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First photographic record of the Persian leopard in Kurdistan, northern Iraq

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Under a grant from the Conservation Leadership Programme (CLP), the Nature Iraq team has been researching wild goat Capra aegagrus in the Kurdistan region of northern Iraq. On the IUCN Red List (Weinberg et al. 2008), wild goat is listed as Vul-nerable, and our research will help to enrich international understanding about the status of this animal in Iraq and conserve this species. During the CLP field surveys the team set one camera trap in a habitat where it was thought to be a suitable place for different carnivorous animals. The Persian leopard Panthera pardus was our main target species because our team was already working on their prey species, wild goats, and because we have seen a few incidents of the killing of leopards, which made us want to clarify their status in our study areas. Persian leopard is an Endangered species whose presence in Iraq, according to IUCN Red List (Khorozyan et al. 2008) is uncertain
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ISSN 1027-2992
CAT
news
N° 56 | SPRING 2012
CATnews 56 Spring 2012
02
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Cover Photo: Persian leopard in Bafgh, Iran
Photo DoE Iran/CACP/UNDP/
ICS/Panthera
The designation of the geographical entities in this publication, and the representation of the material, do not imply the expression of any
opinion whatsoever on the part of the IUCN concerning the legal status of any country, territory, or area, or its authorities, or concerning
the delimitation of its frontiers or boundaries.
CATnews 56 Spring 2012
Fig. 1. View of the survey site facing south (Photo Saman A. Ahmad).
HANA A. RAZA1*, SAMAN A. AHMAD2, NABEEL A. HASSAN1, KORSH ARARAT1 MARIWAN
QADIR3 AND LAITH ALI1
First photographic record of
the Persian leopard in
Kurdistan, northern Iraq
Under a grant from the Conservation Leadership Programme (CLP), the Nature Iraq
team has been researching wild goat Capra aegagrus in the Kurdistan region of
northern Iraq. On the IUCN Red List (Weinberg et al. 2008), wild goat is listed as Vul-
nerable, and our research will help to enrich international understanding about the
status of this animal in Iraq and conserve this species. During the CLP field surveys
the team set one camera trap in a habitat where it was thought to be a suitable place
for different carnivorous animals. The Persian leopard Panthera pardus was our
main target species because our team was already working on their prey species,
wild goats, and because we have seen a few incidents of the killing of leopards,
which made us want to clarify their status in our study areas. Persian leopard is an
Endangered species whose presence in Iraq, according to IUCN Red List (Khorozyan
et al. 2008) is uncertain.
According to Hatt (1959), leopards were few
and scattered. He reported only two skins ob-
tained from leopards in the Kurdistan Moun-
tains; one recorded by Pocock (1930) from
Rawanduz, and the other by Field (1955) from
Bradost Mountain. These are very old sour-
ces and no information on the presence of
this animal was documented since this time.
Studies on biodiversity in this area have been
undertaken by Nature Iraq for six years as
part of the Key Biodiversity Areas (KBA) Pro-
ject undertaken in partnership with the Iraqi
Ministry of Environment and with the support
of the Italian Ministry of Environment, Land &
Sea. Nature Iraq has only had confirmation of
the species presence in Iraq through animals
it has been proposed as a protected area by
Nature Iraq. It is located within the Zagros
Mountain Forest Steppe eco-region, which is
critically threatened and part of a biodiversity
hotspot. The specific study area is approxi-
mately 310 km2 within the Qara Dagh range,
which is part of the Zagros foldbelt that ex-
tends from southern Turkey through northern
Iraq and down to southwest Iran (Fig. 2). The
highest peak within the range is reaching
1,800 meters. This area is very important
due to the richness of its flora and fauna. The
habitat is mainly oak forest with vegetation
coverage of about 90%. There are more than
500 species of plants, including the national
or regional endemic species such as Galium
qaradaghense, Cousinia kopi-karadaghensis,
Symphytum kurdicum, Onosma albo-roseum,
Cephalaria syriaca, Astragalus spinosus, Ce-
phalanthera kurdica and Silybum marianum
(Al-Rawi 1988). This area contains about
12% of flora species found in Iraq. Based on
Nature Iraq's rapid assessments this area is
home to more than 8 mammal species and
over 87 bird species. Data for amphibians and
reptiles are scarce.
