016). Djibouti was listed by Hassler (2016) and there seem to be records in APD (2016), but the species is not explicitly indicated as present in the country. ...
Article
This paper presents notes on new records and the taxonomy of 12 vascular plant species from several continental African countries and Madagascar. Datura wrightii (Solanaceae) is a new species for Africa reported from Morocco. Amaranthus muricatus (Amaranthaceae) is reported for the first time in Tunisia, Barleria oenotheroides (Acanthaceae) – from Central African Republic and Senegal, Burkea africana (Fabaceae) – from Guinea-Bissau, Caylusea abyssinica (Resedaceae) – from Zambia, Digitaria ciliaris (Poaceae) – from Morocco. New data on distribution in Africa is provided for Camptorrhiza strumosa (Colchicaceae) and four Resedaceae taxa:Caylusea hexagyna, Ochradenus baccatus, Reseda arabica and Reseda villosa. The taxonomy of the enigmatic Madagascan endemic Mollugo caespitosa (Molluginaceae) is reviewed, and it is synonymized with Mollugo decandra. The lectotypes are designated for both Mollugo species.
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Ranunculus species grouped in the section Batrachium (water crowfoots) indicate the most valuable fluvial habitat in Europe under the EU Habitats Directive, and recognition of their environmental preferences plays a crucial role in efficient conservation. Moreover, water crowfoots are used in various monitoring systems throughout Europe. The aim of this study was to identify the distribution patterns of two common water crowfoot species ( Ranunculus fluitans , R. circinatus ) and their hybrid ( R. circinatus × R. fluitans ) in relation to environmental variables in rivers. Various chemical and hydromorphological parameters were estimated for 54 river sites where 58 different Batrachium populations were recorded. Our study revealed the most distinct positive reaction of R. fluitans to rhithral conditions characterized by rapid currents and coarse substrate. R. circinatus × R. fluitans preferred larger channels, although this taxon also flourished in rhithral rivers. The preferences of R. circinatus were not so evident confirming its occurrence under a wide range of habitat conditions. The ecological reaction of the hybrid was compared with its parental species, revealing their ecological specificity, which is useful in designing appropriate conservation programmes.
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Central Asia, which is composed of five countries e Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, is an important global biodiversity hotspot, but a botanically poorly-known part of Asia. As part of the Mapping Asia Plants (MAP) project, we reviewed the history of botanical investigations, floristic works and publications in this region, as well as key floras, checklists, herbaria and online databases in Central Asia. The published Floras and Checklists dating back to 1851 lay a solid foundation for botanical research in Central Asia today. Eight Floras (61 vol) and four Cheklists for this region were listed and introduced in this paper. At the regional level, the full eleven volumes Conspectus Florae Asiae Mediae, containing 9341 species of vascular plants belonging to 1245 genera and 161 families across Central Asia, were published during 1968e2015. At the country level, the national Floras of five countries which were published during 1932 e1991, listed 5631, 3576, 4445, 2607 and 4148 species of vascular plants in Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, respectively. The species counts were updated to 5658, 3927 and 2800 in the national Checklists of vascular plant in Kazakhstan (1999), Kyrgyzstan (2014) and Turkmenistan (1988), respectively. We also simply introduced the key herbaria in this region which contain about 2.5 million specimens in total. However, some major gaps and limitations, i.e. the outdate publications, national Floras mainly in local language, the massive non-digitized specimens, and the lack of an online database or platform, should be addressed in terms of their global applications.
