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Coléoptères Staphylinidae de la région paléarctique occidentale. V. Sous famille Paederinae Tribu Paederini 2, Sous famille Euaesthetinae

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... This species was only known from Goran cave, Cap Cantin (Ras Beddouza) in Morocco (Peyerimhoff de Fontenelle 1949;Coiffait 1982;Hernando 2007). Aedeagus illustrations of this species provided by Coiffait (1982, fig. ...
... This species was only known from Goran cave, Cap Cantin (Ras Beddouza) in Morocco (Peyerimhoff de Fontenelle 1949;Coiffait 1982;Hernando 2007). Aedeagus illustrations of this species provided by Coiffait (1982, fig. 99D-F). ...
... Domene gaudini is confined to the Atlantic Pyrenees in France and Spain (Jeannel 1938;Coiffait 1982;Struyve 2018). ...
... Distribution: The ssp. gridellii Jarrige, 1949 is present in France, Georgia, Italy and Spain (Schülke & Smetana, 2015). Coiffait, 1982 Pseudobium gridellii ibericum Coiffait, 1982: 262 Subspecies described by Coiffait (1982) from Algarve, Alportel, no date, 1 ♂, Type, (cCoi) and 4 ex., Paratypes near Tavira, no date (cCoi) and 1 ex., near Silves, no date (cCoi). Drugman & Outerelo (1997) repeated these localities. ...
... Distribution: The ssp. gridellii Jarrige, 1949 is present in France, Georgia, Italy and Spain (Schülke & Smetana, 2015). Coiffait, 1982 Pseudobium gridellii ibericum Coiffait, 1982: 262 Subspecies described by Coiffait (1982) from Algarve, Alportel, no date, 1 ♂, Type, (cCoi) and 4 ex., Paratypes near Tavira, no date (cCoi) and 1 ex., near Silves, no date (cCoi). Drugman & Outerelo (1997) repeated these localities. ...
... Throbalium lesnei (Koch, 1937) Throbalium lesnei (Koch, 1937a): 27 Coiffait (1982) reported this species from southeastern Portugal. Boieiro et al. (1999) did it from Castro Marim (Algarve). ...
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The revision of the subfamily Paederinae (Coleoptera: Staphylinidae) of the Portuguese fauna is concluded with this contribution, in which data of 69 species within 22 genera belonging to eight subtribes are provided. In this article, one genus and 10 species are newly recorded from Portugal, while three of them are in need of confirmation. Resumo: Sobre alguns géneros de Paederinae da fauna portuguesa (Coleoptera: Staphylinidae: Paederinae). A revisão da subfamília Paederinae (Coleoptera: Staphylinidae) da fauna portuguesa fica terminada com esta contribuição, na qual se procuram dados de 69 espécies distribuídas em 22 géneros pertencentes a oito subtribos. Neste artigo, um género e 10 espécies são assinalados como novidade para Portugal, enquanto três delas aguardam confirmação.
... Identificarea: Specia notată cu simbolul (*) prezintă statutul de specie nouă pentru fauna țării noastre, semnalată în premieră. Definirea trăsăturilor morfologice ale speciilor s-a bazat pe cheile sinoptice (Assing, & Schulke, 1999;Coiffait, 1974 ...
... Este specie micetobiontă, coprobiontă, prădătoare şi coprofagă. Geografic, este distribuit ca element holarctic (Coiffait, 1974). În colecția muzeului de Entomologie se păstrează în cutia 27. ...
... Dezvoltă o generație în perioada de sezon. Este specie micetobiontă, coprobiontă şi prădătoare (Coiffait, 1974 eri. Manifestă o extindere sporadică şi populează diverse tipuri de substraturi, precum dejecțiile animaliere, salcâmul în faza de înflorire, grămezile de plante uscate şi /sau în proces de descompunere, litiera pădurilor cu amestec de plante tip gorun, conifere, glădiță cu frasin etc. Prezența sa sub plantele de romaniță poate fi explicată prin atracția față de umiditatea solului şi densitatea plantelor. ...
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This study provides the first-ever data on staphylinids inhabiting the wild chamomile (Matricaria chamomilla) biotope. The research was conducted in April and May 2022, within a 3-are field with wild chamomile plants in the flowering stage. The findings, outlined in this paper, encompass 26 species from 4 subfamilies and 19 genera. Notably, a new species for the fauna of the Republic of Moldova was reported: Tachyporus dispar (Paykull, 1789), belonging to the Tachyporinae subfamily. The research presented in this paper completes the chapter on staphylinid faunal diversity within the country’s territory and highlights the potential for exploring new biotopes with other medicinal plants, thereby enhancing the attraction of this group of Coleoptera.
... Systematics of Paederus species is rather problematic and is relied on the male main and secondary sexual morphological characters (20). This has ended a very complex history for Paederus taxonomy and has reformed it intensely (21) and some species are regarded as synonyms of each other and or lowered to a single subspecies/species (22 and references herein). ...
... The pictorial keys of Coiffait (1982) were used to determine the specimens to genus level (20) and then we send them to the specialist (Dr Sinan Anlaş laboratory) in Turkey for species identification. The specimens were identified on the base of habitus and sound structure of male basal and secondary sexual characters ( Fig. 2). ...
... The pictorial keys of Coiffait (1982) were used to determine the specimens to genus level (20) and then we send them to the specialist (Dr Sinan Anlaş laboratory) in Turkey for species identification. The specimens were identified on the base of habitus and sound structure of male basal and secondary sexual characters ( Fig. 2). ...
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Background: A combined morphological and molecular survey was performed to determine the agent of human linear dermatitis Paederus Fabricius, 1775 (Coleoptera: Staphylinidae, Paederinae) species composition in Mazandaran Prov­ince in the Caspian Sea coast in northern Iran, where most of linear dermatitis cases of the country occurred. Methods: Altogether, 397 Paederus specimens were collected from May to August 2021 and classified using morphological characters and ITS2-rDNA sequence analysis. Results: Morphological investigation revealed that all the specimens were Paederus fuscipes. ITS2 polymerase chain reaction (PCR) direct-sequences and the profiles of restriction fragment length polymorphism (RFLP) derived from digestion of PCR products by HinfI, HpaII, and SalI enzymes were identical confirming the morphological results, implying that all specimens belonged to a single taxon. Conclusion: Paederus fuscipes (Fabricius, 1775) is considered the dominant taxon and responsible for linear dermatitis in Ma­zandaran Province. To our knowledge, we have provided the first molecular typing of Paederus beetles at the species level, suggesting that ITS2-rDNA characterization is an alternative tool for species discrimination of Paederus spp.
... Eurytop obviously prefers open landscapes [Schillhammer, 2012]. Ambigusoulsy recorded from "Caucasus" [Fauvel, 1874;Coiffait, 1967aCoiffait, , 1974, or ST [Schülke, Smetana, 2015], no exact records from PSER. ...
... Distributed from Northern Europe eastward to Central Asia [Schülke, Smetana, 2015]; mainly in decaying substances, especially dung and compost [Shillhammer, 2012]. In PSER not recorded but potentially expected because of unclear records from "Caucasus" [Horion, 1965;Coiffait, 1974]. ...
... Müller, 1925)] Notes. Known only from Turkey and the Caucasus [Coiffait, 1974;Schülke, Smetana, 2015]. Inhabits decaying organics and dung [Altunsoy et al., 2017]; in Turkey collected mostly from mountains [Özdemir, Sert, 2009;Altunsoy et al., 2017]. ...