Methods
There have been no field studies on leopards
or their prey species in this area or Iraq as a
whole. This discovery was made in the course
of Nature Iraq’s wild goat survey, in particular
thanks to a camera trap set in the same loca-
tion over two survey periods: the first session
of camera trapping started from 1 October to
20 October 2011, where two camera traps
were set in two different mountains of Qara
Dagh Area. The first camera (Stealth Cam
STC-I540IR model) was left on Tasha Loos
Mountain for 19 days and the second came-
ra (Keep Guard KG 550 model) was left on
Jazhna Mountain, also for 19 days and both
cameras were retrieved on the same day (20
October 2011). The second session of camera
trapping started from 2 January to 13 Febru-
ary 2012, where only one camera trap (Keep
Guard KG 550 model) was set on only Jazhna
Mountain and the camera was left there for a
period of approximately one month and was
retrieved on 13 February 2012. The cameras
were not checked until they were retrieved at
the end of each survey.
Results & Discussion
During the first session of camera trapping in
6 October 2011, one picture of a male leopard
was taken (Supporting Online Material, SOM
Photo 1) with the Keep Guard KG 550 camera
that were killed. One leopard was killed by
landmine in 2009 near the village of Mortka
east of Darbandikhan Lake. This leopard was
examined by Nature Iraq staff as a mounted
specimen in 2010. A second was killed in
Diyala in 2008 (Nature Iraq & Iraqi Ministry
of Environment 2011). No live sightings have
been made in Iraq until the Nature Iraq ca-
mera trap photographed a male leopard in
October 2011. This project identifies globally,
regionally, and nationally important areas for
biological diversity within Iraq.
Study area
The area where the leopard was found (Fig. 1)
is a key biodiversity area (KBA) and therefore
34
short communication
CATnews 56 Spring 2012
on Jazhna Mountain, while the Stealth Cam
STC-I540IR was unsuccessful to photograph
any wildlife on Tasha Loos Mountain, and on
23 January 2012 during the second session of
camera trapping on Jazhna Mountain, ano-
ther picture of a leopard (possibly the same
individual) was caught (Fig. 3). Besides the
Persian leopard, six different species of mam-
mals were photographed by the camera trap:
wild cat Felis silvestris (SOM Photo 2), red fox
Vulpes vulpes, golden jackal Canis aureus, In-
dian crested porcupine Hystrix indica, Persian
squirrel Sciurus anomalus, and wild boar Sus
scrofa (Raza et al. 2011). In addition to this
finding, the team was successful in observing
and photographing 49 individual wild goats,
during the CLP summer and winter surveys.
Although this finding is only a beginning of
research on Persian leopard status in Iraq, it
is a great finding. As leopards were last re-
ported by locals (interviewed by Nature Iraq
staff at different sites in the Kurdistan region
in 2010 and as part of the Key Biodiversity
Areas (KBA) project) in the 1980’s, this could
spur further research on this species and its
prey species to improve our knowledge on
their conservation status in Iraq. Leopards and
wildlife in general are under threat from over-
exploitation of their main prey species, and
although the leopard is not usually hunted for
its fur or other means of use but rather killed
in self defense, as claimed by a local who re-
cently killed a leopard in one of the mountains
of Kurdistan, Northern Iraq. More studies are
needed to understand the status of leopard in
this area and all over Iraq and regulations are
needed to better control the hunting pressure.
Acknowledgement
Our thanks go to the Conservation Leadership Pro-
gramme fund for helping us to initiate a project on
wild goat conservation in Iraq. We would like to
thank the members of the Nature Iraq team who
have been dedicated for making this project a gre-
at success. We would also like to thank Amir Hus-
sein Khaleghi of the Plan4Land Society in Iran for
providing advice. Special thanks go to the forestry
police, who have been very helpful in our survey
trips to the area, for their efforts to control the hun-
ting pressure in this area. We also acknowledge
Anna Bachmann and James Wudel for editing.