Article
Full-text available
Central Asia, which is composed of five countries – Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, is an important global biodiversity hotspot, but a botanically poorly-known part of Asia. As part of the Mapping Asia Plants (MAP) project, we reviewed the history of botanical investigations, floristic works and publications in this region, as well as key floras, checklists, herbaria and online databases in Central Asia. The published Floras and Checklists dating back to 1851 lay a solid foundation for botanical research in Central Asia today. Eight Floras (61 volumes) and four Cheklists for this region were listed and introduced in this paper. At the regional level, the full eleven volumes Conspectus Florae Asiae Mediae, containing 9341 species of vascular plants belonging to 1245 genera and 161 families across Central Asia, were published during 1968 to 2015. At the country level, the national Floras of five countries which were published during 1932 to 1991, listed 5631, 3576, 4445, 2607 and 4148 species of vascular plants in Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, respectively. The species counts were updated to 5658, 3927 and 2800 in the national Checklists of vascular plant in Kazakhstan (1999), Kyrgyzstan (2014) and Turkmenistan (1988), respectively. We also simply introduced the key herbaria in this region which contain about 2.5 million specimens in total. However, some major gaps and limitations, i.e. the outdate publications, national Floras mainly in local language, the massive non-digitized specimens, and the lack of an online database or platform, should be addressed in terms of their global applications.
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The genus Stipa L. comprises over 150 species, all native to the Old World, where they grow in warm temperate regions throughout Europe, Asia, and North Africa. It is one of the largest genera in the family Poaceae in Middle Asia, where one of its diversity hotspots is located. However, identification of Middle Asian Stipa species is difficult because of the lack of new, comprehensive taxonomic studies including all of the species recorded in the region. We present a critical review of the Mid-Asian representatives of Stipa, together with an identification key and taxonomic listing. We relied on both published and unpublished information for the taxa involved, many of which are poorly known. For each taxon, we present a taxonomic and nomenclatural overview, habitat preferences, distribution, altitudinal range, and additional notes as deemed appropriate. We describe four new nothospecies: S. ×balkanabatica M. Nobis & P. D. Gudkova, S. ×dzungarica M. Nobis, S. ×pseudomacroglossa M. Nobis, S. ×subdrobovii M. Nobis & A. Nowak, one subspecies S. caucasica Schmalh. subsp. nikolai M. Nobis, A. Nobis & A. Nowak, and eight varieties: S. araxensis Grossh. var. mikojanovica M. Nobis, S. caucasica var. fanica M. Nobis, P. D. Gudkova & A. Nowak, S. drobovii (Tzvelev) Czerep. var. jarmica M. Nobis, S. drobovii var. persicorum M. Nobis, S. glareosa P. A. Smirn. var. nemegetica M. Nobis, S. kirghisorum P. A. Smirn. var. balkhashensis M. Nobis & P. D. Gudkova, S. richteriana Kar. & Kir. var. hirtifolia M. Nobis & A. Nowak, and S. ×subdrobovii var. pubescens M. Nobis & A. Nowak. Additionally, 12 new combinations, Achnatherum haussknechtii (Boiss.) M. Nobis, A. mandavillei (Freitag) M. Nobis, A. parviflorum (Desf.) M. Nobis, Neotrinia chitralensis (Bor) M. Nobis, S. badachschanica Roshev. var. pamirica (Roshev.) M. Nobis, S. borysthenica Klokov ex Prokudin var. anomala (P. A. Smirn.) M. Nobis, S. holosericea Trin. var. transcaucasica (Grossh.) M. Nobis, S. kirghisorum P. A. Smirn. var. ikonnikovii (Tzvelev) M. Nobis, S. macroglossa P. A. Smirn. var. kazachstanica (Kotuchov) M. Nobis, S. macroglossa var. kungeica (Golosk.) M. Nobis, S. richteriana var. jagnobica (Ovcz. & Czukav.) M. Nobis & A. Nowak, and S. zalesskii Wilensky var. turcomanica (P. A. Smirn.) M. Nobis are proposed, and the lectotypes for 14 taxa (S. arabica Trin. & Rupr., S. bungeana Trin. ex Bunge, S. caspia K. Koch, S. ×consanguinea Trin. & Rupr., S. effusa Mez, S. ×heptapotamica Golosk., S. jacquemontii Jaub. & Spach., S. kungeica Golosk., S. margelanica P. A. Smirn., S. richteriana, S. rubentiformis P. A. Smirn., S. sareptana A. K. Becker, S. tibetica Mez, and Timouria saposhnikovii Roshev.) are designated. In Middle Asia the genus Stipa comprises 98 taxa, including 72 species, four subspecies, and 22 varieties. Of the 72 species of feather grasses, 23 are of hybrid origin (nothospecies). In Middle Asia, feather grasses can be found at elevations from (0 to)300 to 4500(to 5000) m, but most are montane species. The greatest species richness is observed at altitudes between 1000 and 2500 m. Nineteen species grow above 3000 m, but only nine above 4000 m. The number of taxa (species and subspecies) growing in each country also varies considerably, with the highest noted in Kazakhstan (42), Tajikistan (40), and Kyrgyzstan (35). Of the 76 taxa of Stipa (species and subspecies) recorded in Middle Asia, 41 are confined to the region, with some being known only from a single country or mountain range. Distribution maps of selected species are provided.