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Based on the exhaustive literature survey we provide the annotated catalogue with 874 species of Staphylinidae beetles relevant for the fauna of the open plains of the South European Russia (PSER). PSER is a convenience study region with some ecological integrity as it coincides with the Russian part of the Eurasian belt of steppe and semi-desert biomes. Longitudinally it stretches from the border with Donbass in the west to southern Fore Urals in the east. Latitudinally it is extended from the southern border of the forest-steppe zone in the north to the Sea of Azov, Black Sea, northern foothills of the Caucasus, northern shores of the Caspian Sea and the border with Kazakhstan in the south. Along with the zonal grasslands, PSER is a network of intra-zonal riverine woodlands, wetlands, as well as anthropogenic landscapes. It covers, fully or partly, several administrative regions of the Russian Federation. Our review shows the more complete knowledge of the rove beetle fauna for the western areas of PSER and for the Volga River basin, while several large regions like Republic of Kalmykia, Stavropol or Orenburg regions, on the contrary, are extremely poorly known. Taxonomic groups like Paederinae, Oxytelinae or Staphylininae are better explored, while many taxa including entire big subfamilies like Aleocharinae are hardly known in PSER. To facilitate further investigation of the PSER fauna, we provide keys to all rove beetle genera either recorded or expected in this territory.
... Nomenclature of all species follows . All specimens were identified by the author using the keys by Coiffait (1970Coiffait ( , 1984 and are deposited in the author"s collection (RNFC), unless otherwise stated. New data includes: province, district, town, village or place. ...
... Head slightly longer than wide, pronotum oblong narrower than head and elytra slightly longer than wide but wider than the head. For a more complete description see Coiffait (1984). Aedeagus as in Fig. 1. ...
... Head without mouthparts wider than long, and as width as elytra. For a more complete description see Coiffait (1984). Aedeagus as in Fig. 2. The species is found all year round with two peaks, one from March to April and the other from June to August. ...
... Additional phylogenetically isolated lineages, important to the higher systematics of Staphylininae and study of its evolution, may still be hidden within the morphologically heterogeneous genus Quedius, further emphasizing the need for comprehensive taxon sampling. Much morphological diversity previous efforts for North America (Smetana, 1971) and the West Palaearctic (Coiffait, 1978) such that this diversity is easier to navigate for phylogenetic sampling using informal species groups and other morphological groupings elucidated by keys. However, only a fraction of the hundreds of taxa needed for adequate representation of Quediini are optimally preserved for the high quality of DNA required for the PCR-based workflows that have become standard in the aforementioned phylogenetic studies. ...
... Clade F was resolved as sister to a clade containing all other members of the Microsaurus lineage and named here the 'core Microsaurus' clade ( Fig. 3). Clade G was recovered by all analyses and was composed of the erythrogaster group (clade G1) (sensu Brunke et al., 2020a), the Nearctic caseyi, desertus and limbifer groups of Smetana (1971) (clade G2), and clade G3 containing the West Palaearctic microps group (Coiffait, 1978) and the East Palaearctic kiangiensis group (Smetana, 2017). Clade H was recovered by all analyses and contained representatives of the criddlei, mnemon, przewalskii and spelaeus groups (Smetana, 1971(Smetana, , 2017Solodovnikov & Hansen, 2016;Salnitska & Solodovnikov, 2018b) and Quedius (Megaquedius). ...
... The Raphirus lineage was resolved as a group consisting of large-eyed Quedius species forming sister clades V and W, and genus Euryporus sister to the smaller-eyed and mostly West Palaearctic Quedius of clade X (Fig. 6). Clade V was recovered by all analyses and consisted of the probus and vulpinus groups of Q. (Raphirus) (Smetana, 1971) and species treated as Microquedius (= Raphirus) by Coiffait (1978). Clade W was recovered by all analyses and consisted of species from the boops and muscicola groups of Q. (Raphirus) (Smetana, 1971(Smetana, , 2017. ...
Article
Rove beetles of the tribe Quediini are abundant predators in humid micro-habitats of forested, open, synanthropic or subterranean ecosystems, with just over 800 species distributed across the temperate and subtropical regions of the Northern Hemisphere. Previous molecular phylogenies included only a limited representation of this diversity but have already indicated that Quedius, containing the majority of Quedi-ini species, is polyphyletic. Six genera, historically associated with Quediini but now Staphylininae incertae sedis, are known only from few pinned specimens and have never been sequenced. Recent synergy between target enrichment phylogenomics, low-input sequencing of dry, pinned insect specimens and advances in alpha taxonomic knowledge have made comprehensive sampling of Quediini tractable. Here we developed a novel probe set specialized for anchored hybrid enrichment of 1229 single-copy orthol-ogous loci in Staphylinidae. In one of the largest target enrichment phylogenies of insects to-date, we sequenced 201 ingroup taxa to clearly delimit monophyletic Quediini within Staphylininae and resolve relationships within this tribe, with 46% of sampled taxa derived from pinned specimens (0-45 years old). Maximum likelihood and coalescent phylogenetic analyses produced well-resolved, congruent topologies that will serve as a framework for further exploration of this radiation and its necessary generic revision. The inclusion of nearly all remaining Staphylininae incertae sedis genera, all known only from pinned specimens, resulted in the creation of Quelaestrygonini Brunke, trib. n. and revised concepts for Cyrtoquediini and Indoquediini. Quediini was resolved as mono-phyletic with the transfer of Q. elevatus and Q. nigropolitus to other tribes but Quedius and its subgenera Microsaurus, Distichalius and Raphirus were shown to be para-or polyphyletic. Based on the results of our analyses, Velleiopsis Fairmaire, 1882 syn. n. and Megaquedius Casey, 1915 syn. n. are synonymized with Microsaurus Dejean.
... From L. longulum by the different coloration of the body (in L. longulum: body uniformly black or blackish brown), less oblong head, shorter elytra, and by the different shape of the male sternite VIII (in L. longulum: posterior margin of sternite VIII distinctly convex). For illustrations of L. longulum, see Coiffait (1982: Figures 83e and 83f). ...
... From the transPalaearctic L. brunnipes by the different coloration of the body (in L. brunnipes: head, pronotum and abdomen black or blackish brown), smaller body (in L. brunnipes: 8-9 mm), less oblong head, shorter elytra, relatively longer antennae and by the different shapes of the male sternite VII and VIII. For illustrations of L. brunnipes, see Coiffait (1982: Figures 81g and 81h). ...
... fulvipenne: body black, with elytra dark reddish brown or reddish yellow),less oblong head, less distinct punctation of the body, shorter elytra, and by the completely different shape and chaetotaxy of the male sternite VII and VIII (in L. fulvipenne: sternite VII with weakly concave posterior margin and without modified black setae; sternite VIII with medioposterior marginrelatively large concave). For illustrations of L. fulvipenne, see Coiffait (1982: Figures 81a and 81b). ...
... C'est le cas d'Astenus bimaculatus bimaculatus (Erichson, 1840), seule sousespèce de l'espèce typique connue de France qui, bien que possédant une très vaste distribution paléarctique depuis l'Europe occidentale, la péninsule ibérique, l'Afrique du Nord, Madère et les Canaries, le Caucase jusqu'à l'Asie (SMETANA, 2015), reste encore peu signalée sur notre territoire. Les autres sous-espèces sont auliensis Coiffait, 1984, du COIFFAIT (1960COIFFAIT ( , 1984. ...