References
Al-Rawi A. 1988. Wild Plants of Iraq with their dis-
tribution. 3rd edition. Ministry of Agriculture &
Irrigation, State board for Agricultural & Water
Resources Research National Herbarium of
Iraq. Baghdad, Iraq.
Field H. 1955. Animals of Jebel Bradost, Iraq.
Amer. Doc. Inst., Doc. No. 4427, 60-62.
Hatt R. T. 1959. The mammals of Iraq. Museum of
Zoology, Univ. of Michigan No. 106.
Khorozyan I. 2008. Panthera pardus ssp. saxicolor.
In: IUCN 2011. IUCN Red List of Threatened
Species. Version 2011.2. <www.iucnredlist.
org>. Downloaded on 15 February 2012.
Nature Iraq & Iraqi Ministry of Environment. 2011.
Key Biodiversity Areas of Iraq 2010 Site Re-
view. Sulaimani: Nature Iraq. Publication No.
NI-0311-01
Pocock R. I. 1930. The Panthers and Ounces of
Asia. Ibid., pp. 64-82, 307-36.
Raza H.A., Qadir M., Ali L., Ararat K., & Abdulha-
san N. 2011. The Search for the Vulnerable
wild goat (Capra aegagrus) Continues with
Great Success. Conservation Leadership Pro-
gramme (CLP), BirdLife International, Conser-
vation International (CI), and Fauna and Flora
International (FFI): Projects in the middle east.
Retrieved from http://maildogmanager.com/
page.html?p=000001XDDtjL2KurMdfWs%2Fz
6ktHaRjkw%3D%3D on 16 February 2012.
Weinberg P., Jdeidi T., Masseti M., Nader I., de
Smet K. & Cuzin F. 2008. Capra aegagrus. In:
IUCN 2011. IUCN Red List of Threatened Spe-
cies. Version 2011.2. <www.iucnredlist.org>.
Downloaded on 15 February 2012.
Supporting Online Material at www.catsg.org/
catnews.
1 Nature Iraq
*<hana.ahmad@natureiraq.org >
2 Nature Iraq & University of Sulaimani
3 Nature Iraq & Sulaimani Environm. Directorate
Fig. 2. Google Earth Map showing Qara Dagh Area’s location within Iraq and as part
of the Zagros belt.
Fig. 3. Persian leopard caught on camera trap on January 2012 (Photo Nature Iraq/CLP).
35
leopard in Iraq
Raza et al. 2012. First photographic record of the Persian leopard in Kurdistan, northern Iraq. Cat
News 56, 34-35.
Supporting Online Material Additional photos
SOM Photo 1: Male Persian Leopard Panthera pardus caught on camera trap on October 2011
(Photo Nature
SOM Photo 2. Wild Cat Felis silvestris captured by camera trap in the same area on January 2012
(Photo Nature Iraq/CLP)
... Since then, no further records have been made until 2008 when a Persian leopard was killed near Mandli in Diyala (Al-Sheikhly, 2012) and another was killed by a landmine from Iraq-Iran war ruminants near the village of Mortka, to the east of Darbandikhan Lake in 2009 (Nature Iraq, 2017). Regardless, the status of Persian leopard in the mountains of northern Iraq was enigmatic and no sightings of live leopards have been made until a male leopard was captured by a camera trap in Qara Dahg area in Sulymaniyah Province in October 2011 (Raza et al., 2012). In 2001-2014, 10 confirmed records of Persian leopard from northern Iraq and southeastern Turkey were investigated and hypothesized as a long dispersal of males from the Iranian populations (Avgan et al., 2016). ...
... 2).The distribution of Persian leopard in Bamo Mountain could be attributed to an expansion of the home range of some individuals that previously reported from Qara Dagh Mountain (e.g. Raza et al., 2012) or from the western Iranian populations (e.g. Moqanaki et al., 2013;Parchizadeh and Adibi, 2019;Farhadinia et al., 2019). ...