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The species of genus Allium growing naturally in Uzbekistan have been revised. 103 species and one subspecies are listed in a conspectus arranged according to the preliminary system of the genus by Kamelin (1973). Several nomenclatural changes have been made including the acceptation of a new subsect. Erectopetala.
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We report new floristic records of some rare Orobanche and Phelipanche species (Orobanchaceae) in Croatia. Orobanche salviae and O. alsatica are reported for the first time in 100 years, and O. laserpitii-sileris, and P. lavandulacea are rare species with only few localities known.
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The paper presents new records for 20 vascular plant species from 13 Eurasian countries. Four taxa (Calamagrostis sichuanensis, Klasea dissecta, Ptilagrostis milleri and Stipa klimesii) are reported from China, four (Aconogonon valerii, Carex siderosticta, Poa tanfiljewii and Potamogeton × subobtusus) from Russia, three (Amorpha fruticosa, Carduus acanthoides and Plantago minuta) from Tajikistan, two (Achillea sergievskiana and Delphinium barlykense) from Kazakhstan, one (Calamagrostis effusiflora) from Bhutan and India, one (Campanula wolgensis) from Mongolia, one (Orobanche coerulescens) from Georgia, two (Dysphania geoffreyi and Ptilagrostis milleri) from Nepal, one (Stipa × alaica) from Afghanistan, one (Stipa × manrakica) from Kyrgyzstan, one (Ranunculus × gluckii) from Poland and one (Sporobolus cryptandrus) from Italy. Four of the taxa presented (Amorpha fruticosa, Carduus acanthoides, Carex siderosticta and Sporolobus cryptandrus) are regarded as alien to the studied areas, whereas the remaining 16 are native elements to the flora of the countries. For each species synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition and differentiation of the species from the most similar taxa occurring in a given country, as well as a list of recorded localities (often far from the previously known areas) are presented. Based on the spikelet morphology, we have proposed in Ptilagrostis a new section Barkworthia M. Nobis, A. Nobis & A. Nowak, which comprises two species Ptilagrostis yadongensis and Ptilagrostris milleri, with distinctly unequal glumes, lemmas and paleas.
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The paper presents new records for 18 vascular plant species from six Eurasian countries. Three taxa (Lepidium densiflorum, Stipa czerepanovii, Xanthium albinum) are reported from Kazakhstan, one (Ranunculus schmakovii) from Mongolia, three (Dianthus campestris, Ranunculus kauffmannii, Viola suavis) from Poland, five (Eragrostis amurensis, Linum catharticum, Ludwigia peploides subsp. stipulacea, Pyrethrum mikeschinii, Solidago canadensis) from Tajikistan, five (Clinopodium menthifolium, Juncus effusus, Mollugo cerviana, Poa sphondylodes, Vulpia myuros) from Russia, and one (Orobanche alba subsp. xanthostigma) from Georgia. Ten of the taxa presented (Clinopodium menthifolium, Dianthus campestris, Eragrostis amurensis, Juncus effusus, Lepidium densiflorum, Mollugo cerviana, Solidago canadensis, Viola suavis, Vulpia myuros and Xanthium albinum) are regarded as alien to the studied areas, whereas the remaining eight are native elements to the flora of the countries. For each species, synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition and differentiation of the species from the most similar occurring in a given country, as well as a list of localities recorded (often far from the previously known areas), are presented.