... L'espèce se reconnaît de prime abord à sa forme fine et élancée, à ses élytres portant une grosse tache foncée touchant le bord latéral (Fig. 1) et surtout à son édéage à lame ventrale longuement rétrécie au sommet en une pointe aiguë retroussée du côté dorsal, fortement dentée en dessous, assez loin avant l'apex (COIFFAIT, 1984). ...
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The rove beetle Astenus bimaculatus bimaculatus (Erichson, 1840) was again observed in Gironde, in the Médoc, in 2021, 61 years after his last mention.
... Gli Staphylinidae si rinvengono in molti tipi di ambienti, tra cui ad esempio la lettiera, i suoli forestali, sotto le pietre, in prossimità delle aree umide o lungo i litorali, ma anche nello sterco, nelle carcasse di animali morti e all'interno dei nidi e delle tane di uccelli e mammiferi. Bibliografia per determinazione: Assing 2003aAssing , 2003bAssing , 2003cAssing , 2007aAssing , 2007bAssing , 2009Assing , 2010aAssing , 2010bAssing & Schülke 2012;Assing & Wunderle 1997Benick 1954;Bordoni 1982;Coiffait 1972Coiffait , 1974Coiffait , 1978Coiffait , 1982Coiffait , 1984Freude et al. 1974;Jászay & Hlaváč 2006;Makranczy 2014;Pace 1975Pace , 1989Pace , 1996Porta 1926;Tronquet 2007;Zanetti 1987. characters: Body usually rather elongate or less frequently ovate, brown to black, sometimes yellowish, with red-orange markings; last palpomere rarely much larger than previous ones; antennae usually 11-segmented, usually filiform or moniliform, sometimes dentate, not clubbed, more rarely weakly clubbed; sometimes antennomere 1 elongate as a scape; elytra usually very short, exposing 2 to 7 abdominal segments; abdomen free, mobile and dorsoventrally very flexible; tarsi usually pentamerous, but many other tarsal formula can occur; last palpomere never strongly enlarged nor much broader than previous one; 2 tarsal claws simple and subequal. ...
... Staphylinidae occur in a great variety of habitats, such as litter, forest soil, under stones, near wetlands margins or along seashores, but also in dung, in carrion or inside nests and burrows of birds and mammals. identification references: Assing 2003aAssing , 2003bAssing , 2003cAssing , 2007aAssing , 2007bAssing , 2009Assing , 2010aAssing , 2010bAssing & Schülke 2012;Assing & Wunderle 1997Benick 1954;Bordoni 1982;Coiffait 1972Coiffait , 1974Coiffait , 1978Coiffait , 1982Coiffait , 1984Freude et al. 1974;Jászay & Hlaváč 2006;Makranczy 2014;Pace 1975Pace , 1989Pace , 1996Porta 1926;Tronquet 2007;Zanetti 1987. 34. ...
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L’esigenza di studiare la fauna dei Coleotteri delle 10 Riserve Naturali dello Stato gestite dal Reparto Carabinieri Biodiversità di Belluno, al fine di approfondirne le conoscenze, ha messo in evidenza la necessità di disporre di uno strumento “agile” per la determinazione delle numerose famiglie di Coleotteri della nostra fauna. Da questa necessità è nata l’iniziale idea di sviluppare una chiave dicotomica ricca di immagini e schemi di dettaglio, che in corso d’opera è stata poi arricchita anche dalle tavole anatomiche e dal glossario. A completamento del progetto, è sembrato utile allegare un catalogo di tutte le 13047 specie di Coleotteri ad oggi segnalate per il nostro Paese, oltre ad una sintetica descrizione delle famiglie esaminate, corredata da note bio-ecologiche e dall’indicazione di alcuni strumenti bibliografici utili per la determinazione a livello specifico degli esemplari in studio. Quest’opera rappresenta principalmente uno strumento adatto a suddividere i Coleotteri della fauna d’Italia in un totale di 148 famiglie, permettendo talvolta di discriminare tra generi o specie diverse, soprattutto nel caso di famiglie per le quali la fauna italiana risulta composta da poche specie. Tutte le informazioni e i dati che sono qui riportati (caratteri anatomici, note biologiche ed ecologiche, numero di specie ...) fanno riferimento esclusivamente alle specie di Coleotteri presenti in Italia, con aggiornamento al mese di novembre 2021. La struttura di questa chiave dicotomica si basa sullo studio e la rielaborazione, con adattamento per le specie della fauna italiana, di numerosi lavori scientifici e chiavi dicotomiche che vengono riportati in bibliografia, oltre all’analisi diretta dei caratteri diagnostici, effettuata sul materiale della collezione entomologica del Reparto Carabinieri Biodiversità di Belluno e delle collezioni private degli autori. The need to study the Beetle fauna of the 10 State Natural Reserves managed by the Reparto Carabinieri Biodiversità of Belluno, in order to increase their knowledge, has highlighted the need to have a "smart" tool for determining the numerous families of Beetles of our fauna. From this need, the initial idea of developing a dichotomous key, rich in images and detailed schemes, was born. The key was then enriched with anatomical tables and a glossary. To complete the project, it seemed useful to add a catalogue of all 13047 species of Beetles reported to date for our Country, as well as a brief description of the examined families, with bio-ecological notes and a list of some bibliographic tools to be used for the determination of the specimens under study at the specific level. This work mainly represents a suitable tool for dividing the Beetles of the Italian fauna into a total of 148 families, sometimes allowing to distinguish different genera or species, especially in the case of families composed of a few italian species. Information and data here given (anatomical characters, biological and ecological notes, number of species ...) refer exclusively to the species of Coleoptera recorded for Italy, updated on November 2021. The structure of this dichotomous key is based on the study and re-elaboration of numerous scientific works and dichotomous keys, that are listed in bibliography, with adaptation for the species of the Italian fauna, in addition to the direct analysis of the diagnostic characters of the specimens of the entomological collection of the Reparto Carabinieri Biodiversità of Belluno and of the authors' private collections.
... All that is currently known is scattered in some old taxonomic works, regional species list, limited revisions, species descriptions and summaries of the results of individual field trips (e. g. Jarrige, 1952Jarrige, , 1971Coiffait, 1960Coiffait, , 1971Coiffait, , 1981Coiffait, , 1984Assing, 2008;Anlaş & Rose, 2009;Anlaş, 2017). As a consequence, the status of numerous species and subgeneric names, particularly those from the Mediterranen, is doubtful. ...
... The map (Fig. 82) was made using the software Google Earth Pro (2020). Primary and secondary sexual characters of the species are described following the terminology of Coiffait (1984). ...
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Based on the study of types and additional material, new taxonomic and faunistic data on some species of the genus Astenus from the Mediterranean and adjacent regions are presented. Astenus henrii sp. n. from Turkey is described, illustrated, and distinguished from related congeners. The following new synonyms are proposed: Astenus procerus (Gravenhorst, 1806) = A. rufopacus Reitter, 1909 syn. n. = A. circumflexus circumflexus Jarrige, 1952 syn. n. = A. circumflexus scutellaris Coiffait, 1970 syn. n. = A. obliquus Jarrige, 1952 syn. n. = A. baali Coiffait, 1960 syn. n.; A. melanurus (Küster, 1853) = A. adonis Coiffait, 1960 syn. n.; A. thoracicus (Baudi di Selve, 1857) =A. jordanicus Coiffait, 1981 syn. n.