Article
Full-text available
Mammals are under threat worldwide due to deforestation, hunting, and other human activities. In Iraq, a total of 93 species of wild mammals have been recorded including species with global conservation concern. Bamo Mountain is situated within the Zagros Mountains in northern Iraq which is a suitable habitat for wild mammals. Due to scarcity of the field survey efforts and cryptic behavior, monitoring of the wild mammals fauna in Zagros Mountain seems challenging. Therefore, we used a camera trap which seems to be an ideal way to determine species diversity of wild mammals in Bamo Mountain. Moreover, interviews with local villagers were performed. The mammalian diversity of Bamo Mountain is not fully explored but seemed threatened by local extinction due to poaching and wildlife trafficking, minefields, and annual fires. In this study, a total of eight species of wild mammals were recorded for the first time in Bamo Mountain using camera trap method including the Persian leopard Panthera pardus saxicolor Pocock, 1927, and the wild goat Capra aegagrus Erxleben, 1777, flagship and key species of conservation concern. As far as it is concerned, the major threats on the wild mammals were discussed and some important points were highlighted towards the establishment of the protected area in Bamo Mountain.
... Felis s. iraki was recorded from: Sheikh Saad (Cheesman 1920); Baghdad and Diwaniya . Felis s. nesterovi was recorded from Nachr-Chasasch (Harrison 1968; and Peramagroon (Raza et al. 2012 ; between Al-Qurna and Azir, Shatt al Adhaim, and Ad Dawr (Pitman 1922); Hilla Desert ; Abu Ghreb, near Akkarkuf (Bodenheimer 1959); Hila and Dujail Caracal Caracal caracal (Schreber, 1776) Subspecies: C. c. schmitzi (Matschie, 1912). Distribution: Confined to the desert and arid steppes of the extreme western and southern Iraq. ...
... Distribution: Abundant along the Tigris and Euphrates valleys, deserts and arid steppes, and mountains of northern Iraq. Recorded from many localities by , Harrison (1959), Smielowski (1978, Nadachowski et al. (1990) and Kadhim (1997); Peramagroon (Raza et al 2012); Hawraman Mountain (Lahony et al. 2013). RR: Jabal Makhool, Al-Alam, Al-Cidir, Al-Tharthar Valley (Sa-13); Rutba, Ga'ara, Wadi Horan, Khan Al-Baghdadi, Al-Jezera (An-13); Jabal Himreen and near Hor Shwija (Di-13), Hor Dalmaj (Qa-13); Central Marshes (Th-14). ...
Article
Full-text available
A checklist of the mammals of Iraq including their current status and geographical distribution is provided. Data were collected from literature and combined with field observations obtained during 2013–2014. The checklist comprises 93 species of wild mammals occurring in Iraq and belonging to eight orders, 28 families, and 65 genera. The status and distribution of three Erinaceomorphs, three Soricomorphs, 20 Bats, 23 Carnivores, eight Artiodactyls, two Lago-morphs, 28 Rodents, and six Cetaceans are reported. The checklist includes 24 species evaluated by the International Union for the Conservation of Nature and Natural Resources: one was listed as Extinct, one as Critically Endangered, four as Endangered, eight as Vulnerable, seven as Near Threatened, and three as Data Deficient. One subspecies of car-nivores (Smooth-coated Otter Lutrogale perspicillata maxwelli) and one rodent species (Long-tailed Nesokia Nesokia bunnii) endemic to Mesopotamian marshlands were highlighted as taxa with conservation priority to Iraq. We also fo-cused on main threats to the most common wild mammals with special attention to impacts of illegal hunting and poaching .
... However, habitat is suitable and wild goat as the main leopard prey species is available in the region. In a neighbouring habitat in Iraq close to the Kusalan and Shahu PA, a Persian leopard was photo captured via camera trapping (Raza et al., 2012). Furthermore, frequent signs of the leopards (voices in mating season in 2010), livestock losses and leopard scratches on the trees are frequently detected. ...