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The paper presents new records for 18 vascular plant species from six Eurasian countries. Three taxa (Lepidium densiflorum, Stipa czerepanovii, Xanthium albinum) are reported from Kazakhstan, one (Ranunculus schmakovii) from Mongolia, three (Dianthus campestris, Ranunculus kauffmannii, Viola suavis) from Poland, five (Eragrostis amurensis, Linum catharticum, Ludwigia peploides subsp. stipulacea, Pyrethrum mikeschinii, Solidago canadensis) from Tajikistan, five (Clinopodium menthifolium, Juncus effusus, Mollugo cerviana, Poa sphondylodes, Vulpia myuros) from Russia, and one (Orobanche alba subsp. xanthostigma) from Georgia. Ten of the taxa presented (Clinopodium menthifolium, Dianthus campestris, Eragrostis amurensis, Juncus effusus, Lepidium densiflorum, Mollugo cerviana, Solidago canadensis, Viola suavis, Vulpia myuros and Xanthium albinum) are regarded as alien to the studied areas, whereas the remaining eight are native elements to the flora of the countries. For each species, synonyms, general distribution, habitat preferences, taxonomy with remarks on recognition and differentiation of the species from the most similar occurring in a given country, as well as a list of localities recorded (often far from the previously known areas), are presented.
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The Panicum capillare complex includes several taxa, among them P. capillare L., which is usually considered to be an established alien throughout Europe, whereas other species are recorded only as casuals. A new representative of the complex, P. riparium H. Scholz was described from Germany in 2002, and shortly after its description was recorded in several countries on the continent. In the course of herbarium revisions and recent fi eld studies the authors documented several localities of the species in Croatia as well. The paper presents a new key for the determination of Croatian species of the complex and anticipates the invasion of P. riparium in the sub-Mediterranean regions of the Balkan Peninsula.
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A new, probably endemic, species, Orobanche zajaciorum (Orobanchaceae) is described from Georgia, in the Lesser Caucasus. The species grows on rocky slopes and is a parasite Scutellaria sosnowskyi. It is a small pale pink-yellowish plant, characterised by strongly curved, geniculate, white-pink flowers. A detailed description, diagnosis, and illustrations are provided.
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Pollen grains of 450 samples of 25 species of the genus Orobanche and Phelipanche occurring in Central Europe were investigated using light and scanning electron microscopy. Palynological data on 18 species are reported here for the first time. The usefulness of micromorpho-logical studies on pollen of Orobanche and Phelipanche is demonstrated. Previous research showed the separation of Phelipanche from Orobanche, a finding which is also supported by pollen aperture type and exine ornamentation. The pollen of Phelipanche is tricolpate, while that of Orobanche is inaperturate. Our research showed that one of the species included so far within Orobanche, namely O. coerulescens, has tricolpate pollen with microreticulate sculpture. As a consequence of the finding of tricolpate pollen and intermediate macromorphological characteristics of this species between Orobanche and Phelipanche, it is proposed to transfer it to a new genus, Orobanchella, described in the paper. A comparison of the main characteristics distinguishing Orobanche, Phelipanche, Boulardia and Orobanchella is also presented and discussed.
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Orobanche coerulescens has a Eurasian distribution. The species is classified as extinct at most of its localities at the western limit of its range. Its populations are very scarce and critically endangered in Central Europe. This work presents the current distribution of O.coerulescens in Poland, based on a critical revision of herbarium and literature data as well as results of original field research, and reviews its distribution in Central Europe (partly in Eastern Europe). Habitats, plant communities, and migration routes of O.coerulescens in Central Europe are discussed. The species was initially known in Poland from now historical localities in Pomerania and the valley of the lower Vistula. In 2000-2011 it was recorded at 9 localities in Podlasie, the Małopolska Upland (Wyżyna Małopolska), and the Łódź Hills (Wzniesienia Łódzkie). Its abundance at the localities ranged from a few to over 1000 shoots. These are the largest populations of O.coerulescens at its western and north-western range limits.