... Várias espécies têm sido referenciadas para a Península Ibérica, mas postas em dúvida como existentes na região, como Edaphus beszedesi Reitter, 1914 e Euaesthetus ruficapillus Lacordaire, 1835 (Outerelo, 1976), espécie mal identificada na altura e depois descrita como uma nova espécie-E. brevelytratus Outerelo & Gamarra, 1986-e anos depois considerada por Puthz (1994) como variedade de E. hispanicus Coiffait, 1984. Cremos contudo que Outerelo & Gamarra (1986) Este género pode distinguir-se bem de Euaesthetus por possuir os olhos mais pequenos e menos salientes. ...
... Género Octavius Fauvel, 1873 Este género foi descrito por Fauvel (1873) e as suas espécies são consideradas como relíquias (Coiffait, 1972(Coiffait, , 1984 de acordo com as suas características morfológicas. Compreende mais de 250 espécies espalhadas pelas regiões Paleártica, Etiópica, Madagascariense, Oriental, Oceânica e Neotropical (Herman, 2001) às quais se juntam duas espécies fósseis. ...
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Resumo: Estuda-se neste artigo a fauna portuguesa da subfamília Euaesthetinae Thomson, 1859 (Coleoptera, Staphylinidae), apresentando-se chaves para identificação dos géneros existentes e de possível ocorrência, bem como registos de duas novas espécies para Portugal. Abstract: Contributions to the knowledge of the Portuguese fauna of Staphylinidae (Coleoptera). II-The Portuguese Euaesthetinae. An overall view of the Portuguese fauna of the subfamily Euaesthetinae Thomson, 1859 (Coleoptera, Staphylinidae) is provided, including identification keys to the existing genera and of likely occurrence, as well as records of two species new for Portugal. Introdução A subfamília Euaesthetinae Thomson, 1859 pertence à família Staphylinidae Latreille, 1802 e é representada por espécies de pequeno porte (1,0-2,0 mm) com cabeça saliente e uma ligeira constrição na parte posterior dos olhos, salientes e com o pescoço largo. Antenas inseridas a pequena distância e em frente dos olhos por cima da base das mandíbulas, formadas por 11 artículos (antenómeros). Lábio curto e mandíbulas compridas e curvas. Palpos maxilares de quatro artículos com o primeiro mais comprido ou tão comprido como o segundo e o terceiro juntos. O quarto é pequeno e afilado na ponta (subulado). Os palpos labiais, formados por três segmentos, em que o primeiro é curto, o segundo oval e o terceiro subulado. Abdómen com um sulco na parte inferior junto à base e os lados com paratergitos em que o primeiro é mais comprido que os restantes. Tíbias ciliadas e tarsos da forma 4-4-4 ou 5-5-5 ou ainda 5-5-4 em alguns géneros exóticos. Grupo com distribuição global e em que o conhecimento ecológico das espécies é muito pobre.
... In the Palaearctic region, the genus Tetartopeus Czwalina, 1888 is represented by 28 species, three of which occur in Turkey: T. adanensis Assing, 2004 from Adana and Osmaniye provinces, T. czwalinai (Jakobson, 1909) from Izmir province, and T. stylifer (Reitter, 1909) from Erzurum province (Coiffait 1982;Smetana 2004;Assing 2004Assing , 2007a. Presence of two additional species in Turkey requires confirmation. ...
... Presence of two additional species in Turkey requires confirmation. Coiffait (1982) reported T. rufonitidus (Reitter, 1909) (as T. confusus Coiffait, 1972) from Asia Minor, but without specification of locality. Also, T. scutellaris (Nordmann, 1837) has been recorded from Adana province by E. Lokay (Horion 1965). ...
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Tetartopeus tezcani sp. n. from Anatolia, Turkey, is described, illustrated and distinguished from related congeners. Tetartopeus persicus Coiffait is reported from Turkey for the first time. A key to the Turkish species of Tetartopeus is presented.
... This beetle has been observed partially buried in the surface sand layer beneath wrack, exposing only its extended mandibles to catch passing preys [58]. Remus sericeus, another staphylinid with similar habits but more sensitive to environmental alterations, is smaller and preys on dipteran larvae and collembolans [50,59] (p. 334). ...
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Anthropogenic development has historically concentrated in coastal areas to exploit resources from fishing and commercial navigation. In recent centuries, intensive tourism has added pressure on sandy shorelines, leading to their modification. This development model has led to the disappearance of most coastal sand dunes and their rich biodiversity, which includes specialized plant and animal species adapted to sandy substrates, harsh arid conditions, and variable levels of salinity. The European Community’s conservation policies, particularly the Habitats Directive (Council Directive 92/43/EEC), have facilitated the preservation and restoration of the few remaining dune systems. However, these policies have unfortunately overlooked the protection of the adjacent beaches, which are integral to the coastal ecosystem. The loss of biodiversity typical of the beach–dune ecosystems is examined in relation to the anthropogenic disturbance factors, with particular attention to mechanical beach cleaning. Indeed, the metabolizable energy generated by this decomposer biomass is crucial for supporting a diverse trophic network of predators, ranging from insects to birds. The rapid disappearance of the specialized beetle fauna is examined, and some essential criteria for defining standard biotic indices suitable for monitoring these ecosystems are suggested. This approach aims to support more effective conservation programs for these fragile environments. We recommend revising the regulatory framework for safeguarding beach–dune ecosystems, while also proposing some key management principles to be incorporated into the protection guidelines.
... The collected samples were brought to the Biodiversity Laboratory in Department of Zoology, Government College University, Faisalabad. The samples were sorted through visual observation and then identified under microscope (M33OO-D) in the laboratory with the help of available keys (Scheerpeltz, 1960;Abdullah & Qadri, 1970;Coiffait, 1982Coiffait, , 1984Lobl, 1986;Pace, 1986;Herman, 2001;Smetana, 2004), web sites and entomological articles. ...
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Field experiment was conducted to collect/sample staphylinid beetles with four different traps (flight intercept trap, Berlese funnel trap, light trap and pit fall trap) and net/hand collection from eleven selected locations of Punjab (Pakistan) in 2013 and 2014. Each locality was sampled for 4 days with an interval of two months. Different abiotic factors were noted and Shannon diversity index was calculated for each locality. A total of 4386 specimens (beetles) were collected. Pitfall traps were found most conducive and effective in sampling beetles followed by Berlese funnel traps and net/hand collection while light traps showed least efficiency. Maximum value of species richness and abundance was observed during Monsoon season (July-August). Paederus fuscipes was the most common species. The highest value of α-diversity index was observed from Sargodha during both years while in case of Shannon-Wiener index value, Murid Wala was the highest during 2013 and Gutwala during 2014. Changa Manga was the place with highest evenness value. The results of Generalized Linear Mixed Model (GLMM) also indicated that the abundance/number of beetles sampled with different collection methods had significant effects with locality and crop type while insignificant effects with time (years). We conclude that methods of trapping need refinement by installing traps for large duration in all study location keeping all conditions (biotic & abiotic) in view to enhance the efficiency of collection methods and exploration of staphylinid beetles.