Chapter
This chapter is dedicated to assessing the Persian leopard potential distribution in Iran on a regional basis that aims to address four objectives and a null hypothesis. Objectives are concerning (1) estimation of the leopard potential distribution, (2) possibility of a major fragmentation in the Persian leopard range in Iran as first mentioned by Sanei et al. (2016), (3) prediction of landscape corridors which can improve the distribution pattern connectivity and (4) the main environmental variables that contribute to assessing the predictive maps. The null hypothesis addresses the variability of permutation importance of the environmental factors in accordance with the regional variability of environmental characteristics. Due to the variability of the environmental characteristics across the country and the leopard putative range which includes almost 30 provinces out of 31, the area has been innovatively divided into five significantly dissimilar regions as discussed in the previous chapter. Subsequently, MaxEnt modelling is conducted in a regional context using a total of 17 variables including 12 natural and 5 human factors together with more than 550 well distributed leopard occurrence data in all regions. Environmental variables have been tested for possible correlation prior to the modelling procedures. Area under the curve (AUC) was used to test the model fit to the data set. Jackknife test was performed to assess the contribution of environmental variables to the MaxEnt models. Fifteen replications with test percentage of 20% were used for validation. Additional evaluation of the predictive models was conducted by assessing the potential habitat distribution maps via the expert/local knowledge of 150 individuals from all five regions. Findings support that the Persian leopard range in Iran is in the process of a major fragmentation to the northern and the southern parts. Accordingly, two landscape corridors providing vital linkages to connect leopard potential habitats in a metapopulation scale are identified. Developed predictive maps in this chapter are a basis for the researches presented in Chaps. 5, 6 and 7. Authors believe that MaxEnt modeling on a regional basis has considerably improved the accuracy of the predictive maps that eventually formed the countrywide potential distribution of the Persian leopard potential habitats in Iran.
... However, habitat is suitable and wild goat as the main leopard prey species is available in the region. In a neighbouring habitat in Iraq close to the Kusalan and Shahu PA, a Persian leopard was photo captured via camera trapping (Raza et al., 2012). Furthermore, frequent signs of the leopards (voices in mating season in 2010), livestock losses and leopard scratches on the trees are frequently detected. ...
Chapter
Even though the Persian leopard Panthera pardus saxicolor is an endangered subspecies with the main population inhabiting in Iran (Khorozyan and Abramov, Zool Middle East 41:11–24, 2007; Kiabi et al., Zool Middle East 26(1):41–47, 2002), earlier studies (Sanei et al., Assessment of the Persian leopard mortality rate in Iran. In: Proceedings from UMT 11th International Annual Symposium on Sustainability Science and Management (pp. 1458–1462, 2012). Terengganu, Malaysia: Universiti Malaysia Terengganu) demonstrated that the majority of leopard mortalities are recorded to be as a result of intentional hunting, revenge killing, and poisoning of the specimens. To mitigate livestock–carnivore conflicts and reduce the subsequent revenge killings, an innovative model including a medium and a long-term insurance schemes together with awareness raising, trust building, and participatory conservation strategies is designed. Accordingly, the medium term insurance scheme addresses three main subjects of (1) improving conservation practices in the areas of leopard mortality hot spots, (2) medical payments and wergild for possible human injuries/maim/death because of human–leopard conflicts and (3) recompensing livestock depredation. Also, since the wolf Canis lupus distribution is comparable with the leopard range in the country, because of conservation concerns, damages caused by wolf depredation are also planned to be recompensed partially in the first type (i.e. medium term) insurance scheme and fully recompensed in the long term (i.e. second type) insurance program. Introducing sessions about the relative regulations and instructions were conducted for provincial wildlife wardens and DoE staff who are well familiar with wildlife sign surveys and have a quick access to the habitats in each region. Subsequently, they took the responsibility for identification of wildlife species in livestock–carnivore conflicts. Improvements in husbandry practices, linking the payments to the acceptable husbandry enhancements and participation in reducing risk of damages by local people are some of the instructions considered in the model to improve the efficacy and outcomes. So far, Department of Environment of Iran together with a private insurance company has partially launched the short term insurance scheme since 2016 and launching other sections of this model is in progress.