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Pollen grains of 450 samples of 25 species of the genus Orobanche and Phelipanche occurring in Central Europe were investigated using light and scanning electron microscopy. Palynological data on 18 species are reported here for the first time. The usefulness of micromorphological studies on pollen of Orobanche and Phelipanche is demonstrated. Previous research showed the separation of Phelipanche from Orobanche, a finding which is also supported by pollen aperture type and exine ornamentation. The pollen of Phelipanche is tricolpate, while that of Orobanche is inaperturate. Our research showed that one of the species included so far within Orobanche, namely O. coerulescens, has tricolpate pollen with microreticulate sculpture. As a consequence of the finding of tricolpate pollen and intermediate macromorphological characteristics of this species between Orobanche and Phelipanche, it is proposed to transfer it to a new genus, Orobanchella, described in the paper. A comparison of the main characteristics distinguishing Orobanche, Phelipanche, Boulardia and Orobanchella is also presented and discussed.
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Panicum riparium, in Central Europe established, at least since 1990, on the Elbe and the Oder riversides, is described as new to science. Regarded as a neogen species, probably having originated in situ from the introduced NorthAmerican P. capillare, its relationships with several species of the New World P. capillare complex are discussed, results from comparative cultivation experiments presented, and a key to the most frequent species of this complex in Europe is provided.
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The first treatment of the family Chenopodiaceae for the flora of Nepal (Central and Eastern Himalaya) has been recently published (Sukhorukov & Kushunina 2014). However, after a detailed investigation of original material concerning Chenopodium pallidum Moquin-Tandon (1840: 30), which are part of Jacquemont’s collection from India (Herbarium P), we can state that all these specimens indeed belong to Atriplex Linnaeus (1753: 1052). According to Art. 11 of ICN (McNeill et al. 2012), the name Chenopodium pallidum appears to be an older name at specific rank for Atriplex schugnanica Iljin (1936: 123), and thus a new combination is proposed in the present paper. Besides, new Chenopodium species, previously named Chenopodium pallidum [now Atriplex pallida Moquin-Tandon (1840: 30)], is described from Nepal and dedicated to the prominent Japanese botanist Hiroshi Hara (Chenopodium harai Sukhor.).
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The book describes richness and diversity of Georgia's vegetation. Contrasting ecosystems coexist on the relatively small territory of the country and include semi-deserts in East Georgia, Colchic forests with almost sub-tropical climate in West Georgia and subnival plant communities in high mountains. West Georgia lacks xerophilous vegetation zone and mesophilous forest vegetation spreads from the sea level to subalpine zone. The Colchic refugium (West Georgia) ensured survival of the Tertiary's mesophilous forest flora. Vertical profile of the vegetation is more complex in East Georgia with semi-desert, steppe and arid open forest zone. In South Georgia the montane zone represented by montane steppe is devoid of forests.