... Pensant avoir des immatures ou des individus aberrants et devant la difficulté qu'aurait pu procurer l'identification de ces insectes, les choses en étaient restées là. Devant Coiffait [1984], il s'avère rapidement que l'insecte étudié n'est pas présent dans cette faune. L'utilisation de la clé d 'Assing, [2012] ainsi que la publication de Gay [2023] permettent sans aucun doute d'affirmer qu'il s'agit alors de la deuxième commune connue actuellement en France où R. ceylanensis est présent. ...
Article
Authors list a new locality of Rugilus ceylanensis (Kraatz, 1859) for France in Eure-et-Loir and its maintaining in Haute-Savoie. Illustrations are provided with the aim from separating this Rugilus from other french species.
... Google Earth Pro was used to create the map. Primary and secondary sexual characters of the species are described following the terminology of Assing (2004Assing ( , 2006 and Coiffait (1982). The material referred to in this study is stored in the following collections: ...
Article
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In the study, Medon turcicus sp. nov. from Yıldız Mountain in Sivas province, central northern Anatolia, Turkey, is described, illustrated, and separated from related congeners. The distribution of the new species is mapped. Additional Turkish records of seven Medon species, all earlier known from Turkey, are reported. Accounting for a new species, a total of 23 species of Medon are now known from Turkey.
... The nomenclature and classification suggested by was followed in this study. Morphological studies were conducted by using a Wild M5 Stereomicroscope and the keys for identification by Coiffait (1982). All specimens are deposited in the author's collection (RNFC). ...
... Pero no siempre, como por ejemplo la primera cita para Cataluña de Lachnaia (Lachnaia) paradoxa (Olivier, 1808) , especie perteneciente a los Chrysomelidae, familia de la que existe un catálogo en la región catalana (Petitpierre, 2009). (Pandellé, 1869;Portevin, 1929;Coiffait, 1974Coiffait, , 1978Coiffait, , 1984Outerelo y Gamarra, 1985;Dauphin, 1991Dauphin, , 1994Drugmand, 1994;Cuccodoro y Löbl, 1997;Li et al., 2000;Bordoni, 2004), Lucanidae, Trogidae, Geotrupidae, Scarabaeidae (Baraud, 1992; Martín-Piera y López-Colón, 2000), Buprestidae (Cobos, 1986), Elateridae (Leseigneur, 1972), Lampyridae (Olivier, 1884), Cantharidae (Kiesenwetter, 1852;Wittmer, 1974;Dahlgren, 1975;Svihla, 1997) 1990), Sphindidae (Vogt, 1967), Coccinellidae (Fürsch et al., 1967;Plaza Infante, 1977, 1986a, 1986b, 1987Cardoso Raimundo y Gomes Alves, 1986;Eizaguirre, 2004;Bensusan et al., 2006), Corylophidae (Bowestead, 1999), Mordellidae (Méquignon, 1946;Ermisch, 1965;Plaza Infante y Compte Sart, 1980;Plaza Infante, 1985;Leblanc, 2002), Tenebrionidae (Español, 1961), Oedemeridae (Vázquez, 1993(Vázquez, , 2002, Meloidae (Pardo Alcaide, 1950Alcaide, , 1956Alcaide, , 1958Bologna, 1991 ...
... El resto ha sido determinado por el autor mediante las siguientes fuentes bibliográficas: Carabidae (Jeannel, 1941-42;Sciaky, 1987;Gañán & Novoa, 2005;Ortuño & Toribio, 2005;Coulon et al. 2011), Staphylinidae (Portevin, 1929;Jeannel, 1950;Freude et al., 1974;Coiffait, 1974Coiffait, , 1978Coiffait, , 1984Outerelo & Ga marra, 1985;Dauphin, 1991Dauphin, , 1994Dauphin, , 2005Assing & Wunderle, 1995;Drugmand, 1994;Bordoni, 2004;Assing & Schülke, 2011), Lucanidae, Scarabaeidae (Baraud, 1992), Clambidae (Johnson, 1966;Viñolas & Masó, 2013), Buprestidae (Cobos, 1986), Elateridae (Leseigneur, 1972), Drilidae (Bahillo de la Puebla & López-Colón, 2005), Cantharidae (Kiesenwetter, 1852;Wi ttmer, 1974;Da hlgren, 1975;Svihla, 1997;Diéguez Fernández, 2011), Dermestidae (Lepesme, 1950;Pla ta Negrache, 1972;Herrmann & Baena, 2004), Ptinidae (Bellés, 1978;Español, 1992), Cleridae (Bahillo de la Puebla & López-Colón, 2001), Prionoceridae (Ba hillo de la Puebla & López-Colón, 2003), Melyridae (Pla ta Negrac he & Santiago Hernández, 1990;Constantin & Liberti, 2011), Ka teretidae, Ni tidulidae (Audisio, 1993), Silvanidae (Vogt, 1967a), Phalacridae (Vogt, 1967b;Svec, 1992aSvec, , 1992bSvec & Angelini, 1996), Cryptophagidae (Otero, 2011), Erotylidae (Iablokoff-Khnzorian, 1975, Coccinellidae (Fursch et al., 1967;Plaza Infante, 1977, 1986a, 1986b, 1987Cardoso Raimundo & Gomes Alves, 1986;Eizaguirre, 2004;Bensusan et al., 2006), Corylophidae (Bowestead, 1999); La tridiidae (Johnson, 1975;Rücker, 1992;Bouget & Vincent, 2008), Mycetophagidae (Vogt, 1967c), Ciidae (Strand, 1965), Zopheridae (Dajoz, 1977), Mordellidae (Méquignon, 1946;Ermisch, 1965;Plaza Infante & Compte Sart, 1980;Leblanc, 2002), Tenebrionidae (Portevin, 1934;Español, 1957Español, , 1961Weise, 1974;Viñolas, 1984;Viñolas & Cartagena, 2003, 2005, Oedemeridae (Vázquez, 1993(Vázquez, , 2002, Meloidae (Bologna, 1991), Mycteridae (Vázquez, 1993), Anthicidae (Bonadona, 1991;Uhmann, 1992), Aderidae (Gompel & Barrau, 2002), Scraptiidae (Emery, 1876;Ermi sch, 1969;Levey, 2009), Cera mbycidae (Vives, 2000), Bruc hidae (Yus Ramos, 2007), Chrysomelidae (Doguet, 1994;Peti tpierre, 2000;Warchalowski, 2003;Winkel man & Debreuil, 2008), Apionidae (Ehret, 1990), Dryophthoridae (Alonso-Zarazaga & Sánchez-Ruiz, 2009), Curculionidae (Hoffmann, 1950(Hoffmann, , 1954Dieckmann, 1968) y Rhynchi tidae (Hoff mann, 1958). ...
Article
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Resumen: Se presentan los primeros resultados de un catálogo de los coleópteros de la Sierra de Collserola (Barcelona, España), que incluye 305 especies pertenecientes a 41 familias. Palabras clave: Coleoptera, Sierra de Collserola, Barcelona, España, Catálogo. Abstract: Catalogue of the Coleoptera of Collserola Mountain-chain (Barcelona, NE Spain): first results. The first results of a catalogue of the Coleoptera of Collserola Mountain-chain (Barcelona, NE Spain) is presented, which includes 305 species belonging to 41 families.