... However, habitat is suitable and wild goat as the main leopard prey species is available in the region. In a neighbouring habitat in Iraq close to the Kusalan and Shahu PA, a Persian leopard was photo captured via camera trapping (Raza et al., 2012). Furthermore, frequent signs of the leopards (voices in mating season in 2010), livestock losses and leopard scratches on the trees are frequently detected. ...
Chapter
Similar to the other big cats of Iran (i.e., the Asiatic lion Panthera leo persica and the Caspian tiger Panthera tigris virgata that are extinct in the region), the Persian leopard Panthera pardus saxicolor has a unique importance in the Iranian art, history, and literature. Symbolization of these species over centuries to emphasize on strength, intelligence, and bravery of the kings and national heroes indicates the significance of the big cats including the leopard, for the Iranian society. The Persian leopard has been also the center of attention for several local and nationwide researches in Iran since 2002. Seeing this introductory chapter of the book mainly consisted of an overview to the prior knowledge about the subspecies, the cultural importance, results of an assessment to determine the leopard conservation requirements in various regions of Iran as well as problem trees addressing habitat-related issues (e.g., habitat destruction and degradation) and high leopard mortality rate.
Article
Full-text available
As the largest extant cat species in west Asia, the leopard (Panthera pardus) shows high morphological variation, which has led to the description of seven different subspecies in the region. Different investigations have tried to clarify its phylogenetic structure; however, sample size and spatial distribution insufficiently represent the Iranian population, the largest remaining bulk of the Persian leopard (P. p. saxicolor) in the Middle East that probably functioned as a source for the subspecies' range. We examined sequence variation in the mitochondrial NADH-5 gene for 25 leopards from different parts of Iran. Also, we examined 49 adult male skulls to understand the morphological variation of the Iranian leopard population. Our craniometrical results revealed that while no differentiation is seen based on size or shape characteristics from different parts of Iran, larger individuals normally belong to the northern range. Time-calibrated Bayesian phylogenetic analysis suggested that the Iranian female lineage is a monophyletic group that diverged from a group of Asian leopards in the second half of the Pleistocene. Three closely related haplotypes were identified for the entire country: one commonly found haplotype throughout Iran, south Caucasus and Turkmenistan and two localized haplotypes were sequenced from southern Zagros and eastern Alborz. Accordingly, the Persian leopard population in Iran as well as in neighbouring countries can be protected as a large management unit through large-scale conservation planning. Moreover, the available captive stock of the Persian leopard represents an invaluable source for reintroduction for countries interested in restoring their locally extinct population. © 2015 The Linnean Society of London, Biological Journal of the Linnean Society, 2015, ●●, ●●–●●.
Article
Full-text available
Designation of subspecies in wild cats (Carnivora: Felidae) often results from arbitrary description of morphological characters and it can be confounded by geographic variation and sexual dimorphism. To test the contribution of these factors to subspecies grouping, I studied 22 cranial shape variables in 55 skulls of the common leopard (Panthera pardus) from the Middle East and estimated their differences between the three subspecies groups in this region, namely the Persian leopard (P.p. ciscaucasica Satunin, 1914), Anatolian leopard (P.p. tulliana Valenciennes, 1856) and Arabian leopard (P.p. nimr Hemprich et Ehrenberg, 1833), among sexes. Male and female skulls increased in size with latitude (Bergmann’s rule) and female skull size also significantly increased with longitude. Results suggest that taxonomic grouping was determined only by female skulls which showed high variation between the groups, unlike male skulls which varied more within the groups. Inter-group variation of female skulls, presence of only four cranial variables which are significantly different between the groups in all skulls, and the lack of historical barriers for subspeciation may suggest the existence of one leopard subspecies in the Middle East. Independent genetic and ecological studies are required to settle this taxonomic issue. This study urges for securing connectivity between the two remnant, taxonomically identical populations in the region, namely in (i) Iran and neighbouring countries and (ii) four Arab countries and Israel, with more research and conservation efforts in these populations and in the Fertile Crescent lying between them.