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Holoparasitic genera within family Orobanchaceae are characterised by greatly reduced vegetative organs; therefore, seed micromorphology has proved to be a useful complementary taxonomic criterion. Seeds of 160 samples from 54 localities of 26 taxa of the Orobanche and Phelipanche genera occurring in central Europe, specifically from Poland, the Czech Republic, Austria, and Slovakia, supplemented by samples from Spain, France and Ukraine, were investigated using scanning electron microscopy. Thirteen quantitative or qualitative morphological characters of seeds were analysed. The following three types of periclinal wall sculpture of seeds were identified: veined and fibrillar in Phelipanche; with oval or elliptic perforations (pitted) in almost all species of Orobanche; with outer periclinal wall smooth, granular or rugged (very rarely visibly pitted), impeding vision of the inner one, occurring only in O. gracilis Sm. and O. coerulescens Stephan in Willd. The influence of different hosts on the features of seeds of eight species is also presented, as well as relationships between seed morphology and taxonomic classification, including problematic taxa. The best diagnostic features include type of ornamentation of the periclinal wall, perforation diameter (in pitted sculpture), fibrillar diameter (in fibrillar sculpture) and width of anticlinal walls. Size and shape of the seeds and cells and the presence of median troughs are variable, however, these features can be helpful when using larger samples. The usefulness of micromorphological studies on seeds of Orobanche and Phelipanche is demonstrated.
Chapter
Plants annual, biennial or perennial, herbaceous or suffrutescent, rarely climbers, shrubs or trees. Stems erect, ascending or prostrate. Leaves opposite or alternate, entire, rarely divided or pinnatifid to pinnate, leaf-bases often fused. Inflorescence a raceme or thyrse,without terminal flower. Flowers zygomorphic, prophylls present or absent. Calyx with sepals free to base or almost fused, actinomorphic or zygomorphic, often tetramerous by reduction of the median sepal. Corolla sympetalous, zygomorphic, rarely actinomorphic, the 2 upper (adaxial) petals often fused, forming an upper lip opposite to the lower (abaxial) lip with 3 lobes, throat of corolla often closed by a palate, abaxial base sometimes calcarate or at least gibbose to saccate. Stamens usually inserted on corolla-tube, sometimes 5, usually 4, the fifth (median) stamen reduced to staminode or absent, sometimes also the anterior (abaxial) stamens reduced to staminodes or lacking, anthers with two well-developed thecae or with 1 fertile theca and a smaller, reduced, sterile theca, the latter sometimes lacking.
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The Panicum capillare complex includes several taxa, among them P. capillare L., which is usually considered to be an established alien throughout Europe, whereas other species are recorded only as casuals. A new representative of the complex, P. riparium H. Scholz was described from Germany in 2002, and shortly after its description was recorded in several countries on the continent. In the course of herbarium revisions and recent fi eld studies the authors documented several localities of the species in Croatia as well. The paper presents a new key for the determination of Croatian species of the complex and anticipates the invasion of P. riparium in the sub-Mediterranean regions of the Balkan Peninsula.
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A taxonomic revision of the family Chenopodiaceae in Nepal with new diagnostic keys, descriptions and detailed distribution patterns is here presented for the first time. 24 species from 13 genera are reported. The apophytes in Himalaya are found in both ruderal and segetal plant communities in contrast to Chenopodiaceae in temperate parts of Eurasia which prefer ruderal places only. The Himalaya and West Tibet are considered to be the regions with the greatest taxonomic diversity of native Dysphania in Eurasia. Chenopodium perttii is described as new for science, and its differences from other related taxa are described, including its unique minute-papillate (almost smooth when dry) pericarp. Chenopodium bengalense seems to be an older name for the taxa known as C. giganteum or C. album subsp. amaranticolor. The lectotypes of Chenopodium giganteum, C. album subsp. yunnanense, C. aristatum f. muticum (here synonymized with Dysphania kitiae) and Microgynoecium tibeticum are designated. Both lectotype and epitype for Atriplex bengalensis (basionym of C. bengalense), C. giganteum and the re-instated species C. pallidum are designated.
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Scutellaria ×ketenoglui M.Cicek & Yaprak, a natural hybrid between S. tortumensis and S. sosnowskyi subsp. sosnowskyi is described as new to science from northeastern Anatolia, Turkey, and illustrated. The morphological characteristics of S. ×ketenoglui are compared with those of its parents.
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Agrostis pendryi, a new species from Nepal and northwest India, is described and illustrated. This species is morphologically similar to A. pilosula and A. munroana, but differs from them by its shorter, weak and straight awn, and palea less than half the length of the lemma.