... After Scheerpeltz (1962) rejected Gridelli's subgeneric division of Scymbalium, it included no subgenera, whereas Schatzmayria was placed in synonymy of Scymbalium. Finally, Coiffait (1980Coiffait ( , 1982 reinstated Micrillus as a genus distinct from Scymbalium and considered Schatzmayria a synonym of the former. In 2008, Assing began the revision of the Palaearctic and Oriental members of Micrillus, with some attention to the similar-looking Scymbalium, which led to a significant reduction of the number of species in Micrillus because of the high rate of synonymies and to the diagnoses of both genera based on an incomplete sample of described species. ...
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Paederinae, a diverse subfamily of rove beetles (Staphylinidae), is poorly explored with an outdated subtribal and generic classification lacking proper phylogenetic perspective. Therefore, the discovery of two Baltic amber fossil specimens resembling the genera Micrillus and Scymbalium, which are particularly challenging in terms of systematics, called for a thorough analysis to infer their phylogenetic position and their ecological requirements. The fossils were examined with light microscopy supplemented by X-ray micro-computed tomography, and then scored into a Paederinae-specific matrix of 99 morphological characters, along with a broad sample of recent Paederinae and non-paederine outgroups. Morphological phylogenetic analyses were conducted, using Bayesian Inference and Maximum Parsimony. The obtained phylogeny confirmed that the genera Scymbalium and Micrillus form a lineage outside Lathrobiina; therefore, both genera are now classified as Lathrobiini incertae sedis pending a more inclusive phylogenetic work on Paederinae. The analysis firmly placed both fossils in that recent lineage, albeit rendering Micrillus paraphyletic with respect to Scymbalium. Without a more extensive analysis based on the revised world fauna, any systematic changes would be premature. Thus, the fossils are described as Micrillus electrus, sp. nov. and Scymbalium phaethoni, sp. nov. in accordance with the current diagnoses of both genera. Given that the recent species of Micrillus and Scymbalium are predominantly thermophilic and mainly confined to dry open landscapes in Africa, southern Eurasia and Australia, the finding of Baltic amber representatives implies the very diverse landscape and the equable (sub)tropical palaeoclimatic conditions of the Eocene amberiferous ‘forest’, the latter being the subject of continued debates.
... Quedionuchus plagiatus is distributed in forested parts of Europe, non-European Russia, Kazakhstan and North America, avoiding only the forests of the southeastern USA and the southern West Palaearctic (Smetana 1971;Schülke and Smetana 2015). Across its range, the species demonstrates considerable morphological variation in colouration, body proportions, sculpture and the male aedeagus (e.g., Coiffait 1978;Smetana 1967Smetana , 1971), which has led to a number of currently recognized synonyms (Schülke and Smetana 2015). Additionally, two currently valid and likely sympatric East Palaearctic species, Q. planatus Sharp, 1888 and Q. samuraicus (Bernhauer and Schubert 1916), have never been clearly differentiated from Q. plagiatus, and it is not known whether the latter species extends into China, where another species, Q. reitterianus (Bernhauer 1934), has been described. ...
Article
The recognition of Holarctic species, those shared between Nearctic and Palaearctic regions, often implies continuous or recent events of gene flow across the 85-km-wide Bering Strait between Alaska and Russia. During the Pleistocene (2.8–0.012 Mya), the Bering land bridge has provided frequent episodes of continuous, tundra habitat across this barrier, while the taiga forests of the northern hemisphere has been separated for much longer, at least 5.4 Mya. This more ancient divergence has led to allopatric speciation in nearly all forest-specialized organisms, including all tree species, and casts doubt on the taxonomic validity of the few subcortical beetle species that are considered to be Holarctic. Here we test the apparent Holarctic distribution of one such species, the morphologically variable rove beetle Quedionuchus plagiatus. Drawing upon morphological and molecular evidence, including morphometric analysis of male genitalia and phylogenetic and cluster analyses of DNA barcodes, we demonstrate that species-level diversity has been greatly underestimated in this lineage and conclude that none of its members are Holarctic. We propose complete allopatric divergence across Beringia in obligate forest beetles and discuss the role of biological constraints as barriers to Holarctic geneflow. We describe Quedionuchus caucasicus Brunke, sp. nov., Q. deceptor Brunke sp. nov., Quedionuchus gilaensis Brunke sp. nov., and Quedionuchus yunnanensis Brunke sp. nov.; revalidate Quedionuchus glaber (O. Müller) and Quedionuchus longipennis (Mannerheim); and propose the following: Quedius longipennis Mannerheim, 1846 = Quedius rufipennis Mäklin, 1853 syn. nov. (previous synonym of Q. plagiatus Mannerheim); Staphylinus glaber O. Müller, 1776 = Quedius planatus Sharp, 1884 syn. nov.
... Given the large volume and diversity of Staphylininae, it is not surprising that the composition of this subfamily has been a matter of discussion from the early days of rove beetle systematics to the present. Staphylininae has been shifting from only containing the group we know now as a tribe Staphylinini (Lohse, 1964;Coiffait, 1972Coiffait, , 1974Coiffait, , 1978Coiffait, , 1982Coiffait, , 1984Smetana, 1982) to a more inclusive taxon also containing the current tribes Xantholinini, Othiini, Diochini and Platyprosopini (and, later on, Maorothiini, Arrowinini and extinct †Thayeralinini as well) (Newton & Thayer, 1992;Assing, 2000;Solodovnikov & Newton, 2005;Solodovnikov et al., 2013). The unknown sister-group relationships among these tribes gave way to solutions like Moore's (1964) equal subfamily rank for Xantholinini, Diochini, Platyprosopini, Staphylininae and Paederinae. ...
Article
We provide the first multilocus molecular phylogeny of a group corresponding to the former subfamily Staphylininae. Results are corroborated by the morphological, biogeographical and palaeobiological evidence to serve as a baseline for an updated suprageneric classification. The former subfamily Staphylininae is proven to be a lineage sister to the monophyletic Paederinae and reclassified according to a robust phylogeny resolving a number of long-standing controversies. The subfamily Xantholinina (revised status) is reinstated to contain the tribes Xantholinini, Othiini, Maorothiini and Diochini. Subfamily Platyprosopinae (revised status) is reinstated for the tribes Platyprosopinini, Arrowinini and †Thayeralinini. For a highly peculiar genus Coomania Cameron, formerly in Diochini, a new subfamily Coomaniinae subfam.n. is established and the composition of Diochini (revised status) is changed accordingly. The subfamily Staphylininae (revised status) is reduced to contain the former tribe Staphylinini only. Elevating this mega-diverse tribe to the subfamily rank opened up an opportunity for its more fractional classification by raising several subtribes to the tribal level as follows: Acylophorini, Afroquediini, Amblyopinini, Antimerini, †Baltognathini, Cyrtoquediini, Erichsoniini, Hyptiomini, Indoquediini, Quediini and Tanygnathinini (revised status for all). As a result, the most species-rich tribe Staphylinini (revised status) is reduced to the more homogeneous lineage containing the subtribes Algonina, Anisolinina, Philonthina, Philothalpina, Staphylinina and Xanthopygina. Morphological synapomorphies and diagnostic characters supporting all newly defined higher taxa are provided.
... Therefore further transfers from these genera to Pseudolathra are likely. A key to west Palaearctic species was provided by Coiffait (1982), and east Palaearctic and Oriental species were revised and keyed by Assing (2012Assing ( , 2013. ...