Article
Full-text available
Under a grant from the Conservation Leadership Programme (CLP), the Nature Iraq team has been researching wild goat Capra aegagrus in the Kurdistan region of northern Iraq. On the IUCN Red List (Weinberg et al. 2008), wild goat is listed as Vulnerable, and our research will help to enrich international understanding about the status of this animal in Iraq and conserve this species. During the CLP field surveys the team set one camera trap in a habitat where it was thought to be a suitable place for different carnivorous animals. The Persian leopard Panthera pardus was our main target species because our team was already working on their prey species, wild goats, and because we have seen a few incidents of the killing of leopards, which made us want to clarify their status in our study areas. Persian leopard is an Endangered species whose presence in Iraq, according to IUCN Red List (Khorozyan et al. 2008) is uncertain.
Article
http://deepblue.lib.umich.edu/bitstream/2027.42/56350/1/MP106.pdf
Animals of Jebel Bradost
  • H Field
Field H. 1955. Animals of Jebel Bradost, Iraq. Amer. Doc. Inst., Doc. No. 4427, 60-62.
Panthera pardus ssp. saxicolor
  • I Khorozyan
Khorozyan I. 2008. Panthera pardus ssp. saxicolor. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist. org>. Downloaded on 15 February 2012.
Wild Plants of Iraq with their distribution
  • A Al-Rawi
Al-Rawi A. 1988. Wild Plants of Iraq with their distribution. 3rd edition. Ministry of Agriculture & Irrigation, State board for Agricultural & Water Resources Research National Herbarium of Iraq. Baghdad, Iraq.
The Search for the Vulnerable wild goat (Capra aegagrus) Continues with Great Success BirdLife International, Conservation International (CI), and Fauna and Flora International (FFI): Projects in the middle east
  • H A Raza
  • M Qadir
  • L Ali
  • K Ararat
  • N Abdulhasan
Raza H.A., Qadir M., Ali L., Ararat K., & Abdulhasan N. 2011. The Search for the Vulnerable wild goat (Capra aegagrus) Continues with Great Success. Conservation Leadership Programme (CLP), BirdLife International, Conservation International (CI), and Fauna and Flora International (FFI): Projects in the middle east. Retrieved from http://maildogmanager.com/ page.html?p=000001XDDtjL2KurMdfWs%2Fz 6ktHaRjkw%3D%3D on 16 February 2012.
Key Biodiversity Areas of Iraq
Nature Iraq & Iraqi Ministry of Environment. 2011. Key Biodiversity Areas of Iraq 2010 Site Review. Sulaimani: Nature Iraq. Publication No. NI-0311-01
Key Biodiversity Areas of Iraq 2010 Site Review
Nature Iraq & Iraqi Ministry of Environment. 2011. Key Biodiversity Areas of Iraq 2010 Site Review. Sulaimani: Nature Iraq. Publication No. NI-0311-01
The Search for the Vulnerable wild goat (Capra aegagrus) Continues with Great Success. Conservation Leadership Programme (CLP), BirdLife International, Conservation International (CI), and Fauna and Flora International (FFI): Projects in the middle east
  • H A Raza
  • M Qadir
  • L Ali
  • K Ararat
  • N Abdulhasan
Raza H.A., Qadir M., Ali L., Ararat K., & Abdulhasan N. 2011. The Search for the Vulnerable wild goat (Capra aegagrus) Continues with Great Success. Conservation Leadership Programme (CLP), BirdLife International, Conservation International (CI), and Fauna and Flora International (FFI): Projects in the middle east. Retrieved from http://maildogmanager.com/ page.html?p=000001XDDtjL2KurMdfWs%2Fz 6ktHaRjkw%3D%3D on 16 February 2012.