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A review of all species of the genus Pseudolathra Casey, 1905 known from Réunion is presented. Five species are recorded and one species, Pseudolathra janaki Kocian & Hlaváč, sp. n., is described. New records for Pseudolathra gomyi (Lecoq, 1986) are given. Four species are illustrated and a key to all species is given.
... The present paper is based mainly on material collected during recent field studies in Turkey, conducted by S. Örgel, S. Yaman, E. A. Yağmur and the author. Primary and secondary sexual characters are termed following Coiffait (1982) and Assing (2001Assing ( , 2007. The morphological studies were conducted using a Stemi 2000-C microscope (Zeiss Germany). ...
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A new species of microphthalmous Lathrobium Gravenhorst from Yozgat province in central northern Anatolia is described and its diagnostic characters are illustrated: Lathrobium newtoni sp. n. Additional records of four microphthalmous Lathrobium species are reported. A key to the microphthalmous Lathrobium species of Turkey is presented. Distributions of microphthalmous Lathrobium species in Anatolia are mapped.
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The behavior of organisms is very difficult to observe and document, especially direct interactions such as predation. As a result, there are few systematic studies of such phenomena on a large scale and the food webs of organisms that are formed by fleeting and opportunistic interactions are largely unknown. Understanding food webs is essential for addressing the Eltonian shortfall in biodiversity knowledge and revealing ecosystem dynamics. The rise of citizen science in recent years offers unique opportunities to study food webs on a global scale, which has been demonstrated for larger animals but not for small ones like insects. Here we explore the potential of iNaturalist as a cost-effective citizen science platform to obtain data about the prey choices of predatory rove beetles (family Staphylinidae) from the subtribe Staphylinina, as an alternative to traditional, labor-intense laboratory studies. We manually mined the dietary evidence of Staphylinina worldwide through over 48,000 observations on iNaturalist and 159 records of predation were found. Our findings show that citizen science data not only supports the published studies, but also provides direct and novel field-based evidence of rove beetle prey specialization with numbers of observations that exceed the amount of previously available data by an order of magnitude. We confirmed that some Staphylinina are generalist predators and discovered that some genera and species exhibit specific prey preferences, as documented by iNaturalist. This approach demonstrates that citizen science platforms offer an innovative, scalable, and cost-effective solution to filling global biodiversity knowledge gaps.
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Two new species of rove beetles belonging to the genus Domene Fauvel, 1873, subgenus Lathromene Koch, 1938 from Portugal are described: Domene (Lathromene) boavidae sp. nov. and Domene (Lathromene) pedroi sp. nov. These subterranean species are diagnosed and compared with closely related species from the Iberian endemic subgenus Lathromene. The external morphology and male genitalia are illustrated and information on the species` ecology is presented. A dichotomous key for identifying species of the Lathromene subgenus, based on male genitalia, is presented. New faunistic and ecological data on Domene (Lathromene) viriatoi Serrano & Boieiro, 2015 are also given.
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An unexpected troglobitic staphylinid is described from a dolomite cave in western China as Domene lizeyui Wang & He, sp. nov. (Coleoptera, Staphylinidae, Paederinae). The habitus of both sexes and important diagnostic features are illustrated. Brief notes on the habitat, biology and taxonomic status of the new species are provided. This is the first discovery of a troglobitic representative of Paederinae from China, the first record of a troglobitic Domene species, and only the third cavernicolous species of Paederinae from eastern Asia.
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A key to the 12 Afrotropical genera of Cryptobiina is presented. The type species. the diagnosis. phylogenetical and morphometrical definiions, the general distribulion and the catalogus of the actually known species is given for each genus. The new genus Longiscapus nov. and its type species, L. parallelopipedus (BERNHAUER), are describe. Allocotobium SCHEERPELTZ, subgenus of the invalid genus Cryptobium MANNERHEIM, was found to be a synonym of the genus Tracypum FAGEL. Résumé Une clé des 12 genres afrotropicaux composant la sous-tribu des Cryptobiina est présentée. Pour chaque genre, l'espèce-type, la diagnose, les définitions phylogénétique et morphométrique, la répartition générale et le catalogue des espèces sont donnés. Le genre Longiscapus nov. et son espièce-type, L. parallelopipedus (BERNHAUER), sont décrits. Le sous-genre Allocotobium SCHEERPELTZ est mis en synonymie avec le genre Tracypum FAGEL.
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Les Afrobium F AGEL de Madagascar: systématique; éléménts de biogéographie et de phylogénie sur la sous-tribu (Coleoptera Staphylinidae Paederinae Cryptobiina) par Didier DRUGMAND Résumé Les Afrobium FAGEL, 1977 de Madagascar sont revus. Une espèce, A. fairmairei (FAUVEL) est redécrite; un lectotype ainsi qu'un paralectotype sont désignés. Une seconde espèce, A. julieae sp.n., est décrite. Une clé pour ces deux espèces est donnée; leur édéage, les caractères sexuels secondaires du mâle et l'urite sexuel de la femelle sont figurés. Quelques considérations sur la phylogénie et la biogéographie de la so us-tribu à Madagascar sont données. Abstract Malagasy Afrobium FAGEL, 1977 are reviewed. One species, A. fairmairei (FAUVEL), is redescribed; a lectotype and a paralec-totype are designated. Another species, A. julieae sp.n. is described as new. These two species occur in the northern and northeastern parts of Madagascar. A key to the species is given; aedeagus, secondary sexual characters of the male and the genüal urite of the female are figured. Sorne cornments on the biogeography and the phylogeny of the subtribe from Madagascar are given.
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In this study, between 2009 and 2021, totally more than thausand specimens belonging to the genus Paederus Fabricius, 1775, Paederidus Mulsant & Rey, 1878 and Uncopaederus Korge, 1969 were collected in Türkiye. As a result of field studies, a total of eight species belonging to three genera were recorded. These are Paederidus rubrothoracicus (Goeze, 1777); P. ruficollis (Fabricius, 1777); Paederus balcanicus Koch, 1938, P. fuscipes Curtis, 1826, P. riparius Linnaeus, 1758, P. littoralis Gravenhorst, 1802, P. mesopotamicus Eppelsheim, 1889 and Uncopaederus signiventris (Smetana, 1962). Among these species, P. rubrothoracicus and P. ruficollis from the Marmara and Central Black Sea Regions, P. fuscipes from the Western Black Sea Region, and P. riparius from the Marmara Region of Turkey were reported for the first time. The presence of P. balcanicus in Türkiye has been confirmed. U. signiventris is endemic and distributed only in the Black Sea Region. In addition, P. balcanicus and U. signiventris are figured for the first time.
Conference Paper
In the paper are presented two new species from the subfam. Paederinae (order Coleoptera, family Staphylinidae) found in the natural habitats of the Republic of Moldova. 1. Medon piceus (Kraatz, 1858), a species with eurytopic distribution, populates forest areas, appearing sporadically. 2. Astenus discopunctatus (Say, 1831), adventive species, detected in the oak forest litter mixed with hornbeam. For each species are described, the collection points, status, morphological features, geographical distribution, accentuation of the bioecological category, the place of maintenance in the collection: Coleoptera, Staphylinidae.
Article
The generic classifications of the paederine subtribes Scopaeina Mulsant and Rey, 1878, and Sphaeronina Casey, 1905, are revised. Sphaeronina, revised status, is resurrected from synonymy. Keys to the included genera of both subtribes are included. Newly discovered characters in both subtribes are discussed and illustrated. The Scopaeina now includes Scopaeus, Hyperscopaeus, Micranops, Orus, and Trisunius. The account for each genus includes its diagnostic characters, a description, summary of the general distribution, and list of the included species and specimens examined. Scopaeus Erichson, 1839, has a revised definition and is now restricted to species that have not only a constricted neck and a trichobothrium adjacent to and at about the middorsal margin of the eye, but also a metathoracic/mesofemoral stridulum comprised of a lateral, metaventral file and mesofemoral plectral ridges, slender, apically acute, metakatepisternal processes, and a middorsally fused median lobe of the aedeagus. The stridulum, redefined herein as a file and plectrum that when rubbed together produce stridulation in insects. The metaventral file and mesofemoral plectral ridges of Scopaeus, is, heretofore, unknown in the Staphylinidae or perhaps, even the Coleoptera. Variations of the stridulum and metakatepisternal processes are illustrated and described for each species group. Five genus-group names in the Western Hemisphere, Scopaeomerus Sharp, 1886, and Euscopaeus Sharp, 1886, are new synonyms of Scopaeus; Scopaeodera Casey, 1886, Scopaeoma Casey, 1905, and Scopaeopsis Casey, 1905, are revised status junior synonyms of Scopaeus. The species in those generic groups are now included in species groups of Scopaeus. Hyperscopaeus Coiffait, 1984, new status, is elevated to genus from subgeneric status in Scopaeus. Trisunius Assing, 2011, new subtribal assignment, is moved from the Medonina to the Scopaeina. Typhloscopaeus Jarrige, 1951, incertae sedis, formerly a subgenus of Scopaeus, is of unknown placement, but the species and generic names are retained in Scopaeus awaiting study of the type. Orus cervicula Casey, 1905, revised combination, is returned to Orus from Scopaeus. Orus femoralis (Sharp, 1887), new combination, is transferred from Scopaeus. There are now three named species of Orus with narrow necks. Scopaeus chiriquensis (Sharp, 1886), S. guatemalensis (Sharp, 1886), S. obscurus (Sharp, 1886), and S. palmatus (Sharp, 1886), new combinations, are transferred to Scopaeus from Scopaeomerus. Medon mexicanus (Bernhauer, 1910), new combination, is transferred to Medon from Scopaeomerus. Scopaeus crassitarsis (Sharp, 1886), S. gracilicornis (Sharp, 1886), S. impar (Bierig, 1935), new combinations, are transferred to Scopaeus from Euscopaeus. The following names are transferred from Scopaeus to Hyperscopaeus as new combinations: Hyperscopaeus admixtus (Fagel, 1973), H. albertvillensis (Fagel, 1973), H. allardianus (Fagel, 1973), H. andrewesi(Cameron, 1931), H. angolanus (Fagel, 1973), H. bamaniaensis (Fagel, 1973), H. borneensis (Cameron, 1941), H. bredoanus (Fagel, 1973), H. calidus (Bernhauer, 1932), H. confusoides (Fagel, 1973), H. confusus (Fagel, 1973), H. consimilis (Fagel, 1973), H. convexiceps (Bernhauer, 1932), H. corpulentus(Fagel, 1973), H. decelleanus (Fagel, 1973), H. dolosus (Fagel, 1973), H. endrodyanus (Fagel, 1973), H. errans (Fagel, 1973), H. erraticus (Fagel, 1973), H. fageli (Levasseur, 1981), H. fallaciosus (Fagel, 1973), H. filicornis (Fagel, 1973), H. flavidulus (Fagel, 1973), H. flavocastaneus (Lea, 1923), H. fluviatilis (Fagel, 1973), H. fossiceps (Eppelsheim, 1885), H. fuliginosus (Fagel, 1973), H. fulvescens (Motschulsky, 1858), H. fusculus (Motschulsky, 1858), H. gigantulus (Bernhauer, 1929), H. girardianus (Fagel, 1973), H. hova (Fauvel, 1905), H. hulstaertianus (Fagel, 1973), H. intermixtus (Fagel, 1973), H. kaszabianus (Fagel, 1973), H. katanganus (Fagel, 1973), H. kivuanus (Fagel, 1973), H. lamtoensis (Fagel, 1973), H. leleupianus (Fagel, 1973), H. leopoldvillensis (Fagel, 1973), H. lescuyeri (Delaunay, Coache, and Rainon, 2019), H. levasseuri (Lundgren, 1982), H. longiusculus (Fagel, 1973), H. machadoanus(Fagel, 1973), H. major (Eppelsheim, 1885), H. methneri (Bernhauer, 1932), H. minutulus (Fagel, 1973), H. mulongoensis (Fagel, 1973), H. nitidiceps (Fagel, 1973), H. nitidicollis (Fagel, 1973), H. opacicollis (Bernhauer, 1942), H. overlaetianus (Fagel, 1973), H. parvicornis (Fauvel, 1900), H. procerus (Kraatz, 1859), H. pruinosulus (Eppelsheim, 1885), H. pseudomethneri (Fagel, 1973), H. puberulus (Kraatz, 1859), H. reduncus (Fagel, 1973), H. ripicola (Fagel, 1973), H. rubricollis (Fagel, 1973), H. rubrotestaceus (Kraatz, 1859), H. ruguliceps (Fagel, 1973), H. ruziziensis (Fagel, 1973), H. semifuscus (Kraatz, 1859), H. senegalensis (Fagel, 1973), H. seydeli (Cameron, 1952), H. simillimus (Fagel, 1973), H. simulator (Fagel, 1973), H. spathiferus (Coiffait, 1970), H. spinosophallatus (Frisch, 2012), H. subconfusus (Fagel, 1973), H. subprocerus (Coiffait, 1978), H. surdus (Fagel, 1973), H. suspectus (Fauvel, 1907), H. tchapembanus (Fagel, 1973), H. thoracicus (Motschulsky, 1858), H. tristis (Bernhauer, 1929), H. vagans (Fagel, 1973, and H. voltae (Fagel, 1973). Sphaeronina, revised status, is resurrected from synonymy and now includes Sphaeronum Sharp, 1876, Tripectenopus Lea, 1918, Typhloleleupius Fagel, 1964, and Coecoscopaeus Coiffait, 1982; the last three genera are new assignments to the subtribe. Sphaeronina is redefined by the presence of a hypopharyngeal peg, an enlarged protibial concavity with combs, a ventral denticle on the left mandible, and a groove on the outer edge of the mandibles; additional possible diagnostic characters are discussed. Sphaeronum, Tripectenopus, Typhloleleupius, and Coecoscopaeus are redescribed; the genera are found, respectively, in the American tropical and subtropical regions, Australia, southern Africa and perhaps Madagascar, and Tunisia. Few African and Australian were available for study. Scopaeodracus Scheerpeltz, 1935, is a new synonym of Tripectenopus. Tripectenopus handschini (Scheerpeltz, 1935), new combination, is transferred from Scopaeodracus; Tripectenopus australiae (Fauvel, 1878), T. microps (Lea, 1923), T. pectinatrix (Lea, 1923), and T. torrensensis (Blackburn, 1891), new combinations, are transferred from Domene.
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